taxonID	type	description	language	source
9B1687F8FFDC25003288FC53FEDFFA98.taxon	materials_examined	TYPE MATERIAL. — Gulf of California. San José I., coll. L. Diguet, 5. VII. 1898, 1 syntype ♂ 5.8 mm; 1 syntype ♀ 2.2 mm (MNHN-Pg 632). OTHER MATERIAL EXAMINED (see Remarks for meaning of asterisks). — Gulf of California. Espíritu Santo I., trawl, 15 - 25 m, coll. L. Diguet, 1. VIII. 1897, 1 ♂ 6.0 mm; 2 ♀♀ 4.2 - 4.7 mm (MNHN-Pg 633, labelled as “ type ” but not in Bouvier 1898). — Point Lobos, coll. & don. E. F. Ricketts, 20. III. 1940, 3 ♂♂ 2.3 - 6.6 mm; 2 ♀♀ 3.4 - 4.0 mm (USNM 81950). — Pulmo Reef, coll. & don. E. F. Ricketts, 19. III. 1940, 3 ♂♂ 1.7 - 3.0 mm; 1 ♀ 2.9 mm (USNM 81948); 2 ♂♂ 2.4 - 2.8 mm; 2 ♀♀ 2.2 - 2.3 mm (USNM 1076349). — San Josef, L. C. Gulf, coll. Albatross, 31. III. 1911, 2 ♂♂ 4.6 - 4.7 mm; 1 ovig. ♀ 2.1 mm; 1 ♀ 2.3 mm (USNM 1076375). — “ Basse Californie ”, 2 ♂♂ 6.3 - 6.5 mm; 2 ovig. ♀♀ 4.2 - 4.8 mm; 1 ♀ 5.9 mm (* MNHN-Pg 636). Mexico. Sinaloa, Maztlán, Ola Atlas Bay, N of Derecke Point, 1 ovig. ♀ 4.2 mm (USNM 1076386); Nayarit, Isabel I., shore, coll. S. A. Glassell, 2. VII. 1938, 1 ♂ 5.3 mm (USNM), 19. III. 1933, 2 ♂♂ 2.3 - 3.6 mm; 3 ovig. ♀♀ 2.0 - 2.8 mm; 1 juvenile (* USNM 1076364). — Acapulco, Roqueta I., Jean-Louis Étienne Expedition, coll. J. Poupin, snorkeling 1 - 10 m, 8. III. 2005, 5 ♂♂ 2.0 - 5.0 mm; 3 ovig. ♀♀ 2.6 - 2.8 mm; 2 ♀♀ 2.0 - 3.0 mm (MNHN-Pg 7616). — Unknown locality, coll. “ Mexico, Sec. agricultura 1926 ”, det. W. L. Schmitt, 2 ♂♂ 6.1 - 7.8 mm (* USNM 62385); 1 ♂ 3.6 mm (* USNM 62415); 1 ♂ 4.0 mm; 2 ♀♀ 2.3 - 3.4 mm (* USNM 62436). DISTRIBUTION. — Gulf of California, mainland Mexico from Puerto Peñasco, Sonora (31 ° 20 ’ N) to Huatulco Bay, Oaxaca, and El Salvador (about 13 ° 30 ’ N). Insular records from Revillagigedo (Socorro I.) and tip of Baja California Sur, could be C. mclaughlinae n. sp. (see Remarks under that species). HABITAT. — Lower intertidal, sometimes found in large aggregations (Glassell 1937; Kerstitch 1989), and shallow water to 15 - 25 m, often on Pocillopora coral. DIAGNOSIS Ocular acicle terminating in two to five spines, or rarely one. Anterodorsal plate of branchiostegite smooth, at most with one to two spinules subproximally. Upper margin of right chela smooth (Fig. 1 D). Outer face of left chela regularly convex (Fig. 1 C). Dactyls of P 2 - P 3 with similar setation consisting of sparse, widely-spaced tufts of setae (Fig. 1 E). Propodus of P 3 with dorsolateral margin weakly angled; upper half of outer face slightly concave, forming faint sulcus. Dactyl of P 3 0.6 - 0.7 times length of propodus, with six to eight small spines on ventral margin. Telson with posterior lobes each with single terminal spine. Coloration (Fig. 1) In life, shield black with red orange margins, turning quickly to orange after few days in preservative. Ocular acicles with orange hue at bases. Ocular peduncles dark brown to black, with narrow white band close to corneas. Antennular peduncles and flagella orange. Antennal peduncles, flagella, and antennal acicles bright red, with white-tipped spines. Chelipeds black on meri, carpi, palm of chelae, margins bright red. Dactyl of left cheliped black on inner and outer faces, margins bright red; dactyl of right cheliped uniformly bright red; tip and cutting edges of fingers white. Ambulatory legs uniformly bright red; claws of dactyls black. Abdomen cream to white. REMARKS Calcinus californiensis is morphologically close to C. mclaughlinae n. sp. and, although this still must be proved by examination of living coloration, specimens of C. californiensis from west coast of Baja California (Haig et al. 1970; Ball & Haig 1974) and Socorro I. (Hernández Aguilera 2002) could belong in fact to C. mclaughlinae n. sp. Differences between the two are discussed under the latter species. The former is also morphologically close to C. explorator Boone, 1930 but the two can be easily separated by differences in coloration (Figs 1; 3). In C. californiensis, the distal white ring on the ocular peduncle is larger than in C. explorator; the chelipeds are black with bright red margins; the P 2 and P 3 are red. In C. explorator, the chelipeds are totally black, including the margins; the P 2 and P 3 meri, carpi, propodi are black; the dactyls are orange with a large black median ring, and a narrow black proximal ring. In the absence of coloration, the two species can usually be separated by the armature of the ocular acicles, with several spines in C. californiensis, whereas there is one in C. explorator; the P 3 propodus with an upper margin less angled and the upper half of the outer face less concave in C. explorator. However, because of intraspecific variations, identification of preserved museum specimens based exclusively on morphology is not always possible. For this reason, a few specimens (marked with an asterisk under Other material examined) are herein only tentatively attributed to C. californiensis. Calcinus chilensis (H. Milne Edwards, 1836) (Fig. 2)	en	Poupin, Joseph, Bouchard, Jean Marie (2006): The eastern Pacific species of the genus Calcinus Dana, 1851, with description of a new species from Clipperton Island (Decapoda, Anomura, Diogenidae). Zoosystema 28 (2): 465-486, DOI: 10.5281/zenodo.5393881
9B1687F8FFDC25003288FC53FEDFFA98.taxon	materials_examined	TYPE MATERIAL. — Chile. 1 syntype ♂ 5.2 mm, in poor condition (missing left P 3) (MNHN-Pg 635). OTHER MATERIAL EXAMINED (see Remarks). — Peru. Callao, 2 ♂♂ 7.5 - 7.6 mm (USNM 1024522).	en	Poupin, Joseph, Bouchard, Jean Marie (2006): The eastern Pacific species of the genus Calcinus Dana, 1851, with description of a new species from Clipperton Island (Decapoda, Anomura, Diogenidae). Zoosystema 28 (2): 465-486, DOI: 10.5281/zenodo.5393881
9B1687F8FFDC25003288FC53FEDFFA98.taxon	distribution	DISTRIBUTION. — Chile (unknown locality),? Peru. HABITAT. — Not known, probably intertidal.	en	Poupin, Joseph, Bouchard, Jean Marie (2006): The eastern Pacific species of the genus Calcinus Dana, 1851, with description of a new species from Clipperton Island (Decapoda, Anomura, Diogenidae). Zoosystema 28 (2): 465-486, DOI: 10.5281/zenodo.5393881
9B1687F8FFDC25003288FC53FEDFFA98.taxon	diagnosis	DIAGNOSIS Ocular acicle terminating in spine. Anterodorsal plate of branchiostegite smooth, with one or two spinules subproximally. Upper margin of right chela smooth. Outer face of left chela excavated on lower half, convex on upper half (Fig. 2 B). Left P 3 unknown. Ventrodistal pilosity of right P 2 and P 3 similar, sparse (Fig. 2 D). Propodus of right P 3 with upper margin and upper half of outer face rounded; dactyl 0.75 times length of propodus, with eight spines on ventral margin. Telson with posterior lobes each with single terminal spine. Coloration In life, unknown. The coloration illustrated by H. Milne Edwards (1837: pl. 22, fig. 9), herein reproduced (Fig. 2 A), is probably based on a preserved specimen.	en	Poupin, Joseph, Bouchard, Jean Marie (2006): The eastern Pacific species of the genus Calcinus Dana, 1851, with description of a new species from Clipperton Island (Decapoda, Anomura, Diogenidae). Zoosystema 28 (2): 465-486, DOI: 10.5281/zenodo.5393881
9B1687F8FFDC25003288FC53FEDFFA98.taxon	discussion	REMARKS	en	Poupin, Joseph, Bouchard, Jean Marie (2006): The eastern Pacific species of the genus Calcinus Dana, 1851, with description of a new species from Clipperton Island (Decapoda, Anomura, Diogenidae). Zoosystema 28 (2): 465-486, DOI: 10.5281/zenodo.5393881
9B1687F8FFDC25003288FC53FEDFFA98.taxon	description	Two discoloured specimens from Peru are herein tentatively attributed to C. chilensis because of the similar left chela (see Fig. 2 E), the presence of only one spine on the ocular acicle, and collection locality (Callao, 12 ° S). A number of sibling species have been recognized in Calcinus recently, raising the possibility of the existence of new color variants along the eastern Pacific, especially in latitudes ranging from 10 ° to 25 ° S where scarce collections or observations have been made (Fig. 7). Such is needed to make comparisons of any new variants with C. chilensis.	en	Poupin, Joseph, Bouchard, Jean Marie (2006): The eastern Pacific species of the genus Calcinus Dana, 1851, with description of a new species from Clipperton Island (Decapoda, Anomura, Diogenidae). Zoosystema 28 (2): 465-486, DOI: 10.5281/zenodo.5393881
9B1687F8FFDC25003288FC53FEDFFA98.taxon	materials_examined	TYPE MATERIAL (not seen, see Remarks). — Galápagos. Española I. (= Hood I.), Gardner Bay, stn 54, 1925 Arcturus Oceanographic Expedition, coll. W. Beebe, diving 4.5 m, possible holotype 1 ♂ (AMNH 12244), possible paratypes 8 specimens (AMNH 12245). Others AMNH lots labeled as paratypes are: Galápagos. Eden I., 4 specimens (AMNH 12582), Genevosa I. (= Tower I.), 15 specimens (AMNH 6312). — Cocos I. 1 specimen (AMNH 6309), 2 specimens (AMNH 6313), 8 specimens (AMNH 12577). MATERIAL EXAMINED (see Remarks for meaning of asterisks). — Galápagos. Santiago I. (= James I.), 1975, coll. H. Kühl, poor condition, 2 ♂♂ 3.0 - 4.5 mm; 3 ♀♀ 2.3 - 4.0 mm; 2 juveniles 2.0 - 2.2 mm (RMNH 31343). — Santa Fe I. (= Barrington I.), 10. I. 1972, M. L. Azzaroli leg., don. B. Lanza, 3 ♂♂ 4.0 - 6.0 mm (RMNH 31948). — Isabela I., Elizabeth Bay, 26. VII. 1938, coll. W. L. Schmitt, 1 ♂ 2.3 mm; 4 juveniles in shells (USNM 77820). — Santa Cruz I., coll. Galápagos New York Expedition 1923, 2 ♂♂ 3.9 - 4.9 mm; 1 ♀ 4.4 mm (USNM 57722). — Isabela I., Tagus Cove, 29. X. 1961, coll. J. Joseph and E. Forsbergh, Swansong cruise Inter- AmericanTropical Fish Commission, 2 ♂♂ 4.9 - 5.0 mm; 1 ♀ 2.2 mm; 8 specimens in shells (USNM 1076311); 17. II. 1933, 1 ♂ 3.8 mm (* USNM 1076330), 26. I. 1933, 1 ♂ 1.6 mm (* USNM 1076328); 24. I. 1933, 8 ♂♂ 3.0 - 6.0 mm; 6 ovig. ♀♀ 3.0 - 5.3 mm; 3 ♀♀ 2.6 - 4.6 mm; 6 juveniles (* USNM 1076325); 8. II. 1933, 1 ♂ 2.0 mm; 1 juvenile (* USNM 1076329); 14. II. 1933, 1 juvenile (* USNM 1076331). — Marchena I., 2. XII. 1934, coll. W. L. Schmitt, poor condition, 5 ♂ 2.3 - 5.0 mm; 14 ovig. ♀♀ 2.6 - 4.0 mm; 3 ♀♀ 3.0 - 4.0 mm; 3 juveniles (USNM 1076327). — Santiago I. (= James I.), Sullivan Bay, stn 15, 24. VII. 1938, coll. W. L. Schmitt, 1 ♂ 2.6 mm (USNM 77830). — Fernandina I., coll. W. L. Schmitt, 25. VII. 1938, 1 specimen in shell (USNM 77824); 1 ♂ 4.6 mm (USNM 1076324). — San Cristóbal (= Chathman I.), coll. W. H. Jones, VIII. 1884, 1 ♂ 8.6 mm (USNM 9332). — Pinzon I. (= Duncan I.), 13. IV. 1888, coll. Albatross, 3 ♂♂ 4.2 - 6.5 mm (USNM 1076326). — Santa Maria I. (= Charles I.), 6. XII. 1934, coll. W. L. Schmitt, 1 ♀ 1.9 mm; 1 juvenile (* USNM 1076332). — “ Galápagos ”, 4 ♂♂ 5.1 - 6.1 mm (USNM 33504) Clipperton I. Side reef flat at low tide, depth 0 - 0.3 m (1 ft), 13. IX. 1958, coll. C. Limbaugh, T. Chess, A. Hambly, 1 ♂ 2.8 mm (USNM 110971); rocky intertidal, 7 - 26. VIII. 1958, coll. E. S. Reese, 85 ♂♂ 1.7 - 6.8 mm; 56 ovig. ♀♀ 2.3 - 5.0 mm; 42 ♀♀ 1.6 - 3.5 mm (USNM 110990); coll. Service de Santé des Armées, 4 ♂♂ 6.0 - 7.6 mm (MNHN-Pg 564); 1966, coll. Le Chuiton, 1 ♂ 6.9 mm (MNHN-Pg 651); leg. Chace, Smithsonian, 10 specimens (MNHN-Pg 653). Jean-Louis Étienne Expedition to Clipperton 2004 - 2005, outer reef at low tide, I-II. 2005, coll. J. Poupin, J. M. Bouchard, L. Albenga, 19 ♂♂ 1.9 - 8.5 mm; 4 ovig. ♀♀ 2.3 - 3.0 mm; 1 ♀ 2.2 mm; 1 juvenile 1.1 mm; 22 specimens in shells (MNHN-Pg 7617). — Outer reef, low tide, stn 4, 10 ° 18.11 ’ N, 109 ° 14 ’ W, coll. L. Albenga, J. M. Bouchard, L. Dugrais, 9. I. 2005, 3 specimens in shells (MNHN-Pg 7618). — Outer reef, low tide, stn 33, 10 ° 18.27 ’ N, 109 ° 14 ’ W, coll. L. Albenga, L. Dugrais, 25. I. 2005, 5 ♂♂ 2.2 - 7.5 mm; 1 ♀ 3.5 mm; 5 specimens in shells (MNHN-Pg 7619). Clarión I. Stn 2, coll. A. W. Anthony, 3 ♂♂ 4.4 - 5.6 mm; 1 ♀ 5.3 mm (* USNM 42545). — Sulphur Bay, coll. W. L. Schmitt, stn 141 - 34, 1 ♂ 4.0 mm (* USNM 1076333).	en	Poupin, Joseph, Bouchard, Jean Marie (2006): The eastern Pacific species of the genus Calcinus Dana, 1851, with description of a new species from Clipperton Island (Decapoda, Anomura, Diogenidae). Zoosystema 28 (2): 465-486, DOI: 10.5281/zenodo.5393881
9B1687F8FFDC25003288FC53FEDFFA98.taxon	description	Socorro I. Braithwaite Bay, shore collecting at landing place, 20. VII. 1938, coll. W. L. Schmitt, 3 specimens in shells (USNM 78212). — Crayson Cove, intertidal zone, 7. III. 1937, coll. W. Williams, “ M. S. Stranger ”, 5 ♂♂ 2.2 - 4.5 mm (USNM 1076323). — Coll. Albatross, 8 ♂♂ 2.7 - 6.1 mm; 2 ovig. ♀ 3.9 - 5.1 mm; 2 ♀♀ 2.8 - 3.4 mm (USNM 33505).	en	Poupin, Joseph, Bouchard, Jean Marie (2006): The eastern Pacific species of the genus Calcinus Dana, 1851, with description of a new species from Clipperton Island (Decapoda, Anomura, Diogenidae). Zoosystema 28 (2): 465-486, DOI: 10.5281/zenodo.5393881
9B1687F8FFDC25003288FC53FEDFFA98.taxon	materials_examined	Mexico. Nayarit, Isabel I. shore, 2. VII. 1938, coll. S. H. Glassell, “ Isabel Is. Mex. ”, 2 ♂♂ 4.0 - 6.0 mm (USNM 1076312). Cocos I. I. 1902, coll. P. Biolley, 1 ♂ 5.8 mm (* USNM 33503). DISTRIBUTION. — Revillagigedo (Socorro I., Clarión I.); Clipperton I.; Cocos I.; Galápagos (Española I., Fernandina I., Genovesa I., Isabela I., Marchena I., Santa Cruz I., Santa Maria I., Santa Fe I.); Gulf of California (Carmen I., Cabo Pulmo); Mexico (Tres Marías Is., Isabel I., Punta Mita, Chamela, Tenacatita). The species is also reported from Colombia by Snyder-Conn (1980: 278, undefined place) but it must have been confused here with C. obscurus. HABITAT. — Collected at low tide in coral rubble; common in upper and median part of intertidal area, occasionally in lower part up to depth of few meters. DIAGNOSIS Ocular acicle terminating in one or rarely two spines, on one side only. Anterodorsal plate of branchiostegite smooth with one or two spinules subproximally. Upper margin of right chela smooth (Fig. 3 D). Outer face of left chela regularly convex (Fig. 3 C). Dactyls of P 2 and P 3 with similar setation consisting of sparse, widely-spaced tufts of setae (Fig. 3 E). Propodus of P 3 with dorsolateral margin rounded or weakly angled; upper half of outer face evenly convex or slightly depressed. Dactyl of P 3 0.7 - 0.8 times length of propodus, with five to seven spines on ventral margin. Telson with posterior lobes each with single terminal spine. Coloration (Fig. 3) In life, shield black, paler on proximal half, fading quickly to orange after few days in preservative. Ocular acicles white on distal half, black at bases. Ocular peduncles black, with narrow white band close to corneas. Antennular peduncles black with distal half of terminal segments and flagella orange. Antennal peduncles black on first, to third segments, and orange on fourth and fifth segments, flagella, and antennal acicles. Chelipeds black on meri, carpi, and chelae, with white on cutting margins and tip of fingers (Fig. 3 D). Ambulatory legs uniformly black on meri, carpi and propodi. Dactyls bright orange with large black median ring, and narrow black proximal ring (sometimes unclear); terminal claws black. Abdomen cream to white. Eggs red. REMARKS The type series of Calcinus explorator is defined by Boone (1932: 22) as: “ The type, an adult male specimen and four additional specimens, one ovigerous ”. This material was checked for us in AMNH collections by C. B. Boyko (pers. comm.). According to place of collection, one male (AMNH 12244) labelled as “ Paratype ” must be the holotype. The paratypes are in AMNH 12245 but this lot includes eight specimens instead of four cited by Boone. Five others AMNH lots are labeled as “ Paratypes ”: Galápagos, Eden I., 6. IV. 1923, 4 specimens (AMNH 12582); Genovesa I. (= Tower I.), C. Hay leg., 15. IV. 1930, 15 specimens (AMNH 6312); Cocos I., J. Chapin leg., 17. IV. 1930, 1 specimen (AMNH 6309), 2 specimens (AMNH 6313); 31. III- 2. IV. 1925, 8 specimens (AMNH 12577). AMNH collection includes two additional uncataloged lots from the Vanderbilt Museum labeled as “ Pagurus explorator ”, probably those cited by Boone (1930: 28): Galápagos, Genovesa I. (= Tower I.), Darwin Bay, 13. III. 1926; Española I. (= Hood I.), 5. II. 1928. The black coloration of Calcinus explorator is very similar to that of C. obscurus. Some authors have discussed their possible synonymy (Chace 1962: 626; Hendrickx & Harvey 1999: 368). The two species, however, can be separated by the armature of the ocular acicles, one spine in C. explorator, and two to four spines in C. obscurus; and by subtle colour differences. The chelae are totally black in C. explorator, whereas there are orange tints on the margins in C. obscurus (Figs 3 B; 5 A); the dactyls of P 2 and P 3 are orange with a large median black ring and narrow proximal black ring in C. explorator, whereas this color pattern is variable in C. obscurus (Fig. 6); the distal white band on the ocular peduncles is narrower in C. explorator than in C. obscurus. Calcinus explorator is also morphologically close to C. mclaughlinae n. sp. The two can be distinguished based on coloration (Figs 3; 4), if still present on specimens. The specimens from Clarión I., Cocos I., and Galápagos, marked by an asterisk in Material examined, had lost coloration, and thus are A E assigned to C. explorator based on the morphological differences cited by Chace (1962: 628). The distal margin of palm of left chela is oblique and almost continuous with the upper margin of fixed finger in C. explorator, whereas the distal margin of the palm is nearly transverse and makes an angle with the upper margin of fixed finger in C. mclaughlinae n. sp. (see Chace 1962: figs 4, 6). The upper part of the outer surface of the propodus of P 3 is less flattened and concave in C. explorator than in C. mclaughlinae n. sp. However, due to intraspecific variation these two characters are often difficult to use reliably. Calcinus explorator has been frequently reported from eastern Pacific Oceanic Islands such as Revillagigedo, Clipperton, Cocos, and Galápagos. Snyder-Conn (1980: 278), based on J. Haig unpublished identifications (M. Hendrickx pers. comm.), listed other localities where C. explorator co-occurs with C. californiensis, such as Gulf of California (Cabo Pulmo), Mexican continental islands (Tres Maria Is., Isabel I.), and Mexican coast (Bay Tenacatita). The presence of C. explorator in the Gulf of California has been confirmed by Rodríguez Almaraz et al. 2002 (Carmen I.), and along the Mexican coast by Hernández Aguilera et al. 1986 (Tres Maria Is., Chamela Bay), and Hendrickx & Harvey 1999 (Punta Mita). Cooccurrence of Calcinus explorator and C. californiensis along the Mexican mainland is confirmed in this study. Both species were positively identified based on morphology and color, in material examined from Nayarit, Isabel I. (coll. Glassell, 2. VII. 1938, USNM).	en	Poupin, Joseph, Bouchard, Jean Marie (2006): The eastern Pacific species of the genus Calcinus Dana, 1851, with description of a new species from Clipperton Island (Decapoda, Anomura, Diogenidae). Zoosystema 28 (2): 465-486, DOI: 10.5281/zenodo.5393881
9B1687F8FFD72519328DFAEFFE63F93C.taxon	description	(Fig. 4)	en	Poupin, Joseph, Bouchard, Jean Marie (2006): The eastern Pacific species of the genus Calcinus Dana, 1851, with description of a new species from Clipperton Island (Decapoda, Anomura, Diogenidae). Zoosystema 28 (2): 465-486, DOI: 10.5281/zenodo.5393881
9B1687F8FFD72519328DFAEFFE63F93C.taxon	materials_examined	TYPE MATERIAL. — Clipperton I. Jean-Louis Étienne Expedition, outer reef, low tide and snorkeling 1 - 2 m, II. 2005, coll. J. Poupin, holotype, 1 ♂ 4.4 mm (MNHN- Pg 7620). Paratypes: same data, 10 ♂♂ 2.0 - 4.4 mm, 7 ovig. ♀♀ 2.0 - 3.0 mm; 2 ♀♀ 1.6 - 2.2 mm; 4 specimens in shells (MNHN-Pg 7621). TYPE LOCALITY. — Clipperton Island.	en	Poupin, Joseph, Bouchard, Jean Marie (2006): The eastern Pacific species of the genus Calcinus Dana, 1851, with description of a new species from Clipperton Island (Decapoda, Anomura, Diogenidae). Zoosystema 28 (2): 465-486, DOI: 10.5281/zenodo.5393881
9B1687F8FFD72519328DFAEFFE63F93C.taxon	etymology	ETYMOLOGY. — This species is dedicated to Patsy A. McLaughlin for her invaluable contributions to the systematics of the anomura. Her warm support to the first author when working at MNHN was a strong encouragement to continue in this field of research. OTHER MATERIAL EXAMINED. — Clipperton I. Jean- Louis Étienne Expedition, outer reef, low tide, stn 4, 10 ° 18.11 ’ N, 109 ° 14 ’ W, 9. I. 2005, coll. L. Albenga, J. M. Bouchard, L. Dugrais, and J. E. Blatteau, 2 ♂♂ 4.4 - 5.0 mm; 1 ovig. ♀ 2.5 mm; 5 specimens in shells (MNHN-Pg 7622). — Outer reef, scuba dive, 15 m, stn 8, 10 ° 18.49 ’ N, 109 ° 14.1 ’ W, 14. I. 2005, 2 specimens in shells (MNHN-Pg 7623). — Outer reef, stn 20, 10 ° 17.50 ’ N, 109 ° 13.55 ’ W, 22. I. 2005, 1 specimen in shell (MNHN-Pg 7624). — Scuba dive, 20 m, stn 41, 10 ° 19.01 ’ N, 109 ° 13.75 ’ W, 29. I. 2005, 1 ♂ 1.5 mm (MNHN-Pg 7625). — Scuba dive 22 m, stn 45, 10 ° 17.49 ’ N, 109 ° 13.52 ’ W, 3. II. 2005, 1 specimen in shell (MNHN-Pg 7626). — Coll. Centre de Recherche et Service de Santé des Armées, 2 ♂♂ 2.3 - 2.4 mm (MNHN-Pg 634). — Leg. Chace USNM, coll. 1958, 5 ♂♂; 5 ♀♀ (MNHN-Pg 631). — NW end reef, 14. VIII. 1958, coll. Limbaugh, Chess, Reese, Hambly, and Wintersteen, size range males 1.3 - 4.2 mm, ovig. females 2.0 - 4.7 mm, females 1.4 - 3.0 mm, 1 ♂; 1 ovig. ♀ (USNM 110896). — Rocky intertidal areas, 7 - 26. VIII. 1958, 39 ♂♂; 2 ovig. ♀♀; 30 ♀♀; 1 juvenile (USNM 110987). — NE side, 14 m (45 ft), 28. VIII. 1958, 4 ♂♂; 1 ovig. ♀; 1 ♀ (USNM 110900. — NE corner, 14 m (45 ft), 30. VIII. 1958, 2 ♂♂; 1 ovig. ♀ (USNM 110897). — NE side, low intertidal to 2 m (6 ft), 4. IX. 1958, 1 ♀; 2 juveniles (USNM 110898). — Reef flat 0 - 0.3 m (1 ft), 13. IX. 1958, 1 ♂ (USNM 110901). — East side reef flat, 14. IX. 1958, 1 ovig. ♀ (USNM 110899).	en	Poupin, Joseph, Bouchard, Jean Marie (2006): The eastern Pacific species of the genus Calcinus Dana, 1851, with description of a new species from Clipperton Island (Decapoda, Anomura, Diogenidae). Zoosystema 28 (2): 465-486, DOI: 10.5281/zenodo.5393881
9B1687F8FFD72519328DFAEFFE63F93C.taxon	description	West coast of Baja California Sur (see Remarks). Coll. A. W. Anthony, 23 ♂♂ 3.2 - 7.0 mm; 10 ovig. ♀♀ 2.8 - 5.6 mm; 4 ♀♀ 3.7 - 5.0 mm (USNM 42550).	en	Poupin, Joseph, Bouchard, Jean Marie (2006): The eastern Pacific species of the genus Calcinus Dana, 1851, with description of a new species from Clipperton Island (Decapoda, Anomura, Diogenidae). Zoosystema 28 (2): 465-486, DOI: 10.5281/zenodo.5393881
9B1687F8FFD72519328DFAEFFE63F93C.taxon	distribution	DISTRIBUTION. — Clipperton Island, possibly Socorro I., and west coast of Baja California Sur. HABITAT. — Lower intertidal to 22 m; on coral Pocillopora, and rubble. At Clipperton, usually found deeper than C. explorator, although the two can be found together.	en	Poupin, Joseph, Bouchard, Jean Marie (2006): The eastern Pacific species of the genus Calcinus Dana, 1851, with description of a new species from Clipperton Island (Decapoda, Anomura, Diogenidae). Zoosystema 28 (2): 465-486, DOI: 10.5281/zenodo.5393881
9B1687F8FFD72519328DFAEFFE63F93C.taxon	diagnosis	DIAGNOSIS Ocular acicle terminating in one or occasionally two or three spines. Upper margin of right chela smooth. Outer face of left chela regularly convex. Dactyls of P 2 - P 3 with similar setation consisting of sparse, widely-spaced tufts of setae. Propodus of P 3 with upper part of outer face flattened or slightly concave, forming low sulcus. Dactyl of P 3 0.7 times length of propodus, with six or seven small spines on ventral margin. Telson with posterior lobes each with single terminal spine. Ocular peduncles dark brown to black with white distal band adjacent to corneas. Chelipeds black on meri, carpi, and palms of chelae, with margins bright red; dactyls of left and right chelae bright red, cutting edges white. Ambulatory legs bright red, with distal half of dactyls orange.	en	Poupin, Joseph, Bouchard, Jean Marie (2006): The eastern Pacific species of the genus Calcinus Dana, 1851, with description of a new species from Clipperton Island (Decapoda, Anomura, Diogenidae). Zoosystema 28 (2): 465-486, DOI: 10.5281/zenodo.5393881
9B1687F8FFD72519328DFAEFFE63F93C.taxon	description	DESCRIPTION Shield about 1.1 as long as broad. Rostrum acute, clearly overreaching level of broadly subtriangular lateral projections; anterior margins between rostrum and lateral projections concave. Anterodorsal plate of branchiostegite with dorsal margin smooth. Ocular peduncles 0.8 - 1.0 times as long as shield, slightly constricted medially; peduncles each six times as long as diameter of corneas. Corneas weakly dilated. Ocular acicles well developed, subtriangular, terminating in simple spine, or rarely two or three spines. Antennular peduncles reaching to distal 0.25 - 0.3 of ocular peduncles. Ultimate segment 0.3 - 0.4 times as long as shield. Antennal peduncles shorter than antennular peduncle, reaching between midpoint and distal 0.3 of ocular peduncles. First segment with ventrolateral angle produced, bearing two to four spines. Second segment with laterodistal angle produced, terminating in bifid spine; distomesial angle unarmed or with spine. Third segment with strong spine at ventrodistal angle. Fourth segment with dorsodistal spine. Fifth segment unarmed. Antennal acicle reaching beyond proximal margin of fifth antennal segment, terminating in strong spine; dorsolateral margin with one or two spines distally, dorsomesial margin with row of three or four spines. Left chela (Fig. 4 C) 0.5 - 0.7 times as high as long. Dactyl slightly shorter than palm, covered with low granules; cutting edge smooth or with one or two obtuse calcareous teeth; dorsolateral margin smooth. Fixed finger with one submedian obtuse tooth on cutting edge; outer and lower faces with low granules. Palm with outer face regularly convex, finely granular; lower face with larger tubercles; inner face almost smooth; upper margin smooth. Carpus with prominent submedian tubercle on outer face; upper margin angled, dorsodistal angle unarmed. Merus stout, triangular in cross-section; ventromesial margin unarmed distally; ventrolateral margin with one stout spine distally; upper margin unarmed. Right chela (Fig. 4 D) smooth on upper margin of palm. Dactyl about as long as palm; cutting edge with one or two obtuse teeth proximally, distal half spoon-shaped; dorsal surface with low granules. Fixed finger with obtuse tooth on proximal cutting edge; distally hoof-shaped. Palm with outer face weakly convex, slightly granulated; inner face flat, almost smooth. Carpus with smooth upper margin, dorsodistal angle unarmed; outer face with 1 low submedian tubercle. Merus less broad than merus of left cheliped, outer face with numerous small pits, dorsal, ventromesial and ventrolateral margins unarmed distally. Ambulatory legs similar from left to right P 2 (Fig. 4 E), reaching to tip of left cheliped when fully extended; dactyl 0.5 - 0.7 times as long as propodus; ventral margin with widely-spaced, sparse tufts of setae and six or seven minute spines; propodus with few long setae, and with or without minute movable spine at ventrodistal angle; carpus 0.5 - 0.6 times as long as propodus, with dorsodistal spine; merus about as long as propodus, with obtuse laterodistal spine. P 3 (Fig. 4 E) shorter than P 2, reaching to base of fixed finger of left chela when fully extended; ventrodistal pilosity similar to that of P 2, not brush-like; dactyl 0.6 - 0.7 times as long as propodus, ventral margin armed with six to eight minute spines; propodus with few setae and one or two minute movable spines at ventrodistal angle, dorsolateral margin angled, upper half of outer face flat to slightly concave, forming a low sulcus; carpus 0.6 - 0.7 times as long as propodus, with dorsodistal spine; merus 0.8 - 0.9 times as long as propodus, with obtuse laterodistal spine, sometimes absent. Fourth pereopod semichelate, bordered with long setae on dorsal and ventral margins. Dactyl terminating in corneous claw; ventrolateral margin with row of spinules. Propodus with broad rasp on outer face consisting of several rows of corneous scales; ventral margin with row of stout spines. Carpus with dorsodistal spine. Merus unarmed. Fifth pereopod chelate, with rasp on propodus and dactyl; carpus and merus unarmed, subovate in cross-section. Abdomen with four unpaired biramous left pleopods in both sexes. Sixth abdominal tergite calcified, with dorsal face divided into four subequal areas by longitudinal and transverse furrows. Telson with posterior lobes asymmetrical, left more elongated than right; lobes with long setae marginally, each with one posterior spine slightly curved ventrally. Size range sl 1.3 - 5.0 mm; smallest ovigerous ♀ 2.0 mm. Coloration (Fig. 4) In life, shield black with bright red on margins. Ocular acicles bright red at base, white at tip. Ocular peduncles dark brown to black with white distal band adjacent to corneas. Antennular peduncles dark brown to black, with orange at articulations of segments and distal 0.5 of ultimate segment; flagella orange. Antennal peduncles, flagella, and antennal acicles orange overall. Chelipeds black on meri, carpi, palm of chelae, margins bright red; dactyls of left and right chelae bright red, white on cutting edges. Ambulatory legs bright red, distal half of dactyls orange, terminal claws black. Fifth and fourth pereopods cream to pale orange with darker orange hues on segments. Abdomen tan, sixth abdominal tergite, uropods and telson with scattered orange patches. In preservative coloration of shield (Fig. 4 B) and outer face of chelae fading quickly to bright red or orange, but color pattern on P 2 - P 3 dactyls still clear after several months.	en	Poupin, Joseph, Bouchard, Jean Marie (2006): The eastern Pacific species of the genus Calcinus Dana, 1851, with description of a new species from Clipperton Island (Decapoda, Anomura, Diogenidae). Zoosystema 28 (2): 465-486, DOI: 10.5281/zenodo.5393881
9B1687F8FFD72519328DFAEFFE63F93C.taxon	discussion	REMARKS	en	Poupin, Joseph, Bouchard, Jean Marie (2006): The eastern Pacific species of the genus Calcinus Dana, 1851, with description of a new species from Clipperton Island (Decapoda, Anomura, Diogenidae). Zoosystema 28 (2): 465-486, DOI: 10.5281/zenodo.5393881
9B1687F8FFD72519328DFAEFFE63F93C.taxon	materials_examined	The coloration of Calcinus mclaughlinae n. sp. is distinct from that of C. explorator (Figs 3; 4). However, uncoloured specimens of the two species can be confused because of same aspect of ocular acicle (one terminal spine). The few uncoloured specimens examined herein from the Revillagigedo (Clarión I.) and Cocos I. are very similar to C. mclaughlinae n. sp. by their overall morphology although tentatively assigned to C. explorator (see Remarks under C. explorator). For the time being, C. mclaughlinae n. sp. can be considered as endemic to Clipperton Island, although it is potentially an “ insular endemic ” sensu Garth (1960, 1991, 1992) i. e. distributed at least in the Revillagigedo and possibly also in the west coast of Baja California Sur (Punta Cala, Roca de la Vela, Punta Tosca, Magdalena Bay). Calcinus californiensis has previously been reported from these insular localities but has not been compared with live material from Clipperton (Haig et al. 1970; Ball & Haig 1974; Hernández Aguilera 2002). During the present study, a lot with 37 preserved specimens (USNM 42550) lacking colour were examined from Baja Magdalena, and are tentatively attributed to C. mclaughlinae n. sp. based on the presence of one spine on the ocular acicles. Two to four spines were observed in all specimens of C. californiensis from the Gulf and Mexican mainland.	en	Poupin, Joseph, Bouchard, Jean Marie (2006): The eastern Pacific species of the genus Calcinus Dana, 1851, with description of a new species from Clipperton Island (Decapoda, Anomura, Diogenidae). Zoosystema 28 (2): 465-486, DOI: 10.5281/zenodo.5393881
9B1687F8FFD72519328DFAEFFE63F93C.taxon	materials_examined	TYPE MATERIAL. — Not seen; probably not extant. Stimpson (1859: 49) indicated that specimens he used had been chiefly supplied from the Museum of the Smithsonian Institution. His syntypes have not been found during this study, and probably were lost in the great Chicago fire of 1871 (see Manning 1993: 112). MATERIAL EXAMINED (see Remarks for meaning of asterisks). — El Salvador. La Libertad, leg. G. Kruseman, summer 1952, 21 ♂♂ 2.6 - 8.3 mm; 9 ovig. ♀♀ 3.1 - 5.4 mm; 7 ♀♀ 2.2 - 5.4 mm (ZMA De 100.355); 9 ♂♂ 2.5 - 7.0 mm; 3 ovig. ♀♀ 3.3 - 5.0 mm; 1 ♀ 3.0 mm (RMNH 9399). — La Libertad, coll. and leg. John Boursot, 10. XII. 1953, 3 ♂♂ 7.5 - 9.6 mm (USNM 96289); VI. 1953, 1 ♂ 3.4 mm; 1 ♀ 3.7 mm (USNM 96316). — La Libertad, shore collecting, 18. I. 1933, 12 ♂♂ 1.8 - 5.3 mm; 12 ovig. ♀♀ 2.3 - 3.5 mm; 8 ♀♀ 2.8 - 4.0 mm (USNM 1076239); 21. I. 1933, 2 ♂♂ 5.5 - 5.8 mm; 1 ♀ 4.4 mm (USNM 1076226). Costa Rica. Puerto Culebra, shore collecting along S slough, 12. III. 1933, 1 ovig. ♀ 2.0 mm (* USNM 1076234). — San Lucas I., coll. & don. M. Valerico, 15. I. 1930, 2 ♂♂ 5.4 - 6.2 mm (* USNM 64134). Panama. Perico I., coll. Albatross, 26. X. 1904, 21 ♂♂ 3.2 - 6.2 mm; 6 ovig. ♀♀ 2.9 - 4.9 mm; 5 ♀♀ 3.1 - 4.9 mm; several others specimens damaged or in shells (USNM 40545). — Chopilla I., coll. L. G. Abele, 26. V. 1969, 3 ♂♂ 7.8 - 9.8 mm; 1 ♀ 7.8 mm (USNM 267603); 3 ♂♂ 6.4 - 7.6 mm (USNM 267600); 1 ♂ 4.7 mm (USNM 267593); 1 specimen in shell (USNM 267590). — Taboguilla I., low tide, coll. L. G. Abele, A. Clarke, T. Dana, J. Graham, G. Powell, and A. Rodaniche, 30. IV. 1969, 3 ♂♂ 4.4 - 6.4 mm; 1 ♀ 4.9 mm; 1 specimen damaged (USNM 267556); 2 ♂♂ 1.8 - 3.5 mm; 1 ovig. ♀ damaged; 2 ♀♀ 2.0 - 3.2 mm (USNM 267596); 12. VI. 1969, several juveniles in shells (USNM 267602); 1 ♂ 3.4 mm (USNM 267554); 7. IV. 1969, several juveniles, poor condition (USNM 267598). — East end of Panama Boulevard, intertidal, coll. L. G. Abele, 17. II. 1969, 3 ♂♂ 1.7 - 3.7 mm (USNM 267599). — Intertidal, beyond Fort Kobke, Venado Beach, coll. L. G. Abele, A. Clarke and J. Graham, 8. V. 1969, 1 ♂ 1.7 mm (USNM 267594); 1. VII. 1969, 1 ♂ 5.6 mm; 1 specimen in shell (USNM 267597). — Panama city, Punta Paitilla, low tide, coll. L. G. Abele, J. Graham, N. Powell, and A. Soler, 7. VII. 1969, 2 juveniles in shells (USNM 267595); 7. VII. 1969, 16 ♂♂ 2.3 - 8.1 mm; 4 ovig. ♀♀ 3.3 - 4.3 mm; 2 ♀♀ 3.0 - 3.8 mm; several other specimens in shells (USNM 267557); 16. IX. 1967, 2 ♂♂ 5.2 - 5.6 mm (USNM 267555); 14. IV. 1969, 4 ♂♂ 2.5 - 4.2 mm; 3 ovig. ♀♀ 4.0 - 4.2 mm; 2 ♀♀ 3.5 - 3.6 mm (USNM 267592); 8. VI. 1969, 1 ♂ 7.6 mm (USNM 267553); 31. XII. 1968, 1 ovig. ♀ 3.9 mm (USNM 267601); 1. VII. 1969, 6 ♂♂ 3.0 - 6.5 mm; several others specimens in shells (USNM 267589). — Bay of Panama, coll. F. H. Bradley, 5 ♂♂ 5.0 - 8.4 mm; 1 ♀ 4.2 mm (USNM 265349). — Taboga, coll. Meek and Hildebrand, 12. V. 1915, 1 ♂ 6.8 mm; 1 ovig. ♀ 5.3 mm; 1 ♀ 5.2 mm (USNM 44192); coll. E. Deichmann, VII. 1924, 1 ♂ 7.0 mm; 1 ♀ 5.3 mm (USNM 1076231); shore, low tide rocks, coll. E. Deichmann, V-VII. 1924, 1 ♂ 7.8 mm; 1 ♀ 3.0 mm (USNM 1076243); 4 specimens in shells (USNM 1076233). — Panama city, coll. and don. “ J. Zetek ”, I. (?) 1914, 5 ♂♂ 3.0 - 7.3 mm (USNM 48794). — “ Panama, Geay 1887, det. Forest 1949 ”, 3 ♂♂ 5.8 - 6.2 mm (MNHN-Pg 804). — “ Panama ”, 1 ♂ 4.6 mm (USNM 1076225); 1 ♂ 8.6 mm (MNHN-Pg 802); 1 ♂ 6.0 mm; 1 ♀ 7.0 mm (MNHN-Pg 803). — Bahia Honda, coll. W. L. Schmitt, stn 111, 9. III. 1933, 1 ♂ 1.9 mm (* USNM 1076242); stn 114, 10. III. 1933, 1 ♂ 1.8 mm (* USNM 1076241). — “ Panama, Cocos Island ”, coll. Pinchot Exp. 1929, 1 ♂ 4.2 mm; 1 ovig. ♀ 4.2 mm (* USNM 63162). Colombia. Port Utria, shore, coll. W. L. Schmitt, stn 418, 23. I. 1935, 2 ♂♂ 4.5 - 5.3 mm; 2 ovig. ♀♀ 2.7 - 4.5 mm; 1 ♀ 2.8 mm (* USNM 1076232). — Octavia Bay, stn 433, 27. I. 1935, 1 ♂ 4.5 mm (* USNM 1076240). Ecuador. Salinas, 14. IX. 1926, 9 ♂♂ 3.8 - 8.3 mm; 2 ovig. ♀♀ 6.5 - 6.8 mm; 3 ♀♀ 2.9 - 5.2 mm (USNM 1076227). Peru. Máncora, intertidal, coll. W. L. Klawe, Inter. Am. Tropic. Tuna Com., 14. VI. 1960, 1 ♂ 5.8 mm; 1 ovig. ♀ 6.4 mm (USNM 106445); Máncora, intertidal among stones, “ W. K. Weyrauch no. 12.636 ”, 23. IV. 1959, 4 ♂♂ 4.4 - 5.7 mm (RMNH 14833). West coast of South America. “ Côtes ouest de l’Amérique du Sud, ancien envoi, Patria ignota, 1905, det. G. Nobili ”, 1 ♂ 9.0 mm; 3 ♀♀ 5.5 - 7.2 mm (* MNHN-Pg 805). DISTRIBUTION. — Eastern Pacific, from El Salvador (La Libertad, 13 ° 29 ’ N) to Peru (Máncora, 4 ° 07 ’ S), including Ecuador, Costa Rica, Panama, Colombia and Gorgona Is. Holthuis (1954: 22) reported a doubtful record (reference not indicated) from South California, probably based on C. californiensis. Lemaitre & Alvarez-León (1992: 46) listed “ California ” in the distribution of C. obscurus, based on Holthuis’ reference. As suggested by Hendrickx & Harvey (1999: 368), this obscure record must be considered erroneous, and the northern limit of C. obscurus more confidently is La Libertad (13 ° 29 ’ N). HABITAT. — Lower intertidal, among rocks, rubble, or algae; commonly found in large numbers (Stimpson 1859; Ball & Haig 1974). DIAGNOSIS Ocular acicle terminating in two to four spines, or rarely one. Anterodorsal plate of branchiostegite smooth, with one or two spinules subproximally. Upper margin of right chela smooth (Fig. 5 C). Outer face of left chela regularly convex (Fig. 5 B). Dactyls of P 2 - P 3 with similar setation consisting of sparse, widely-spaced tufts of setae (Fig. 5 D). Propodus of P 3 with dorsolateral margin weakly angled, upper half of outer face weakly concave. Dactyl of P 3 0.5 - 0.7 times length of propodus, with 6 - 8 spines on ventral margin. Telson with posterior lobes each with single terminal spine. Coloration (Fig. 5 A) In life, shield black, posterior shield gray; ocular acicles orange with broad black band at bases. Ocular peduncles black, with narrow orange band proximally and narrow white band close to cornea. Antennular peduncles black with distal half of terminal segments and flagella orange. Antennal peduncles, antennal acicles, and flagella orange. Chelipeds with meri and carpi black, bordered with orange on upper and anterior margins; chelae black with orange on upper margins of palms and dactyls, tip of dactyls pale orange to white, cutting edges of fingers white. Ambulatory legs uniformly black on meri, carpi and propodi. Dactyls black with bright orange rings varying as follows: with single, narrow ring distally (Fig. 6 A), or with submedian and distal rings (Fig. 6 D), or with distal orange ring and variably developed intermediate submedian ring (Fig. 6 B, C). Abdomen tan with orange hues, sixth abdominal tergite and telson gray, white at margins, uropods white. In preservative, coloration of shield, ocular peduncles and chelae fading quickly to orange. However, coloration of ambulatory legs is still visible after 36 years in some specimens, and color pattern of dactyls is still visible after 100 years in the case of specimens collected during a cruise of the USFC Albatross (USNM 40545). REMARKS Calcinus obscurus is closely allied, and easily confused with C. explorator. The differences between the two species are reported under the latter species. The easiest way to separate the two is by the armature of the ocular acicles, with two to four spines in C. obscurus instead of usually one spine in C. explorator. However, 3 % of C. obscurus specimens examined have only one terminal spine on the ocular acicle, and thus this character should be used with caution to separate the two species. Coloration is overall black and very similar in C. obscurus and C. explorator, differing in minor details of the ocular peduncles, chelae, and dactyls of ambulatory legs (see Remarks under C. explorator). The most obvious color difference is the presence of orange margins on chelae in C. obscurus only (Figs 5 A; 3 C). The geographic distributions of the two are also different, with C. obscurus known from the mainland only from 13 ° 29 ’ N (La Libertad, El Salvador) to 4 ° 07 ’ S (Máncora, Peru), whereas C. explorator is insular (Cocos, Galápagos, Clipperton, Revillagigedo), and has been recorded at higher latitudes on the mainland between 25 ° 55 ’ N (Carmen I., Gulf of California) to 19 ° 16 ’ N (Tenacatita, Mexico).	en	Poupin, Joseph, Bouchard, Jean Marie (2006): The eastern Pacific species of the genus Calcinus Dana, 1851, with description of a new species from Clipperton Island (Decapoda, Anomura, Diogenidae). Zoosystema 28 (2): 465-486, DOI: 10.5281/zenodo.5393881
