taxonID	type	description	language	source
96247A4783207C034C9EFE59FD52F877.taxon	materials_examined	Material examined. Holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR- 97), in Crassostrea madrasensis (Preston, 1916), from Pamban Fishing Harbor, Tamil Nadu, India, 9 ° 16 ' 56 " N, 79 ° 12 ' 31 " E, 2 February 2016, coll. J. Trivedi & D. Patel. Paratypes: 1 female (CL 3.43 mm, CW 3.30 mm), 1 male (CL 3.50 mm, CW 4.14 mm), ZL-AR- CR- 98, same data as holotype.	en	Trivedi, Jigneshkumar N., Campos, Ernesto, Vachhrajani, Kauresh D. (2018): A new pea crab species of the genus Arcotheres Manning, 1993 (Crustacea: Decapoda: Brachyura: Pinnotheridae) from India. Zootaxa 4433 (1): 195-200, DOI: 10.11646/zootaxa.4433.1.13
96247A4783207C034C9EFE59FD52F877.taxon	description	Description. Female: carapace sub-trapezoidal (Fig. 1 A), slightly wider than long, regions moderately defined, anterolateral margin cristate, front slightly projecting, relatively convex, posterior margin slightly concave. Eyes small, pigmented, not visible in dorsal view; orbits oval. Antennules folded obliquely, antennular cavities slightly larger than ocular cavities (Fig. 1 B); antennae 6 - segmented, basal segment with large, quadrate urinal article (Fig. 1 B), fused with epistome, distal segment setose, scarcely reaching beyond inner edge of orbit. Epistome, endostome separated, margin of first thoracic sternite emarginated. MXP 3 obliquely placed in buccal cavity, propodus rhomboidal about twice as long as high, longer than carpus, dorsal margin with fringe of short setae, digitiform dactylus distally setose, inserted medially in notch on ventral margin of propodus, distally not overreaching tip of propodus but overreaching anteromesial angle of merus; ischiomerus completely fused (Fig. 1 F), 1.80 times as long as wide, lateral margin convex, with short, scattered setae; mesial margin angular at the widest point forming anteromesial angle, with long, slender setae. Exopod slightly longer than half length of ischiomerus, flagellum with three articles before aesthetascs, small distal segment inconspicuously articulated (Fig. 1 F). Chelipeds (P 1) stouter than ambulatory legs (P 2 – 5), merus slightly longer than carpus (Fig. 1 A); chela slightly globose (Figs. 1 D, E), ventral margin gently convex, dactylus shorter than dorsal margin of palm, longer than greatest height of palm, small scattered patches of granules present on outer surface, distal tip pointed, strongly curved (Fig. 1 D), cutting edge with one large proximal tooth, few small teeth distally, with long setae on inner surface. Pollex with two large, proximal teeth separated by denticulate notch, patch of short setae present just below cutting edge visible on outer surface, row of long setae running parallel to cutting edge visible on inner surface (Fig. 1 E), outer surface of palm slightly convex, inner surface highly convex. Ambulatory legs (P 2 – 5) asymmetrical in length (Figs. 3 A – D), those on left side longer than corresponding legs on right side (Fig. 1 A, Figs. 3 A – D; A’ – C’); P 2, P 3 (Figs. 3 A – B; A’ – B’) similar, with merus, carpus, propodus, dactylus glabrous; propodus longer than carpus, slightly curved on both margins; dactylus conical, shorter than carpus, propodus, hooked at tip. Fourth pereopods markedly asymmetrical (Figs. 3 C, C’), longer P 4 (Fig. 3 C) slender than P 2, P 3; carpus outer margin curved; dactylus sickle-like, ending in small curved tip, slightly smaller than propodus, longer than dactyli of P 2, P 3, P 5, ventral margin with long fine setae, few short setae also present on dorsal distal margin of curved tip. P 5 more slender than P 2 – P 4, all articles similar in shape to those of P 2, P 3; except longer sickle-like dactylus, dactylus much longer than those of P 2 and P 3 but shorter than that of P 4, few long setae present on distal half of ventral (inner) margin. Relative length of ambulatory legs P 4> P 3> P 2> P 5 and dactyli of longer pereopods P 4> P 5> P 3> P 2 (Figs. 3 A – D; A’ – C’). Pleon wide (Fig. 1 C), with six free somites and telson, margin with tomentum, pleonal somites increasing in length distally, widening from somite 1 to 4, then decreasing towards telson. Male: carapace subovate (Fig. 2 A), slightly longer than wide, dorsal surface smooth, front projecting anteriorly, emarginated, eyes visible in dorsal view. MXP 3 as in female. Chela stouter than in female, dorsal and ventral margins strongly convex (Fig. 2 B), inner margin of fingers with long setae, outer surface without setae, cutting edges of both fingers same as female chela. Ambulatory legs symmetrical, length of ambulatory legs P 4> P 3> P 2> P 5 (Figs. 3 E – H). P 2 – P 4 with a fringe of long swimming setae on outer face of carpus and propodus increasing in size distally. P 5 with propodus ellipsoidal, dactylus very curved, ventral margin of both with long setae. Relative length of dactyli of ambulatory legs P 5> P 4> P 3> P 2. Pleon with six free somites and telson (Fig. 2 C), widest at somites 2, 3, decreasing towards evenly rounded telson. G 1 long (Fig. 2 D), sharply curved, with short angular tip, ventral surface with long setae; G 2 (Fig. 2 E) stout with pointed angular tip, exopod absent.	en	Trivedi, Jigneshkumar N., Campos, Ernesto, Vachhrajani, Kauresh D. (2018): A new pea crab species of the genus Arcotheres Manning, 1993 (Crustacea: Decapoda: Brachyura: Pinnotheridae) from India. Zootaxa 4433 (1): 195-200, DOI: 10.11646/zootaxa.4433.1.13
96247A4783207C034C9EFE59FD52F877.taxon	biology_ecology	Host. The specimens of A. shahi n. sp. were collected from the oyster, Crassostrea madrasensis (Preston, 1916).	en	Trivedi, Jigneshkumar N., Campos, Ernesto, Vachhrajani, Kauresh D. (2018): A new pea crab species of the genus Arcotheres Manning, 1993 (Crustacea: Decapoda: Brachyura: Pinnotheridae) from India. Zootaxa 4433 (1): 195-200, DOI: 10.11646/zootaxa.4433.1.13
96247A4783207C034C9EFE59FD52F877.taxon	distribution	Distribution. So far known only from the coastal waters of Pamban, Gulf of Mannar, Tamil Nadu state, India.	en	Trivedi, Jigneshkumar N., Campos, Ernesto, Vachhrajani, Kauresh D. (2018): A new pea crab species of the genus Arcotheres Manning, 1993 (Crustacea: Decapoda: Brachyura: Pinnotheridae) from India. Zootaxa 4433 (1): 195-200, DOI: 10.11646/zootaxa.4433.1.13
96247A4783207C034C9EFE59FD52F877.taxon	etymology	Etymology. The new species is named in honor of Professor Raman Vadilal Shah, former Head of the Department of Zoology, The Maharaja Sayajirao University of Baroda, in Vadodara, Gujarat, India, for his valuable contributions in the field of zoology.	en	Trivedi, Jigneshkumar N., Campos, Ernesto, Vachhrajani, Kauresh D. (2018): A new pea crab species of the genus Arcotheres Manning, 1993 (Crustacea: Decapoda: Brachyura: Pinnotheridae) from India. Zootaxa 4433 (1): 195-200, DOI: 10.11646/zootaxa.4433.1.13
96247A4783207C034C9EFE59FD52F877.taxon	discussion	Remarks. On the basis of the relative lengths of the dactyli of the longer P 4 and P 5, it is possible to recognize two subgroups within the genus Arcotheres. The larger subgroup, contains 23 species: viz. A. alcocki (Rathbun, 1909), A. acrophilus (Bürger, 1895), A. atrinae (Sakai, 1939); A. borradailei (Nobili, 1905), A. casta Antony & Kuttyamma, 1971, A. coarcatus (Bürger, 1895), A. cyclinus (Shen, 1932); A. exiguus (Bürger, 1895), A. guinotae Campos, 2001, A. latifrons (Bürger, 1895), A. latus (Bürger, 1895), A. modiolicola (Bürger, 1895), A. nudifrons (Bürger, 1895), A. palaensis (Bürger, 1895), A. placunae (Hornell & Southwell, 1909), A. purpureus (Alcock, 1900), A. rhombifer (Bürger, 1895), A. ridgewayi (Southwell, 1911), A. rotundatus (Bürger, 1895), A. similis (Bürger, 1895), A. sinensis (Shen, 1932), A. spinidactylus (Gordon, 1936) and A. tivelae (Gordon, 1936) (Bürger 1895; Hornell & Southwell 1909; Gordon 1936; Ahyong & Ng 2007; Ng et al. 2017). The smaller subgroup contains only three species — A. pernicola (Bürger, 1895), A. winckworthi (Gordon, 1936) and A. rayi Ahyong & Ng, 2007 (Campos 2001). Arcotheres shahi n. sp. belongs to the smaller subgroup on the basis of the relative lengths of the dactyli of the P 4 and P 5.	en	Trivedi, Jigneshkumar N., Campos, Ernesto, Vachhrajani, Kauresh D. (2018): A new pea crab species of the genus Arcotheres Manning, 1993 (Crustacea: Decapoda: Brachyura: Pinnotheridae) from India. Zootaxa 4433 (1): 195-200, DOI: 10.11646/zootaxa.4433.1.13
