identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
B199F4E7403A5378871300271D48F54C.text	B199F4E7403A5378871300271D48F54C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronychini Gistel 1848	<div><p>Macronychini Gistel, 1848</p><p>Macronychini Gistel, 1848: unnumbered page between columns 400 and 409. Type genus: Macronychus Müller, 1806 .</p><p>Notes.</p><p>No modern classification of Elmidae has been published so far. Zaitzevia and several other closely related genera are currently placed in the tribe Macronychini . But due to the lack of comprehensive phylogenetic studies, it is currently not possible to provide a valid subfamily classification.</p></div>	https://treatment.plazi.org/id/B199F4E7403A5378871300271D48F54C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Bian, Dong-Ju;Jäch, Manfred A.	Bian, Dong-Ju, Jäch, Manfred A. (2025): The species of the genus Zaitzevia Champion, 1923 (Coleoptera, Elmidae) from Shaanxi Province, China. ZooKeys 1264: 129-157, DOI: 10.3897/zookeys.1264.156144
D38B58A772065B2392938BD4E30D1EAC.text	D38B58A772065B2392938BD4E30D1EAC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zaitzevia Champion 1923	<div><p>Genus Zaitzevia Champion, 1923</p><p>Zaitzevia Champion, 1923: 170 . Type species: Zaitzevia solidicornis Champion, 1923 . Awadoronus Kôno, 1934: 127 (synonymized by Nomura 1958). Type species: Awadoronus awanus Kôno, 1934 .</p></div>	https://treatment.plazi.org/id/D38B58A772065B2392938BD4E30D1EAC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Bian, Dong-Ju;Jäch, Manfred A.	Bian, Dong-Ju, Jäch, Manfred A. (2025): The species of the genus Zaitzevia Champion, 1923 (Coleoptera, Elmidae) from Shaanxi Province, China. ZooKeys 1264: 129-157, DOI: 10.3897/zookeys.1264.156144
F71BA7B71D295530809672D840A5AD95.text	F71BA7B71D295530809672D840A5AD95.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zaitzevia chenzhitengi Jiang & Wang 2020	<div><p>Zaitzevia cf. chenzhitengi Jiang &amp; Wang, 2020</p><p>Figs 4 A, B, 10 D – F</p><p>Zaitzevia chenzhitengi Jiang and Wang 2020: 232; Bian and Zhang 2022: 265; Jiang and Chen 2023: 204 (key).</p><p>Material examined.</p><p>China, Shaanxi • 1 ♂ (IAECAS): “ China: Shaanxi, \ Qinling Northwest \ A &amp; F University forestry center | Huoditang \ 1974 m, 2005.6. 12, Leg. Wang M. ” ; Sichuan • 2 ♂♂, 2 ♀♀ (NMW): “ China: Sichuan \ Emeishan \ 160 km SSW Chengdu \ 1530 m (4 a) [CWBS 52] \ 22. 6. 1994 \ leg. Schillhammer ” ; Yunnan • 1 ♂ (IAECAS): “ China: Yunnan (20) \ Nujiang, \ Lushui, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.66167&amp;materialsCitation.latitude=26.000834" title="Search Plazi for locations around (long 98.66167/lat 26.000834)">Pianma</a> | 26°0'3"N, 98°39'42"E \ 2392 m, 2018.10. 25. \ Leg Peng, Zhu, Dong ”; 3 ♂♂, 3 ♀♀ (IAECAS) : “ CHINA: Yunnan (25) \ Nujiang, Lushui \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.66167&amp;materialsCitation.latitude=26.000834" title="Search Plazi for locations around (long 98.66167/lat 26.000834)">Pianma</a> | 26°0'26"N, 98°38'54"E \ 2172 m, 2019.7. 6 \ Leg. Peng, Zhu, Dong ” • 2 ♂♂, 4 ♀♀ (NMW): “ China NW-Yunnan \ 15 km N <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.66167&amp;materialsCitation.latitude=26.000834" title="Search Plazi for locations around (long 98.66167/lat 26.000834)">Lijiang</a> \ 2800 m, 6. 7. 1994 \ leg. JI (16) [CWBS 60] ” .</p><p>Taxonomy.</p><p>Zaitzevia chenzhitengi was described by Jiang and Wang (2020) based on one male and one female from northern Sichuan. Two years later, Bian and Zhang (2022) newly recorded this species from Shaanxi and Yunnan and published photographs of the male habitus and the aedeagus of a specimen from Shaanxi. However, after having examined additional material, which we compared with the original description, we want to express our doubts about the identity of the specimens listed above. Although not explicitly mentioned in the written description by Jiang and Wang (2020), the elytral apices of the holotype are distinctly produced and conjointly acuminate in the figures (see Jiang and Wang 2020: figs 1 A, 2 F), while none of the nine males from the three provinces examined by us has distinctly acuminate elytral apices (see Fig. 4 A). Furthermore, the body form of the holotype of Z. chenzhitengi is clearly more slender and parallel-sided, and the anterior angles of the pronotum seem to be less acute. Despite the external differences, the aedeagi are remarkably similar, but the endophallic teeth appear to be smaller and situated further proximally in the holotype of Z. chenzhitengi . More material from the type area of Z. chenzhitengi and / or DNA sequencing will be necessary to clarify the identity of “ Z. cf. chenzhitengi ”.</p><p>Distribution.</p><p>Shaanxi, Sichuan, Yunnan.</p></div>	https://treatment.plazi.org/id/F71BA7B71D295530809672D840A5AD95	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Bian, Dong-Ju;Jäch, Manfred A.	Bian, Dong-Ju, Jäch, Manfred A. (2025): The species of the genus Zaitzevia Champion, 1923 (Coleoptera, Elmidae) from Shaanxi Province, China. ZooKeys 1264: 129-157, DOI: 10.3897/zookeys.1264.156144
C97BBB869AEE5623AF73446170D26E7A.text	C97BBB869AEE5623AF73446170D26E7A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zaitzevia coronifer Bian & Jäch 2025	<div><p>Zaitzevia coronifer sp. nov.</p><p>Figs 2 C, D, 7 A – C, 11 D – F</p><p>Material examined.</p><p>(117 exs) Holotype: China • ♂ (IAECAS): “ China – Shaanxi \ Zhouzhi County l. w [leg. Wang] | Houzhenzi Ca 1200 m \ 2. VI 1998 ” [CWBS 308] . Paratypes: China, Shaanxi • 5 ♀♀ (IAECAS), the same data as holotype • 35 exs (NMW): “ China: Shaanxi, 2. 6. 1998 \ Zhouzhi County, ca. 1200 m \ 2 km W Houzhenzi Nat. Res. \ leg. M. Wang (CWBS 308) ” • 2 ♂♂, 3 ♀♀ (IAECAS): “ China: Shaanxi, Ankang City, \ Ningshan County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.5468&amp;materialsCitation.latitude=33.557884" title="Search Plazi for locations around (long 108.5468/lat 33.557884)">Xunyang Dam</a> | 33°33.473′N, 108°32.808′E, \ 1355 m, 2019.8. 20 \ Leg. Tong Y. F ” • 1 ♂, 1 ex. (IAECAS): “ China-Shaanxi \ Mei County L [leg.]. W [Wang] | Tangyu ca. 1100 m \ 1. VI. 1998 ” [CWBS 307] • 1 ♂ (NMW): “ China: Shaanxi, 4. 6. 1998 \ Foping County, ca. 1 300 m \ 5 km S Longcaoping \ leg. Wang (CWBS 312) ” • 2 ♂♂, 8 ♀♀ (IAECAS): “ China-Shaanxi \ Ningshan L. [leg.] W [Wang] | Xunyangba ca. 1500 m \ 6. VI. 1998 ” [CWBS 315] • 40 exs (NMW): “ China: Shaanxi, 6. 6. 1998 \ Ningshan Co., ca. 1 500 m \ 10 km NE Xunyangba \ leg. M. Wang (CWBS 315) ” • 1 ♂, 1 ♀ (NMW): “ China: Shaanxi, 8. 6. 1998 \ 2 km SE Liuba, 1 400 m \ ( Zhangliangmiao) \ leg. M. Wang (CWBS 317) ” • 22 exs (CBC, NMW): “ China, 17. – 22. VI. \ Shaanxi prov. 1991 \ Hua Shan peak env. \ 100 km E of Xi’an \ Z. Kejval lgt. ” ; Hubei • 3 exs (NMW): “ China: Hubei, 13. 10. 2004 \ Shennongjia Forest Distr. \ 5 km E Muyu, Tong Mu \ 1 250 m, leg. Schönmann \ &amp; Wang (CWBS 531) ” • 2 ♂♂, 4 ♀♀ (IAECAS): “ China: Hubei \ Shennongjia For. Dist. | 1600 m, 2004.10. 15 \ Leg. Wang (CWBS 536) ” .</p><p>Differential diagnosis.</p><p>In Shaanxi, this species is similar to Z. pilosa sp. nov. in body size (BL 2.3–2.7 mm) and habitus, but it can be easily distinguished from the latter by the less pronounced dorsal pilosity, the granules on the elytral interval 5 not reaching the elytral base and by the glabrous shoulders; the aedeagus differs mainly in the more strongly produced apex and the absence of endophallic teeth.</p><p>Zaitzevia coronifer also resembles Z. tangliangi, but it can be easily distinguished from the latter, e. g., by the stronger pronotal punctation, more distinct sublateral pronotal carinae, stronger tibiae (especially foretibiae), and by the clearly different aedeagus, being less elongate, apically more acuminate, with characteristic endophallic sclerotizations.</p><p>The new species is externally also somewhat similar to Z. tsushimana, which is, however, markedly smaller (BL 1.8–2.2 mm), with its median pronotal groove more or less reaching the pronotal base, and its elytral carinae being located on intervals 5–7 (see Jung et al. 2015; Iwata et al. 2022: fig. 3 G).</p><p>Description.</p><p>Male (holotype). Body broadly obovate. BL 2.6 mm, BW 1.2 mm. Antennae yellowish brown, dorsum black, ventral side brown to black, legs reddish brown to dark brown except tarsi light brown.</p><p>Labrum microreticulate in basal 0.1, distal 0.9 smooth and shiny, sparsely punctate and pubescent. Anterior margin broadly rounded, not emarginate, densely pubescent laterally. Clypeus coarse, densely pubescent, sparsely granulate. Frons densely pubescent, with a few small granules near inner sides of eyes.</p><p>Pronotum (PL 0.7 mm, PW 0.9 mm) broadest at base, gradually narrowing anteriad. Lateral margin slightly rimmed. Anterior corners acute, slightly produced, posterior corners right-angled. Disc smooth and shiny, densely punctate, and sparsely pubescent. Area near anterior and posterior corners and in front of scutellum coarse, finely granulate. Median sulcus reaching from basal 0.2–0.5, narrow, not broadened medially. Sublateral carinae extending from base to basal 0.4, slightly raised.</p><p>Elytra (EL 1.9 mm, EW 1.2 mm) broadest at distal 0.33, slightly narrowed anteriorly and distinctly attenuate posteriorly. Strial punctures moderately large, especially in the middle area, separated by ~ 0.5–1.0 × their diameters, becoming smaller and shallower towards declivity, separated by 2–3 × their diameters. Intervals flat, more or less smooth and glabrous, hardly punctate, and hardly pubescent. Intervals 5, 7 and 8 carinate; carinae on interval 5 extending from basal 0.2 to apex; carinae on intervals 7 and 8 extending from basal 0.1 to apex. Elytral apices densely granulate, separately rounded (in posterior view).</p><p>Prosternum laterally densely pubescent; prosternal process gradually narrowed from base to distal 0.2, distinctly narrowed in distal 0.2, apex narrowly rounded, lateral margin distinctly rimmed; disc coarse with some large granules, sparsely punctate. Disc of metaventrite broadly and distinctly impressed, smooth and shiny, with a few very small punctures and large granules; with a row of large punctures behind of mesocoxa and another row of small punctures in front of metacoxa; median sulcus complete, gradually narrowing anteriad; lateral margin of disc in the median part (between meso- and metacoxae) with a large hump covered with a few distinct granules; lateral areas densely pubescent and sparsely granulate.</p><p>Ventrite I moderately densely punctate, with a pair of admedian carinae; disc more or less impressed, anterior margin raised. Middle of ventrites II – IV smooth and shiny, sparsely and finely punctate. Lateral areas of ventrites I – V densely pubescent. Ventrite V densely pubescent and sparsely granulate; lateral margins V fringed with short spines; apical margin weakly emarginate medially; lateral corners not forming distinct teeth.</p><p>Protibiae slightly enlarged.</p><p>Aedeagus. 1.2 mm long, elongate, cylindrical. Penis ~ 1.9 × as long as phallobase, gradually broadened in basal 0.33, then almost subparallel to basal 0.67, distinctly acuminate in distal 0.33, apex acute. Endophallus with characteristic sclerotizations resembling a crown. Parameres very slim, reaching apical 0.2 of penis.</p><p>Females. Metaventrite not distinctly impressed; pair of humps between meso- and metacoxae smaller, without granules. Disc of ventrite I glabrous, very sparsely punctate, partly and faintly microreticulate, concave in posterior 0.5. Apex of ventrite V straight or weakly rounded, not emarginate. Protibiae not enlarged.</p><p>Measurements.</p><p>Males: BL 2.50–2.65 mm (n = 10), BW 1.1–1.2 mm (n = 6); females: BL 2.3–2.7 mm (n = 10), BW 1.1–1.2 mm (n = 5).</p><p>Distribution.</p><p>Hubei, Shaanxi.</p><p>Etymology.</p><p>The epithet is derived from the Latin noun coronifer (carrier of a crown) and refers to the crown-like sclerotizations of the endophallus.</p></div>	https://treatment.plazi.org/id/C97BBB869AEE5623AF73446170D26E7A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Bian, Dong-Ju;Jäch, Manfred A.	Bian, Dong-Ju, Jäch, Manfred A. (2025): The species of the genus Zaitzevia Champion, 1923 (Coleoptera, Elmidae) from Shaanxi Province, China. ZooKeys 1264: 129-157, DOI: 10.3897/zookeys.1264.156144
5B246B76E0325B2EA681F05C9119A4AC.text	5B246B76E0325B2EA681F05C9119A4AC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zaitzevia disparilis Bian & Jäch 2025	<div><p>Zaitzevia disparilis sp. nov.</p><p>Figs 3 A, B, 8 A – C, 12 A – C</p><p>Material examined.</p><p>(822 exs) Holotype: China • ♂ (IAECAS): “ China: Shaanxi, \ Foping County \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.96555&amp;materialsCitation.latitude=33.571667" title="Search Plazi for locations around (long 107.96555/lat 33.571667)">Gaozhuanggou</a> | 33°34′18″N, 107°57′56″E \ 1035 m, 2019. VI. 21 \ Leg. Tong ” . Paratypes: China, Shaanxi • 6 ♂♂, 3 ♀♀, 32 exs (IAECAS), the same data as holotype • 3 exs (NMW): “ China: Shaanxi, 1. 6. 1998 \ 130 km SE Baoji, ca. 1 100 m \ Taibai Shan Forest Park \ leg. M. Wang (CWBS 307) ” • 10 exs (NMW): “ China: Shaanxi, 7. 6. 1998 \ 7 km SE Liuba, ca. 1300 m \ Taoyuanpu Village env. \ leg. M. Wang (CWBS 316) ” • 1 ♀ (CPE): “ China: Shaanxi, Qin Ling Shan \ 108.49E, 33.55N [coordinates obviously unprecise], River Valley \ 40 km S Xian, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.49&amp;materialsCitation.latitude=33.55" title="Search Plazi for locations around (long 108.49/lat 33.55)">Autoroute km 50</a> \ River bank, 1200 m \ 31. 08. 1995, leg. A. Pütz ” • 11 exs (CBC, NMW): “ China, 17. – 22. VI. \ Shaanxi prov. 1991 \ Hua Shan peak env. \ 100 km E of Xi’an \ Z. Kejval lgt. ” .</p><p>Additional material.</p><p>China, Gansu • 1 ♀ (NMW): “ China: Gansu, 15. 6. 1998 \ Wen Co., Bikou env. \ Dong Gou River, ca. 950 m \ leg. M. Wang (CWBS 326) ” ; Anhui • 437 exs (NMW): “ China: Anhui, Dabie Shan \ 40 km N Yuexi, 5. 11. 1997 \ env. Gui Xing Di, 800 m \ leg. Schönmann [respectively: “ leg. M. Wang ”] (CWBS 295) ” • 220 exs (NMW): “ China: Anhui, Dabie Shan \ 20 km N Yuexi, 6. 11. 1997 \ env. Shi Guan, 900 m \ leg. Schönmann [respectively: “ leg. M. Wang ”] (CWBS 296) ” • 27 exs (NMW): “ China: Anhui, Dabie Shan \ 20 km N Yuexi, 6. 11. 1997 \ env. Shi Guan, 950–1000 m \ leg. Schönmann [respectively: “ leg. M. Wang ”] (CWBS 297) ” • 82 exs (NMW): “ China: Anhui, Dabie Shan \ 50 km NW Yuexi, 7. 11. 1997 \ Huang Liyan / Baojia, 1050 m \ leg. Schönmann [respectively: “ leg. M. Wang ”] (CWBS 298) ” • 8 exs (NMW): “ China: Anhui, Dabie Shan \ 50 km NW Yuexi, 8. 11. 1997 \ Huang Liyan / Baojia, 1050 m \ leg. Schönmann [respectively: “ leg. M. Wang ”] (CWBS 299) ” • 17 exs (NMW): “ China: Anhui, Dabie Shan \ 25 km N Yuexi, 9. 11. 1997 \ env. Shi Guan, 1000 m \ leg. Schönmann [respectively: “ leg. M. Wang ”] (CWBS 301) ” ; Hunan • 16 exs (NMW): “ China, NW-Hunan 1993 \ Wulingyuan, N Dayong \ Zangjiajie [ Zhangjiajie], 30. 10., 450 m \ leg. Schönmann (4) [CWBS 23] ” • 12 exs (NMW): “ China, NW-Hunan 1993 \ Wulingyuan, N Dayong \ Suoxiyu, 31. 10., 400 m \ leg. Schönmann (5) [CWBS 24] ” ; Guizhou • 1 ♀ (NMW): “ China: Guizhou, Zunyi Pref. \ 25 km NE Zunyi City \ ca. 1000 m, 3. 8. 1997 \ leg. M. Wang (CWBS 278) ” • 1 ♀ (NMW): “ China: Guizhou, Leishan Co. \ SE Kaili, NE Leishan \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.268&amp;materialsCitation.latitude=26.435167" title="Search Plazi for locations around (long 108.268/lat 26.435167)">Leigong Shan</a>, E - slope \ 26°26.11'N, 108°16.08'E \ Wunang River \ 13. 6. 2001, ca. 9-00 m [sic] \ leg. Schillhammer &amp; Wang (CWBS 432) ” .</p><p>Differential diagnosis.</p><p>Zaitzevia disparilis sp. nov. is a rather small species (BL 1.46–2.00 mm), characterized by the often remarkably unequal size of the elytral punctures, by the elytral carinae being located on intervals 5–7, and the conspicuous sclerotizations of the endophallus.</p><p>In habitus and body size it somewhat resembles Z. tsushimana from Jilin, which is on average longer (BL 1.8–2.2 mm) and differs, among other characters, in the longer median pronotal sulcus (see Jäch and Boukal 1995: fig. 59 [as “ Zaitzevia sp. (Jilin) ”]; Iwata et al. 2022: fig. 3 G), the flat elytral intervals (only rarely flat in Z. disparilis sp. nov.) and the structure of the endophallic sclerotizations.</p><p>Description.</p><p>Male (holotype). BL 1.8 mm, BW 0.8 mm. Body elongate obovate. Dorsum and ventral side dark brown, antennae yellowish brown. Legs dark brown but tarsi reddish brown. Labrum wider than long. Basal 0.33 microreticulate, distal 0.67 densely punctate and pubescent, with fringe of long setae laterally. Clypeus and frons densely pubescent and sparsely granulate.</p><p>Pronotum (PL 0.55 mm, PW 0.6 mm) broadest at base, gradually narrowing anteriad, lateral margin narrowly rimmed. Anterior corners acute, slightly produced, posterior corners rectangular. Disc smooth and shiny, punctures sparser than on lateral parts. Areas near anterior and posterior corners coarse, densely and finely granulate. Median groove moderately wide, extending from basal 0.2–0.6. Lateral carinae present in basal 0.4.</p><p>Elytra (EL 1.25 mm, EW 0.8 mm) subparallel in basal 0.67, distal 0.33 distinctly attenuate. Strial punctures in basal 0.5 unequal in size and depth, separated by 1.0–1.5 × their diameters, being deepest in the third stria, becoming smaller and shallower on declivity (separated by 2–3 × their diameters); striae not reaching elytral apex. Elytral intervals weakly convex, smooth and shiny, with only a few small setal punctures; interval 4 very slightly more convex than intervals 1–3; intervals 5–7 carinate (due to fusion of elytral striae 5 and 6), carina on interval 5 extending from basal 0.1 to apex, carinae on intervals 6 and 7 complete, merging at basal ~ 0.1. Elytral apices narrowly conjointly rounded.</p><p>Prosternal disc coarse, sparsely punctate and pubescent, lateral areas densely tomentose. Prosternal process gradually narrowed in basal 0.8, then distinctly narrowed in distal 0.2, apex narrowly rounded. Disc of prosternal process sparsely punctate and pubescent. Metaventrite shiny, with a few larger elongate punctures, slightly impressed medially; median sulcus present in basal 0.7, distinctly narrowed from base to apex. Lateral areas densely pubescent. Posterior margin of metaventrite impressed, with two large impressions behind of mesocoxae, and a row of middle-sized punctures in front of metacoxae.</p><p>Disc of ventrite I flat, smooth, and shiny, with only a few punctures, and a transverse row of few granules posteriorly; anteriorly impressed and wrinkled; laterally bordered by strong admedian carinae. Middle of ventrites II – IV and basal 0.33 of ventrite V smooth and shiny, with only a few punctures. Distal 0.67 of ventrite V densely pubescent and granulate. Lateral areas of ventrites I – IV densely pubescent. Lateral margins of ventrite V fringed with short spines; apex subtruncate, lateral apical corners covered by a brush of stiff setae.</p><p>Aedeagus. 0.8 mm long, elongated, and cylindrical. Penis ~ 2.2 × as long as phallobase. Penis gradually broadened in basal 0.25, then almost subparallel to apical 0.25, then distinctly narrowed toward cuspidal apex. Sclerotizations of endophallus in median part of penis (basal 0.3–0.7) resembling a “ bowknot ” (in ventral / dorsal view), consisting of a pair of ovoid structures, each with an apical curved tooth and a moderately long, thin basal rod. Parameres short, clinging to penis, reaching apical 0.45 of penis.</p><p>Females. Disc of metaventrite more evenly convex or evenly flattened than in males, without larger punctures or granules; ventrite I less impressed anteriorly, without distinct wrinkles; ventrite V very similar to male, but more sparsely granulate; apex subtruncate or weakly rounded.</p><p>Measurements.</p><p>(type specimens) Males: BL 1.68–1.90 mm (n = 10), BW 0.75–0.80 mm (n = 7); females: BL 1.80–1.96 mm (n = 10), BW 0.75–0.80 mm (n = 3).</p><p>Including the additional material, the BL of Z. disparilis is 1.46–2.00 mm. The smallest male (Anhui, CWBS 299) is 1.50 mm long, the smallest female (Anhui, CWBS 301) 1.46 mm, and the largest female (Hunan, CWBS 24) 2.00 mm.</p><p>Distribution.</p><p>Anhui, Gansu, Guizhou, Hunan, Shaanxi. Remarkably, there are no records from Hubei at present.</p><p>Etymology.</p><p>The name is derived from the Latin adjective disparilis (disparate, different, unequal), referring to the differently sized and variably deep impressed elytral punctures.</p><p>Remarks.</p><p>We think that all specimens recorded from these five provinces belong to the same species, but due to the morphological variability (e. g., body length, size and depth of elytral punctures, granules of elytral intervals sometimes reaching elytral base) we do not designate the specimens from Anhui, Gansu, Guizhou, and Hunan as paratypes. Confirmation of the conspecific molecular data would be highly desired, especially in case of the material from Gansu and Guizhou from where only females are currently known.</p></div>	https://treatment.plazi.org/id/5B246B76E0325B2EA681F05C9119A4AC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Bian, Dong-Ju;Jäch, Manfred A.	Bian, Dong-Ju, Jäch, Manfred A. (2025): The species of the genus Zaitzevia Champion, 1923 (Coleoptera, Elmidae) from Shaanxi Province, China. ZooKeys 1264: 129-157, DOI: 10.3897/zookeys.1264.156144
1015FD05253F54C68A74BD25083B13C0.text	1015FD05253F54C68A74BD25083B13C0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zaitzevia hybrida Bian & Jäch 2025	<div><p>Zaitzevia hybrida sp. nov.</p><p>Figs 3 C, D, 9 A – C, 12 D – F</p><p>Material examined.</p><p>(45 exs) Holotype: China • ♂ (IAECAS): “ China, Shaanxi, Qingling \ Ningshan County | Yaowangtang, 1286 m \ 2005.6. 10, Wangm [leg. Wang Miao] ” . Paratypes: China, Shaanxi • 4 ♂♂, 3 ♀♀ (IAECAS): “ China-Shaanxi \ Zhouzhi County I. [leg.] W [Wang] | Houzhenzi ca. 1200 m \ 2. VI. 1998 ” [CWBS 308] • 8 ♂♂, 8 ♀♀ (NMW): “ China: Shaanxi, 2. 6. 1998 \ Zhouzhi County, ca. 1200 m \ 2 km W Houzhenzi Nat. Res. \ leg. M. Wang (CWBS 308) ” • 1 ♀ (NMW): “ China: Shaanxi, 4. 6. 1998 \ Foping County, ca. 1 300 m \ 5 km S Longcaoping \ leg. Wang (CWBS 312) ” • 2 ♂♂, 6 ♀♀ (NMW): “ China: Shaanxi, 6. 6. 1998 \ Ningshan Co., ca. 1 500 m \ 10 km NE Xunyangba \ leg. M. Wang (CWBS 315) ” • 1 ♂ (NMW): “ China, 17. – 22. VI. \ Shaanxi prov. 1991 \ Hue Shan peak env. \ 100 km E of Xi’an \ Z. Kejval lgt. ” .</p><p>Additional material.</p><p>China, Gansu • 2 ♂♂ (NMW): “ China: Gansu, 13. 6. 1998 \ Wen Co., ca. 1100 m \ 4 km N Shangdan Village \ leg. M. Wang (CWBS 322) ” • 1 ♀ (NMW): “ China: Gansu, 14. 6. 1998 \ Wen Co., ca. 1 150 m \ 1 km N Shangdan Village \ leg. M. Wang (CWBS 323) ” ; Sichuan • 2 ♂♂ (NMW): “ China: Sichuan, 13. 6. 1996 \ W Ya’an, 20 km W Tianquan \ Dayuxi stream, 1200 m \ leg. Ji &amp; Wang (CWBS 235) ” • 1 ♀ (NMW): “ China: Sichuan, 29. 7. 1998 \ Mao Xian Co., Jiuding Shan \ 10 km NE Mao Xian, ca. 1950 m \ Schönmann, Ji, Wang (CWBS 336) ” • 1 ♀ (NMW): “ China: Sichuan, 30. 7. 1998 \ Mao Xian Co., Jiuding Shan \ 20 km NE Mao Xian, ca. 1650 m \ Schönmann, Ji, Wang (CWBS 337) ” • 1 ♀ (NMW): “ China: Sichuan, 30. 7. 1998 \ Mao Xian Co., Jiuding Shan \ 15 km NE Mao Xian, ca. 1950 m \ Schönmann, Ji, Wang (CWBS 338) ” • 1 ♀ (NMW): “ China: Sichuan, 30. 7. 1998 \ Mao Xian Cty., Jiuding Shan \ 6 km NE Mao Xian, ca. 1750 m \ Schönmann, Ji, Wang (CWBS 340) ” ; Yunnan • 1 ♂ (NMW): “ China-Yunnan 24. – 29.6. \ 50 km N Lijiang, 1993 \ Yulongshan Nat. Res. \ E. Jendek &amp; O. Sausa leg. ” • 3 ♂♂, 6 ♀♀ (NMW): “ China-Yunnan 14. – 21.6. \ 100 km W Baoshan, 1993 \ Gaoligongshan Nat. Res. \ E. Jendek &amp; O. Sausa leg. ” .</p><p>Differential diagnosis.</p><p>Zaitzevia hybrida sp. nov. is a small species (BL 1.50–1.86 mm), which differs significantly from all other species described in the genus Zaitzevia in possessing characters that are typically found in Zaitzeviaria Nomura, 1959, i. e., elytra with only two instead of three carinae, and elytral the plastron being confined to the area between the lateral margin and interval 7 (therefore not reaching interval 5). This species is furthermore characterized by the enlarged male protibiae.</p><p>Description.</p><p>Male (holotype). BL 1.7 mm, BW 0.7 mm. Dorsum dark brown, ventral side and legs reddish brown except tarsi. Antennae, mouth parts, and tarsi yellowish brown.</p><p>Labrum wider than long, anterior margin not emarginate, basal 0.5 microreticulate, distal 0.5 smooth and shiny, sparsely punctate and pubescent laterally. Clypeus and frons densely punctate and pubescent.</p><p>Pronotum (PL 0.5 mm, PW 0.6 mm) broadest at basal 0.3, slightly narrowed posteriorly and distinctly attenuate anteriorly, posterior corners rectangular. Disc smooth and shiny, densely pubescent shallowly punctate. Areas near anterior and posterior corners coarsely granulate. Median groove short and shallowly impressed, extending from basal 0.4–0.7. Lateral carinae present in basal 0.4, joined by a distinct mesial groove. Base of pronotum with a group of granules medially.</p><p>Elytra (EL 1.2 mm, EW 0.75 mm) broadest at basal 0.67, slightly narrowed anteriorly and distinctly attenuate posteriorly, apices narrowly conjointly rounded. Strial punctures large in basal 0.5, separated by 0.5–1.0 × their diameters; punctures becoming smaller and well separated in distal 0.5. Intervals smooth and shiny, each with a row of setae; intervals 7 and 8 distinctly carinate, extending from base to declivity. Base of elytra narrowly granulate. Lateral margins distinctly denticulate.</p><p>Prosternum coarsely punctate and pubescent. Prosternal process distinctly narrowed from base to narrowly rounded apex; lateral margin distinctly rimmed. Disc convex, coarse, sparsely punctate and pubescent. Metaventrite slightly and broadly impressed, disc smooth and shiny, sparsely punctate and pubescent; median groove extending from base to anterior 0.1, gradually narrowed towards apex; lateral area densely pubescent and sparsely granulate; each side with two large elongate pits behind mesocoxae and a row of large punctures is in front of metacoxae.</p><p>Disc of ventrite I convex, largely smooth and shiny, sparsely punctate; along anterior margin slightly impressed with densely arranged, deeply impressed grooves; laterally bordered by strong admedian carinae. Discs of ventrites II – IV and basal 0.2 of ventrite V smooth and shiny; sparsely punctate and pubescent. Lateral sides of ventrites I – IV densely pubescent and sparsely granulate. Apex of ventrite V subtruncate, lateral apical corners rounded, bearing some stiff setae. Protibia distinctly enlarged.</p><p>Aedeagus. 0.69 mm long, elongate. Penis ~ 3.5 × as long as phallobase, very slightly asymmetrical, subparallel in basal 0.4, then gradually narrowed; apex subacute. Sclerotizations of endophallus in basal 0.5 of penis, consisting of a pair of elongate ovoid structures (in basal 0.3–0.5) and a pair of long thin contiguous basal rods. Parameres short, reaching apical 0.33 of penis.</p><p>Females. Protibiae not distinctly enlarged. Sexual dimorphism of metaventrite and abdominal ventrites poorly developed; ventrite I anteriorly less strongly impressed and less distinctly wrinkled; apex of ventrite V variable in both sexes: weakly emarginate, subtruncate or weakly rounded.</p><p>Measurements.</p><p>Males: BL 1.50–1.86 mm (n = 18), BW 0.75–0.80 mm (n = 6); females: BL 1.68–1.86 mm (n = 17), BW 0.75–0.80 mm (n = 3).</p><p>Distribution.</p><p>Gansu, Shaanxi, Sichuan, Yunnan,? Himalaya.</p><p>Comparison with Himalayan specimens (Himachal Pradesh, Nepal, Bhutan) indicates that Zaitzevia hybrida seems to occur there as well. However, it will be necessary to confirm this assumption by molecular data.</p><p>Variability.</p><p>Due to the morphological variability of the material examined (e. g., protibiae, elytral intervals, aedeagi), we do not designate the specimens from Gansu, Sichuan, and Yunnan as paratypes. Examination by molecular methods would be highly desired. The single male from Yulongshan (Yunnan) is 1.68 mm long and its aedeagus measures 0.69 mm; its external morphology agrees quite well with specimens of Z. hybrida from Gansu, Shaanxi, and Sichuan. However, the three males from Gaoligongshan (Yunnan) are 1.46–1.64 mm long, and their aedeagi are slimmer and shorter (0.61–0.63 mm) than in the males from the other populations (0.68–0.70 mm). Furthermore, they differ externally in the distinctly less densely punctate elytral striae and the flatter elytral declivity; genetically, they differ in the ovoid sclerotization of the endophallus being distinctly smaller. Therefore, these nine specimens from Gaoligongshan (Yunnan) listed above may well represent a different species, which should be confirmed by DNA sequencing.</p><p>The enlargement of the male protibiae is obviously allometric. In larger specimens the tibiae are disproportionately larger than in smaller ones.</p><p>As in other species of the genus Zaitzevia, the size and number of the elytral punctures, and especially the width and convexity of the elytral intervals 2–6 are quite variable; these intervals can be flat, moderately convex or carinate, at least in the anterior 0.33. In some specimens, the elytral apices are separately rounded.</p><p>The shape of the aedeagus of Z. hybrida is normally symmetrical; in the holotype it is hardly noticeably asymmetrical, and in the two males from Gansu (CWBS 322), one is very slightly asymmetrical and the other one even more distinctly so.</p><p>Etymology.</p><p>The epithet is derived from the Latin noun hybrida (hybrid) referring to the fact that Zaitzevia hybrida and a number of other undescribed species from China and the Himalaya combine diagnostic characters of Zaitzevia and Zaitzeviaria .</p><p>Remarks.</p><p>At a first glance, Zaitzevia hybrida seems to belong to the genus Zaitzeviaria, based on the small size and the elytral carinae being located on intervals 7 and 8. On the other hand, it shares a number of characters with Zaitzevia . Therefore, and in the absence of any molecular data, we have decided to place this species tentatively in the genus Zaitzevia . For further information, see the Discussion.</p></div>	https://treatment.plazi.org/id/1015FD05253F54C68A74BD25083B13C0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Bian, Dong-Ju;Jäch, Manfred A.	Bian, Dong-Ju, Jäch, Manfred A. (2025): The species of the genus Zaitzevia Champion, 1923 (Coleoptera, Elmidae) from Shaanxi Province, China. ZooKeys 1264: 129-157, DOI: 10.3897/zookeys.1264.156144
0661EB0E5132553891AAFD57374E301B.text	0661EB0E5132553891AAFD57374E301B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zaitzevia pilosa Bian & Jäch 2025	<div><p>Zaitzevia pilosa sp. nov.</p><p>Figs 2 A, B, 6 A – C, 11 A – C</p><p>Zaitzevia sp.: Jäch and Boukal 1995: figs 36, 37.</p><p>Material examined.</p><p>(237 exs) Holotype: China • ♂ (IAECAS): “ China: Shaanxi \ Qinling, Ningshan County \ Ningdong Forest Agency | Dacigou, 2005. VI. 11 \ 1437 m, Leg. Wangm [Wang Miao] ” . Paratypes: China, Shaanxi • 3 ♀♀ (IAECAS), the same data as holotype • 1 ♀ (IAECAS): “ China: Shaanxi \ Qinling, NingShan County, \ Ningdong Forest Agency | Dachigou \ 2005.5. 11 \ 1398 m \ Leg. Bian ” • 3 ♂♂, 1 ♀ (IAECAS): “ China: Shaanxi, Ankang City, \ Ningshan County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.5468&amp;materialsCitation.latitude=33.557884" title="Search Plazi for locations around (long 108.5468/lat 33.557884)">Xunyang Dam</a> | 33°33.473′N, 108°32.808′E, \ 1355 m, 2019.8. 20 \ Leg. Tong Y. F ” • 2 ♂♂, 3 ♀♀ (NMW): “ China: Shaanxi, 5. 6. 1998 \ Ningshan Co., ca. 1900 m \ 5 km NW Huoditang \ leg. M. Wang (CWBS 313) ” • 18 exs (NMW): “ China: Shaanxi, 5. 6. 1998 \ Ningshan Co., ca. 1650 m \ 7 km NW Huoditang \ leg. M. Wang (CWBS 314) ” • 27 exs (NMW): “ China: Shaanxi, 6. 6. 1998 \ Ningshan Co., ca. 1 500 m \ 10 km NE Xunyangba \ leg. M. Wang (CWBS 315) ” ; Hubei • 11 exs (IAECAS): “ China: Hubei \ Shennongjia For. Dist. | 1600 m, 2004.10. 9 \ Leg. Wang (CWBS 522) ” • 10 exs (NMW): “ China: Hubei, 9. 10. 2004 \ Shennongjia Forest Distr. \ 10 km E Muyu, Chaiqi \ 1600 m, leg. Schönmann, \ &amp; Wang (CWBS 522) ” • 4 exs (NMW): “ China: Hubei, 10. 10. 2004 \ Shennongjia Forest Distr. \ 3 km N Muyu, Duanjiang \ 1300 m, leg. Schönmann \ &amp; Wang (CWBS 523) ” • 3 exs (NMW): “ China: Hubei, 10. 10. 2004 \ Shennongjia Forest Distr. \ 5 km SW Muyu \ 1350 m, leg. Schönmann \ &amp; Wang (CWBS 525) ” • 4 ♂♂, 4 ♀♀ (IAECAS): “ China: Hubei \ Shennongjia Forest Dist. | 2004.10. 11 \ Leg. Wang (CWBS 527) ” • 7 exs (NMW): “ China, Hubei, 11. 10. 2004 \ Shennongjia Forest Distr. \ 4 km NW Muyu \ 1800 m, leg. Schönmann \ &amp; Wang (CWBS 527) ” • 5 exs (NMW): “ China: Hubei, 11. 10. 2004 \ Shennongjia Forest Distr. \ 3 km N Muyu, Chaiqi env. \ 1 700 m, leg. Schönmann \ &amp; Wang (CWBS 528) ” • 2 exs (NMW): “ China: Hubei, 12. 10. 2004 \ Shennongjia Forest Distr. \ 20 km E Muyu, 10 km E \ Chaiqi, 2150 m, leg. Wang \ &amp; Schönmann (CWBS 529) ” • 4 exs (NMW): “ China: Hubei, 15. 10. 2004 \ Shennongjia Forest Distr. \ 25 km W pass Muyu - \ Shennongjia \ 1650 m, leg. Schönmann \ &amp; Wang (CWBS 536) ” • 2 exs (NMW): “ China: Hubei, 17. X. 2004 \ Shennongjia Forest Distr. \ 35 km N Muyu \ 1600 m, leg. Schönmann \ &amp; Wang (CWBS 539) ” • 23 exs (IAECAS): “ China: Hubei, Enshi \ 35 km N Muyu | 1600 m, 2004.10. 17 \ Leg. Wang (CWBS 540) ” • 5 ♂♂, 48 exs (IAECAS): “ China: Hubei, Enshi \ Lichuan | 1600 m, 2004.10. 23 \ Leg. Wang (CWBS 549) ” • 33 exs (NMW): “ China: Hubei, 23. 10. 2004 \ 50 km W Enshi \ 1 600 m, leg. Schönmann \ &amp; Wang (CWBS 549) ” ; Hunan • 10 exs (NMW): “ China, NW-Hunan 1993 \ Wulingyuan, N Dayong \ Zangjiajie [ Zhangjiajie], 29.10., 650 m \ leg. Schillhammer (1) [CWBS 20] ” • 12 exs (NMW): same locality data, but “ leg. Schönmann ” • 1 ♂, 1 ♀ (IAECAS): “ China: Hunan, Zhangjiajie \ Suoxiyu Nat. Res. | 650 m, 1993.10. 29 \ Leg. L. Ji (CWBS 21) ” • 1 ♂, 1 ♀ (NMW): “ China, NW-Hunan 1993 \ Wulingyuan, N Dayong \ Zangjiajie [ Zhangjiajie], 29.10., 650 m \ leg. L. Ji (2) [CWBS 21] ” .</p><p>Differential diagnosis.</p><p>This species is well characterized by its size (BL 2.4–2.9 mm) in combination with the conspicuous dorsal pilosity (if not rubbed off, as in many older specimens) and the granules on the elytral interval 5 more or less reaching the elytral base.</p><p>In habitus and body size the new species vaguely resembles Z. yingzuijieensis . It can be distinguished from the latter mainly by the conspicuous pubescence, the black dorsum (dark brown in Z. yingzuijieensis), the narrow median pronotal sulcus extending from basal 0.2–0.6 (&lt;0.33 length of pronotum in Z. yingzuijieensis), the distinct sublateral pronotal carinae (faint in Z. yingzuijieensis), the apex of the prosternal process being narrowly rounded (broadly rounded in Z. yingzuijieensis), the symmetrical aedeagus (asymmetrical in Z. yingzuijieensis), and several differences in the endophallus.</p><p>Description.</p><p>Male (holotype). BL 2.5 mm, BW 1.1 mm. Body slender. Dorsal surface black, ventral surface dark brown to black. Legs dark brown to black but tarsi reddish brown. Antennae yellowish brown.</p><p>Labrum wider than long, microreticulate in basal 0.33 but smooth and shiny in distal 0.67; punctures and pubescence moderately dense. Anterior margin arched, not emarginate, densely pubescent laterally. Clypeus and frons densely pubescent and granulate. Clypeal suture shallowly impressed.</p><p>Pronotum (PL 0.7 mm, PW 0.8 mm) subparallel in basal 0.4, distinctly attenuate anteriorly. Lateral margin narrowly rimmed, anterior 0.5 slightly serrate. Anterior corners acute, slightly produced; posterior corners almost right-angled. Disc smooth and shiny, densely punctate and covered by golden setae. Median sulcus extending from basal 0.2–0.6. Sublateral carinae present in basal 0.4. Areas near anterior and posterior corners densely and finely granulate.</p><p>Elytra (EL 1.8 mm, EW 1.1 mm) broadest at anterior 0.67, slightly narrowed anteriorly and distinctly attenuate posteriorly, apices separately rounded. Strial punctures moderately large in basal 0.5, separated by 1–2 × their diameters, distinctly smaller and well separated in distal 0.5. Intervals flat, sparsely punctate, each with a row of moderately long setae. Carinae on intervals 5, 7 and 8 complete.</p><p>Prosternum laterally densely pubescent; prosternal process gradually narrowed from base to basal 0.8, distinctly narrowed in distal 0.2, apex narrowly rounded; lateral margin distinctly rimmed; disc rugose, sparsely punctate and pubescent. Metaventrite broadly and shallowly impressed, disc smooth and shiny, sparsely punctate, and pubescent, somewhat rugose in posterior 0.5; median sulcus broad in posterior 0.8, and narrowed in anterior 0.2; with a row of large punctures behind of mesocoxae and a shallow groove in front of metacoxae; base with a pair of large pits on each side of median sulcus.</p><p>Ventrite I with a pair of admedian carinae; disc distinctly impressed, microreticulate and sparsely punctate. Middle of ventrites II – IV and anterior 0.25 of ventrite V smooth and shiny, sparsely punctate and sparsely pubescent. Lateral areas of ventrites I – V densely pubescent. Distal 0.75 of ventrite V densely granulate and sparsely pubescent. Lateral margins of ventrite V fringed with short spines; apex of ventrite V widely and deeply emarginate medially; lateral corners forming long teeth.</p><p>Aedeagus. 1.1 mm long, elongate, cylindrical. Penis ~ 2.5 × as long as phallobase, subparallel in basal 0.7, distinctly narrowed in distal 0.3, apex acute. Sclerotizations of endophallus somewhat resembling a pair of overlapping banana leaves; apex of endophallus with a pair of large curved double-pointed teeth. Parameres very slim, reaching apical 0.3 of penis.</p><p>A specimen from Hunan (CWBS 20) with everted endophallus has been illustrated by Jäch and Boukal (1995: figs 36, 37); unfortunately, this particular specimen could not be traced and is therefore not designated as paratype.</p><p>Females. Secondary sexual dimorphism rather poorly developed. Elytra apically often, but not always, more acuminate than in males. Metaventrite, including discrimen, a little less strongly or less comprehensively impressed. Disc of ventrite I less distinctly impressed. Ventrite V less widely and less deeply excised apically, lateral apical teeth distinctly shorter and smaller than in males; apical 0.33 of ventrite V less strongly granulate. Femora on average insignificantly slimmer than in males.</p><p>Measurements.</p><p>Males: BL 2.5–2.8 mm, BW 1.1–1.2 mm (n = 10); females: BL 2.4–2.9 mm, BW 1.1–1.2 mm (n = 12).</p><p>Variability.</p><p>Elytral apices of males usually conjointly rounded, but sometimes they are separately rounded or slightly excised. In some specimens from Hubei (e. g., CWBS 529, 549), the elytral intervals are tendentially more convex, and specimens from Hunan (CWBS 20) differ from the specimens from Shaanxi in the usually more glabrous surface, larger and deeper elytral punctures, and more convex elytral intervals, and the larger aedeagus (~ 1.30–1.36 mm long).</p><p>Distribution.</p><p>Hubei, Hunan, Shaanxi.</p><p>Habitat.</p><p>This species is normally found in cold mountains streams, but 23 specimens were surprisingly collected in a hydrothermal spring in Hubei (CWBS 540).</p><p>Etymology.</p><p>The epithet is derived from the Latin adjective pilosus (hairy, pilose) and refers to the pronounced pilosity on the dorsal body surface often observed in fresh specimens.</p></div>	https://treatment.plazi.org/id/0661EB0E5132553891AAFD57374E301B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Bian, Dong-Ju;Jäch, Manfred A.	Bian, Dong-Ju, Jäch, Manfred A. (2025): The species of the genus Zaitzevia Champion, 1923 (Coleoptera, Elmidae) from Shaanxi Province, China. ZooKeys 1264: 129-157, DOI: 10.3897/zookeys.1264.156144
74FFDBBBBD765F82ABB5B71AF1F41256.text	74FFDBBBBD765F82ABB5B71AF1F41256.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zaitzevia robusta Bian & Jäch 2025	<div><p>Zaitzevia robusta sp. nov.</p><p>Figs 1 A – D, 5 A – C, 10 A – C</p><p>Material examined.</p><p>(41 exs) Holotype: China • ♂ (IAECAS): “ China: Shaanxi, \ Foping County \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.96555&amp;materialsCitation.latitude=33.571667" title="Search Plazi for locations around (long 107.96555/lat 33.571667)">Gaozhuanggou</a> | 33°34′18″N, 107°57′56″E \ 1035 m, 2019. VI. 21 \ Leg. Tong ” . Paratypes: China, Shaanxi • 1 ♂, 5 ♀♀ (IAECAS): “ China: Shaanxi \ Qinling, NingShan County, \ Ningdong Forest Agency | Dacigou \ 2005.5. 11 \ 1398 m \ Leg. Wang ” • 5 ♀♀ (IAECAS): “ China: Shaanxi \ Qinling, Ningshan County \ Ningdong Forest Agency | Dacigou, 2005. VI. 11 \ 1437 m, Leg. Wangm [Wang Miao] ” • 2 ♂♂ (IAECAS): “ China – Shaanxi \ Zhouzhi County l. w [leg. Wang] | Houzhenzi Ca 1200 m \ 2. VI 1998 ” [CWBS 308] • 2 ♀♀ (NMW): “ CHINA: Shaanxi, 2. 6. 1998 \ Zhouzhi County, ca. 1200 m \ 2 km W Houzhenzi Nat. Res. \ leg. M. Wang (CWBS 308) ” • 1 ♀ (NMW): “ China: Shaanxi, 3. 6. 1998 \ Houzhenzi County, ca. 1300 m \ 2 km E Houzhenzi Nat. Res. \ leg. M. Wang (CWBS 309) ” • 1 ♂, 2 ♀♀ (NMW): “ China: Shaanxi, 9. 6. 1998 \ Tiantai Shan Forest Park \ Feng Co., 7 km NE Qinling \ Train Station, ca. 2000 m \ leg. M. Wang (CWBS 319) ” • 1 ♂ (IAECAS): “ China – Shaanxi \ Feng County I. [leg.] W [Wang] | Qinling Train Station \ ca. 1900 m; 9. VI. 1998 ” [CWBS 320] • 1 ♂ (NMW): “ China: Shaanxi, 10. 6. 1998 \ Tiantai Shan Forest Park \ Feng Co., 7 km NE Qinling \ Train Station, ca. 1850 m \ leg. M. Wang (CWBS 321) ” • 2 ♂♂, 5 ♀♀ (NMW), 1 ♂, 1 ♀ (CPE): “ China: Shaanxi, Qin Ling Shan \ 110.06E, 34.27N \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.06&amp;materialsCitation.latitude=34.27" title="Search Plazi for locations around (long 110.06/lat 34.27)">Hua Shan Mt. N Valley</a>, 1200–1400 m \ 180 km E Xian, sifted [crossed out with black pen] 18. / 20.08. 1995, leg. A. Pütz ” – type locality of Elmomorphus catenatus Selnekovič, Jäch &amp; Kodada, 2024 ( Dryopidae) ; Sichuan • 1 ♂ (IAECAS): “ China: Sichuan \ Aba Pref., Mao County \ Daguan Town | <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.67188&amp;materialsCitation.latitude=31.93547" title="Search Plazi for locations around (long 103.67188/lat 31.93547)">Dadian Vill.</a> 2012.04 [day not mentioned on label] \ 31°56'7.7"N, 103°40'18.8"E \ ca. 1830 m, leg. Wang &amp; Guo ” • 1 ♂, 3 ♀♀ (NMW): “ China: Sichuan, 29. 7. 1998 \ Mao Xian Co., Jiuding Shan \ 7 km NE Mao Xian, ca. 1850 m \ Schönmann, Ji, Wang (CWBS 335) ” • 1 ♂ (IAECAS): “ China-Sichuan \ Mao cty Jiudingshan | 1950 m leg Ji &amp; W [Wang] \ 7.29. 1998 ” [CWBS 336] • 1 ♂, 1 ♀ (NMW): “ China: Sichuan, 29. 7. 1998 \ Mao Xian Co., Jiuding Shan \ 10 km NE Mao Xian, ca. 1950 m \ Schönmann, Ji, Wang (CWBS 336) ” • 1 ♂ (NMW): “ China: Sichuan, 30. 7. 1998 \ Mao Xian Cty., Jiuding Shan \ 6 km NE Mao Xian, ca. 1750 m \ Schönmann, Ji, Wang (CWBS 340) ” ; Hubei • 1 ♂ (NMW): “ China: Hubei, 19. 10. 2004 \ Tienchanghuang Forest Park \ Enshi [Pref.], 60 km SSW Badong \ 1 600 m, leg. Schönmann \ &amp; Wang (CWBS 543) ” .</p><p>1 Zaitzevia fengtongzhaiensis is obviously wide-spread in China. We have examined specimens from three provinces, deposited in the NMW and CPE: Gansu (first record, 1 ♂, CWBS 322), Sichuan (37 exs, CWBS 227, 332, 337–339; 3 exs, Ganzi Pref., Gongga Shan, 29.36°N, 102.04°E, 2,100 m a. s. l., 28. / 31. V. 1997, leg. E. Pütz; 3 exs, Qingchengshan, 65 km NW Chengdu, 10 km W Taiping, ca 600 m a. s. l., 4. V. 1997, leg. E. Pütz), and Yunnan (first record, 1 ♂, 2 ♀♀, CWBS 56). We have examined one male from Hubei (CWBS 522), which agrees in most principal characters with the specimens from Gansu, Sichuan, and Yunnan, but its endophallic teeth are larger and straighter, and the apex of the penis is slightly wider. Probably, this specimen represents a closely related, undescribed species. More material and molecular data are needed to verify this assumption.</p><p>Differential diagnosis.</p><p>Zaitzevia robusta sp. nov. is the largest species (BL 3.14–3.80 mm) of the genus described so far. The new species resembles Z. chenzhitengi, Z. fengtongzhaiensis, Z. muchenae, and Z. sichuanensis . Males can be distinguished from Z. chenzhitengi and Z. fengtongzhaiensis mainly by the longer aedeagus, which is distally not acuminate but apically rounded in ventral / dorsal views, and the elytral apices are more acuminate in Z. fengtongzhaiensis and the holotype of Z. chenzhitengi; males are distinguished from Z. muchenae by the more robust body, the longer and broader pronotal median sulcus and by the distinctly longer and broadly rounded aedeagus; males differ from Z. sichuanensis in the larger and more robust body, and the straight and longer, apically broadly rounded aedeagus; from Z. triangularis (BL: 2.9–3.2 mm) males differ in the slightly larger body size, the less acuminate elytral apices, and in the broadly rounded, not triangular aedeagal apex.</p><p>Description.</p><p>Male (holotype). BL 3.6 mm, BW 1.5 mm. Body elongate obovate. Antennae yellowish brown, dorsum black, ventral surface reddish brown to dark brown, and legs reddish brown.</p><p>Labrum smooth and shiny, with only a few punctures and scarce pubescence, lateral margins densely covered with long setae; anterior margin broadly rounded, not emarginate. Clypeus and frons densely punctate and pubescent, sparsely granulate.</p><p>Pronotum (PL 0.9 mm, PW 1.1 mm) broadest at basal 0.3, slightly narrowed posteriorly and distinctly attenuate anteriorly. Disc in basal 0.3 rugose, sparsely punctate and pubescent, in distal 0.7 smooth and shiny, densely punctate and sparsely pubescent. Lateral areas densely granulate and sparsely pubescent. Anterior corners acute, slightly produced, posterior corners almost right-angled. Lateral margin narrowly rimmed. Median sulcus extending from basal 0.2–0.8, broadest at basal 0.3, gradually narrowed anteriorly and posteriorly. Sublateral carinae present in basal 0.4 bending outwards at basal 0.3.</p><p>Elytra (EL 2.7 mm, EW 1.5 mm) broadest at distal 0.33, slightly narrowed anteriorly and distinctly attenuate posteriorly. Lateral margin serrate. Strial punctures small in basal 0.5, separated by 0.5 their diameters, punctures in distal 0.5 smaller and well separated (1–3 × their diameters). Intervals rugose, sparsely punctate and pubescent. Intervals 2–4 slightly elevated in basal 0.2. Intervals 5, 7 and 8 carinate. Elytral apices separately rounded, densely granulate.</p><p>Prosternum with disc rugose, almost glabrous; lateral parts of prosternum granulate and pubescent. Prosternal process gradually narrowed from base to broadly rounded apex; disc glabrous, surface coarse, with a few small setae and scattered granules; lateral margins distinctly rimmed. Metaventrite broadly impressed medially, disc smooth and shiny, with some longitudinal rugosities; median sulcus broad in posterior 0.8 and narrow in anterior 0.2; lateral areas densely pubescent and sparsely granulate; with a row of large punctures behind of mesocoxa and another row of small punctures in front of metacoxa; base with a pair of pits on each side of median sulcus.</p><p>Ventrite I with a pair of admedian carinae; area between these carinae (disc) flat or slightly concave, anteriorly rugose, remaining parts with small punctures, interstices micropunctate. Lateral areas of ventrites I – V densely pubescent and sparsely granulate. Middle of ventrites II – IV and basal 0.5 of ventrite V glabrous and sparsely punctate. Distal 0.5 of ventrite V densely granulate; lateral margins sometimes fringed with very short, inconspicuous spines; apex deeply emarginate medially, lateral corners produced, tooth-like and slightly bent ventrad.</p><p>Mesal margin of mesotibia slightly expanded around middle.</p><p>Aedeagus. 1.9 mm long, elongate, penis ~ 2.2 × as long as phallobase. Lateral margins of penis in ventral / dorsal view more or less subparallel except for a very inconspicuous swelling and / or a very shallow emargination at apical 0.3; dorsal wide with a conspicuous gibbosity of variable size at apical 0.25 (best seen in lateral view); apex of penis broadly and evenly rounded in ventral / dorsal view, distinctly curved dorsad in lateral view. Endophallus in distal 0.33 with a pair of small curved teeth. Parameres short and inconspicuous, clinging to penis.</p><p>Females. Elytral apices not separately rounded but distinctly acuminate and produced to a remarkably variable extent; they vary from short to long and beak-like, and sometimes they are asymmetrically produced, being longer on one side than on the other. Metaventrite flat or slightly depressed, distinctly less strongly impressed than in males, smoother. Disc of ventrite I more or less convex. Ventrite V less widely and less deeply excised apically, lateral apical teeth shorter and smaller than in males; apical 0.33 of ventrite V less strongly granulate. The femora often appear to be more strongly-built in the males, but this depends largely on the body size (in larger specimens the femora are proportionately thicker).</p><p>Measurements.</p><p>Males: BL 3.50–3.80 mm (n = 14), BW 1.40–1.55 mm (n = 5); females: BL 3.14–3.78 mm (n = 25), BW 1.40–1.60 mm (n = 11).</p><p>Distribution.</p><p>Hubei, Shaanxi, Sichuan.</p><p>Etymology.</p><p>The name robustus, a Latin adjective, refers to the massive body.</p></div>	https://treatment.plazi.org/id/74FFDBBBBD765F82ABB5B71AF1F41256	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Bian, Dong-Ju;Jäch, Manfred A.	Bian, Dong-Ju, Jäch, Manfred A. (2025): The species of the genus Zaitzevia Champion, 1923 (Coleoptera, Elmidae) from Shaanxi Province, China. ZooKeys 1264: 129-157, DOI: 10.3897/zookeys.1264.156144
