taxonID	type	description	language	source
8870B305E3334C159699FF02896F8A05.taxon	materials_examined	Material examined. Holotype: 1 ovigerous female (8.3 mm) (MNHN-IU- 2013 - 1352), stn PD 31, Alexishafen, 05 ° 05.3 ' S 145 ° 48.1 ' E, 1 – 6 m, 17.12.2012. Paratypes: 1 immature female (6.3 mm) (MNHN-IU- 2013 - 1193), stn PD 66, south Yabob I., 05 ° 15.5 ' S 145 ° 47.3 ' E, 2 – 6 m, 3.12.2012; 1 immature female (6.6 mm) (ZRC 2015.269), stn PD 73, Madang, 6.12.2013.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3334C159699FF02896F8A05.taxon	description	Description. Carapace (Fig. 6 A – D) laterally expanded, 1.5 times as wide as long, irregularly rimmed, dorsal surface prominently sculpted, pitted, irregularly covered with coalesced flattened granules, resembling eroded coral rubble. Front narrow, produced, upcurved, bilobed, medially deeply pitted. Anterior margin of carapace vertical. Antennule well developed, basal antennular article operculiform, rugose, entirely sealing subovate antennular fossa. Antennae folded in orbital hiatus without gap. Orbits small, rounded, visible in dorsal view; ocular peduncle nearly seals orbit when retracted. Third maxillipeds concealing buccal cavity, closely set with flattened granules (Fig. 8 A); merus with pointed apex, outer margin convex, surface with median shallow depression; ischium twice as long as merus along inner margin. Anterior, lateral margins of carapace upturned to form granular rim; posteriorly, margin forms granular ridge merging with branchial protuberances. Anterolateral margin with 2 closed fissures, subhepatic margin with angular, granular, facet medially, visible in dorsal view. Posterolateral margin with prominently granular angular facet submedially. Posterior margin produced, narrow, bilobate. Postfrontal median longitudinal ridge narrow, distinct, merging into cardiac region; on either side of ridge slightly anterior to branchial regions a granular protuberance, pitted anteriorly. Bottom of depression between anterolateral rim, swollen branchial regions lined with flattened granules. Branchial regions swollen, with irregular cluster of granules. Granule-lined grooves separate cardiac, intestinal regions from branchial regions. Intestinal region swollen. Branchiostegal region covered with obsolete granules, medially demarcated by faint longitudinal suture. Chelipeds (Fig. 6 E) subequal, robust, covered with flattened granules. Cheliped palm inflated, minutely granular; fingers 1.5 as long as palm; dactylus with longitudinal rows of closely spaced granules on outer surface; pollex, wider than dactylus, with carinate ventral margin; cutting edges with minute teeth. Ambulatory legs stout, short, closely granular, diminishing in size posteriorly; meri with single granular row dorsally, 2 rows ventrally; propodi with row of conic granules ventrally; dactyli slender, longer than propodi, studded with conic granules, terminating in incurved cornute claw. Thoracic sternum closely granular. Female abdominal cavity deep, reaching buccal cavity. Abdomen closely covered with flattened granules; abdominal somites 1, 2 narrow, horizontal; somites 3 – 6 fused shield-like, divided by 2 submedian furrows; telson laciniate. Vulvar sternal cover oval, large kidney-shaped opening directed interiorly. Colour in life. Carapace bone-coloured, carapacial pits and grooves brown, female abdomen with 2 pairs of brown dots; dorsal margin of cheliped merus medially with 3 dark red granules, fingers spotted dark red; dorsal margins of ambulatory legs with dark red blotches (Fig. 2 A, B).	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3334C159699FF02896F8A05.taxon	discussion	Remarks. Alox naispela sp. nov. closely resembles A. patella (Alcock, 1896) (type locality: Andaman Is.) in the general appearance of the carapace, with its distinct rim formed by the upturned margins. It is distinguished from the latter, however, by its proportionately longer fingers (1.5 as compared to 1.2 as long as palm in A. patella, Tan & Ng 1995: fig. 11 A), dorsal surface of carapace with coalesced flattened granules rather than large mushroom-like granules, groove parallel to anterior margin of carapace nearly obsolete rather than distinct; and branchiostegal region covered with obsolete granules rather than large mushroom-like granules (Tan & Ng 1995: pl. 6 A – C).	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3334C159699FF02896F8A05.taxon	etymology	Etymology. From naispela for “ beautiful ” in Tok Pisin, the Pidgin language spoken in Papua New Guinea. Used as a noun in apposition. Geographical distribution. This species is known only from the type location in Papua New Guinea.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3354C139699FF478DBF8A9A.taxon	materials_examined	Material examined. 1 male (7.3 mm) (MNHN-IU- 2013 - 8854), 1 female (8.0 mm) (MNHN-IU- 2013 - 17888), stn PR 69, Rempi Area, south of lagoon inside bay, 05 ° 01.6 ' S 145 ° 48.1 ' E, 2 – 15 m, 20.11.2012; 1 male (5.8 mm) (ZRC 2015.267), stn PB 12, Kranket I., west side, 05 ° 11.8 ' S 145 ° 48.8 ' E, 7 – 15 m, 11.11.2012; 1 immature female (5.9 mm) (ZRC 2015.268), stn PD 19, Alexishafen, south Megas Islet, 05 ° 05.4 ' S 145 ° 48.5 ' E, 3 – 10 m, 13.11.2012; 1 female (10.0 mm) (MNHN-IU- 2013 - 17896) stn PD 57, Gosern I., 05 ° 09,4 ' S 145 ° 48,3 ' E, 1 – 4 m, 29.11.2012.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3354C139699FF478DBF8A9A.taxon	description	Description. Carapace (Fig. 7 A – G) laterally expanded, 1.5 times as wide as long. Carapace irregularly rimmed, dorsal surface prominently sculpted, pitted, irregularly covered with coalesced flattened granules, resembling coral rubble. Front narrow, produced, upcurved, bilobed, medially deeply pitted. Anterior margin of carapace vertical. Antennule well developed, basal antennular article operculiform, rugose, entirely sealing subovate antennular fossa. Antennae folded in orbital hiatus without gap. Orbits small, rounded, visible in dorsal view; ocular peduncle nearly seals orbit when retracted. Third maxillipeds concealing buccal cavity, closely set with flattened granules (Fig. 8 B); merus with pointed apex, outer margin convex, surface with median shallow depression; ischium twice as long as merus along inner margin. Subhepatic region with narrow groove parallel to anterior margin of carapace, fungiform granules inserted in groove, margin with obtuse, granular, facet medially, visible in dorsal view. Lateral margins of carapace subquadrate, swollen, forming granular rim. Posterolateral margin with granular angular facet submedially. Posterior margin slightly produced, narrow, bilobate. Postfrontal median longitudinal ridge of irregular width. Bottom of irregular depression between marginal rim, swollen branchial regions lined with fungiform granules. Branchial regions swollen, irregularly pitted. Granule-lined irregular grooves separate cardiac, intestinal regions from branchial regions. Intestinal region swollen. Branchiostegal region covered with obsolete granules externally, large fungiform granules interspaced with small rounded granules interiorly. Chelipeds (Fig. 7 E) subequal, robust, covered with flattened granules. Cheliped merus with large tubercle distally on posterior margin; palm inflated, granular, distally pitted; fingers 1.5 as long as palm; dactylus with longitudinal rows of closely spaced granules on outer surface; pollex, wider than dactylus, with carinate ventral margin; cutting edges with minute teeth. Ambulatory legs stout, short, closely granular, diminishing in size posteriorly; meri with single granular row dorsally, 2 rows ventrally; propodi with row of conic granules ventrally; dactyli slender, longer than propodi, studded with conic granules, terminating in incurved cornute claw (Fig. 8 D). Thoracic sternites with large, granule-lined fossae. Male, female abdominal cavity deep, reaching buccal cavity. Male abdomen with granular interrupted grooves on either side of median ridge, telson triangular, elongated (Fig. 8 E). Gl straight along entire length, dorsoventrally flattened, distally slightly dilated, setose, apex pointed, with slit-like opening (Fig. 8 F, G). Female abdominal somites 1, 2 narrow, horizontal, distinctly granular; somites 3 – 6 fused, shield-like, divided into 3 parts by 2 shallow submedian furrows; telson laciniate. Vulvae large, nearly rectangular; directed vertically. Colour in life. Carapace and legs bone-coloured with pale orange blotches, female abdomen with 2 pairs of brown dots (Fig. 2 C, D).	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3354C139699FF478DBF8A9A.taxon	discussion	Remarks. We refer the present specimens to A. rugosum (Stimpson, 1858) (type locality: “ Loo Choo ”, see discussion Tan & Ng 1995: 131) with doubt. In the general appearance of the carapace, with dorsal surface prominently sculpted, pitted, and irregularly covered with coalesced flattened granules, it is closest to this species. The present specimens nevertheless differ slightly from most of the specimens previously referred to A. rugosum (cf. Tan & Richer de Forges 1993: fig. 4 C, D; Tan & Ng 1995: fig. 12 D, pl. 6 D) in possessing a more distinct depression medially on the dorsal frontal surface; the male abdomen has more distinct granular interrupted grooves on either side of the median ridge, the rounded obtuse angle on hepatic margin with a shallow oval depression is not distinct, and the male telson is proportionately shorter. The specimens from Papua New Guinea most closely resemble in carapace form those previously reported from Gisser I. near Ceram in the Moluccas, Indonesia and figured as A. rugosum by Tan & Ng (1995: pl. 7 A, B). Considering the substantial variation in the carapace form in A. rugosum, as well as the similarity of the G 1 structures of the present specimens (Fig. 8 F, G, I, J) to those figured for A. rugosum s. str. (cf. Tan & Ng 1995: fig. 12 H, I), it would be premature to recognize the present material as a new species. Geographical distribution. This species is known only from Japan to New Caledonia (Tan & Ng 1995). New record for Papua New Guinea.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3344C129699FF47897A8B43.taxon	materials_examined	Material examined. Holotype: ovigerous female (8.3 mm) (MNHN-IU- 2013 - 8855), inner slope, stn PR 152, Tab I., 05 ° 10.1 ' S 145 ° 50.2 ' E, 3 – 25 m, 1.12.2012.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3344C129699FF47897A8B43.taxon	description	Description. Carapace laterally expanded (Fig. 6 F – I), 1.4 times as wide as long. Dorsal surface of carapace prominently sculpted, pitted, irregularly covered with coalesced flattened granules, resembling coral rubble. Front narrow, produced, upcurved, bilobed, horizontally grooved. Anterior margin of carapace vertical. Antennule well developed, basal antenular article operculiform, rugose, entirely sealing triangular antennular fossa. Antennae folded in orbital hiatus without gap. Orbits small, rounded, visible in dorsal view; when retracted, ocular peduncle nearly seals orbit. Third maxillipeds concealing buccal cavity, rugose; merus with pointed apex, outer margin convex; ischium 1.5 as long as merus along inner margin (Fig. 8 C). Obtuse angle medially on hepatic margin with shallow oval depression at apex. Subhepatic region with narrow groove parallel to anterior margin of carapace, fungiform granules inserted in groove; margin with prominent, granular, triangular facet medially, visible in dorsal view. Lateral margins of carapace extended, irregular. Posterolateral margins nearly horizontal in dorsal view. Posterior margin slightly produced, obsoletely bilobate. Postfrontal median longitudinal ridge widening posteriorly, merging into cardiac region. Postorbital prominence planar. Bottom of meandering depression between marginal rims, swollen branchial regions lined with fungiform granules. Branchial regions swollen, extending to posterolateral angle, irregularly pitted distally. Three pairs of irregularly shaped pits separate cardiac from branchial regions. Intestinal region raised, rimmed with fungiform granules. Branchiostegal region covered with small flattened granules separated externally by narrow cleft from larger granules interiorly. Chelipeds subequal, robust, rugose. Cheliped merus with large proximal tubercle on anterior margin, 2 distally on posterior margin; palm inflated, granular crests on dorsal, ventral margins; fingers 1.5 as long as palm; dactylus with longitudinal rows of closely spaced granules dorsally; pollex, wider than dactylus, with carinate ventral margin; cutting edges with minute teeth (Fig. 6 J). Ambulatory legs stout, short, closely granular, diminishing in size posteriorly; meri with single granular row dorsally, 2 rows ventrally; propodi with conic granules; dactyli slender, longer than propodi, studded with conic granules, terminating in incurved cornute claw. Thoracic sternites rugose. Male abdominal cavity deep, reaching buccal cavity. Female abdominal somites 1, 2 narrow, horizontal, distinctly granular; somites 3 – 6 fused, shield-like, with 2 shallow submedian furrows; telson elongate, laciniate. Vulvae large, oval-shaped, on sternal vulvar cover. Colour in life. Carapace and legs bone-coloured with pale orange blotches and spots (Fig. 2 E).	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3344C129699FF47897A8B43.taxon	discussion	Remarks. Alox liklik sp. nov. closely resembles A. rugosum (Stimpson, 1858) in the general appearance of the carapace, with its dorsal surface prominently sculpted, pitted, and irregularly covered with coalesced flattened granules. It is distinguished from the latter, however, in possessing posterolaterally extended swollen branchial regions (Tan & Ng 1995: pl. 6 D), relatively more elongated third maxilliped merus and laciniate telson, and vulvae which are large, oval-shaped, rather than ‘ comma’ shaped.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3344C129699FF47897A8B43.taxon	etymology	Etymology. From liklik for “ small ” in Tok Pisin, the Pidgin language spoken in Papua New Guinea. Used as a noun in apposition. Geographical distribution. This species is known only from the type location, Papua New Guinea.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3344C129699F91B8F5C8ABF.taxon	materials_examined	Material examined. 1 male (6.1 mm), 1 ovigerous female (6.4 mm) (MNHN-IU- 2013 - 8883); 2 ovigerous females (7.1 mm, 7.4 mm), 1 female (6.9 mm) (MNHN-IU- 2013 - 8884); 1 male (6.6 mm), 1 ovigerous female (7.4 mm) (ZRC 2014.0163), stn CP 4016, Dampier Strait, east Umboi Island, 05 ° 40 ' S 148 ° 14 ' E, 280 – 285 m, Solomon Sea, 12.12.2012. Geographical distribution. New Caledonia, Coral Sea, Fiji, Tonga, Sala-y-Gómez and Nazca ridges; 218 – 618 m (Galil 2004). New record for Papua New Guinea.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3374C119699FF028DBF8E30.taxon	materials_examined	Material examined. 1 male (9.1 mm) (MNHN-IU- 2013 - 458), stn PD 14, Madang Harbour, east Nui Is., 05 ° 12.3 ' S 145 ° 47.9 ' E, 10 – 15 m, 11.11.2012; 1 male (10.3 mm) (MNHN-IU- 2013 - 12766), stn PD 37, Rempi Area, north Tadwai I., 04 ° 58.5 ' S 145 ° 47.5 ' E, 5 – 10 m, 20.11.2012; 1 male (14.9 mm) (ZRC 2014.0170), stn PD 38, Rempi Area, north Tadwai I., 04 ° 58.8 ' S 145 ° 47.4 ' E, 10 – 15 m, 20.11.2012; 1 male (13.9 mm) (MNHN-IU- 2013 - 12769), stn PD 40, Rempi Area, east Tadwai I., 04 ° 59.3 ' S 145 ° 47.7 ' E, 21.11.2012; 1 immature male (9.4 mm) (MNHN-IU- 2013 - 12767), 1 immature specimen (6.2 mm) (MNHN-IU- 2013 - 17894), stn PD 54, west Panab I., 05 ° 10.3 ' S 145 ° 48.4 ' E, 15 m, 28.11.2012; 1 female (13.4 mm) (ZRC 2014.0171), 1 immature specimen (5.7 mm) (MNHN-IU- 2013 - 17893), stn PD 62, north Banap Damon Point, 05 ° 09.8 ' S 145 ° 48.4 ' E, 1 – 3 m, 1.12.2012; 1 immature specimen (4.1 mm) (MNHN-IU- 2013 - 12765), stn PD 66, S Yabob I., 05 ° 15.5 ' S 145 ° 47.3 ' E, 2 – 6 m, 3.12.2012; 1 immature specimen (6.4 mm) (MNHN-IU- 2013 - 17895); 4 immature specimens (6.2 – 9.5 mm) (MNHN-IU- 2013 - 12768); 2 immature specimens (6.9 mm, 6.8 mm) (MNHN-IU- 2013 - 12725), stn PD 73, Yomba I., 05 ° 14.7 ' S 145 ° 47.3 ' E, 20 – 25 m, 6.12.2012. Colour in life. Carapace and chelipeds pale orange; ambulatory legs, tips of spines, fingers, white (Fig. 3 A). Geographical distribution. Japan, China, Taiwan, Hong Kong, Singapore; 1 – 150 m (Galil 2001 a). New record for Papua New Guinea.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3374C119699FC2E8D2588FD.taxon	materials_examined	Material examined. 1 male (8.7 mm) (ZRC 2014.0172), stn PS 22, north Tadwai I., 04 ° 59.1 ' S 145 ° 47.6 ' E, 11 m, 22.11.2012; 1 female (13.1 mm) (ZRC 2014.0168); 1 female (9.5 mm) (MNHN-IU- 2013 - 1195), stn PD 66, south Yabob I., 05 ° 15.5 ' S 145 ° 47.3 ' E, 2 – 6 m, 3.12.2012; 1 male (8.1 mm) (MNHN-IU- 2013 - 17892), stn PS 42, S Urembo I., outer slope, 05 ° 15,9 ' S 145 ° 47,1 ' E, 18 – 27 m, 6.12.2012; 1 broken female (MNHN-IU- 2013 - 1494), stn CP 4035, east Kotakot, 04 ° 31 ' S 145 ° 31 ' E, 380 – 382 m, 17.12.2012. Colour in life. Carapace bone-coloured, posteriorly mottled red-brown; cheliped merus proximally stained red-brown, fingers mottled with pale brown; ambulatory legs proximally reddish-brown (Fig. 3 B – D). Geographical distribution. India to Philippines (Tan 1996), Vanuatu (Galil & Ng 2010). New record for Papua New Guinea.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3374C119699F9F18DBF8AE2.taxon	materials_examined	Material examined. 1 juvenile male (7.8 mm) (MNHN-IU- 2013 - 12748), stn PD 30, Budup, 05 ° 04.7 ' S 145 ° 48.3 ' E, 1 – 8 m, 17.11.2012; 1 female (12.4 mm) (MNHN-IU- 2013 - 12742); 1 ovigerous female (14.0 mm) (ZRC 2014.0173), stn PD 38, Rempi Area, north Tadwai I., 04 ° 58.8 ' S 145 ° 47.4 ' E, 10 – 15 m, 20.11.2012. Colour in life. Carapace and legs olive-coloured, granules on margins of carapace and cheliped merus duncoloured; inner margin of fingers orange, tips white (Fig. 3 E). Geographical distribution. New Caledonia, Australia (Queensland, Western Australia) (Galil 2009). New record for Papua New Guinea.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3394C1F9699FF07886B8FD5.taxon	materials_examined	Material examined. 1 immature specimen (6.6 mm) (MNHN-IU- 2013 - 462), stn PD 10, west Kranket I., 05 ° 11.7 ' S 145 ° 48.8 ' E, 16 – 28 m, 9 – 10.11.2012; 1 immature female (6.7 mm) (MNHN-IU- 2013 - 620), stn PD 22, S Sek I., 05 ° 06.3 ' S 145 ° 49.2 ' E, 12 – 20 m, 14 – 15.11.2012; 1 immature specimen (9.3 mm) (MNHN-IU- 2013 - 17891), stn PS 25, Barag I., outer slope, 05 ° 01,2 ' S 145 ° 48,0 ' E, 24.11.2012; 1 immature female (9.4 mm) (MNHN-IU- 2013 - 1226), stn PD 74, Yomba I. Beach, 05 ° 14.7 ' S 145 ° 47.4 ' E, 1 – 3 m, 7.12.2012. Colour in life. Carapace bone-coloured, mottled with irregular brown patches and spots, front mauve, pale crescent on intestinal region, median posterior spine white. Cheliped merus bone-coloured with broad brown bands medially and distally, 2 dark dots between bands; dark bands proximally and distally on palms; fingers mottled brown. Ambulatory legs with brown bands distally on meri, proximally on carpi, propodi (Fig. 3 F). Geographical distribution. Philippines, Indonesia, Australia (Western Australia), Vanuatu, New Caledonia, Fiji, 1 – 120 m (Galil 2001 b; Galil & Ng 2010). New record for Papua New Guinea.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3394C1F9699FCDD8CBD8913.taxon	materials_examined	Material examined. 1 male (19.6 mm) (MNHN-IU- 2013 - 2010), stn PP 06, Rempi Area east Tadwai I., muddy, 04 ° 59 ' S 145 ° 48 ' E, 180 m, 19.11.2012. Colour in life. Carapace mottled with irregular reddish brown patches and spots, lateral posterior denticles bone-coloured, median posterior spine proximally mottled orange, distally white. Cheliped merus mottled reddish brown, with broad brown bands medially and distally; palms proximally white; fingers mottled brown, distally white. Ambulatory legs with 2 brown bands across meri, proximally on carpi, propodi (Fig. 3 G). Geographical distribution. South China Sea, Timor Sea; 50 – 180 m (Galil 2004). New record for Papua New Guinea.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3394C1F9699FB1E8DC08BE0.taxon	materials_examined	Material examined. 1 immature specimen (7.1 mm) (MNHN-IU- 2013 - 609), stn PR 47, Siar I., 05 ° 11.2 ' S 145 ° 48.4 ' E, 16.11.2012; 1 immature specimen (8.1 mm) (MNHN-IU- 2013 - 12728), stn PD 53, south Panab I., 05 ° 10.3 ' S 145 ° 48.5 ' E, 2 – 10 m, 28.11.2012; 1 male (17.9 mm) (MNHN-IU- 2013 - 17890), stn PD 79, between Taush I. and Malamal I., 05 ° 07 ' S 145 ° 48.5 ' E, 20 m, 10.12.2012. Colour in life. immature: carapace pale bone, granules pale orange. Cheliped merus with pale orange bands medially and distally; palm white, fingers darker. Ambulatory legs with brown bands distally on meri, proximally on carpi, propodi (Fig. 3 H). Geographical distribution. Gulf of Aden, Andaman Is., Thailand (Gulf of Thailand), Vietnam, Philippines, Indonesia, Arafura Sea, New Guinea, Torres Strait, Australia (western Australia), Chesterfield Is., New Caledonia; 8 – 69 m (Galil 2001 b).	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3394C1E9699F8FE8D798C28.taxon	materials_examined	Material examined. 1 ovigerous female (10.3 mm) (MNHN-IU- 2013 - 834), stn PD 53, south Panab I., 05 ° 10.3 ' S 145 ° 48.5 ' E, 2 – 10 m, 28.11.2012; 1 male (7.2 mm) (ZRC 2014.0165), 1 female (6.7 mm) (MNHN-IU- 2013 - 1224), stn PD 75, Madang South hospital, 05 ° 14.2 ' S 145 ° 47.8 ' E, 10 – 25 m, 7.12.2012. Colour in life. Carapace bone-white, medially mottled reddish-brown, bisected by narrow white stripe running from bidentate front to posterior margin. Cheliped merus with pale orange band medially; distal part of palm, fingers mottled pale brown. Ambulatory legs with 2 orange bands on meri, proximally on carpi, propodi (Fig. 4 A). Geographical distribution. Red Sea to Fiji (Tan 1996), Vanuatu (Galil & Ng 2010). New record for Papua New Guinea.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3384C1E9699FE068EFD8FA0.taxon	materials_examined	Material examined. 1 male (4.3 mm) (MNHN-IU- 2013 - 17889), stn PB 34, N Wonad I., N Mililat Passage, 05 ° 07.1 ' S 145 ° 49.4 ' E, 15 m, 3.12.2012. Geographical distribution. Chesterfield Is., New Caledonia (Tan & Richer de Forges 1993); Amami-Oshima I. Ryukyu Is., Japan (Takeda & Komatsu 2005).	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3384C1E9699FCBE8E7B8938.taxon	materials_examined	Material examined. 1 immature male (35.1 mm) (MNHN-IU- 2013 - 1044), stn CP 3968, east Kranket I., 05 ° 12 ' S 145 ° 51 ' E, 380 m, 2.12.2012. Geographical distribution. Japan, Taiwan, Philippines, Indonesia, Solomon Is., 100 – 425 m (Galil 2007). New record for Papua New Guinea.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3384C1E9699FB368DCC8B2D.taxon	materials_examined	Material examined. 1 ovigerous female (10.5 mm) (ZRC 2014.0169), stn PD 45, Rempi Area, west Barag I., 05 ° 01.2 ' S 145 ° 47.9 ' E, 5 – 10 m, 22 – 23.11.2012; 1 ovigerous female (10.3 mm) (MNHN-IU- 2013 - 8853), stn PD 48, Wonad I., 05 ° 08.2 ' S 145 ° 49.4 ' E, 10 – 20 m, 26.11.2012. Colour in life. Dorsal surface of carapace grayish pink, posterior tubercles white; cheliped merus with brown band medially (Fig. 4 C). Geographical distribution. Indonesia, Vanuatu (Galil & Ng 2010), New Caledonia (Galil 2001 c). New record for Papua New Guinea.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3384C1C9699F9478DBF8DBD.taxon	materials_examined	Material examined. 1 young female (10.4 mm) (MNHN-IU- 2013 - 117), stn PR 07, Madang Harbour, 05 ° 12.5 ' S 145 ° 48.5 ' E, 2 – 17 m, 7.11.2012. Colour in life. Dorsal surface of carapace deep gold-coloured, frontal region laterally framed with dark lines, intestinal tubercle white, orange crescents posteriorly to submedian branchial spines. Cheliped merus dark orange, palm and fingers pale orange. Ambulatory legs with 2 orange bands on meri, proximally on carpi, propodi (Fig. 4 D). Geographical distribution. Madagascar to New Caledonia (Galil 2001 c), Vanuatu (Galil & Ng 2010). New record for Papua New Guinea.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E33A4C1C9699FEB288AC8BB5.taxon	materials_examined	Material examined. Holotype: female (11.5 mm) (MNHN-IU- 2013 - 12749), stn PD 72, between Yabob & Bil Bil I., 05 ° 16.2 ' S 145 ° 46.5 ' E, 20 – 25 m, 6.12.2012. Paratypes: 1 ovigerous female (9.6 mm) (ZRC 2015.270), same data as holotype; 1 ovigerous female (12.5 mm) (MNHN-IU- 2013 - 12750), stn PD 68, south Kesup Gun River mouth, 05 ° 16 ' S 145 ° 46.5 ' E, 2 – 3 m, Papua New Guinea, 4.12.2012.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E33A4C1C9699FEB288AC8BB5.taxon	description	Description. Carapace discoidal, with shallow branchio-cardiac grooves (Fig. 9 A, B). Dorsal surface of carapace minutely granular anteriorly, with closely-spaced pearliform granules on branchial, cardial, intestinal regions. Frontal margin horizontal, setose (Fig. 9 C). Margins of carapace closely beaded, granules smaller anteriorly (Fig. 9 E). Hepatic facet distinct. Pterygostomian region prominently granular (Fig. 9 F). Epimeral, posterior surfaces minutely granular. Antennular fossa sealed by basal antennular article. Antennae short, inserted between antennular fossa, orbit. Eyes retractable. Epistome continuous with expanded, beaded subhepatic margin, projecting beyond frontal margin, medially notched. Anterior margin of efferent branchial channel forms part of lower orbital margin, entire, fissure situated ventrally to orbit. Third maxilliped exopod semicircular (Fig. 9 F), expanded, densely setose; endopod triangular, merus densely setose, shorter than ischium along inner margin, apex invisible in dorsal view. Chelipeds subequal. Cheliped merus 0.6 as long as carapace in female; entirely covered with pearliform granules, decreasing in size distally (Fig. 10 A – C). Carpus with row of minute granules on inner margin; patch of minute granules distally on dorsal margin. Dorsal margin of chela cristate, with 2 parallel rows of granules, ventral margin minutely granular. Fingers slightly longer than palm, cutting edges cristate, distally denticulate. Ambulatory legs slender, short, meri subcylindrical, longer than carpi, propodi, ventral margin minutely granular; dorsal, ventral propodal margins carinate; dactyli longer than propodi, lanceolate, terminating in cornute tips. Female abdomen with first 2 somites transversely narrow, yoke-like, granular; somites 3 – 6 fused, greatly enlarged, shield-like, proximally granular; telson laciniate (Fig. 10 E). Thoracic sternites with closely spaced pearliform granules. Vulvae submedially near suture 6 / 5; small, rounded, opening interiorly, hooded by calcified pear-shaped dome (Fig. 10 D). Colour in life. Dorsal surface of carapace and chelipeds brown, granules bright orange-red, pale blotches on post orbital regions, and on both sides of cardiac region. Rounded knob distally on cheliped merus bright red, finger tips white. Ambulatory legs pale, spotted red on merus and carpus (Fig. 4 E).	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E33A4C1C9699FEB288AC8BB5.taxon	discussion	Remarks. Ryphila bertrandi sp. nov. shares with it congeners R. cancellus (Herbst, 1783) and R. verrucosa (Henderson, 1893) a projecting epistome and ventral margin of hepatic facet and reniform third maxilliped exopod. It is clearly distinguished from both, however, in having cheliped merus entirely covered with pearliform granules dorsally rather than with several rows of pearliform granules, and the carapace dorsally minutely granular anteriorly, with larger pearliform granules on the branchial, cardiac and intestinal regions.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E33A4C1C9699FEB288AC8BB5.taxon	etymology	Etymology. In honour of our good friend, Bertrand Richer de Forges, intrepid explorer and collector of crabs from all over the Indo-West Pacific. Geographical distribution. This species is known only from type location, Papua New Guinea.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E33A4C1B9699F8B4897A8B60.taxon	materials_examined	Material examined. Holotype: male (33.0 mm) (MNHN-IU- 2013 - 7963), stn CP 3984, Bismarck Sea, northwest Long I., 05 ° 12 ' S 146 ° 59 ' E, 500 m, 6.12.2012. Paratypes: 3 males (32.4 mm, 33.8 mm, 32.5 mm) (MNHN-IU- 2013 - 17899), 1 male (33.1 mm) (ZRC 2015.271), same data as holotype; 1 immature male (18.9 mm) (MNHN-IU- 2013 - 8886), stn CP 3979, north Bagabag Is, 04 ° 44 ' S 146 ° 11 ' E, 540 – 580 m, Bismarck Sea, 5.12.2012. Comparative material. Tanaoa distinctus (Rathbun, 1894): 1 male (37.5 mm), 1 female (ZRC 2000.535), station 34, TC 33, Hawaiian Islands, coll. Townsend Cromwell. Tanaoa serenei (Richer de Forges, 1983): 1 male (33.2 mm) (ZRC 2000.565), Tumon Bay, Guam, 400 m, in fishtrap, coll. S. Annesbury, 10.1999.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E33A4C1B9699F8B4897A8B60.taxon	description	Description. Carapace rounded, slightly longer than wide, globose; cardiac, intestinal regions laterally demarcated by grooves (Fig. 13 A). Dorsal surface covered densely by rounded granules of various sizes, granules somewhat larger, conical, on branchial regions; 4 pairs of pits along branchiocardiac line (Fig. 13 A, B). Intestinal region swollen, medially marked with granular tubercle, directed posteriorly (Fig. 13 A). Front narrow, slightly produced, upturned, closely set with granules, divided into 2 subtriangular lobes by minutely setose gap (Fig. 13 A – C). Eyes small, retractable within orbit, setose eyestalk exposed. Outer orbital margin with 3 sutures. V-shaped gap proximally on ventral margin. Antennular fossae below frontal lobes oblique, antennules obliquely folded, basal antennular operculiform, sealing lower half of antennular aperture when retracted. Antennae small, slender, basal antennal article inserted in orbital hiatus. Postorbital region concave. Anterior margin of efferent branchial channel convex, produced, bilobed, separated by narrow groove from lower orbital margin. Outer surface of third maxillipeds granular, granules more closely set, larger anteriorly, forming granular ridge mesially on ischium (Fig. 13 B, C). Subhepatic region produced, angular, furnished with subconical granules, medially with granular tubercle (Fig. 13 B, C). Epibranchial margin with 3 equidistant granular tubercles (Fig. 13 A). Posterolateral margins rounded. Posterior margin of carapace narrow, laterally with 2 prominently granular tubercles (Fig. 13 A). Chelipeds slender, subequal, covered with small granules on all articles, including fingers (Fig. 13 A, F). Cheliped merus, subcylindrical, 1.2 as long as carapace; palm subcylindrical, 0.6 as long as merus, fingers 0.6 as long as dorsal margin of palm, inner margins obsoletely denticulate. Ambulatory legs slender; decreasing in size posteriorly; merus, carpus, propodus granular, granules more prominent dorsally; dorsal surface of dactylus setose, dactylar tips corneous (Fig. 13 G). Thoracic sternites (Fig. 13 D) closely set with flattened granules. Male abdominal cavity deep, nearly reaching buccal cavity, anterior margin ogive, slightly raised (Fig. 13 C, D). Male abdomen triangular, elongated; abdominal somites 1, 2 transversely narrow; somite 1 yoke-like, somite 2 medially convex as it fits into cavity of somite 1; somites 3 – 6 fused, proximo-lateral regions inflated, granular, distal margin with triangular denticle; telson slender, 1 / 3 as long as fused somites, not reaching tip of abdominal cavity (Fig. 13 D). G 1 elongated, slightly sinuous, attenuate, with small but distinct preapical bifurcated snout-like process perpendicular to tip, opening facing dorsally (Fig. 15 A – D); G 2 short, distally scoop-like (Fig. 15 E). Colour in life. Dorsal surface of carapace and chelipeds bright orange-red. Ambulatory legs pale, meri banded in red (Fig. 4 F).	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E33A4C1B9699F8B4897A8B60.taxon	etymology	Etymology. Kuka for “ crab ” in Tok Pisin, the Pidgin language spoken in Papua New Guinea. The name is used as a noun in apposition.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E33A4C1B9699F8B4897A8B60.taxon	discussion	Remarks. See discussion for T. retpela sp. nov. Geographical distribution. This species is known only from the type location, Papua New Guinea.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E33D4C0F9699F98389768ECD.taxon	materials_examined	Material examined. Holotype: male (36.5 mm) (MNHN-IU- 2013 - 7961), stn CP 3984, Bismarck Sea, northwest Long I., 05 ° 12 ' S 146 ° 59 ' E, 500 m, 6.12.2012. Paratypes: 1 male (39.0 mm) (ZRC 2015.272), stn CP 3978, Bismarck Sea, north Bagabag Is., 04 ° 45 ' S 146 ° 12 ' E, 456 – 582 m, 5.12.2012. Additional material: BIOPAPUA – 1 female (29.5 mm) (MNHN-IU- 2011 - 3835), stn CP 3653, west of New Hanover, 02 ° 13 ’ S 150 ° 23 ’ E, 680 – 700 m, 28.08.2010; 2 males (38.4 mm, 29.8 mm) (MNHN-IU- 2011 - 914), stn CP 3655, west of New Hanover, 02 ° 15 ’ S 150 ° 16 ’ E, 402 – 440 m, 28.08.2010; 1 male (39.1 mm) (MNHN-IU- 2011 - 2098), stn CP 3669, north of Rabaul, 04 ° 08 ’ S 151 ° 56 ’ E, 382 – 389 m, 24.09.2010; 1 female (28.7 mm) (MNHN-IU- 2011 - 2481), stn CP 3681, Vitu I., 04 ° 38 ’ S 149 ° 27 ’ E, 564 – 712 m, 27.09.2010; 1 male (28.6 mm) (MNHN-IU- 2011 - 2478), stn CP 3682, Vitu I., 04 ° 38 ’ S 149 ° 28 ’ E, 515 – 812 m, 27.09.2010; 3 juveniles (11.7 – 12.5 mm) (MNHN-IU- 2011 - 2563), stn DW 3748, seamounts near Bougainville, 05 ° 37 ’ S 154 ° 01 ’ W, 398 – 399 m, 12.10.2010; 1 male (21.4 mm) (MNHN-IU- 2011 - 2710), stn CP 3695, Feni Is., 02 ° 10 ’ S 147 ° 15 ’ W, 198 m, 14.10.2010; 3 females (19.5 – 21.0 mm) (MNHN-IU- 2011 - 2299), stn CP 3760, Is., 03 ° 58 ’ S 153 ° 43 ’ W, 613 – 660 m, 14.10.2010.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E33D4C0F9699F98389768ECD.taxon	description	Description. Carapace rounded, slightly wider than long, globose; gastric, cardiac, intestinal regions laterally demarcated by deep grooves (Fig. 14 A). Dorsal surface covered densely by rounded, pearliform granules of various sizes; 4 pairs of pits along branchiocardiac line, additional pair on mesogastric region (Fig. 14 A). Intestinal region swollen, anteriorly with rounded median carina, posteriorly with granular tubercle (Fig. 14 A). Front narrow, slightly produced, upturned, closely set with granules, divided into 2 rounded lobes (Fig. 14 A – C). Eyes small, retractable within orbit, minutely granulose eyestalk exposed (Fig. 14 B). Outer orbital margin with 3 sutures. Vshaped gap proximally on ventral margin. Antennular fossae below frontal lobes oblique, antennules obliquely folded, basal antennular operculiform, sealing lower half of antennular aperture when retracted. Antennae small, slender, basal antennal article inserted in orbital hiatus. Postorbital region concave. Anterior margin of efferent branchial channel convex, produced, bilobed, separated by narrow groove from lower orbital margin (Fig. 14 B, C). Outer surface of third maxillipeds granular, granules more prominent anteriorly, forming granular ridge mesially on endopod (Fig. 14 C). Hepatic margin medially with granular tubercle (Fig. 14 B, C). Subhepatic region produced, inflated, with horizontal line medially free of granules (Fig. 14 B, C). Epibranchial margin with 3 equidistant granular tubercles (Fig. 13 A). Posterolateral margins rounded. Posterior margin of carapace narrow, laterally with 2 prominently granular tubercles (Fig. 14 A). Chelipeds slender, subequal, covered with small granules on all articles, including fingers (Fig. 14 A, F). Cheliped merus, subcylindrical, not nearly as long as carapace; palm subcylindrical, 0.6 as long as merus; fingers longer than dorsal margin of palm, granules arraigned in longitudinal lines, cutting edges denticulate (Fig. 14 A, F). Ambulatory legs slender; decreasing in size posteriorly; merus, carpus, propodus granular, granules more prominent dorsally; dorsal surface of dactylus setose, dactylar tips corneous (Fig. 14 A, G). Thoracic sternites set with granules of various sizes; sternite 4 anteriorly inflated (Fig. 14 D). Male abdominal cavity deep, nearly reaching buccal cavity, anterior margin of cavity ogive, slightly raised (Fig. 14 D). Male abdomen triangular, elongated; abdominal somites 1, 2 transversely narrow; somite 1 yoke-like, somite 2 medially convex as it fits into cavity of somite 1; somites 3 – 6 fused, proximo-lateral regions greatly inflated, granular; subterminally with horizontal ridge, anterior margin with quadrate denticle; telson slender, 1 / 3 as long as fused somites, not reaching tip of abdominal cavity (Fig. 14 D, E). G 1 elongated, slightly sinuous, attenuate, with preapical digitate process perpendicular to tip, opening facing dorsally (Fig. 15 F – J); G 2 short, distally scoop-like (Fig. 15 K). Colour in life. Carapace bright orange-red mottled bone-white. Cheliped merus pale orange, distal part bright orange; palm, fingers mottled pale orange. Ambulatory legs pale (Fig. 5 A).	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E33D4C0F9699F98389768ECD.taxon	etymology	Etymology. From retpela for “ red ” or “ orange ” in Tok Pisin, the Pidgin language spoken in Papua New Guinea. The name is used as a noun in apposition.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E33D4C0F9699F98389768ECD.taxon	discussion	Remarks. Tanaoa distincta, T. serenei, T. kuka sp. nov. and T. retepela sp. nov. are all superficially similar in external appearance. Adults, however, are easy to separate using several reliable characters. Ng & Richer de Forges (2007) appraised the taxonomy of T. distincta and T. serenei, noting that the granulation degree and pattern on the carapace was reliable as a character, and their G 1 differ. Among the four species, the granules on the carapace are finest in T. kuka sp. nov. (Fig. 13 A, B), T. distincta slightly coarser (Fig. 11 A, B). In T. serenei, the granules are large and densely packed (Fig. 12 A, B), while in T. retepela sp. nov. the granules are spaced further apart (Fig. 14 A, B). Tanaoa kuka sp. nov. is distinguished from the named species by its proportionately elongated chela and palm (Figs. 13 A, F, 11 A, F, 12 A, F, 14 A, F). With regards to the ambulatory legs, the meri of T. serenei and T. kuka sp. nov. are proportionately the most slender and longest; but whereas in T. serenei, the meri bear sharp granules (Fig. 12 A, G) in T. kuka sp. nov. the meri are smoother (Fig. 13 A, G). The ambulatory meri of T. distincta and T. retepela sp. nov. are proportionately shorter (Figs. 11 A, G, 14 A, G). The G 1 of T. distincta is distinct in that the distal part has broad folds or is only subtruncate, lacking subdistal processes (Ng & Richer de Forges 2007: fig. 4 A – D). The G 1 of T. kuka sp. nov. bears a subdistal bifurcate process (Fig. 15 C, D). The G 1 of T. serenei and T. retepela sp. nov. are superficially similar (Fig. 15 F – J; Ng & Richer de Forges 2007: fig. 4 E, F) in that there is a long subdistal process which tapers to a sharp tip. The overall gonopod structure of T. retepela sp. nov. is nevertheless relatively stouter and the subdistal process relatively shorter (Fig. 15 F – J) compared to the more slender one of T. serenei which has a relatively longer subdistal process (Ng & Richer de Forges 2007: fig. 4 E, F). Geographical Distribution. This species is known only from the type locality, Papua New Guinea.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3294C0F9699FBE28E2A8B6D.taxon	materials_examined	Material examined. 1 male (11.1 mm) (MNHN-IU- 2013 - 651), stn CP 3948, west Kranket I., 05 ° 12 ' S 145 ° 51 ' E, 363 – 388 m, 26.11.2012; 1 female (12.2 mm) (MNHN-IU- 2013 - 8939), 1 female (12.5 mm) (MNHN-IU- 2013 - 8932), stn CP 3949, west Kranket I., 05 ° 12 ' S 145 ° 51 ' E, 380 – 407 m, 26.11.2012; 1 male (11.1 mm), 1 female (12.8 mm) (MNHN-IU- 2013 - 11291), stn CP 4023, Astrolabe Bay, 05 ° 22 ' S 145 ° 48 ' E, 340 – 385 m, 14.12.2012; 1 female (10.6 mm) (MNHN-IU- 2013 - 1504), stn DW 4031, Cape Croisiles, 04 ° 53 ' S 145 ° 49 ' E, 370 m, coll. 16 December 2012; 1 male (9.2 mm) (MNHN-IU- 2013 - 8885), stn CP 4035, east Kotakot, 04 ° 31 ' S 145 ° 31 ' E, 380 – 382 m, 17.12.2012; 1 male (11.5 mm) (MNHN-IU- 2013 - 1556), 3 males (10.5 mm, 12.1 mm, 11.5 mm) (ZRC 2014.0164), stn CP 4055, north Aitape, 03 ° 03 ' S 142 ° 18 ' E, 370 – 374 m, Bismarck Sea, 20.12.2012. Geographical distribution. Philippines, Solomon Is., Vanuatu, New Caledonia, Fiji, 230 – 620 m (Galil 2003, 2007). New record for Papua New Guinea.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3294C0F9699F9428EEC8ABD.taxon	materials_examined	Material examined. 1 broken juvenile female (MNHN-IU- 2013 - 1494), stn CP 4035, East Kotakot, 03 ° 31 ' S 145 ° 31 ' E, 380 – 382 m, 17.12.2012; 1 male (36.9 mm) (ZRC 2014.0166), 1 female (38.6 mm) (MNHN-IU- 2013 - 7923), stn CP 4055, north Aitape, 03 ° 03 ' S 142 ° 18 ' E, 370 – 374 m, Bismarck Sea, 20.12.2012; 1 male (28.6 mm) (MNHN-IU- 2013 - 7965), stn CP 4066, Dogreto Bay, 03 ° 18 ' S 143 ° 02 ' E, 440 – 440 m, Bismarck Sea, 22.12.2012. Geographical distribution. Japan, Taiwan, Philippines, Indonesia, Australia, Solomon Is., 85 – 839 m (Galil 2003, 2007). New record for Papua New Guinea.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3284C0E9699FEFD8E7B8FCA.taxon	materials_examined	Material examined. 1 male (20.8 mm) (MNHN-IU- 2013 - 1199), stn PR 176, Madang, Madang Resort, 05 ° 12.5 ' S 145 ° 48.5 ' E, 3 – 17 m, Papua New Guinea, 4.12.2012. Colour in life. Carapace ochre-orange, darker anteriorly, 2 orange spots posteriorly near lateral corners of posterior margin; dorsal surface of cheliped merus, carpus ochre, granules pale orange, bright-orange patch on inner surface of palm. Epimeral ledge white. Ambulatory legs white, banded with orange on merus, propodus (Fig. 5 E). Geographical distribution. Singapore, Philippines, Indonesia, New Caledonia (Galil 2003; Galil & Ng 2007). New record for Papua New Guinea.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3284C0B9699FCE588AC8CC5.taxon	materials_examined	Material examined. Holotype: 1 male (12.6 mm) (MNHN-IU- 2013 - 12758), stn PD 73, Yomba I., 05 ° 14.7 ' S 145 ° 47.3 ' E, 20 – 25 m, 6.12.2012. Paratypes: 1 male (12.4 mm) (ZRC 2015.273), 1 male (10.5 mm), 1 female (13.2 mm) (MNHN-IU- 2013 - 12756), stn PD 73, Yomba I., 05 ° 14.7 ' S 145 ° 47.3 ' E, 20 – 25 m, 6.12.2012; 1 male (11.6 mm), 2 females (11.8, 11.5 mm) (MNHN-IU- 2013 - 12755), stn PD 63, north Banap Damon Point, 05 ° 09.9 ' S 145 ° 48.3 ' E, 0 – 15 m, 1.12.2012.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3284C0B9699FCE588AC8CC5.taxon	description	Description. Carapace subpentagonal, glabrous; regions of carpace indistinct; dorsal surface smooth, finely punctate (Figs. 16 A, 17 A). Frontal region constricted, produced, laterally concave; frontal margin mucronate. Antennular fossae continuous with orbit, sealed by basal antennular article; antennules fold obliquely within fossa. Antennae short, inserted between antennular fossa, orbit. Eyes retractable. Orbits small, rounded, outer orbital margin entire, anterior margin of efferent branchial channel forms part of lower orbital margin. Third maxillipeds concealing trapezoid buccal cavity; endopod merus triangulate, as long as ischium, planar (Fig. 16 C); in female, endopod with setose fringe lengthwise, ischium medially keeled (Fig. 17 B). Anterolateral margins of carapace oblique, margin minutely beaded (Fig. 16 B). Lateral angle of carapace prominent, overhanging tomentose thoracic sinus, margin sinuous, distinctly milled. Thoracic sinus shallow, anteriorly defined by straight, smooth margin of pterygostomian region (Fig. 16 C, D); ventral margin of sinus, above base of cheliped, with row of granules continuous with epimeral ridge. Posterolateral margins of carapace devoid of granulation. Epimeral margin visible in dorsal view, slightly notched medially, shallow gutter-like anteriorly, narrowing posteriorly, continuous with posterior margin (Figs. 16 A, 17 A). Posterior margin nearly horizontal in male, rounded in female (Figs. 16 A, 17 A), slightly deflexed, surface granular (Fig. 16 H, I). Chelipeds subequal, robust, slightly longer in adult male than in female (Figs. 16 A, 17 A). Cheliped merus trigonal in cross-section, with pearliform granules on anterior, posterior margins; patch of spongy pubescence proximally on dorsal surface, bordered basally by coalesced cluster of granules (Fig. 16 A, E), 3 pearliform granules anteriorly; ventral surface pitted proximally, nearly smooth (Fig. 16 F). Carpus rounded, cluster of granules on inner margin. Propodus somewhat inflated, dorsal margin obscurely beaded, ventral surface with prominently granular rows at inner, outer margins, continuing onto pollex. Fingers as long as dorsal margin of palm (Fig. 16 E). Ambulatory legs slender, short. Meri of first and second legs with 2 minutely beaded lines on dorsal margin, posterior line short; 2 beaded lines on ventral margin. Meri of third and fourth legs with single beaded line dorsally; 2 beaded lines ventrally (Fig. 16 G), P 5 merus proximally constricted in female, posterior margin prominently crested. Dorsal margin of carpi of ambulatory legs prominently carinate, propodi dorsally, ventrally carinate. Dactyli longer than propodi, dorsoventrally flattened, lanceolate (Fig. 16 G). Male abdominal cavity deep, elongated, nearly reaching buccal cavity (Fig. 16 C). Male abdomen with somite 2 minute; somites 3 – 6 fused, tapering distally, with a non-articulating suture, constriction at commissure of somites 5, 6, opposite ridge on lateral wall of abdominal cavity, triangular denticle fronted by an indentation, halfway lengthwise; telson triangular (Fig. 16 D). Female abdomen with somites 3 – 6 fused, greatly enlarged, shield-like, telson triangular (Fig. 17 B, C). G 1 elongated, shaft straight, twisted once on its axis, distally club-like (Fig. 18 A – D). G 2 short, curved, apex scoop-like (Fig. 18 E). Vulvae prominent, ovoid; anterolaterally hooded by calcified peaked semi-dome; opening, directed posteriorly, kidney-shaped (Fig. 17 D). Colour (alcohol preserved specimens). Dorsal surface of carapace and chelipeds grayish-taupe peppered with fine dark grey dots, paler posteriorly; pearliform granules on cheliped merus yellowish ivory, fingers proximally barred pale yellow. Ambulatory legs whitish, meri tinted yellow distally.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3284C0B9699FCE588AC8CC5.taxon	discussion	Remarks. The new species is superficially similar to the three Seulocia species with a granular deflexed surface of the posterior carapace margin: S. cristata Galil, 2005, S. rhomboidalis (De Haan, 1841) and S. vittata (Stimpson, 1858). Yet, it is easily distinguished from these congeners. The front of S. cristata is strongly deflexed and medially cristate, whereas the front of the new species is neither deflexed nor cristate, but mucronate. The carapace of S. rhomboidalis is only slightly longer than wide, its palms are lenticular, and it bears five red spots on each side of the median line of the carapace, arranged in arcuated diverging lines, whereas the carapace of the new species is 1.3 as long as wide, its palm distinctly longer than wide, and it lacks the red spots. Seulocia vittata has the thoracic sinus anteriorly defined by the granular, overhanging, margin of the pterygostomian region and smooth dorsal margin of the palm, whereas the pterygostomian margin in the new species is smooth, and the dorsal margin of the palm indistinctly beaded.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E3284C0B9699FCE588AC8CC5.taxon	etymology	Etymology. This new species is named after Janet Gagul, Project Officer of the Papua Niugini Expedition in Port Moresby in 2012 – 2013. Janet was instrumental in facilitating permits and logistics for the expedition, skillfully and efficiently liaising with PNG authorities, administrations, service providers, and scientists. Geographical distribution. This species is known only from type location, Papua New Guinea.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E32D4C0A9699F99A8EED8DBD.taxon	materials_examined	Material examined. 1 male (10.6 mm) (MNHN-IU- 2013 - 615); 1 female (11.6 mm) (MNHN-IU- 2013 - 12759), stn PD 22, south Sek I., 05 ° 06.3 ' S 145 ° 49.2 ' E, 12 – 20 m, 14 – 15.11.2012; 1 male (11.3 mm) (ZRC 2014.0167), stn PD 45, Rempi Area, west Barag I., 05 ° 01.2 ' S 145 ° 47.9 ' E, 5 – 10 m, 22 – 23.11.2012. Colour in life. Dorsal surface of carapace bone coloured, minutely dotted red on post frontal ridge, pale brownish blotches on hepatic regions, dotted red, pale brownish stripe medially on gastric, cardiac regions, pale brownish blotches laterally near branchial margins. Cheliped pale pink, granules slightly darker pink; fingers proximally red. Ambulatory legs with pale orange spots (Fig. 5 D). Geographical distribution. South China Sea, Philippines, Singapore, Indonesia, Arafura Sea, Australia (western coast), Papua New Guinea (Galil 2005).	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E32C4C089699FEF789248CE0.taxon	materials_examined	Material examined. Holotype female (8.3 mm) (IRSNB-RBINS IG. 26253), stn 69, Purar Reef, Hansa Bay, Madang Province, 10 m, 4 ° 11 ’ 47 ” S 144 ° 55 ’ 05 ” E, coll. J. Pierret, 29.10.1980. Additional material: 1 female (8.3 mm) (MNHN-IU- 2013 - 17898), 1 male (7.6 mm) (MNHN-IU- 2013 - 17897), 1 ovigerous female (8.9 mm) (ZRC 2015.274). Madang area, no precise data, coll. Papua Niugini Biodiversity Expedition, 2012, coll. 10 – 12.2012.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E32C4C089699FEF789248CE0.taxon	discussion	Remarks. The type specimen, a female, was examined and it agrees very well with several specimens collected by the expedition from Papua New Guinea. Unfortunately, there was no station data with the recent specimens other than a “ PNG ” code and they were with the rest of the expedition material. So while we are certain they were obtained in the expedition area, we have no precise data. In any case, the type locality, Hansa Bay, is within the expedition area. A detailed description of the type specimen is given in Ovaere (1987). The vulvae, which were not described, are prominent, large, anteriorly overreaching suture 5 / 4; ovoid, and interiorly bordered by a calcified, rounded raised rim, narrowing anteriorly, with the outer margin crested anteriorly and the opening facing interiorly, occluded, fibula-shaped, with the narrow part pointing anteriorly (Fig. 19 H). The male is recorded for the first time, and it is figured in detail here (Figs. 20, 23 E – H). Urnalana purarensis is closest to U. flammea sp. nov. but differs in a number of carapace, ambulatory leg and gonopod characters (see discussion for next species). Geographical distribution. Papua New Guinea: Madang Province, Hansa Bay, Purar Reef (Ovaere 1987).	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E32E4C359699FE0388AC8B38.taxon	materials_examined	(Figs. 5 F, 21, 22 A – G, 23 C, D, I – M) Material examined. Holotype: male (9.3 mm) (MNHN-IU- 2013 - 1341), stn PR 227, Madang Resort, 05 ° 12.5 ' S 145 ° 48.5 ' E, 10.12.2012. Paratypes: 1 female (7.6 mm) (ZRC 2015.275), stn PR 14, Biliau I., 05 ° 12 ' S 145 ° 48.1 ' E, 2 – 3 m, 9.11.2012; 1 male (9.0 mm) 1 ovigerous female (10.7 mm) (MNHN-IU- 2013 - 13123) stn PR 28, Madang Resort, 05 ° 12.5 ' S 145 ° 48.5 ' E, 13.11.2012; 1 male (9.2 mm) (ZRC 2015.276), stn PR 245, Tabad I., 05 ° 08.2 ' S 145 ° 48.7 ' E, 12 m, 12.12.2012.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E32E4C359699FE0388AC8B38.taxon	description	Description. Carapace subpentagonal, urn-shaped, globose; regions of carapace ill defined (Fig. 21 A). Dorsal surface of carapace sparsely setose, punctate. Front somewhat produced, upturned, medially concave, lateral lobes rounded (Fig. 21 D). Postfrontal region laterally concave. Anterolateral margin convex, smooth, medially inflated in dorsal view; swollen distal margin of pterygostomian region invisible in dorsal view. Lateral angle of carapace rounded, overhanging thoracic sinus, lacking setose fringe (Fig. 21 B). Thoracic sinus deep, anteriorly defined by vaulted margin of pterygostomian region (Fig. 21 D, G); cluster of minute granules overgrown by club-like setae anteriorly; row of pearliform granules on ventral surface of epibranchial angle; another row above base of first pair of ambulatory legs. Epimeral ledge visible over its entire length in dorsal view, continuous with posterior margin, margin beaded, granules larger anteriorly. Posterolateral margin sparsely setose. Posterior margin minutely beaded (Fig. 21 A). Antennular fossa sealed by basal antennular article. Antenna short, nestled underneath antennular fossa (Fig. 21 C). Orbits small, rounded, outer orbital margin with 1 suture. Anterior margin of efferent branchial channel forms part of lower orbital margin, short suture below eye. Eyes retractable. Third maxillipeds concealing trapezoid buccal cavity; smooth, planar, lacking denticle anterolaterally on ischium (Fig. 21 D), setose fringe lengthwise on endopod of female. Chelipeds subequal, robust, longer in adult male than in female specimens (Fig. 21 A). Cheliped merus trigonal, merus 2 / 3 as long as carapace. Dorsal surface proximally with cluster of granules partly obscured by patch of club-shaped setae; anterior margin with 3 large, pearliform granules proximally, row of small granules distally (Fig. 21 E, 22 F, G); posterior margin with row of lozenge-shaped granules, largest medially; ventral surface proximally pitted, granular. Carpus subglobular, granules proximally on inner margin. Palm elongated, 0.7 times as wide as long; dorsal margin of palm distinctly carinate, smooth; inner, outer basal lobe granular, ventral margin with milled carina, continuing into pollex (Fig. 21 E). Cutting edges of fingers meeting only distally. Ambulatory legs slender, short; meri ventrally concave, meri 1 – 3 with 2 granular rows on ventral, dorsal surface, merus of fourth pair with single row dorsally, 2 rows ventrally. Carpi of ambulatory legs dorsally carinate, propodi dorsally, ventrally carinate. Dactyli as long as propodi, lanceolate. Male abdominal cavity deep, elongated, nearly reaching buccal cavity; lateral margin with distinct ridge fitting into suture between abdominal somites. Male abdomen with somite 2 minute, spindle-shaped; somites 3 – 5 fused; somite 6 large, rounded, narrowing distally, minute median denticle; telson laciniate (Fig. 21 F, G). Female abdomen with somite 2 yoke-shaped, somites 3 – 6 fused, greatly enlarged, shield-like; telson laciniate (Fig. 22 C, D). G 1 shaft somewhat sinuous, tip slightly swollen, setose. G 2 short, curved, apex scoop-like. Vulvae prominent, large, anteriorly reaching suture 5 / 4; ovoid; interiorly hooded by calcified semi-dome, opening, directed laterally, slit-like, crescentic, bordered by shallow groove (Fig. 22 E). Colour in life. Dorsal surface of carapace and chelipeds bone-coloured, densely set with orange-red markings; finger tips white. Dorsal surface of meri, carpi and propodi of ambulatory legs with orange blotches (Fig. 5 F). Examination of the ethanol-preserved specimen revealed that the pterygostomian region and third maxillipeds are suffused with pale orange; in male, the sternites and abdomen are pale ivory except for 2 round orange markings on the third sternite and telson; female abdomen with 2 orange patches submedially near distal margin, telson orange.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E32E4C359699FE0388AC8B38.taxon	discussion	Remarks. The new species is superficially similar to U. purarensis (Ovaere, 1987), also described from Madang, Papua New Guinea. The holotype female (and only previously known specimen) was re-examined, and it differs in several key aspects. The merus of the third maxilliped of U. purarensis has a tubercle on the inner distal angle of the ischium (Fig. 23 A) (merus proportionately longer and the distal margin of the ischium unarmed in U. flammea sp. nov., Fig. 23 C). The ambulatory legs of U. purarensis, notably the meri, are relatively shorter and stouter (Fig. 23 B) than in to U. flammea (Fig. 23 D). The merus of the cheliped and chela of U. purarensis (Figs. 19 E, 20 E, 22 H) are also proportionately shorter than those of U. flammea sp. nov. (Figs. 20 A, 21 A, E, 22 B, F, G), even when females of similar sizes are compared; and the posterior margin of the merus has two diverging rows of granules (Fig. 19 A) (single row of more lozenge-shaped granules in U. flammea sp. nov., Figs. 21 A, 22 A). In addition, the frontal region of U. purarensis is less produced (Figs. 19 A, 20 A) (more projected in U. flammea sp. nov., Fig. 21 A), the posterolateral margin of the pterygostomian region adjacent to the buccal cavity is angled (Fig. 19 D) (more rounded in U. flammea sp. nov., Fig. 21 D), the posterior carapace margin is proportionately broader in U. purarensis (Figs. 19 A, 20 A) (narrower in U. flammea sp. nov., Figs. 21 A, 22 A), the distal part of the fused male abdominal somites 3 – 5 is noticeably convergent (Fig. 20 G) (less convergent and relatively broader in U. flammea sp. nov., Fig. 21 F), and the G 1 is more sinuous with the distal part distinctly dilated (Fig. 23 I – L) (less sinuous with the distal part not dilated in U. flammea sp. nov., Fig. 23 E – G).	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
8870B305E32E4C359699FE0388AC8B38.taxon	etymology	Etymology. From the Latin, flamma for “ flame ”, alluding to the flame-coloured markings of the carapace and chelipeds. Geographical distribution. This species is known only from type location, Papua New Guinea.	en	Galil, Bella S., Ng, Peter K. L. (2015): Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa 4027 (4): 451-486, DOI: 10.11646/zootaxa.4027.4.1
