taxonID	type	description	language	source
8D4F87E02E35FFD124A34F4141EEF7B3.taxon	description	(Figs. 1, 3, 5, 6)	en	Migliavacca, Paulo Presti, Simone, Luiz Ricardo L. (2020): Morphological comparison between Doryteuthis pleii and D. sanpaulensis (Cephalopoda, Myopsida, Loliginidae) from Brazil. Papéis Avulsos de Zoologia 60: 1-14, DOI: 10.11606/1807-0205/2020.60.01
8D4F87E02E35FFD124A34F4141EEF7B3.taxon	distribution	Type locality: Martinique, West Indies, western North Atlantic Ocean.	en	Migliavacca, Paulo Presti, Simone, Luiz Ricardo L. (2020): Morphological comparison between Doryteuthis pleii and D. sanpaulensis (Cephalopoda, Myopsida, Loliginidae) from Brazil. Papéis Avulsos de Zoologia 60: 1-14, DOI: 10.11606/1807-0205/2020.60.01
8D4F87E02E35FFD124A34F4141EEF7B3.taxon	materials_examined	Holotype: Muséum National d’Histoire Naturelle, Paris [despite the label of the specimen thought to be the holotype is missing (Cohen, 1976)]. Material examined: Brazil; Rio de Janeiro: off São João da Barra, MZSP 15294, 24 spm. (“ Exp. Dep. Zool. ” coll., July 1963). off Farol de São Thomé, 22 ° 08 ′ S, 41 ° 00 ′ W, MZSP 27095, 22 m, 1 spm. (R / V “ W. Besnard ” coll., 12 March 1971). off Cabo Frio, 22 ° 51 ′ S, 41 ° 52 ′ W, MZSP 27092, 48 m, 17 spm. (R / V “ W. Besnard ” coll., 01 December 1970). 22 ° 59 ′ S, 42 ° 06 ′ W, MZSP 102761, 25 - 33 m, 2 spm. (1 June 1987). off Saquarema, 23 ° 02 ′ S, 42 ° 37 ′ W, MZSP 101893, 50 - 62 m, 1 spm. (19 November 2007). off Niterói, 23 ° 05 ′ S, 43 ° 06 ′ W, MZSP 110029, 30 - 50 m, 2 spm. (Mendonça Jr. coll., 05 December 2012). off Angra dos Reis, Vila Velha Beach, MZSP 27063, 1 spm. (“ Exp. Dep. Zool. ” coll., 21 July 1966). off Paraty, 23 ° 11 ′ S, 44 ° 38 ′ W, MZSP 58954, 2 spm. (Simone coll., 12 January 2006). MZSP 27061, 13 spm. (July 1983). MZSP 32965, 2 spm. (Fishermen coll., March 2001). São Paulo: off Ubatuba, MZSP 27076, 8 spm. (Boffi coll., 08 February 1970). MZSP 101892, 1 spm. (Caires coll., 2011). MZSP 27084, 2 spm. (Local fishermen coll., 12 March 1970). Lázaro Beach, MZSP 27087, 2 spm. (Sazima coll., 04 April 1970). off Maranduba, 23 ° 39 ′ S, 45 ° 05 ′ W, MZSP 27101, 28 - 32 m, 4 spm. (R / V “ W. Besnard ” coll., 05 December 1970). off Santos, MZSP 90936,2 spm. (Cunha coll., March 2009). MZSP 27097, 42 m, 4 spm. (R / V “ W. Besnard ” coll., 05 December 1970). MZSP 86427, 8 spm. (R / V “ Miragem Boat ” coll., March 1988). off Praia Grande, Boqueirão, 24 ° 01 ′ S, 46 ° 25 ′ W, MZSP 80342, 1 spm. (Simone coll., 1997). 24 ° 05 ′ S, 46 ° 20 ′ W, MZSP 110145, 2 spm. (10 April 2012). off São Sebastião, Alcatrazes archipelago. 24 ° 07 ′ S, 45 ° 40 ′ W, MZSP 102400, 46 m, 3 spm. (Amaral coll., 04 October 2011). 24 ° 07 ′ S, 45 ° 40 ′ W, MZSP 102401, 46 m, 2 spm. (Amaral coll., 04 October 2011). 24 ° 24 ′ S, 46 ° 05 ′ W, MZSP 27159, 52 m, 25 spm. (R / V “ W. Besnard ” coll., 21 January 1971). off Itanhaém, 24 ° 24 ′ S, 46 ° 18 ′ W, MZSP 27093, 45 m, 13 spm. (R / V “ W. Besnard ” coll., 05 March 1971). off Peruíbe, 24 ° 20 ′ S, 46 ° 49 ′ W, MZSP 27094, 79 m, 28 spm. (R / V “ W. Besnard ” coll., 06 March 1971). 24 ° 38 ′ S, 45 ° 05 ′ W, MZSP 27102, 102 m, 6 spm. (R / V “ W. Besnard ” coll., 06 March 1971). off Barra do Una, 24 ° 40 ′ S, 46 ° 49 ′ W, MZSP 27096, 38 m, 10 spm. (R / V “ W. Besnard ” coll., 06 December 1970). off Cananéia, Castilho Island – ESEC Tupiniquins, 25 ° 00 ′ S, 47 ° 55 ′ W, MZSP 83165, 5 spm. (Guimarães, Lima & Amaral coll., 24 January 2007). 25 ° 26 ′ S, 45 ° 35 ′ W, MZSP 27091, 125 - 128 m, 10 spm. (R / V “ W. Besnard ” coll., 05 July 1970). 25 ° 44 ′ S, 46 ° 37 ′ W, MZSP 27089,111 m, 6 spm. (R / V “ W. Besnard ” coll., 04 March 1971). Paraná: off Pontal do Sul, 25 ° 36 ′ S, 47 ° 30 ′ W, MZSP 27104, 50 m, 2 spm. (R / V “ W. Besnard ” coll., 07 December 1970). 25 ° 59 ′ S, 47 ° 15 ′ W, MZSP 27105, 77 - 82 m, 4 spm. (R / V “ W. Besnard ” coll., 07 December 1970). Santa Catarina: São Francisco do Sul Island, 26 ° 17 ′ S, 48 ° 22 ′ W, MZSP 27162, 10 spm. (R / V “ W. Besnard ” coll., February 1977). off Itajaí, MZSP 92949, 51 m, 2 spm. (R / V “ NPq Soloncy Moura ” coll., 09 November 2009). off Imbituba, 28 ° 18 ′ S, 48 ° 34 ′ W, MZSP 27090, 50 m, 12 spm. (R / V “ W. Besnard ” coll., 01 March 1971). Rio Grande do Sul: off Balneário Pinhal, 30 ° 37 ′ S, 49 ° 25 ′ W, MZSP 27058, 150 m, 4 spm. (GEDIP Project coll., 05 August 1972).	en	Migliavacca, Paulo Presti, Simone, Luiz Ricardo L. (2020): Morphological comparison between Doryteuthis pleii and D. sanpaulensis (Cephalopoda, Myopsida, Loliginidae) from Brazil. Papéis Avulsos de Zoologia 60: 1-14, DOI: 10.11606/1807-0205/2020.60.01
8D4F87E02E35FFD124A34F4141EEF7B3.taxon	distribution	Distribution: Cape Hatteras (36 ° N) to northern Argentina (35 ° S) usually associated with Brazilian warm current. Also including Gulf of Mexico, Caribbean Sea, Bermuda and Bahamian and Caribbean islands (Jereb & Roper, 2010).	en	Migliavacca, Paulo Presti, Simone, Luiz Ricardo L. (2020): Morphological comparison between Doryteuthis pleii and D. sanpaulensis (Cephalopoda, Myopsida, Loliginidae) from Brazil. Papéis Avulsos de Zoologia 60: 1-14, DOI: 10.11606/1807-0205/2020.60.01
8D4F87E02E35FFD124A34F4141EEF7B3.taxon	description	Description Mantle: Cylindrical; muscular (~ 91 mm; Table 1); usually long and slender (Fig. 1 A), ~ 4 - times longer than wide. Head: Eye diameter approximately 13 % of mantle length (Table 2). Width similar to length values. Chromatophores: Widely distributed, with higher densities on dorsal region including head (around eyes), dorsal midline of mantle (highly concentrated beginning of fins), fins and arms / tentacles. Mature males presenting striped pattern along ventral surface of mantle (Fig. 1 A). Fins: Rhomboid, slightly ovate comprising ~ 48 % ML (Table 2), connected posteriorly to posterior end of mantle (Fig. 1 A). Arms: Muscular, broad at base and gradually tapering into thin tips; formula III> IV> II> I (~ lengths: 33 mm, 30 mm, 29 mm, 23 mm); biserial suckers. Hectocotylized portion ~ 39 % of left ventral arm length reaching arm’s tip (Table 2), only left row of suckers modified into small peduncles. Bar between row of peduncles and row of suckers connecting them in ladder pattern (Fig. 1 E). Tentacle: Club long and thin, ~ 28 % ML (Table 2), carpus, manus and dactylus easily defined; 4 rows of suckers diagonally distributed along club, more visible at manus and dactylus regions; 2 central rows at manus region much bigger than other suckers. Each sucker with small chitin ring of teeth inside. Longitudinal muscle around sucker visible (Figs. 1 B, 1 D). Circulatory system: Normal Loliginidae form (Fig. 6 A). One central systemic croissant-shaped heart; two small- er peripheral branchial round hearts associated to gills. Two aortae: (i) anterior aorta delivering blood to head, arms and tentacles; (ii) posterior aortic trunk, divided in: (a) visceral aorta, sending blood to posterior part of mantle and visceral organs; (b) mantle aorta, distributing blood to anterior region of mantle. Blood return to heart (i) from posterior vena cava and (ii) from lateral nephridial vessel, both ending in a branchial heart. Branchial artery leaving branchial heart, and ending in gills complex; after this, returning to heart through branchial vein (Fig. 3 A). Digestive system (Figs. 5, 6): Pigmented tip of upper beak short and robust (Fig. 1 C). Radula usual for the family; rachidian and first lateral teeth bearing primary large projection and secondary smaller one (s) (Fig. 4 A). Usual digestive system for Loliginidae: buccal bulb occupying ~ half of inner head volume. Esophagus narrow, running straight posteriorly along ~ ⅕ of mantle length. Stomach spherical, wall weakly muscular, occupying ~ 20 % of visceral volume, located in middle level of visceral sac; posterior caecum wide, extending up to posterior mantle space. Intestine narrow, originating in YY region of stomach, performing simple loop in anterior region of visceral sac. Anus with two papillae, located in middle level of pallial cavity, preceded by short flexible stalk. Digestive gland wide and located between four retractile muscles of head and funnel. Esophagus passing through digestive gland ventro-dorsally. Male genital system: Normal genital system for Loliginidae, including characters of penis, Needham sac, spermatophoric gland and a single testicle (Figs. 6 A, 6 B). Hectocotylus described above. Female genital system: Normal genital system for Loliginidae, including characters of vagina, oviduct, oviducal gland, ovary, a pair of nidamental glands and pair of accessory nidamental glands (Fig. 5). Nidamental glands more uniform, with no slender tip (Fig. 5 B). Central nervous system (Fig. 3 B): Central mass slightly divided into anterior and posterior regions. Giant fibers running from posterior end, along dorsal portion of mantle; stellate ganglion visible beneath gills. One pair of optic lobes mushroom-shaped, flanking central mass on each side. Several nerves running anteriorly towards arms and tentacles, forming ring around buccal bulb (Fig. 3 B: ib).	en	Migliavacca, Paulo Presti, Simone, Luiz Ricardo L. (2020): Morphological comparison between Doryteuthis pleii and D. sanpaulensis (Cephalopoda, Myopsida, Loliginidae) from Brazil. Papéis Avulsos de Zoologia 60: 1-14, DOI: 10.11606/1807-0205/2020.60.01
8D4F87E02E31FFDD24FD4F614306F873.taxon	description	(Figs. 2, 3, 5, 6)	en	Migliavacca, Paulo Presti, Simone, Luiz Ricardo L. (2020): Morphological comparison between Doryteuthis pleii and D. sanpaulensis (Cephalopoda, Myopsida, Loliginidae) from Brazil. Papéis Avulsos de Zoologia 60: 1-14, DOI: 10.11606/1807-0205/2020.60.01
8D4F87E02E31FFDD24FD4F614306F873.taxon	distribution	Type locality: San Matias Gulf, Argentina, 41 ° 47 ′ S, 63 ° 35 ′ W.	en	Migliavacca, Paulo Presti, Simone, Luiz Ricardo L. (2020): Morphological comparison between Doryteuthis pleii and D. sanpaulensis (Cephalopoda, Myopsida, Loliginidae) from Brazil. Papéis Avulsos de Zoologia 60: 1-14, DOI: 10.11606/1807-0205/2020.60.01
8D4F87E02E31FFDD24FD4F614306F873.taxon	materials_examined	Holotype: USNM 815464. Material examined: Brazil; Rio de Janeiro: off Farol de São Thomé, MZSP 101894, 1 spm. (Nunan coll., 26 January 2007). off Maricás Island, 23 ° 01 ′ S, 42 ° 50 ′ W, MZSP 101891, 44 - 50 m, 1 spm. (17 - 18 November 2002). off Niterói, 23 ° 05 ′ S, 43 ° 06 ′ W, MZSP 110019, 30 - 50 m, 1 spm. (Mendonça Jr coll., 05 December 2012). 23 ° 05 ′ S, 43 ° 06 ′ W, MZSP 110025, 30 - 50 m, 20 spm. (Mendonça Jr coll., 05 December 2012). off Paraty, MZSP 32966, 2 spm. (Fishermen coll., March 2001). São Paulo: off Ubatuba, Itaguá Beach, MZSP 26409, 2 spm. (Figueiredo coll.). MZSP 27083, 19 spm. (Monfouchet coll., 12 March 1970). off São Sebastião, Alcatrazes Island, MZSP 27075, 2 spm. (16 March 1964). 24 ° 02 ′ S, 44 ° 42 ′ W, MZSP 27160, 80 - 90 m, 25 spm. (R / V “ W. Besnard ” coll., 11 August 1970). MZSP 47280, 2 spm. off Santos, MZSP 34506, 10 - 15 m, 6 spm. (Cunha coll., May 1998). MZSP 34645, 130 - 150 m, 1 spm. (Cunha coll., February 2002). MZSP 47289, 18 m, 13 spm. (R / V “ Emilia ” coll., November 1976). MZSP 91389, 3 spm. (Cunha coll., June 2009). MZSP 107571, 20 m, 30 spm. (Cunha coll., June 2009). off Boqueirão, 24 ° 01 ′ S, 46 ° 25 ′ W, MZSP 80343, 11 spm. (Simone coll., 1997). MZSP 27321, 6 m, 3 spm. (Simone coll., 10 January 1990). off Iguape, 25 ° 10 ′ S, 46 ° 45 ′ W, MZSP 18639, 7 spm. (R / V “ W. Besnard ” coll., 30 May 1970). Santa Catarina: off Itajaí, MZSP 92950, 51 m, 11 spm. (R / V “ NPq Soloncy Moura ” coll., 09 November 2009). off Florianópolis, 27 ° 09 ′ S, 47 ° 41 ′ W, MZSP 27135, 115 m, 2 spm. (R / V “ W. Besnard ” coll., 16 August 1970). 27 ° 56 ′ S, 48 ° 29 ′ W, MZSP 18680, 8 spm. (R / V “ W. Besnard ” coll., 17 December 1970). off Laguna, 28 ° 32 ′ S, 48 ° 11 ′ W, MZSP 18638, 4 spm. (R / V “ W. Besnard ” coll., 03 June 1970). off Balneário Gaivota, 29 ° 13 ′ S, 49 ° 35 ′ W, MZSP 27133, 20 m, 2 spm. (R / V “ W. Besnard ” coll., 31 January 1972). 29 ° 13 ′ S, 49 ° 35 ′ W, MZSP 27148, 19 m, 10 spm. (R / V “ W. Besnard ” coll., 06 April 1972). 29 ° 16 ′ S, 49 ° 14 ′ W, MZSP 27143, 51 m, 3 spm. (R / V “ W. Besnard ” coll., 30 January 1972). 29 ° 23 ′ S, 49 ° 16 ′ W, MZSP 27137, 50 m, 4 spm. (R / V “ W. Besnard ” coll., 06 April 1972). Rio Grande do Sul: off Arroio do Sal, 29 ° 52 ′ S, 48 ° 41 ′ W, MZSP 18603, 4 spm. (R / V “ W. Besnard ” coll., 05 March 1969). off Capão da Canoa, 29 ° 58 ′ S, 49 ° 18 ′ W, MZSP 27144, 92 m, 5 spm. (R / V “ W. Besnard ” coll., 29 January 1972). off Tramandaí, MZSP 26410, 3 spm. (Phonlor coll., 1974). off Cidreira, MZSP 26411, 10 m, 1 spm. (Phonlor coll., July 1974). MZSP 26408, 10 m, 1 spm. (Phonlor coll., 08 October 1974). off Balneário Pinhal, 30 ° 16 ′ S, 50 ° 09 ′ W, MZSP 27138, 22 m, 2 spm. (R / V “ W. Besnard ” coll., 28 January 1972). 30 ° 22 ′ S, 49 ° 36 ′ W, MZSP 18589, 18 spm. (R / V “ W. Besnard ” coll., 05 December 1968). 30 ° 27 ′ S, 49 ° 48 ′ W, MZSP 27130, 71 m, 6 spm. (R / V “ W. Besnard ” coll., 28 January 1972). off Solidão, 30 ° 47 ′ S, 50 ° 28 ′ W, MZSP 27145, 17 m, 8 spm. (R / V “ W. Besnard ” coll., 26 January 1972). 30 ° 53 ′ S, 50 ° 16 ′ W, MZSP 27132, 60 m, 2 spm. (R / V “ W. Besnard ” coll., 26 January 1972). off Mostardas, 31 ° 09 ′ S, 50 ° 43 ′ W, MZSP 27131, 20 m, 12 spm. (R / V “ W. Besnard ” coll., 07 August 1972). 31 ° 12 ′ S, 50 ° 35 ′ W, MZSP 27141, 59 m, 5 spm. (R / V “ W. Besnard ” coll., 29 January 1972). off Tavares, 31 ° 30 ′ S, 51 ° 00 ′ W, MZSP 27139, 22 m, 13 spm. (R / V “ W. Besnard ” coll., 25 January 1972). off Rio Grande, 32 ° 13 ′ S, 50 ° 35 ′ W, MZSP 27136, 84 m, 2 spm. (R / V “ W. Besnard ” coll., 22 January 1972). 32 ° 20 ′ S, 51 ° 22 ′ W, MZSP 27128, 52 m, 8 spm. (R / V “ W. Besnard ” coll., 21 January 1972). 32 ° 28 ′ S, 51 ° 07 ′ W, MZSP 27155, 61 m, 5 spm. (R / V “ W. Besnard ” coll., 03 November 1972). 32 ° 28 ′ S, 52 ° 15 ′ W, MZSP 47314, 15 m, 10 spm. (R / V “ W. Besnard ” coll., 20 January 1972). MZSP 47292, 2 spm. (R / V “ W. Besnard ” coll., 17 August 1968). MZSP 18579, 3 spm. (R / V “ W. Besnard ” coll., 22 August 1968). off Curral Alto, 33 ° 13 ′ S, 51 ° 58 ′ W, MZSP 27150, 51 m, 2 spm. (R / V “ W. Besnard ” coll., 02 November 1972). 33 ° 16 ′ S, 52 ° 05 ′ W, MZSP 18601, 4 spm. (R / V “ W. Besnard ” coll., 13 December 1968). 33 ° 32 ′ S, 52 ° 18 ′ W, MZSP 18619, 2 spm. (R / V “ W. Besnard ” coll., 13 March 1969). MZSP 27158, 20 spm. (No additional information).	en	Migliavacca, Paulo Presti, Simone, Luiz Ricardo L. (2020): Morphological comparison between Doryteuthis pleii and D. sanpaulensis (Cephalopoda, Myopsida, Loliginidae) from Brazil. Papéis Avulsos de Zoologia 60: 1-14, DOI: 10.11606/1807-0205/2020.60.01
8D4F87E02E31FFDD24FD4F614306F873.taxon	distribution	Distribution: Southwestern Atlantic Ocean; usually from southern Brazil (20 ° S) to northern Patagonia, Argentina (46 ° S) (Jereb & Roper, 2010).	en	Migliavacca, Paulo Presti, Simone, Luiz Ricardo L. (2020): Morphological comparison between Doryteuthis pleii and D. sanpaulensis (Cephalopoda, Myopsida, Loliginidae) from Brazil. Papéis Avulsos de Zoologia 60: 1-14, DOI: 10.11606/1807-0205/2020.60.01
8D4F87E02E31FFDD24FD4F614306F873.taxon	description	Description Mantle: Cylindrical and muscular ranging to about 89 mm (Table 3; Fig. 2 A); ~ 3 times longer than wide. Head: Eye diameter approximately 12 % ML (Table 4). Length and width very similar to each other. Chromatophores: Distributed throughout body, especially on dorsal region including head (around eyes); dorsal midline of mantle forming an expressive solid line in region free from fins; few chromatophores clustering in middle of fins and at arms / tentacles (Fig. 2 A). Mature males do not present stripped chromatophores along ventral portion of mantle. Fins: Rhomboid, posteriorly connected to mantle, slightly ovate and moderately long, comprising ~ 59 % ML (Table 4; Fig. 2 A). Arms: Muscular, thick at base and distally tapering into slender tips; formula III> IV> II> I (~ lengths: 41 mm, 38 mm, 36 mm, 28 mm); two rows of suckers. Hectocotylus consists of ~ 40 % of left ventral arm length extending to arm tip (Table 4). Left row of suckers modified into a simple peduncle whereas right row remaining as suckers. No connection between modified row and suckers, forming valley pattern throughout hectocotylus (Fig. 2 E). Tentacle: Club long and broad, ~ 37 % ML (Table 4). Carpus, manus and dactylus uniformly distributed, with boundaries not very sharp, hindering their definition (especially between manus and dactylus); 4 diagonal rows of suckers along club as stated for D. pleii; 2 central rows at manus not so different in size from other suckers (Fig. 2 B). Each sucker with small ring of teeth apparently at surface of sucker’s aperture. Longitudinal muscle around suckers present (Fig. 2 D). Circulatory and central nervous system: Similar to preceding species. No apparent morphological differences (Fig. 3 A). Digestive system: Pigmented tip of upper beak is long and slender (Fig. 2 C). Remaining structures similar to preceding species. Male genital system: Similar to preceding species. No apparent morphological differences (Figs. 6 A, 6 B) except for hectocotylus above described. Female genital system: Similar to preceding species except for nidamental glands with prominent thin tip (Fig. 5 D).	en	Migliavacca, Paulo Presti, Simone, Luiz Ricardo L. (2020): Morphological comparison between Doryteuthis pleii and D. sanpaulensis (Cephalopoda, Myopsida, Loliginidae) from Brazil. Papéis Avulsos de Zoologia 60: 1-14, DOI: 10.11606/1807-0205/2020.60.01
