taxonID	type	description	language	source
8FC0F6608760531E944D48839959073A.taxon	description	Description. Size and form: Body length 2.0 mm. Body elongate oval, strongly convex (Fig. 8 E). Color and punctation: Dorsal and ventral surfaces of body dark brown, with lateral margins of prothorax only very slightly paler (Fig. 8 D, E); mouthparts and antennae yellow, with slightly darker antennal club and apical third of maxillary palpomere IV; legs orange with paler tarsi (Fig. 8 F). Ground punctation on head, pronotum and elytra moderately marked (Fig. 8 D). Head: Eyes in dorsal view with anterior margin oblique (anteriorly directed), and outer margins slightly bulging from outline of head; in lateral view, eyes not emarginate. Thorax: Elytra with well-defined rows of serial punctures, moderately impressed, forming grooves along apical 3 / 4 of elytra (Fig. 8 D); interserial punctures somewhat irregularly distributed (Fig. 8 D). Metafemora mostly glabrous on anterior face, with narrow band of pubescence along basal third of dorsal margin (Fig. 8 D). Elevation of mesoventrite forming a low transverse carina (Fig. 8 D). Metaventrite with distinct median, longitudinal, narrow glabrous area extending along posterior half (Fig. 8 D). Abdomen: Abdominal ventrites uniformly and very densely pubescent. Aedeagus (Fig. 11 K). Basal piece 0.4 x the length of a paramere; parameres nearly 1 / 3 as narrow as greatest width of median lobe, with outer margins widely and uniformly convex, and rounded apex; median lobe roughly triangular, rounded and slightly pinched at apex; gonopore situated nearly at midlength of median lobe.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
8FC0F6608760531E944D48839959073A.taxon	etymology	Etymology. Noun in apposition. Named after the Akoerio, an indigenous nomadic tribe, with only few people remaining in the South of Suriname.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
8FC0F6608760531E944D48839959073A.taxon	distribution	Distribution. The species is only known from an exposed rocky summit in the Grensgebergte Mountains along the border between Suriname and Brazil. See Fig. 13.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
5429D85BD30B57C3BB2AE8D7FC4BE1AF.taxon	description	Description. Size and form: Body length 1.8 - 2.0 mm. Body elongate oval, moderately convex (Fig. 7 B). Color and punctation: Dorsal and ventral surfaces of body, dark brown, with slightly paler margins of pronotum (Fig. 7 A, B); mouthparts yellowish brown; antennae light brown; legs orange with yellow tarsi (Fig. 7 C). Ground punctation on head, pronotum and elytra rather shallowly marked. Head: Eyes in dorsal view with anterior margin oblique (anteriorly directed; e. g., Fig. 2 D), and outer margins slightly bulging from outline of head; in lateral view, eyes not anteriorly emarginate (e. g., Fig. 2 E). Thorax: Elytra with all kinds of punctures similar in size and degree of impression (Fig. 7 A); serial punctures aligned in rows, slightly impressed, forming shallow longitudinal grooves; interserial punctures somewhat irregularly distributed in a single row (Fig. 7 A). Metafemora mostly glabrous on anterior face, with hydrofuge pubescence along basal third of antero-dorsal margin (Fig. 7 C). Elevation of mesoventrite forming a low longitudinal bulge (Fig. 7 C). Metaventrite with distinct median, longitudinal, narrow glabrous area extending along posterior half (Fig. 7 C). Abdomen: Abdominal ventrites uniformly and very densely pubescent. Aedeagus (Fig. 11 F). Basal piece 0.3 x the length of a paramere; greatest width of a paramere nearly 0.5 x greatest width of median lobe; outer margins of parameres nearly straight, only slightly curved inwards along apical region; apex of paramere rounded; median lobe roughly triangular, widely rounded at apex; gonopore situated at apical fourth of median lobe.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
5429D85BD30B57C3BB2AE8D7FC4BE1AF.taxon	etymology	Etymology. Named after Becky Anthony, program and meetings manager at the Entomological Society of America (ESA), in recognition of all her hard work in service to the society and the entomological community.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
5429D85BD30B57C3BB2AE8D7FC4BE1AF.taxon	distribution	Distribution. Only known from a single locality just south of the Orinoco River along the northwestern edge of the Guiana Shield. See Fig. 13.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
373AD26035E55DE88C1F496B981C165A.taxon	description	Description. Size and form: Body length 1.6 - 1.8 mm. Body elongate oval, strongly convex (Fig. 10 A, B). Color and punctation: Dorsal and ventral surfaces of body dark brown, with lateral margins of prothorax and elytra only slightly paler (Fig. 10 A, B); mouthparts yellow, with slightly darker apical third of maxillary palpomere IV; antennae brown; legs reddish to dark brown with paler tarsi (Fig. 10 C). Ground punctation on head, pronotum and elytra moderately marked (Fig. 10 A, B). Head: Eyes in dorsal view with anterior margin oblique (anteriorly directed), and outer margins slightly bulging from outline of head; in lateral view, eyes not emarginate (see Fig. 2 E). Thorax: Elytra with slightly defined rows of shallow serial punctures, not forming grooves (Fig. 10 A); interserial punctures somewhat irregularly distributed in two or three rows (Fig. 3 I). Elevation of mesoventrite forming a very low transverse carina (Fig. 10 C). Metaventrite with distinct median, longitudinal, narrow glabrous area extending along posterior half (Fig. 10 C). Abdomen: Abdominal ventrites uniformly and densely pubescent. Aedeagus (Fig. 11 M). Basal piece 0.4 x the length of a paramere; parameres nearly 1 / 3 as narrow as greatest width of median lobe, with outer margins widely and uniformly convex, and rounded apex; median lobe roughly triangular, rounded and slightly pinched at apex; gonopore situated nearly at midlength of median lobe.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
373AD26035E55DE88C1F496B981C165A.taxon	etymology	Etymology. Noun in apposition. Named after the Arawak, an indigenous tribe of northern South America.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
373AD26035E55DE88C1F496B981C165A.taxon	distribution	Distribution. Tobochares arawak is only known from the Upper Potaro region in Guyana. See Fig. 13.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
21DDD52DBD4F5409AE901B50F0236358.taxon	description	Description. Size and form: Body length 2.0 - 2.2 mm. Body elongate oval, moderately convex (Fig. 7 D). Color and punctation: Dorsal and ventral surfaces of body dark brown, with prothorax and lateral margins of elytra slightly paler (Fig. 7 D, E); mouthparts and antennae yellow, with slightly darker apical third of maxillary palpomere IV; legs orange to reddish brown with paler tarsi (Fig. 7 E). Ground punctation on head, pronotum and elytra rather shallowly marked. Head: Eyes in dorsal view with anterior margin oblique (anteriorly directed), and outer margins slightly bulging from outline of head (e. g., Fig. 2 D); in lateral view, eyes not emarginate (Fig. 2 E). Thorax: Elytra with longitudinal rows of shallow punctures, not forming grooves; punctures in two different sizes: serial punctures larger, interserial punctures smaller and denser (Fig. 3 B, G). Metafemora mostly glabrous on anterior face (Fig. 7 F). Elevation of mesoventrite forming a low transverse carina (Fig. 7 F). Metaventrite with distinct median, longitudinal, narrow glabrous area extending along posterior half (Fig. 7 F). Abdomen: Abdominal ventrites uniformly and very densely pubescent (Fig. 7 F). Aedeagus (Fig. 11 G). Basal piece nearly 0.5 x the length of a paramere; parameres nearly 0.4 x greatest width of median lobe, with outer margins weakly and uniformly convex, and rounded apex; median lobe roughly triangular, widely rounded at apex; gonopore situated nearly at apical third of median lobe.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
21DDD52DBD4F5409AE901B50F0236358.taxon	etymology	Etymology. Noun in apposition. Named after Atures, the municipality where the type locality is situated.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
21DDD52DBD4F5409AE901B50F0236358.taxon	distribution	Distribution. This species is known from several localities along the northwestern edge of the Guiana Shield in Venezuela. See Fig. 13.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
8D05A782189D587D80604D432D27B950.taxon	description	Description. Size and form: Body length 1.6 - 2.0 mm. Body elongate oval, strongly convex (Fig. 4 A, B). Color and punctation. Head dark brown, pronotum and elytra uniformly orange brown; antennae, mouthparts and legs orange to yellowish brown, with paler tarsi (Fig. 4 C). Ground punctation on head, pronotum and elytra moderately marked. Head: Eyes in dorsal view with anterior margin oblique, posteriorly directed, and outer margins continuous with outline of head (as in Fig. 2 A); in lateral view, eyes emarginate to about half the length of eye (as in Fig. 2 B). Maxillary palps slender, nearly as long as the width of the head, uniformly yellow in color (as in Fig. 2 A). Thorax: Elytra with slightly defined rows of punctures, not forming grooves (Fig. 4 A). Elevation of mesoventrite forming a somewhat transverse bulge (Fig. 4 C). Metaventrite with distinct median, broad, ovoid glabrous area extending along posterior two thirds (Fig. 4 C). Abdomen: Abdominal ventrites uniformly and very densely pubescent. Aedeagus (Fig. 11 A) with basal piece 0.25 x the length of a paramere; widest point of parameres (near base) nearly 2 / 3 greatest width of median lobe (near base), with outer margins slightly sinuate, and rounded apex; median lobe roughly triangular, dorsally concave, apical region nearly half as wide as base, broadly rounded at apex; gonopore situated at apex of median lobe.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
8D05A782189D587D80604D432D27B950.taxon	etymology	Etymology. Named after Cesar J. Benetti, Brazilian specialist on aquatic beetles, in honor of his contributions to Neotropical beetle taxonomy and for all his assistance in the field.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
8D05A782189D587D80604D432D27B950.taxon	distribution	Distribution. Only known from the type locality in Amazonas State, Brazil, situated slightly north of the Amazon River (Fig. 13).	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
4EF0B01948C7579E8E3FC4EE79583DCF.taxon	description	Description. Size and form: Body length 1.9 - 2.1 mm. Body elongate oval, moderately convex (Fig. 10 E). Color and punctation: Dorsal and ventral surfaces of body dark brown, with anterolateral margins of prothorax slightly paler (Fig. 10 D, E); mouthparts yellow to orange; antennae light brown; legs reddish to dark brown, distally paler (orange), with paler (yellow) tarsi (Fig. 10 F). Ground punctation on head, pronotum and elytra sharply marked; pronotal punctation dense (Fig. 10 D). Head: Eyes in dorsal view with anterior margin oblique (anteriorly directed; e. g., Fig. 2 D), and outer margins slightly bulging from outline of head; in lateral view, eyes not emarginate (e. g., Fig. 2 E). Thorax: Elytra with serial punctures similar in size and degree of impression to interserial punctures, and only seemingly aligned longitudinally, not forming grooves; interserial punctures irregularly distributed in two or three rows (Fig. 10 D). Metafemora mostly glabrous on anterior face (Fig. 10 F). Elevation of mesoventrite forming a very low transverse carina (Fig. 10 F). Metaventrite with distinct median, longitudinal, narrow glabrous area extending along posterior half (Fig. 10 F). Abdomen. Abdominal ventrites uniformly and very densely pubescent (Fig. 10 F). Aedeagus (Fig. 11 N). Basal piece nearly 0.45 x the length of a paramere; greatest width of a paramere nearly 0.6 x greatest width of median lobe; outer margins of parameres straight to widely and uniformly convex; apex of paramere obliquely rounded; median lobe roughly sagittate, rounded at apex; gonopore situated nearly at midlength of median lobe.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
4EF0B01948C7579E8E3FC4EE79583DCF.taxon	etymology	Etymology. Noun in apposition. Named after the Canaima National Park in Venezuela, where the type locality is situated.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
4EF0B01948C7579E8E3FC4EE79583DCF.taxon	distribution	Distribution. This species is known from the famous Auyan-tepui, which is also home to Angel Falls, the highest waterfall in the world. Collected at elevations of 1700 - 2170 m, this species is one of the relatively few water beetle taxa known from the " Pantepui Province ", which comprises areas of the Guiana Shield which are greater than 1500 m in elevation. See Fig. 13.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
533C525AE3A4500AB7AA779059209867.taxon	description	Description. Size and form: Body length 1.8 - 2.0 mm. Body elongate oval, somewhat parallel sided, moderately convex (Fig. 6 A, B). Color and punctation: Dorsal and ventral surfaces of body dark brown, with anterior and lateral margins of prothorax, and outer margins of elytra slightly paler (Fig. 6 A); mouthparts yellow (especially maxillary palps) to orange; antennae yellowish brown; legs orange brown with paler (yellow) tarsi (Fig. 6 C). Ground punctation on head, pronotum and elytra sharply marked. Head: Eyes in dorsal view with anterior margin oblique (anteriorly directed), and outer margins slightly bulging from outline of head (Fig. 2 H); in lateral view, eyes not emarginate (e. g., Fig. 2 E). Thorax: Elytra with all kinds of punctures similar in size and degree of impression, all seemingly longitudinally aligned and uniformly distributed, not forming grooves (Fig. 6 A). Metafemora mostly glabrous on anterior face (Fig. 6 C). Elevation of mesoventrite forming a longitudinal bulge (Fig. 3 D). Metaventrite with distinct median, longitudinal, narrow glabrous area extending along posterior half (Fig. 3 D). Abdomen: Abdominal ventrites uniformly and very densely pubescent (Fig. 6 C). Aedeagus (Fig. 11 H). Basal piece nearly 0.4 x the length of a paramere; greatest width of a paramere nearly 0.6 x greatest width of median lobe; outer margins of parameres nearly straight along basal 2 / 3, then curved inwards; apex of paramere rounded; median lobe roughly sagittate, narrow along apical third, rounded at apex; gonopore situated nearly at midlength of median lobe.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
533C525AE3A4500AB7AA779059209867.taxon	etymology	Etymology. Named with the Latin word Tobochares communis meaning common, highlighting the abundance and wide distribution of the species, which is the most commonly found.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
533C525AE3A4500AB7AA779059209867.taxon	distribution	Distribution. This species is the most widely distributed of all known Tobochares, occurring from the northwest margin of the Guiana Shield in Venezuela all the way to its eastern edge in the state of Amapa, Brazil (Fig. 12).	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
E1FB953C41AE5865B61B61D56B7AFAD8.taxon	diagnosis	Diagnosis. The Tobochares communis species group can be recognized by the straight anterior margin of the eye in lateral view (e. g., Fig. 2 E), the usually longitudinally aligned elytral punctures, the short and narrow glabrous patch on the metaventrite (Fig. 3 D, E), the high density of the hydrofuge pubescence on the abdominal ventrites, and the well-developed and numerous metatibial spines.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
B9756193795A551DA3A8B914A9482A0A.taxon	diagnosis	Diagnosis. The Tobochares emarginatus species group can be recognized by the oblique and posteriorly directed anterior margin of the eye in lateral view, which emarginates the eye to about a quarter of its dorsal width (Fig. 2 C; Kohlenberg and Short 2017: fig. 6 C-F), the presence of a broad, ovoid to diamond-shaped glabrous patch on the metaventrite (Fig. 5 F), the dense and uniform distribution of the hydrofuge pubescence on the abdominal ventrites (Fig. 5 F), and the moderate (in number and size) metatibial spines.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
745AB8B13AFA55B7BAF80954868FD0F5.taxon	description	Description. Size and form: Body length 1.7 - 2.0 mm. Body elongate oval, moderately convex (Fig. 5 D, E). Color and punctation: Dorsal surfaces of body dark brown, with slightly paler pronotum (Fig. 5 D); antennae, mouthparts, and legs yellowish brown, with orange tibiae; ventral surfaces of meso-, metathorax and abdomen dark brown (Fig. 5 F). Ground punctation on head, pronotum and elytra moderately marked. Head: Eyes in dorsal view with anterior margin oblique, posteriorly directed; Tobochares canthus emarginating eye to about a quarter of its dorsal width in lateral view (Fig. 2 C). Maxillary palps 0.7 x width of head. Thorax: Elytra with all kinds of punctures similar in size and degree of impression, seemingly evenly distributed, not forming grooves (Fig. 5 D). Elevation of mesoventrite forming a low transverse ridge (Fig. 5 F). Metaventrite with distinct median, broad, diamond-shaped glabrous area extending along posterior two thirds (Fig. 5 F). Abdomen: Abdominal ventrites uniformly and densely pubescent (Fig. 5 F). Aedeagus (Fig. 11 C). Basal piece nearly 0.4 x length of a paramere; greatest width of a paramere nearly 0.7 x greatest width of median lobe; outer margins of parameres diverging along basal half, then broadly bending inward and tapering along apical fifth; apex of paramere oblique, pointing towards longitudinal midline of aedeagus; median lobe roughly triangular, apically rounded; gonopore situated at apex of median lobe.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
745AB8B13AFA55B7BAF80954868FD0F5.taxon	etymology	Etymology. Named with the Latin word Tobochares fusus, meaning fusiform, in reference to the shape of the aedeagus of this species.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
745AB8B13AFA55B7BAF80954868FD0F5.taxon	distribution	Distribution. This species is known from two closely situated localities on either side of the Oiapoque River, the boundary between French Guiana and the Brazilian state of Amapa. See Fig. 13.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
BD22AB85DE46525EA309E7B921CBCFAA.taxon	description	Description. Dorsal surfaces of body dark brown with paler outer margins of pronotum and elytra (Fig. 4 D, E); head slightly darker; mouthparts and antennae yellowish; legs yellowish to brown with paler tarsi (Fig. 4 F). Ground punctation on head, pronotum and elytra moderately marked. Head: Eyes in dorsal view with anterior margin oblique, posteriorly directed, and outer margins continuous with outline of head; in lateral view, eyes emarginate to about half the length of eye (Fig. 2 A, B). Maxillary palps slender, slightly shorter than width of head, uniformly yellowish in color (Fig. 2 A). Thorax: Elytra with slightly defined rows of punctures, not forming grooves (Fig. 4 D). Elevation of mesoventrite with a low transverse ridge (Fig. 4 F). Metaventrite with distinct median, broad, ovoid glabrous area extending along posterior two thirds (Fig. 4 F). Abdomen: Abdominal ventrites uniformly and very densely pubescent. Aedeagus (Fig. 11 B) with basal piece 0.2 x the length of a paramere; widest point of parameres (near base) nearly as wide as basal width of median lobe, with outer margins very slightly sinuate, and rounded apex; median lobe fusiform, with widest point slightly apicad of midpoint, broadly rounded at apex; gonopore situated at apex of median lobe.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
BD22AB85DE46525EA309E7B921CBCFAA.taxon	etymology	Etymology. Named after the Brazilian state of Goias, from which the species is known.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
BD22AB85DE46525EA309E7B921CBCFAA.taxon	distribution	Distribution. Only known from the type locality in the central Brazilian state of Goias. This is the first and currently only species of Tobochares reported from south of the Amazon River (Fig. 12).	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
070C7F3CDA6D59618BFD6D730015238A.taxon	description	Description. Size and form: Body length 1.6 - 1.9 mm. Body elongate oval, strongly convex (Fig. 9 A, B). Color and punctation: Dorsal and ventral surfaces of body dark brown, prothorax (especially its lateral areas) and outer margins of elytra slightly paler (Fig. 9 A, B); mouthparts yellow (especially maxillary palps); antennae yellowish brown; legs orange brown with paler (yellow) tarsi (Fig. 9 C). Ground punctation on head, pronotum and elytra moderately marked. Head: Eyes in dorsal view with anterior margin oblique (anteriorly directed; e. g., Fig. 2 D), and outer margins slightly bulging from outline of head; in lateral view, eyes not emarginate (e. g., Fig. 2 E). Thorax: Elytral punctation uniform in size and degree of impression, not impressed to form elytral striae; serial punctures seemingly aligned in rows; interserial punctures somewhat irregularly distributed in one or two rows (Fig. 9 A). Metafemora mostly glabrous on anterior face (Fig. 9 C). Elevation of mesoventrite forming a very low transverse carina (Fig. 9 C). Metaventrite with distinct median, longitudinal, narrow glabrous area extending along posterior half (Fig. 9 C). Abdomen: Abdominal ventrites uniformly and very densely pubescent (Fig. 9 C). Aedeagus (Fig. 11 L). Basal piece nearly 0.5 x the length of a paramere; greatest width of a paramere nearly 0.5 x greatest width of median lobe; outer margins of parameres weakly convex; apex of paramere narrowly rounded; median lobe roughly sagittate, pinched and rounded at apex; gonopore situated nearly at midlength of median lobe.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
070C7F3CDA6D59618BFD6D730015238A.taxon	etymology	Etymology. Noun in apposition. Named after the Kappel airstrip, the locality where the species has been collected.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
070C7F3CDA6D59618BFD6D730015238A.taxon	distribution	Distribution. Known from two very closely situated localities adjacent to Kappel Airstrip, at the foot to Tafelberg Tepui in Suriname. See Fig. 13.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
E49F01331C215F70A43A6DE50639C97D.taxon	description	Description. Size and form: Body length 1.9 mm. Body elongate oval, strongly convex (Fig. 9 E). Color and punctation: Dorsal and ventral surfaces of body dark brown, with prothorax (especially its anterolateral margins) slightly paler (Fig. 9 D, E); mouthparts yellow (especially maxillary palps) to orange brown; antennae yellowish brown; legs orange brown with paler (yellow) tarsi (Fig. 9 F). Ground punctation on head, pronotum and elytra moderately marked. Head: Eyes in dorsal view with anterior margin oblique (anteriorly directed; e. g., Fig. 2 D), and outer margins slightly bulging from outline of head; in lateral view, eyes not emarginate (e. g., Fig. 2 E). Thorax: Elytral punctation uniform in size and degree of impression, not impressed to form elytral striae (Fig. 9 D); serial punctures seemingly aligned in rows; interserial punctures somewhat irregularly distributed in one or two rows (e. g., Fig. 3 H). Metafemora mostly glabrous on anterior face (Fig. 9 F). Elevation of mesoventrite forming a very low transverse carina (Fig. 9 F). Metaventrite with distinct median, longitudinal, narrow glabrous area extending along posterior half (Fig. 9 F). Abdomen: Abdominal ventrites uniformly and very densely pubescent (Fig. 9 F).	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
E49F01331C215F70A43A6DE50639C97D.taxon	etymology	Etymology. Noun in apposition. Named with the Surinamese word Tobochares kolokoe meaning lucky, as this species is known from a single female specimen.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
E49F01331C215F70A43A6DE50639C97D.taxon	distribution	Distribution. This species is only known from the summit of Tafelberg Tepui, a low-elevation sandstone massif in the center of Suriname (Fig. 13).	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
B8A4BDC02CC758F49A78635F7623A4C4.taxon	materials_examined	New material examined. Brazil: Roraima: Caroebe Municipality, Reservoir by Usina de Jatapu, 0.872953 °, - 59.282170 °, 185 m, 17. i. 2018, large wall seep with algae, leg. Short, Benetti, and Santana, BR 18 - 0117 - 01 A (7, INPA, SEMC including DNA Voucher SLE 1501).	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
96950BE29DDC521FA024F6DA894AE37B.taxon	description	Description. Size and form: Body length 1.7 - 2.1 mm. Body elongate oval, moderately convex (Fig. 5 B). Color and punctation: Dorsal surfaces of body dark brown, with clypeus (Fig. 2 A), outer margins of pronotum, and outer margins of elytra yellowish brown (Fig. 5 A, B); maxillary and labial palps yellow, remainder mouthparts orange; antennae yellowish brown; legs orange brown with paler (yellow) tarsi; ventral surfaces brown (Fig. 5 C). Ground punctation on head, pronotum and elytra moderately marked (Figs 2 A, 5 A). Head: Eyes in dorsal view with anterior margin oblique and anteriorly directed (Fig. 2 A, B), and outer margins slightly bulging from outline of head; in lateral view, eyes not anteriorly emarginate (Fig. 2 B). Maxillary palps as long as 0.8 x width of head. Thorax: Elytra with all kinds of punctures similar in size and degree of impression, moderately aligned in rows, not forming grooves (Fig. 5 A). Elevation of mesoventrite forming a very low transverse carina (Fig. 5 C). Metaventrite with distinct median, longitudinal, glabrous area, 3 x longer than wide, extending along posterior half (Fig. 5 C). Abdomen: Abdominal ventrites uniformly and very densely pubescent (Fig. 5 C). Aedeagus (Fig. 11 D). Basal piece 2 x longer than a paramere; greatest width of a paramere nearly 0.6 x greatest width of median lobe; outer margins of parameres slightly sinuate, inner margins strongly sinuate; apex of paramere sharply acute, pointing towards longitudinal midline of aedeagus; median lobe roughly sagittate, medially with narrow emargination extending along apical fourth; gonopore situated nearly at apical third of median lobe.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
96950BE29DDC521FA024F6DA894AE37B.taxon	etymology	Etymology. Named with the Greek words luteo, meaning yellow, and margo meaning margin, in reference to the striking yellow band surrounding the marginal areas of these beetles.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
96950BE29DDC521FA024F6DA894AE37B.taxon	distribution	Distribution. This species is known from several localities along the northwestern edge of the Guiana Shield in Bolivar State, Venezuela. See Fig. 13.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
B9E3BC4184895B7CBC486CA8F4B9F637.taxon	diagnosis	Diagnosis. Members of this species group can be recognized by the straight anterior margin of the eye in lateral view (Fig. 2 E), the longitudinally aligned and undifferentiated elytral punctures with about the same size and degree of impression (Fig. 5 A), the short and narrow glabrous patch on the metaventrite (Fig. 5 C), the scantiness of the hydrofuge pubescence on the abdominal ventrites (Fig. 5 C), and the reduced (in number and size) metatibial spines.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
73BC8FFD76D05FB5B78E4FBB26389077.taxon	description	Description. Size and form: Body length 1.6 - 1.7 mm. Body elongate oval, moderately convex (Fig. 6 E). Color and punctation: Dorsal and ventral surfaces of body dark brown, with anterior and lateral margins of prothorax slightly paler (Fig. 6 D, E); mouthparts yellow (especially maxillary palps) to orange; antennae yellowish brown; legs orange brown with paler (yellow) tarsi (Fig. 6 F). Ground punctation on head, pronotum and elytra moderately marked. Head: Eyes in dorsal view with anterior margin slightly oblique (anteriorly directed), and outer margins nearly half the greatest length of eye, slightly bulging from outline of head (Fig. 2 I); in lateral view, eyes not anteriorly emarginate (e. g., Fig. 2 E). Thorax: Elytra with all punctures about the same size and degree of impression, all seemingly aligned in rows and uniformly distributed, not forming grooves (Fig. 6 D). Metafemora mostly glabrous on anterior face (Fig. 6 F). Elevation of mesoventrite forming a broad bulge (Fig. 6 F). Metaventrite with distinct median, longitudinal, narrow glabrous area extending along posterior half (Fig. 6 F). Abdomen: Abdominal ventrites uniformly and very densely pubescent. Aedeagus (Fig. 11 I). Basal piece nearly 0.4 x the length of a paramere; greatest width of a paramere nearly 0.5 x greatest width of median lobe; outer margins of parameres uniformly weakly convex; apex of paramere rounded; median lobe roughly sagittate, narrow along apical third, rounded at apex; gonopore situated basad of midlength of median lobe.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
73BC8FFD76D05FB5B78E4FBB26389077.taxon	etymology	Etymology. Named with the combination of the Latin words micro, meaning small, and ops, meaning eyes, highlighting the small eyes of the members of the species.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
73BC8FFD76D05FB5B78E4FBB26389077.taxon	distribution	Distribution. This species is only known from the summit of Tafelberg Tepui, a low-elevation sandstone massif in the center of Suriname (Fig. 13).	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
94F0E5CDE2685078952A6F7AD3EA1C0B.taxon	description	Description. Size and form: Body length 1.7 - 1.8 mm. Body elongate oval, moderately convex (Fig. 7 H). Color and punctation: Dorsal surfaces of body orange brown, with lateral margins of prothorax slightly paler (Fig. 7 G, H); ventral surfaces of body (except prosternum) dark brown; mouthparts orange brown; antennae light brown; legs, including tarsi orange brown (Fig. 7 I). Ground punctation on head, pronotum and elytra rather shallowly marked. Head: Eyes in dorsal view with anterior margin slightly oblique (anteriorly directed; Fig. 2 D); in lateral view, eyes not anteriorly emarginate (e. g., Fig. 2 E). Thorax: Elytra with all kinds of punctures similar in size and degree of impression; serial punctures aligned in rows, not forming grooves; interserial punctures somewhat irregularly distributed in a single row (Fig. 3 F). Metafemora mostly glabrous on anterior face (Fig. 7 I). Elevation of mesoventrite forming a low transverse carina (Fig. 7 I). Metaventrite with distinct median, longitudinal, narrow glabrous area extending along posterior half (Fig. 7 I). Abdomen: Abdominal ventrites uniformly and very densely pubescent (Fig. 7 I). Aedeagus (Fig. 11 E) with basal piece nearly 0.6 x the length of a paramere; greatest width of a paramere nearly 0.7 x greatest width of median lobe; outer margins of parameres straight and slightly converging along basal 2 / 5, then uniformly and widely convex; apex of paramere rounded; median lobe roughly rectangular, with wide and short medial emargination at apex; gonopore situated at apical fourth of median lobe.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
94F0E5CDE2685078952A6F7AD3EA1C0B.taxon	etymology	Etymology. Noun in apposition. Named after the Pemon, an indigenous tribe that inhabits La Gran Sabana region in Venezuela, where Auyan Tepui is located.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
94F0E5CDE2685078952A6F7AD3EA1C0B.taxon	distribution	Distribution. This species is known from the famous Auyan-tepui, which is also home to Angel Falls, the highest waterfall in the world. Collected at an elevation of 1700 m, this species is one of the relatively few water beetle taxa known from the " Pantepui Province ", which comprises areas of the Guiana Shield which are greater than 1500 m in elevation (Fig. 13).	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
85C628544A8D5C25B4E992A035135F9E.taxon	description	Description. Size and form: Body length 2.6 mm. Body elongate oval, moderately convex. Color and punctation: Dorsal and ventral surfaces of body, antennae and legs (except yellowish tarsi) dark brown; mouthparts light brown (Fig. 8 A-C). Ground punctation on head, pronotum and elytra moderately marked. Head: Eyes in dorsal view with anterior margin oblique (anteriorly directed), and outer margins bulging from outline of head; in lateral view, eyes not anteriorly emarginate. Thorax: Elytra with all kinds of punctures similar in size and degree of impression; serial punctures aligned in rows, impressed as to form longitudinal grooves (Fig. 8 A, B); elytral stria I more strongly impressed along posterior half of elytra, resembling well-developed sutural stria (Fig. 8 A); interserial punctures somewhat irregularly distributed (Fig. 8 A, B). Metafemora mostly glabrous on anterior face, with hydrofuge pubescence along basal half of antero-dorsal margin (Fig. 8 C). Elevation of mesoventrite forming a very low transverse carina (Fig. 8 C). Metaventrite with distinct median, longitudinal, narrow glabrous area extending along posterior half (Fig. 8 C). Abdomen: Abdominal ventrites uniformly and very densely pubescent (Fig. 8 C). Aedeagus (Fig. 11 J). Basal piece nearly 0.5 x the length of a paramere; greatest width of a paramere nearly 0.4 x greatest width of median lobe; outer margins of parameres very weakly convex; apex of paramere rounded; median lobe roughly triangular, rounded at apex; gonopore situated at midlength of median lobe.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
85C628544A8D5C25B4E992A035135F9E.taxon	etymology	Etymology. Named after Rosina Romano, Director of Meetings and Membership at the Entomological Society of America (ESA), in recognition of all her hard work and dedication to the society and entomological community.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
85C628544A8D5C25B4E992A035135F9E.taxon	distribution	Distribution. This species is only known from the summit of Serra do Tepequem, a low-elevation sandstone massif in northern Brazil, near the border with Venezuela (Fig. 13).	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
93E066E5CD5D5316A45CC5D4F1BEE31C.taxon	materials_examined	New material examined. Brazil: Roraima: Amajari Municipality, Serra do Tepequem, Igarape Preto Negro, Cachoeira Laje Preta, 3 ° 36.381 ' N, 61 ° 42.878 ' W, 618 m, 14. i. 2018, leg. Short and Benetti, BR 18 - 0114 - 04 B (12, INPA, SEMC, including DNA voucher SLE 1496); Caroebe Municipality, Reservoir by Usina de Jatapu, 0.872953, - 59.282170, 185 m, 17. i. 2018, large wall seep with algae, leg. Short, Benetti, and Santana, BR 18 - 0117 - 01 A (1, SEMC including DNA Voucher SLE 1502).	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
D91EC0BFB8FD5C509475FEC68807C861.taxon	materials_examined	New material examined. Suriname: Sipaliwini: Sipaliwini Savanna Nature Reserve, 2 ° 00.240 ' N, 55 ° 58.259 ' W, 374 m, 4 - Brothers Mountains, leg. Short and Baca, 30. iii. 2017, seeps on granite SR 17 - 0330 - 04 A (1, SEMC including DNA Voucher SLE 1244).	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
C005666E376B50FC8C4C154284A560E4.taxon	diagnosis	Diagnosis. This species group can be recognized by the oblique and posteriorly directed anterior margin of the eye in lateral view, which emarginates the eye in lateral view (Fig. 2 A-C), the broad, ovoid to diamond-shaped glabrous patch on the metaventrite (Kohlenberg and Short 2017: fig. 10), the dense and uniform distribution of the hydrofuge pubescence on the abdominal ventrites (Kohlenberg and Short 2017: fig. 13 A), and the moderate (in number and size) metatibial spines.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93-140, DOI: http://dx.doi.org/10.3897/zookeys.1019.59881, URL: http://dx.doi.org/10.3897/zookeys.1019.59881
