identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
EDDDF3A648CA54AABF1EF10CC8EDACD8.text	EDDDF3A648CA54AABF1EF10CC8EDACD8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Attaphila aptera Bolivar 1905	<div><p>4.1. Attaphila aptera Bolivar, 1905</p><p>Figs 1A-C, 2B, E, F, 3A, B, 5A, B, J, 6A-E, 12A, 13C, 14A, I, 15F-H, 17C, D, G, 18A, B, 19A, 22A, B, 24A, B, 28H, S1A, B, S2A-C, S3A</p><p>Attaphila aptera Bolívar, 1905: 137; Princis 1963: 111.</p><p>Material studied.</p><p>Type material. Holotype, 1L ♂, Columbien, Esperanza, Dibulla, b. Atta 8-spinosa Reiche (Forel!) (completely on two slides: Bo 1232) (M. Maastricht). - Other material. Panama, Gamboa, IV-VI.2009, leg. V. Nehring, in nest of Acromyrmex octospinosus: 1♂ (completely on one slide: Bo 1224), 1♀ (completely on one slide: Bo 1225) (M. Maastricht, NHMM 2021 001, 002); 1♂ (completely on one slide: Bo 1227), 1♂ (abdomen on slide: Bo 1256, remains for DNA), 1♀ (abdomen on one slide: Bo 1257, remains for DNA), 1L ♂ (head on one slide: Bo 1457), 2L ♂ (each completely on one slide: Bo 1242, Bo 1291), 1L ♂ (head and abdomen on two slides: Bo 1292), 3L ♀ (each completely on one slide: Bo 1243, Bo 1289, Bo 1290) (M. Dresden) . - Panama, Gamboa, IV-VI.2009, leg. V. Nehring, in nest of Acromyrmex echinatior: 1♂ (abdomen, tegmina, and head on three slides: Bo 1252), 1♀ (abdomen and head on two slides: Bo 1253) (ZS Munich) .</p><p>Diagnosis.</p><p>Male Well characterised by several features unique in Attaphila: the specialisation on T1 involving moderately long bristles (msp1), the bristle distribution on T6,7 with numerous bristles on the surface but none along the posterior border, and the specially shaped virga (with a moderate sinusoidal curvature). Female: Surface bristles of T2-5 dispersed, of T6 numerous, relatively long and strong. Unique among Attaphila species by presence of a lateral gonangulum sclerite (unknown for A. sexdentis). From A. bergi distinguished by S7 having a median gap in the transversal ridge. A. sexdentis also has numerous, but smaller bristles on T6 and a differently shaped S7.</p><p>Description.</p><p>Size: Length of body (in alcohol): male 2.81-3.31 mm, female 3.16-3.36 mm. Surface bristles of tergites 2-5 (definition in 2.4.) dispersed, not arranged in transversal rows (Fig. 6A, C, D). Transversal ridges tr2-5 without distinct excurvations (Fig. 6A, D, compare grey arrows in Fig. 11A, and in Fig. 9B for male tr2).</p><p>Male. Tegmina (Fig. 3A, B) widest at about 2/3 of length; overall shape roughly triangular; posterior border convex from tegmen base (fwar) onward, its wide curvature very uniform throughout, joining the apical border far apically in a much tighter curvature (also tighter than in A. bergi); apical border transversal, fairly straight; surface bristles moderately strong. Hindwings fairly rhombic, with obtuse apex (similar to Fig. 1H). Glandular pores on T1-5 in the area between the transversal ridge and the anterior border, numerous and tiny, especially densely arranged in median third of tergite (Fig. 5A, B). Tergite 1 without long bristles on surface and borders (in contrast to long bristles on T2-5); medially, immediately posteriorly of the transversal ridge tr, with a specialisation (msp1) consisting of a more or less rounded, weakly sclerotised area with two groups of medium-sized bristles pointing anteriorly; on both sides of this area with some much smaller bristles of varying orientation (Figs 5A, B, 6A). Ridge tr1 almost continuous across midline, but more or less weakened and fragmented. Tergite 2 without specialisations. Tergites 6,7: (Figs 13C, 14I, 15F, H (larval)) Median lobe of T7 (definition in 3.6.) very short, but distinct; transversal ridges tr6 and tr7 well developed; posterior borders of T6 and T7 without bristles, surfaces with dispersed bristles of moderate to small size, on T6 larger than on T7, in radiating orientation. Subgenital lobe: (Fig. 24A, B) distal part with a deep excavation only along left side; lobe posterior to level of excavation widely tongue-shaped; with two styli, the larger left one (sll) conical, inserted at base of excavation, not reaching tip of lobe, the much smaller right one (slr) knob-like, situated subterminally on right flank of tip; stylus and borders of distal lobe with numerous bristles, most of them of moderate size, some rather long and strong. Phallomeres: (Fig. 24A, B) Sclerotised part of hook (h) from the long, wide base (b) very gradually narrowing into a slender, fairly short neck (n; shorter than in A. fungicola) which is hardly curved and bends almost rectangularly into the slightly wider claw part (cl). Endophallic apodeme (ea) not narrowed at base (forking site of sclerite). Relative to the axis of the endophallic apodeme (ea), the virga (vi) shows a distinct but quite shallow sinusoidal excurvation to the left, its apical part being moderately curved (back into the ea -axis) and gradually narrowed to a rather stout tip; virga longitudinally grooved. Paraprocts: Right paraproct with a sclerotised hook-like projection (hmp in Figs 5J, 6B), left one without a projection.</p><p>Female. Tergites 6,7: (Figs 6E, 12A, 14A, 15G (larval)) Median lobe of T7 rather long; transversal ridges tr6 and tr7 complete, tr6 sublaterally at most with a very weak bend; surface bristles (definition in 2.4.) numerous and rather long and strong; focused to central (= longitudinal and transversal middle) part of surface area (between transversal ridge and posterior border). Subgenital plate (Figs 17C, D, 18A, B) rounded-rectangular, with parallel lateral borders; lateral parts of transversal ridge (sr7-l) in the middle with a shallow, but distinct mesally directed curvation, lateral terminal parts fairly straight; ridge mesally ending shortly after having reached a transversal orientation, with a very wide median gap, partly recurved sr7-m ends indicating the ridge to be at least slightly bisinuate. Genitalia: Spermathecal plate sp large (reaching far to the left), pouch gcp moderately sized (Fig. 19A). In laterosternal shelf sclerite (ls in Fig. 22A, B) central part (c) moderately long and posteriorly more or less transversally cut, arms (a) moderately wide, wing parts (w) moderately to very wide, their base restricted to posterior half of tubes (blue arrowhead); tubes (lst) slightly curved mesad and slightly narrowed towards the anterior. Lateral and mesal gonangulum sclerites distinct (gg-l, gg-m in Fig. 19A).</p><p>Host species.</p><p>Acromyrmex octospinosus (Reich, 1793) ( Bolívar 1905), Acr. echinatior (Forel, 1899).</p><p>Distribution.</p><p>Panama (Gamboa); Colombia (Dibulla).</p><p>Remarks.</p><p>Attaphila aptera was described on the basis of a single specimen, a last-stage larval male which Bolívar (1905) interpreted as an adult; the corresponding adult stages were hitherto not known and their identification is problematic due to the scarcity of species-specific characters in larvae. A study of larval characters in four species (see 3.13.) has shown that the bristle patterns of tergites may contribute to an identification of the corresponding imagines: the bristle pattern of male (and female) larvae is very similar to that of the imaginal female.</p><p>The type specimen of Attaphila aptera has dispersed bristles on T2-5, thus ruling out the specimens herein classified as A. multisetosa, A. paucisetosa, and A. sinuosocarinata as its conspecifics. The bristles on the surface of T6,7 are numerous and relatively long and strong (Fig. 15H), comparable to those observed in larvae and females of A. bergi (Figs 14B, 15I, J) and of the specimens from Gamboa (Panama) found in the nests of Acromyrmex octospinosus and Acr. echinatior (Figs 14A, 15F, G). There is no similarity with the bristle pattern of the remaining four species, which, therefore, can also be eliminated from the list of candidates for conspecifity with the A. aptera type: A. sexdentis has considerably smaller bristles in much higher numbers and density, A. fungicola and A. schuppi have fewer and/or smaller bristles on T6,7. A. flava is only known from the male sex, but the high similarity with the male of A. fungicola justifies the assumption of a corresponding similarity between the females of the two species.</p><p>The long distance between the localities of the larval A. aptera type (Colombia) and the localities where A. bergi (Argentina, Uruguay) was found, and the different host species, Acr. octospinosus versus Acr. lundii (which are phylogenetically disjunct: Cristiano et al. 2020; Fig. 27), argue against the assumption that both could belong to the same species. The A. aptera type is much more likely conspecific with (or, at least, most closely related to) the species from Panama: their localities are much closer together and the host species reported for the A. aptera type is among the two host species known for the species from Panama. We did not find a character contradicting the assumption of conspecifity.</p><p>Because of the still existing uncertainties it may be seen as premature to assign the specimens from Gamboa to A. aptera . But the possible alternative, to describe them as a new species, appears to be less appropriate. If some day it turns out that the assignment to A. aptera is wrong, the species from Gamboa has to get a new name; if the alternative fails, the system is enriched by a new synonym.</p></div>	https://treatment.plazi.org/id/EDDDF3A648CA54AABF1EF10CC8EDACD8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter	Bohn, Horst, Nehring, Volker, G., Jonathan Rodriguez, Klass, Klaus-Dieter (2021): Revision of the genus Attaphila (Blattodea: Blaberoidea), myrmecophiles living in the mushroom gardens of leaf-cutting ants. Arthropod Systematics & amp; Phylogeny 79: 205-280, DOI: http://dx.doi.org/10.3897/asp.79.e67569, URL: http://dx.doi.org/10.3897/asp.79.e67569
4A668C721AD95502974FA7553955584A.text	4A668C721AD95502974FA7553955584A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Attaphila bergi Bolivar 1901	<div><p>4.2. Attaphila bergi Bolivar, 1901</p><p>Figs 1E, G, 2G, 3C, D, 5C, D, 7A-E, 12B, 13D, 14B, 15A, I, J, 16A, B, 17H, 18C, D, 19B, 22C, D, 24C-G, S1C, S3B-D</p><p>Attaphila bergi Bolívar, 1901: 335, pl. 6; Princis 1963: 111.</p><p>Attaphila bergi var. minor Bruch, 1916: 329, fig. 17A-F.</p><p>Material.</p><p>Type material (presumably lost). Syntypes, 2♂, 6♀, Argentina (Buenos Aires?), Uruguay. - Material studied. 2♀, [Argentina], B[ueno]s. A[ire]s (abdomen of each on one slide: Bo 1284, Bo 1440); 4L ♀, Argentina, [Prov.] B[ueno]s Aires, Castelar, M. Viana and R.Maniglia, # 41175 (together with a worker of Acromyrmex lundii on one pin); 4L ♀, same data as preceding (together with a worker of Acromyrmex lundii on one pin) (abdomen of 1L ♀ on one slide: Bo 1285) ; 3♂, 1♀, [Argentina], [ Prov.] Entre Ríos, Paranacito, [1931, acc. to loan form], Daguerre, # 28 994 (together with a worker of Acromyrmex lundii on one pin) (1♂ completely on two slides: Bo 1286; abdomen and tegmina of 1♂ on two slides: Bo 1443; abdomen of ♀ on one slide: Bo 1275) (M. Buenos Aires) . - 1♂, 1♀, [Argentina, Prov. Buenos Aires], Moreno BA, 30.XI.[19]38, M.D.Jurado, # MACN-En 7781/7780 (♂ completely on two slides: Bo 1283; abdomen and part of legs of the ♀ on one slide: Bo 1282) (M. Maastricht, NHMM 007, 008); 10L, Argentina, [Prov. Buenos Aires], La Plata, VII.1918, C.Bruch (on 4 pins, each with a worker of Acromyrmex lundii) (1L ♂ completely on two slides: Bo 1230; 1L ♀ completely on two slides: Bo 1231) (M. Maastricht) . - 1♂ [Argentina], [ Prov.] Entre Ríos, Paranacito, [1931, acc. to loan form], Daguerre, # 28 944 (from a pin with a worker of Acromyrmex lundii) (completely on two slides: Bo 1274) ; 1♀, [Argentina, Prov. Santa Fé], Rosario, Coll. Hubrich, # 15 A 16 (completely on two slides: Bo 1239); 6L ♀, [Argentina], Rosario (?), Coll. Hubrich, # R.66, B.p.77 (together with a worker of Acromyrmex lundii on one cardboard) (1L ♀ completely on one slide: Bo 1238) (ZS Munich) .</p><p>Diagnosis.</p><p>Male Well characterised by several features unique in Attaphila: the rather inconspicuous specialisation anteromedially on T1 with a pair of areas showing strongly developed microreticulation (msa1), the bristle distribution on T6,7 with some bristles on the surface of T6 and long bristles along the posterior borders, and the specially shaped virga (with a very strong sinusoidal curvature). With A. aptera it shares the absence of long bristles on surface and borders of T1. Female: Distinguished from all other species by S7 having a continuous, uninterrupted transversal ridge.</p><p>Description.</p><p>Size Length of body (dried, type specimens after Bolívar 1901): male 2.8 mm, female 2.8 mm. Surface bristles of tergites 2-5 arranged in about two irregular transversal rows (Fig. 7A, D). Transversal ridges tr2-5 without distinct excurvations (Fig. 7A, D, compare grey arrows in Fig. 11A, and in Fig. 9B for male tr2).</p><p>Male Tegmina (Fig. 3C, D) widest at about 2/3 of length; overall shape more rectangular than triangular; posterior border straight or slightly concave near tegmen base (fwar), slightly converging with the apical border, soon bending into a rather tight curvation (as compared to A. aptera), finally joining the apical border in a still tighter curvation (though less tight than in A. aptera); apical border transversal, fairly straight; surface bristles strong (stronger than in A. aptera). Hindwings lanceolate, with narrowly rounded apex (Fig. 1G). Glandular pores only on T1, in the area of the specialisation and some more around (Fig. 5C, D). Tergite 1 without long bristles on surface and borders (in contrast to long bristles on T2-5); medially, about at the level of the medially obsolete ridge tr1 (far anteriorly on T1-a), with an inconspicuous specialisation (msa1) consisting of a pair of small areas showing a net-like pattern produced by delicate furrows or ridges along which tiny glandular pores are sporadically arranged; area in between either unstructured or with dispersed small pores, occasionally also with few small bristles (Fig. 5C, D). Ridge tr1 interrupted by the specialisation. Tergite 2 without specialisations. Tergites 6,7: (Figs 7B, 13D, 15A, I (larval)) Median lobe of T7 very short, but distinct; transversal ridge tr6 well developed, tr7 rather weakly developed, perhaps with gaps in lateral parts; posterior border of T6 and T7 with a row of rather long and strong bristles, almost reaching the size of those of the lateral borders; surface of T6 in the posterior half with dispersed bristles of slightly smaller size, sometimes distributed in two groups, of T7 only with few isolated and usually much smaller bristles. Subgenital lobe: (Fig. 24C, D) distal part with a deep excavation only along left side; lobe posterior to level of excavation widely tongue-shaped; with two styli, the larger left one (sll) conical, inserted at base of excavation, not reaching tip of lobe, the much smaller right one (slr, relatively larger than in A. aptera) knob-like, situated subterminally on right flank of tip; stylus and borders of distal lobe with numerous bristles, most of them large (on average longer and stronger than in A. aptera), some rather long and strong. Phallomeres: (Fig. 24D-G) Sclerotised part of hook (h) from the long, wide base (b) very gradually narrowing into a slender, very short neck (n; shorter than in A. aptera) which is evenly curved into the slightly wider claw part (cl; neck thus hardly set off from both the base and the claw part). Endophallic apodeme (ea) not narrowed at base (forking site of sclerite). Relative to the axis of the endophallic apodeme (ea), the virga (vi) shows a distinct, deep sinusoidal curvature to the left, the apical part being very strongly and tightly curved (back into the ea -axis), and gradually narrowed to a rather acute tip ending shortly after the termination of the recurvation, hence hook-like (presence of a groove on the virga unclear). Paraprocts: Both lacking a sclerotised projection (Fig. 7C).</p><p>Female Tergites 6,7: (Figs 7E, 12B, 14B, 15J (larval)) Median lobe of T7 rather long; transversal ridges tr6 and tr7 complete, tr6 sublaterally with a distinct bend; surface bristles numerous and rather long and strong (in length comparable to those of A. aptera, but less strong), strongly focused to central part of surface area. Subgenital plate (Figs 16A, B, 18C, D) rounded-rectangular, with parallel lateral borders; lateral parts of transversal ridge (sr7-l) almost straight; median part (sr7-m) complete, slightly (Fig. 16A) to strongly (Fig. 16B) bisinuate. Genitalia: Spermathecal plate sp rather small, pouch gcp rather small (Fig. 19B). In laterosternal shelf sclerite (Fig. 22C, D) central part (c) moderately long and posteriorly transversally cut, arms (a) very narrow (narrower than in A. aptera), wing parts (w) moderately wide, their base reaching far into anterior half of tubes (blue arrowhead); tubes (lst) straight, of fairly uniform width throughout, anterior end widely rounded. Mesal gonangulum sclerites (gg-m) distinct, lateral ones absent (Fig. 19B).</p><p>Host species.</p><p>Acromyrmex lundii ( Guérin-Méneville, 1838); host species of var. Acromyrmex lundii minor according to Bruch (1916) Acr. lobicornis (Emery, 1888), and Amoimyrmex silvestrii (Emery, 1905).</p><p>Distribution.</p><p>Argentina: Prov. Buenos Aires (Castelar, La Plata, Moreno), Prov. Entre Rios (Paranacito), Prov. Santa Fé (Rosario). The Attaphila bergi var. minor is reported from the Provinces San Luis and Catamarca; the report of A. bergi from Córdoba (Bruch 1929) most likely is a mistake and refers to the occurrence of Attaphila bergi var. minor in San Luis, not mentioned in this paper; Uruguay.</p><p>Remarks.</p><p>The loss of the type specimens of A. bergi and the insufficient original description of the species prevent an unequivocal identification. But for several reasons the determination of the available Argentine specimens as A. bergi is most probably correct: They have the same host ant species, Acromyrmex lundii; no other species of Attaphila has so far been found in nests of this ant. The specimens studied are from at least 5 different localities in rather close distance, not very far from the localities of the type specimens (Argentina: Buenos Aires?, Uruguay), without showing remarkable differences among each other.</p><p>Bruch (1916) described a Acromyrmex silvestrii var. minor occurring in nests of Amoimyrmex silvestrii (Prov. San Luis) and Acromyrmex lobicornis (Prov. Catamarca; belonging to Amoimyrmex or Acromyrmex clade not yet phylogenetically tested), of smaller size and paler colour than the nominate form. This characterisation and the missing of males raise the suspicion that the type specimens were larvae, possibly of normal A. bergi . On the other hand, the complete or partial association with Amoimyrmex hosts, which form the sister taxon of Acromyrmex + Atta and are thus phylogenetically far remote from Acromyrmex lundii (Cristiano et al. 2020) raises doubt on Acromyrmex lundii var. minor being conspecific with A. bergi . The type specimens of Acromyrmex bergi var. minor, formerly deposited in the Museo de La Plata, are no longer present there and presumably lost.</p></div>	https://treatment.plazi.org/id/4A668C721AD95502974FA7553955584A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter	Bohn, Horst, Nehring, Volker, G., Jonathan Rodriguez, Klass, Klaus-Dieter (2021): Revision of the genus Attaphila (Blattodea: Blaberoidea), myrmecophiles living in the mushroom gardens of leaf-cutting ants. Arthropod Systematics & amp; Phylogeny 79: 205-280, DOI: http://dx.doi.org/10.3897/asp.79.e67569, URL: http://dx.doi.org/10.3897/asp.79.e67569
3F54CA4203F85F8B9B2103DD37006B60.text	3F54CA4203F85F8B9B2103DD37006B60.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Attaphila flava Gurney 1937	<div><p>4.3. Attaphila flava Gurney, 1937</p><p>Figs 3E, F, 5I, 8D, 15C, 25C, 26A</p><p>Attaphila flava Gurney, 1937: 106, fig. 9; Princis 1963: 111.</p><p>Material studied.</p><p>Holotype, 1♂, British Honduras, Belize, Botanic Gardens, 11.VII.1904, P.G.Goll Type No. 52014 U.S. N.M. (abdomen and legs on two slides: Bo 1280) (M. Washington).</p><p>Diagnosis.</p><p>Male The type specimen of A. flava is very similar to the male of A. fungicola and only shows slight differences in few characters: tegmina with apical border slightly convex, less oblique, surface bristles less strong; hook of left phallomere from the wide sclerotised base more gradually narrowing into a shorter neck, endophallic apodeme near its posterior base less strongly narrowed; and it has a different, though unknown host species.</p><p>Description.</p><p>Size Length of body (dried): male 2.83 mm. Surface bristles of tergites 2-5 dispersed, not arranged in transversal rows (Fig. 8D). Transversal ridges tr2-5 without distinct small excurvations to the anterior (Fig. 8D; compare grey arrows in Fig. 11A), but male tr2 posteriorly of the specialisations with a wide excurvation to the posterior, mesally followed by a wide, very shallow excurvation to the anterior (grey arrows in Fig. 8D).</p><p>Male Tegmina (Fig. 3E, F) widest at about 1/2 of length; slightly obtusely wedge-shaped due to the converging course of the basalmost part of the posterior border and the apical border, both connected in a rather wide curvature; posterior border with a slight concavity near the tegmen base, oblique apical border throughout slightly convex; surface bristles fine. Hindwings fairly rhombic, with obtuse apex. Glandular pores on T2-5 (and perhaps T1) in the area between the transversal ridge and the anterior border, dispersed, in moderately high density (Fig. 5I). Tergite 1 without specialisations (occurrence of long bristles could not be studied for T1; they are present on T2-5). Tergite 2 along anterior border with a pair of specialisations (msl2) consisting of a shallow, narrow transversal trough the bottom of which is patterned by low crossing ridges (Figs 5I, 8D; very similar to those of A. fungicola). Tergites 6,7: (Fig. 15C) Median lobe of T7 very short, but distinct; transversal ridge tr6 well developed, tr7 absent; posterior border of T6 and T7 with a relatively dense row of very short and thin bristles much smaller than those of the respective lateral borders; surface of T6 and T7 only with few isolated and very small bristles, still smaller than those of the posterior borders. Subgenital lobe: (Figs 25C, 26A) distal part with a deep excavation each along left side and right side; lobe posterior to level of excavations narrowly tongue-shaped and presumably inclined leftward (tip of lobe not retained in specimen); only left stylus present (sll), which is quite conical, inserted at base of left excavation; at the base of the right excavation with a group of short and strong bristles; stylus with few rather long and thin bristles. Phallomeres: (Figs 25C, 26A) Sclerotised part of hook (h) from the short, wide base (b) gradually narrowing into a slender neck (n; shorter than in A. fungicola) with a hardly curved proximal part, terminating in a slightly wider claw part (cl). Endophallic apodeme (ea) slightly narrowed at base (forking site of sclerite). Relative to the axis of the endophallic apodeme (ea), the virga (vi) shows a weak, very shallow sinusoidal curvature to the left, its apical part being hardly curved (back into the ea -axis) and gradually narrowed to a rather acute tip, distinctly more acute than in A. paucisetosa; virga likely longitudinally grooved. Paraprocts: Both lacking a sclerotised projection.</p><p>Female Unknown.</p><p>Distribution.</p><p>Only known from the type locality Belize, Belize City.</p><p>Host species.</p><p>Unknown, certainly not Atta texana as in A. fungicola since the distribution of this ant species does not reach further south than northeast Mexico. In view of the great similarity of A. flava with A. fungicola, its host is likely an Atta species. It could be Atta cephalotes, the only species of Atta, Acromyrmex, and Amoimyrmex so far reported for Belize (according to https://antwiki.org and https://www.antweb.org, both accessed on 19.i.2021), although Acromyrmex echinatior is known from all neighbouring countries (https://www.antweb.org).</p><p>Remarks.</p><p>In the description of A. flava Gurney (1937) only noted one difference to A. fungicola, the very short subgenital plate (his fig. 9). But the subgenital plate of the type specimen is damaged and missing its apical part (Figs 25C, arrow in 26A); it might have been of the same length and shape as in A. fungicola (Fig. 26B). In view of the weak differences between the males of A. flava and A. fungicola one might have doubts whether the type specimen of A. flava really represents a separate species. But as long as neither the corresponding female nor the full shape of the subgenital plate is known, synonymisation with A. fungicola appears unjustified.</p></div>	https://treatment.plazi.org/id/3F54CA4203F85F8B9B2103DD37006B60	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter	Bohn, Horst, Nehring, Volker, G., Jonathan Rodriguez, Klass, Klaus-Dieter (2021): Revision of the genus Attaphila (Blattodea: Blaberoidea), myrmecophiles living in the mushroom gardens of leaf-cutting ants. Arthropod Systematics & amp; Phylogeny 79: 205-280, DOI: http://dx.doi.org/10.3897/asp.79.e67569, URL: http://dx.doi.org/10.3897/asp.79.e67569
90567B49330A5CAFBEFA87D6C9B05DA7.text	90567B49330A5CAFBEFA87D6C9B05DA7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Attaphila fungicola Wheeler 1900	<div><p>4.4. Attaphila fungicola Wheeler, 1900</p><p>Figs 1D, 3G, H, 5G, H, 8A-C, 12C, 13F, 14C, 15B, 16C, D, 18E, F, 19C, D, 22E, F, 25D, E, 26B</p><p>Attaphila fungicola Wheeler, 1900: 860, figs 3-6; Hebard 1916: 214, pl. X, figs 5-6;? Wheeler 1928: 255 (reports from British Guiana and Panama); Princis 1963: 110;? Brossut 1976: 167, figs 1-9; Roth 1968: 135, fig. 17; Roth 1971: 130, fig. 21.</p><p>Material studied.</p><p>Type material. Lectotype, 1♂, Texas, Austin, XI.00, Atta fervens with Attaphila fungicola, Nov. 20.1900, W.M.Wheeler Coll. (left tegmen and abdomen on two slides: Bo 1265) (M. New York) . - Paralectotypes: 7L, same data as Lectotype (M. New York); 2L, Texas, Austin, 4/1900, Wheeler!, b. Atta fervens (M. Maastricht) . - Other material. 1♂, 1♀, Texas, Milam Co., Sugarloaf Mt., 300', 4 mi. N Gause, 19.IX.1992, Godwin, Quinn, Riley et al. (each completely on two slides: Bo 1229, Bo 1228) (M. Maastricht, NHMM 2021 004, 005) . - 4♀, Texas, Freestone Co., Old Spring Seat Church, nr. Donie, pit-fall in Atta nest, 26.-31.V.[19]95, Wm.Godwin and E.Riley ; 1♀, 9L, Texas, Milam Co., Sugarloaf Mt., 300', 4 mi. N Gause, 19.IX.1992, Godwin, Quinn, Riley et al. (3L ♀, each completely on one slide: Bo 1435, Bo 1436, Bo 1437); 4L, LA [Louisiana], Natchitoches Par. Red Dirt Wdlf. Mn. Ar., nr. Red Buff Campgr., 26.IX.1992, E.G. and T.J.Riley et al. (1L ♀ completely on slide: Bo 1438) ; 1♀, USA, Texas, Guadalupe <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-97.85017&amp;materialsCitation.latitude=29.48282" title="Search Plazi for locations around (long -97.85017/lat 29.48282)">Co.</a>, 14.5 km SE Seguin, 29.48282°N 97.85017°W, ± 5 m, 4.XII.2014, A. Graf, B. Hays, B. Lyons, J. Oswald, E.Riley and W. Ryan, ex nest of Atta texana from depth of 2-8 ft. (Coll. TAMU) . - 1♀, Texas, Freestone Co., Old Spring Seat Church, nr. Donie, pit-fall in Atta nest, 26.-31.V.[19]95, Wm.Godwin and E.Riley (completely on two slides: Bo 1236) ; 1♀, USA, Texas, Travis Co., Austin, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-97.78194&amp;materialsCitation.latitude=30.28444" title="Search Plazi for locations around (long -97.78194/lat 30.28444)">University</a> of Texas, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-97.78194&amp;materialsCitation.latitude=30.28444" title="Search Plazi for locations around (long -97.78194/lat 30.28444)">Brackenridge Field Lab.</a>, Atta texana nest, 30.28444 N 97.78194 W, 1.VII.2010, leg. U.Mueller (abdomen on one slide: Bo 1264) (ZS Munich) .</p><p>Diagnosis.</p><p>Male As in A. paucisetosa with specialisations on T2 (msl2, but these are narrower, with simpler ridges), distinguished by having dispersed surface bristles on T2-5. For differences to A. flava, see 4.3. Female: Well characterized by the combined occurrence of two characters: T2-5 with dispersed surface bristles, T6,7 with only few and small surface bristles. The latter feature is also shared by A. paucisetosa, in which, however, the surface bristles of T2-5 are arranged in one line.</p><p>Description.</p><p>Size Length of body (dried): male 2.65-3.5 mm, female 2.45-3.5 mm (after Hebard 1916). Surface bristles of tergites 2-5 dispersed, not arranged in transversal rows (Fig. 8A, C). Transversal ridges tr2-5 without distinct small excurvations to the anterior (Fig. 8A, C; compare grey arrows in Fig. 11A), but male tr2 posteriorly of the specialisations with a wide excurvation to the posterior, mesally followed by a wide, very shallow excurvation to the anterior (grey arrows in Fig. 8A).</p><p>Male Tegmina (Fig. 3G, H) widest at about 1/2 of length; obtusely wedge-shaped due to the converging course of the basalmost part of the posterior border and the apical border, connecting curvature shorter than in A. flava thanks to the slightly more oblique course of the borders; likewise, posterior border near the tegmen base less concave, oblique apical border fairly straight; surface bristles rather fine (stronger than in A. flava). Hindwings fairly rhombic, with obtuse apex. Glandular pores on T2-5 in the area between the transversal ridge and the anterior border, dispersed, in moderately high density (Fig. 5G, H). Tergite 1 with long bristles on surface and borders (like T2-5); without specialisations. Tergite 2 along the anterior border with a pair of specialisations (msl2) consisting of a shallow, narrow transversal trough the bottom of which is patterned by low crossing ridges (Fig. 5G, H). Tergites 6,7: (Figs 8A, 13F, 15B) Median lobe of T7 scarcely visible; transversal ridge tr6 well developed, tr7 absent; posterior border of T6 and T7 with a relatively dense row of very short and thin bristles, much smaller than those of the respective lateral borders; surface of T6 and T7 only with few isolated and very small bristles, still smaller than those of the posterior borders. Subgenital lobe: (Figs 25D, 26B) distal part with a deep excavation each along left side and right side; lobe posterior to level of excavations narrowly tongue-shaped and inclined leftward; only left stylus present (sll), which is conical, inserted at base of left excavation, not reaching tip of lobe; at the base of the right excavation with a group of short and strong bristles; distal lobe and stylus with few rather long and thin bristles. Phallomeres: (Fig. 25D, E, 26B) Sclerotised part of hook (h) from the short, wide base (b) almost gradually narrowing (not as gradually as in A. flava, but with a weak, yet distinct shoulder, Fig. 25C) into a slender neck (n) with a hardly curved proximal part, terminating in a slightly wider claw part (cl). Endophallic apodeme (ea) strongly narrowed at base (forking site of sclerite). Relative to the axis of the endophallic apodeme (ea), the virga (vi) shows a weak, very shallow sinusoidal curvature to the left, its apical part being hardly curved (not fully back into the ea -axis) and gradually narrowed to a rather acute tip, distinctly more acute than in A. paucisetosa; virga likely longitudinally grooved. Paraprocts: Both lacking a sclerotised projection (Fig. 8B).</p><p>Female Tergites 6,7: (Figs 12C, 14C) Median lobe of T7 rather long; transversal ridges tr6 and tr7 complete, tr6 sublaterally at most with a very weak bend; surface bristles numerous, very few of medium size, most between very small and tiny; dispersed all over the surface between transversal ridge and posterior border. Subgenital plate (Figs 16C, D, 18E, F) rounded-rectangular, with parallel lateral borders; lateral parts of transversal ridge (sr7-l) fairly straight; median part (sr7-m) with gap(s) of various size and number: two cases with one rather wide gap, and one case with two quite narrow gaps, then indicated to be slightly bisinuate. Genitalia: Spermathecal plate (sp) moderately large, pouch gcp present (size not determinable; Fig. 19C, D). In laterosternal shelf sclerite (Figs 19C, 22E, F) central part (c) fairly short and posteriorly rounded, posterior and lateral margins of central part and arms together forming a quite evenly curved horseshoe arch, arms (a) fairly narrow, wing parts (w) moderately wide, their base restricted to posterior half of tubes (blue arrowhead); tubes (lst) with angular bend exceptionally far anteriorly (at red arrowhead: near midlength of lateral border), from there towards the anterior rather strongly narrowed and curved mesad. Mesal gonangulum sclerites (gg-m) distinct, lateral ones absent (Fig. 19D).</p><p>Host species.</p><p>Atta texana (Buckley, 1860). [The current taxonomic status (according to Bolton 2021) is that (1) Atta fervens (Drury, 1782), the name to which the sampling of Attaphila fungicola type specimens in "Material studied" above most likely refers, is a synonym of Atta cephalotes (Linnaeus, 1758), but that (2) the " Atta fervens " populations from Texas, the area of this sampling, were described as a separate species Atta texana (Buckley, 1860), which is a currently valid name.]</p><p>Distribution.</p><p>USA: Texas (Austin, Donie, Gause, Seguin), Louisiana (Natchitoches). The reports of Wheeler (1928) from British Guiana and Panama and of Brossut (1976) from Trinidad most likely concern other species.</p><p>Remarks.</p><p>In his description of the species Wheeler (1900) reported that he had collected four males, two females, and about seventy immature specimens from a nest of " Atta fervens " (i.e. Atta texana) (excavated on the 10th of April). It is not indicated which of the specimens he considered as type specimens.</p><p>Specimens labelled as types of A. fungicola are present in the Collection Wheeler, which is preserved in the American Museum of Natural History (New York). It comprises one male and seven larvae, each labelled with "Austin, Texas, W.M.Wheeler, Coll."; additionally, there is a common handwritten label: " Atta fervens with Attaphila fungicola, Austin, Nov. 20. 1900".</p><p>A second sample of pretended type specimens is deposited in the Wasman Collection in the Natuurhistorisch Museum in Maastricht, consisting of two immatures. They are labelled - in Wasmann’s handwriting - with " Attaphila fungicola Wheel. (Typen), b. Atta fervens, 4/1900 Wheeler!, Austin (Tex.)".</p><p>The agreement in the date shows that the specimens from Maastricht undoubtedly belong to the series which Wheeler described in his paper, while the New York series was collected later in the year. But the only specimen among the two series which could serve as an informative type is the sole adult specimen, the male from the New York series. Since both series were collected by Wheeler, in the same year, at the same locality, in nests of the same ant species, it appears legitimate to consider the two series together as a Syntype series, from which the male specimen is here selected as the Lectotype, while all other specimens are designated as Paralectotypes.</p></div>	https://treatment.plazi.org/id/90567B49330A5CAFBEFA87D6C9B05DA7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter	Bohn, Horst, Nehring, Volker, G., Jonathan Rodriguez, Klass, Klaus-Dieter (2021): Revision of the genus Attaphila (Blattodea: Blaberoidea), myrmecophiles living in the mushroom gardens of leaf-cutting ants. Arthropod Systematics & amp; Phylogeny 79: 205-280, DOI: http://dx.doi.org/10.3897/asp.79.e67569, URL: http://dx.doi.org/10.3897/asp.79.e67569
53C5A9FF8D925D82B10140AAF23773D1.text	53C5A9FF8D925D82B10140AAF23773D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Attaphila multisetosa sp. nov. Bohn and Klass	<div><p>4.5. Attaphila multisetosa sp. nov. Bohn and Klass</p><p>Figs 10A, B, 12D, 14D, 17E, 18G, 21A, 22G</p><p>A. aptera Bolívar, 1905 in Bruijning 1959: 23, figs 7, 8 (misidentification).</p><p>Material studied.</p><p>Holotype, 1♀, Suriname, [Distr. Wanica], Lelydorp, in Atta -nest, 20.V.1938, Geijskes (completely on two slides: Bo 1270) (M. Leiden) . - Paratype, 1L ♀, same data as holotype (terminalia on one slide Bo 1449) (M. Leiden) .</p><p>Diagnosis.</p><p>Female Arrangement of bristles on T2-5 similar to A. paucisetosa and A. sinuosocarinata, but much more irregular, only partly in one line; from the former distinguished by having numerous and rather long and strong bristles on T6 and a larger spermathecal plate, from the latter by the course of the T6 transversal ridge sublaterally not having an angular bend, and from both by the unique structure of the middle part of the laterosternal shelf sclerite (ls).</p><p>Etymology.</p><p>The species name refers to the numerous (Latin: multi) bristles (Latin: setae) present on T6,7.</p><p>Description.</p><p>Size Length of body (dried): female 2.8 mm. Surface bristles of tergites 2-5 at least in median third of tergites arranged in one fairly regular transversal row (though almost two-rowed on T5), towards laterally distribution less regular, appearing two-rowed (Fig. 10A). Transversal ridges tr2-5 medially and in part sublaterally with a very weak small excurvation to the anterior (grey arrows in Fig. 10A; weaker than in A. schuppi, Fig. 11A).</p><p>Male Unknown.</p><p>Female Tergites 6,7: (Figs 10B, 12D, 14D) Median lobe of T7 rather long; transversal ridges tr6 and tr7 complete, tr6 sublaterally without angular bend; surface bristles numerous on T6, rather few on T7, of medium size (slightly smaller than in A. bergi), strongly focused to central part of surface area. Subgenital plate (Figs 17E, 18G) rounded-rectangular, with parallel lateral borders; lateral parts of transversal ridge (sr7-l) fairly straight; median part (sr7-m) with three small gaps, weakly bisinuate. Genitalia: Spermathecal plate (sp) large, pouch gcp large (Fig. 21A). In laterosternal shelf sclerite (Fig. 22G) central part (c) very long and posteriorly widely rounded, arms (a) moderately wide, wing parts (w) very wide, their base reaching far into anterior half of tubes (blue arrowhead; but interpretation not entirely clear); tubes (lst) with an evenly convex lateral border and a straight mesal border, narrowed towards the anterior. Mesal gonangulum sclerites (gg-m) distinct, lateral ones absent (Fig. 21A).</p><p>Host species.</p><p>Atta, species unknown. (Three Atta species known from Suriname: A. cephalotes, A. laevigata, and A. sexdens; Bolton 2021)</p><p>Distribution.</p><p>So far only known from the type locality Lelydorp, Distr. Wanica, Suriname.</p></div>	https://treatment.plazi.org/id/53C5A9FF8D925D82B10140AAF23773D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter	Bohn, Horst, Nehring, Volker, G., Jonathan Rodriguez, Klass, Klaus-Dieter (2021): Revision of the genus Attaphila (Blattodea: Blaberoidea), myrmecophiles living in the mushroom gardens of leaf-cutting ants. Arthropod Systematics & amp; Phylogeny 79: 205-280, DOI: http://dx.doi.org/10.3897/asp.79.e67569, URL: http://dx.doi.org/10.3897/asp.79.e67569
31348359E8745C70B5E357A00B36B7DB.text	31348359E8745C70B5E357A00B36B7DB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Attaphila paucisetosa sp. nov. Bohn and Klass	<div><p>4.6. Attaphila paucisetosa sp. nov. Bohn and Klass</p><p>Figs 1F, H, 2A, C, D, H, I, 3I, J, 4A-D, 5E, F, K, 9A-F, 12E, 13E, 14E, 15D, E, K, L, 16E, F, 18H, I, 20A, B, 21D, 22H, 25A, B, 26C, S2D, E</p><p>Material studied.</p><p>Type material. Holotype, 1♂, Panama, Gamboa, in nest of Atta colombica, IV.-VI.2009, leg. V. Nehring (abdomen on slide: Bo 1258) (M. Maastricht, NHMM 2021 006). - 5 Paratypes, same data as holotype: 1♀ (completely on two slides: Bo 1226) (M. Maastricht, NHMM 2021 003), 1♂ (abdomen, tegmina, and head on three slides: Bo 1441), 1♀ (abdomen on two slides: Bo 1241) (ZS Munich), 1♂ (abdomen on one slide: Bo 1254; remains for DNA), 1♀ (abdomen on one slide: Bo 1255; remains for DNA) (M. Dresden) . - Other material. Same data as holotype: 1♂ (completely on two slides: Bo 1240) , 1♂ (abdomen and head on two slides: Bo 1442) , 2♂ (each with abdomen on one slide: Bo 1444, Bo 1445), 1L ♂ (abdomen and head on two slides: Bo 1433), 1L ♂ (abdomen on one slide: Bo 1434), 1L ♂ (head on one slide: Bo 1458), 1L ♀ (abdomen on one slide: Bo 1293; remains for DNA), 2L ♀ (each with abdomen on one slide: Bo 1295, Bo 1446), 2L ♀ (each with abdomen and head on two slides: Bo 1294, Bo 1296) (M. Dresden) . - 1♀, Panama, Panamá Par. Nac. Soberiana, Pipeline Rd., km 2, at entrance to Atta [ Atta colombica] nest at night, 16.V.1993, E. Riley (completely on two slides: Bo 1235) (Coll. TAMU) . - 1♀, Panama, Gamboa, 14.VI.2010, attached to a queen of Atta cephalotes, leg. R.Adams (abdomen on one slide: Bo 1268) (ZS Munich) . - 1♂, 4♀, 3L, Colombia, Valle del Cauca, Cali, Vereda los Andes, El Cabuyal, 3°24′50.37″N 76°35′38.34″W, 1360 m, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-76.59399&amp;materialsCitation.latitude=3.4139917" title="Search Plazi for locations around (long -76.59399/lat 3.4139917)">Nido</a> de Atta cephalotes, colecta manual, 25.XI.2012, leg. Jonathan Rodríguez G. (abdomen and head of ♂ on two slides: Cb 4/1, legs of a female on slide Cb 4/2, head of a L on slide Cb 4/3) ; 1♂, 1♀, 3L, Colombia, Valle del Cauca, Cali, Vereda el Peón, Loma larga, 3°20′23.69″N 76°35′13.17″W, 1399 m, Nido de Atta cephalotes, colecta manual, 24.X.2013, leg. Jonathan Rodríguez G. (abdomen and head of ♂ and ♀ each on two slides: Cb 2/1, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-76.587&amp;materialsCitation.latitude=3.3399138" title="Search Plazi for locations around (long -76.587/lat 3.3399138)">Cb</a> 2/2) ; 1♀, Colombia, Valle del Cauca, Buenaventura, Vereda el Salto, La Vibora, 3°50′29.5″N 76°47′32.0″W, 600 m, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-76.79222&amp;materialsCitation.latitude=3.8415277" title="Search Plazi for locations around (long -76.79222/lat 3.8415277)">Nido</a> de Atta cephalotes, colecta manual, 7.V.2014, leg. Jonathan Rodríguez G. (Cb 3) (ZS Munich) .</p><p>Diagnosis.</p><p>Male As in A. fungicola and A. flava with specialisations on T2 (but these are wider, with more complicated ridges), distinguished by the arrangement of the surface bristles of T2-5 in one transversal line. Female: Well characterized by the combined occurrence of two features: bristles of T2-5 arranged in one strict transversal line, and T6,7 with only few and rather small bristles. The latter feature is also shared by A. fungicola, which, however, has dispersed surface bristles on T2-5. The species A. sinuosocarinata and A. multisetosa resemble A. paucisetosa in the first feature, but are distinguished by having numerous surface bristles on T6,7 and by differences in the laterosternal shelf area.</p><p>Etymology.</p><p>The species name refers to the few (Latin: pauci) bristles (Latin: setae) being present on T6,7.</p><p>Description.</p><p>Size Length of body (in alcohol): male 2.7-3.43 mm, female 2.61-3.49 mm. Surface bristles of tergites 2-5 strictly arranged in one straight transversal row, only on T2 near the lateral borders bristles often less regularly distributed (Fig. 9A, B, E). Transversal ridges tr2-5 without distinct small excurvations to the anterior (Fig. 9B, E; compare grey arrows in Fig. 11A), but male tr2 posteriorly of the specialisations with a wide excurvation to the posterior, mesally followed by a wide, very shallow excurvation to the anterior (grey arrows in Fig. 9B).</p><p>Male Tegmina (Fig. 3I, J) widest at about 1/2 of length; obtusely wedge-shaped due to the roughly converging course of the basalmost part of the posterior border and the apical border, connecting curvature similarly short as in A. fungicola; basalmost part of posterior border and the oblique apical border slightly concave; surface bristles rather fine (as in A. flava but longer and at a lower density). Hindwings fairly rhombic, with obtuse apex (Fig. 1H). Glandular pores on T1-5 in the area between the transversal ridge and the anterior border, dispersed, in moderately high density (Fig. 5E, F). Tergite 1 with long bristles on surface and borders (like T2-5); without specialisations. Tergite 2 along the anterior border with a pair of specialisations (msl2) consisting of a shallow, fairly wide transversal trough the bottom of which is patterned by low crossing ridges that are more complicated than in A. flava and A. fungicola, forming a net-like relief (Figs 5E, F, 9B). Tergites 6,7: (Figs 9C, 13E, 15D, E, K (larval)) Median lobe of T7 very short, but distinct; transversal ridge tr6 well developed, tr7 absent; posterior border of T6 and T7 with a row of relatively long and thin bristles, distinctly shorter and thinner than those at the respective lateral borders; surface of T6 and T7 only with few isolated and small bristles, still smaller than those at the posterior borders. Subgenital lobe: (Figs 25A, B, 26C) distal part with a deep excavation each along left side and right side; lobe posterior to level of excavations narrowly tongue-shaped and inclined leftward; only left stylus present (sll), which is conical, inserted at base of excavation, not reaching tip of lobe; at the base of the right excavation with a group of short and strong bristles; distal lobe and stylus with few rather long and thin bristles. Phallomeres: (Figs 25A, B, 26C) Sclerotised part of hook (h) from the short, wide base (b) suddenly narrowing into a slender neck (n) widely and uniformly curved from the beginning, terminating in a slightly wider claw part (cl). Endophallic apodeme (ea) not narrowed at base (forking site of sclerite). Relative to the axis of the endophallic apodeme (ea), the virga (vi) shows a weak, angular proximal bend to the left, its apical part being hardly curved (not fully back into the ea -axis) and gradually narrowed to a fairly stout tip; virga likely longitudinally grooved. Paraprocts: Both lacking a sclerotised projection (Figs 5K, 9D).</p><p>Female Tergites 6,7: (Figs 9F, 12E, 14E, 15L (larval)) Median lobe of T7 rather long; transversal ridges tr6 and tr7 complete, tr6 sublaterally with a weak, but distinct bend; surface bristles very few, of medium size or smaller; focused to central part of surface area. Subgenital plate (Figs 16E, F, 18H, I) rounded-rectangular, with parallel lateral borders; lateral parts of transversal ridge (sr7-l) with a weak mesally directed curvation, lateral terminal parts with a weak laterally directed curvation; ridge mesally ending well after having reached a transversal orientation, close to a bristle-bearing point upon it, with a moderately wide median gap, recurved sr7-m ends indicating the ridge to be bisinuate. Genitalia: (Figs 20A, B, 22H) Spermathecal plate sp small, pouch gcp fairly small (Fig. 20A, B). In laterosternal shelf sclerite (Figs 20A, 22H) central part (c) fairly long and posteriorly rounded, posterior and lateral margins of central part and arms together forming a quite evenly curved horseshoe arch, arms (a) fairly wide, wing parts (w) fairly narrow, their base restricted to posterior half of tubes (blue arrowhead); tubes (lst) narrowed towards the anterior and curved mesad. Mesal gonangulum sclerites (gg-m) distinct, lateral ones absent (Fig. 20B).</p><p>Host species.</p><p>Atta colombica Guérin-Méneville, 1844, Atta cephalotes (Linnaeaus, 1758).</p><p>Distribution.</p><p>Panama (Gamboa); Colombia: Depart. Valle del Cauca (Cali, Buenaventura).</p></div>	https://treatment.plazi.org/id/31348359E8745C70B5E357A00B36B7DB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter	Bohn, Horst, Nehring, Volker, G., Jonathan Rodriguez, Klass, Klaus-Dieter (2021): Revision of the genus Attaphila (Blattodea: Blaberoidea), myrmecophiles living in the mushroom gardens of leaf-cutting ants. Arthropod Systematics & amp; Phylogeny 79: 205-280, DOI: http://dx.doi.org/10.3897/asp.79.e67569, URL: http://dx.doi.org/10.3897/asp.79.e67569
BA4D3E16229A56FA8322212D7AC846F6.text	BA4D3E16229A56FA8322212D7AC846F6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Attaphila schuppi Bolivar 1905	<div><p>4.7. Attaphila schuppi Bolivar, 1905</p><p>Figs 11A, B, 12F, 14F, 16G, H, 18J, K, 20C, D, 22J</p><p>Attaphila schuppi Bolívar, 1905: 138; Princis 1963: 111.</p><p>Material studied.</p><p>Type material. Syntypes, 2♀, [Brazil, Estado Rio Grande do Sul], Porto Alegre, acc. by Atta nigra Schupp (each on two slides: Bo 1234 [labelled "Lectoholotype, det. A.B.Gurney, 1971"] and Bo 1237) (M. Maastricht) .</p><p>Diagnosis.</p><p>Female Characterised by a series of unique features: Transversal ridge of T2-5 mesally and laterally with a small but distinct anterior excurvation (much weaker present also in A. multisetosa), transversal ridge absent on T7, cerci longer than wide.</p><p>Description.</p><p>Size Length of body (dried): female 3.5 mm. Surface bristles of tergites 2-5 approximately arranged in two transversal rows (Fig. 11A). Transversal ridges tr2-5 medially and sublaterally with a small but distinct excurvation to the anterior (grey arrows in Fig. 11A; much stronger than in A. multisetosa, Fig. 10A).</p><p>Male Unknown.</p><p>Female Tergites 6,7: (Figs 11B, 12F, 14F) Median lobe of T7 short, but distinct; transversal ridge tr6 well developed, tr7 missing except for few short remnants including also the two tr7 -associated bristles, tr6 sublaterally with a weak, but distinct angular bend; surface bristles numerous on T6, small to very small, focused to wider central part of surface area; absent on T7. Subgenital plate (Figs 16G, H, 18J, K) rounded-rectangular, with parallel lateral borders; posterior border rather weakly trilobed; lateral parts of transversal ridge (sr7-l) fairly straight, ridge mesally ending before having reached a transversal orientation (near anterior border of S7, and hardly curved mesad), with a very wide median gap. Genitalia: (Figs 20C, D, 22J) Spermathecal plate sp likely small (not clearly identifiable), pouch gcp likely absent (Fig. 20C, D). In laterosternal shelf sclerite (Fig. 22J) central part (c) moderately long and posteriorly transversally cut, arms (a) fairly wide, wing parts (w) fairly wide, their base reaching far into anterior half of tubes; tubes (lst) straight, of fairly uniform width throughout, anterior end widely rounded. Mesal gonangulum sclerites (gg-m) distinct, lateral ones absent (Fig. 20D).</p><p>Host species.</p><p>Acromyrmex niger (F. Smith, 1858) (as " Atta nigra Schupp" in Bolívar 1905; originally placed in Atta, since 1913 attributed to Acromyrmex, and placed deeply in Acromyrmex according to Cristiano et al. 2020).</p><p>Distribution.</p><p>Brazil: Est. Rio Grande do Sul, only known from the type locality Porto Allegre.</p><p>Remarks.</p><p>The Wasmann Collection in Maastricht keeps two female specimens (on one pin) labelled by Wasmann as follows: " Attaphila schuppi (m) Boliv., Typen"; there are no labels concerning locality and species of the ant host. Bolívar’s description contains, after a short morphological characterisation, the following data: "♂ Long. 3,5 mill. Hab. Porto Alegre. Elle se trouve en compagnie de Atta nigra Schupp. ...". The discrepancy between the two data sets raises the question whether the females from Maastricht had really been the subjects of Bolívar’s description.</p><p>The absence of a number ahead of the sex symbol does not necessarily mean that Bolívar had only one specimen for study; in the descriptions of some other new species in the same paper Bolívar never noted the number of treated specimens. The strongest doubts in considering the Maastricht specimens as type specimens of A. schuppi concern the sex determination. It is extremely unlikely that Bolívar should have confused the two sexes. In the description of A. aptera in the same paper Bolívar emphasises the exceptional case of that species having wingless males (a wrong assumption, as a larval male is concerned, see 4.1.). On the other hand, it appears unlikely that Bolívar had males in his hands when he described the species A. schuppi. The description does not contain any remarks concerning wings, and the posterior border of the last sternite is described as being "transverso trisinuato", as is typical for the subgenital plate of Attaphila females. The simplest solution for the conflicting pieces of information would be to assume an error in the printing of the sex symbol.</p><p>The last remaining issue is the incomplete labelling of the type specimens. Wasmann had got the specimens from R.P. Schupp, possibly already without a label and only with a verbal information about the collecting data, which he might have passed on to Bolívar . After getting back the specimens he might have forgotten to label them accordingly. In spite of the dubious circumstances, the authors are convinced that the Maastricht specimens represent the type specimens of A. schuppi .</p><p>A.B. Gurney had labelled one of the specimens as lectotype: "Top specimen (mature ♀) designated lectoholotype Attaphila schuppi . det. A.B.Gurney 1971". In fact, both specimens are mature females, and since the designation was not published, it is ignored.</p></div>	https://treatment.plazi.org/id/BA4D3E16229A56FA8322212D7AC846F6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter	Bohn, Horst, Nehring, Volker, G., Jonathan Rodriguez, Klass, Klaus-Dieter (2021): Revision of the genus Attaphila (Blattodea: Blaberoidea), myrmecophiles living in the mushroom gardens of leaf-cutting ants. Arthropod Systematics & amp; Phylogeny 79: 205-280, DOI: http://dx.doi.org/10.3897/asp.79.e67569, URL: http://dx.doi.org/10.3897/asp.79.e67569
7A973A062FFE514487AED823723C301D.text	7A973A062FFE514487AED823723C301D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Attaphila sexdentis Bolivar 1905	<div><p>4.8. Attaphila sexdentis Bolivar, 1905</p><p>Figs 11C, D, 13A, 14G, 17F, 18L, 21C, 22I</p><p>Attaphila sexdentis Bolívar, 1905: 137; Princis 1963: 111.</p><p>Material studied.</p><p>Type material. Holotype, 1♀, [Brazil], Rio Grande do Sul, San Leopoldo, b. Atta sexdens, (Dr. Dutra!) (completely on three slides: Bo 1233) (M. Maastricht).</p><p>Diagnosis.</p><p>Female Characterised by the unique shape of S7: posterior parts of the lateral borders anteriorly converging, in all other species (more or less) parallel.</p><p>Description.</p><p>Size Length of body (dried): female 3 mm. Surface bristles of tergites 2-5 approximately arranged in two transversal rows (Fig. 11C). Transversal ridges tr2-5 without distinct excurvations (Fig. 11C, compare grey arrows in Fig. 11A).</p><p>Male Unknown.</p><p>Female Tergites 6,7: (Figs 11D, 13A, 14G) Median lobe of T7 rather long; transversal ridges tr6 and tr7 complete, tr6 sublaterally with a weak, but distinct bend; surface bristles very numerous on T6, only few on T7, small (slightly smaller than the larger ones in A. paucisetosa), focused to central part of surface area. Subgenital plate (Figs 17F, 18L) semicircular, lateral borders distinctly converging to the anterior; lateral parts of transversal ridge (sr7-l) fairly straight, ridge mesally ending well after having reached a transversal orientation, with a fairly narrow median gap (end parts of ridge only slightly more developed than in A. aptera, but lateral parts more strongly inclined mesally, thus leaving a narrower gap), hardly recurved sr7-m ends indicating the ridge to be at most weakly bisinuate. Genitalia: (Figs 21C, 22I) Spermathecal plate sp and pouch gcp not clearly identifiable. In laterosternal shelf sclerite (Fig. 22I) central part (c) moderately long and posteriorly widely truncate (slightly rounded), arms (a) moderately wide, wing parts (w) moderately wide, their base restricted to posterior half of tubes; tubes (lst) with a straight mesal border and slightly narrowed towards the anterior, anteriorly more or less transversally cut. Mesal gonangulum (gg-m) sclerites distinct (presence of lateral ones questionable; Fig. 21C).</p><p>Host species.</p><p>Atta sexdens (Linnaeus, 1758).</p><p>Distribution.</p><p>Brazil: Est. Rio Grande do Sul, only known from the type locality San Leopoldo.</p></div>	https://treatment.plazi.org/id/7A973A062FFE514487AED823723C301D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter	Bohn, Horst, Nehring, Volker, G., Jonathan Rodriguez, Klass, Klaus-Dieter (2021): Revision of the genus Attaphila (Blattodea: Blaberoidea), myrmecophiles living in the mushroom gardens of leaf-cutting ants. Arthropod Systematics & amp; Phylogeny 79: 205-280, DOI: http://dx.doi.org/10.3897/asp.79.e67569, URL: http://dx.doi.org/10.3897/asp.79.e67569
60B57AB4EDE85E7E9DFC9726AC297AEB.text	60B57AB4EDE85E7E9DFC9726AC297AEB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Attaphila sinuosocarinata sp. nov. Bohn and Klass	<div><p>4.9. Attaphila sinuosocarinata sp. nov. Bohn and Klass</p><p>Figs 10C-E, 13B, 14H, 17A, B, 18M, N, 21B, 22K, L</p><p>Material studied.</p><p>Type material. Holotype, 1♀, [Brazil], São Paulo, Baxnery, XI.20., A. Hempel leg., #20 242 (head missing, otherwise completely on one slide: Bo 1273) (M. São Paulo) . - Paratypes, 9♀: 1♀, same data as holotype (completely on two slides: Bo 1439) (ZS Munich) . 1♀, same data as holotype (abdomen on one slide: Bo 1288) (M. Maastricht, NHMM 2021 009) . 2♀, same data as holotype (M. São Paulo) . 5♀, [Brazil], São Paulo , Hinanga, # 6287 A (abdomen and legs of 1♀ on one slide: Bo 1272; abdomen of 1♀ on one slide: Bo 1287) (M. São Paulo) .</p><p>Diagnosis.</p><p>Female Surface bristles of T2-5 similar as in A. paucisetosa and A. multisetosa fairly in one line, distinguished from the former by the more numerous bristles on T6, and from both by the more strongly bent transversal ridge of S7 and the unique shape of tubes lst (see key).</p><p>Etymology.</p><p>The species name refers to the sine-shaped (Latin: sinuosus) transversal ridge (Latin: carina) of S7.</p><p>Description.</p><p>Size Length of body (dried): female 2.75-3.2 mm. Surface bristles of tergites 2-5 arranged in one transversal row (less regular than in A. paucisetosa; Fig. 10C). Transversal ridges tr2-5 without distinct excurvations (Fig. 10C, compare grey arrows in Fig. 11A).</p><p>Male Unknown.</p><p>Female Tergites 6,7: (Figs 10D, E, 13B, 14H) Median lobe of T7 rather long; transversal ridges tr6 and tr7 complete, tr6 sublaterally with a weak, but distinct bend; surface bristles very numerous on T6, only few on T7, very small, focused to central part of surface area. Subgenital plate (Figs 17A, B, 18M, N) rounded-rectangular, with parallel lateral borders; lateral parts of transversal ridge (sr7-l) with a strong mesally directed curvation, lateral terminal parts with a distinct laterally directed curvation; at least in part of the cases ridge mesally ending after having reached a transversal orientation, with a fairly wide median gap, partly recurved sr7-m ends indicating the ridge to be bisinuate (course of the transveral ridge very similar as in A. aptera, but lateral parts more strongly inclined mesally, thus leaving a narrower gap). Genitalia: (Figs 21B, 22K, L) Spermathecal plate sp large, pouch gcp quite large (Fig. 21B). In laterosternal shelf sclerite (Fig. 22K, L) central part (c) moderately long and posteriorly widely rounded-truncate, arms (a) moderately wide, wing parts (w) moderately wide, their base restricted to posterior half of tubes; tubes (lst) with a straight mesal border, slightly narrowed towards the anterior, anteriorly more or less transversally cut. Mesal gonangulum sclerites (gg-m) distinct, lateral ones absent (Fig. 21B).</p><p>Host species.</p><p>Unknown.</p><p>Distribution.</p><p>Brazil: Est. São Paulo (the possible localities “Baxnery” and “Hinanga” indicated on the labels could not be identified).</p></div>	https://treatment.plazi.org/id/60B57AB4EDE85E7E9DFC9726AC297AEB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter	Bohn, Horst, Nehring, Volker, G., Jonathan Rodriguez, Klass, Klaus-Dieter (2021): Revision of the genus Attaphila (Blattodea: Blaberoidea), myrmecophiles living in the mushroom gardens of leaf-cutting ants. Arthropod Systematics & amp; Phylogeny 79: 205-280, DOI: http://dx.doi.org/10.3897/asp.79.e67569, URL: http://dx.doi.org/10.3897/asp.79.e67569
