taxonID	type	description	language	source
9A3E87FEEB62FFDCD2F7FF2EFA5CFC56.taxon	materials_examined	Type species: Chaetozone setosa Malmgren, by monotypy.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB62FFDCD2F7FF2EFA5CFC56.taxon	diagnosis	Diagnosis. Prostomium blunt to conical, peristomium elongate to short, usually lacking eye spots, with a pair of small nuchal slits or depressions at posterior edge; with a single pair of grooved dorsal tentacles arising from posterior edge of peristomium, or sometimes more posterior on an achaetous anterior segment, or rarely an anterior setiger. First pair of branchiae arising from an achaetous segment or first setiger; or sometimes with first two pairs of branchiae on a single anterior segment. Body basically thick and fusiform over many segments, rarely with middle or posterior body segments beaded or moniliform. Setae include capillaries on most setigers and acicular spines in neuropodia and / or notopodia, spines typically concentrated in posterior segments, forming distinct cinctures with spines emerging from elevated membranes; cinctures with few to many spines and with none to many alternating capillaries; some species with posterior noto- and neuropodial sigmoid acicular spines numerous, encircling entire posterior parapodia; bidentate spines sometimes present in juveniles or occasionally in ventral most position of far posterior setigers of adults accompanying unidentate spines in cinctures; some species with long, natatory-like capillaries, sometimes limited to gravid individuals. Pygidium a simple lobe, disk like, or with long, terminal cirrus.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB62FFDCD2F7FF2EFA5CFC56.taxon	discussion	Remarks. With an increasing number of species of Chaetozone being described, further details of the peristomium, anterior segmentation, position of the dorsal tentacles and branchiae, nature of the posterior cinctures and associated noto- and neuropodial acicular spines, and pygidium are changing the manner in which species of this genus are viewed. Distinct groups of species that share contrasting suites of characters are now evident. The generic diagnosis presented here is slightly altered from that of Blake (1996, 2006) but does not entirely reflect the diverse morphology now evident in this genus. Some aspects of this morphology are reviewed in the discussion at the end of this paper. The following description of the lectotype of C. setosa selected by Petersen (1999) is intended to better define the type-species of the genus to allow for comparison with the numerous global records referred to the species.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB62FFD9D2F7FB23FC38F9D0.taxon	materials_examined	Material examined. Norwegian Arctic, Spitsbergen, Isfjord, 30 fathoms (55 m), coll. 0 6 June 1864, A. J. Malmgren syntype lot SMNH 1493: Lectotype, female, SMNH 1493 - 03 examined by Dr. Mary E. Petersen and designated in print (Petersen 1999); 38 Paralectotypes from SMNH 1493: 1493 - 01, 1493 - 02, 1493 - 04 ‒ 36; data on these collections provided with illustrations, sketches, and descriptive data from preliminary manuscript by Dr Petersen, dated 0 3 April 2001; eight prepared slides of parapodia and illustrations from paralectotype 1493 - 33 also provided by Dr. Petersen; additional study of prepared slides including preparation of photomicrographs, by J. A. Blake, September 2014.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB62FFD9D2F7FB23FC38F9D0.taxon	description	Description. A moderately sized species, lectotype a complete ovigerous female, 20.2 mm long, 1.7 mm wide for 90 setigerous segments; complete paralectotypes up to 28 mm long, 2 mm wide for 94 setigerous segments (Table 1). Body of most preserved specimens curled into a C-shape, but not strongly coiled. Body thickened in middle, narrowing anteriorly and posteriorly. Anterior setigers short, wide, becoming up to 2 times longer in middle body segments, but always narrower than wide except for some segments on ovigerous specimens. Dorsal groove weakly developed, narrow, often limited to anterior setigers; ventral groove well-developed, visible along most of body, absent in far posterior cinctured segments. Color in alcohol brown or grey; no distinct pigmentation. Prostomium conical, narrow, bluntly pointed anteriorly (Fig. 1 A); eyes absent; nuchal organs narrow diagonal slits, not pigmented. Peristomium with two large, distinct rings best visible laterally, overlain dorsally by swollen peristomial crest with peristomial annulations weakly developed or not apparent on crest (Fig. 1 A), crest overlapping prostomium anteriorly, narrowing posteriorly, extending to near anterior margin of achaetous segment 1. Dorsal tentacles arising from notch at posterior margin of peristomium (Fig. 1 A); first pair of branchiae typically positioned posterior to tentacles on posterior margin of incomplete achaetous segment; second pair of branchiae on setiger 1 (Fig. 1 A). Setiger 1 of approximately same size as preceding achaetous segment and subsequent setigers (Fig. 1 A); parapodial lobes reduced, inconspicuous ridges in anterior and middle setigers; enlarged with elevated ridges in posterior cinctured segments bearing conspicuous armature; posterior segments separated by deeply cut intersegmental furrows and with highly elevated membranous podial lobes from which spines and capillaries emerge, forming full cinctures; notopodial spines directed ventrally, neuropodial spines directed dorsally (Fig. 2 A). Noto- and neurosetae from setiger 1, setae of anterior segments all limbate capillaries, numbering about 7 – 10 per fascicle (Fig. 1 D); long, natatory-like notosetae present from about setiger 18 ‒ 21, continuing posteriorly. Capillaries thin throughout, some with fibrils along edge, but not consistent; natatory-like setae capillaries, very long, flattened in cross section numbering 2 ‒ 5 per notopodium mainly restricted to lower part of setal fascicle (Fig. 1 E ‒ F). Based on data from 16 types in Table 1, with 63 ‒ 93 setigerous segments (mean = 83.9 ± SD 7.7), acicular spines begin from setiger 35 ‒ 65 in neuropodia (mean = 51.7 ± SD 7.6) and setiger 43 ‒ 71 in notopodia (mean = 58.7 ± SD 7.1). Lectotype with neuroacicular spines from setiger 57 and notoacicular spines from setiger 63 (Table 1). Spines numbering 1 ‒ 3 at first, accompanied by narrow limbate capillaries, increasing to 10 ‒ 13 in each ramus in fully developed and complete posterior cinctures, with 20 ‒ 26 spines on a side with alternating capillaries (Fig. 1 F, 2 A); spines sometimes overlapping at dorsal midline; when long natatory-like setae occur within posterior cinctures, they accompany ventral-most notopodial spines and sometimes dorsal-most neuropodial spines (Fig. 1 E ‒ F). Spines brownish or brassy in appearance, round in cross section with weak narrow notch at point of emergence, with slightly curved or sigmoidal shape narrowing to a bluntly pointed tip (Figs. 1 B ‒ C, 2 B ‒ D); shafts with thick borders and fine internal striations. Last few cinctured setigers tapering to narrow posterior end; pygidium with terminal anus and small flattened ventral lobe. Methyl Green staining pattern. MG stains the peristomial area with heavy to sparse concentrations of small rows or stripes of stained cells, sometimes very intense; a pattern of stripes on anterior thoracic segments and posterior cinctured segments are most prominent on the ventral surface, but are not intersegmental.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB62FFD9D2F7FB23FC38F9D0.taxon	biology_ecology	Biology. The syntype collection comes from coastal fjords and shelf depths around Spitsbergen of 20 ‒ 60 fathoms (36.5 ‒ 110 m), in clayey or muddy sediments. Numerous specimens were observed with oocytes in various stage of maturity (Table 1); largest ova observed were 120 ‒ 125 µm in diameter (SMNH 1493 - 09). Some males confirmed with sperm platelets (Table 1).	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB62FFD9D2F7FB23FC38F9D0.taxon	discussion	Remarks. Chaetozone setosa is readily distinguished from most related species by having a separate achaetous segment anterior to setiger 1 that bears the first pair of branchiae in combination with an enlarged dorsal swelling or crest on the peristomium. In addition, acicular spines of C. setosa begin in the posterior third of the body and have fully developed cinctures in the far posterior parapodia with 22 ‒ 24 spines on each side. Further, C. setosa has a MG staining pattern that includes stain on the peristomium and ventral segmental stripes in anterior and far posterior parapodia. Other species of Chaetozone described with an inflated lobe or crest overlying the peristomium as in C. setosa include: C. corona Berkeley & Berkeley, 1941, C. pugettensis Blake n. sp., C. platycera Hutchings & Murray, 1984, and C. carpenteri McIntosh, 1911.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB62FFD9D2F7FB23FC38F9D0.taxon	distribution	Distribution. Pending further study of historical records, C. setosa is here limited to Arctic and subarctic areas around Spitsbergen and other areas of northern Europe in shelf depths of 30 ‒ 110 m. Dr. Petersen’s notes indicate that the species was also found offshore East Greenland in muddy sediments.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB67FFD4D2F7F8A1FCD4FBE7.taxon	materials_examined	Material examined. Canadian Arctic, offshore Baffin Island, East Angiak Island, R / V Hero Sta. 20, 132 – 245 m, 65 ° 43 ʹN, 62 ° 05 ʹW, coll. J. A. Blake, 14 Aug 1968, holotype (USNM 51221), 45 paratypes (USNM 1263246). — Off Labrador, Hebron Fjord, Blue Dolphin Sta. 27, 174 m, 58 ° 11.4 ʹN, 62 ° 34.2 ʹW, coll. D. C. Nutt, 8 Aug 1949, Otter trawl, mud bottom, 16 specimens (USNM 22815); Sta. 28, 229 m, 58 ° 09 ʹN, 62 ° 45.7 ʹW, coll. D. C. Nutt, 8 Aug 1949, Otter trawl, 30 + specimens (USNM 22816). — Alaskan Arctic, Off Point Barrow, 104 m, offshore, coll. G. E. McGinitie, 11 Oct 1949, dredged, rocks, stones, gravel, 1 specimen (USNM 22803); 138 m, offshore, coll. G. E. McGinitie, 11 Oct. 1949, dredged, rocks and stones, 1 specimen (USNM 22804); 128 m, 7 mi offshore, coll. G. E. McGinitie, 0 9 Aug 1949, dredged, stones and gravel, 3 specimens (USNM 22805); 138 m, 8 mi offshore, coll. G. E. McGinitie, 11 Oct 1949, dredged, rocks, stones, gravel, 4 specimens (USNM 22806); 66 m, 7.5 mi offshore, coll. G. E. McGinitie, 6 September 1949, dredged, removed from rock and growth on rock, 4 specimens (USNM 22807); 226 m, 12.1 mi offshore, coll. G. E. McGinitie, 17 Aug 1950, dredged, mass of worm tubes, 1 specimen (USNM 22808); 38 m, offshore, coll. G. E. McGinitie, 0 9 Sep 1948, dredged, 1 specimen (USNM 22809); 40 m, offshore, 4 miles out, coll. G. E. McGinitie, 0 9 Aug 1949, stones, gravel, from surface of tunicates, 1 specimen (USNM 22811); 36.5 m, offshore, coll. G. E. McGinitie, 15 Sep 1948, 2 specimens (USNM 22813).	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB67FFD4D2F7F8A1FCD4FBE7.taxon	description	Description. A moderately sized species, holotype complete, 8.2 mm long, 0.8 mm wide across setigers 15 – 30, for 80 setigerous segments. Some complete paratypes up to 14 mm long, 1.2 mm wide for about 100 setigers; largest specimens from Baffin Island collection. Body widest in anterior 15 – 30 setigers, narrowing posteriorly; all anterior thoracic segments short, crowded, at least 15 – 20 times wider than long; middle body segments of some specimens larger, inflated on some specimens; posterior segments narrow, about 4 – 5 times wider than long. A weakly developed narrow dorsal groove runs along body from about setiger 30 (Fig. 4 A); a prominent ventral ridge formed of ventromedial bulges arising from each segment runs along entire length of body (Figs. 3 C, 4 C, E); this feature present in all specimens examined. Color in alcohol light brown with body segments covered with numerous brown to black pigment speckles imparting distinctive background coloration to most specimens (Figs. 3 A – B; 4 A – C, F); in some paratypes this pigment becomes prominent on prostomium and peristomium, forming bands across dorsum of some anterior segments, and often concentrated between segments; some specimens very darkly pigmented. Prostomium triangular, narrowing anteriorly to rounded tip (Fig. 3 A); without eyes, with nuchal organ narrow slit on posterior margin of prostomium, sometimes pigmented resembling eyes (Fig. 3 B); peristomium with one large and one narrow achaetous ring, followed by an achaetous segment between peristomium and setiger 1; narrow peristomial ring incomplete dorsally, overlain medially by posterior extension of large ring; tentacles arising from narrow peristomial ring; achaetous segment similar in form to setiger 1, bearing first pair of branchiae on posterior margin; second pair of branchiae on setiger 1, dorsal to notosetae, branchiae continuing on subsequent setigers (Fig. 3 A). Anterior setae all capillaries arranged in single rows in both noto- and neuropodia; notosetae numbering 6 – 9 per row, neurosetae numbering 7 – 10 per row; about half of specimens with additional long, natatory notosetae along most of body, these specimens sexually mature with many specimens having coelom full of eggs (Fig. 4 B, D). Notopodial acicular spines from setigers 30 – 50 in all specimens; neuropodial acicular spines from anterior third of body or setigers 12 – 26 in Baffin Island specimens having 50 – 85 total setigers and setigers 5 – 15 in specimens from Point Barrow and Labrador having total 29 – 65 setigers; neuroacicular spines 1 – 2 per neuropodium at first, increasing to 6 – 7 near posterior end of body; each neuropodial acicular spine with curved blunt-tips or weakly pointed (Figs. 3 E, 4 G), these alternating with capillaries; notoacicular spines 2 – 5 per segment, narrower and more pointed than neuroacicular spines (Fig. 1 F), also with additional capillaries; noto- and neuroacicular spines and capillaries of posterior segments hooked, forming weakly developed cinctures with low membranes, leaving broad dorsal gap between opposite parapodia (Fig. 3 C). Pygidium simple, with terminal anus and single ventral lobe (Fig. 3 D). Variability. Some specimens from Point Barrow and Labrador with posterior part of first peristomial ring enlarged, bulbous, extending dorsally over two rings bearing the tentacles and first pair of branchiae; this variation is believed due to contraction during preservation. All specimens with distinct ventral line of ridges and reduced development of posterior segments; posterior segments bearing noto- and neuropodial acicular spines never developed into full cinctures as in related species, with only low parapodial membranes or none evident. Neuropodial acicular spines begin earlier in the Point Barrow and Labrador specimens, on setigers 5 – 15 vs. setigers 12 – 26 in the Baffin Island specimens. This difference appears to be size related, with larger Baffin Island specimens having spines beginning later, suggesting a replacement of spines by capillaries with growth. However, all specimens have acicular spines first appearing in the anterior third of the body. Methyl Green staining pattern. No pattern.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB67FFD4D2F7F8A1FCD4FBE7.taxon	discussion	Remarks. Based on available collections, Chaetozone pigmentata n. sp. is the only species found to range across the entire North American Arctic. While there is some variability, all of these specimens share several unique features not found in other species encountered as part of this study. These features include a heavily pigmented body, a prominent mid-ventral ridge line that extends along the entire body, a very weak mid-dorsal groove in middle body segments, the first occurrence of the neuropodial acicular spines in the anterior one-third of the body (setigers 5 – 26) with the smallest specimens having spines in anteriormost locations and the overall distribution being size dependent, and most importantly a reduced posterior armature of spines where the parapodia have only weakly developed cinctures and parapodial membranes. C. pigmentata n. sp. bears some resemblance to C. brunnea Blake, 2009 from deep-sea sediments off California in the nature of the peristomium and presence of body pigment. However, C. brunnea has an unusual body shape denoted by an enlarged and often dark mid-body stomach; in addition, C. brunnea has an achaetous segment bearing two pairs of branchiae instead of one pair and the posterior spines are more numerous and formed into well-developed parapodial cinctures more typical of most Chaetozone species. Some specimens of C. pigmentata n. sp. also have enlarged mid-body segments, but these appear to be associated with reproductive development. C. pigmentata n. sp. is readily distinguished from other Chaetozone species by the heavily pigmented body, weakly developed posterior cinctures with a reduced number of spines, the unusual ventral ridge with mid-ventral segmental bulges, and separate achaetous segments with the first bearing tentacles and the second bearing the first pair of branchiae.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB67FFD4D2F7F8A1FCD4FBE7.taxon	biology_ecology	Biology and ecology. Some Baffin Island specimens have eggs 120 – 125 µm in diameter (Fig. 4 D). Sediments at the collecting locality consisted of well-sorted brown-grey sticky mud with a mean particle size of 150 µm. In addition to C. pigmentata n. sp., 16 other polychaete species were identified (Blake & Dean 1973). Of these, Galathowenia oculata (Zachs, 1923) was the most abundant followed by C. pigmentata n. sp. and Myriochele heeri (Malmgren, 1867). Labrador specimens were found within mud-filled Pectinaria tubes; Point Barrow specimens were dredged from bottoms with stones and gravel.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB67FFD4D2F7F8A1FCD4FBE7.taxon	etymology	Etymology. The species name is based on the pigment spots found on the bodies of all specimens.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB67FFD4D2F7F8A1FCD4FBE7.taxon	distribution	Distribution. Widely distributed across the North American Arctic, Baffin Island, Labrador, and the Beaufort Sea, offshore Point Barrow, in shelf depths of 38 – 245 m.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB6AFFD3D2F7FB17FC42F86C.taxon	materials_examined	Material examined. Canadian subarctic: Western Hudson Strait, CSS Hudson cruise 90 - 023, Sta. 100, collected between 23 September and 16 October 1991, van Veen grab, 63 ° 04.07 ʹN, 74 ° 34.00 ʹW, 393 m, Western Hudson Strait, holotype (LACM-AHF Poly 6535), 19 paratypes (LACM-AHF Poly 6536). Southern Baffin Bay, R / V Hero Sta. 26 A, 1745 m, 67 ° 49 ʹN, 60 ° 46 ʹW, coll. J. A. Blake, 16 Aug 1968, 8 specimens (USNM 51222).	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB6AFFD3D2F7FB17FC42F86C.taxon	description	Description. A moderate-sized species, holotype complete, 11.5 mm long, 0.8 mm wide across thoracic region, with 80 setigerous segments; all paratypes incomplete but of similar size as holotype. Body widest anteriorly, with short, crowded segments (Figs. 5 A – B, 6 A), narrowing posteriorly with segments becoming nearly as long as wide (Fig. 6 D); one of two ovigerous paratypes with abdominal segments nearly moniliform, with body widest anteriorly, narrow crowded segments at least 8 – 10 times wider than long; segments then becoming about as wide as long in middle body with posterior segments narrowing to about 3 – 4 times wider than long. Without any distinct dorsal groove along body; with weak ridge along ventral midline (Figs. 5 B, 6 B). Color in alcohol light brown with anterior segments sometimes with black pigment pattern (Fig. 6 A), faded in other specimens; Baffin Bay specimens more darkly pigmented. Prostomium triangular, narrowing to pointed anterior end (Figs. 5 A – B, 6 A – C), without eyes; nuchal grooves on posterior margin of prostomium, sometimes pigmented in Baffin Bay specimens. Peristomium complex, with three incomplete anterior annulations producing two relatively large achaetous rings and a narrower posterior ring bearing dorsal tentacles (Fig. 5 A – B); peristomium extending posteriorly to setiger 1, overlying a narrow achaetous segment bearing first pair of branchiae (Fig. 5 A); second pair of branchiae on posterior margin of setiger 1 dorsal and medial to notosetae, subsequent setigers with branchiae arranged in same manner. Neuropodial acicular spines first appear in anterior third of body: setigers 20 – 25 on the 80 - setiger holotype and setigers 18 – 20 on an incomplete 50 - setiger paratype; spines single at first, then increasing to 3 – 4 in middle and posterior segments, and up to 10 – 12 per neuropodium in far posterior cinctured setigers. Notopodial acicular spines first present in far posterior setigers, rapidly increasing to 8 – 10 per notopodium in far posterior cinctures; cinctures fully developed, with elevated membranes. Acicular spines numbering 18 – 22 per posterior parapodium, alternating with thin capillaries in dorsal-most and ventral-most positions in notopodia and neuropodia of cinctures, respectively (Fig. 5 C ‒ D). Individual spines somewhat geniculate, curving toward narrow, bluntly pointed tip; internal striae clearly visible; alternating capillaries narrow (Figs. 5 E, 6 F). Two paratypes with long notopodial natatory-like capillaries (Fig. 6 D) and distended abdominal segments with coelom packed with ova; ova measuring up to 156 µm in longest dimension (Fig. 6 G), but average proportions of 137 x 106 µm; other 18 paratypes without natatory-like capillaries or any evidence of gametes. Pygidium with terminal anal opening and with short, protruding dorsal lobe (Figs. 5 C, 6 E). Methyl Green staining pattern. With distinct MG staining pattern on prostomium and peristomium; all of prostomium except tip staining, with most of peristomium staining forming “ mask ” over the head region (Fig. 6 C), with grooves separating peristomial rings either staining poorly or not at all.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB6AFFD3D2F7FB17FC42F86C.taxon	biology_ecology	Biology and ecology. Based on currently available material, C. bathyala n. sp. is limited to deep, cold waters in the Canadian Arctic from 383 to 1745 m. Specimens from shallower depths are referred to other species. The Hudson Strait survey report by McLean et al. (1991) has only limited information on the results of the grab sampling. Sediments are noted to consist of sand and mud, but no actual grain size information is included. The fauna to a depth of 400 m includes several bivalves, polychaetes, and echinoderms, but only bivalves were identified, not specific to any one station. The specimens from southern Baffin Bay, R / V Hero Station 26 were recorded in Blake & Dean (1973). Bottom temperatures were 0.1 ° C and sediment consisted of thick sticky mud. The most abundant polychaetes occurring with C. bathyala n. sp. were Aricidea suecica Eliason, 1920 (Paraonidae), Cryptosclerocheilus baffinensis Blake, 1972 (Scalibregmatidae), and Jasmineira schudiinni Augener, 1912 (Sabellidae).	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB6AFFD3D2F7FB17FC42F86C.taxon	discussion	Remarks. C. bathyala n. sp. is similar to the type species, C. setosa, in the nature of the peristomium, anterior achaetous segments, fully developed cinctures of posterior spines, and in having a MG staining pattern. However, C. setosa has not been observed with body pigmentation and the ventral surface has a deep groove instead of a ridge; further, C. setosa has capillaries alternating with all spines in the cinctures, whereas C. bathyala n. sp. has them limited to the upper and lower parts of the noto- and neuropodia, respectively. In addition, C. setosa has long, natatory-like setae present in all specimens, whereas in C. bathyala n. sp. they are limited to sexually mature specimens. C. bathyala n. sp. closely resembles C. pigmentata n. sp., also described in this paper, in having a narrow peristomial ring bearing the dorsal tentacles, followed by the first pair of branchiae on a subsequent achaetous segment and the second pair of branchiae appearing on setiger 1. Both species bear body pigment, but in C. pigmentata n. sp. it consists of numerous speckles or patches instead of the diffuse dusky pigment of C. bathyala n. sp. Further, C. bathyala n. sp. has full posterior cinctures of spines with up to 18 ‒ 22 spines on a side, whereas C. pigmentata n. sp. has only weak cinctures with no more than 11 spines on a side. C. bathyala n. sp. has a MG staining pattern, whereas C. pigmentata n. sp. has none. The pygidium of C. bathyala n. sp. differs from other species of the genus in having a small dorsal lobe overlying the anal opening instead of the distinct ventral lobe or disk found in most species of Chaetozone. Among specimens examined by the late Dr. Mary E. Petersen from the collections of the Swedish Museum of Natural History (SMNH) was a single incomplete 39 - setiger specimen collected off Newfoundland at coordinates 52 ° 05 ʹN, 52 ° 19 ʹW, in sand at a depth of 294 m (SMNH 1451). Dr. Petersen’s notes indicate that this specimen was a female with brown pigment on the anterior dorsum to about setiger 17, being darkest on setigers 4 ‒ 16. The first neuroacicular spines were on setiger 30 and increased to three or four by setiger 39; notoacicular spines were not yet present and natatory-like capillaries were absent although oocytes were present and measured 85 µm in diameter. These characteristics are mostly consistent with those of C. bathyala n. sp. except for the lack of natatory-like capillaries on a female with eggs.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB6AFFD3D2F7FB17FC42F86C.taxon	etymology	Etymology. The epithet is derived from the Greek, bathys, for deep, denoting the bathyal depths from which this species was collected.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB6AFFD3D2F7FB17FC42F86C.taxon	distribution	Distribution. Canadian Arctic and subarctic, in deep water of the Hudson Strait, 393 m and offshore, Baffin Bay, 1745 m; likely record from offshore Newfoundland in 294 m.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB6EFFCDD2F7FF2EFE7EFDA7.taxon	materials_examined	Material examined. Alaskan Arctic, Beaufort Sea, coll. off Pitt Point, 20 May 1976, 71 ° 19 ʹN, 152 ° 38.5 ʹW, 55 m, 0.1 m 2 Smith-McIntyre grab, Bell 205 helicopter, Sta. PPB- 55, coll. A. G. Carey Jr., holotype (LACM-AHF Poly 6537), 1 paratype (LACM-AHF-Poly 6538); Beaufort Sea, coll. off Narwhal Island, 28 Aug 1976, 70 ° 24.3 ʹN, 147 ° 29.2 ʹW, 10 m, 0.1 m 2 Smith-McIntyre grab, R / V Aluminak Sta. NIB- 15, coll. P. A. Montagna, 10 paratypes (LACM-AHF-Poly 6539). — Canadian subarctic, Hudson Strait, Deception Bay, Quebec, coll. Jésica Goldsmit, Sta. 4 B, 0 2 Aug 2012, 62 ° 13.187 ʹN, 74 ° 52.187 ʹW, intertidal, in sand, 5 specimens (CMNA 2014 - 0015); Sta. 3 D, 0 2 Aug 2012, 62 ° 30.137 ʹN, 74 ° 48.614 ʹW, 6.7 m, in sand and silt, 2 specimens (CMNA 2014 - 0016).	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB6EFFCDD2F7FF2EFE7EFDA7.taxon	description	Description. A moderate-sized species, holotype complete, 11 mm long, 0.6 mm wide across thoracic region, with 118 setigerous segments; most paratypes complete, smaller than holotype; Deception Bay specimens of similar size, up to 11 mm long, 0.8 mm wide, for 90 setigers. Body relatively sleek in appearance, not expanded in anterior region, consistent in width along most of body, narrowing in far posterior setigers. Dorsal surface of body with weak longitudinal groove (Fig. 8 A), venter with distinct groove along entire length of body (Fig. 8 B). Color in alcohol light tan with no body pigment. Prostomium triangular, acutely pointed on anterior margin (Figs. 7 A, 8 A); eyes absent; nuchal organs not pigmented, narrow slits at posterior lateral margin of prostomium; peristomium elongate, with 3 – 4 rings (Fig. 7 A); peristomium extending dorsally over achaetous segment 1; dorsal tentacles arising over segment 2 (setiger 1) (Figs. 7 A, 8 A); first pair of branchiae arising from achaetous segment 1; second pair of branchiae arising from segment 2 (setiger 1) lateral to paired medial dorsal tentacles and dorsal to notosetae; branchiae continuing on subsequent setigers (Fig. 7 A, 8 A). Anterior noto- and neurosetae all simple capillaries arranged in single rows of 5 – 6 notosetae and neurosetae; capillaries increasing to 6 – 8 per noto- and neuropodium in middle body segments. Some middle body parapodia with natatory-like capillary setae. Neuropodial acicular spines first present from setigers 60 – 65, or about posterior one-third of body; notopodial acicular spines from about setiger 90 or near posterior end; spines single at first, then increasing to 5 – 7 in notopodia and 7 – 10 in neuropodia; spines forming distinct cinctures in posterior segments, with moderately developed elevated membranes and up to 17 spines on a side (Figs. 7 C, 8 D). All acicular spines in cinctured segments alternating with long, thin capillaries; each spine thickened basally, curved, and tapered to blunt tip (Figs. 7 D, 8 E). Pygidium with elongate ventral lobe (Figs. 7 B, 8 C). Methyl Green staining pattern. No pattern.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB6EFFCDD2F7FF2EFE7EFDA7.taxon	discussion	Remarks. The characteristic that most defines C. careyi n. sp. is a shift in the position of the paired dorsal tentacles posteriorly over the first setiger and posterior to the first pair of branchiae. In most species of Chaetozone, including the other species described in this paper, the tentacles arise posteriorly on the peristomium but well anterior to the first branchiae and any of the setigerous segments. The situation in C. careyi n. sp. is similar to but not as extreme as that in C. bansei Blake, 1996, where the first tentacles are shifted posteriorly to setigers 4 – 7. C. careyi n. sp. and C. bansei are also similar in the cinctured posterior setigers, shape of the acicular spines, and the nature of the pygidium. However, C. careyi n. sp. further differs from C. bansei in lacking thickened anterior capillaries and a MG staining pattern, which is characteristic of C. bansei. In addition, in C. bansei the neuropodial acicular spines begin on setigers 28 – 29 or the anterior third of the body, whereas in C. careyi n. sp. they begin on setigers 60 – 65 or near the posterior third of the body; notoacicular spines begin about setiger 80 in C. bansei and setiger 90 in C. careyi n. sp. Originally described from shallow shelf depths offshore San Francisco, California, C. bansei ranges north to at least off Oregon (Blake unpublished), whereas C. careyi n. sp. appears to be limited to the North American Arctic and subarctic. In correspondence and among the notes and illustrations left by Dr. Mary E. Petersen, was data on specimens of a species of Chaetozone from the Aleutian Islands, Dutch Harbor, Alaska coll. 30 September 1980, from shallow water grab samples provided by the University of Alaska, Fairbanks. These specimens were attributed to C. bansei by Dr. Petersen and were noted to have the dorsal tentacles shifted posteriorly over an anterior setiger and with the neuroaciculars from the posterior third of the body instead of the anterior third in C. bansei. This information agrees well with the characters of C. careyi n. sp. described here.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB6EFFCDD2F7FF2EFE7EFDA7.taxon	biology_ecology	Biology and Ecology. The data from the Beaufort Sea project is only available in a report to the U. S. government (Carey et al. 1984). From this, C. careyi (as C. setosa) was part of an assemblage dominated by a suite of polychaetes including Capitella sp., Chone cf. murmanica Lukasch, 1910, Prionospio cirrifera Wirén, 1883, and Pholoe minuta (Fabricius, 1780).	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB6EFFCDD2F7FF2EFE7EFDA7.taxon	etymology	Etymology. This species is named for Dr. Andrew (Drew) G. Carey, Oregon State University, Corvallis, Oregon, benthic ecologist and friend. Dr. Carey’s surveys in the Alaskan Arctic and in deep-sea habitats inspired several students, some of whom have developed their own careers in benthic ecology and taxonomy.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB6EFFCDD2F7FF2EFE7EFDA7.taxon	distribution	Distribution. Known from the Beaufort Sea, Alaska, to Hudson Strait, Canada, in shallow depths; Aleutian Islands; intertidal to 55 m.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB73FFCAD2F7FDD0FCCFFEAC.taxon	materials_examined	Material examined. Alaskan Arctic, Beaufort Sea, Prudhoe Bay, 0 5 Aug 1983, 70 ° 23.9 ʹN, 148 ° 26.0 ʹW, 3 m, push cores, coll. M. Busdosh, Waterflood 6 project, R / V Annika Marie, Sta. 15 - 1, holotype (LACM-AHF Poly 6540), 3 paratypes (LACM-AHF Poly 6541).	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB73FFCAD2F7FDD0FCCFFEAC.taxon	description	Description. A moderate-sized species, holotype 12 mm long, 1.0 mm wide with 96 setigerous segments; paratypes similar in size with 90 – 100 setigers. Body expanding slightly in anterior segments, narrowing gradually along body; posterior third of body with full cinctures of spines. Middle body dorsoventrally flattened, with deep ventral groove continuing to near posterior end; mid-dorsal groove present along most of body (Figs. 9 A, 10 A – B), becoming deeper and more distinct in posterior setigers where elevated membranes and cinctures of spines overlap and enhance groove. Color in alcohol light tan with no body pigment. Prostomium narrow, pointed on anterior margin (Fig. 9 A – B), with slit-like nuchal organ on posterolateral margin (Fig. 9 B); eyes absent. Peristomium broad, with 2 – 3 weak annulations, forming one large anterior ring and two narrow posterior rings; all rings weakly defined (Fig. 9 A – B) and variable in size and development between specimens; distinct achaetous segment not apparent; last peristomial ring bearing tentacles and first pair of branchiae, with latter arising lateral to tentacles (Fig. 9 A – B). Second pair of branchiae arising from setiger 1, continuing on subsequent segments; branchiae of anterior third of body long, thick, branchiae of middle and posterior segments, short and thin where present. Anterior noto- and neurosetae all simple capillaries arranged in single rows of 6 – 8 notosetae and neurosetae; capillaries increasing in middle body segments to 15 or more in notopodia and 10 – 12 in neuropodia; some capillaries long, but extremely long natatory-like capillary setae absent in mature individuals. Neuropodial acicular spines from setigers 56 – 57 or posterior third of body, first spines single, in ventral-most position in fascicle, then increasing to 9 – 11 over next 20 – 22 setigers replacing all but dorsal-most capillaries; very thin capillaries appearing in last 15 or so posterior-most segments, alternating with spines. Notopodial acicular spines from setigers 58 – 65, with first spines in dorsal-most location of fascicle, increasing to 9 – 11 posteriorly, but not replacing capillaries as in neuropodia, instead, capillaries retained and alternate with spines but become very thin in far posterior cinctures. Posterior segments with well-developed elevated membranes and 10 – 11 spines in both noto- and neuropodia, forming full cinctures on each segment, with notosetae overlapping at dorsal midline (Fig. 10 B – D). All cinctured segments with spines alternating with long, thin capillaries; each spine thickened basally, curving, tapering to blunt tip (Figs. 9 D – E, 10 D – E); notopodial spines generally longer than neuropodial spines, with neuropodial spines in ventral-most position shorter, curved, weakly geniculate. Pygidium with terminal anus and small, rounded ventral lobe (Figs. 9 C, 10 C). Methyl Green staining pattern. No pattern.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB73FFCAD2F7FDD0FCCFFEAC.taxon	biology_ecology	Biology and ecology. This species occurred in shallow, sandy silt sediments of Prudhoe Bay. Other commonly associated invertebrates included the polychaetes Tharyx alaskensis n. sp., Marenzellaria viridis (Verrill, 1873), Prionospio cirrifera, Ampharete vega (Wirén, 1883), and the amphipod Monoporeia affinis (Lindström, 1855). Coelomic oocytes present in the holotype and paratypes, ca. 60 ‒ 80 µm in longest dimension.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB73FFCAD2F7FDD0FCCFFEAC.taxon	discussion	Remarks. Due to weakly developed peristomial annuli, a distinct achaetous segment between the peristomium and setiger 1 could not be discerned in Chaetozone ruffi n. sp.; therefore, both the paired tentacles and first pair of branchiae are interpreted as arising from the posterior margin of the peristomium. Both the neuropodial and notopodial spines begin in the posterior third of the body rather than more anteriorly as is typical for related species. The posterior cinctures form highly modified posterior segments in this species; the notopodial spines and notopodial membranes overlap mid-dorsally enhancing the dorsal groove. Among species known from the NE Pacific and the North American Arctic, the species closest to C. ruffi n. sp. are C. columbiana Blake 1996 from off Oregon and Washington and C. hobsonae n. sp. from British Columbia and SE Alaska. Each of these species lack a distinct achaetous segment anterior to setiger 1 and the dorsal tentacles and first pair of branchiae arise from the posterior margin of the peristomium. Chaetozone ruffi n. sp. has up to 22 posterior acicular spines on each side; C. columbiana has 11 ‒ 12; and C. hobsonae n. sp. has 17 – 19. Further, both C. columbiana and C. hobsonae have distinctive MG staining patterns while C. ruffi n. sp. has none. In lacking a distinct achaetous segment that can be discerned on the peristomium, C. ruffi n. sp. is also similar to C. hystricosa Doner & Blake, 2006, from off New England; however, C. hystricosa has only 12 – 13 acicular spines on each side in posterior setigers, whereas C. ruffi n. sp. has up to 22 acicular spines.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB73FFCAD2F7FDD0FCCFFEAC.taxon	etymology	Etymology. This species is named for my friend and long-time colleague, Mr. R. Eugene Ruff. Gene provided much of the material and data presented in this paper and contributed valuable information relative to his extensive experience with the benthos of northeastern Pacific and Alaska.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB73FFCAD2F7FDD0FCCFFEAC.taxon	distribution	Distribution. Alaskan Arctic in shallow water, 3 m.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB74FFC8D2F7FED3FB4AF91B.taxon	materials_examined	Material examined. Oregon, Clapsop County, off Tillamook Head, Sta. SMG 1937, Scallops Island, 45 ° 53.6 ʹN, 124 ° 11.9 ʹW, 109 m, 0.1 m 2 Smith-McIntyre grab, coll. 7 Nov 1981, A. G. Carey, Jr. holotype (LACM-AHF Poly 6542), one paratype (LACM-AHF Poly 6543).	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB74FFC8D2F7FED3FB4AF91B.taxon	description	Description. A large species, two complete specimens; holotype largest, 25 mm long, 2.5 mm wide for 106 setigers; paratype 18 mm long, 1 mm wide for 82 setigers. Color in alcohol light tan to brown, lacking any distinctive body pigment. Body long, arched dorsally, with shallow, narrow, mid-dorsal groove along entire length of body (Figs. 11 A, 12 A), most prominent in anterior setigers, but reemphasized in far posterior segments by overlap of setae and membranes of dorsal cinctures; venter flattened, with deep prominent mid-ventral groove (Fig. 12 B) continuing along entire body through cinctured posterior segments (Fig. 11 F). Prostomium long, narrow, pointed on anterior end (Figs. 11 A – B, 12 B, D); eyes absent; nuchal organs elongated slits on posterior margin of prostomium (Fig. 11 B); peristomium with large anterior inflated buccal region followed by two narrow annulations, second annulation interpreted as an achaetous segment bearing a pair of dorsal tentacles and a pair of branchiae positioned laterally (Fig. 11 A – B). Subsequent setigers with branchiae dorsal to notosetae (Figs. 11 A – B, 12 D); anterior branchiae thicker than those of middle segments. Setiger 1 of approximately same size as preceding achaetous segment and following segments; podial lobes reduced to inconspicuous ridges in anterior setigers, becoming inflated and conspicuous in middle setigers, greatly enlarged with elevated ridges in posterior setigers on deeply cinctured segments bearing conspicuous armature (Fig. 12 A – C); posterior segments separated by deeply cut intersegmental furrows and with highly elevated membranous podial lobes from which spines and capillaries emerge, forming full cinctures (Figs. 11 F, 12 A – C); small, ciliated organ present between ventral-most notopodial spine and dorsal-most neuropodial spine (Fig. 11 E); notopodial spines directed ventrally, neuropodial spines directed dorsally. Noto- and neurosetae from setiger 1 all capillaries, numbering about 6 – 7 per fascicle; each seta thickened, with short fibrils usually apparent along broadest edge; these capillaries remaining thick and long until far posterior setigers. Acicular spines first present from setigers 20 – 35 in neuropodia and 40 – 45 in notopodia, numbering 2 – 3 at first, accompanied by an equal number of thickened capillaries; neuropodial spines numbering 8 – 9 per fascicle and notopodial spines numbering 7 – 8 per fascicle in far posterior setigers, spines forming complete setal cinctures with up to 17 spines on a side (Fig. 9 F); spines accompanied by alternating thin capillaries (Fig. 11 F); spines blunttipped, slightly curved, with weak node or notch at point of emergence from podial lobe (Fig. 11 E); spines with thick borders and fine internal striations. Last few cinctured setigers narrowing to pointed posterior end (Fig. 12 A – C); pygidium with terminal anus and small flattened ventral lobe (Figs. 11 C – D, 12 C). Methyl Green staining pattern. Stain imparts distinct pattern on prostomium, peristomium, and anterior parapodia (Fig. 12 D). Anterior tip of prostomium unstained, then with transverse band, then clear again; peristomium lightly stained, but with deeper stain on dorsal surface of last achaetous ring. Anterior parapodia deeply stained laterally, with bands extending across venter and sometimes dorsum.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB74FFC8D2F7FED3FB4AF91B.taxon	discussion	Remarks. Chaetozone malmgreni n. sp. is characterized by having one large and one narrow achaetous peristomial ring followed by another narrow achaetous segment that bears both the tentacles and first pair of branchiae. In addition, this species has deep dorsal and ventral grooves and a distinct MG staining pattern. Other species having the dorsal tentacles and first pair of branchiae on a single achaetous segment include C. lunula Blake, 1996 from northern California, C. corona Berkeley & Berkeley, 1941, from southern California, C. allanotai Blake, 2006 from deep-water offshore California, and C. michellae Magalhães & Bailey-Brock, 2013 from Hawaii. C. lunula, however, has the peristomium shifted posteriorly so that the dorsal tentacles and first pair of branchiae are at the level of setiger 1; further differences with C. malmgreni n. sp. are that the ventral-most neuropodial acicular spine of C. lunula is typically bidentate and the pygidium has a terminal cirrus. C. corona differs from all Chaetozone species in North America in having neuropodial acicular spines beginning on setiger 1. C. allanotai belongs to a group of species having a distinct type of acicular spine where the fine tip curves back and fuses with the shaft. C. michellae has a deep dorsal groove along the entire body, whereas C. malmgreni has both a shallow dorsal groove and a deep ventral groove along the body; the two species have very different MG staining patterns. There is also some similarity between C. malmgreni n. sp. and C. anasima Doner & Blake, 2006 and C. hystricosa Doner & Blake, 2006, both described from New England waters. However, both of these species have indistinct annulations on the peristomium and the exact origin of the tentacles and first pair of branchiae are vague. Etymology. The species is named for Dr. A. J. Malmgren, who described the first species of Chaetozone, C. setosa, in his classic work on polychaetes from Greenland, Spitsbergen, Iceland, and Scandinavia in 1867.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB74FFC8D2F7FED3FB4AF91B.taxon	distribution	Distribution. Known only from continental shelf sediments, offshore Oregon, 109 m.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB76FFC5D2F7F949FC4FFA54.taxon	materials_examined	Material examined. Washington, Puget Sound, San Juan Islands, Decatur Island, Puget Sound Environmental Monitoring Program (PSEMP) Spatial Survey, R / V Kittiwake Sta. 753, coll. 89 Jul 2012, 48 ° 30.1315 ʹN, 122 ° 49.4996 ʹW, grab, 9 m, silt and clay, holotype (LACM-AHF Poly 6544), 17 paratypes (LACM-AHF Poly 6545).	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB76FFC5D2F7F949FC4FFA54.taxon	description	Description. A moderate-sized species, holotype and larger paratypes 12 – 13 mm long, 0.9 – 1.0 mm wide across anterior fourth of body, with 85 – 90 setigers. Color in alcohol light tan to brown, lacking any distinctive body pigment. Body with narrow crowded segments for two-thirds of body, without any prominent swelling or anterior enlargement (Fig. 14 C – D); dorsal groove absent or weakly developed anteriorly (Fig. 13 A), emphasized in far posterior segments by overlap of dorsal cinctures (Figs. 13 A, 14 F); venter flattened, with deep prominent midventral groove continuing along entire body through cinctured posterior segments (Fig. 14 B, D). Prostomium broadly triangular, narrowing anteriorly to bluntly rounded tip (Figs. 13 A, 14 A – B); eyes absent; nuchal organ at posterior lateral margin, consisting of narrow slit, sometimes weakly pigmented (Fig. 13 B). Peristomium with two large rings, both overlain dorsally by an inflated ridge or crest extending posteriorly to level of setiger 1; these peristomial rings followed by narrow achaetous segment having a similar shape as setiger 1. Dorsal tentacles arising from groove at posterior end of second large peristomial ring (Figs. 13 A – B, 14 A); first pair of branchiae arising posterior to tentacles on posterior margin of achaetous segment; achaetous segment approximately same size as subsequent setigers, sometimes with a lateral furrow (Figs. 13 A – B, 14 A); second pair of branchiae arising dorsal to notosetae on first setiger immediately posterior to achaetous segment 1 (Figs. 13 A, 14 A); subsequent setigers with branchiae dorsal to notosetae; branchiae of anterior setigers thicker than those in middle segments. Setiger 1 of approximately same size as preceding achaetous segment and subsequent setigers; podial lobes reduced to inconspicuous ridges in anterior setigers; becoming inflated and conspicuous in middle setigers and greatly enlarged with elevated ridges in posterior setigers, with conspicuous armature; posterior segments separated by deeply cut intersegmental furrows and with highly elevated membranous podial lobes from which spines and capillaries emerge, forming full cinctures (Figs. 13 C, 14 E); notopodial spines directed ventrally, neuropodial spines directed dorsally (Fig. 14 F). Noto- and neurosetae from setiger 1 all capillaries, numbering 8 – 10 per fascicle; each seta thickened, with fibrils usually apparent along broadest edge; these capillaries remaining thick and long until far posterior setigers; long natatory-like capillary setae not observed; acicular spines first present from about setigers 50 – 53 in neuropodia and 60 in notopodia, or the posterior one-third of the body on specimens with 85 ‒ 90 setigers; spines numbering 2 – 3 at first, accompanied by an equal number of thickened capillaries; in far posterior setigers notopodial spines numbering 8 – 9, and neuropodial spines numbering 9 – 10, formed into complete setal cinctures, with spines alternating with thin capillaries (Fig. 13 D – F) with up to 19 spines on a side; spines blunt-tipped, slightly curved, with weak node or notch at point of emergence from podial lobe; with thick borders and fine internal striations (Fig. 14 G – H). Last few cinctured setigers tapering to narrow posterior end; pygidium with terminal anus and small flattened ventral plate or disk (Figs. 13 C, 14 E). Methyl Green staining pattern. No distinct pattern.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB76FFC5D2F7F949FC4FFA54.taxon	discussion	Remarks. Chaetozone pugettensis n. sp. closely resembles the type species, C. setosa in the nature of the two large peristomial rings, overlaid dorsally by an inflated and sometimes bulbous dorsal ridge or crest; the form and placement of the achaetous segment anterior to setiger 1; position of the dorsal tentacles and first pair of branchiae; and the number and structure of the posterior spines. Both species also have the noto- and neuropodial acicular spines from the posterior third of the body and both have weakly developed dorsal grooves and well-developed ventral grooves along the body. C. pugettensis n. sp. differs from C. setosa in that the type species exhibits long natatory-like capillaries on most body segments including among the posterior spines whereas no specimens of C. pugettensis n. sp. in the present collection have been observed with these setae. Another important difference is that C. setosa has a distinct MG staining pattern, whereas C. pugettensis n. sp. has none.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB76FFC5D2F7F949FC4FFA54.taxon	etymology	Etymology. This species is named for its collection in the Puget Sound, offshore Decatur Island.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB76FFC5D2F7F949FC4FFA54.taxon	distribution	Distribution. Puget Sound, in silt and clay, shallow water, 9 m.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB7BFFC3D2F7F98EFC2AFF65.taxon	materials_examined	Material examined. Canada, British Columbia, Prince Rupert, off Ridley Island, coll. September 2012, AECOM, by P. Winchell & S. Doner, Van Veen grab: Sta. B 05, 19 Sep 2012, 54 ° 12.352 ʹN, 130 ° 19.966 ʹW, 17.7 m, holotype (LACM-AHF Poly 6546), 10 paratypes (LACM-AHF Poly 6547); Sta. B 09, 19 Sep 2012, 54 ° 11.698 ʹN, 130 ° 18.848 ʹW, 7.5 m, 10 paratypes (LACM-AHF Poly 6548); Sta. B 23, 20 Sep 2012; 54 ° 11.766 ʹN, 130 ° 18.691 ʹW, 5.9 m, ca. 200 paratypes (LACM-AHF Poly 6549); Sta. B 25, 21 Sep 2012, 54 ° 11.949 ʹN, 130 ° 18.672 ʹW, 20.1 m, 8 paratypes (LACM-AHF Poly 6550); Sta. B 26, 19 Sep 2012, 54 ° 12.070 ʹN, 130 ° 18.801 ʹW, 12.4 m, 7 paratypes (CMNA 2014 - 0 0 17); Sta. B 35, 21 Sep 2012, 54 ° 12.281 ʹN, 130 ° 18.089 ʹW, 26.7 m, 10 paratypes (CMNA 2014 - 0018). Southeastern Alaska, Boca de Quadra, Cruise 3 BQ, R / V Redoubt, Sta. 100 - 2, 55 ° 19.2 ʹN, 130 ° 29.2 ʹW, 95 m, coll. Dec 1979, R. L. Cimberg, Van Veen grab, 6 specimens (LACM-AHF Poly 6551).	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB7BFFC3D2F7F98EFC2AFF65.taxon	description	Description. A moderate-sized species, holotype and larger paratypes 12 – 13 mm long, 0.6 – 0.7 mm wide across anterior setigers and ca. 0.8 mm wide in middle setigers, with about 80 – 85 setigers. Body typically breaks mid-body, hence many specimens incomplete. Color in alcohol light tan to brown, lacking any distinctive body pigment. Body with narrow crowded segments for two-thirds of body; posterior cinctured setigers not as crowded; body without any prominent swelling or anterior enlargement; with narrow, weakly developed mid-dorsal groove from about setiger 6 ‒ 8, continuing posteriorly for about first third of body, not apparent posteriorly; venter flattened, with narrow ventral groove from just posterior to mouth continuing over anterior, middle, and most posterior setigers, absent from far posterior cinctured segments. Prostomium swollen, enlarged, tapering to triangular and pointed anterior end (Fig. 15 A – B); eyes absent; nuchal organ a narrow slit bordered by thickened pigmented cells, superficially appearing to be an eye (Fig. 15 B). Peristomium elongate with two large rings, best seen laterally, separated from prostomium by deep lateral groove (Fig. 15 A – B). Dorsal tentacles arise near posterior margin of peristomium; first pair of branchiae arising posterior to tentacles just anterior to setiger 1; second pair of branchiae on posterior border of setiger 1, dorsal to notosetae, subsequent setigers with branchiae similarly dorsal to notosetae. Setiger 1 merging with peristomium approximately same size as subsequent setigers; podial lobes reduced to inconspicuous ridges in anterior setigers, inflated and conspicuous in middle setigers, greatly enlarged with highly elevated ridges and conspicuous armature in posterior setigers; posterior segments separated by deeply cut intersegmental furrows. Noto- and neurosetae from setiger 1 all capillaries, numbering about 6 – 8 per fascicle; each seta thickened, relatively smooth, with fibrils sometimes apparent along broadest edge; these capillaries remaining thick and long until far posterior setigers; about half of specimens examined with long, thin, natatory-like notosetae beginning from about setigers 15 – 20 and continuing along most of body, most prominent in middle setigers. Acicular spines first present from about setiger 50 in notopodia and 35 ‒ 40 in neuropodia, numbering 1 – 3 at first, accompanied by capillaries; in far posterior setigers notopodial spines numbering 7 – 8, neuropodial spines numbering 10 – 11, forming complete setal cinctures with 17 – 19 spines on a side and accompanied by alternating, long, thin capillaries (Figs. 15 D, 16 A); spines pointed, slightly curved, notopodial spines directed ventrally, neuropodial spines directed dorsally, spines with weak node or notch at point of emergence from podial lobe (Fig. 15 E – F), with thick borders and fine internal striations (Fig. 16 B – C). Far posterior setigers narrowing, pygidium with terminal anus and small, flattened ventral disk (Fig. 15 C). Methyl Green staining pattern. Stain concentrating on dorsum of last peristomial ring and on setiger 1, some stain dorsally on setigers 2 – 3; some anterior setigers retaining stain ventrally.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB7BFFC3D2F7F98EFC2AFF65.taxon	discussion	Remarks. Chaetozone hobsonae n. sp. is similar to C. columbiana and C. ruffi n. sp. in that a distinct achaetous segment between the peristomium and setiger 1 is not readily apparent due to weakly developed peristomial annuli; because of this, the species is interpreted as lacking an achaetous segment. The species is however, readily recognized in benthic samples due to the triangular prostomium, the pigmented nuchal organs that sometimes resemble eyes, no apparent achaetous segment between the peristomium and setiger 1, acicular spines from mid-body segments, and fully developed posterior cinctures bearing 17 ‒ 19 spines on a side, as well as a distinct MG staining pattern. Among, the three closely related species, C. columbiana has 11 ‒ 12 spines per posterior cincture, a MG staining pattern, and a terminal cirrus on a short pygidial lobe; C. hobsonae n. sp. has 17 ‒ 19 spines per posterior cincture, a MG staining pattern, and pygidial lobe without an cirrus; C. ruffi n. sp. has 20 ‒ 22 spines per posterior cincture, no MG staining pattern, and a pygidial lobe without a cirrus. Further, the acicular spines of C. hobsonae n. sp. begin in middle body segments whereas in C. columbiana and C. ruffi n. sp., they begin in the posterior one-third of the body.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB7BFFC3D2F7F98EFC2AFF65.taxon	biology_ecology	Biology and Ecology. Chaetozone hobsonae n. sp. was part of a rich benthic macrofaunal assemblage consisting of more than 400 species of invertebrates from shallow waters (5 ‒ 30 m) off Ridley Island, Prince Rupert, BC (Blake, unpublished data). The sediments varied throughout the study area with some sites dominated by sands and others with silt; some samples had shell hash, others wood chips. The habitat diversity contributed to the high faunal diversity. The bivalve Nutricola lordi (Baird, 1863) was the most abundant species in the study area followed by the polychaetes Levinsenia gracilis (Tauber, 1879), Mediomastus californiensis Hartman, 1944, Cossura pygodactylata Jones, 1956, Leitoscoloplos pugettensis Pettibone, 1957, Apistobranchus ornatus Hartman, 1965, and Scoletoma zonata (Johnson, 1901).	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB7BFFC3D2F7F98EFC2AFF65.taxon	etymology	Etymology. This species is named for the late Katharine D. Hobson, in recognition of her work on the polychaete fauna of British Columbia.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB7BFFC3D2F7F98EFC2AFF65.taxon	distribution	Distribution. British Columbia and SE Alaska, low water to 95 m; this species is likely more widely distributed in the northeastern Pacific (R. E. Ruff, personal communication).	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB7CFFC0D2F7FC46FB2FFE45.taxon	materials_examined	Material examined. Southeastern Alaska, Boca de Quadra, Cruise 3 BQ, R / V Redoubt, Sta. 100 - 2, 55 ° 19.2 ʹN, 130 ° 29.2 ʹW, 95 m, coll. Dec 1979, R. L. Cimberg, Van Veen grab, holotype (LACM-AHF Poly 6552), one paratype (LACM-AHF Poly 6553). Prince Rupert, British Columbia, off Ridley Island, coll. September 2012, AECOM by P. Winchell & S. Doner, few specimens mixed with C. hobsonae n. sp., mostly incomplete, 12 ‒ 20 m (JAB).	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB7CFFC0D2F7FC46FB2FFE45.taxon	description	Description. A moderate-sized species, holotype complete, 6.0 mm long, 0.4 mm wide for 65 setigerous segments; paratype in two pieces, 10.5 mm long, 0.5 mm wide for 64 setigerous segments; Prince Rupert specimens small, mostly incomplete. Color in alcohol light tan to brown, lacking any distinctive body pigment. Body generally thick, with narrow segments throughout, middle body segments widest; last 12 ‒ 15 setigers formed into distinct cinctures with high membranes bearing spines. Dorsum rounded, with narrow and shallow dorsal groove apparent from about setiger 20, continuing along body until cinctured posterior segments; venter somewhat flattened, with well-developed ventral groove present from about setiger 20, continuing through middle body segments, not apparent posteriorly. Prostomium swollen posteriorly, narrowing anteriorly to triangular, blunted margin (Fig. 17 A); eyes absent; small slit-like nuchal organ present, not pigmented; peristomium with three nearly equal rings, merging with posterior margin of prostomium (Fig. 17 A); achaetous segment absent; dorsal tentacles arising from posterior margin of posterior ring; first pair of branchiae lateral and slightly posterior to tentacles on anterior margin of setiger 1 (Fig. 17 A); second pair of branchiae on posterior edge of setiger 1, dorsal to notosetae; subsequent setigers with branchiae in similar location. Setiger 1 of approximately same size as last peristomial annulation and following segments; podial lobes reduced to inconspicuous ridges in anterior setigers; inflated and conspicuous in middle setigers, greatly enlarged with elevated ridges and conspicuous armature in posterior setigers (Fig. 17 B); posterior segments separated by deeply cut intersegmental furrows (Fig. 17 B). Noto- and neurosetae from setiger 1 all capillaries; notosetae 9 – 10 per fascicle, neurosetae 6 – 8 per fascicle; each capillary thickened, with no distinct fibrils apparent along edge; middle body segments with long, natatory-like notosetae. Acicular spines first present from about setiger 30 in neuropodia and 40 in notopodia of holotype and setiger 40 in neuropodia and 45 in notopodia of paratype; spines numbering 2 – 3 at first, accompanied by an equal number of thin capillaries; in far posterior setigers notopodial spines numbering 9 – 11 and neuropodial spines numbering 11 – 12, forming nearly complete cinctures with spines numbering 20 – 23 on a side and accompanied by alternating thin capillaries (Fig. 18 A, B); spines with sharply pointed tip that curves back and adheres to shaft (Fig. 17 C), with weak node or notch at point of emergence from podial lobe (Fig. 18 C). Last few cinctured setigers narrowing to posterior end; pygidium with terminal anus and small flattened ventral lobe (Fig. 17 B). Methyl Green staining pattern. Tip of prostomium staining, last two peristomial rings staining, with some streaks extending dorsally onto expanded anterior peristomial ring.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB7CFFC0D2F7FC46FB2FFE45.taxon	biology_ecology	Biology. Paratype with body full of oocytes about 70 µm in diameter.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB7CFFC0D2F7FC46FB2FFE45.taxon	discussion	Remarks. The curved tip of the posterior spines of C. camasetosa n. sp. occurs in a small group of Chaetozone species that includes C. curvata Hartmann-Schröder, 1965, from Chile, C. commonalis Blake, 1996 from California shelf depths, C. allanotai from California deep-water slope depths, and C. anasima Doner & Blake, 2006 from offshore New England. Of these, C. curvata and C. commonalis have the first pair of branchiae on setiger 1, whereas C. allanotai, C. anasima, and C. camasetosa n. sp. have an extra pair of branchiae lateral and posterior to the dorsal tentacles on the anterior margin of setiger 1, as well as a pair on the posterior margin of the same setiger. This condition suggests that a segment has been lost or fused with setiger 1. These three species differ from one another in that C. anasima lacks distinct peristomial rings including any demarcation or annulation between the peristomium and setiger 1; whereas, both C. camasetosa n. sp. and C. allanotai have two distinct rings. The latter two species appear to be closely related to one another and may be a shallow-water to deep-water sibling species pair. In C. camasetosa n. sp., the two peristomial rings are strongly set off from one another by deep annulations and also separated from the swollen posterior margin of the prostomium. In C. allanotai, the peristomial rings are not strongly demarcated and, in addition, the first pair of branchiae actually occurs on setiger 1. Both species have different MG staining patterns on the prostomium and peristomium. C. allanotai appears to be limited to continental slope depths of 1800 ‒ 3100 m and is a dominant species in the 2700 ‒ 2850 m depth range from sites offshore northern California (Blake 2006). The specimens of C. camasetosa n. sp. examined here are from shelf depths of up to 95 m.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB7CFFC0D2F7FC46FB2FFE45.taxon	etymology	Etymology. The epithet is from the Latin camur, for crooked or curving inward, combined with seta referring to the manner in which the tip of the posterior spines curve inward merging on the concave side of the shaft and forming an apparent blunt tip.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB7CFFC0D2F7FC46FB2FFE45.taxon	distribution	Distribution. Southeastern Alaska to British Columbia, subtidal, 12 ‒ 95 m. The specimens here suggest that they fragment easily and as such cannot be readily identified without the posterior modified spines.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB7EFFFFD2F7FA85FC58FF48.taxon	materials_examined	Type species: Tharyx acutus Webster & Benedict, 1887, by monotypy.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB7EFFFFD2F7FA85FC58FF48.taxon	diagnosis	Diagnosis. Prostomium conical; peristomium elongate, with pair of grooved dorsal tentacles arising on posterior margin anterior to setiger 1; first pair of branchiae typically arising immediately posterior to dorsal tentacles either on posterior margin of peristomium or on setiger 1; abdominal segments sometimes beadlike. Noto- and neurosetae arising close to one another, not widely separated. Setae include simple capillaries in anterior and middle setigers, acicular spines in posterior setigers with irregular notched tips, sometimes appearing more or less bidentate, with pair of stunted or rounded knobs but never with distinct, sharply pointed teeth; spines present either in both posterior noto- and neuropodia or only in neuropodia. Pygidium with terminal anus and small ventral lobe or disk.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB7EFFFFD2F7FA85FC58FF48.taxon	discussion	Remarks. Blake (1991) determined that the type species of Tharyx, T. acutus Webster & Benedict, 1887 had knob-tipped acicular spines in addition to capillaries. At the time, Tharyx species were defined as having only capillaries (Hartman 1961). Blake (1991) limited Tharyx to those species having knob-tipped spines and moved species having only simple or serrated capillaries to the genera Aphelochaeta Blake, 1991 and Monticellina Laubier, 1961 respectively. The genus Tharyx superficially appears most closely related to Caulleriella in that both genera have species with modified spines that are more or less bidentate. Caulleriella and Tharyx are easily distinguished, however, by the position of the noto- and neuropodia. In Caulleriella, the setal fascicles are widely separated from one another, so much so that in cross section of some species, they appear to be positioned at four corners. In Tharyx, on the other hand, the setal fascicles are close together. Two new species of T haryx are described as part of this study. One species from Alaska had been previously referred to the genus Chaetozone. The second species, from the Puget Sound, Washington, was known in Puget Sound monitoring programs under a provisional Tharyx designation.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB41FFFCD2F7FEFAFB7CFC88.taxon	materials_examined	Material examined. Alaskan Arctic, Beaufort Sea, Prudhoe Bay, coll. M. Busdosh, 0 6 Aug 1983, 70 ° 24.1 ʹN, 148 ° 32.3.0 ʹW, 2.5 m, push cores, Waterflood 6 project, R / V Annika Marie, Sta. 50 - 1, holotype (LACM-AHF Poly 6554), 10 paratypes (LACM AHF Poly 6555). Off Point Barrow, coll. G. E. McGinitie, 0 9 Aug 1949, 7 miles offshore, 128 m, stones and gravel, 1 specimen (USNM 1263247); coll. 0 6 September 1949, 7.5 miles offshore, 66 m, removed from growth on rocks, 2 specimens (USNM 1263248); 14 Oct 1949, 4 miles offshore, 53 m, gravel and small stones 1 specimen (USNM 22810).	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB41FFFCD2F7FEFAFB7CFC88.taxon	description	Description. Holotype 11 mm long, 0.7 mm wide for ca. 75 setigerous segments; paratypes of similar size. Body light tan in alcohol, without pigment. Body with dorsum of anterior and middle segments slightly elevated above parapodia; venter somewhat flattened throughout body, with weak ventral groove in posterior 30 segments. First 12 ‒ 15 segments narrower, more crowded than middle segments, no moniliform segments except in some juveniles; far posterior expanded region again with narrow, crowded segments (Fig. 20 C). Prostomium triangular, narrow, pointed on anterior margin; eyes absent (Figs. 19 A, 20 A); nuchal organs small, crescent shaped notch at posterior lateral margin of prostomium, not pigmented. Peristomium longer than wide, with 2 ‒ 3 distinct annulations visible dorsally, depending on preservation (Fig. 20 A), with 2 – 3 weak divisions ventrally; dorsal midline with a long ridge, extending to end of peristomium (Figs. 19 A, 20 A), better developed on some specimens. Paired dorsal tentacles arising from posterior margin of peristomium, first pair of branchiae arising lateral and posterior to tentacles on setiger 1 (Fig. 19 A); second pair of branchiae on posterior margin of setiger 1, subsequent branchiae on following setigers. Parapodia low mounds from which setal fascicles arise; noto- and neuropodial setal fascicles positioned close to one another throughout. Notosetae all capillaries for first 55 setigers with 5 – 7 capillaries in noto- and neuropodia; notosetae of far posterior setigers becoming spinous, straight, with blunt tips (Fig. 19 D – E); natatory setae present in mature individuals; neurosetae transitioning from capillaries in anterior and middle setigers to short curved spines at about mid body, setiger 34 in holotype. Neuropodial spines curved, somewhat geniculate with tip blunt; shortest spines ventralmost (Fig. 19 D, F). Pygidium with dorsal anus and flattened ventral lobe (Figs. 19 B ‒ C, 20 B). Methyl Green staining pattern. No distinct staining pattern apparent.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB41FFFCD2F7FEFAFB7CFC88.taxon	discussion	Remarks. Tharyx alaskensis was one of the dominant polychaetes in surveys offshore Prudhoe Bay in shallow water as part of benthic monitoring associated with oil and gas development in the early 1980 s (Busdosh 1984). At the time, the species was tentatively referred to Chaetozone cf. gracilis Moore, probably because posterior neuropodial spines appeared to be limited to the neuropodia. In actuality, there are spinous notosetae in far posterior notopodia, but they are stiff, narrow, thinner and with a blunt point instead of short, thick, and curved neuropodial spines. Based on Blake’s (1991) revision of some cirratulid genera, these specimens should be referred to the genus Tharyx because the posterior spines are few, blunt-tipped, and not formed into cinctures. The species is unusual in that the posterior notopodial and neuropodial spines differ from one another: the notopodial spines are straight, stiff, thin, and elongate, whereas the neuropodial spines are curved, somewhat geniculate, short, and thick; both sets of spines are blunt-tipped and not pointed, but are not of the typical knob-tipped form of related species. A few specimens of T. alaskensis n. sp. were encountered among the cirratulids identified as Chaetozone setosa by Pettibone (1954) from off Point Barrow, Alaska, in deeper water and are described as C. pigmentata n. sp. elsewhere in this paper. Pettibone’s Figure 33 D is most definitely T. alaskensis n. sp. because it clearly depicts two pairs of branchiae on setiger 1. The Point Barrow specimens agree very well with the shallower water specimens from Prudhoe Bay. Tharyx alaskensis n. sp. appears most similar to T. killariensis (Southern, 1914), which was originally described from Ireland as a species of Caulleriella and is likely widespread in northern Europe (Blake & Göransson, in preparation). In T. killariensis, however, the posterior noto- and neuropodial spines are few in number and are accompanied by capillaries, which are not present in the far posterior parapodia of T. alaskensis n. sp. A major difference between T. alaskensis n. sp. and other species is that the first and second pairs of branchiae both occur on setiger 1. Other species either have the first pair of branchiae on the posterior margin of the peristomium or on an achaetous segment anterior to setiger 1 and the second pair on the subsequent setigerous segment.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB41FFFCD2F7FEFAFB7CFC88.taxon	biology_ecology	Biology. Tharyx alaskensis n. sp. (as Chaetozone cf. gracilis) was the single most abundant benthic invertebrate collected during a monitoring survey in the Beaufort Sea, Prudhoe Bay, in August 1983 (Busdosh 1984). A total of 3,458 specimens were identified, accounting for 30.6 % of the entire infaunal abundance. Other commonly associated species included the polychaetes Eteone longa (Fabricius, 1780), Capitella capitata (Fabricius, 1780), and Chaetozone ruffi n. sp. (as C. setosa), the cumacean Diastylis sulcata Calman, 1912, and the isopod Saduria entomon (Linnaeus, 1758). Tharyx alaskensis occurred in soft sediments, often with debris. One specimen with natatory setae was gravid, with the body filled with eggs of about 150 µm in diameter. The few specimens available from offshore Point Barrow in deeper water were largely collected from coarse rocky substrates.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB41FFFCD2F7FEFAFB7CFC88.taxon	etymology	Etymology. The name refers to Alaska, where the species occurs and appears to be endemic.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB41FFFCD2F7FEFAFB7CFC88.taxon	distribution	Distribution. Known only from Alaska, Beaufort Sea; shallow subtidal to 128 m offshore.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB42FFFAD2F7FC3AFC36F8FB.taxon	materials_examined	Material examined. — Washington, Strait of Juan de Fuca, east central Port Angeles Harbor, coll. WA Department of Ecology, Puget Sound Ecosystem Monitoring Program (PSEMP), Sta. 1121, 18 June 2013, 48 ° 07.9026 ʹ N, 123 ° 23.2853 ʹ W, 29.5 m depth, in sandy silt, holotype (LACM-AHF Poly 6556), 12 paratypes (LACM-AHF Poly 6557).	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB42FFFAD2F7FC3AFC36F8FB.taxon	description	Description. Holotype 14.8 mm long, 0.43 mm wide for ca. 70 setigerous segments; largest paratype 16.4, mm long, 0.32 mm wide with 95 setigers. Body light tan in alcohol; with a cluster of lateral black pigment spots on posterior peristomium, otherwise without pigment except for a few cells surrounding nuchal organs on some specimens. Body long, slender, with first 15 ‒ 20 segments expanded, widest part of body with narrow, crowded segments each four times wider than long; following segments becoming as long as wide, with some in middle of body almost moniliform; far posterior segments narrowing to small pygidial segment. Body mostly cylindrical in cross section, with weakly developed ventral groove in far posterior segments. Prostomium triangular, tapering to pointed anterior margin; eyes absent (Figs. 21 A – B, 22 A ‒ B); nuchal organs narrow slits surrounded by pigmented cells (Fig. 21 B). Peristomium wider than long, with one annulation visible laterally and dorsally (Figs. 21 A ‒ B, 22 A); dorsal midline with a low, weakly developed ridge extending to end of peristomium (Figs. 21 A, 22 A). Paired dorsal tentacles arising from posterior margin of peristomium; first pair of branchiae arising lateral and posterior to tentacles on posterior edge of peristomium (Figs. 21 A, 22 A); second pair of branchiae arising from posterior margin of setiger 1, dorsal to notosetae. Parapodia low ridges from which setal fascicles arise; noto- and neuropodial setal fascicles positioned close to one another throughout. Notosetae all capillaries throughout (Fig. 21 D ‒ E); 5 – 7 in anterior setigers, reduced to 4 – 5 in far posterior parapodia. Neurosetae all capillaries in anterior setigers, with ventral-most setae transitioning to shorter, recurved spines in middle body (setiger 38 in holotype and setiger 40 in largest paratype); spines curved, somewhat geniculate with blunt tip; shortest spines in ventral position (Fig. 21 F ‒ H); spines accompanied by 2 – 4 dorsal capillaries at first (Fig. 21 D), far posterior setigers with 3 – 5 spines and no capillaries (Fig. 21 E). Pygidium with dorsal anus and small ventral lobe (Figs. 21 C, 22 C ‒ D). Methyl Green staining pattern. Stain retained in intersegmental grooves of some anterior and middle body segments; weak mid-ventral stain sometimes evident.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB42FFFAD2F7FC3AFC36F8FB.taxon	discussion	Remarks. The nature of the prostomium and peristomium of Tharyx circacutus n. sp. are typical for most species of Tharyx where branchiae begin posterior to the tentacles at the posterior margin of the peristomium and the second pair of branchiae begin on setiger 1. Curved posterior spines are limited to the neuropodia and begin about three-fourths of the way along the body; in far posterior segments the spines assume the blunt, knob-tipped appearance of other Tharyx spp. T. circacutus n. sp. most closely resembles the type species, T. acutus, from the northeastern United States in having the posterior spines limited to the neuropodia (Blake 1991). However, the pigmented nuchal organs, the very narrow almost moniliform posterior end, and the very narrow, slender body differ noticeably from T. acutus, which lacks pigmented nuchal organs and in which the entire body and posterior end are more robust with none of the segments appearing moniliform. Further, the body of T. circacutus n. sp. is nearly round in cross section whereas the body of T. acutus is thicker and somewhat dorsoventrally flattened.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB42FFFAD2F7FC3AFC36F8FB.taxon	etymology	Etymology. The Latin circa for around or near is combined with the Latin term acutus for sharp pointed and the species name of the type-species Tharyx acutus, with which this new species is closely related.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
9A3E87FEEB42FFFAD2F7FC3AFC36F8FB.taxon	distribution	Distribution. Known from shallow subtidal habitats in the Puget Sound.	en	Blake, James A. (2015): New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitsbergen in the Norwegian Arctic. Zootaxa 3919 (3): 501-552, DOI: 10.11646/zootaxa.3919.3.5
