taxonID	type	description	language	source
1451431600C7573188AC72B2838E26D3.taxon	distribution	Distribution. The Pristimantis lacrimosus group is distributed in Central America, the Guianan Shield, Pacific Basin of Ecuador, and the Amazon Basin. Its species richness peaks in the Ecuadorian Andes (n = 19) and Amazon basin of Ecuador and Peru (n = 14).	en	Carrion-Olmedo, Julio C., Ron, Santiago R. (2021): A new cryptic species of the Pristimantis lacrimosus group (Anura, Strabomantidae) from the eastern slopes of the Ecuadorian Andes. Evolutionary Systematics 5: 151-175, DOI: http://dx.doi.org/10.3897/evolsyst.5.62661, URL: http://dx.doi.org/10.3897/evolsyst.5.62661
77498C97373659109C0C81229C58BD55.taxon	diagnosis	Diagnosis. The assignment of the new species to the genus Pristimantis is based on the phylogeny (Fig. 1). Pristimantis petersioides sp. nov. is characterized by the following combination of characters: (1) Skin on dorsum smooth to shagreen with or without scattered small tubercles, head with or without one interorbital small tubercle, skin of venter shagreened to weakly areolate; discoidal fold present, ill-defined; dorsolateral folds absent; (2) tympanic membrane and tympanic annulus present, round, its length 2 / 5 to 1 / 2 of eye diameter; its upper border weakly concealed by inconspicuous supratympanic fold; (3) snout rounded to truncate in dorsal view, truncate in lateral view, bearing a small rostral papilla; (4) interorbital space flat, broader than upper eyelid; upper eyelid with one distinct subconical tubercle surrounded by lower, indistinct rounded tubercles; cranial crests absent; (5) vomerine odontophores low to prominent, oblique, moderately separated, posteromedial to choanae; (6) males with prominent, subgular vocal sac and vocal slits; (7) first finger shorter than second; all fingers long, discs broadly expanded, rounded to truncate; all fingers bearing a hyperdistal tubercle (Fig. 4 B); (8) fingers with narrow lateral fringes; (9) few ulnar tubercles; (10) no knee and heel tubercles, outer tarsal fold bearing one to three indistinct tubercles; (11) two metatarsal tubercles, inner oval, 3 x the size of outer conical and elliptical metatarsal tubercle; supernumerary plantar tubercles numerous; (12) all toes with hyperdistal tubercles; toes with narrow lateral fringes; basal toe webbing absent, discs broadly expanded, Toe IV much longer than Toe III (disc on Toe III reaches proximal edge of penultimate subarticular tubercle on Toe IV, disc on Toe V exceeds the distal edge of penultimate subarticular tubercle on Toe IV), discs as expanded as those on fingers (Fig. 4 A); (13) SVL 22.8 + / - 1.4 mm (20.4 - 24.8 mm; n = 15) in females, 18.5 + / - 1.5 mm (15.8 - 23.9 mm; n = 39) in males.	en	Carrion-Olmedo, Julio C., Ron, Santiago R. (2021): A new cryptic species of the Pristimantis lacrimosus group (Anura, Strabomantidae) from the eastern slopes of the Ecuadorian Andes. Evolutionary Systematics 5: 151-175, DOI: http://dx.doi.org/10.3897/evolsyst.5.62661, URL: http://dx.doi.org/10.3897/evolsyst.5.62661
77498C97373659109C0C81229C58BD55.taxon	description	Description of the holotype. Adult female (QCAZ 58939). Measurements (in mm): SVL 22.02; tibia length 12.07; foot length 10.72; head length 8.82; head width 9.09; eye diameter 2.96; tympanum diameter 1.35; interorbital distance 2.52; upper eyelid width 2.44; internarial distance 1.59; eye-nostril distance 2.59; tympanum-eye distance 0.71. Body slender; head slightly wider than long, wider than body; snout rounded to truncate with rostral papilla in dorsal view, truncate in lateral profile; canthus rostralis distinct, slightly curved in dorsal view; loreal region concave; interorbital space flat, no cranial crests; eye large, protuberant; upper eyelid about 97 % of interorbital distance, bearing one subconical tubercle. Tympanic membrane and annulus distinct, rounded, with inconspicuous supratympanic fold, partially obscuring anterodorsal edge; horizontal diameter of tympanum about 13 % of head length, separated from eye by a distance about one half tympanum length; choanae large, rounded, not concealed by palatal shelf of maxillary arc; dentigerous processes of vomers prominent, oblique, bearing a transverse row of five teeth; tongue big, elliptical, posterior border slightly notched, 40 % of the anterior surface adherent to floor of mouth. Skin on dorsum smooth to shagreen; dorsolateral folds absent; skin on upper flanks bearing scattered low tubercles; skin on belly weakly areolate; skin on throat and chest smooth; discoidal fold ill-defined; skin in upper cloacal region shagreen. Forearms slender bearing low antebrachial tubercle and one subconical ulnar tubercle at the distal half of the forearm; fingers large and slender, all with broadly expanded pads, all fingers with discs; fingers bearing narrow lateral fringes; relative lengths of fingers I <II <IV <III; three subarticular tubercles on finger III (Fig. 4 B), the most distal we refer as hyperdistal, all the tubercles well defined, round in ventral and lateral view; several supernumerary tubercles present, prominent at the base of the fingers and lower, indistinct at the palmar surface; palmar tubercle bifid, heart-shaped, about the same length and twice the width of elliptical thenar tubercle (Fig. 4 B). Hindlimbs slender; tibia length about 55 % of SVL; upper surfaces of hindlimbs smooth; foot length about 48 % of SVL, posterior surfaces of thighs smooth, ventral surfaces of thighs slightly areolate; knee and heel lacking tubercles; outer surface of tarsus bearing three low, inconspicuous tubercles, equally distributed along tarsus; toes bearing narrow lateral fringes; webbing between toes absent; discs on toes broadly expanded as those on fingers, rounded; relative lengths of toes: I <II <III <V <IV; Toe V much longer than Toe III (disc on Toe III reaches proximal edge of penultimate subarticular tubercle on Toe IV, disc on Toe V exceeds the distal edge of penultimate subarticular tubercle on Toe IV), subarticular tubercles rounded, simple, elevated; plantar surface with low supernumerary tubercles, bearing four subarticular tubercles (Fig. 4 A), inner metatarsal tubercle prominent, elliptical, approximately 3 x size of oval and conical outer metatarsal tubercle (Fig. 4 A). Color of holotype in preservative. (Fig. 3 C, D) Background color pale grayish cream with scattered, irregular dark brown chevrons, head bearing dark brown supratympanic and canthal stripe, upper lip bearing ill-defined stripe formed by irregular dark brown dots; upper flanks bearing dark brown, irregular flecks and blotches densely distributed; venter, ventral surfaces of forearms and hindlimbs pale creamy white, chest and throat with diminutive dark brown dots uniformly distributed (visible under magnification); ventral surfaces of hands and foot with dense minute dark brown dots, posterior surfaces of thighs pale cream to dark brown; iris reddish coppery with fine, dense, black reticulation. Color of holotype in life. (Fig. 3 A, B) Dorsal surfaces yellowish green with scattered, irregular dark brown chevrons; canthal stripe and supratympanic fold black, upper flanks pale cream with dark brown irregular flecks and blotches; venter creamy white; axils pinkish white; ventral surfaces of limbs, thighs yellowish green; iris reddish copper with dark bronze faint horizontal streak and thin irregular black reticulations.	en	Carrion-Olmedo, Julio C., Ron, Santiago R. (2021): A new cryptic species of the Pristimantis lacrimosus group (Anura, Strabomantidae) from the eastern slopes of the Ecuadorian Andes. Evolutionary Systematics 5: 151-175, DOI: http://dx.doi.org/10.3897/evolsyst.5.62661, URL: http://dx.doi.org/10.3897/evolsyst.5.62661
77498C97373659109C0C81229C58BD55.taxon	distribution	Distribution and natural history. Pristimantis petersioides sp. nov. is known from six localities in the eastern Andean slopes of central Ecuador between 1221 - 2300 m (Fig. 9). It inhabits the Eastern Andean Foothills Forest and Eastern Montane Forest natural regions (as defined by Ron et al. 2019). It has been recorded in primary forest and, less frequently, in secondary forest. Individuals were found during nocturnal surveys, usually perching on ferns, herbs, or Heliconia leaves, branches, or inside bromeliads up to 350 cm above the ground, usually near water bodies. Three amplectant pairs were found on January and February 2015 in Sardinayacu and Zarentza.	en	Carrion-Olmedo, Julio C., Ron, Santiago R. (2021): A new cryptic species of the Pristimantis lacrimosus group (Anura, Strabomantidae) from the eastern slopes of the Ecuadorian Andes. Evolutionary Systematics 5: 151-175, DOI: http://dx.doi.org/10.3897/evolsyst.5.62661, URL: http://dx.doi.org/10.3897/evolsyst.5.62661
77498C97373659109C0C81229C58BD55.taxon	etymology	Etymology. The specific epithet is a masculine noun in apposition. The suffix oides is derived from the Greek eidos meaning similar. The name makes reference to the similarity between the new species and its sister species, Pristimantis petersi.	en	Carrion-Olmedo, Julio C., Ron, Santiago R. (2021): A new cryptic species of the Pristimantis lacrimosus group (Anura, Strabomantidae) from the eastern slopes of the Ecuadorian Andes. Evolutionary Systematics 5: 151-175, DOI: http://dx.doi.org/10.3897/evolsyst.5.62661, URL: http://dx.doi.org/10.3897/evolsyst.5.62661
