taxonID	type	description	language	source
8D0987B7D572FFAEFC187E0829FAFDA6.taxon	description	(Fig. 1 A-D) ZooBank: http: // zoobank. org / 8 EAA 0977 - F 5 A 1 - 4 F 1 B- 8207 - 68926110 E 12 F	en	Ahrens, Dirk (2022): Tetrasericatakakuwai sp. nov. - a further new species ofTetraserica Ahrens, 2004 from Thailand (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Faunitaxys 10 (33): 1-4, DOI: 10.57800/faunitaxys-10(33), URL: http://dx.doi.org/10.5281/zenodo.15933344
8D0987B7D572FFAEFC187E0829FAFDA6.taxon	materials_examined	Holotype, ♂, “ Phu Rua NP (900 m alt.), Loei O. NE Thai. 26 - 30 IV. 2006 Takakuwa, M. leg. / 1154 Sericini Asia spec. ” (ZFMK).	en	Ahrens, Dirk (2022): Tetrasericatakakuwai sp. nov. - a further new species ofTetraserica Ahrens, 2004 from Thailand (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Faunitaxys 10 (33): 1-4, DOI: 10.57800/faunitaxys-10(33), URL: http://dx.doi.org/10.5281/zenodo.15933344
8D0987B7D572FFAEFC187E0829FAFDA6.taxon	description	Description of the holotype. General appearance. – Length of body: 7.4 mm; – Length of elytra: 5.4 mm; – Maximum width: 4.4 mm. Dorsal surface reddish brown; surface of labroclypeus and disc of frons glabrous. Head. – Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex and convergent to strongly rounded anterior angles, anterior margin weakly sinuate medially, margins moderately reflexed. – Surface weakly convex, moderately shiny, finely and densely punctate. – Frontoclypeal suture indistinctly incised, flat and weakly curved medially. – Ocular canthus short and triangular, impunctate, with a single terminal seta. Smooth area anterior to eye twice as wide as long. – Frons dull, Reviewers: Denis Keith (France) - ZooBank: http: // zoobank. org / C 922 B 63 D- 331 E- 44 BF-B 386 - 9 A 70 AE 05 CA 28 with sparse, fine punctures, with single erect setae beside each eye. – Antenna yellowish, with ten antennomeres; club composed of four antennomeres in male, straight, rarely longer than 1.5 times as the remaining antennomeres combined. – Eyes moderatelysmall, ratio of diameter / interocular width: 0.71. – Mentum elevated and slightly flattened anteriorly. Thorax. – Pronotum moderately wideandstronglyconvex, lateralmargins evenly convex, more strongly narrowed anteriorly towards sharp and slightly produced anterior angles. – Anterior margin of pronotum slightly convex, with fine complete marginal line. – Posterior angles blunt or strongly rounded. – Surface finely and densely punctate, except minute setae glabrous, lateral and lateral anterior margins sparsely setose. – Hypomeron not carinate. – Scutellum triangular, finely and densely punctate. Elytra. – Oblong, widest just behind middle, striae distinctly impressed, finely and moderately densely punctate, intervals distinctly convex, with coarse and dense punctures concentrated along striae, with very minute setae in punctures. – Epipleural edge robust, ending at weakly curved and slightly blunt external apical angle of elytra, epipleura densely setose, apical border with a broad fringe of microtrichomes (100 x). Ventral. – Surface weakly shiny, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae, metacoxa glabrous, with a few single setae laterally. – Abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. – Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.67. – Pygidium weakly convex and dull, densely punctate, without smooth midline, almost glabrous, but with a few longer setae along apical margin; pygidium without strong sexual dimorphism. Legs. – Moderately wide. – Femora finely and sparsely punctate. – Metafemur wide and moderately shiny or dull, anterior margin acute, posterior margin smooth ventrally and only weakly widened in apical half, posterior margin smooth dorsally, with a few short setae basally. Posterior margin of metafemur generally straight or slightly convex. – Metatibia short and wide, ratiowidth / length: 1 / 2.8; basal group of dorsal spines of metatibia at first third of metatibial length. Robust spine at middle of ventral margin extremely prolonged and s-shaped, exceeding distal margin of metatibia. – Tarsomeres with fine, very dense setae ventrally on distal half, neither laterally nor dorsally carinate, dorsally smooth. – Metatarsomeres with a strongly serrated ridge ventrally and glabrous; first metatarsomere slightly shorter than two following tarsomeres combined, one third of its length longer than dorsal tibial spine. – Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Male genitalia. – Aedeagus: Fig. 1 A – C.	en	Ahrens, Dirk (2022): Tetrasericatakakuwai sp. nov. - a further new species ofTetraserica Ahrens, 2004 from Thailand (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Faunitaxys 10 (33): 1-4, DOI: 10.57800/faunitaxys-10(33), URL: http://dx.doi.org/10.5281/zenodo.15933344
8D0987B7D572FFAEFC187E0829FAFDA6.taxon	discussion	Remarks. – Female unknown.	en	Ahrens, Dirk (2022): Tetrasericatakakuwai sp. nov. - a further new species ofTetraserica Ahrens, 2004 from Thailand (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Faunitaxys 10 (33): 1-4, DOI: 10.57800/faunitaxys-10(33), URL: http://dx.doi.org/10.5281/zenodo.15933344
8D0987B7D572FFAEFC187E0829FAFDA6.taxon	diagnosis	Diagnosis. – Tetraserica takakuwai sp. nov. is in shape of aedeagus most similar with Tetraserica bansanpakiana Fabrizi, Dalstein & Ahrens, 2019 and T. margheritae Fabrizi, Dalstein & Ahrens, 2019 (Fig. 2). It shares with both species: - the robust metatibial spine in middle of ventral margin being extremely elongate and s-shaped, exceeding distal margin of metatibia, - short median lamina of phallobase, being subequal to at maximum half of length of phallobase, - the unicolored dorsal surface, - the left paramere being at base simple, and the strongly asymmetric phallobase (dorsal view). Tetrasericatakakuwai sp. nov. differs from T. bansanpakiana by: - the longer and narrower median lamina of phallobase which does not possess an apical dorsal tooth (Fig. 2 A, C), - the basal uncleft portion of the left paramere is much longer (Fig. 1 A), although if the apical branches appear to be broken in the holotype of T. takakuwai sp. nov. From T. margheritae the new species differs by: - the dorsoventrally wider median lamina of phallobase whose dorsal margin is weakly convex and not straight as in T. margheritae (Fig. 2 E); - the left paramere is basally not bent as in T. margheritae (Fig. 2 E; lateral view) but straight in basal third (Fig. 1 A), and over its length (lateral view) weakly curved instead of straight as in T. margheritae; - in dorsal view, the left paramere is in T. takakuwai sp. nov. straight and not curved externally (Fig. 1 B) as in T. margheritae (Fig. 2 F).	en	Ahrens, Dirk (2022): Tetrasericatakakuwai sp. nov. - a further new species ofTetraserica Ahrens, 2004 from Thailand (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Faunitaxys 10 (33): 1-4, DOI: 10.57800/faunitaxys-10(33), URL: http://dx.doi.org/10.5281/zenodo.15933344
8D0987B7D572FFAEFC187E0829FAFDA6.taxon	etymology	Etymology. – The new species is named after its collector, M. Takakuwa (nouningenitivesingular).	en	Ahrens, Dirk (2022): Tetrasericatakakuwai sp. nov. - a further new species ofTetraserica Ahrens, 2004 from Thailand (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Faunitaxys 10 (33): 1-4, DOI: 10.57800/faunitaxys-10(33), URL: http://dx.doi.org/10.5281/zenodo.15933344
