taxonID	type	description	language	source
FC9BE9CA6F4951E2ABAF35E5710C1519.taxon	etymology	Etymology. Cicadoforma refers to how people in South America confuse this group with cicadas.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
FC9BE9CA6F4951E2ABAF35E5710C1519.taxon	diagnosis	Diagnosis. Cicadoforma is similar to Cicadomorphus, not only externally, but internally as well; however, the phylogenetic analysis results showed enough evidence to separate Cicadoforma in a different genus from Cicadomorphus (Fig. 1). Nevertheless, both genera can be distinguished morphologically by the forewing, which is hyaline with scales only on margins and veins in Cicadoforma, whereas in Cicadomorphus has scales are more widely distributed on the forewing. Genitalia have some small indentations on outer margin of valva that are not present in Cicadomorphus; apex and lobe on valva is much wider in Cicadoforma; upper side of vesica with one patch of spines, and a narrow band of spines near the base are present in Cicadoforma. Female genitalia with square-shaped anal papillae in Cicadoforma; more rounded in Cicadomorphus. DNA barcodes show a closer relationship with Cicadomorphus (~ 5 % divergent than with Gaujonia (~ 6 %).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
FC9BE9CA6F4951E2ABAF35E5710C1519.taxon	description	Description. Sexually dimorphic mainly in size, female slightly larger than male; forewing in male with some hyaline areas and with poorly developed pattern, whereas female forewing semi-hyaline with pattern better defined. Antenna filiform, black, or dark brown in both sexes; antenna with yellow basal line of scales; eye large, covered by long interfacetal setae; palp with black upper side and yellow underside; haustellum dark brown and reduced, but functional. Forewing with orbicular reniform spots small. Hindwing semi-hyaline with scales only on margin and veins. Male genitalia with simple, lightly-sclerotized valva; valva wide apically with some small indentations on outer margin; clasper absent, cucullus wide, with a broad lobe extended in front of sacculus, ear-like in shape; sacculus with a small foot-like process; uncus crooked, long and wide; aedeagus short, with a simple long vesica with a broad patch of spines on upper side and a narrow band of spines at base. Female genitalia with a large square-shaped and lightly sclerotized sterigma, and a rugose, sclerotized appendix bursae; corpus bursae unsclerotized.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
FC9BE9CA6F4951E2ABAF35E5710C1519.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
A1529BC638E6586EAC5CF4FBD6407849.taxon	etymology	Etymology. The species name ocelotus is derived from the characteristic yellowish orange coloration on its body, reminiscent of the color of an ocelot.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
A1529BC638E6586EAC5CF4FBD6407849.taxon	diagnosis	Diagnosis. Cicadoforma ocelotus can be distinguished from its only congener, C. vau-nigrum, by its remarkably brighter ground color, its larger orbicular spot, and shorter space between forewing veins R 3 and R 4. The valva of the male genitalia of C. ocelotus have concave outer margins, in contrast with the straight margin of C. vau-nigrum.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
A1529BC638E6586EAC5CF4FBD6407849.taxon	description	Description. Head. Palp short with a combination of black, yellow, and orange scales; ground color of frons orange or yellow with scattered black hair-like scales; antenna dark brown. Thorax. Yellow or orange with small spots scattered throughout dorsum. Wings. Forewing length: male 19 - 21 mm; female 25 - 27 mm; forewing yellow or orange scales covering veins and margins with semi-hyaline areas between them; subterminal line slightly visible; basal, antemedial, medial, and postmedial lines poorly developed, present only as small black dots on veins; reniform spot narrow with top and bottom surrounded by black scales and a small black dot in middle; orbicular spot small, elongated; female with antemedial, medial, postmedial, and subterminal lines slightly defined; reniform and orbicular spots similar to those of male; R 4 almost entirely black; black V-shaped marking at base of CuA 2; hindwing semi-hyaline with yellow or orange veins paler than forewing; fringe composed of short yellow or orange hair-like scales except on posterior margin, which has long, pale-yellow scales. Leg. Yellow or orange with some irregular brown spots that decrease in number from prothoracic legs to metathoracic legs. Abdomen. Covered by brownish orange scales that are paler than those of thorax; brown tufts in A 2 - A 7 with tips with same color as remainder of abdomen. Male genitalia. Cucullus wide, lobe narrow; lobe apex rounded; apex clothed with short setae that expand over entire costal margin; sacculus base narrow; saccus relatively long, V-shaped; tegumen fairly flat; juxta U-shaped on upper side; aedeagus 3 3 / 4 x longer than wide with opening to vesica same width as aedeagus; vesica 1 1 / 2 x longer than aedeagus with a patch of spines on upper side with some conspicuous basal spines. Female genitalia. Anal papilla wide, petal shaped, clothed with relatively long setae; A 8 membranous, relatively short; posterior apophysis almost same length as anal papilla; sterigma enlarged, lightly sclerotized, fused above ostium; anterior apophysis ⅞ x shorter than posterior apophyses; ductus bursae short, strongly sclerotized and wide; posterior 3 / 4 of appendix bursae strongly sclerotized, remainder including corpus bursae unsclerotized; corpus bursae 1 1 / 4 x longer than the appendix bursae.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
A1529BC638E6586EAC5CF4FBD6407849.taxon	distribution	Distribution. This species is only known from northern Colombia from moderate elevations from 2000 - 3000 m, especially in cloud forests (Fig. 92).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
A1529BC638E6586EAC5CF4FBD6407849.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
30BBF736A0925DA4A95BEF2BFB8BBFA3.taxon	etymology	Etymology. George F. Hampson likely named this species " vau-nigrum " because of the black V-shaped mark at the base of CuA 2.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
30BBF736A0925DA4A95BEF2BFB8BBFA3.taxon	diagnosis	Diagnosis. Cicadoforma vau-nigrum is the largest species in this genus; males have a forewing length of ca. 24 - 26 mm, whereas that of females is ca. 30 - 32 mm. This species is similar to C. ocelotus, but easy to distinguish by the size, and also by the orbicular spot, which is larger in C. vau-nigrum. Another important difference is that C. vau-nigrum males have hyaline wings where only the veins and margins are covered with yellow and black scales; the abdomen has dark brown tufts in the middle of A 4 - A 7. Cicadoforma vau-nigrum can be differentiated from other species by the male genitalia; the vesica has two patches of spines, one near to the aedeagus and the other on the upper side of the vesica. Female genitalia have heavily sclerotized ductus bursae and appendix bursae, with the corpus bursae ca. 2 x longer than the appendix bursae and 11 / 2 x wider; the anterior apophyses are remarkably short, whereas the posterior apophyses are almost as long as the valva.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
30BBF736A0925DA4A95BEF2BFB8BBFA3.taxon	distribution	Distribution. Cicadoforma vau-nigrum is endemic to Venezuela, known only from the states of Vargas and Aragua at moderate elevation in the northern part of the northwestern Andes (Fig. 92).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
30BBF736A0925DA4A95BEF2BFB8BBFA3.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
79E999A1528A50C293CAF1F65B70ECF1.taxon	etymology	Etymology. The word chicharra means cicada in the Quechua language.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
79E999A1528A50C293CAF1F65B70ECF1.taxon	diagnosis	Diagnosis. Cicadomorphus chicharra is one of the most easily identified species in the group due to its whitish yellow coloration, but also the wing pattern is thinner and paler than in related species, such as C. lilianae. The male genitalia have a wider vesica and narrower spine band on the upper side. Additionally, barcoding showed 2 % divergence from C. lilianae.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
79E999A1528A50C293CAF1F65B70ECF1.taxon	description	Description. Head. Palp with last segment black with a small white dot; frons with greenish yellow scales; antenna brownish orange. Thorax. Greenish yellow, with small black dots dorsally; collar with margins black. Wings. Forewing length: male 23 - 25 mm; forewing greenish yellow, semi-hyaline areas clothed by fewer greenish yellow scales; subterminal, medial, and antemedial lines slightly visible; reniform spot poorly defined; orbicular spot long; hindwing: semi-hyaline with greenish yellow veins and fringe. Leg. Prothoracic legs black with some patches of same color as body; mesothoracic legs and metathoracic legs greenish yellow. Abdomen. Greenish yellow, paler than thorax; tufts in male on A 1 - A 6 with characteristic small black spots on A 1, A 5 and A 6. Male genitalia. Cucullus wide with a wide lobe; apex, outer margin, and lobe covered by long setae; apex slightly squared; sacculus wide with needle-shaped process; tegumen narrow, unsclerotized around uncus; saccus relatively wide; juxta with U-shaped concave depression on upper side; aedeagus ⅔ x as long as vesica; opening to vesica as wide as aedeagus; vesica wide with narrow transverse band of spines.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
79E999A1528A50C293CAF1F65B70ECF1.taxon	distribution	Distribution. The three specimens were found in the western zone of Bolivia at moderate elevations ca. 1500 m (Fig. 92).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
79E999A1528A50C293CAF1F65B70ECF1.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
B32B3358D4A958C8B8752757B7002B27.taxon	etymology	Etymology. Ch'uya means pure, transparent, or glassy in the Quechua language.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
B32B3358D4A958C8B8752757B7002B27.taxon	diagnosis	Diagnosis. Although Cicadomorphus chuya shares some characters with C. chicharra, it is easy to distinguish by wing pattern; C. chuya is the only species that has both the terminal and subterminal lines serrated. In addition, the genitalia have the lobe reduced, and the juxta is pentagonal-shaped. The base of the lobe opens externally diagonally from the tornus to the cucullus.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
B32B3358D4A958C8B8752757B7002B27.taxon	description	Description. Head. Third segment of palp divided with upper side black, underside white, but white areas with a few brown spots; antenna dark brown basally, distal segments paler. Thorax. Whitish yellow coated with small black dots dorsally. Wings. Pattern in both sexes well defined and visible; forewing length: male 22 - 24 mm; female 29 - 31 mm; forewing whitish yellow with pattern somewhat blurred; nearly all lines visible and slightly blurred with exception of postmedial line, which is formed by black dots on wing veins plus some dispersed scales; subterminal and terminal lines zigzag; reniform spot relatively wide, outline poorly defined; orbicular spot medium sized and elongate; hindwing hyaline with yellow scales on fringe paler than thorax; wing veins darkened. Leg. Prothoracic and mesothoracic legs whitish yellow with two patches of brown scales on femur. Abdomen. Dorsally gray, black, and whitish yellow tufts in middle segment with A 1 and A 3 whitish yellow; whitish yellow ventrally. Male genitalia. Cucullar region wider close to apex; lobe small, almost completely covered by long setae from middle of costal margin to lobe; tooth-like protuberance barely visible; lobe diagonal to tornus of valva; saccular region relatively short with tip of process just touching tooth-like protuberance externally; saccus thin, rhomboid shaped; juxta pentagonal shaped; tegumen narrow with a small hood on base of uncus; aedeagus short and wide; opening to vesica square shaped ventrally; vesica tapered, with a large band of spines. Female genitalia. Anal papilla square shaped with short setae; posterior apophysis almost same size as anal papilla; anterior apophysis short; sterigma V-shaped, lightly sclerotized, above ostium; ductus bursae wide posteriorly, narrow toward appendix bursae; appendix bursae 1 / 4 x shorter than corpus bursae; corpus bursae semi-transparent.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
B32B3358D4A958C8B8752757B7002B27.taxon	distribution	Distribution. Males were found in Colombia and Peru, whereas the female was found in Bolivia, all in deciduous forest habitats at moderate to high elevations from 1800 - 3000 m (Fig. 92).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
B32B3358D4A958C8B8752757B7002B27.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
3AB0F17E52EB5263B50F9629138DC35E.taxon	etymology	Etymology. Falkasiska is the combination of the names from Falk Zahlaus who collect the first female specimen and Toni Kasiske, who reared the eggs obtained from that female.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
3AB0F17E52EB5263B50F9629138DC35E.taxon	diagnosis	Diagnosis. Cicadomorphus falkasiska and C. chicharra share certain characters, especially in coloration and wing pattern. However, C. falkasiska is paler and the orbicular spot larger and outlined in black; also, the subterminal line is quite visible in C. chicharra, whereas in C. falkasiska it is inconspicuous. The male genitalia have a relatively wide aedeagus; the vesica is large and semi-rounded; the female genitalia have a wide appendix bursae and corpus bursae.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
3AB0F17E52EB5263B50F9629138DC35E.taxon	description	Description. Head. Third segment of palp in both sexes black with a pale-yellow stripe ventrally and a dot of same color dorsally; frons darker-yellow than rest of body; antenna dark brown. Thorax. Pale yellow with some black blurred spots, which are more visible in male than female. Wings. Forewing length: male 23 - 25 mm; female 30 - 32 mm; forewing pale yellow; hyaline areas in male nearly without scales, whereas forewing slightly more covered with scales in female; postmedial and subterminal lines inconspicuous in male, whereas basal, antemedial, and medial lines well defined; female with all lines weakly defined; orbicular spot large and elongate; reniform spot slightly wide, outlined with black scales and with a black dot in middle; hindwing hyaline in male, semi-hyaline in female with fringe paler yellow than forewing. Leg. Pale yellow with prothoracic legs with some brown patches. Abdomen. Whitish gray with some tufts over middle of abdomen; tufts on A 1 - A 3 yellow with a small black dot on A 2, whereas remainder tufts on other segments are black with some yellow and white scales; female with yellow tufts on A 1 - A 4. Male genitalia. Cucullar region relatively wide with an axe-shaped lobe; apex round with a lobe and apex covered with quite long setae; saccular region wide with process of sacculus thin and curved; juxta flat on upper side but with a small V-shaped depression in middle; aedeagus relatively wide at opening to vesica; aedeagus ca. as long as vesica; vesica rounded with narrow transverse band of spines. Female genitalia. Anal papilla long and wide with posterior apophysis ca. as long as anal papilla; anterior apophysis short; sterigma large, fused above ostium; ductus bursae wide and short; and appendix bursae large, well sclerotized; corpus bursae not sclerotized 11 / 4 x longer than appendix bursae.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
3AB0F17E52EB5263B50F9629138DC35E.taxon	distribution	Distribution. All specimens were found in central Peru at a high altitude of ca. 2500 m or above (Fig. 92).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
3AB0F17E52EB5263B50F9629138DC35E.taxon	biology_ecology	Biology. Cicadomorphus falkasiska is the only species in the genus for which immature stages are known. Adults are active throughout the year, but especially so in spring and summer. Cicadomorphus falkasiska feeds on Prunus spp. especially on P. subcorymbosa Ruiz ex Koehne (JIM, pers. obs.).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
0BA64973144F54879A3162105E8A3C1E.taxon	etymology	Etymology. This species is named in honor of my sister Lilian Martinez Canto (1989 - 2017) for her love, charisma, and support offered during all her beautiful life.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
0BA64973144F54879A3162105E8A3C1E.taxon	diagnosis	Diagnosis. Cicadomorphus lilianae is small and the wing pattern is blurred, causing the lines to appear shapeless. The males have darker coloration than the females, and the line pattern is better defined. This species shares some features in genitalia with C. chuya, such as the tapered vesica.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
0BA64973144F54879A3162105E8A3C1E.taxon	description	Description. Head. Palp black, terminal segment admixed with brown and white scales; frons dark yellow basally with some black scales; female ground color pale yellow with gray scales; antenna brownish orange. Thorax. Ground color dark yellow with some black tufts; female same color but with gray tufts; collar with black ground color with margins yellow, female with ground color gray. Wings. Both sexes dark yellow, similar in pattern, nevertheless female is paler, and wing pattern is basally almost imperceptible; forewing length: male 25 - 27 mm; female 31 - 33 mm; forewing yellow with pattern in black, but blurrier, male darker; both sexes with semi-hyaline areas with some scattered dark yellow scales; both sexes with blurry lines; orbicular spot elongate, ca. same size in both sexes; reniform spot heavily outlined with black scales; hindwing hyaline with fringe yellow, but gray on posterior margin. Legs. Yellow, except prothoracic legs, which are brown with some black scales. Abdomen. Gray with yellow tufts in middle of abdomen and on each side; brown tufts on A 3 - A 7; female abdomen yellow with some brown scales. Male genitalia. Cucullus wide with apex wide; costal margin without setae; outer margin sharply bent; tooth-like protuberance small; lobe with external apex large; posterior margin of lobe curved; sacculus wide with narrow process touching tooth-like protuberance; saccus thin; tegumen narrow, lightly sclerotized, barely visible around uncus; juxta wide, upper side flat; aedeagus relatively wide, V-shaped in opening; tapered vesica; band of spines with a rounded ending. Female genitalia. Anal papilla small; relatively wide posterior apophysis 1 ⅓ x longer than anal papilla; sterigma open trapezoid shaped; anterior apophysis short; ductus bursae small, well-sclerotized; appendix bursae broad, sclerotized; corpus bursae ⅓ x longer than appendix bursae.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
0BA64973144F54879A3162105E8A3C1E.taxon	distribution	Distribution. This species has been found in Ecuador and Colombia at very variable elevations from 1100 - 2700 m (Fig. 92).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
0BA64973144F54879A3162105E8A3C1E.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
BAD2CC02668F547B995785234CB2CFFA.taxon	etymology	Etymology. Cicadomorphus refers to how similar is this genus to Cicadoforma.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
BAD2CC02668F547B995785234CB2CFFA.taxon	diagnosis	Diagnosis. Cicadomorphus is closed related to Cicadoforma genetically and morphologically (see Cicadoforma diagnosis).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
BAD2CC02668F547B995785234CB2CFFA.taxon	description	Description. Sexually dimorphic in size, female slightly larger than male; cells of forewing with some semi-hyaline areas in male, whereas in female forewing covered by scales; line pattern conspicuous in both sexes. Antenna in both sexes dark brown, filiform with a stripe of sulfur-yellow scales basally; haustellum functional but reduced. Forewing with orbicular spot varying in size, reniform spot narrow. Hindwing semi-hyaline presenting scales only on margins and veins. Male genitalia with simple valva, lightly sclerotized lacking clasper; cucullar region with apex and ear-shaped lobe extremely narrow; saccular region with a long and narrow harpoon-shaped process; uncus hooked, broad and long; aedeagus short with simple vesica with a narrow patch of spines on upper side. Female genitalia with sterigma large, rounded, lightly sclerotized; appendix bursae rugose and sclerotized; corpus bursae transparent, not sclerotized.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
BAD2CC02668F547B995785234CB2CFFA.taxon	biology_ecology	Biology. Currently known only for Cicadomorphus falkasiska (see C. falkasiska diagnosis).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
A20BC6C54A355DBCAFFA8B36E804FB06.taxon	etymology	Etymology. Paul Dognin probably named this species arbosi after Fr. Mariano Arbos, a friend of Fr. Theophile Gaujon.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
A20BC6C54A355DBCAFFA8B36E804FB06.taxon	diagnosis	Diagnosis. Gaujonia arbosi is similar to G. chiqyaq, but it can be identified by coloration, which is dark yellow in G. arbosi. Both sexes are similar with the only difference being that the female is covered with brighter yellow scales from the fold to the posterior margin of the forewing, accenting the lines. For males the forewing length ranges from 17 - 19 mm and for females from 23 - 25 mm. Palp short and black; antenna has a stripe of dark yellow scales from the base to the seventh antennomere; antennae are black, and longer than in G. chiqyaq. Male thorax dark yellow with some patches of black. Forewing with pattern similar to that of G. chiqyaq, but the lines on Sc + R 1 and posterior margin are thicker; there is a small dot in the middle of the base of the cell M 1, which is not present in G. chiqyaq; the V-shaped mark at the base of CuA 2 smaller and thicker than in G. chiqyaq. The hindwing, base of M 2 is angled diagonally forward to the base of the wing, fused to the base of CuA 1; in G. chiqyaq M 2 is squared and not fused with the base of CuA 1; the fringe is black with few yellow scales making minute spots at the end of the veins, except the cell A 2, which is completely outlined in black. The male genitalia have the cucullar region wide, slightly diagonal to the base, whereas the cucullar region in G. chiqyaq is wide, opened, almost touching the saccular region; the juxta is semicircular in shape; the saccus is short and wide. Female genitalia, the sterigma is peanut shaped, and the corpus bursae is ⅛ x larger than the appendix bursae. DNA Barcoding showed that G. arbosi and G. chiqyaq are sister species with ca. ~ 0.3 % of difference, but the morphological characters mentioned above distinguish both species.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
A20BC6C54A355DBCAFFA8B36E804FB06.taxon	distribution	Distribution. This species is endemic to coniferous and deciduous forest of Ecuador and Colombia and can be found at high elevations above 2500 m (Fig. 93).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
A20BC6C54A355DBCAFFA8B36E804FB06.taxon	biology_ecology	Biology. Guevara and Romero (2008) were the first to record the larval stages in Gaujonia arbosi feeding on Alnus acuminata Kunth but they misidentified it as Gaujonia arbosioides (now Millerana arbosioides), even though the immature stages differ greatly between both genera (see Millerana immature stages). Additionally, the illustrations from Guevara and Romero (2008) lack of good quality to use for identification.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F46C233F0A1C59B3B6BFBDE35919E693.taxon	etymology	Etymology. The term bichu is used to refer to an insect or small animal in the Quechua language. Since this is the smallest species of Gaujonia, this name seems appropriate.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F46C233F0A1C59B3B6BFBDE35919E693.taxon	diagnosis	Diagnosis. Gaujonia bichu is closely related to G. sourakovi, but G. bichu is the smaller. The male genitalia have the valva and the saccular region narrow, and the process of the sacculus is longer and thinner.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F46C233F0A1C59B3B6BFBDE35919E693.taxon	description	Description. Head. Palp black; frons yellow with some black scales. Thorax. Dorsally covered by large black patches. Wings. Male yellow with thin spots and lines on venation; forewing length: male 15 - 17 mm; forewing with two V-shaped marks on base of R 1 + R 2 and R 4 + R 5; base of cell with small black dot on M 1; five black dots on Sc + R 1 with a line right on apex; half of posterior margin black; hindwing with fringe black, interrupted by a yellow dot at end of each vein. Leg. Prothoracic leg black with femur and tarsi presenting yellow spots; mesothoracic legs yellow with some black spots; metathoracic legs yellow. Abdomen. Grayish yellow with yellow and black tufts in middle of abdomen dorsally on A 1 - A 5, remainder of abdomen covered in yellow scales. Male genitalia. Valva long and narrow a little swollen from middle to apex externally; sacculus narrow; sacculus process long and narrow; juxta shield-like; tegumen wide, narrower near valva; aedeagus short and narrow; vesica elongated with two oval patches, one 2 x longer than the other.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F46C233F0A1C59B3B6BFBDE35919E693.taxon	distribution	Distribution. Gaujonia bichu specimens were found in Ecuador in coniferous and deciduous forests at high elevations above 2500 m (Fig. 93).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F46C233F0A1C59B3B6BFBDE35919E693.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
6DCEF0E953A5508BA1344103BF5EE01F.taxon	etymology	Etymology. The name chiqyaq means green in the Quechua language, referencing the diagnostic green coloration of this species.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
6DCEF0E953A5508BA1344103BF5EE01F.taxon	diagnosis	Diagnosis. Beside the features mentioned above to differentiate Gaujonia chiqyaq from G. arbosi, there are also other informative characters to identify this species. The male has a narrow reniform spot outlined by black scales along the upper and lower sides, but not in the middle region. The orbicular spot is small and greenish yellow, outlined in black. The thorax has large black tufts. The female has a larger reniform spot with the same yellow coloration. Forewing transverse lines are more sharply defined and thicker than in G. arbosi.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
6DCEF0E953A5508BA1344103BF5EE01F.taxon	description	Description. Head. Palp with last segment black; frons with a combination of black and greenish yellow scales. Thorax. Covered with long black tufts. Wings. Both sexes greenish yellow with thin spots and lines; forewing length: male: 20 - 22 mm; female: 28 - 30 mm; forewing ground color greenish yellow with small, thin lines on venation forming pattern, female with thicker and well-defined lines; orbicular spot in male small, black, but almost imperceptible; orbicular spot larger in female and elongated; reniform spot in both sexes narrow, greenish yellow, outlined in black; both sexes with hindwing venation green, and fringe black with some greenish yellow dots. Leg. Metathoracic and metathoracic legs black with some greenish yellow patches; hind legs greenish yellow with black shading. Abdomen. Brown dorsally with green tufts on A 1 - A 3 and some black scales in middle of the green tufts; ventrally greenish yellow; female similar to male but paler in color. Male genitalia. Cucullar region extended close to saccular region, valva relatively narrow, slightly ridged externally on apex; saccular region rounded at base and process wide; saccus wide V-shaped; juxta rectangular concave on top; tegumen wide; aedeagus short 3 x longer than wide; basal area almost same size as apical part of vesica with two patches of spines, one 2 x narrower than the other. Female genitalia. Anal papilla wide, slightly ridged; posterior apophysis ⅓ x longer than anal papilla; sterigma large oval shaped, fused above ostium; anterior apophysis short; ductus bursae heavily sclerotized and connected to appendix bursae, which is sclerotized as well; appendix bursae 1 / 2 x narrower than corpus bursae.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
6DCEF0E953A5508BA1344103BF5EE01F.taxon	distribution	Distribution. This species has a broad distribution from Ecuador to Peru between 2100 - 3200 m (Fig. 93).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
6DCEF0E953A5508BA1344103BF5EE01F.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
04D4F827BEFA5B008074585E7A984459.taxon	etymology	Etymology. Paul Dognin probably named this genus Gaujonia after Fr. Theophile Gaujon a Catholic Lazarist missionary priest and entomologist residing in Loja, Ecuador, who helped by collecting specimens for him.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
04D4F827BEFA5B008074585E7A984459.taxon	diagnosis	Diagnosis. Gaujonia is similar to Oculicattus morphologically, but it is most closely aligned genetically with Cicadoforma and Cicadomorphus (Fig. 1). The interfacetal setae of Gaujonia are longer than those in Cicadoforma and Cicadomorphus. Male genitalia are similar to those of Oculicattus, however, the cucullus is wider and slightly square in shape, whereas in Oculicattus the cucullus is thinner and rounded. Gaujonia has a short vesica and two patches of spines, whereas in Oculicattus the vesica is longer with three patches of spines. Female genitalia of Gaujonia are similar to that of Oculicattus, but that of Oculicattus is larger in size, however, the sterigma is smaller in that genus than in Gaujonia. DNA barcoding corroborated that Gaujonia is closer to Cicadoforma and Cicadomorphus (~ 6 % divergent) than Oculicattus (~ 8 %).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
04D4F827BEFA5B008074585E7A984459.taxon	description	Description. Sexually dimorphic in size, female larger than males; forewing and hindwing hyaline with scales only on wing veins; orbicular and reniform spots in male inconspicuous or absent, whereas female has an orbicular spot and narrow reniform spot. Antenna filiform and long haustellum dark brown; eyes hairy with long interfacetal setae. Male genitalia with saccular and cucullar regions separated and clasper absent; short-beaked uncus; aedeagus wide and vesica short with two sclerotized patches of spines on each side. Female genitalia with sterigma open wide, ductus bursae and appendix bursae are heavily sclerotized, whereas corpus bursae is not.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
04D4F827BEFA5B008074585E7A984459.taxon	biology_ecology	Biology. The biology is known only from one species in Gaujonia (see G. kanakusika immature stages) (JIM, pers. obs.).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F5BE7255BB81583EBF5660D37140F969.taxon	etymology	Etymology. The name is formed from the Quechua words kanaku (fire) and sika (caterpillar), based on the immature stages of this species (Fig. 91 B).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F5BE7255BB81583EBF5660D37140F969.taxon	diagnosis	Diagnosis. This species can be differentiated from other species by the sulfur-yellow coloration of most of the body, including the abdomen.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F5BE7255BB81583EBF5660D37140F969.taxon	description	Description. Head. Palp with basal and second palpal segment sulfur-yellow, last segment with white tip; frons with small black line; female with similar wing pattern, but brighter in coloration. Thorax. Sulfur-yellow, covered with small black dots. Wings. Forewing sulfur-yellow; length: male 18 - 20 mm; female 27 - 29 mm; forewing with lines visible from posterior margin to fold; basal and antemedial lines almost completely developed; no presence of a V-shaped mark; Hind wing fringe same yellow color as body. Legs. Prothoracic and mesothoracic legs black with some sulfur-yellow spots on joints; tarsal scales yellow and black. Abdomen. Sulfur-yellow with a small area of gray in middle of dorsum divided by black and yellow tufts. Male genitalia. Cucullus large, rounded apically; sacculus narrow with a long process; juxta shell-shaped with apices pointed; tegumen wide; saccus narrow; aedeagus short and wide; vesica 1 1 / 2 x as long as aedeagus with two large spine patches ca. same size. Female genitalia. Anal papilla small, semicircular; posterior apophysis ⅓ x longer than anal papilla; sterigma narrow and large, semi-rectangular; anterior apophysis relatively long and heavily sclerotized; corpus bursae 1 1 / 2 x longer than appendix bursae.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F5BE7255BB81583EBF5660D37140F969.taxon	distribution	Distribution. All specimens were found in deciduous forests from the central-north Colombia between 2900 - 3300 m (Fig. 93).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F5BE7255BB81583EBF5660D37140F969.taxon	biology_ecology	Biology. Gaujonia kanakusika is the second species in this genus known to feed on Alnus acuminata Kunth. Adults fly year-round, mainly in summer (JIM, pers. obs.).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
D7E01EB215015EAC93E9A8D818451487.taxon	etymology	Etymology. This species was found by my colleague and friend Andrei Sourakov when he was helping me locate additional specimens deposited in the MGCL, and thus I named it in his honor for his wonderful help.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
D7E01EB215015EAC93E9A8D818451487.taxon	diagnosis	Diagnosis. Gaujonia sourakovi differs from other species mainly in coloration, having the most lightly marked forewing in this group. The male genitalia have the apex of the cucullus rounded, the saccular region is wide, and two patches of long hair-like spines on the vesica, differing from the others which have wider and well-developed spines.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
D7E01EB215015EAC93E9A8D818451487.taxon	description	Description. Head. Last segment of palp black with some yellow scales; frons yellow, shaded with black scales. Thorax. Dark yellow with black dorsally. Wings. Forewing length: male 19 - 21 mm; forewing scales only on venation and margins; V-shaped mark at base of CuA 2 widely open; dot in the middle of base of cell M 1 elongated; orbicular spot small; veins lined with black; hindwing fringe black with dark yellow scales at end of each vein, whereas area on posterior margin of hindwing brown; a black line at base of vein M 3. Leg. Prothoracic and mesothoracic legs dark brown with yellow scales on joints, whereas metathoracic legs dark yellow with some dark brown spots. Abdomen. Brown with scattered black scales dorsally, paler ventrally; long yellow scales laterally; black tufts with some yellow and brown scales on A 1 - A 4. Male genitalia. Cucullar area narrow and curved with apex rounded; saccular region and process wide; juxta flat on upper side; tegumen wide; aedeagus 2 ⅔ x longer than wide; basal area of vesica 4 x longer than wide and vesica almost same width with two large patches of hair-like spines, one ovoid.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
D7E01EB215015EAC93E9A8D818451487.taxon	distribution	Distribution. The species Gaujonia sourakovi was collected in a deciduous forest at 1800 m elevation in southeastern Peru (Fig. 93).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
D7E01EB215015EAC93E9A8D818451487.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
3E47E8A82A805EBC902EB2FF45A8BD1D.taxon	etymology	Etymology. Amsa is a Quechuan word that means dark, opaque, or confused.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
3E47E8A82A805EBC902EB2FF45A8BD1D.taxon	diagnosis	Diagnosis. The species Gaujoptera amsa has similar morphological characters with species of Millerana, however, it is easy to identify because the thorax has no spots or patches, but is completely marbled with black, gray, brown, and sulfur-yellow scales. The forewing is dark sulfur-yellow with a blurry pattern in gray. Additionally, the genitalia have a simple valva without any lobe or protuberances, and a small clasper is present.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
3E47E8A82A805EBC902EB2FF45A8BD1D.taxon	description	Description. Head. Palp with last segment black with few yellow scales on tip; frons marbled. Thorax. Marbled in black, gray, brown, and sulfur-yellow. Wings. Forewing length: male 18 - 20 mm; forewing marbled with wing pattern in gray; pattern blurry; orbicular spot barely visible in yellow; a narrow lunate marking on reniform spot; terminal lines zigzag; hindwing hyaline with sulfur-yellow fringe. Leg. Prothoracic and mesothoracic legs sulfur-yellow with some brown spots; tarsi brown and yellow; metathoracic legs marbled in brown and yellow with tarsi sulfur-yellow. Abdomen. Sulfur-yellow with a black line of tufts in middle of abdomen on A 1 - A 5. Male genitalia. Valva rectangular, simple; saccular region wide; apex densely covered by setae; clasper small, triangular; juxta shell-like; tegumen narrow; uncus small; aedeagus short, almost same width as vesica; vesica ovoid with a line of minute spines in middle, positioned diagonally.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
3E47E8A82A805EBC902EB2FF45A8BD1D.taxon	distribution	Distribution. Gaujoptera amsa occurs from central Peru to northern Bolivia at high elevations ca. 3000 m or above (Fig. 93).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
3E47E8A82A805EBC902EB2FF45A8BD1D.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
7477128CB284571CA57C6ED98C41D09A.taxon	etymology	Etymology. Gaujoptera is derived from the similarities with the genera Gaujonia and Lichnoptera.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
7477128CB284571CA57C6ED98C41D09A.taxon	diagnosis	Diagnosis. Gaujoptera is only known from two well-preserved male specimens and it has a close relationship with Millerana, not only genetically, but also by distinct morphological characters. Nevertheless, the wing pattern is more defined in Millerana than in Gaujoptera, which has a blurry pattern that is difficult to discern. In addition, the thorax is marbled with black, gray, brown, and sulfur-yellow scales, differing from species of Gaujonia, which have the black spots and patches well defined on the thorax. Gaujoptera does not have a V-shaped mark at the base of CuA 2. The most important feature to differentiate Gaujoptera from Gaujonia is the presence of small clasper on the valva, similar to that of Lichnoptera.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
7477128CB284571CA57C6ED98C41D09A.taxon	description	Description. Forewing and hindwing dark sulfur-yellow with an inconspicuous pattern, only reniform spot visible. Hindwing hyaline with scales only on margins. Antenna filiform, with a short brownish orange haustellum; eye hairy with short interfacial setae. Male genitalia simple in male with small triangular clasper; small, curved uncus; aedeagus shorter than vesica; vesica with small spines in middle. Female unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
7477128CB284571CA57C6ED98C41D09A.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
007255D1ABAB5C758C709E8F3DCC0A37.taxon	etymology	Etymology. Paul Dognin likely name this species arbosioides in reference to its similarities to Gaujonia arbosi, which was described previously.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
007255D1ABAB5C758C709E8F3DCC0A37.taxon	diagnosis	Diagnosis. Millerana arbosioides is closely related to three species: M. austini, M. cajas, and M. cundinamarquensis; however, M. arbosioides can be recognized by the black antenna and haustellum. Forewing length in males is 16 - 18 mm. Millerana arbosioides shares more external characters with M. cundinamarquensis, but it can be distinguished from it by the antemedial line, which is well developed in M. arbosioides, the reniform spot is better defined, and the lunate marking is small; these features in M. cundinamarquensis are barely visible, but it presents four black tufts in the middle of 1 A- 4 A, whereas M. arbosioides has three on A 1 - 3 A. The genitalia in M. arbosioides have a semi-squared cucullar area and some protuberances in the outer margin that are present in M. tigrina and M. cajas as well however, both species differ from M. arbosioides by their apexes that are rounded and their outer margins of the cucullar area, which are straighter.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
007255D1ABAB5C758C709E8F3DCC0A37.taxon	distribution	Distribution. The species is known only from coniferous forests in Ecuador at very high elevations ca. 2500 - 3500 m (Fig. 94).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
007255D1ABAB5C758C709E8F3DCC0A37.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
A3B3851B3FEF5986AAC7A74D44BE7CE4.taxon	etymology	Etymology. The species was named after my friend Kyhl Austin, an American lepidopterist, who found this species in the dark corners of the CUIC.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
A3B3851B3FEF5986AAC7A74D44BE7CE4.taxon	diagnosis	Diagnosis. Millerana austini is similar to M. cajas, but it can be differentiated by the antemedial line, which is formed by a single line, whereas M. cajas has two. The male genitalia are without protuberances on the outer margin, except for two extensions; one near the apex and the other on the lobe; they are similar to those of M. tigrina; however, the outer margin of the valve is concave in M. tigrina.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
A3B3851B3FEF5986AAC7A74D44BE7CE4.taxon	description	Description. Head. Palp black with a small line with pale yellow underneath; frons yellow. Thorax. Dark yellow with dots in black. Wings. Forewing dark yellow with black pattern; length 17 - 19 mm. All lines well developed with exception of terminal line; orbicular spot circular; reniform spot incomplete, with a small lunate marking; V-shaped mark on base of CuA 2 thick, black; hind wing whitish orange, veins yellowish orange. Leg. Prothoracic legs black with some yellow spots on joints; mesothoracic legs with some black dots on tibia; metathoracic legs yellow. Abdomen. Paler yellow than thorax; small tufts in black from 1 A- 2 A. Male genitalia. Cucullar area wide; apex and lobe with small extensions; outer margin flat; sacculus notably wide; tegumen broad; juxta inverted, triangle-shaped; aedeagus 3 ⅓ x longer than wide; vesica 1 1 / 2 x wider than aedeagus, gradually reduced at tip; band with large spines and thick patch of spines near apex.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
A3B3851B3FEF5986AAC7A74D44BE7CE4.taxon	distribution	Distribution. All specimens were found in a cloud forest at the north-central region of Ecuador at high elevations above 3500 m (Fig. 94).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
A3B3851B3FEF5986AAC7A74D44BE7CE4.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
823F30C1D06E5F53BF65CF12EC4D9CE2.taxon	etymology	Etymology. The only two specimens found in El Cajas National Park, Ecuador, are known.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
823F30C1D06E5F53BF65CF12EC4D9CE2.taxon	diagnosis	Diagnosis. Millerana cajas is similar to M. austini. The best way to identify M. cajas externally is to examine the medial and postmedial lines, which are fused on the forewing. Moreover, the genitalia are very distinctive. The valva is enlarged, especially in the costal region; Millerana cajas has a wide apical protuberance on the valve, similar to that of M. matthewsae; the vesica has a distal diverticulum surrounded by a spine band.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
823F30C1D06E5F53BF65CF12EC4D9CE2.taxon	description	Description. Head. Palp short, black and yellow, except last segment that is black with some small yellow scales; frons yellow. Thorax. Covered with light lemon-yellow hair-like scales with some black spots. Wing. Light lemon yellow with black and gray pattern; forewing length: male 18 - 20 mm; forewing antemedial, medial, and subterminal lines well developed; whereas basal, postmedial, and terminal lines defined by small dots; medial line wide; orbicular spot slightly flattened; reniform spot large, incomplete, with lunate marking at base of M 1 cell; V-shaped mark at base of CuA 2 narrow; medial and postmedial lines fused by black scales between V-shaped mark in CuA 2 and fold; hindwing whitish yellow with veins whitish orange. Leg. Prothoracic legs black with fuscous joints; mesothoracic and metathoracic legs yellow with a black spot on tibia; tarsi black. Abdomen. Paler yellow than remainder of body; tuft on middle area on A 1 - A 5; A 1 - A 3 black and A 4 - A 5 yellow. Male genitalia. Cucullus wide, mainly in on costal area; apex with a wide extension followed by some protuberances on outer margin; saccular region wide; tegumen wide; juxta dentate, slightly curved on upper side; aedeagus 3 1 / 2 x longer than wide; opening larger than rest of aedeagus; vesica long with a distal diverticulum, which is surrounded by a band of long spines; medial area with a large patch of minute spines.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
823F30C1D06E5F53BF65CF12EC4D9CE2.taxon	distribution	Distribution. This species was found in a deciduous forest at a high elevation in southwestern Ecuador ca. 3600 m (Fig. 94).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
823F30C1D06E5F53BF65CF12EC4D9CE2.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
93F5844AE5B551A7A4372746F1C51789.taxon	etymology	Etymology. The name is derived from the place (Department of Cundinamarca in Colombia) where this species was found.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
93F5844AE5B551A7A4372746F1C51789.taxon	diagnosis	Diagnosis. Compared with other species in Millerana, M. cundinamarquensis differs in the genitalia, in which the valva is petal-like, whereas the other two species have a rectangular valva.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
93F5844AE5B551A7A4372746F1C51789.taxon	description	Description. Head. Palpi short, divided by black hair-like scales on upper side and yellow underneath; last segment coated with a mix of black, white, and yellow scales; frons with sulfur-yellow scales. Thorax. Covered by sulfur-yellow scales with some large black spots. Wing. Forewing length: male 16 - 18 mm; forewing sulfur-yellow with black scales defining transverse lines; antemedial, medial, and subterminal lines barely developed; basal, postmedial, and terminal lines defined by series of dots on veins; orbicular spot well-defined, oval; reniform spot blurry, with a small triangular lunate marking; CuA 2 with a black V-shaped mark at base, with inferior line longer than superior line; hindwing with yellowish orange veins; fringe with pale-yellow hair-like scales; base of M 2 + M 3 rounded. Legs. Yellow except prothoracic legs, which are black with pale yellow spots. Abdomen. Pale yellow with light-gray hair-like scales; four black tufts in the middle of A 1 - A 4; a small pale-yellow tuft on A 8 at terminus. Male genitalia. Cucullar area with rounded apex, apex extension claw-like; costa ovoid, covered with setae; saccular area wide; tegumen narrow and rounded; juxta with concave V-shaped depression on top; aedeagus 4 x longer than wide; opening to vesica % x total length of aedeagus; vesica long, 2 x wider than aedeagus, decreasing in width as it approaches to apex; a band of medium-size spines near basal area of vesica.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
93F5844AE5B551A7A4372746F1C51789.taxon	distribution	Distribution. Only two specimens are known from a cloud forest in Cundinamarca, Colombia (Fig. 94).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
93F5844AE5B551A7A4372746F1C51789.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F87312EB4A735280A96C39711D7AE0D9.taxon	etymology	Etymology. Millerana is dedicated to my mentor and friend who is like a family member Dr. Jacqueline Y. Miller, an American entomologist who has worked on Lepidoptera, especially Castniidae, for nearly 40 years. However, her most important legacy has been her guidance to lepidopterists throughout the world.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F87312EB4A735280A96C39711D7AE0D9.taxon	diagnosis	Diagnosis. Millerana is the most genetically distant genus in the Gaujonia genus group and is more closely related to Gaujoptera (Fig. 1). Externally, Millerana is similar to Cicadoforma and Cicadomorphus, but smaller in size and a slightly different forewing pattern. The genitalia valva is simple as for Cicadoforma and Cicadomorphus, but much wider, with relatively long protuberances at the apex and outer margin. The vesica has a band of spines surrounding the middle area. DNA barcoding reveals that Millerana is distant from the other genera: Gaujoptera (~ 9 % divergent), Cicadoforma, and Cicadomorphus (~ 18 %).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F87312EB4A735280A96C39711D7AE0D9.taxon	description	Description. Orbicular spot well developed; reniform spot with small lunate marking. Forewing pale yellow with black scales, which form the forewing pattern. Hindwing with yellow scales restricted to veins and margin, extending to fringe. Antenna dark brown or brownish orange, serrate, with a stripe of pale-yellow scales on basal % of antenna. Mouthparts reduced; eyes covered by black interfacetal setae. Male genitalia: moderately sclerotized; valva wide, without clasper; apex small with a pointed extension; uncus broad, beak-like; aedeagus short, vesica with narrow band of spines around middle part of vesica.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F87312EB4A735280A96C39711D7AE0D9.taxon	biology_ecology	Biology. Millerana tigrina is the only species known and is considered a pest of pine trees in Ecuador (Bollino and Onore 2001).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
AE610BCE43F850A298139339A5848954.taxon	etymology	Etymology. In honor of my colleague and friend Dr. Deborah Matthews-Lott, who has supported me during my travels in entomology, especially on Lepidoptera trips.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
AE610BCE43F850A298139339A5848954.taxon	diagnosis	Diagnosis. Millerana matthewsae is closely related to M. tigrina; it has brownish orange antenna and can be differentiated by the wide well-defined pattern of foreign lines, and the large orbicular and reniform spots. The most distinctive character of the male genitalia is the costal margin of the valva is strongly curved upward.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
AE610BCE43F850A298139339A5848954.taxon	description	Description. Head. Palp with long black and yellow hair-like scales; frons dark yellow. Thorax. Dark yellow with some black patches on dorsal area; Wing. Forewing length: male 18 - 20 mm; forewing paler yellow coloration than thorax; antemedial, medial, and subterminal lines complete; postmedial line almost complete except in area between cells of CuA 1 and half of CuA 2; V-shaped mark at base of CuA 2 with upper line longer than lower line; terminal area covered by scattered black scales except in M 3 and CuA 1 cells; orbicular spot very large, barely touching top and bottom of discal cell; reniform spot with large line marking inside of discal cell; hindwing white with veins and fringe pale yellow. Leg. Prothoracic and mesothoracic legs brownish orange and black with joints yellow; metathoracic legs yellow. Abdomen. Yellow, paler than remainder of body; dorsally with thin black tufts in middle of A 1 - A 6 and with dark yellow tufts at A 2 - A 3. Male genitalia. Cucullus wide and curved upward on costal margin; apex rounded; some protuberances in the outer margin with the upper one the largest and curved downward; sacculus wide and short; upper side of juxta has a shallow concave depression; tegumen narrow laterally; aedeagus narrow, ⅓ x longer than vesica; vesica long rounded basally; Transverse band of spines near middle of vesica.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
AE610BCE43F850A298139339A5848954.taxon	distribution	Distribution. This species has been collected only in the western zones of Peru at very high elevations almost 4000 m (Fig. 94).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
AE610BCE43F850A298139339A5848954.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F0B099DAE73D5AB1B02185541549686F.taxon	etymology	Etymology. The name refers to the tigrina, Leopardus guttulus (Hensel), in keeping with the wild feline names, as with the other genera, since they are known as " the jaguar moths. "	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F0B099DAE73D5AB1B02185541549686F.taxon	diagnosis	Diagnosis. Millerana tigrina can be distinguished from M. matthewsae by the large, well-rounded orbicular spot. Millerana tigrina also has distinctive genitalia, with the valva presenting a pronounced concavity on the outer margin, forming a moon-shaped crescent.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F0B099DAE73D5AB1B02185541549686F.taxon	description	Description. Head. Palp covered with hair-like scales with dorsal side black and ventral side whitish yellow; last segment black with small whitish yellow dot internally; frons yellow. Thorax. Covered with yellow hair-like scales with black patches. Wing. Forewing length: male 17 - 19 mm; forewing yellow with black scales forming wing pattern; basal line barely visible; antemedial and medial lines disrupted at discal cell; reniform spot large, with small lunate marking in middle; orbicular spot rounded, almost as high as discal cell; two V-shaped marks, one at base of R 1 + R 2, other one at base of CuA 2; hindwing whitish silver with yellow fringe; veins yellow. Leg. Prothoracic and mesothoracic legs dark brown with some patches of black and pale yellow; metathoracic legs pale yellow with dark brown from tibia to claws. Abdomen. Covered with pale-yellow, hair-like scales; three small black tufts on A 1 - A 3; a pale-yellow terminal tuft. Male genitalia. Outer margin of cucullar area concave, with two small protuberances, one subapical and lateral to apex and other one smaller, on lower end of lobe; saccular area wide; tegumen wider dorsally and squared; juxta concave posteriorly; aedeagus 3 x longer than wide; a diagonal opening to vesica ⅓ length of aedeagus; vesica almost as long as aedeagus; transverse band of spines near base.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F0B099DAE73D5AB1B02185541549686F.taxon	distribution	Distribution. The two specimens of Millerana tigrina were found in the highest elevations of the Andes in Ecuador (Fig. 94).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F0B099DAE73D5AB1B02185541549686F.taxon	biology_ecology	Biology. Only one species is known in this genus, Millerana tigrina, which was discussed first by Bollino and Onore (2001). Adults are active throughout the year, but mainly during summer. Larvae resemble some species of the genus Panthea, which also feed on pine trees. Larvae have been observed to feed on Podocarpus magnifolius J. Buchholz & N. E. Grayand and Quercus humboldtii Bonpl as well (O. Mahecha-Jimenez pers. comm.).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
2BD9F0AD3DB55112A15FFF03305598F9.taxon	etymology	Etymology. This species is only found in Bolivia, hence the proposed name.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
2BD9F0AD3DB55112A15FFF03305598F9.taxon	diagnosis	Diagnosis. This species can be distinguished from O. renifera, and O. schmidti by the large lunate mark in the reniform spot, being lightning bolt shaped, and by the small orbicular spot. The male genitalia have the saccular process with a sharply pointed tip; uncus thin. Oculicattus uturunku and O. boliviana share almost identical mtDNA, however, distribution and biology, in addition to morphology demonstrate that they are different species.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
2BD9F0AD3DB55112A15FFF03305598F9.taxon	description	Description. Head. Segments of palp divided in black upper side and yellow underside; two large black spots posterior to antenna. Thorax. Almost entirely light yellow with some small black spots on dorsal area. Wing. Light yellow with wide black lines; forewing length: male 17 - 19 mm; forewing yellow, like rest of body, with black stripes defining pattern; reniform spot with large lightning bolt-like lunate marking; orbicular spot black, small, outlined in pale yellow; open V-shaped mark on CuA 2 at base wide; fringe on hindwing light yellow with black terminal line, interrupted at the veins; posterior margin with fringe whitish yellow; vein yellow with three lines on each vein from M 1 to CuA 2. Leg. Prothoracic leg black with tibia and metatarsi yellow, whereas tarsi black, even at joints. Abdomen. Bright yellow with dorsal region clothed with dark brown scales; dorsal abdomen with yellow tufts and a thin black line on A 1 - A 5. Male genitalia. Cucullar area wide, densely covered by hair-like setae; apex tapered; saccular area relatively narrow, ends in sharp process with a sharply pointed tip; saccular process with setae mainly confined to upper side; saccus V-shaped, considerably flat on tip; juxta flat on upper side and narrow on under side; tegumen wide; uncus thin; aedeagus 1 ⅔ x longer than wide; vesica base 1 / 2 x as long as vesica; two oval-shaped subapical spines patches, with small cluster of spines is near the middle of the dorsal wall of the vesica.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
2BD9F0AD3DB55112A15FFF03305598F9.taxon	distribution	Distribution. The specimens were collected in south-central Bolivia at high elevations above 2000 m (Fig. 95).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
2BD9F0AD3DB55112A15FFF03305598F9.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
414F14EE7ADC5DA19F942DBB83F98FC6.taxon	etymology	Etymology. The name is in honor of a great lepidopterist, Dr. Gunnar Brehm, who loaned some specimens for this research.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
414F14EE7ADC5DA19F942DBB83F98FC6.taxon	diagnosis	Diagnosis. Oculicattus brehmi is the smallest species in this genus; pattern formed by relative pale, thin markings. Orbicular spot small, black. There V-shaped mark at the base of CuA 2 is small. Male genitalia have a tapered cucullar region, wider at the base; the saccus smaller and more tapered than in other species, and the saccular process is the second shortest in the genus with only O. uturunku being shorter than O. brehmi.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
414F14EE7ADC5DA19F942DBB83F98FC6.taxon	description	Description. Head. Wide, palp large, with last segment marbled in light yellow and black; frons pale yellowish gray; female similar to male, even in coloration. Thorax. Covered with dark sulfur-yellow scales and with small gray spots on dorsum. Wing. Pale yellow with pattern of thin dark gray lines; forewing length: male 16 - 18 mm; female: 21 - 23 mm; forewing with thin dots and stripes forming lines; lunate marking in reniform spot narrowly outlined by a thin yellow line; orbicular spot black, small; V-shaped mark at base of CuA 2 small; hindwing with fringe yellow with long yellow lines between veins terminally. Leg. Prothoracic leg brown with joints pale yellow; mesothoracic legs marbled in brown and yellow, tarsi brown with each joint yellow; metathoracic legs yellow. Abdomen. Pale yellow with dorsal area gray; dorsal tufts sulfur-yellow along middle of abdomen, smaller on A 1 - A 2, whereas A 5 - A 8 are wide and combined with gray scales. Male genitalia. Cucullus tapered with small rounded apex and base wide, heavily covered with setae; sacculus wide with process long and densely clothed with setae; saccus V-shaped, and more narrowly tapered towards the end; juxta wide deeply concave posteriorly ,, with the outer edges sharply pointed; tegumen wide; aedeagus 1 1 / 4 x length of vesica to medial cluster of spines; vesica with small patch of spines near middle; apical part of vesica bulbous, with large tapered patch of spines on each side. Female genitalia. anal papilla rounded posteriorly; posterior apophysis ⅓ x longer than anal papilla; anterior apophysis ⅔ x shorter than posterior apophysis; sterigma crescent moon shaped; corpus bursae 1 / 4 x longer than appendix bursae.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
414F14EE7ADC5DA19F942DBB83F98FC6.taxon	distribution	Distribution. This species has been recorded only in Ecuador (Fig. 95).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
414F14EE7ADC5DA19F942DBB83F98FC6.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
B941DDAEF551544F89058993B8BFF808.taxon	etymology	Etymology. This is species is named after the Inca Empire, which originated in the area around Cusco.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
B941DDAEF551544F89058993B8BFF808.taxon	diagnosis	Diagnosis. Oculicattus inca is similar to O. renifera; however, there are some morphological features that are useful to differentiate them. O. inca is slightly smaller, darker in coloration, the orbicular spot smaller, and wing lines are thinner than in O. renifera; the lunate marking in the reniform spot is not outlined in yellow. Genitalia have a wider cucullus; the juxta is flattened posteriorly, whereas in O. renifera the posterior margin has a V-shaped invagination. The DNA barcodes differ by 5 % between the two species.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
B941DDAEF551544F89058993B8BFF808.taxon	description	Description. Head. Last segment of palp divided in black upper side and yellow under side; frons yellow with some gray scales. Thorax. Grayish yellow with some small gray patches. Wing. Forewing length: male: 19 - 21 mm; forewing grayish yellow with markings wide; orbicular spot small, elongated, grayish yellow, outlined with black scales and with some black scales inside orbicular spot; reniform spot with lunate marking not outlined; V-shaped mark short; hindwing with fringe black, vein ends yellow, with some black scales on vein CuA 2. Leg. Prothoracic and mesothoracic legs black anteriorly, yellow posteriorly, whereas metathoracic legs all yellow. Abdomen. Dorsal area brown with some yellow scales between each tergite; a large yellow tuft on A 3; last abdominal segment with long yellow scales. Male genitalia. Cucullar and saccular regions large; saccus triangular; juxta V-shaped with posterior part flattened; vesica 1 1 / 2 x size of aedeagus; Vesica with small patch of spines near middle of vesica; bulbous apical part of vesica with large dense patch of spines posteriorly and smaller, less dense patch anteriorly.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
B941DDAEF551544F89058993B8BFF808.taxon	distribution	Distribution. Broadly distributed in the cloud forests in Peru and Bolivia (Fig. 95).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
B941DDAEF551544F89058993B8BFF808.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
D11289FFF7735B33BEA8D3FDFFAAFB10.taxon	etymology	Etymology. Oculicattus refers to the reniform spot, which is black and surrounded with yellow scales, giving it the appearance of a cat's eye.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
D11289FFF7735B33BEA8D3FDFFAAFB10.taxon	diagnosis	Diagnosis. Oculicattus can be differentiated from Gaujonia externally by the presence of the large reniform spot in Oculicattus, as well as by other features (see Gaujonia diagnosis).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
D11289FFF7735B33BEA8D3FDFFAAFB10.taxon	description	Description. Sexually dimorphic in size, females larger than males; forewing and hindwing hyaline with sulfur-yellow and black scales covering veins and wing margins. Forewing with a small black or sulfur-yellow orbicular spot, sometimes barely perceptible; reniform spot elongated, outlined in black; elongated black central line surrounded by sulfur-yellow outline, except for O. raizae and O. uturunku, in which the spot is entirely black. Antenna filiform, brownish orange with a sulfur-yellow band on basal to three quarters of antenna; eyes with coppery interfacetal setae. Male genitalia slightly sclerotized; valva with saccular and cucullar regions separated, without clasper; uncus long and narrow ending in beak-like tip; vesica has spine patch; vesica wider than its base, which has two patches of spines with one patch of spines larger than other. Female genitalia medium sized; lightly sclerotized rectangular-shaped sterigma; appendix bursae elongate and rugose; corpus bursae for most species approximately half size of appendix bursae.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
D11289FFF7735B33BEA8D3FDFFAAFB10.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
2E45AF379CF45CA0BABBFA0B0513F8B6.taxon	etymology	Etymology. The species is dedicated to my wife, Raiza Castillo, for her love and support since the beginning of my career.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
2E45AF379CF45CA0BABBFA0B0513F8B6.taxon	diagnosis	Diagnosis. Oculicattus raizae is closely related to O. uturunku; they differ from the other Oculicattus species by the darker color, and the unusual reniform spot that is completely brown or black. Oculicattus raizae can be identified by its brownish yellow coloration, and the brown forewing pattern. In the male genitalia the cucullus is longer and wider than in O. uturunku. Additionally, DNA barcodes are ca. 4 % different.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
2E45AF379CF45CA0BABBFA0B0513F8B6.taxon	description	Description. Head. Palp reduced in size; dorsal surface covered by brown scales, ventral surface by dark sulfur-yellow scales; frons covered by dark sulfur-yellow scales, mixed with black; female similar to male externally. Thorax. Dark sulfur-yellow with some brown patches. Wing. Forewing with dark sulfur-yellow and brown scales covering venation and margins; forewing length: male: 22 - 24 mm; female: 27 - 29 mm; antemedial, postmedial, and subterminal lines not defined; orbicular spot small, poorly-defined oblong brown spot; reniform spot brown, narrow, and barely defined; base of CuA 2 with Y-shaped mark large, brown; a brown line across fold on cell at CuA 2; hindwing hyaline with some dark sulfur-yellow scales on venation, but more notable along margins; a Y-shape mark at base of CuA 2; some small brown lines scattered through M 3 + CuA 1 + CuA 2. Leg. Dark sulfur-yellow with brown patches on anterior area of prothoracic legs, which is lighter on mesothoracic and metathoracic legs. Abdomen. Clothed in grayish yellow hair-like scales with white scales at terminus and with a dark sulfur-yellow tuft on A 8. Male genitalia. Cucullus wide, parallel sided with the apex rounded; costal margin swollen basally; sacculus triangular with saccular process entirely coated with setae; tegumen wide; saccus broadly V-shaped; juxta small with parallel sides and a V-shaped notch posteriorly; aedeagus 3 x longer than wide, with a wide opening to vesica ca. 1 / 4 x total length of aedeagus; vesica with basal area ca. same width that of aedeagus, and it is ventrally curved; basal area with a large patch of spines; vesica with two patches of spines on each side, one small slightly beyond middle of dorsal wall of the vesica, and truncated anteriorly, whereas other larger apically narrow, and broader towards the aedeagus. Female genitalia. Small and truncated apically; anal papilla with posterior apophysis 1 / 4 x shorter than anal papilla; anterior apophysis short; rectangular sterigma that is fused above ostium; ductus bursae wide and long; appendix bursae ⅔ x shorter than corpus bursae which is partially sclerotized at base.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
2E45AF379CF45CA0BABBFA0B0513F8B6.taxon	distribution	Distribution. Recorded from cloud forests in Colombia, Ecuador, Peru, and Bolivia in middle and high elevations from 2000 m to above 3000 m (Fig. 95).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
2E45AF379CF45CA0BABBFA0B0513F8B6.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
11DA460FFD6A592C84F214E792767D9B.taxon	etymology	Etymology. George F. Hampson probably named this species renifera based on the characteristic reniform spot.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
11DA460FFD6A592C84F214E792767D9B.taxon	diagnosis	Diagnosis. Regarding Ocullicatus renifera, there is one species that is particularly similar to it (O. inca); however, O. renifera can be identifiable by its remarkably large yellow orbicular spot. The male is pale yellow, brighter yellow in the female. The forewing length in males is 19 - 21 mm and females 24 - 26 mm. Palp reduced, black with white tips; frons yellow and gray; antenna dark orange; male thorax dark yellow with scattered brownish black spots. Pattern on the forewing in male is created by black small lines and dots; the reniform spot has slightly curved and wide lunate marking. The hindwing has dark scales on the veins. The abdomen is brown with yellow scales dorsally, differing from O. inca in having a line of black tufts over the middle of the dorsum of the abdomen; black tufts on A 1 - A 3 combined with yellow scales. Genitalia of male with the cucullar region narrow and the apex rounded; saccular area with the process almost same size that the cucullus; aedeagus long, almost the same size of the vesica; large patch of spines on the base of the vesica; the two spine patches large and wide similar size covering 3 / 4 of the vesica. Genitalia in female with sterigma relatively open, with the corpus bursae almost the same size as the appendix bursae.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
11DA460FFD6A592C84F214E792767D9B.taxon	distribution	Distribution. Oculicattus renifera is restricted to Peru and Bolivia (Fig. 95).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
11DA460FFD6A592C84F214E792767D9B.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
B4251AC323D753C98CD0D0E8E39C9BCD.taxon	etymology	Etymology. The name of this species is in honor of my colleague and friend Dr. B. Christian Schmidt, who shared his knowledge with me about Noctuoidea without hesitation.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
B4251AC323D753C98CD0D0E8E39C9BCD.taxon	diagnosis	Diagnosis. Oculicattus schmidti has a set of remarkable characters that separate it from other species, such as the orbicular spot is rounded; the lunate marking of the reniform spot is incomplete, being similar to those of Gaujonia species; the forewing is more stylized with straighter margins. The male genitalia are similar to those of O. boliviana, but can be distinguished from them by the cucullar region, which is narrower than those of other species; the saccular is large and densely covered with setae; also the uncus is wider and the saccus longer than the other species of Oculicattus. The DNA barcode is similar to that of O. renifera; however, external and internal morphology reveal enough differences to identify the two species.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
B4251AC323D753C98CD0D0E8E39C9BCD.taxon	description	Description. Head. Palp marbled in black and white with frons yellowish gray; black patch between antennae small. Thorax. Covered in yellow with some black patches on dorsum. Wing. Forewing length: male 17 - 19 mm; forewing, dark yellow with well-developed lines from posterior margin to fold that look similar to those of other Oculicattus species; reniform spot with lunate marking narrowed in middle of base of cell M 1; orbicular spot small, outlined in black, rounded; V-shaped mark on CuA 2 base small, with upper line longer, extending to lower side of reniform spot; hindwing with fringe marbled in yellow and black; posterior margin with a combination of yellow and brown hair-like scales; veins yellow with two black lines in middle of veins from M 2 to CuA 2; black line posterior to base of CuA 2. Leg. Prothoracic and mesothoracic legs marbled in yellow and black; metathoracic legs yellow. Abdomen. Dark yellow with dorsum clothed with dark brown and black scales; yellow tufts in middle of A 1 - A 4 with a small vertical line on each tuft. Male genitalia. Cucullus bullhorn-like, with pointed apex heavily coated with setae; sacculus narrow with its process long and wide, densely covered with setae; terminus of saccular process tapered to apex; saccus long rhomboid-shaped, tip barely rounded; juxta shield-shaped and upper side concave; juxta with an expanded upper side; wide tegumen; aedeagus 1 / 4 x longer than basal area of vesica, remarkable curved inwards; vesica ⅔ x longer than wide; one of patches of spines on tip small slightly squared and other covering ⅓ of whole vesica.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
B4251AC323D753C98CD0D0E8E39C9BCD.taxon	distribution	Distribution. This species is distributed from Central region of Peru and to the east region of Bolivia (Fig. 95).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
B4251AC323D753C98CD0D0E8E39C9BCD.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F3C8831AB2545931959332D2CB89819C.taxon	etymology	Etymology. The name uturunku makes reference to the jaguar Panthera onca (Linnaeus) in the Quechua language.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F3C8831AB2545931959332D2CB89819C.taxon	diagnosis	Diagnosis. There is only one species that has similar characters to O. uturunku, which is O. raizae. Nevertheless, they are easy to separate because the color black is predominant in O. uturunku and the yellow is secondary. The reniform spot is black and quite smaller, converging with long black lines making it look like an eyelash. Genitalia of the male are remarkably smaller than O. raizae, mainly the cucullus, which is also remarkably narrower.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F3C8831AB2545931959332D2CB89819C.taxon	description	Description. Head. Palp with last segment divided into three parts, base and tip yellow and middle area black; frons dark yellow, with a large black band between antennae. Thorax. Marbled in black and sulfur-yellow dorsally, and sulfur-yellow ventrally. Wing. Forewing length, male 20 - 22 mm; forewing black with some regions of sulfur-yellow; enormous black lines on veins define forewing pattern; black orbicular spot small and elongated; unusual reniform spot eyelash-like, black; black line through inferior region of discal cell barely touching base of CuA 2; hindwing with black fringe and some sulfur-yellow scales at end of each vein paler than forewing, whereas that from posterior margin completely gray; veins black with some spots of sulfur-yellow. Leg. Prothoracic and mesothoracic legs black with some sulfur-yellow on joints, and metathoracic legs in yellow. Abdomen. Black with segments sulfur-yellow ventrally, whereas dorsally dark gray, paler on first three segments; A 1 - A 3 with tufts in yellow and with some black scales. Male genitalia. Cucullus wider on base and apex small; costal margin curved; sacculus and process wide; saccus narrow and rhomboid-shaped; juxta square-shaped with base narrower; tegumen narrow; aedeagus 3 x longer than wide; basal area of vesica 1 1 / 2 x longer than vesica itself; large slightly curved patch of spines close to basal area; one of patches of spines on tip small with triangular terminus and another larger covering almost 1 / 2 of vesica.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F3C8831AB2545931959332D2CB89819C.taxon	distribution	Distribution. Oculicattus uturunku occurs mainly in the Western Cordillera of the Andes in Ecuador. It is found at high to very high elevations (Fig. 95).	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
F3C8831AB2545931959332D2CB89819C.taxon	biology_ecology	Biology. Unknown.	en	Martinez, Jose I. (2020): Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species. ZooKeys 985: 71-126, DOI: http://dx.doi.org/10.3897/zookeys.985.51622, URL: http://dx.doi.org/10.3897/zookeys.985.51622
