taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
8F2F2F2DA727FF9EFF1F4A163540FC32.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16982622/files/figure.png	https://doi.org/10.5281/zenodo.16982622	FIGURE 2. Pygmarrhopalites arbaicus sp. nov.: A, chaetotaxy of head; B, chaetotaxy of labial palp; C, labrum and maxillary outer lobe; D, chaetotaxy of antenna and details of the apical subsegment.	FIGURE 2. Pygmarrhopalites arbaicus sp. nov.: A, chaetotaxy of head; B, chaetotaxy of labial palp; C, labrum and maxillary outer lobe; D, chaetotaxy of antenna and details of the apical subsegment.	2025-08-07	Vargovitsh, Robert S.		Zenodo	biologists	Vargovitsh, Robert S.			
8F2F2F2DA727FF9EFF1F4A163540FC32.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16982624/files/figure.png	https://doi.org/10.5281/zenodo.16982624	FIGURE 3. Pygmarrhopalites arbaicus sp. nov.: A, chaetotaxy of foreleg; B, chaetotaxy of midleg; C, chaetotaxy of hindleg; D–L, shape of setae: D, D2 of head dorsum; E, m1 of mesothorax; F, a4 of fore tibiotarsus; G, dI-1 of great abdomen; H, Ie of dens; I, IIpe of dens; J, IIIpi of dens; K, mpi2 of Abd VI; L, mi5 of Abd VI (appendices anales).	FIGURE 3. Pygmarrhopalites arbaicus sp. nov.: A, chaetotaxy of foreleg; B, chaetotaxy of midleg; C, chaetotaxy of hindleg; D–L, shape of setae: D, D2 of head dorsum; E, m1 of mesothorax; F, a4 of fore tibiotarsus; G, dI-1 of great abdomen; H, Ie of dens; I, IIpe of dens; J, IIIpi of dens; K, mpi2 of Abd VI; L, mi5 of Abd VI (appendices anales).	2025-08-07	Vargovitsh, Robert S.		Zenodo	biologists	Vargovitsh, Robert S.			
8F2F2F2DA727FF9EFF1F4A163540FC32.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16982626/files/figure.png	https://doi.org/10.5281/zenodo.16982626	FIGURE 4. Pygmarrhopalites arbaicus sp. nov.: A, chaetotaxy of great abdomen; B, chaetotaxy of furca; C, tenaculum; D, chaetotaxy of female Abd VI, lateral view; E, сhaetotaxy of male Abd VI, posterior view.	FIGURE 4. Pygmarrhopalites arbaicus sp. nov.: A, chaetotaxy of great abdomen; B, chaetotaxy of furca; C, tenaculum; D, chaetotaxy of female Abd VI, lateral view; E, сhaetotaxy of male Abd VI, posterior view.	2025-08-07	Vargovitsh, Robert S.		Zenodo	biologists	Vargovitsh, Robert S.			
8F2F2F2DA727FF9EFF1F4A163540FC32.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16982628/files/figure.png	https://doi.org/10.5281/zenodo.16982628	FIGURE 5. Pygmarrhopalites arbaicus sp. nov.: A, habitus of mounted and alcohol specimen; B, appendices anales; C, Ant IV of male; D, chaetotaxy of head dorsum; E, eye; F, subapical organite of Ant IV; G, apex of Ant IV with apical sensillum; H, Ant III organ; I, labrum and maxillary outer lobe; J, head of maxilla; K–M, foot complex: K, foreleg; L, midleg; M, hindleg.	FIGURE 5. Pygmarrhopalites arbaicus sp. nov.: A, habitus of mounted and alcohol specimen; B, appendices anales; C, Ant IV of male; D, chaetotaxy of head dorsum; E, eye; F, subapical organite of Ant IV; G, apex of Ant IV with apical sensillum; H, Ant III organ; I, labrum and maxillary outer lobe; J, head of maxilla; K–M, foot complex: K, foreleg; L, midleg; M, hindleg.	2025-08-07	Vargovitsh, Robert S.		Zenodo	biologists	Vargovitsh, Robert S.			
8F2F2F2DA727FF9EFF1F4A163540FC32.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16982630/files/figure.png	https://doi.org/10.5281/zenodo.16982630	FIGURE 6. Pygmarrhopalites arbaicus sp. nov.: A, chaetotaxy of trichobothrial complex; B, circumanal setae of female Abd VI; C–D, chaetotaxy of furca: C, anterior side; D, posterior side; E–F, mucro: E, posterior side; F, anterior side; G–I, spine-like setae of dens: G, Ie; H, IIpe; I, IIpi; J, trochanteral organ on midleg; K, sensilla on coxa of midleg.	FIGURE 6. Pygmarrhopalites arbaicus sp. nov.: A, chaetotaxy of trichobothrial complex; B, circumanal setae of female Abd VI; C–D, chaetotaxy of furca: C, anterior side; D, posterior side; E–F, mucro: E, posterior side; F, anterior side; G–I, spine-like setae of dens: G, Ie; H, IIpe; I, IIpi; J, trochanteral organ on midleg; K, sensilla on coxa of midleg.	2025-08-07	Vargovitsh, Robert S.		Zenodo	biologists	Vargovitsh, Robert S.			
8F2F2F2DA72EFF86FF1F4A3835AEFB36.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16982634/files/figure.png	https://doi.org/10.5281/zenodo.16982634	FIGURE 7. Pygmarrhopalites trisetosus sp. nov.: A, chaetotaxy of head; B, chaetotaxy of labial palp; C, labrum and maxillary outer lobe; D, chaetotaxy of antenna and details of the apical subsegment.	FIGURE 7. Pygmarrhopalites trisetosus sp. nov.: A, chaetotaxy of head; B, chaetotaxy of labial palp; C, labrum and maxillary outer lobe; D, chaetotaxy of antenna and details of the apical subsegment.	2025-08-07	Vargovitsh, Robert S.		Zenodo	biologists	Vargovitsh, Robert S.			
8F2F2F2DA72EFF86FF1F4A3835AEFB36.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16982636/files/figure.png	https://doi.org/10.5281/zenodo.16982636	FIGURE 8. Pygmarrhopalites trisetosus sp. nov.: A, chaetotaxy of foreleg; B, chaetotaxy of midleg; C, chaetotaxy of hindleg; D–O, shape of setae: D, D2 of head dorsum; E, m1 of mesothorax; F, a4 of fore tibiotarsus; G, dI-1 of great abdomen; H, ms1 of Abd VI; I, mps3 of Abd VI; J, mpi1 of Abd VI; K, mpi2 of Abd VI; L, mi5 of Abd VI (appendices anales); M, Ie of dens; N, IIpe of dens; O, Ipi of dens.	FIGURE 8. Pygmarrhopalites trisetosus sp. nov.: A, chaetotaxy of foreleg; B, chaetotaxy of midleg; C, chaetotaxy of hindleg; D–O, shape of setae: D, D2 of head dorsum; E, m1 of mesothorax; F, a4 of fore tibiotarsus; G, dI-1 of great abdomen; H, ms1 of Abd VI; I, mps3 of Abd VI; J, mpi1 of Abd VI; K, mpi2 of Abd VI; L, mi5 of Abd VI (appendices anales); M, Ie of dens; N, IIpe of dens; O, Ipi of dens.	2025-08-07	Vargovitsh, Robert S.		Zenodo	biologists	Vargovitsh, Robert S.			
8F2F2F2DA72EFF86FF1F4A3835AEFB36.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16982638/files/figure.png	https://doi.org/10.5281/zenodo.16982638	FIGURE 9. Pygmarrhopalites trisetosus sp. nov.: A, chaetotaxy of great abdomen; B, chaetotaxy of furca; C, tenaculum; D, chaetotaxy of male Abd VI; E–F, chaetotaxy of female Abd VI: E, lateral view; F, posterior view, schematic.	FIGURE 9. Pygmarrhopalites trisetosus sp. nov.: A, chaetotaxy of great abdomen; B, chaetotaxy of furca; C, tenaculum; D, chaetotaxy of male Abd VI; E–F, chaetotaxy of female Abd VI: E, lateral view; F, posterior view, schematic.	2025-08-07	Vargovitsh, Robert S.		Zenodo	biologists	Vargovitsh, Robert S.			
8F2F2F2DA72EFF86FF1F4A3835AEFB36.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16982640/files/figure.png	https://doi.org/10.5281/zenodo.16982640	FIGURE 10. Pygmarrhopalites trisetosus sp. nov.: A, habitus of mounted and alcohol specimen; B, chaetotaxy of head dorsum; C–D, Ant IV: C, female; D, male; E–F, Ant III organ; G, head of maxilla; H, trochanteral organ of midleg; I–K, foot complex: I, foreleg; J, midleg; K, hindleg.	FIGURE 10. Pygmarrhopalites trisetosus sp. nov.: A, habitus of mounted and alcohol specimen; B, chaetotaxy of head dorsum; C–D, Ant IV: C, female; D, male; E–F, Ant III organ; G, head of maxilla; H, trochanteral organ of midleg; I–K, foot complex: I, foreleg; J, midleg; K, hindleg.	2025-08-07	Vargovitsh, Robert S.		Zenodo	biologists	Vargovitsh, Robert S.			
8F2F2F2DA72EFF86FF1F4A3835AEFB36.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16982644/files/figure.png	https://doi.org/10.5281/zenodo.16982644	FIGURE 11. Pygmarrhopalites trisetosus sp. nov. A, chaetotaxy of trichobothrial complex; B,Abd VI of female with circumanal setae; C–F, mucro; G–H, spine-like setae of dens: G, Ie; H, Ipi; I, tenaculum; J, appendices anales.	FIGURE 11. Pygmarrhopalites trisetosus sp. nov. A, chaetotaxy of trichobothrial complex; B,Abd VI of female with circumanal setae; C–F, mucro; G–H, spine-like setae of dens: G, Ie; H, Ipi; I, tenaculum; J, appendices anales.	2025-08-07	Vargovitsh, Robert S.		Zenodo	biologists	Vargovitsh, Robert S.			
8F2F2F2DA72EFF86FF1F4A3835AEFB36.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16982617/files/figure.png	https://doi.org/10.5281/zenodo.16982617	FIGURE 1. Type locality of Pygmarrhopalites arbaicus sp. nov. and P. trisetosus sp. nov.: A, location of the Arabika Massif in the Caucasus; B, location of ‘Central Plateau’ on the Arabika Massif; C, ‘Central Plateau’ of the Arabika Massif with marked entrances to caves: Nadezhda (1), Sirota (2), Moskva (3), Tochka Zh (4); D–E, landscape of ‘Central Plateau’ of the Arabika Massif: D, view from Arabika Mt.; E, view from locality of Nadezhda Cave; F–G, Nadezhda Cave; H, Moskva Cave; I, Sirota Cave; J–K, Tochka Zh Cave.	FIGURE 1. Type locality of Pygmarrhopalites arbaicus sp. nov. and P. trisetosus sp. nov.: A, location of the Arabika Massif in the Caucasus; B, location of ‘Central Plateau’ on the Arabika Massif; C, ‘Central Plateau’ of the Arabika Massif with marked entrances to caves: Nadezhda (1), Sirota (2), Moskva (3), Tochka Zh (4); D–E, landscape of ‘Central Plateau’ of the Arabika Massif: D, view from Arabika Mt.; E, view from locality of Nadezhda Cave; F–G, Nadezhda Cave; H, Moskva Cave; I, Sirota Cave; J–K, Tochka Zh Cave.	2025-08-07	Vargovitsh, Robert S.		Zenodo	biologists	Vargovitsh, Robert S.			
