taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
EDDDF3A648CA54AABF1EF10CC8EDACD8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563270	https://doi.org/10.3897/asp.79.e67569.figure1	Figure 1. A - D: Head structures of Attaphila species, head made transparent by treatment with KOH. A: Entire head with focus on antennae and circumantennal pit (inset: median part, darkened). B: Entire head with focus on mouthparts (cardo and stipes seen behind circumantennal pit; inset: scheme of mandibular dentition in anterior view, mdl = left, mdr = right, elements above transversal line belonging to mola). C: Entire head with left antenna rectangularly bent between scapus and pedicellus. In A-C different levels of tentorium visible internally. D: Base of right antenna and surrounding parts of head capsule at higher magnification, also showing dorsal excavation at distal end of scapus allowing the strong bend shown in C. E - H: Thoracic nota of Attaphila species. E: All thoracic nota of a female, stretched. F: Pronotum of a male. G, H: Right part of metanotum of males with the strongly reduced hindwing. - Species: A - C: A. aptera, male (A, B: Bo 1224), larval female (C: Bo 1243); D: A. fungicola, male (Bo 1229); E, G: A. bergi, female (Bo 1239), male (Bo 1283); F, H: A. paucisetosa, male (Bo 1240). - Abbreviations: ata anterior tentorial arm; atb anterior tentorial bridge; atp anterior tentorial pit; car cardo of maxilla; cap circumantennal pit (with sharp dorsal edge capd visible and bottom capb of the pit shining through); cc coronal cleavage line; cpe compound eye; ct corpotentorium; fc frontal cleavage line; fl fagellum (all antennomeres following pedicellus, numbered from base as fl 1 etc.); hw hindwing; hwar level of (reduced) hindwing articulation; li lacinia; lp labial palpus (I-III: palpomeres); md mandible (left mdl, right mdr; I-IV: tip and incisivi); mo mola of mandible; mp maxillary palpus (I-V: palpomeres); N thoracic notum (N 1 pro-, N 2 meso-, N 3 metanotum); pc pedicellus; pta posterior tentorial arm; sc scapus; sc-pc scapus-pedicellus articulation; scex dorsal excavation at distal end of scapus; s-chA field of sensilla chaetica A; s-cpf row of sensilla campaniformia; sti stipes of maxilla; tp tentorial perforation.	Figure 1. A - D: Head structures of Attaphila species, head made transparent by treatment with KOH. A: Entire head with focus on antennae and circumantennal pit (inset: median part, darkened). B: Entire head with focus on mouthparts (cardo and stipes seen behind circumantennal pit; inset: scheme of mandibular dentition in anterior view, mdl = left, mdr = right, elements above transversal line belonging to mola). C: Entire head with left antenna rectangularly bent between scapus and pedicellus. In A-C different levels of tentorium visible internally. D: Base of right antenna and surrounding parts of head capsule at higher magnification, also showing dorsal excavation at distal end of scapus allowing the strong bend shown in C. E - H: Thoracic nota of Attaphila species. E: All thoracic nota of a female, stretched. F: Pronotum of a male. G, H: Right part of metanotum of males with the strongly reduced hindwing. - Species: A - C: A. aptera, male (A, B: Bo 1224), larval female (C: Bo 1243); D: A. fungicola, male (Bo 1229); E, G: A. bergi, female (Bo 1239), male (Bo 1283); F, H: A. paucisetosa, male (Bo 1240). - Abbreviations: ata anterior tentorial arm; atb anterior tentorial bridge; atp anterior tentorial pit; car cardo of maxilla; cap circumantennal pit (with sharp dorsal edge capd visible and bottom capb of the pit shining through); cc coronal cleavage line; cpe compound eye; ct corpotentorium; fc frontal cleavage line; fl fagellum (all antennomeres following pedicellus, numbered from base as fl 1 etc.); hw hindwing; hwar level of (reduced) hindwing articulation; li lacinia; lp labial palpus (I-III: palpomeres); md mandible (left mdl, right mdr; I-IV: tip and incisivi); mo mola of mandible; mp maxillary palpus (I-V: palpomeres); N thoracic notum (N 1 pro-, N 2 meso-, N 3 metanotum); pc pedicellus; pta posterior tentorial arm; sc scapus; sc-pc scapus-pedicellus articulation; scex dorsal excavation at distal end of scapus; s-chA field of sensilla chaetica A; s-cpf row of sensilla campaniformia; sti stipes of maxilla; tp tentorial perforation.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
EDDDF3A648CA54AABF1EF10CC8EDACD8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563271	https://doi.org/10.3897/asp.79.e67569.figure2	Figure 2. A-I: Antennae of Attaphila species of various age. A, H, I: Presumably second stage larva (head width 0.52 mm), with enlarged details of the proximal (H) and distal regions (I, distal end to the right), both from (A); B - E: intermediate and late larval stages, head width 0.66 (B), 0.61 (C), 0.76 (D), 0.73 mm (E); F, G: imagines. J-M: Imaginal antennae of Blattellidae species, in (J) showing parts of the proximal, intermediate and distal region, in (K-M) only of the proximal and distal region. - Species: A, C, E, H, I: Attaphila paucisetosa (A, H, I: Colombia, Cb 4 / 3; C: Bo 1458; E: Bo 1433); B, D, F: A. aptera (B: Bo 1457; D: Bo 1292; F: Bo 1225); G: A. bergi (Bo 1274); J: Blattella germanica (ex cult.); K: Symploce pallens (ex cult.); L: Xestoblatta cantralli (Costa Rica, CR 15); M: Ischnoptera sp. (Costa Rica, CR 13 / 1). - Abbreviations and symbols: sc scapus; pc pedicellus; me meriston (first flagellomere of current stage); + new proximal flagellomeres generated by a division of the meriston into two or three flagellomeres during the preceding intermoult period in B, E, H, in C, D the division had presumably already occurred one moult earlier; L - early larval stage; L + intermediate or late larval stage; Im imago.	Figure 2. A-I: Antennae of Attaphila species of various age. A, H, I: Presumably second stage larva (head width 0.52 mm), with enlarged details of the proximal (H) and distal regions (I, distal end to the right), both from (A); B - E: intermediate and late larval stages, head width 0.66 (B), 0.61 (C), 0.76 (D), 0.73 mm (E); F, G: imagines. J-M: Imaginal antennae of Blattellidae species, in (J) showing parts of the proximal, intermediate and distal region, in (K-M) only of the proximal and distal region. - Species: A, C, E, H, I: Attaphila paucisetosa (A, H, I: Colombia, Cb 4 / 3; C: Bo 1458; E: Bo 1433); B, D, F: A. aptera (B: Bo 1457; D: Bo 1292; F: Bo 1225); G: A. bergi (Bo 1274); J: Blattella germanica (ex cult.); K: Symploce pallens (ex cult.); L: Xestoblatta cantralli (Costa Rica, CR 15); M: Ischnoptera sp. (Costa Rica, CR 13 / 1). - Abbreviations and symbols: sc scapus; pc pedicellus; me meriston (first flagellomere of current stage); + new proximal flagellomeres generated by a division of the meriston into two or three flagellomeres during the preceding intermoult period in B, E, H, in C, D the division had presumably already occurred one moult earlier; L - early larval stage; L + intermediate or late larval stage; Im imago.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
EDDDF3A648CA54AABF1EF10CC8EDACD8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563272	https://doi.org/10.3897/asp.79.e67569.figure3(a-f)	Figure 3 (A-J). Tegmina (forewings) of males of Attaphila species, dorsal views, left tegmen at the left. - Species: A, B: A. aptera (Bo 1252); C, D: A. bergi (Bo 1443); E, F: A. flava (HT Bo 1280); G, H: A. fungicola (Bo 1229); I, J: A. paucisetosa (Bo 1441). - Abbreviation: fwar forewing articulation area. In A borders specified as addressed in the text.	Figure 3 (A-J). Tegmina (forewings) of males of Attaphila species, dorsal views, left tegmen at the left. - Species: A, B: A. aptera (Bo 1252); C, D: A. bergi (Bo 1443); E, F: A. flava (HT Bo 1280); G, H: A. fungicola (Bo 1229); I, J: A. paucisetosa (Bo 1441). - Abbreviation: fwar forewing articulation area. In A borders specified as addressed in the text.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
EDDDF3A648CA54AABF1EF10CC8EDACD8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563274	https://doi.org/10.3897/asp.79.e67569.figure5	Figure 5. A - I: Tergite specialisations of the males of Attaphila species, medially on T 1 (A - D), or laterally at the anterior border of T 2 (E - I), in E-G from right side of tergite, in H and I from left side. - Species: A, B: A. aptera (Bo 1227, HT Bo 1258); C, D: A. bergi (Bo 1283, Bo 1274); E, F: A. paucisetosa (PT Bo 1254, Bo 1256); G, H: A. fungicola (LT Bo 1265); I: A. flava (HT Bo 1280). - Abbreviations: T 1 - a, T 2 - a anterior borders of tergites T 1,2; T 1 - p posterior border of T 1; tr 1, tr 2 transversal ridges of T 1,2; msa 1 anterior median specialisation of tergite T 1; msl 2 lateral specialisation of tergite T 2; msp 1 posterior median specialisation of tergite T 1; pg pores of glands, tiny in A, B, between T 1 - a and tr 1, larger in E as part of msl 2 specialisations. - J, K: Paraprocts, tergite 10 and cerci in ventral view, of J: A. aptera (Bo 1224) and K: A. paucisetosa (HT Bo 1258). - Abbreviations and arrows: Ca sclerite at mesal base of cercus; ce cercus; hmp hook-like mesal projection of paraproct; PPl, PPr left and right paraproct; T 9 tergite 9; T 10 tergite 10 with its ventrally bent lateral (paratergal) parts T 10 p (larger median parts of T 10 out of focus); white arrows: articulation between mesal end of T 10 p and lateral end of paraproct.	Figure 5. A - I: Tergite specialisations of the males of Attaphila species, medially on T 1 (A - D), or laterally at the anterior border of T 2 (E - I), in E-G from right side of tergite, in H and I from left side. - Species: A, B: A. aptera (Bo 1227, HT Bo 1258); C, D: A. bergi (Bo 1283, Bo 1274); E, F: A. paucisetosa (PT Bo 1254, Bo 1256); G, H: A. fungicola (LT Bo 1265); I: A. flava (HT Bo 1280). - Abbreviations: T 1 - a, T 2 - a anterior borders of tergites T 1,2; T 1 - p posterior border of T 1; tr 1, tr 2 transversal ridges of T 1,2; msa 1 anterior median specialisation of tergite T 1; msl 2 lateral specialisation of tergite T 2; msp 1 posterior median specialisation of tergite T 1; pg pores of glands, tiny in A, B, between T 1 - a and tr 1, larger in E as part of msl 2 specialisations. - J, K: Paraprocts, tergite 10 and cerci in ventral view, of J: A. aptera (Bo 1224) and K: A. paucisetosa (HT Bo 1258). - Abbreviations and arrows: Ca sclerite at mesal base of cercus; ce cercus; hmp hook-like mesal projection of paraproct; PPl, PPr left and right paraproct; T 9 tergite 9; T 10 tergite 10 with its ventrally bent lateral (paratergal) parts T 10 p (larger median parts of T 10 out of focus); white arrows: articulation between mesal end of T 10 p and lateral end of paraproct.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
EDDDF3A648CA54AABF1EF10CC8EDACD8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563275	https://doi.org/10.3897/asp.79.e67569.figure6	Figure 6. Abdominal tergites and terminalia of Attaphila aptera. A, B: Male. A: T 1 - 7 (Bo 1227; see also Fig. S 1 A), T 1 with median specialisation including its bristles, otherwise without long bristles (the two long bristles on the left have no base and are certainly contaminations from other tergites); B: Terminalia (Bo 1256), ventral view, with T 9 and T 10, cerci, and paraprocts. C - E: Female. C: Parts of T 1,2 (Bo 1257); D: T 2 - 5 (Bo 1253; see also Fig. S 1 B); E: T 6,7 (Bo 1253). - Abbreviations: ce cercus; hmp hook-like mesal projection on right paraproct; ltga 9 lateral tergal apodeme of tergite T 9; msp 1 posterior median specialisation of T 1; PPr right paraproct; T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered).	Figure 6. Abdominal tergites and terminalia of Attaphila aptera. A, B: Male. A: T 1 - 7 (Bo 1227; see also Fig. S 1 A), T 1 with median specialisation including its bristles, otherwise without long bristles (the two long bristles on the left have no base and are certainly contaminations from other tergites); B: Terminalia (Bo 1256), ventral view, with T 9 and T 10, cerci, and paraprocts. C - E: Female. C: Parts of T 1,2 (Bo 1257); D: T 2 - 5 (Bo 1253; see also Fig. S 1 B); E: T 6,7 (Bo 1253). - Abbreviations: ce cercus; hmp hook-like mesal projection on right paraproct; ltga 9 lateral tergal apodeme of tergite T 9; msp 1 posterior median specialisation of T 1; PPr right paraproct; T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered).	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
EDDDF3A648CA54AABF1EF10CC8EDACD8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563281	https://doi.org/10.3897/asp.79.e67569.figure12	Figure 12. Abdominal tergites T 6,7 of females of Attaphila species. A: A. aptera (Bo 1257). B: A. bergi (Bo 1282). C: A. fungicola (Bo 1264). D: A. multisetosa (HT Bo 1270). E: A. paucisetosa (PT Bo 1255). F: A. schuppi (ST Bo 1237), transversal ridge tr 7 almost completely missing, only tiny remnants present near the associated bristles (see Fig. 14 F). - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 12. Abdominal tergites T 6,7 of females of Attaphila species. A: A. aptera (Bo 1257). B: A. bergi (Bo 1282). C: A. fungicola (Bo 1264). D: A. multisetosa (HT Bo 1270). E: A. paucisetosa (PT Bo 1255). F: A. schuppi (ST Bo 1237), transversal ridge tr 7 almost completely missing, only tiny remnants present near the associated bristles (see Fig. 14 F). - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
EDDDF3A648CA54AABF1EF10CC8EDACD8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563282	https://doi.org/10.3897/asp.79.e67569.figure13	Figure 13. Abdominal tergites T 6,7 of females (A, B) and males (C - F) of Attaphila species. A: A. sexdentis, female (Bo 1233). B: A. sinuosocarinata, female (HT Bo 1273). C: A. aptera, male (Bo 1256). D: A. bergi, male (Bo 1274), transversal ridge tr 7 weakly developed, sublaterally with a large gap. E: A. paucisetosa, male (Bo 1240), transversal ridge tr 7 missing. F: A. fungicola, male (Bo 1229), transversal ridge tr 7 missing. - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 13. Abdominal tergites T 6,7 of females (A, B) and males (C - F) of Attaphila species. A: A. sexdentis, female (Bo 1233). B: A. sinuosocarinata, female (HT Bo 1273). C: A. aptera, male (Bo 1256). D: A. bergi, male (Bo 1274), transversal ridge tr 7 weakly developed, sublaterally with a large gap. E: A. paucisetosa, male (Bo 1240), transversal ridge tr 7 missing. F: A. fungicola, male (Bo 1229), transversal ridge tr 7 missing. - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
EDDDF3A648CA54AABF1EF10CC8EDACD8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563283	https://doi.org/10.3897/asp.79.e67569.figure14	Figure 14. Size and distribution of bristles in the median part of tergites T 6,7 in females (A-H) and males (I) of Attaphila species, phase contrast images. A: A. aptera, female (Bo 1257). B: A. bergi, female (Bo 1282). C: A. fungicola, female (Bo 1264). D: A. multisetosa, female (Bo 1270). E: A. paucisetosa, female (PT Bo 1255). F: A. schuppi, female (ST Bo 1237), with only short remnants of transversal ridge tr 7. G: A. sexdentis, female (Bo 1233). H: A. sinuosocarinata, female (HT Bo 1273). I: A. aptera, male (Bo 1256). - Abbreviations: T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 14. Size and distribution of bristles in the median part of tergites T 6,7 in females (A-H) and males (I) of Attaphila species, phase contrast images. A: A. aptera, female (Bo 1257). B: A. bergi, female (Bo 1282). C: A. fungicola, female (Bo 1264). D: A. multisetosa, female (Bo 1270). E: A. paucisetosa, female (PT Bo 1255). F: A. schuppi, female (ST Bo 1237), with only short remnants of transversal ridge tr 7. G: A. sexdentis, female (Bo 1233). H: A. sinuosocarinata, female (HT Bo 1273). I: A. aptera, male (Bo 1256). - Abbreviations: T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
EDDDF3A648CA54AABF1EF10CC8EDACD8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563284	https://doi.org/10.3897/asp.79.e67569.figure15	Figure 15. Size and distribution of bristles in the median part of tergites T 6,7 in males (A - E) and in larval males and females (F - L) of Attaphila species, phase contrast images. A: A. bergi, male (Bo 1274). B: A. fungicola, male (LT Bo 1265). C: A. flava, male (HT Bo 1280). D: A. paucisetosa, male (Cb 2 / 1). E: A. paucisetosa, male, with slightly shorter bristles than in preceding specimen (Bo 1445). F: A. aptera, larval male (Bo 1291). G: A. aptera, larval female (Bo 1289). H: A. aptera, larval male (HT Bo 1232). I: A. bergi, larval male (Bo 1230); medium sized bristles belong to T 6, the very long ones are from T 5 covering most of T 6. J: A. bergi, larval female (Bo 1285). K: A. paucisetosa, larval male (Bo 1433). L: A. paucisetosa, larval female (Bo 1293). - Abbreviations: T 5 - p, T 6 - p, T 7 - p posterior borders of tergites T 5, T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 15. Size and distribution of bristles in the median part of tergites T 6,7 in males (A - E) and in larval males and females (F - L) of Attaphila species, phase contrast images. A: A. bergi, male (Bo 1274). B: A. fungicola, male (LT Bo 1265). C: A. flava, male (HT Bo 1280). D: A. paucisetosa, male (Cb 2 / 1). E: A. paucisetosa, male, with slightly shorter bristles than in preceding specimen (Bo 1445). F: A. aptera, larval male (Bo 1291). G: A. aptera, larval female (Bo 1289). H: A. aptera, larval male (HT Bo 1232). I: A. bergi, larval male (Bo 1230); medium sized bristles belong to T 6, the very long ones are from T 5 covering most of T 6. J: A. bergi, larval female (Bo 1285). K: A. paucisetosa, larval male (Bo 1433). L: A. paucisetosa, larval female (Bo 1293). - Abbreviations: T 5 - p, T 6 - p, T 7 - p posterior borders of tergites T 5, T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
EDDDF3A648CA54AABF1EF10CC8EDACD8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563286	https://doi.org/10.3897/asp.79.e67569.figure17	Figure 17. Subgenital plate (sternite S 7) of females (A - F) and larval females (G, H) of Attaphila species. A, B: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). C, D: A. aptera (Bo 1225, Bo 1253). E: A. multisetosa (HT Bo 1270). F: A. sexdentis (HT Bo 1233). G: A. aptera, larval female (Bo 1289). H: A. bergi, larval female (Bo 1231). - Abbreviations and arrows: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars). Arrows in B, C, F pointing to lateral borders of S 7, which are parallel or slightly divergent (to the anterior) in A-E, but convergent in F.	Figure 17. Subgenital plate (sternite S 7) of females (A - F) and larval females (G, H) of Attaphila species. A, B: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). C, D: A. aptera (Bo 1225, Bo 1253). E: A. multisetosa (HT Bo 1270). F: A. sexdentis (HT Bo 1233). G: A. aptera, larval female (Bo 1289). H: A. bergi, larval female (Bo 1231). - Abbreviations and arrows: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars). Arrows in B, C, F pointing to lateral borders of S 7, which are parallel or slightly divergent (to the anterior) in A-E, but convergent in F.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
EDDDF3A648CA54AABF1EF10CC8EDACD8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563287	https://doi.org/10.3897/asp.79.e67569.figure18	Figure 18. Anterior half of subgenital plate (sternite S 7) of females of Attaphila species, shape of sternal transversal ridge, phase contrast images (same objects as shown in Figs 16, 17, except for Bo 1236 in F). A, B: A. aptera (Bo 1225, Bo 1253). C, D: A. bergi (Bo 1275, Bo 1282). E, F: A. fungicola (Bo 1264, Bo 1236). G: A. multisetosa (HT Bo 1270). H, I: A. paucisetosa (Cb 2 / 2, PT Bo 1255). J, K: A. schuppi (ST Bo 1234, ST Bo 1237). L: A. sexdentis (HT Bo 1233). M, N: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). - Abbreviations and symbols: sr 7 transversal ridge of subgenital plate S 7; short interruptions of this ridge indicated by arrows, larger gaps delimited by double bars.	Figure 18. Anterior half of subgenital plate (sternite S 7) of females of Attaphila species, shape of sternal transversal ridge, phase contrast images (same objects as shown in Figs 16, 17, except for Bo 1236 in F). A, B: A. aptera (Bo 1225, Bo 1253). C, D: A. bergi (Bo 1275, Bo 1282). E, F: A. fungicola (Bo 1264, Bo 1236). G: A. multisetosa (HT Bo 1270). H, I: A. paucisetosa (Cb 2 / 2, PT Bo 1255). J, K: A. schuppi (ST Bo 1234, ST Bo 1237). L: A. sexdentis (HT Bo 1233). M, N: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). - Abbreviations and symbols: sr 7 transversal ridge of subgenital plate S 7; short interruptions of this ridge indicated by arrows, larger gaps delimited by double bars.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
EDDDF3A648CA54AABF1EF10CC8EDACD8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563288	https://doi.org/10.3897/asp.79.e67569.figure19	Figure 19. Terminalia with genitalia of females of Attaphila species, ventral view, posterior end on top. A: A. aptera (Bo 1253), without laterosternal shelf area; inset showing median part of genital chamber dorsal wall. B: A. bergi (Bo 1239), without laterosternal shelf area. C, D: A. fungicola (Bo 1236, Bo 1264), C with, D without laterosternal shelf area. - Abbreviations: aa anterior arch; ca central apodeme; ce cercus; gcf fold dividing genital chamber (posterior edge labeled); gcp pouch of genital chamber (one-sided); gg-m, gg-l mesal and lateral gonangulum sclerites, the latter present only in A. aptera; IC intercalary sclerite; ls-p posterolateral extremity of laterosternal-shelf sclerite; lst laterosternal-shelf tube; pl posterior lobe; PP paraproct; pt 8,9 extension of paratergite 9; sp spermathecal plate; T 9, T 10 abdominal tergites 9 and 10; T 10 - p posterior border of tergite 10; v 1, v 3 1 st and 3 rd valves of ovipositor; vlf valvifer of segment 8; A articulations, A 1 between gg-l and pt 8,9, A 2 between gg-m and pl, A 3 between gg-m and basal sclerotisation of v 1, A 5 between pt 8,9 and vlf. Arrow in A showing mesal border of putative weak sclerotisation in dorsal wall of genital chamber. (See Supplement 1 and Supplement 3 Fig. S 2 A, B, C for additional explanations and more complete labeling using different terminologies.)	Figure 19. Terminalia with genitalia of females of Attaphila species, ventral view, posterior end on top. A: A. aptera (Bo 1253), without laterosternal shelf area; inset showing median part of genital chamber dorsal wall. B: A. bergi (Bo 1239), without laterosternal shelf area. C, D: A. fungicola (Bo 1236, Bo 1264), C with, D without laterosternal shelf area. - Abbreviations: aa anterior arch; ca central apodeme; ce cercus; gcf fold dividing genital chamber (posterior edge labeled); gcp pouch of genital chamber (one-sided); gg-m, gg-l mesal and lateral gonangulum sclerites, the latter present only in A. aptera; IC intercalary sclerite; ls-p posterolateral extremity of laterosternal-shelf sclerite; lst laterosternal-shelf tube; pl posterior lobe; PP paraproct; pt 8,9 extension of paratergite 9; sp spermathecal plate; T 9, T 10 abdominal tergites 9 and 10; T 10 - p posterior border of tergite 10; v 1, v 3 1 st and 3 rd valves of ovipositor; vlf valvifer of segment 8; A articulations, A 1 between gg-l and pt 8,9, A 2 between gg-m and pl, A 3 between gg-m and basal sclerotisation of v 1, A 5 between pt 8,9 and vlf. Arrow in A showing mesal border of putative weak sclerotisation in dorsal wall of genital chamber. (See Supplement 1 and Supplement 3 Fig. S 2 A, B, C for additional explanations and more complete labeling using different terminologies.)	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
EDDDF3A648CA54AABF1EF10CC8EDACD8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563291	https://doi.org/10.3897/asp.79.e67569.figure22	Figure 22. Laterosternal shelf area of females of Attaphila species, ventral view, posteriorly on top. A, B: A. aptera (Bo 1225, Bo 1257), in A left tube distorted (ventral part of tube squeezed laterally, as indicated by grey arrow). C, D: A. bergi (Bo 1282, Bo 1239). E, F: A. fungicola (Bo 1264, Bo 1447). G: A. multisetosa (HT Bo 1270). H: A. paucisetosa (PT Bo 1255). I: A. sexdentis (HT Bo 1233). J: A. schuppi (ST Bo 1237). K, L: A. sinuosocarinata (HT Bo 1273, PT Bo 1288). - Abbreviations: isf intersternal folds (of floor of vestibulum); ls laterosternal-shelf sclerite (with central part c, arm part a, wing part w, posterolateral extremity p); lst laterosternal-shelf tube; vfl floor of vestibulum. (See Supplement 3 Fig. S 2 D, E for more complete labeling using different terminologies.) - Arrows: Black arrows in A, D, I: twisting of sclerite at transition between ls-w and ls-p. Grey arrow in A indicates direction of squeezing of tube lst. Blue arrowheads: anterior extremity of wing part ls-w on mesal border of tube lst, ls-w either restricted to posterior half of tubes lst (e. g. K) or reaching far into anterior half (e. g. D). Red arrowheads: area where the posterior margin of ls-a bends dorsally, and where a bend or kink is often present in (inner) lateral border of tube lst (compare white arrowheads in Fig. 23 C, D, H, I, J). Black arrowheads: membranous inner lateral border of tube lst posterior to the bend or kink (compare Fig. 23 C, D, H, I and black arrowheads in sections beside D). - Inserts in B, D, E, K showing right (of picture and animal) tube lst with posterior margin of ls-a (red line; compare Fig. 23 C-E) and anterior margin of ls-w (blue line; compare Fig. 23 F, G).	Figure 22. Laterosternal shelf area of females of Attaphila species, ventral view, posteriorly on top. A, B: A. aptera (Bo 1225, Bo 1257), in A left tube distorted (ventral part of tube squeezed laterally, as indicated by grey arrow). C, D: A. bergi (Bo 1282, Bo 1239). E, F: A. fungicola (Bo 1264, Bo 1447). G: A. multisetosa (HT Bo 1270). H: A. paucisetosa (PT Bo 1255). I: A. sexdentis (HT Bo 1233). J: A. schuppi (ST Bo 1237). K, L: A. sinuosocarinata (HT Bo 1273, PT Bo 1288). - Abbreviations: isf intersternal folds (of floor of vestibulum); ls laterosternal-shelf sclerite (with central part c, arm part a, wing part w, posterolateral extremity p); lst laterosternal-shelf tube; vfl floor of vestibulum. (See Supplement 3 Fig. S 2 D, E for more complete labeling using different terminologies.) - Arrows: Black arrows in A, D, I: twisting of sclerite at transition between ls-w and ls-p. Grey arrow in A indicates direction of squeezing of tube lst. Blue arrowheads: anterior extremity of wing part ls-w on mesal border of tube lst, ls-w either restricted to posterior half of tubes lst (e. g. K) or reaching far into anterior half (e. g. D). Red arrowheads: area where the posterior margin of ls-a bends dorsally, and where a bend or kink is often present in (inner) lateral border of tube lst (compare white arrowheads in Fig. 23 C, D, H, I, J). Black arrowheads: membranous inner lateral border of tube lst posterior to the bend or kink (compare Fig. 23 C, D, H, I and black arrowheads in sections beside D). - Inserts in B, D, E, K showing right (of picture and animal) tube lst with posterior margin of ls-a (red line; compare Fig. 23 C-E) and anterior margin of ls-w (blue line; compare Fig. 23 F, G).	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
EDDDF3A648CA54AABF1EF10CC8EDACD8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563293	https://doi.org/10.3897/asp.79.e67569.figure24	Figure 24. Subgenital plate (sternite S 9) and phallomeres of males of Attaphila species, dorsal view, anteriorly on top. A, B: A. aptera, subgenital plate with phallomeres (Bo 1256, Bo 1224), anterior part of R 3 sclerite out of frame in B. C-G: A. bergi. C: Subgenital plate without phallomeres (Bo 1274); D: Subgenital plate with phallomeres (Bo 1283); E: Isolated hook of left phallomere (Bo 1274); F: Isolated endophallic apodeme (damaged near midlength), virga, and psa-process of left phallomere (Bo 1274); G: Isolated right phallomere (Bo 1274). - Abbreviations: A articulations, A 1 between S 9 and pt 9, A 3 between sclerites R 3 and R 1 P; cs cleft sclerite, composed of the dorsally fused R 1 S and R 2, R 2 - part in contact with arm v of R 3 sclerite (not visible, contact area indicated by white arrow in G); ea endophallic apodeme; h hook (with long membranous base p, wide base b of sclerotised part, neck n, and claw cl); hcl cleft in wall of groove hge; hge groove upon claw part of hook; psa process; pt 9 extension of paratergite 9 contacting sternite 9; R 1 (divided in R 1 S and R 1 P), R 2, R 3 (with anterior arm a, dorsoposterior arm d, ventroposterior arm v) are the principal sclerotisations of the right phallomere; sll, slr left and right stylus; sta 9 anterior apodeme of subgenital plate S 9; vf 9 subgenital lobe; vge grooves along virga (see vge 1, vge 2 in Fig. 32 I); vi virga.	Figure 24. Subgenital plate (sternite S 9) and phallomeres of males of Attaphila species, dorsal view, anteriorly on top. A, B: A. aptera, subgenital plate with phallomeres (Bo 1256, Bo 1224), anterior part of R 3 sclerite out of frame in B. C-G: A. bergi. C: Subgenital plate without phallomeres (Bo 1274); D: Subgenital plate with phallomeres (Bo 1283); E: Isolated hook of left phallomere (Bo 1274); F: Isolated endophallic apodeme (damaged near midlength), virga, and psa-process of left phallomere (Bo 1274); G: Isolated right phallomere (Bo 1274). - Abbreviations: A articulations, A 1 between S 9 and pt 9, A 3 between sclerites R 3 and R 1 P; cs cleft sclerite, composed of the dorsally fused R 1 S and R 2, R 2 - part in contact with arm v of R 3 sclerite (not visible, contact area indicated by white arrow in G); ea endophallic apodeme; h hook (with long membranous base p, wide base b of sclerotised part, neck n, and claw cl); hcl cleft in wall of groove hge; hge groove upon claw part of hook; psa process; pt 9 extension of paratergite 9 contacting sternite 9; R 1 (divided in R 1 S and R 1 P), R 2, R 3 (with anterior arm a, dorsoposterior arm d, ventroposterior arm v) are the principal sclerotisations of the right phallomere; sll, slr left and right stylus; sta 9 anterior apodeme of subgenital plate S 9; vf 9 subgenital lobe; vge grooves along virga (see vge 1, vge 2 in Fig. 32 I); vi virga.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
EDDDF3A648CA54AABF1EF10CC8EDACD8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563297	https://doi.org/10.3897/asp.79.e67569.figure28	Figure 28. Pictures of live Attaphila and their host ants in lab culture, all taken from ant nests in Gamboa, Panama. A - E: A male Attaphila paucisetosa together with Atta colombica ants (white arrow in C, E pointing to cockroach). F, G: Females and larvae of Attaphila aptera together with Acromyrmex octospinosus ants. H: Attaphila aptera female or larva together with Acromyrmex octospinosus worker.	Figure 28. Pictures of live Attaphila and their host ants in lab culture, all taken from ant nests in Gamboa, Panama. A - E: A male Attaphila paucisetosa together with Atta colombica ants (white arrow in C, E pointing to cockroach). F, G: Females and larvae of Attaphila aptera together with Acromyrmex octospinosus ants. H: Attaphila aptera female or larva together with Acromyrmex octospinosus worker.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
EDDDF3A648CA54AABF1EF10CC8EDACD8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563280	https://doi.org/10.3897/asp.79.e67569.figure11	Figure 11. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. schuppi (ST Bo 1237). A: T 2 - 5; B: T 6,7. C, D: A. sexdentis (HT Bo 1233). C: T 1 - 5; D: T 6,7. - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to median and lateral excurvations of transversal ridges to the anterior.	Figure 11. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. schuppi (ST Bo 1237). A: T 2 - 5; B: T 6,7. C, D: A. sexdentis (HT Bo 1233). C: T 1 - 5; D: T 6,7. - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to median and lateral excurvations of transversal ridges to the anterior.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
EDDDF3A648CA54AABF1EF10CC8EDACD8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563278	https://doi.org/10.3897/asp.79.e67569.figure9	Figure 9. Abdominal tergites and terminalia of Attaphila paucisetosa. A - D: Male (HT Bo 1258). A: T 1; B: T 2 - 5, T 2 with lateral specialisations; C: T 6,7. D: Terminalia, ventral view, with T 9 (with lateral extension) and T 10, cerci, and paraprocts. E, F: Female (PT Bo 1255). E: T 1 - 5; F: T 6,7. - Abbreviations: ce cercus; msl 2 lateral specialisation of T 2; PPr right paraproct; T n tergite (numbered); pt 9 extension of paratergite of T 9; T n - a anterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to excurvations of male tr 2.	Figure 9. Abdominal tergites and terminalia of Attaphila paucisetosa. A - D: Male (HT Bo 1258). A: T 1; B: T 2 - 5, T 2 with lateral specialisations; C: T 6,7. D: Terminalia, ventral view, with T 9 (with lateral extension) and T 10, cerci, and paraprocts. E, F: Female (PT Bo 1255). E: T 1 - 5; F: T 6,7. - Abbreviations: ce cercus; msl 2 lateral specialisation of T 2; PPr right paraproct; T n tergite (numbered); pt 9 extension of paratergite of T 9; T n - a anterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to excurvations of male tr 2.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
EDDDF3A648CA54AABF1EF10CC8EDACD8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563296	https://doi.org/10.3897/asp.79.e67569.figure27	Figure 27. A: Phylogenetic tree of leaf-cutting ants (according to Cristiano et al. 2020; Acromyrmex lobicornis added, relationships unknown; terminals with several conspecific exemplars simplified). Ant species reported as Attaphila hosts in red and boldface, their association with Attaphila species indicated by coloured lines (based on specimen labels and partly associated reports in literature, or on own results) or grey line with "? " (based on reported occurrence of only this leaf-cutting ant species in region where Attaphila species was collected). Attaphila divided in Attaphila bergi group (orange, with Acromyrmex; possibly paraphyletic), Attaphila fungicola group (blue, with Atta, but unknown for A. sinuosocarinata; likely monophyletic), and the isolated A. sexdentis (green, with Atta). Status of Attaphila bergi (?) Attaphila bergi var. minor unclear (see text 4.2.). B: Geographic distribution of Attaphila species in Neotropical and southern Nearctic regions (based on specimen labels and partly associated reports in literature, or on own results). Attaphila divided in Attaphila bergi group (orange), Attaphila fungicola group (blue), and A. sexdentis (green).	Figure 27. A: Phylogenetic tree of leaf-cutting ants (according to Cristiano et al. 2020; Acromyrmex lobicornis added, relationships unknown; terminals with several conspecific exemplars simplified). Ant species reported as Attaphila hosts in red and boldface, their association with Attaphila species indicated by coloured lines (based on specimen labels and partly associated reports in literature, or on own results) or grey line with "? " (based on reported occurrence of only this leaf-cutting ant species in region where Attaphila species was collected). Attaphila divided in Attaphila bergi group (orange, with Acromyrmex; possibly paraphyletic), Attaphila fungicola group (blue, with Atta, but unknown for A. sinuosocarinata; likely monophyletic), and the isolated A. sexdentis (green, with Atta). Status of Attaphila bergi (?) Attaphila bergi var. minor unclear (see text 4.2.). B: Geographic distribution of Attaphila species in Neotropical and southern Nearctic regions (based on specimen labels and partly associated reports in literature, or on own results). Attaphila divided in Attaphila bergi group (orange), Attaphila fungicola group (blue), and A. sexdentis (green).	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
EDDDF3A648CA54AABF1EF10CC8EDACD8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563294	https://doi.org/10.3897/asp.79.e67569.figure25	Figure 25. Subgenital plate (sternite S 9) and phallomeres of males of Attaphila species, dorsal view, anteriorly on top. A, B: A. paucisetosa, subgenital plate with phallomeres (PT Bo 1254, Bo 1444), hook and right phallomere removed in B. C: A. flava, subgenital plate with phallomeres (HT Bo 1280). D, E: A. fungicola. D showing subgenital plate with phallomeres (LT Bo 1265), hook not seen in profile; E showing isolated hook seen in profile (Bo 1229). - Abbreviations: T 9 p ventrally bent lateral (paratergal) part of tergite 9; otherwise as for Fig. 24.	Figure 25. Subgenital plate (sternite S 9) and phallomeres of males of Attaphila species, dorsal view, anteriorly on top. A, B: A. paucisetosa, subgenital plate with phallomeres (PT Bo 1254, Bo 1444), hook and right phallomere removed in B. C: A. flava, subgenital plate with phallomeres (HT Bo 1280). D, E: A. fungicola. D showing subgenital plate with phallomeres (LT Bo 1265), hook not seen in profile; E showing isolated hook seen in profile (Bo 1229). - Abbreviations: T 9 p ventrally bent lateral (paratergal) part of tergite 9; otherwise as for Fig. 24.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
4A668C721AD95502974FA7553955584A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563270	https://doi.org/10.3897/asp.79.e67569.figure1	Figure 1. A - D: Head structures of Attaphila species, head made transparent by treatment with KOH. A: Entire head with focus on antennae and circumantennal pit (inset: median part, darkened). B: Entire head with focus on mouthparts (cardo and stipes seen behind circumantennal pit; inset: scheme of mandibular dentition in anterior view, mdl = left, mdr = right, elements above transversal line belonging to mola). C: Entire head with left antenna rectangularly bent between scapus and pedicellus. In A-C different levels of tentorium visible internally. D: Base of right antenna and surrounding parts of head capsule at higher magnification, also showing dorsal excavation at distal end of scapus allowing the strong bend shown in C. E - H: Thoracic nota of Attaphila species. E: All thoracic nota of a female, stretched. F: Pronotum of a male. G, H: Right part of metanotum of males with the strongly reduced hindwing. - Species: A - C: A. aptera, male (A, B: Bo 1224), larval female (C: Bo 1243); D: A. fungicola, male (Bo 1229); E, G: A. bergi, female (Bo 1239), male (Bo 1283); F, H: A. paucisetosa, male (Bo 1240). - Abbreviations: ata anterior tentorial arm; atb anterior tentorial bridge; atp anterior tentorial pit; car cardo of maxilla; cap circumantennal pit (with sharp dorsal edge capd visible and bottom capb of the pit shining through); cc coronal cleavage line; cpe compound eye; ct corpotentorium; fc frontal cleavage line; fl fagellum (all antennomeres following pedicellus, numbered from base as fl 1 etc.); hw hindwing; hwar level of (reduced) hindwing articulation; li lacinia; lp labial palpus (I-III: palpomeres); md mandible (left mdl, right mdr; I-IV: tip and incisivi); mo mola of mandible; mp maxillary palpus (I-V: palpomeres); N thoracic notum (N 1 pro-, N 2 meso-, N 3 metanotum); pc pedicellus; pta posterior tentorial arm; sc scapus; sc-pc scapus-pedicellus articulation; scex dorsal excavation at distal end of scapus; s-chA field of sensilla chaetica A; s-cpf row of sensilla campaniformia; sti stipes of maxilla; tp tentorial perforation.	Figure 1. A - D: Head structures of Attaphila species, head made transparent by treatment with KOH. A: Entire head with focus on antennae and circumantennal pit (inset: median part, darkened). B: Entire head with focus on mouthparts (cardo and stipes seen behind circumantennal pit; inset: scheme of mandibular dentition in anterior view, mdl = left, mdr = right, elements above transversal line belonging to mola). C: Entire head with left antenna rectangularly bent between scapus and pedicellus. In A-C different levels of tentorium visible internally. D: Base of right antenna and surrounding parts of head capsule at higher magnification, also showing dorsal excavation at distal end of scapus allowing the strong bend shown in C. E - H: Thoracic nota of Attaphila species. E: All thoracic nota of a female, stretched. F: Pronotum of a male. G, H: Right part of metanotum of males with the strongly reduced hindwing. - Species: A - C: A. aptera, male (A, B: Bo 1224), larval female (C: Bo 1243); D: A. fungicola, male (Bo 1229); E, G: A. bergi, female (Bo 1239), male (Bo 1283); F, H: A. paucisetosa, male (Bo 1240). - Abbreviations: ata anterior tentorial arm; atb anterior tentorial bridge; atp anterior tentorial pit; car cardo of maxilla; cap circumantennal pit (with sharp dorsal edge capd visible and bottom capb of the pit shining through); cc coronal cleavage line; cpe compound eye; ct corpotentorium; fc frontal cleavage line; fl fagellum (all antennomeres following pedicellus, numbered from base as fl 1 etc.); hw hindwing; hwar level of (reduced) hindwing articulation; li lacinia; lp labial palpus (I-III: palpomeres); md mandible (left mdl, right mdr; I-IV: tip and incisivi); mo mola of mandible; mp maxillary palpus (I-V: palpomeres); N thoracic notum (N 1 pro-, N 2 meso-, N 3 metanotum); pc pedicellus; pta posterior tentorial arm; sc scapus; sc-pc scapus-pedicellus articulation; scex dorsal excavation at distal end of scapus; s-chA field of sensilla chaetica A; s-cpf row of sensilla campaniformia; sti stipes of maxilla; tp tentorial perforation.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
4A668C721AD95502974FA7553955584A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563271	https://doi.org/10.3897/asp.79.e67569.figure2	Figure 2. A-I: Antennae of Attaphila species of various age. A, H, I: Presumably second stage larva (head width 0.52 mm), with enlarged details of the proximal (H) and distal regions (I, distal end to the right), both from (A); B - E: intermediate and late larval stages, head width 0.66 (B), 0.61 (C), 0.76 (D), 0.73 mm (E); F, G: imagines. J-M: Imaginal antennae of Blattellidae species, in (J) showing parts of the proximal, intermediate and distal region, in (K-M) only of the proximal and distal region. - Species: A, C, E, H, I: Attaphila paucisetosa (A, H, I: Colombia, Cb 4 / 3; C: Bo 1458; E: Bo 1433); B, D, F: A. aptera (B: Bo 1457; D: Bo 1292; F: Bo 1225); G: A. bergi (Bo 1274); J: Blattella germanica (ex cult.); K: Symploce pallens (ex cult.); L: Xestoblatta cantralli (Costa Rica, CR 15); M: Ischnoptera sp. (Costa Rica, CR 13 / 1). - Abbreviations and symbols: sc scapus; pc pedicellus; me meriston (first flagellomere of current stage); + new proximal flagellomeres generated by a division of the meriston into two or three flagellomeres during the preceding intermoult period in B, E, H, in C, D the division had presumably already occurred one moult earlier; L - early larval stage; L + intermediate or late larval stage; Im imago.	Figure 2. A-I: Antennae of Attaphila species of various age. A, H, I: Presumably second stage larva (head width 0.52 mm), with enlarged details of the proximal (H) and distal regions (I, distal end to the right), both from (A); B - E: intermediate and late larval stages, head width 0.66 (B), 0.61 (C), 0.76 (D), 0.73 mm (E); F, G: imagines. J-M: Imaginal antennae of Blattellidae species, in (J) showing parts of the proximal, intermediate and distal region, in (K-M) only of the proximal and distal region. - Species: A, C, E, H, I: Attaphila paucisetosa (A, H, I: Colombia, Cb 4 / 3; C: Bo 1458; E: Bo 1433); B, D, F: A. aptera (B: Bo 1457; D: Bo 1292; F: Bo 1225); G: A. bergi (Bo 1274); J: Blattella germanica (ex cult.); K: Symploce pallens (ex cult.); L: Xestoblatta cantralli (Costa Rica, CR 15); M: Ischnoptera sp. (Costa Rica, CR 13 / 1). - Abbreviations and symbols: sc scapus; pc pedicellus; me meriston (first flagellomere of current stage); + new proximal flagellomeres generated by a division of the meriston into two or three flagellomeres during the preceding intermoult period in B, E, H, in C, D the division had presumably already occurred one moult earlier; L - early larval stage; L + intermediate or late larval stage; Im imago.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
4A668C721AD95502974FA7553955584A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563272	https://doi.org/10.3897/asp.79.e67569.figure3(a-f)	Figure 3 (A-J). Tegmina (forewings) of males of Attaphila species, dorsal views, left tegmen at the left. - Species: A, B: A. aptera (Bo 1252); C, D: A. bergi (Bo 1443); E, F: A. flava (HT Bo 1280); G, H: A. fungicola (Bo 1229); I, J: A. paucisetosa (Bo 1441). - Abbreviation: fwar forewing articulation area. In A borders specified as addressed in the text.	Figure 3 (A-J). Tegmina (forewings) of males of Attaphila species, dorsal views, left tegmen at the left. - Species: A, B: A. aptera (Bo 1252); C, D: A. bergi (Bo 1443); E, F: A. flava (HT Bo 1280); G, H: A. fungicola (Bo 1229); I, J: A. paucisetosa (Bo 1441). - Abbreviation: fwar forewing articulation area. In A borders specified as addressed in the text.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
4A668C721AD95502974FA7553955584A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563274	https://doi.org/10.3897/asp.79.e67569.figure5	Figure 5. A - I: Tergite specialisations of the males of Attaphila species, medially on T 1 (A - D), or laterally at the anterior border of T 2 (E - I), in E-G from right side of tergite, in H and I from left side. - Species: A, B: A. aptera (Bo 1227, HT Bo 1258); C, D: A. bergi (Bo 1283, Bo 1274); E, F: A. paucisetosa (PT Bo 1254, Bo 1256); G, H: A. fungicola (LT Bo 1265); I: A. flava (HT Bo 1280). - Abbreviations: T 1 - a, T 2 - a anterior borders of tergites T 1,2; T 1 - p posterior border of T 1; tr 1, tr 2 transversal ridges of T 1,2; msa 1 anterior median specialisation of tergite T 1; msl 2 lateral specialisation of tergite T 2; msp 1 posterior median specialisation of tergite T 1; pg pores of glands, tiny in A, B, between T 1 - a and tr 1, larger in E as part of msl 2 specialisations. - J, K: Paraprocts, tergite 10 and cerci in ventral view, of J: A. aptera (Bo 1224) and K: A. paucisetosa (HT Bo 1258). - Abbreviations and arrows: Ca sclerite at mesal base of cercus; ce cercus; hmp hook-like mesal projection of paraproct; PPl, PPr left and right paraproct; T 9 tergite 9; T 10 tergite 10 with its ventrally bent lateral (paratergal) parts T 10 p (larger median parts of T 10 out of focus); white arrows: articulation between mesal end of T 10 p and lateral end of paraproct.	Figure 5. A - I: Tergite specialisations of the males of Attaphila species, medially on T 1 (A - D), or laterally at the anterior border of T 2 (E - I), in E-G from right side of tergite, in H and I from left side. - Species: A, B: A. aptera (Bo 1227, HT Bo 1258); C, D: A. bergi (Bo 1283, Bo 1274); E, F: A. paucisetosa (PT Bo 1254, Bo 1256); G, H: A. fungicola (LT Bo 1265); I: A. flava (HT Bo 1280). - Abbreviations: T 1 - a, T 2 - a anterior borders of tergites T 1,2; T 1 - p posterior border of T 1; tr 1, tr 2 transversal ridges of T 1,2; msa 1 anterior median specialisation of tergite T 1; msl 2 lateral specialisation of tergite T 2; msp 1 posterior median specialisation of tergite T 1; pg pores of glands, tiny in A, B, between T 1 - a and tr 1, larger in E as part of msl 2 specialisations. - J, K: Paraprocts, tergite 10 and cerci in ventral view, of J: A. aptera (Bo 1224) and K: A. paucisetosa (HT Bo 1258). - Abbreviations and arrows: Ca sclerite at mesal base of cercus; ce cercus; hmp hook-like mesal projection of paraproct; PPl, PPr left and right paraproct; T 9 tergite 9; T 10 tergite 10 with its ventrally bent lateral (paratergal) parts T 10 p (larger median parts of T 10 out of focus); white arrows: articulation between mesal end of T 10 p and lateral end of paraproct.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
4A668C721AD95502974FA7553955584A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563276	https://doi.org/10.3897/asp.79.e67569.figure7	Figure 7. Abdominal tergites and terminalia of Attaphila bergi. A - C: Male (Bo 1274). A: T 1 - 5 (see also Fig. S 1 C), T 1 without long bristles, the median specialisation, around the median scratch, is hardly visible (see Fig. 5 D); B: T 6,7; C: Terminalia, ventral view, with T 8 - 10, cerci, and paraprocts. D, E: Female (Bo 1282). D: T 2 - 5; E: T 6,7. - Abbreviations: ce cercus; ltga 9 lateral tergal apodeme of tergite T 9; msa 1 anterior median specialisation of T 1; PPr right paraproct; T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered).	Figure 7. Abdominal tergites and terminalia of Attaphila bergi. A - C: Male (Bo 1274). A: T 1 - 5 (see also Fig. S 1 C), T 1 without long bristles, the median specialisation, around the median scratch, is hardly visible (see Fig. 5 D); B: T 6,7; C: Terminalia, ventral view, with T 8 - 10, cerci, and paraprocts. D, E: Female (Bo 1282). D: T 2 - 5; E: T 6,7. - Abbreviations: ce cercus; ltga 9 lateral tergal apodeme of tergite T 9; msa 1 anterior median specialisation of T 1; PPr right paraproct; T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered).	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
4A668C721AD95502974FA7553955584A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563281	https://doi.org/10.3897/asp.79.e67569.figure12	Figure 12. Abdominal tergites T 6,7 of females of Attaphila species. A: A. aptera (Bo 1257). B: A. bergi (Bo 1282). C: A. fungicola (Bo 1264). D: A. multisetosa (HT Bo 1270). E: A. paucisetosa (PT Bo 1255). F: A. schuppi (ST Bo 1237), transversal ridge tr 7 almost completely missing, only tiny remnants present near the associated bristles (see Fig. 14 F). - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 12. Abdominal tergites T 6,7 of females of Attaphila species. A: A. aptera (Bo 1257). B: A. bergi (Bo 1282). C: A. fungicola (Bo 1264). D: A. multisetosa (HT Bo 1270). E: A. paucisetosa (PT Bo 1255). F: A. schuppi (ST Bo 1237), transversal ridge tr 7 almost completely missing, only tiny remnants present near the associated bristles (see Fig. 14 F). - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
4A668C721AD95502974FA7553955584A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563282	https://doi.org/10.3897/asp.79.e67569.figure13	Figure 13. Abdominal tergites T 6,7 of females (A, B) and males (C - F) of Attaphila species. A: A. sexdentis, female (Bo 1233). B: A. sinuosocarinata, female (HT Bo 1273). C: A. aptera, male (Bo 1256). D: A. bergi, male (Bo 1274), transversal ridge tr 7 weakly developed, sublaterally with a large gap. E: A. paucisetosa, male (Bo 1240), transversal ridge tr 7 missing. F: A. fungicola, male (Bo 1229), transversal ridge tr 7 missing. - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 13. Abdominal tergites T 6,7 of females (A, B) and males (C - F) of Attaphila species. A: A. sexdentis, female (Bo 1233). B: A. sinuosocarinata, female (HT Bo 1273). C: A. aptera, male (Bo 1256). D: A. bergi, male (Bo 1274), transversal ridge tr 7 weakly developed, sublaterally with a large gap. E: A. paucisetosa, male (Bo 1240), transversal ridge tr 7 missing. F: A. fungicola, male (Bo 1229), transversal ridge tr 7 missing. - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
4A668C721AD95502974FA7553955584A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563283	https://doi.org/10.3897/asp.79.e67569.figure14	Figure 14. Size and distribution of bristles in the median part of tergites T 6,7 in females (A-H) and males (I) of Attaphila species, phase contrast images. A: A. aptera, female (Bo 1257). B: A. bergi, female (Bo 1282). C: A. fungicola, female (Bo 1264). D: A. multisetosa, female (Bo 1270). E: A. paucisetosa, female (PT Bo 1255). F: A. schuppi, female (ST Bo 1237), with only short remnants of transversal ridge tr 7. G: A. sexdentis, female (Bo 1233). H: A. sinuosocarinata, female (HT Bo 1273). I: A. aptera, male (Bo 1256). - Abbreviations: T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 14. Size and distribution of bristles in the median part of tergites T 6,7 in females (A-H) and males (I) of Attaphila species, phase contrast images. A: A. aptera, female (Bo 1257). B: A. bergi, female (Bo 1282). C: A. fungicola, female (Bo 1264). D: A. multisetosa, female (Bo 1270). E: A. paucisetosa, female (PT Bo 1255). F: A. schuppi, female (ST Bo 1237), with only short remnants of transversal ridge tr 7. G: A. sexdentis, female (Bo 1233). H: A. sinuosocarinata, female (HT Bo 1273). I: A. aptera, male (Bo 1256). - Abbreviations: T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
4A668C721AD95502974FA7553955584A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563284	https://doi.org/10.3897/asp.79.e67569.figure15	Figure 15. Size and distribution of bristles in the median part of tergites T 6,7 in males (A - E) and in larval males and females (F - L) of Attaphila species, phase contrast images. A: A. bergi, male (Bo 1274). B: A. fungicola, male (LT Bo 1265). C: A. flava, male (HT Bo 1280). D: A. paucisetosa, male (Cb 2 / 1). E: A. paucisetosa, male, with slightly shorter bristles than in preceding specimen (Bo 1445). F: A. aptera, larval male (Bo 1291). G: A. aptera, larval female (Bo 1289). H: A. aptera, larval male (HT Bo 1232). I: A. bergi, larval male (Bo 1230); medium sized bristles belong to T 6, the very long ones are from T 5 covering most of T 6. J: A. bergi, larval female (Bo 1285). K: A. paucisetosa, larval male (Bo 1433). L: A. paucisetosa, larval female (Bo 1293). - Abbreviations: T 5 - p, T 6 - p, T 7 - p posterior borders of tergites T 5, T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 15. Size and distribution of bristles in the median part of tergites T 6,7 in males (A - E) and in larval males and females (F - L) of Attaphila species, phase contrast images. A: A. bergi, male (Bo 1274). B: A. fungicola, male (LT Bo 1265). C: A. flava, male (HT Bo 1280). D: A. paucisetosa, male (Cb 2 / 1). E: A. paucisetosa, male, with slightly shorter bristles than in preceding specimen (Bo 1445). F: A. aptera, larval male (Bo 1291). G: A. aptera, larval female (Bo 1289). H: A. aptera, larval male (HT Bo 1232). I: A. bergi, larval male (Bo 1230); medium sized bristles belong to T 6, the very long ones are from T 5 covering most of T 6. J: A. bergi, larval female (Bo 1285). K: A. paucisetosa, larval male (Bo 1433). L: A. paucisetosa, larval female (Bo 1293). - Abbreviations: T 5 - p, T 6 - p, T 7 - p posterior borders of tergites T 5, T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
4A668C721AD95502974FA7553955584A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563285	https://doi.org/10.3897/asp.79.e67569.figure16	Figure 16. Subgenital plate (sternite S 7) of females of Attaphila species. A, B: A. bergi (Bo 1275, Bo 1282). C, D: A. fungicola (Bo 1264, Bo 1228). E, F: A. paucisetosa (Cb 2 / 2, PT Bo 1255). G, H: A. schuppi (ST Bo 1237, ST Bo 1234). - Abbreviations: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars); vf 7 ventral fold of segment 7 = subgenital lobe (across entire width of S 7; approximate longitudinal extension given by double-headed arrow).	Figure 16. Subgenital plate (sternite S 7) of females of Attaphila species. A, B: A. bergi (Bo 1275, Bo 1282). C, D: A. fungicola (Bo 1264, Bo 1228). E, F: A. paucisetosa (Cb 2 / 2, PT Bo 1255). G, H: A. schuppi (ST Bo 1237, ST Bo 1234). - Abbreviations: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars); vf 7 ventral fold of segment 7 = subgenital lobe (across entire width of S 7; approximate longitudinal extension given by double-headed arrow).	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
4A668C721AD95502974FA7553955584A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563286	https://doi.org/10.3897/asp.79.e67569.figure17	Figure 17. Subgenital plate (sternite S 7) of females (A - F) and larval females (G, H) of Attaphila species. A, B: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). C, D: A. aptera (Bo 1225, Bo 1253). E: A. multisetosa (HT Bo 1270). F: A. sexdentis (HT Bo 1233). G: A. aptera, larval female (Bo 1289). H: A. bergi, larval female (Bo 1231). - Abbreviations and arrows: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars). Arrows in B, C, F pointing to lateral borders of S 7, which are parallel or slightly divergent (to the anterior) in A-E, but convergent in F.	Figure 17. Subgenital plate (sternite S 7) of females (A - F) and larval females (G, H) of Attaphila species. A, B: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). C, D: A. aptera (Bo 1225, Bo 1253). E: A. multisetosa (HT Bo 1270). F: A. sexdentis (HT Bo 1233). G: A. aptera, larval female (Bo 1289). H: A. bergi, larval female (Bo 1231). - Abbreviations and arrows: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars). Arrows in B, C, F pointing to lateral borders of S 7, which are parallel or slightly divergent (to the anterior) in A-E, but convergent in F.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
4A668C721AD95502974FA7553955584A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563287	https://doi.org/10.3897/asp.79.e67569.figure18	Figure 18. Anterior half of subgenital plate (sternite S 7) of females of Attaphila species, shape of sternal transversal ridge, phase contrast images (same objects as shown in Figs 16, 17, except for Bo 1236 in F). A, B: A. aptera (Bo 1225, Bo 1253). C, D: A. bergi (Bo 1275, Bo 1282). E, F: A. fungicola (Bo 1264, Bo 1236). G: A. multisetosa (HT Bo 1270). H, I: A. paucisetosa (Cb 2 / 2, PT Bo 1255). J, K: A. schuppi (ST Bo 1234, ST Bo 1237). L: A. sexdentis (HT Bo 1233). M, N: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). - Abbreviations and symbols: sr 7 transversal ridge of subgenital plate S 7; short interruptions of this ridge indicated by arrows, larger gaps delimited by double bars.	Figure 18. Anterior half of subgenital plate (sternite S 7) of females of Attaphila species, shape of sternal transversal ridge, phase contrast images (same objects as shown in Figs 16, 17, except for Bo 1236 in F). A, B: A. aptera (Bo 1225, Bo 1253). C, D: A. bergi (Bo 1275, Bo 1282). E, F: A. fungicola (Bo 1264, Bo 1236). G: A. multisetosa (HT Bo 1270). H, I: A. paucisetosa (Cb 2 / 2, PT Bo 1255). J, K: A. schuppi (ST Bo 1234, ST Bo 1237). L: A. sexdentis (HT Bo 1233). M, N: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). - Abbreviations and symbols: sr 7 transversal ridge of subgenital plate S 7; short interruptions of this ridge indicated by arrows, larger gaps delimited by double bars.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
4A668C721AD95502974FA7553955584A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563288	https://doi.org/10.3897/asp.79.e67569.figure19	Figure 19. Terminalia with genitalia of females of Attaphila species, ventral view, posterior end on top. A: A. aptera (Bo 1253), without laterosternal shelf area; inset showing median part of genital chamber dorsal wall. B: A. bergi (Bo 1239), without laterosternal shelf area. C, D: A. fungicola (Bo 1236, Bo 1264), C with, D without laterosternal shelf area. - Abbreviations: aa anterior arch; ca central apodeme; ce cercus; gcf fold dividing genital chamber (posterior edge labeled); gcp pouch of genital chamber (one-sided); gg-m, gg-l mesal and lateral gonangulum sclerites, the latter present only in A. aptera; IC intercalary sclerite; ls-p posterolateral extremity of laterosternal-shelf sclerite; lst laterosternal-shelf tube; pl posterior lobe; PP paraproct; pt 8,9 extension of paratergite 9; sp spermathecal plate; T 9, T 10 abdominal tergites 9 and 10; T 10 - p posterior border of tergite 10; v 1, v 3 1 st and 3 rd valves of ovipositor; vlf valvifer of segment 8; A articulations, A 1 between gg-l and pt 8,9, A 2 between gg-m and pl, A 3 between gg-m and basal sclerotisation of v 1, A 5 between pt 8,9 and vlf. Arrow in A showing mesal border of putative weak sclerotisation in dorsal wall of genital chamber. (See Supplement 1 and Supplement 3 Fig. S 2 A, B, C for additional explanations and more complete labeling using different terminologies.)	Figure 19. Terminalia with genitalia of females of Attaphila species, ventral view, posterior end on top. A: A. aptera (Bo 1253), without laterosternal shelf area; inset showing median part of genital chamber dorsal wall. B: A. bergi (Bo 1239), without laterosternal shelf area. C, D: A. fungicola (Bo 1236, Bo 1264), C with, D without laterosternal shelf area. - Abbreviations: aa anterior arch; ca central apodeme; ce cercus; gcf fold dividing genital chamber (posterior edge labeled); gcp pouch of genital chamber (one-sided); gg-m, gg-l mesal and lateral gonangulum sclerites, the latter present only in A. aptera; IC intercalary sclerite; ls-p posterolateral extremity of laterosternal-shelf sclerite; lst laterosternal-shelf tube; pl posterior lobe; PP paraproct; pt 8,9 extension of paratergite 9; sp spermathecal plate; T 9, T 10 abdominal tergites 9 and 10; T 10 - p posterior border of tergite 10; v 1, v 3 1 st and 3 rd valves of ovipositor; vlf valvifer of segment 8; A articulations, A 1 between gg-l and pt 8,9, A 2 between gg-m and pl, A 3 between gg-m and basal sclerotisation of v 1, A 5 between pt 8,9 and vlf. Arrow in A showing mesal border of putative weak sclerotisation in dorsal wall of genital chamber. (See Supplement 1 and Supplement 3 Fig. S 2 A, B, C for additional explanations and more complete labeling using different terminologies.)	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
4A668C721AD95502974FA7553955584A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563291	https://doi.org/10.3897/asp.79.e67569.figure22	Figure 22. Laterosternal shelf area of females of Attaphila species, ventral view, posteriorly on top. A, B: A. aptera (Bo 1225, Bo 1257), in A left tube distorted (ventral part of tube squeezed laterally, as indicated by grey arrow). C, D: A. bergi (Bo 1282, Bo 1239). E, F: A. fungicola (Bo 1264, Bo 1447). G: A. multisetosa (HT Bo 1270). H: A. paucisetosa (PT Bo 1255). I: A. sexdentis (HT Bo 1233). J: A. schuppi (ST Bo 1237). K, L: A. sinuosocarinata (HT Bo 1273, PT Bo 1288). - Abbreviations: isf intersternal folds (of floor of vestibulum); ls laterosternal-shelf sclerite (with central part c, arm part a, wing part w, posterolateral extremity p); lst laterosternal-shelf tube; vfl floor of vestibulum. (See Supplement 3 Fig. S 2 D, E for more complete labeling using different terminologies.) - Arrows: Black arrows in A, D, I: twisting of sclerite at transition between ls-w and ls-p. Grey arrow in A indicates direction of squeezing of tube lst. Blue arrowheads: anterior extremity of wing part ls-w on mesal border of tube lst, ls-w either restricted to posterior half of tubes lst (e. g. K) or reaching far into anterior half (e. g. D). Red arrowheads: area where the posterior margin of ls-a bends dorsally, and where a bend or kink is often present in (inner) lateral border of tube lst (compare white arrowheads in Fig. 23 C, D, H, I, J). Black arrowheads: membranous inner lateral border of tube lst posterior to the bend or kink (compare Fig. 23 C, D, H, I and black arrowheads in sections beside D). - Inserts in B, D, E, K showing right (of picture and animal) tube lst with posterior margin of ls-a (red line; compare Fig. 23 C-E) and anterior margin of ls-w (blue line; compare Fig. 23 F, G).	Figure 22. Laterosternal shelf area of females of Attaphila species, ventral view, posteriorly on top. A, B: A. aptera (Bo 1225, Bo 1257), in A left tube distorted (ventral part of tube squeezed laterally, as indicated by grey arrow). C, D: A. bergi (Bo 1282, Bo 1239). E, F: A. fungicola (Bo 1264, Bo 1447). G: A. multisetosa (HT Bo 1270). H: A. paucisetosa (PT Bo 1255). I: A. sexdentis (HT Bo 1233). J: A. schuppi (ST Bo 1237). K, L: A. sinuosocarinata (HT Bo 1273, PT Bo 1288). - Abbreviations: isf intersternal folds (of floor of vestibulum); ls laterosternal-shelf sclerite (with central part c, arm part a, wing part w, posterolateral extremity p); lst laterosternal-shelf tube; vfl floor of vestibulum. (See Supplement 3 Fig. S 2 D, E for more complete labeling using different terminologies.) - Arrows: Black arrows in A, D, I: twisting of sclerite at transition between ls-w and ls-p. Grey arrow in A indicates direction of squeezing of tube lst. Blue arrowheads: anterior extremity of wing part ls-w on mesal border of tube lst, ls-w either restricted to posterior half of tubes lst (e. g. K) or reaching far into anterior half (e. g. D). Red arrowheads: area where the posterior margin of ls-a bends dorsally, and where a bend or kink is often present in (inner) lateral border of tube lst (compare white arrowheads in Fig. 23 C, D, H, I, J). Black arrowheads: membranous inner lateral border of tube lst posterior to the bend or kink (compare Fig. 23 C, D, H, I and black arrowheads in sections beside D). - Inserts in B, D, E, K showing right (of picture and animal) tube lst with posterior margin of ls-a (red line; compare Fig. 23 C-E) and anterior margin of ls-w (blue line; compare Fig. 23 F, G).	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
4A668C721AD95502974FA7553955584A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563293	https://doi.org/10.3897/asp.79.e67569.figure24	Figure 24. Subgenital plate (sternite S 9) and phallomeres of males of Attaphila species, dorsal view, anteriorly on top. A, B: A. aptera, subgenital plate with phallomeres (Bo 1256, Bo 1224), anterior part of R 3 sclerite out of frame in B. C-G: A. bergi. C: Subgenital plate without phallomeres (Bo 1274); D: Subgenital plate with phallomeres (Bo 1283); E: Isolated hook of left phallomere (Bo 1274); F: Isolated endophallic apodeme (damaged near midlength), virga, and psa-process of left phallomere (Bo 1274); G: Isolated right phallomere (Bo 1274). - Abbreviations: A articulations, A 1 between S 9 and pt 9, A 3 between sclerites R 3 and R 1 P; cs cleft sclerite, composed of the dorsally fused R 1 S and R 2, R 2 - part in contact with arm v of R 3 sclerite (not visible, contact area indicated by white arrow in G); ea endophallic apodeme; h hook (with long membranous base p, wide base b of sclerotised part, neck n, and claw cl); hcl cleft in wall of groove hge; hge groove upon claw part of hook; psa process; pt 9 extension of paratergite 9 contacting sternite 9; R 1 (divided in R 1 S and R 1 P), R 2, R 3 (with anterior arm a, dorsoposterior arm d, ventroposterior arm v) are the principal sclerotisations of the right phallomere; sll, slr left and right stylus; sta 9 anterior apodeme of subgenital plate S 9; vf 9 subgenital lobe; vge grooves along virga (see vge 1, vge 2 in Fig. 32 I); vi virga.	Figure 24. Subgenital plate (sternite S 9) and phallomeres of males of Attaphila species, dorsal view, anteriorly on top. A, B: A. aptera, subgenital plate with phallomeres (Bo 1256, Bo 1224), anterior part of R 3 sclerite out of frame in B. C-G: A. bergi. C: Subgenital plate without phallomeres (Bo 1274); D: Subgenital plate with phallomeres (Bo 1283); E: Isolated hook of left phallomere (Bo 1274); F: Isolated endophallic apodeme (damaged near midlength), virga, and psa-process of left phallomere (Bo 1274); G: Isolated right phallomere (Bo 1274). - Abbreviations: A articulations, A 1 between S 9 and pt 9, A 3 between sclerites R 3 and R 1 P; cs cleft sclerite, composed of the dorsally fused R 1 S and R 2, R 2 - part in contact with arm v of R 3 sclerite (not visible, contact area indicated by white arrow in G); ea endophallic apodeme; h hook (with long membranous base p, wide base b of sclerotised part, neck n, and claw cl); hcl cleft in wall of groove hge; hge groove upon claw part of hook; psa process; pt 9 extension of paratergite 9 contacting sternite 9; R 1 (divided in R 1 S and R 1 P), R 2, R 3 (with anterior arm a, dorsoposterior arm d, ventroposterior arm v) are the principal sclerotisations of the right phallomere; sll, slr left and right stylus; sta 9 anterior apodeme of subgenital plate S 9; vf 9 subgenital lobe; vge grooves along virga (see vge 1, vge 2 in Fig. 32 I); vi virga.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
4A668C721AD95502974FA7553955584A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563280	https://doi.org/10.3897/asp.79.e67569.figure11	Figure 11. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. schuppi (ST Bo 1237). A: T 2 - 5; B: T 6,7. C, D: A. sexdentis (HT Bo 1233). C: T 1 - 5; D: T 6,7. - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to median and lateral excurvations of transversal ridges to the anterior.	Figure 11. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. schuppi (ST Bo 1237). A: T 2 - 5; B: T 6,7. C, D: A. sexdentis (HT Bo 1233). C: T 1 - 5; D: T 6,7. - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to median and lateral excurvations of transversal ridges to the anterior.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
4A668C721AD95502974FA7553955584A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563278	https://doi.org/10.3897/asp.79.e67569.figure9	Figure 9. Abdominal tergites and terminalia of Attaphila paucisetosa. A - D: Male (HT Bo 1258). A: T 1; B: T 2 - 5, T 2 with lateral specialisations; C: T 6,7. D: Terminalia, ventral view, with T 9 (with lateral extension) and T 10, cerci, and paraprocts. E, F: Female (PT Bo 1255). E: T 1 - 5; F: T 6,7. - Abbreviations: ce cercus; msl 2 lateral specialisation of T 2; PPr right paraproct; T n tergite (numbered); pt 9 extension of paratergite of T 9; T n - a anterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to excurvations of male tr 2.	Figure 9. Abdominal tergites and terminalia of Attaphila paucisetosa. A - D: Male (HT Bo 1258). A: T 1; B: T 2 - 5, T 2 with lateral specialisations; C: T 6,7. D: Terminalia, ventral view, with T 9 (with lateral extension) and T 10, cerci, and paraprocts. E, F: Female (PT Bo 1255). E: T 1 - 5; F: T 6,7. - Abbreviations: ce cercus; msl 2 lateral specialisation of T 2; PPr right paraproct; T n tergite (numbered); pt 9 extension of paratergite of T 9; T n - a anterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to excurvations of male tr 2.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
3F54CA4203F85F8B9B2103DD37006B60.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563272	https://doi.org/10.3897/asp.79.e67569.figure3(a-f)	Figure 3 (A-J). Tegmina (forewings) of males of Attaphila species, dorsal views, left tegmen at the left. - Species: A, B: A. aptera (Bo 1252); C, D: A. bergi (Bo 1443); E, F: A. flava (HT Bo 1280); G, H: A. fungicola (Bo 1229); I, J: A. paucisetosa (Bo 1441). - Abbreviation: fwar forewing articulation area. In A borders specified as addressed in the text.	Figure 3 (A-J). Tegmina (forewings) of males of Attaphila species, dorsal views, left tegmen at the left. - Species: A, B: A. aptera (Bo 1252); C, D: A. bergi (Bo 1443); E, F: A. flava (HT Bo 1280); G, H: A. fungicola (Bo 1229); I, J: A. paucisetosa (Bo 1441). - Abbreviation: fwar forewing articulation area. In A borders specified as addressed in the text.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
3F54CA4203F85F8B9B2103DD37006B60.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563274	https://doi.org/10.3897/asp.79.e67569.figure5	Figure 5. A - I: Tergite specialisations of the males of Attaphila species, medially on T 1 (A - D), or laterally at the anterior border of T 2 (E - I), in E-G from right side of tergite, in H and I from left side. - Species: A, B: A. aptera (Bo 1227, HT Bo 1258); C, D: A. bergi (Bo 1283, Bo 1274); E, F: A. paucisetosa (PT Bo 1254, Bo 1256); G, H: A. fungicola (LT Bo 1265); I: A. flava (HT Bo 1280). - Abbreviations: T 1 - a, T 2 - a anterior borders of tergites T 1,2; T 1 - p posterior border of T 1; tr 1, tr 2 transversal ridges of T 1,2; msa 1 anterior median specialisation of tergite T 1; msl 2 lateral specialisation of tergite T 2; msp 1 posterior median specialisation of tergite T 1; pg pores of glands, tiny in A, B, between T 1 - a and tr 1, larger in E as part of msl 2 specialisations. - J, K: Paraprocts, tergite 10 and cerci in ventral view, of J: A. aptera (Bo 1224) and K: A. paucisetosa (HT Bo 1258). - Abbreviations and arrows: Ca sclerite at mesal base of cercus; ce cercus; hmp hook-like mesal projection of paraproct; PPl, PPr left and right paraproct; T 9 tergite 9; T 10 tergite 10 with its ventrally bent lateral (paratergal) parts T 10 p (larger median parts of T 10 out of focus); white arrows: articulation between mesal end of T 10 p and lateral end of paraproct.	Figure 5. A - I: Tergite specialisations of the males of Attaphila species, medially on T 1 (A - D), or laterally at the anterior border of T 2 (E - I), in E-G from right side of tergite, in H and I from left side. - Species: A, B: A. aptera (Bo 1227, HT Bo 1258); C, D: A. bergi (Bo 1283, Bo 1274); E, F: A. paucisetosa (PT Bo 1254, Bo 1256); G, H: A. fungicola (LT Bo 1265); I: A. flava (HT Bo 1280). - Abbreviations: T 1 - a, T 2 - a anterior borders of tergites T 1,2; T 1 - p posterior border of T 1; tr 1, tr 2 transversal ridges of T 1,2; msa 1 anterior median specialisation of tergite T 1; msl 2 lateral specialisation of tergite T 2; msp 1 posterior median specialisation of tergite T 1; pg pores of glands, tiny in A, B, between T 1 - a and tr 1, larger in E as part of msl 2 specialisations. - J, K: Paraprocts, tergite 10 and cerci in ventral view, of J: A. aptera (Bo 1224) and K: A. paucisetosa (HT Bo 1258). - Abbreviations and arrows: Ca sclerite at mesal base of cercus; ce cercus; hmp hook-like mesal projection of paraproct; PPl, PPr left and right paraproct; T 9 tergite 9; T 10 tergite 10 with its ventrally bent lateral (paratergal) parts T 10 p (larger median parts of T 10 out of focus); white arrows: articulation between mesal end of T 10 p and lateral end of paraproct.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
3F54CA4203F85F8B9B2103DD37006B60.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563277	https://doi.org/10.3897/asp.79.e67569.figure8	Figure 8. Abdominal tergites and terminalia of Attaphila species. A - C: A. fungicola. A, B: Male (Bo 1229). A: T 2 - 7, T 2 with lateral specialisations; B: Terminalia, ventral view, with T 9 (damaged: only right half) and T 10, cerci, and paraprocts. C: Female (Bo 1264), T 4 - 6. D: A. flava, male (HT Bo 1280), T 2 - 5, T 2 with lateral specialisations. - Abbreviations: ce cercus; ltga 9 lateral tergal apodeme of tergite T 9; msl 2 lateral specialisation of T 2; PPr right paraproct; T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to excurvations of male tr 2.	Figure 8. Abdominal tergites and terminalia of Attaphila species. A - C: A. fungicola. A, B: Male (Bo 1229). A: T 2 - 7, T 2 with lateral specialisations; B: Terminalia, ventral view, with T 9 (damaged: only right half) and T 10, cerci, and paraprocts. C: Female (Bo 1264), T 4 - 6. D: A. flava, male (HT Bo 1280), T 2 - 5, T 2 with lateral specialisations. - Abbreviations: ce cercus; ltga 9 lateral tergal apodeme of tergite T 9; msl 2 lateral specialisation of T 2; PPr right paraproct; T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to excurvations of male tr 2.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
3F54CA4203F85F8B9B2103DD37006B60.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563284	https://doi.org/10.3897/asp.79.e67569.figure15	Figure 15. Size and distribution of bristles in the median part of tergites T 6,7 in males (A - E) and in larval males and females (F - L) of Attaphila species, phase contrast images. A: A. bergi, male (Bo 1274). B: A. fungicola, male (LT Bo 1265). C: A. flava, male (HT Bo 1280). D: A. paucisetosa, male (Cb 2 / 1). E: A. paucisetosa, male, with slightly shorter bristles than in preceding specimen (Bo 1445). F: A. aptera, larval male (Bo 1291). G: A. aptera, larval female (Bo 1289). H: A. aptera, larval male (HT Bo 1232). I: A. bergi, larval male (Bo 1230); medium sized bristles belong to T 6, the very long ones are from T 5 covering most of T 6. J: A. bergi, larval female (Bo 1285). K: A. paucisetosa, larval male (Bo 1433). L: A. paucisetosa, larval female (Bo 1293). - Abbreviations: T 5 - p, T 6 - p, T 7 - p posterior borders of tergites T 5, T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 15. Size and distribution of bristles in the median part of tergites T 6,7 in males (A - E) and in larval males and females (F - L) of Attaphila species, phase contrast images. A: A. bergi, male (Bo 1274). B: A. fungicola, male (LT Bo 1265). C: A. flava, male (HT Bo 1280). D: A. paucisetosa, male (Cb 2 / 1). E: A. paucisetosa, male, with slightly shorter bristles than in preceding specimen (Bo 1445). F: A. aptera, larval male (Bo 1291). G: A. aptera, larval female (Bo 1289). H: A. aptera, larval male (HT Bo 1232). I: A. bergi, larval male (Bo 1230); medium sized bristles belong to T 6, the very long ones are from T 5 covering most of T 6. J: A. bergi, larval female (Bo 1285). K: A. paucisetosa, larval male (Bo 1433). L: A. paucisetosa, larval female (Bo 1293). - Abbreviations: T 5 - p, T 6 - p, T 7 - p posterior borders of tergites T 5, T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
3F54CA4203F85F8B9B2103DD37006B60.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563294	https://doi.org/10.3897/asp.79.e67569.figure25	Figure 25. Subgenital plate (sternite S 9) and phallomeres of males of Attaphila species, dorsal view, anteriorly on top. A, B: A. paucisetosa, subgenital plate with phallomeres (PT Bo 1254, Bo 1444), hook and right phallomere removed in B. C: A. flava, subgenital plate with phallomeres (HT Bo 1280). D, E: A. fungicola. D showing subgenital plate with phallomeres (LT Bo 1265), hook not seen in profile; E showing isolated hook seen in profile (Bo 1229). - Abbreviations: T 9 p ventrally bent lateral (paratergal) part of tergite 9; otherwise as for Fig. 24.	Figure 25. Subgenital plate (sternite S 9) and phallomeres of males of Attaphila species, dorsal view, anteriorly on top. A, B: A. paucisetosa, subgenital plate with phallomeres (PT Bo 1254, Bo 1444), hook and right phallomere removed in B. C: A. flava, subgenital plate with phallomeres (HT Bo 1280). D, E: A. fungicola. D showing subgenital plate with phallomeres (LT Bo 1265), hook not seen in profile; E showing isolated hook seen in profile (Bo 1229). - Abbreviations: T 9 p ventrally bent lateral (paratergal) part of tergite 9; otherwise as for Fig. 24.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
3F54CA4203F85F8B9B2103DD37006B60.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563295	https://doi.org/10.3897/asp.79.e67569.figure26	Figure 26. Subgenital plate (sternite S 9) and phallomeres of males of Attaphila species, phase contrast images of the same objects as shown in Fig. 25 A, C, D, anteriorly on top. A: A. flava (HT Bo 1280). B: A. fungicola (LT Bo 1265). C: A. paucisetosa (PT Bo 1254). - Abbreviations: A 1 articulation between S 9 and pt 9; cs cleft sclerite of right phallomere; ea endophallic apodeme; h hook; psa process; pt 9 extension of paratergite 9 contacting sternite 9; R 3 anterior principal sclerite of right phallomere; sll left stylus; sta 9 anterior apodeme of subgenital plate S 9; vf 9 subgenital lobe; vi virga. Arrow: in A indicating that the distal part of the subgenital lobe is broken off.	Figure 26. Subgenital plate (sternite S 9) and phallomeres of males of Attaphila species, phase contrast images of the same objects as shown in Fig. 25 A, C, D, anteriorly on top. A: A. flava (HT Bo 1280). B: A. fungicola (LT Bo 1265). C: A. paucisetosa (PT Bo 1254). - Abbreviations: A 1 articulation between S 9 and pt 9; cs cleft sclerite of right phallomere; ea endophallic apodeme; h hook; psa process; pt 9 extension of paratergite 9 contacting sternite 9; R 3 anterior principal sclerite of right phallomere; sll left stylus; sta 9 anterior apodeme of subgenital plate S 9; vf 9 subgenital lobe; vi virga. Arrow: in A indicating that the distal part of the subgenital lobe is broken off.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
3F54CA4203F85F8B9B2103DD37006B60.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563280	https://doi.org/10.3897/asp.79.e67569.figure11	Figure 11. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. schuppi (ST Bo 1237). A: T 2 - 5; B: T 6,7. C, D: A. sexdentis (HT Bo 1233). C: T 1 - 5; D: T 6,7. - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to median and lateral excurvations of transversal ridges to the anterior.	Figure 11. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. schuppi (ST Bo 1237). A: T 2 - 5; B: T 6,7. C, D: A. sexdentis (HT Bo 1233). C: T 1 - 5; D: T 6,7. - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to median and lateral excurvations of transversal ridges to the anterior.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
90567B49330A5CAFBEFA87D6C9B05DA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563270	https://doi.org/10.3897/asp.79.e67569.figure1	Figure 1. A - D: Head structures of Attaphila species, head made transparent by treatment with KOH. A: Entire head with focus on antennae and circumantennal pit (inset: median part, darkened). B: Entire head with focus on mouthparts (cardo and stipes seen behind circumantennal pit; inset: scheme of mandibular dentition in anterior view, mdl = left, mdr = right, elements above transversal line belonging to mola). C: Entire head with left antenna rectangularly bent between scapus and pedicellus. In A-C different levels of tentorium visible internally. D: Base of right antenna and surrounding parts of head capsule at higher magnification, also showing dorsal excavation at distal end of scapus allowing the strong bend shown in C. E - H: Thoracic nota of Attaphila species. E: All thoracic nota of a female, stretched. F: Pronotum of a male. G, H: Right part of metanotum of males with the strongly reduced hindwing. - Species: A - C: A. aptera, male (A, B: Bo 1224), larval female (C: Bo 1243); D: A. fungicola, male (Bo 1229); E, G: A. bergi, female (Bo 1239), male (Bo 1283); F, H: A. paucisetosa, male (Bo 1240). - Abbreviations: ata anterior tentorial arm; atb anterior tentorial bridge; atp anterior tentorial pit; car cardo of maxilla; cap circumantennal pit (with sharp dorsal edge capd visible and bottom capb of the pit shining through); cc coronal cleavage line; cpe compound eye; ct corpotentorium; fc frontal cleavage line; fl fagellum (all antennomeres following pedicellus, numbered from base as fl 1 etc.); hw hindwing; hwar level of (reduced) hindwing articulation; li lacinia; lp labial palpus (I-III: palpomeres); md mandible (left mdl, right mdr; I-IV: tip and incisivi); mo mola of mandible; mp maxillary palpus (I-V: palpomeres); N thoracic notum (N 1 pro-, N 2 meso-, N 3 metanotum); pc pedicellus; pta posterior tentorial arm; sc scapus; sc-pc scapus-pedicellus articulation; scex dorsal excavation at distal end of scapus; s-chA field of sensilla chaetica A; s-cpf row of sensilla campaniformia; sti stipes of maxilla; tp tentorial perforation.	Figure 1. A - D: Head structures of Attaphila species, head made transparent by treatment with KOH. A: Entire head with focus on antennae and circumantennal pit (inset: median part, darkened). B: Entire head with focus on mouthparts (cardo and stipes seen behind circumantennal pit; inset: scheme of mandibular dentition in anterior view, mdl = left, mdr = right, elements above transversal line belonging to mola). C: Entire head with left antenna rectangularly bent between scapus and pedicellus. In A-C different levels of tentorium visible internally. D: Base of right antenna and surrounding parts of head capsule at higher magnification, also showing dorsal excavation at distal end of scapus allowing the strong bend shown in C. E - H: Thoracic nota of Attaphila species. E: All thoracic nota of a female, stretched. F: Pronotum of a male. G, H: Right part of metanotum of males with the strongly reduced hindwing. - Species: A - C: A. aptera, male (A, B: Bo 1224), larval female (C: Bo 1243); D: A. fungicola, male (Bo 1229); E, G: A. bergi, female (Bo 1239), male (Bo 1283); F, H: A. paucisetosa, male (Bo 1240). - Abbreviations: ata anterior tentorial arm; atb anterior tentorial bridge; atp anterior tentorial pit; car cardo of maxilla; cap circumantennal pit (with sharp dorsal edge capd visible and bottom capb of the pit shining through); cc coronal cleavage line; cpe compound eye; ct corpotentorium; fc frontal cleavage line; fl fagellum (all antennomeres following pedicellus, numbered from base as fl 1 etc.); hw hindwing; hwar level of (reduced) hindwing articulation; li lacinia; lp labial palpus (I-III: palpomeres); md mandible (left mdl, right mdr; I-IV: tip and incisivi); mo mola of mandible; mp maxillary palpus (I-V: palpomeres); N thoracic notum (N 1 pro-, N 2 meso-, N 3 metanotum); pc pedicellus; pta posterior tentorial arm; sc scapus; sc-pc scapus-pedicellus articulation; scex dorsal excavation at distal end of scapus; s-chA field of sensilla chaetica A; s-cpf row of sensilla campaniformia; sti stipes of maxilla; tp tentorial perforation.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
90567B49330A5CAFBEFA87D6C9B05DA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563272	https://doi.org/10.3897/asp.79.e67569.figure3(a-f)	Figure 3 (A-J). Tegmina (forewings) of males of Attaphila species, dorsal views, left tegmen at the left. - Species: A, B: A. aptera (Bo 1252); C, D: A. bergi (Bo 1443); E, F: A. flava (HT Bo 1280); G, H: A. fungicola (Bo 1229); I, J: A. paucisetosa (Bo 1441). - Abbreviation: fwar forewing articulation area. In A borders specified as addressed in the text.	Figure 3 (A-J). Tegmina (forewings) of males of Attaphila species, dorsal views, left tegmen at the left. - Species: A, B: A. aptera (Bo 1252); C, D: A. bergi (Bo 1443); E, F: A. flava (HT Bo 1280); G, H: A. fungicola (Bo 1229); I, J: A. paucisetosa (Bo 1441). - Abbreviation: fwar forewing articulation area. In A borders specified as addressed in the text.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
90567B49330A5CAFBEFA87D6C9B05DA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563274	https://doi.org/10.3897/asp.79.e67569.figure5	Figure 5. A - I: Tergite specialisations of the males of Attaphila species, medially on T 1 (A - D), or laterally at the anterior border of T 2 (E - I), in E-G from right side of tergite, in H and I from left side. - Species: A, B: A. aptera (Bo 1227, HT Bo 1258); C, D: A. bergi (Bo 1283, Bo 1274); E, F: A. paucisetosa (PT Bo 1254, Bo 1256); G, H: A. fungicola (LT Bo 1265); I: A. flava (HT Bo 1280). - Abbreviations: T 1 - a, T 2 - a anterior borders of tergites T 1,2; T 1 - p posterior border of T 1; tr 1, tr 2 transversal ridges of T 1,2; msa 1 anterior median specialisation of tergite T 1; msl 2 lateral specialisation of tergite T 2; msp 1 posterior median specialisation of tergite T 1; pg pores of glands, tiny in A, B, between T 1 - a and tr 1, larger in E as part of msl 2 specialisations. - J, K: Paraprocts, tergite 10 and cerci in ventral view, of J: A. aptera (Bo 1224) and K: A. paucisetosa (HT Bo 1258). - Abbreviations and arrows: Ca sclerite at mesal base of cercus; ce cercus; hmp hook-like mesal projection of paraproct; PPl, PPr left and right paraproct; T 9 tergite 9; T 10 tergite 10 with its ventrally bent lateral (paratergal) parts T 10 p (larger median parts of T 10 out of focus); white arrows: articulation between mesal end of T 10 p and lateral end of paraproct.	Figure 5. A - I: Tergite specialisations of the males of Attaphila species, medially on T 1 (A - D), or laterally at the anterior border of T 2 (E - I), in E-G from right side of tergite, in H and I from left side. - Species: A, B: A. aptera (Bo 1227, HT Bo 1258); C, D: A. bergi (Bo 1283, Bo 1274); E, F: A. paucisetosa (PT Bo 1254, Bo 1256); G, H: A. fungicola (LT Bo 1265); I: A. flava (HT Bo 1280). - Abbreviations: T 1 - a, T 2 - a anterior borders of tergites T 1,2; T 1 - p posterior border of T 1; tr 1, tr 2 transversal ridges of T 1,2; msa 1 anterior median specialisation of tergite T 1; msl 2 lateral specialisation of tergite T 2; msp 1 posterior median specialisation of tergite T 1; pg pores of glands, tiny in A, B, between T 1 - a and tr 1, larger in E as part of msl 2 specialisations. - J, K: Paraprocts, tergite 10 and cerci in ventral view, of J: A. aptera (Bo 1224) and K: A. paucisetosa (HT Bo 1258). - Abbreviations and arrows: Ca sclerite at mesal base of cercus; ce cercus; hmp hook-like mesal projection of paraproct; PPl, PPr left and right paraproct; T 9 tergite 9; T 10 tergite 10 with its ventrally bent lateral (paratergal) parts T 10 p (larger median parts of T 10 out of focus); white arrows: articulation between mesal end of T 10 p and lateral end of paraproct.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
90567B49330A5CAFBEFA87D6C9B05DA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563277	https://doi.org/10.3897/asp.79.e67569.figure8	Figure 8. Abdominal tergites and terminalia of Attaphila species. A - C: A. fungicola. A, B: Male (Bo 1229). A: T 2 - 7, T 2 with lateral specialisations; B: Terminalia, ventral view, with T 9 (damaged: only right half) and T 10, cerci, and paraprocts. C: Female (Bo 1264), T 4 - 6. D: A. flava, male (HT Bo 1280), T 2 - 5, T 2 with lateral specialisations. - Abbreviations: ce cercus; ltga 9 lateral tergal apodeme of tergite T 9; msl 2 lateral specialisation of T 2; PPr right paraproct; T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to excurvations of male tr 2.	Figure 8. Abdominal tergites and terminalia of Attaphila species. A - C: A. fungicola. A, B: Male (Bo 1229). A: T 2 - 7, T 2 with lateral specialisations; B: Terminalia, ventral view, with T 9 (damaged: only right half) and T 10, cerci, and paraprocts. C: Female (Bo 1264), T 4 - 6. D: A. flava, male (HT Bo 1280), T 2 - 5, T 2 with lateral specialisations. - Abbreviations: ce cercus; ltga 9 lateral tergal apodeme of tergite T 9; msl 2 lateral specialisation of T 2; PPr right paraproct; T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to excurvations of male tr 2.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
90567B49330A5CAFBEFA87D6C9B05DA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563281	https://doi.org/10.3897/asp.79.e67569.figure12	Figure 12. Abdominal tergites T 6,7 of females of Attaphila species. A: A. aptera (Bo 1257). B: A. bergi (Bo 1282). C: A. fungicola (Bo 1264). D: A. multisetosa (HT Bo 1270). E: A. paucisetosa (PT Bo 1255). F: A. schuppi (ST Bo 1237), transversal ridge tr 7 almost completely missing, only tiny remnants present near the associated bristles (see Fig. 14 F). - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 12. Abdominal tergites T 6,7 of females of Attaphila species. A: A. aptera (Bo 1257). B: A. bergi (Bo 1282). C: A. fungicola (Bo 1264). D: A. multisetosa (HT Bo 1270). E: A. paucisetosa (PT Bo 1255). F: A. schuppi (ST Bo 1237), transversal ridge tr 7 almost completely missing, only tiny remnants present near the associated bristles (see Fig. 14 F). - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
90567B49330A5CAFBEFA87D6C9B05DA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563282	https://doi.org/10.3897/asp.79.e67569.figure13	Figure 13. Abdominal tergites T 6,7 of females (A, B) and males (C - F) of Attaphila species. A: A. sexdentis, female (Bo 1233). B: A. sinuosocarinata, female (HT Bo 1273). C: A. aptera, male (Bo 1256). D: A. bergi, male (Bo 1274), transversal ridge tr 7 weakly developed, sublaterally with a large gap. E: A. paucisetosa, male (Bo 1240), transversal ridge tr 7 missing. F: A. fungicola, male (Bo 1229), transversal ridge tr 7 missing. - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 13. Abdominal tergites T 6,7 of females (A, B) and males (C - F) of Attaphila species. A: A. sexdentis, female (Bo 1233). B: A. sinuosocarinata, female (HT Bo 1273). C: A. aptera, male (Bo 1256). D: A. bergi, male (Bo 1274), transversal ridge tr 7 weakly developed, sublaterally with a large gap. E: A. paucisetosa, male (Bo 1240), transversal ridge tr 7 missing. F: A. fungicola, male (Bo 1229), transversal ridge tr 7 missing. - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
90567B49330A5CAFBEFA87D6C9B05DA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563283	https://doi.org/10.3897/asp.79.e67569.figure14	Figure 14. Size and distribution of bristles in the median part of tergites T 6,7 in females (A-H) and males (I) of Attaphila species, phase contrast images. A: A. aptera, female (Bo 1257). B: A. bergi, female (Bo 1282). C: A. fungicola, female (Bo 1264). D: A. multisetosa, female (Bo 1270). E: A. paucisetosa, female (PT Bo 1255). F: A. schuppi, female (ST Bo 1237), with only short remnants of transversal ridge tr 7. G: A. sexdentis, female (Bo 1233). H: A. sinuosocarinata, female (HT Bo 1273). I: A. aptera, male (Bo 1256). - Abbreviations: T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 14. Size and distribution of bristles in the median part of tergites T 6,7 in females (A-H) and males (I) of Attaphila species, phase contrast images. A: A. aptera, female (Bo 1257). B: A. bergi, female (Bo 1282). C: A. fungicola, female (Bo 1264). D: A. multisetosa, female (Bo 1270). E: A. paucisetosa, female (PT Bo 1255). F: A. schuppi, female (ST Bo 1237), with only short remnants of transversal ridge tr 7. G: A. sexdentis, female (Bo 1233). H: A. sinuosocarinata, female (HT Bo 1273). I: A. aptera, male (Bo 1256). - Abbreviations: T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
90567B49330A5CAFBEFA87D6C9B05DA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563284	https://doi.org/10.3897/asp.79.e67569.figure15	Figure 15. Size and distribution of bristles in the median part of tergites T 6,7 in males (A - E) and in larval males and females (F - L) of Attaphila species, phase contrast images. A: A. bergi, male (Bo 1274). B: A. fungicola, male (LT Bo 1265). C: A. flava, male (HT Bo 1280). D: A. paucisetosa, male (Cb 2 / 1). E: A. paucisetosa, male, with slightly shorter bristles than in preceding specimen (Bo 1445). F: A. aptera, larval male (Bo 1291). G: A. aptera, larval female (Bo 1289). H: A. aptera, larval male (HT Bo 1232). I: A. bergi, larval male (Bo 1230); medium sized bristles belong to T 6, the very long ones are from T 5 covering most of T 6. J: A. bergi, larval female (Bo 1285). K: A. paucisetosa, larval male (Bo 1433). L: A. paucisetosa, larval female (Bo 1293). - Abbreviations: T 5 - p, T 6 - p, T 7 - p posterior borders of tergites T 5, T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 15. Size and distribution of bristles in the median part of tergites T 6,7 in males (A - E) and in larval males and females (F - L) of Attaphila species, phase contrast images. A: A. bergi, male (Bo 1274). B: A. fungicola, male (LT Bo 1265). C: A. flava, male (HT Bo 1280). D: A. paucisetosa, male (Cb 2 / 1). E: A. paucisetosa, male, with slightly shorter bristles than in preceding specimen (Bo 1445). F: A. aptera, larval male (Bo 1291). G: A. aptera, larval female (Bo 1289). H: A. aptera, larval male (HT Bo 1232). I: A. bergi, larval male (Bo 1230); medium sized bristles belong to T 6, the very long ones are from T 5 covering most of T 6. J: A. bergi, larval female (Bo 1285). K: A. paucisetosa, larval male (Bo 1433). L: A. paucisetosa, larval female (Bo 1293). - Abbreviations: T 5 - p, T 6 - p, T 7 - p posterior borders of tergites T 5, T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
90567B49330A5CAFBEFA87D6C9B05DA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563285	https://doi.org/10.3897/asp.79.e67569.figure16	Figure 16. Subgenital plate (sternite S 7) of females of Attaphila species. A, B: A. bergi (Bo 1275, Bo 1282). C, D: A. fungicola (Bo 1264, Bo 1228). E, F: A. paucisetosa (Cb 2 / 2, PT Bo 1255). G, H: A. schuppi (ST Bo 1237, ST Bo 1234). - Abbreviations: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars); vf 7 ventral fold of segment 7 = subgenital lobe (across entire width of S 7; approximate longitudinal extension given by double-headed arrow).	Figure 16. Subgenital plate (sternite S 7) of females of Attaphila species. A, B: A. bergi (Bo 1275, Bo 1282). C, D: A. fungicola (Bo 1264, Bo 1228). E, F: A. paucisetosa (Cb 2 / 2, PT Bo 1255). G, H: A. schuppi (ST Bo 1237, ST Bo 1234). - Abbreviations: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars); vf 7 ventral fold of segment 7 = subgenital lobe (across entire width of S 7; approximate longitudinal extension given by double-headed arrow).	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
90567B49330A5CAFBEFA87D6C9B05DA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563287	https://doi.org/10.3897/asp.79.e67569.figure18	Figure 18. Anterior half of subgenital plate (sternite S 7) of females of Attaphila species, shape of sternal transversal ridge, phase contrast images (same objects as shown in Figs 16, 17, except for Bo 1236 in F). A, B: A. aptera (Bo 1225, Bo 1253). C, D: A. bergi (Bo 1275, Bo 1282). E, F: A. fungicola (Bo 1264, Bo 1236). G: A. multisetosa (HT Bo 1270). H, I: A. paucisetosa (Cb 2 / 2, PT Bo 1255). J, K: A. schuppi (ST Bo 1234, ST Bo 1237). L: A. sexdentis (HT Bo 1233). M, N: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). - Abbreviations and symbols: sr 7 transversal ridge of subgenital plate S 7; short interruptions of this ridge indicated by arrows, larger gaps delimited by double bars.	Figure 18. Anterior half of subgenital plate (sternite S 7) of females of Attaphila species, shape of sternal transversal ridge, phase contrast images (same objects as shown in Figs 16, 17, except for Bo 1236 in F). A, B: A. aptera (Bo 1225, Bo 1253). C, D: A. bergi (Bo 1275, Bo 1282). E, F: A. fungicola (Bo 1264, Bo 1236). G: A. multisetosa (HT Bo 1270). H, I: A. paucisetosa (Cb 2 / 2, PT Bo 1255). J, K: A. schuppi (ST Bo 1234, ST Bo 1237). L: A. sexdentis (HT Bo 1233). M, N: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). - Abbreviations and symbols: sr 7 transversal ridge of subgenital plate S 7; short interruptions of this ridge indicated by arrows, larger gaps delimited by double bars.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
90567B49330A5CAFBEFA87D6C9B05DA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563288	https://doi.org/10.3897/asp.79.e67569.figure19	Figure 19. Terminalia with genitalia of females of Attaphila species, ventral view, posterior end on top. A: A. aptera (Bo 1253), without laterosternal shelf area; inset showing median part of genital chamber dorsal wall. B: A. bergi (Bo 1239), without laterosternal shelf area. C, D: A. fungicola (Bo 1236, Bo 1264), C with, D without laterosternal shelf area. - Abbreviations: aa anterior arch; ca central apodeme; ce cercus; gcf fold dividing genital chamber (posterior edge labeled); gcp pouch of genital chamber (one-sided); gg-m, gg-l mesal and lateral gonangulum sclerites, the latter present only in A. aptera; IC intercalary sclerite; ls-p posterolateral extremity of laterosternal-shelf sclerite; lst laterosternal-shelf tube; pl posterior lobe; PP paraproct; pt 8,9 extension of paratergite 9; sp spermathecal plate; T 9, T 10 abdominal tergites 9 and 10; T 10 - p posterior border of tergite 10; v 1, v 3 1 st and 3 rd valves of ovipositor; vlf valvifer of segment 8; A articulations, A 1 between gg-l and pt 8,9, A 2 between gg-m and pl, A 3 between gg-m and basal sclerotisation of v 1, A 5 between pt 8,9 and vlf. Arrow in A showing mesal border of putative weak sclerotisation in dorsal wall of genital chamber. (See Supplement 1 and Supplement 3 Fig. S 2 A, B, C for additional explanations and more complete labeling using different terminologies.)	Figure 19. Terminalia with genitalia of females of Attaphila species, ventral view, posterior end on top. A: A. aptera (Bo 1253), without laterosternal shelf area; inset showing median part of genital chamber dorsal wall. B: A. bergi (Bo 1239), without laterosternal shelf area. C, D: A. fungicola (Bo 1236, Bo 1264), C with, D without laterosternal shelf area. - Abbreviations: aa anterior arch; ca central apodeme; ce cercus; gcf fold dividing genital chamber (posterior edge labeled); gcp pouch of genital chamber (one-sided); gg-m, gg-l mesal and lateral gonangulum sclerites, the latter present only in A. aptera; IC intercalary sclerite; ls-p posterolateral extremity of laterosternal-shelf sclerite; lst laterosternal-shelf tube; pl posterior lobe; PP paraproct; pt 8,9 extension of paratergite 9; sp spermathecal plate; T 9, T 10 abdominal tergites 9 and 10; T 10 - p posterior border of tergite 10; v 1, v 3 1 st and 3 rd valves of ovipositor; vlf valvifer of segment 8; A articulations, A 1 between gg-l and pt 8,9, A 2 between gg-m and pl, A 3 between gg-m and basal sclerotisation of v 1, A 5 between pt 8,9 and vlf. Arrow in A showing mesal border of putative weak sclerotisation in dorsal wall of genital chamber. (See Supplement 1 and Supplement 3 Fig. S 2 A, B, C for additional explanations and more complete labeling using different terminologies.)	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
90567B49330A5CAFBEFA87D6C9B05DA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563291	https://doi.org/10.3897/asp.79.e67569.figure22	Figure 22. Laterosternal shelf area of females of Attaphila species, ventral view, posteriorly on top. A, B: A. aptera (Bo 1225, Bo 1257), in A left tube distorted (ventral part of tube squeezed laterally, as indicated by grey arrow). C, D: A. bergi (Bo 1282, Bo 1239). E, F: A. fungicola (Bo 1264, Bo 1447). G: A. multisetosa (HT Bo 1270). H: A. paucisetosa (PT Bo 1255). I: A. sexdentis (HT Bo 1233). J: A. schuppi (ST Bo 1237). K, L: A. sinuosocarinata (HT Bo 1273, PT Bo 1288). - Abbreviations: isf intersternal folds (of floor of vestibulum); ls laterosternal-shelf sclerite (with central part c, arm part a, wing part w, posterolateral extremity p); lst laterosternal-shelf tube; vfl floor of vestibulum. (See Supplement 3 Fig. S 2 D, E for more complete labeling using different terminologies.) - Arrows: Black arrows in A, D, I: twisting of sclerite at transition between ls-w and ls-p. Grey arrow in A indicates direction of squeezing of tube lst. Blue arrowheads: anterior extremity of wing part ls-w on mesal border of tube lst, ls-w either restricted to posterior half of tubes lst (e. g. K) or reaching far into anterior half (e. g. D). Red arrowheads: area where the posterior margin of ls-a bends dorsally, and where a bend or kink is often present in (inner) lateral border of tube lst (compare white arrowheads in Fig. 23 C, D, H, I, J). Black arrowheads: membranous inner lateral border of tube lst posterior to the bend or kink (compare Fig. 23 C, D, H, I and black arrowheads in sections beside D). - Inserts in B, D, E, K showing right (of picture and animal) tube lst with posterior margin of ls-a (red line; compare Fig. 23 C-E) and anterior margin of ls-w (blue line; compare Fig. 23 F, G).	Figure 22. Laterosternal shelf area of females of Attaphila species, ventral view, posteriorly on top. A, B: A. aptera (Bo 1225, Bo 1257), in A left tube distorted (ventral part of tube squeezed laterally, as indicated by grey arrow). C, D: A. bergi (Bo 1282, Bo 1239). E, F: A. fungicola (Bo 1264, Bo 1447). G: A. multisetosa (HT Bo 1270). H: A. paucisetosa (PT Bo 1255). I: A. sexdentis (HT Bo 1233). J: A. schuppi (ST Bo 1237). K, L: A. sinuosocarinata (HT Bo 1273, PT Bo 1288). - Abbreviations: isf intersternal folds (of floor of vestibulum); ls laterosternal-shelf sclerite (with central part c, arm part a, wing part w, posterolateral extremity p); lst laterosternal-shelf tube; vfl floor of vestibulum. (See Supplement 3 Fig. S 2 D, E for more complete labeling using different terminologies.) - Arrows: Black arrows in A, D, I: twisting of sclerite at transition between ls-w and ls-p. Grey arrow in A indicates direction of squeezing of tube lst. Blue arrowheads: anterior extremity of wing part ls-w on mesal border of tube lst, ls-w either restricted to posterior half of tubes lst (e. g. K) or reaching far into anterior half (e. g. D). Red arrowheads: area where the posterior margin of ls-a bends dorsally, and where a bend or kink is often present in (inner) lateral border of tube lst (compare white arrowheads in Fig. 23 C, D, H, I, J). Black arrowheads: membranous inner lateral border of tube lst posterior to the bend or kink (compare Fig. 23 C, D, H, I and black arrowheads in sections beside D). - Inserts in B, D, E, K showing right (of picture and animal) tube lst with posterior margin of ls-a (red line; compare Fig. 23 C-E) and anterior margin of ls-w (blue line; compare Fig. 23 F, G).	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
90567B49330A5CAFBEFA87D6C9B05DA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563294	https://doi.org/10.3897/asp.79.e67569.figure25	Figure 25. Subgenital plate (sternite S 9) and phallomeres of males of Attaphila species, dorsal view, anteriorly on top. A, B: A. paucisetosa, subgenital plate with phallomeres (PT Bo 1254, Bo 1444), hook and right phallomere removed in B. C: A. flava, subgenital plate with phallomeres (HT Bo 1280). D, E: A. fungicola. D showing subgenital plate with phallomeres (LT Bo 1265), hook not seen in profile; E showing isolated hook seen in profile (Bo 1229). - Abbreviations: T 9 p ventrally bent lateral (paratergal) part of tergite 9; otherwise as for Fig. 24.	Figure 25. Subgenital plate (sternite S 9) and phallomeres of males of Attaphila species, dorsal view, anteriorly on top. A, B: A. paucisetosa, subgenital plate with phallomeres (PT Bo 1254, Bo 1444), hook and right phallomere removed in B. C: A. flava, subgenital plate with phallomeres (HT Bo 1280). D, E: A. fungicola. D showing subgenital plate with phallomeres (LT Bo 1265), hook not seen in profile; E showing isolated hook seen in profile (Bo 1229). - Abbreviations: T 9 p ventrally bent lateral (paratergal) part of tergite 9; otherwise as for Fig. 24.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
90567B49330A5CAFBEFA87D6C9B05DA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563295	https://doi.org/10.3897/asp.79.e67569.figure26	Figure 26. Subgenital plate (sternite S 9) and phallomeres of males of Attaphila species, phase contrast images of the same objects as shown in Fig. 25 A, C, D, anteriorly on top. A: A. flava (HT Bo 1280). B: A. fungicola (LT Bo 1265). C: A. paucisetosa (PT Bo 1254). - Abbreviations: A 1 articulation between S 9 and pt 9; cs cleft sclerite of right phallomere; ea endophallic apodeme; h hook; psa process; pt 9 extension of paratergite 9 contacting sternite 9; R 3 anterior principal sclerite of right phallomere; sll left stylus; sta 9 anterior apodeme of subgenital plate S 9; vf 9 subgenital lobe; vi virga. Arrow: in A indicating that the distal part of the subgenital lobe is broken off.	Figure 26. Subgenital plate (sternite S 9) and phallomeres of males of Attaphila species, phase contrast images of the same objects as shown in Fig. 25 A, C, D, anteriorly on top. A: A. flava (HT Bo 1280). B: A. fungicola (LT Bo 1265). C: A. paucisetosa (PT Bo 1254). - Abbreviations: A 1 articulation between S 9 and pt 9; cs cleft sclerite of right phallomere; ea endophallic apodeme; h hook; psa process; pt 9 extension of paratergite 9 contacting sternite 9; R 3 anterior principal sclerite of right phallomere; sll left stylus; sta 9 anterior apodeme of subgenital plate S 9; vf 9 subgenital lobe; vi virga. Arrow: in A indicating that the distal part of the subgenital lobe is broken off.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
90567B49330A5CAFBEFA87D6C9B05DA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563280	https://doi.org/10.3897/asp.79.e67569.figure11	Figure 11. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. schuppi (ST Bo 1237). A: T 2 - 5; B: T 6,7. C, D: A. sexdentis (HT Bo 1233). C: T 1 - 5; D: T 6,7. - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to median and lateral excurvations of transversal ridges to the anterior.	Figure 11. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. schuppi (ST Bo 1237). A: T 2 - 5; B: T 6,7. C, D: A. sexdentis (HT Bo 1233). C: T 1 - 5; D: T 6,7. - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to median and lateral excurvations of transversal ridges to the anterior.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
53C5A9FF8D925D82B10140AAF23773D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563279	https://doi.org/10.3897/asp.79.e67569.figure10	Figure 10. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. multisetosa (HT Bo 1270). A: T 2 - 5; B: T 6,7. C - E: A. sinuosocarinata. C: T 1 - 5 (HT Bo 1273); D: T 6,7 (HT Bo 1273); E: T 6 (PT Bo 1288). - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to weak median and lateral excurvations of transversal ridges to the anterior.	Figure 10. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. multisetosa (HT Bo 1270). A: T 2 - 5; B: T 6,7. C - E: A. sinuosocarinata. C: T 1 - 5 (HT Bo 1273); D: T 6,7 (HT Bo 1273); E: T 6 (PT Bo 1288). - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to weak median and lateral excurvations of transversal ridges to the anterior.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
53C5A9FF8D925D82B10140AAF23773D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563281	https://doi.org/10.3897/asp.79.e67569.figure12	Figure 12. Abdominal tergites T 6,7 of females of Attaphila species. A: A. aptera (Bo 1257). B: A. bergi (Bo 1282). C: A. fungicola (Bo 1264). D: A. multisetosa (HT Bo 1270). E: A. paucisetosa (PT Bo 1255). F: A. schuppi (ST Bo 1237), transversal ridge tr 7 almost completely missing, only tiny remnants present near the associated bristles (see Fig. 14 F). - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 12. Abdominal tergites T 6,7 of females of Attaphila species. A: A. aptera (Bo 1257). B: A. bergi (Bo 1282). C: A. fungicola (Bo 1264). D: A. multisetosa (HT Bo 1270). E: A. paucisetosa (PT Bo 1255). F: A. schuppi (ST Bo 1237), transversal ridge tr 7 almost completely missing, only tiny remnants present near the associated bristles (see Fig. 14 F). - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
53C5A9FF8D925D82B10140AAF23773D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563283	https://doi.org/10.3897/asp.79.e67569.figure14	Figure 14. Size and distribution of bristles in the median part of tergites T 6,7 in females (A-H) and males (I) of Attaphila species, phase contrast images. A: A. aptera, female (Bo 1257). B: A. bergi, female (Bo 1282). C: A. fungicola, female (Bo 1264). D: A. multisetosa, female (Bo 1270). E: A. paucisetosa, female (PT Bo 1255). F: A. schuppi, female (ST Bo 1237), with only short remnants of transversal ridge tr 7. G: A. sexdentis, female (Bo 1233). H: A. sinuosocarinata, female (HT Bo 1273). I: A. aptera, male (Bo 1256). - Abbreviations: T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 14. Size and distribution of bristles in the median part of tergites T 6,7 in females (A-H) and males (I) of Attaphila species, phase contrast images. A: A. aptera, female (Bo 1257). B: A. bergi, female (Bo 1282). C: A. fungicola, female (Bo 1264). D: A. multisetosa, female (Bo 1270). E: A. paucisetosa, female (PT Bo 1255). F: A. schuppi, female (ST Bo 1237), with only short remnants of transversal ridge tr 7. G: A. sexdentis, female (Bo 1233). H: A. sinuosocarinata, female (HT Bo 1273). I: A. aptera, male (Bo 1256). - Abbreviations: T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
53C5A9FF8D925D82B10140AAF23773D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563286	https://doi.org/10.3897/asp.79.e67569.figure17	Figure 17. Subgenital plate (sternite S 7) of females (A - F) and larval females (G, H) of Attaphila species. A, B: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). C, D: A. aptera (Bo 1225, Bo 1253). E: A. multisetosa (HT Bo 1270). F: A. sexdentis (HT Bo 1233). G: A. aptera, larval female (Bo 1289). H: A. bergi, larval female (Bo 1231). - Abbreviations and arrows: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars). Arrows in B, C, F pointing to lateral borders of S 7, which are parallel or slightly divergent (to the anterior) in A-E, but convergent in F.	Figure 17. Subgenital plate (sternite S 7) of females (A - F) and larval females (G, H) of Attaphila species. A, B: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). C, D: A. aptera (Bo 1225, Bo 1253). E: A. multisetosa (HT Bo 1270). F: A. sexdentis (HT Bo 1233). G: A. aptera, larval female (Bo 1289). H: A. bergi, larval female (Bo 1231). - Abbreviations and arrows: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars). Arrows in B, C, F pointing to lateral borders of S 7, which are parallel or slightly divergent (to the anterior) in A-E, but convergent in F.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
53C5A9FF8D925D82B10140AAF23773D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563287	https://doi.org/10.3897/asp.79.e67569.figure18	Figure 18. Anterior half of subgenital plate (sternite S 7) of females of Attaphila species, shape of sternal transversal ridge, phase contrast images (same objects as shown in Figs 16, 17, except for Bo 1236 in F). A, B: A. aptera (Bo 1225, Bo 1253). C, D: A. bergi (Bo 1275, Bo 1282). E, F: A. fungicola (Bo 1264, Bo 1236). G: A. multisetosa (HT Bo 1270). H, I: A. paucisetosa (Cb 2 / 2, PT Bo 1255). J, K: A. schuppi (ST Bo 1234, ST Bo 1237). L: A. sexdentis (HT Bo 1233). M, N: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). - Abbreviations and symbols: sr 7 transversal ridge of subgenital plate S 7; short interruptions of this ridge indicated by arrows, larger gaps delimited by double bars.	Figure 18. Anterior half of subgenital plate (sternite S 7) of females of Attaphila species, shape of sternal transversal ridge, phase contrast images (same objects as shown in Figs 16, 17, except for Bo 1236 in F). A, B: A. aptera (Bo 1225, Bo 1253). C, D: A. bergi (Bo 1275, Bo 1282). E, F: A. fungicola (Bo 1264, Bo 1236). G: A. multisetosa (HT Bo 1270). H, I: A. paucisetosa (Cb 2 / 2, PT Bo 1255). J, K: A. schuppi (ST Bo 1234, ST Bo 1237). L: A. sexdentis (HT Bo 1233). M, N: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). - Abbreviations and symbols: sr 7 transversal ridge of subgenital plate S 7; short interruptions of this ridge indicated by arrows, larger gaps delimited by double bars.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
53C5A9FF8D925D82B10140AAF23773D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563290	https://doi.org/10.3897/asp.79.e67569.figure21	Figure 21. A-C: Terminalia with genitalia of females of Attaphila species, ventral view, posterior end on top. A: A. multisetosa (HT Bo 1270), without laterosternal shelf area, cerci damaged; inset showing median part of genital chamber dorsal wall. B: A. sinuosocarinata (HT Bo 1273), without laterosternal shelf area; inset showing median part of genital chamber dorsal wall. C: A. sexdentis (HT Bo 1233), with laterosternal shelf area. - D: Female of A. paucisetosa carrying an ootheca; length of animal ca. 3 mm. - Abbreviations: aa anterior arch; ca central apodeme; ce cercus; gcf fold dividing genital chamber (posterior edge labeled); gcp pouch of genital chamber (one-sided); gg-m mesal gonangulum sclerite; IC intercalary sclerite; ls laterosternal-shelf sclerite (with posterolateral extremity p); lst laterosternal-shelf tube; otkl ootheca keel; pl posterior lobe; PP paraproct; pt 8,9 extension of paratergite 9; sp spermathecal plate; T 9, T 10 abdominal tergites 9 and 10; T 10 - p posterior border of tergite 10; vlf valvifer of segment 8; A articulations, A 2 between gg-m and pl, A 3 between gg-m and basal sclerotisation of v 1, A 5 between pt 8,9 and vlf. Arrow in A, B showing mesal border of putative weak sclerotisation in dorsal wall of genital chamber. (See Supplement 1 and Supplement 3 Fig. S 2 A, B, C for additional explanations and more complete labeling using different terminologies.)	Figure 21. A-C: Terminalia with genitalia of females of Attaphila species, ventral view, posterior end on top. A: A. multisetosa (HT Bo 1270), without laterosternal shelf area, cerci damaged; inset showing median part of genital chamber dorsal wall. B: A. sinuosocarinata (HT Bo 1273), without laterosternal shelf area; inset showing median part of genital chamber dorsal wall. C: A. sexdentis (HT Bo 1233), with laterosternal shelf area. - D: Female of A. paucisetosa carrying an ootheca; length of animal ca. 3 mm. - Abbreviations: aa anterior arch; ca central apodeme; ce cercus; gcf fold dividing genital chamber (posterior edge labeled); gcp pouch of genital chamber (one-sided); gg-m mesal gonangulum sclerite; IC intercalary sclerite; ls laterosternal-shelf sclerite (with posterolateral extremity p); lst laterosternal-shelf tube; otkl ootheca keel; pl posterior lobe; PP paraproct; pt 8,9 extension of paratergite 9; sp spermathecal plate; T 9, T 10 abdominal tergites 9 and 10; T 10 - p posterior border of tergite 10; vlf valvifer of segment 8; A articulations, A 2 between gg-m and pl, A 3 between gg-m and basal sclerotisation of v 1, A 5 between pt 8,9 and vlf. Arrow in A, B showing mesal border of putative weak sclerotisation in dorsal wall of genital chamber. (See Supplement 1 and Supplement 3 Fig. S 2 A, B, C for additional explanations and more complete labeling using different terminologies.)	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
53C5A9FF8D925D82B10140AAF23773D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563291	https://doi.org/10.3897/asp.79.e67569.figure22	Figure 22. Laterosternal shelf area of females of Attaphila species, ventral view, posteriorly on top. A, B: A. aptera (Bo 1225, Bo 1257), in A left tube distorted (ventral part of tube squeezed laterally, as indicated by grey arrow). C, D: A. bergi (Bo 1282, Bo 1239). E, F: A. fungicola (Bo 1264, Bo 1447). G: A. multisetosa (HT Bo 1270). H: A. paucisetosa (PT Bo 1255). I: A. sexdentis (HT Bo 1233). J: A. schuppi (ST Bo 1237). K, L: A. sinuosocarinata (HT Bo 1273, PT Bo 1288). - Abbreviations: isf intersternal folds (of floor of vestibulum); ls laterosternal-shelf sclerite (with central part c, arm part a, wing part w, posterolateral extremity p); lst laterosternal-shelf tube; vfl floor of vestibulum. (See Supplement 3 Fig. S 2 D, E for more complete labeling using different terminologies.) - Arrows: Black arrows in A, D, I: twisting of sclerite at transition between ls-w and ls-p. Grey arrow in A indicates direction of squeezing of tube lst. Blue arrowheads: anterior extremity of wing part ls-w on mesal border of tube lst, ls-w either restricted to posterior half of tubes lst (e. g. K) or reaching far into anterior half (e. g. D). Red arrowheads: area where the posterior margin of ls-a bends dorsally, and where a bend or kink is often present in (inner) lateral border of tube lst (compare white arrowheads in Fig. 23 C, D, H, I, J). Black arrowheads: membranous inner lateral border of tube lst posterior to the bend or kink (compare Fig. 23 C, D, H, I and black arrowheads in sections beside D). - Inserts in B, D, E, K showing right (of picture and animal) tube lst with posterior margin of ls-a (red line; compare Fig. 23 C-E) and anterior margin of ls-w (blue line; compare Fig. 23 F, G).	Figure 22. Laterosternal shelf area of females of Attaphila species, ventral view, posteriorly on top. A, B: A. aptera (Bo 1225, Bo 1257), in A left tube distorted (ventral part of tube squeezed laterally, as indicated by grey arrow). C, D: A. bergi (Bo 1282, Bo 1239). E, F: A. fungicola (Bo 1264, Bo 1447). G: A. multisetosa (HT Bo 1270). H: A. paucisetosa (PT Bo 1255). I: A. sexdentis (HT Bo 1233). J: A. schuppi (ST Bo 1237). K, L: A. sinuosocarinata (HT Bo 1273, PT Bo 1288). - Abbreviations: isf intersternal folds (of floor of vestibulum); ls laterosternal-shelf sclerite (with central part c, arm part a, wing part w, posterolateral extremity p); lst laterosternal-shelf tube; vfl floor of vestibulum. (See Supplement 3 Fig. S 2 D, E for more complete labeling using different terminologies.) - Arrows: Black arrows in A, D, I: twisting of sclerite at transition between ls-w and ls-p. Grey arrow in A indicates direction of squeezing of tube lst. Blue arrowheads: anterior extremity of wing part ls-w on mesal border of tube lst, ls-w either restricted to posterior half of tubes lst (e. g. K) or reaching far into anterior half (e. g. D). Red arrowheads: area where the posterior margin of ls-a bends dorsally, and where a bend or kink is often present in (inner) lateral border of tube lst (compare white arrowheads in Fig. 23 C, D, H, I, J). Black arrowheads: membranous inner lateral border of tube lst posterior to the bend or kink (compare Fig. 23 C, D, H, I and black arrowheads in sections beside D). - Inserts in B, D, E, K showing right (of picture and animal) tube lst with posterior margin of ls-a (red line; compare Fig. 23 C-E) and anterior margin of ls-w (blue line; compare Fig. 23 F, G).	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
53C5A9FF8D925D82B10140AAF23773D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563280	https://doi.org/10.3897/asp.79.e67569.figure11	Figure 11. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. schuppi (ST Bo 1237). A: T 2 - 5; B: T 6,7. C, D: A. sexdentis (HT Bo 1233). C: T 1 - 5; D: T 6,7. - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to median and lateral excurvations of transversal ridges to the anterior.	Figure 11. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. schuppi (ST Bo 1237). A: T 2 - 5; B: T 6,7. C, D: A. sexdentis (HT Bo 1233). C: T 1 - 5; D: T 6,7. - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to median and lateral excurvations of transversal ridges to the anterior.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
31348359E8745C70B5E357A00B36B7DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563270	https://doi.org/10.3897/asp.79.e67569.figure1	Figure 1. A - D: Head structures of Attaphila species, head made transparent by treatment with KOH. A: Entire head with focus on antennae and circumantennal pit (inset: median part, darkened). B: Entire head with focus on mouthparts (cardo and stipes seen behind circumantennal pit; inset: scheme of mandibular dentition in anterior view, mdl = left, mdr = right, elements above transversal line belonging to mola). C: Entire head with left antenna rectangularly bent between scapus and pedicellus. In A-C different levels of tentorium visible internally. D: Base of right antenna and surrounding parts of head capsule at higher magnification, also showing dorsal excavation at distal end of scapus allowing the strong bend shown in C. E - H: Thoracic nota of Attaphila species. E: All thoracic nota of a female, stretched. F: Pronotum of a male. G, H: Right part of metanotum of males with the strongly reduced hindwing. - Species: A - C: A. aptera, male (A, B: Bo 1224), larval female (C: Bo 1243); D: A. fungicola, male (Bo 1229); E, G: A. bergi, female (Bo 1239), male (Bo 1283); F, H: A. paucisetosa, male (Bo 1240). - Abbreviations: ata anterior tentorial arm; atb anterior tentorial bridge; atp anterior tentorial pit; car cardo of maxilla; cap circumantennal pit (with sharp dorsal edge capd visible and bottom capb of the pit shining through); cc coronal cleavage line; cpe compound eye; ct corpotentorium; fc frontal cleavage line; fl fagellum (all antennomeres following pedicellus, numbered from base as fl 1 etc.); hw hindwing; hwar level of (reduced) hindwing articulation; li lacinia; lp labial palpus (I-III: palpomeres); md mandible (left mdl, right mdr; I-IV: tip and incisivi); mo mola of mandible; mp maxillary palpus (I-V: palpomeres); N thoracic notum (N 1 pro-, N 2 meso-, N 3 metanotum); pc pedicellus; pta posterior tentorial arm; sc scapus; sc-pc scapus-pedicellus articulation; scex dorsal excavation at distal end of scapus; s-chA field of sensilla chaetica A; s-cpf row of sensilla campaniformia; sti stipes of maxilla; tp tentorial perforation.	Figure 1. A - D: Head structures of Attaphila species, head made transparent by treatment with KOH. A: Entire head with focus on antennae and circumantennal pit (inset: median part, darkened). B: Entire head with focus on mouthparts (cardo and stipes seen behind circumantennal pit; inset: scheme of mandibular dentition in anterior view, mdl = left, mdr = right, elements above transversal line belonging to mola). C: Entire head with left antenna rectangularly bent between scapus and pedicellus. In A-C different levels of tentorium visible internally. D: Base of right antenna and surrounding parts of head capsule at higher magnification, also showing dorsal excavation at distal end of scapus allowing the strong bend shown in C. E - H: Thoracic nota of Attaphila species. E: All thoracic nota of a female, stretched. F: Pronotum of a male. G, H: Right part of metanotum of males with the strongly reduced hindwing. - Species: A - C: A. aptera, male (A, B: Bo 1224), larval female (C: Bo 1243); D: A. fungicola, male (Bo 1229); E, G: A. bergi, female (Bo 1239), male (Bo 1283); F, H: A. paucisetosa, male (Bo 1240). - Abbreviations: ata anterior tentorial arm; atb anterior tentorial bridge; atp anterior tentorial pit; car cardo of maxilla; cap circumantennal pit (with sharp dorsal edge capd visible and bottom capb of the pit shining through); cc coronal cleavage line; cpe compound eye; ct corpotentorium; fc frontal cleavage line; fl fagellum (all antennomeres following pedicellus, numbered from base as fl 1 etc.); hw hindwing; hwar level of (reduced) hindwing articulation; li lacinia; lp labial palpus (I-III: palpomeres); md mandible (left mdl, right mdr; I-IV: tip and incisivi); mo mola of mandible; mp maxillary palpus (I-V: palpomeres); N thoracic notum (N 1 pro-, N 2 meso-, N 3 metanotum); pc pedicellus; pta posterior tentorial arm; sc scapus; sc-pc scapus-pedicellus articulation; scex dorsal excavation at distal end of scapus; s-chA field of sensilla chaetica A; s-cpf row of sensilla campaniformia; sti stipes of maxilla; tp tentorial perforation.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
31348359E8745C70B5E357A00B36B7DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563271	https://doi.org/10.3897/asp.79.e67569.figure2	Figure 2. A-I: Antennae of Attaphila species of various age. A, H, I: Presumably second stage larva (head width 0.52 mm), with enlarged details of the proximal (H) and distal regions (I, distal end to the right), both from (A); B - E: intermediate and late larval stages, head width 0.66 (B), 0.61 (C), 0.76 (D), 0.73 mm (E); F, G: imagines. J-M: Imaginal antennae of Blattellidae species, in (J) showing parts of the proximal, intermediate and distal region, in (K-M) only of the proximal and distal region. - Species: A, C, E, H, I: Attaphila paucisetosa (A, H, I: Colombia, Cb 4 / 3; C: Bo 1458; E: Bo 1433); B, D, F: A. aptera (B: Bo 1457; D: Bo 1292; F: Bo 1225); G: A. bergi (Bo 1274); J: Blattella germanica (ex cult.); K: Symploce pallens (ex cult.); L: Xestoblatta cantralli (Costa Rica, CR 15); M: Ischnoptera sp. (Costa Rica, CR 13 / 1). - Abbreviations and symbols: sc scapus; pc pedicellus; me meriston (first flagellomere of current stage); + new proximal flagellomeres generated by a division of the meriston into two or three flagellomeres during the preceding intermoult period in B, E, H, in C, D the division had presumably already occurred one moult earlier; L - early larval stage; L + intermediate or late larval stage; Im imago.	Figure 2. A-I: Antennae of Attaphila species of various age. A, H, I: Presumably second stage larva (head width 0.52 mm), with enlarged details of the proximal (H) and distal regions (I, distal end to the right), both from (A); B - E: intermediate and late larval stages, head width 0.66 (B), 0.61 (C), 0.76 (D), 0.73 mm (E); F, G: imagines. J-M: Imaginal antennae of Blattellidae species, in (J) showing parts of the proximal, intermediate and distal region, in (K-M) only of the proximal and distal region. - Species: A, C, E, H, I: Attaphila paucisetosa (A, H, I: Colombia, Cb 4 / 3; C: Bo 1458; E: Bo 1433); B, D, F: A. aptera (B: Bo 1457; D: Bo 1292; F: Bo 1225); G: A. bergi (Bo 1274); J: Blattella germanica (ex cult.); K: Symploce pallens (ex cult.); L: Xestoblatta cantralli (Costa Rica, CR 15); M: Ischnoptera sp. (Costa Rica, CR 13 / 1). - Abbreviations and symbols: sc scapus; pc pedicellus; me meriston (first flagellomere of current stage); + new proximal flagellomeres generated by a division of the meriston into two or three flagellomeres during the preceding intermoult period in B, E, H, in C, D the division had presumably already occurred one moult earlier; L - early larval stage; L + intermediate or late larval stage; Im imago.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
31348359E8745C70B5E357A00B36B7DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563272	https://doi.org/10.3897/asp.79.e67569.figure3(a-f)	Figure 3 (A-J). Tegmina (forewings) of males of Attaphila species, dorsal views, left tegmen at the left. - Species: A, B: A. aptera (Bo 1252); C, D: A. bergi (Bo 1443); E, F: A. flava (HT Bo 1280); G, H: A. fungicola (Bo 1229); I, J: A. paucisetosa (Bo 1441). - Abbreviation: fwar forewing articulation area. In A borders specified as addressed in the text.	Figure 3 (A-J). Tegmina (forewings) of males of Attaphila species, dorsal views, left tegmen at the left. - Species: A, B: A. aptera (Bo 1252); C, D: A. bergi (Bo 1443); E, F: A. flava (HT Bo 1280); G, H: A. fungicola (Bo 1229); I, J: A. paucisetosa (Bo 1441). - Abbreviation: fwar forewing articulation area. In A borders specified as addressed in the text.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
31348359E8745C70B5E357A00B36B7DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563273	https://doi.org/10.3897/asp.79.e67569.figure4	Figure 4. Legs of left side (A-G) or right side (H) of female Attaphila paucisetosa (A: Bo 1235, B-D, H, G: PT Bo 1226), anterior view, made transparent by treatment with KOH. Hindleg (A, B), midleg (C), and foreleg (D, G [left leg regenerated, with only four tarsomeres], H [right leg with five tarsomeres]), with conventional terminology of tibial spines. E and F show a general scheme of cockroach tibia spination of the foreleg (E) and midleg (F; similar on hindleg) (tibia base on top, tibia cut along ventral midline and spread, spine bases represented by circles filled with different colours according to groups of spines) and distal part of tibia of fore- (E), mid-, and hindleg (F) of Attaphila (from B, C, D, 1.5 x enlarged), with spine terminology according to Klass et al. (2009) and unpublished work by K. - D. Klass and coworkers. G and H show tarsus of left (G) and right (H) foreleg enlarged (compare D); white arrow in G pointing to dorsal vestige of subdivision between tarsomeres 1 and 2. - Abbreviations: co coxa; colb coxal lobelet; tc trochanter; fe femur; fegr ventral groove of femur which can take up part of tibia during strong flexion as shown in (A) (bottom of groove indicated); fane and fpoe antero- resp. posteroventral edge flanking femoral groove; ti tibia; tiaa and tiap anterior resp. posterior articulation of tibia with femur; tivt ventral basal tendon of tibia; dft dorsal fold on apical margin of tibia; itts intertibiotarsal sclerite; pdta posterodorsal tibiotarsal articulation (on opposite side of leg); ta 1 - 5 tarsomeres 1 - 5; ptcl claws of pretarsus; ptar arolium of pretarsus; gs genicular spine of femur. Spines of tibia (see Supplement 1 for complete abbreviations): as apical spines (= terminal spines Tt 1 - 5 plus distal mediodorsal spine Td 1 m [mid- and hindleg] or distal anterodorsal spine Td 1 ma [foreleg]); ds dorsal spines (= dorsal spines Td excluding Td 1 m [mid- and hindleg] and Td 1 ma [foreleg]); vs ventral spines (= ventral spines Tv).	Figure 4. Legs of left side (A-G) or right side (H) of female Attaphila paucisetosa (A: Bo 1235, B-D, H, G: PT Bo 1226), anterior view, made transparent by treatment with KOH. Hindleg (A, B), midleg (C), and foreleg (D, G [left leg regenerated, with only four tarsomeres], H [right leg with five tarsomeres]), with conventional terminology of tibial spines. E and F show a general scheme of cockroach tibia spination of the foreleg (E) and midleg (F; similar on hindleg) (tibia base on top, tibia cut along ventral midline and spread, spine bases represented by circles filled with different colours according to groups of spines) and distal part of tibia of fore- (E), mid-, and hindleg (F) of Attaphila (from B, C, D, 1.5 x enlarged), with spine terminology according to Klass et al. (2009) and unpublished work by K. - D. Klass and coworkers. G and H show tarsus of left (G) and right (H) foreleg enlarged (compare D); white arrow in G pointing to dorsal vestige of subdivision between tarsomeres 1 and 2. - Abbreviations: co coxa; colb coxal lobelet; tc trochanter; fe femur; fegr ventral groove of femur which can take up part of tibia during strong flexion as shown in (A) (bottom of groove indicated); fane and fpoe antero- resp. posteroventral edge flanking femoral groove; ti tibia; tiaa and tiap anterior resp. posterior articulation of tibia with femur; tivt ventral basal tendon of tibia; dft dorsal fold on apical margin of tibia; itts intertibiotarsal sclerite; pdta posterodorsal tibiotarsal articulation (on opposite side of leg); ta 1 - 5 tarsomeres 1 - 5; ptcl claws of pretarsus; ptar arolium of pretarsus; gs genicular spine of femur. Spines of tibia (see Supplement 1 for complete abbreviations): as apical spines (= terminal spines Tt 1 - 5 plus distal mediodorsal spine Td 1 m [mid- and hindleg] or distal anterodorsal spine Td 1 ma [foreleg]); ds dorsal spines (= dorsal spines Td excluding Td 1 m [mid- and hindleg] and Td 1 ma [foreleg]); vs ventral spines (= ventral spines Tv).	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
31348359E8745C70B5E357A00B36B7DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563274	https://doi.org/10.3897/asp.79.e67569.figure5	Figure 5. A - I: Tergite specialisations of the males of Attaphila species, medially on T 1 (A - D), or laterally at the anterior border of T 2 (E - I), in E-G from right side of tergite, in H and I from left side. - Species: A, B: A. aptera (Bo 1227, HT Bo 1258); C, D: A. bergi (Bo 1283, Bo 1274); E, F: A. paucisetosa (PT Bo 1254, Bo 1256); G, H: A. fungicola (LT Bo 1265); I: A. flava (HT Bo 1280). - Abbreviations: T 1 - a, T 2 - a anterior borders of tergites T 1,2; T 1 - p posterior border of T 1; tr 1, tr 2 transversal ridges of T 1,2; msa 1 anterior median specialisation of tergite T 1; msl 2 lateral specialisation of tergite T 2; msp 1 posterior median specialisation of tergite T 1; pg pores of glands, tiny in A, B, between T 1 - a and tr 1, larger in E as part of msl 2 specialisations. - J, K: Paraprocts, tergite 10 and cerci in ventral view, of J: A. aptera (Bo 1224) and K: A. paucisetosa (HT Bo 1258). - Abbreviations and arrows: Ca sclerite at mesal base of cercus; ce cercus; hmp hook-like mesal projection of paraproct; PPl, PPr left and right paraproct; T 9 tergite 9; T 10 tergite 10 with its ventrally bent lateral (paratergal) parts T 10 p (larger median parts of T 10 out of focus); white arrows: articulation between mesal end of T 10 p and lateral end of paraproct.	Figure 5. A - I: Tergite specialisations of the males of Attaphila species, medially on T 1 (A - D), or laterally at the anterior border of T 2 (E - I), in E-G from right side of tergite, in H and I from left side. - Species: A, B: A. aptera (Bo 1227, HT Bo 1258); C, D: A. bergi (Bo 1283, Bo 1274); E, F: A. paucisetosa (PT Bo 1254, Bo 1256); G, H: A. fungicola (LT Bo 1265); I: A. flava (HT Bo 1280). - Abbreviations: T 1 - a, T 2 - a anterior borders of tergites T 1,2; T 1 - p posterior border of T 1; tr 1, tr 2 transversal ridges of T 1,2; msa 1 anterior median specialisation of tergite T 1; msl 2 lateral specialisation of tergite T 2; msp 1 posterior median specialisation of tergite T 1; pg pores of glands, tiny in A, B, between T 1 - a and tr 1, larger in E as part of msl 2 specialisations. - J, K: Paraprocts, tergite 10 and cerci in ventral view, of J: A. aptera (Bo 1224) and K: A. paucisetosa (HT Bo 1258). - Abbreviations and arrows: Ca sclerite at mesal base of cercus; ce cercus; hmp hook-like mesal projection of paraproct; PPl, PPr left and right paraproct; T 9 tergite 9; T 10 tergite 10 with its ventrally bent lateral (paratergal) parts T 10 p (larger median parts of T 10 out of focus); white arrows: articulation between mesal end of T 10 p and lateral end of paraproct.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
31348359E8745C70B5E357A00B36B7DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563278	https://doi.org/10.3897/asp.79.e67569.figure9	Figure 9. Abdominal tergites and terminalia of Attaphila paucisetosa. A - D: Male (HT Bo 1258). A: T 1; B: T 2 - 5, T 2 with lateral specialisations; C: T 6,7. D: Terminalia, ventral view, with T 9 (with lateral extension) and T 10, cerci, and paraprocts. E, F: Female (PT Bo 1255). E: T 1 - 5; F: T 6,7. - Abbreviations: ce cercus; msl 2 lateral specialisation of T 2; PPr right paraproct; T n tergite (numbered); pt 9 extension of paratergite of T 9; T n - a anterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to excurvations of male tr 2.	Figure 9. Abdominal tergites and terminalia of Attaphila paucisetosa. A - D: Male (HT Bo 1258). A: T 1; B: T 2 - 5, T 2 with lateral specialisations; C: T 6,7. D: Terminalia, ventral view, with T 9 (with lateral extension) and T 10, cerci, and paraprocts. E, F: Female (PT Bo 1255). E: T 1 - 5; F: T 6,7. - Abbreviations: ce cercus; msl 2 lateral specialisation of T 2; PPr right paraproct; T n tergite (numbered); pt 9 extension of paratergite of T 9; T n - a anterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to excurvations of male tr 2.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
31348359E8745C70B5E357A00B36B7DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563281	https://doi.org/10.3897/asp.79.e67569.figure12	Figure 12. Abdominal tergites T 6,7 of females of Attaphila species. A: A. aptera (Bo 1257). B: A. bergi (Bo 1282). C: A. fungicola (Bo 1264). D: A. multisetosa (HT Bo 1270). E: A. paucisetosa (PT Bo 1255). F: A. schuppi (ST Bo 1237), transversal ridge tr 7 almost completely missing, only tiny remnants present near the associated bristles (see Fig. 14 F). - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 12. Abdominal tergites T 6,7 of females of Attaphila species. A: A. aptera (Bo 1257). B: A. bergi (Bo 1282). C: A. fungicola (Bo 1264). D: A. multisetosa (HT Bo 1270). E: A. paucisetosa (PT Bo 1255). F: A. schuppi (ST Bo 1237), transversal ridge tr 7 almost completely missing, only tiny remnants present near the associated bristles (see Fig. 14 F). - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
31348359E8745C70B5E357A00B36B7DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563282	https://doi.org/10.3897/asp.79.e67569.figure13	Figure 13. Abdominal tergites T 6,7 of females (A, B) and males (C - F) of Attaphila species. A: A. sexdentis, female (Bo 1233). B: A. sinuosocarinata, female (HT Bo 1273). C: A. aptera, male (Bo 1256). D: A. bergi, male (Bo 1274), transversal ridge tr 7 weakly developed, sublaterally with a large gap. E: A. paucisetosa, male (Bo 1240), transversal ridge tr 7 missing. F: A. fungicola, male (Bo 1229), transversal ridge tr 7 missing. - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 13. Abdominal tergites T 6,7 of females (A, B) and males (C - F) of Attaphila species. A: A. sexdentis, female (Bo 1233). B: A. sinuosocarinata, female (HT Bo 1273). C: A. aptera, male (Bo 1256). D: A. bergi, male (Bo 1274), transversal ridge tr 7 weakly developed, sublaterally with a large gap. E: A. paucisetosa, male (Bo 1240), transversal ridge tr 7 missing. F: A. fungicola, male (Bo 1229), transversal ridge tr 7 missing. - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
31348359E8745C70B5E357A00B36B7DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563283	https://doi.org/10.3897/asp.79.e67569.figure14	Figure 14. Size and distribution of bristles in the median part of tergites T 6,7 in females (A-H) and males (I) of Attaphila species, phase contrast images. A: A. aptera, female (Bo 1257). B: A. bergi, female (Bo 1282). C: A. fungicola, female (Bo 1264). D: A. multisetosa, female (Bo 1270). E: A. paucisetosa, female (PT Bo 1255). F: A. schuppi, female (ST Bo 1237), with only short remnants of transversal ridge tr 7. G: A. sexdentis, female (Bo 1233). H: A. sinuosocarinata, female (HT Bo 1273). I: A. aptera, male (Bo 1256). - Abbreviations: T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 14. Size and distribution of bristles in the median part of tergites T 6,7 in females (A-H) and males (I) of Attaphila species, phase contrast images. A: A. aptera, female (Bo 1257). B: A. bergi, female (Bo 1282). C: A. fungicola, female (Bo 1264). D: A. multisetosa, female (Bo 1270). E: A. paucisetosa, female (PT Bo 1255). F: A. schuppi, female (ST Bo 1237), with only short remnants of transversal ridge tr 7. G: A. sexdentis, female (Bo 1233). H: A. sinuosocarinata, female (HT Bo 1273). I: A. aptera, male (Bo 1256). - Abbreviations: T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
31348359E8745C70B5E357A00B36B7DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563284	https://doi.org/10.3897/asp.79.e67569.figure15	Figure 15. Size and distribution of bristles in the median part of tergites T 6,7 in males (A - E) and in larval males and females (F - L) of Attaphila species, phase contrast images. A: A. bergi, male (Bo 1274). B: A. fungicola, male (LT Bo 1265). C: A. flava, male (HT Bo 1280). D: A. paucisetosa, male (Cb 2 / 1). E: A. paucisetosa, male, with slightly shorter bristles than in preceding specimen (Bo 1445). F: A. aptera, larval male (Bo 1291). G: A. aptera, larval female (Bo 1289). H: A. aptera, larval male (HT Bo 1232). I: A. bergi, larval male (Bo 1230); medium sized bristles belong to T 6, the very long ones are from T 5 covering most of T 6. J: A. bergi, larval female (Bo 1285). K: A. paucisetosa, larval male (Bo 1433). L: A. paucisetosa, larval female (Bo 1293). - Abbreviations: T 5 - p, T 6 - p, T 7 - p posterior borders of tergites T 5, T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 15. Size and distribution of bristles in the median part of tergites T 6,7 in males (A - E) and in larval males and females (F - L) of Attaphila species, phase contrast images. A: A. bergi, male (Bo 1274). B: A. fungicola, male (LT Bo 1265). C: A. flava, male (HT Bo 1280). D: A. paucisetosa, male (Cb 2 / 1). E: A. paucisetosa, male, with slightly shorter bristles than in preceding specimen (Bo 1445). F: A. aptera, larval male (Bo 1291). G: A. aptera, larval female (Bo 1289). H: A. aptera, larval male (HT Bo 1232). I: A. bergi, larval male (Bo 1230); medium sized bristles belong to T 6, the very long ones are from T 5 covering most of T 6. J: A. bergi, larval female (Bo 1285). K: A. paucisetosa, larval male (Bo 1433). L: A. paucisetosa, larval female (Bo 1293). - Abbreviations: T 5 - p, T 6 - p, T 7 - p posterior borders of tergites T 5, T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
31348359E8745C70B5E357A00B36B7DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563285	https://doi.org/10.3897/asp.79.e67569.figure16	Figure 16. Subgenital plate (sternite S 7) of females of Attaphila species. A, B: A. bergi (Bo 1275, Bo 1282). C, D: A. fungicola (Bo 1264, Bo 1228). E, F: A. paucisetosa (Cb 2 / 2, PT Bo 1255). G, H: A. schuppi (ST Bo 1237, ST Bo 1234). - Abbreviations: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars); vf 7 ventral fold of segment 7 = subgenital lobe (across entire width of S 7; approximate longitudinal extension given by double-headed arrow).	Figure 16. Subgenital plate (sternite S 7) of females of Attaphila species. A, B: A. bergi (Bo 1275, Bo 1282). C, D: A. fungicola (Bo 1264, Bo 1228). E, F: A. paucisetosa (Cb 2 / 2, PT Bo 1255). G, H: A. schuppi (ST Bo 1237, ST Bo 1234). - Abbreviations: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars); vf 7 ventral fold of segment 7 = subgenital lobe (across entire width of S 7; approximate longitudinal extension given by double-headed arrow).	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
31348359E8745C70B5E357A00B36B7DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563287	https://doi.org/10.3897/asp.79.e67569.figure18	Figure 18. Anterior half of subgenital plate (sternite S 7) of females of Attaphila species, shape of sternal transversal ridge, phase contrast images (same objects as shown in Figs 16, 17, except for Bo 1236 in F). A, B: A. aptera (Bo 1225, Bo 1253). C, D: A. bergi (Bo 1275, Bo 1282). E, F: A. fungicola (Bo 1264, Bo 1236). G: A. multisetosa (HT Bo 1270). H, I: A. paucisetosa (Cb 2 / 2, PT Bo 1255). J, K: A. schuppi (ST Bo 1234, ST Bo 1237). L: A. sexdentis (HT Bo 1233). M, N: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). - Abbreviations and symbols: sr 7 transversal ridge of subgenital plate S 7; short interruptions of this ridge indicated by arrows, larger gaps delimited by double bars.	Figure 18. Anterior half of subgenital plate (sternite S 7) of females of Attaphila species, shape of sternal transversal ridge, phase contrast images (same objects as shown in Figs 16, 17, except for Bo 1236 in F). A, B: A. aptera (Bo 1225, Bo 1253). C, D: A. bergi (Bo 1275, Bo 1282). E, F: A. fungicola (Bo 1264, Bo 1236). G: A. multisetosa (HT Bo 1270). H, I: A. paucisetosa (Cb 2 / 2, PT Bo 1255). J, K: A. schuppi (ST Bo 1234, ST Bo 1237). L: A. sexdentis (HT Bo 1233). M, N: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). - Abbreviations and symbols: sr 7 transversal ridge of subgenital plate S 7; short interruptions of this ridge indicated by arrows, larger gaps delimited by double bars.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
31348359E8745C70B5E357A00B36B7DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563289	https://doi.org/10.3897/asp.79.e67569.figure20	Figure 20. Terminalia with genitalia of females of Attaphila species, ventral view, posterior end on top. A, B: A. paucisetosa (PT Bo 1226, PT Bo 1255), A with, B without laterosternal shelf area. C, D: A. schuppi (ST Bo 1234, ST Bo 1237), C with, D without laterosternal shelf area. - Abbreviations: aa anterior arch; ca central apodeme; ce cercus; gcf fold dividing genital chamber (posterior edge labeled); gcp pouch of genital chamber (one-sided); gg-m mesal gonangulum sclerite; IC intercalary sclerite; ls laterosternal-shelf sclerite (with central part c, arm part a, wing part w, posterolateral extremity p); lst laterosternal-shelf tube; pl posterior lobe; PP paraproct; pt 8,9 extension of paratergite 9; sp spermathecal plate; T 9, T 10 abdominal tergites 9 and 10; T 10 - p posterior border of tergite 10; v 1, v 3 1 st and 3 rd valves of ovipositor; vlf valvifer of segment 8 (arrow pointing to zone where median widening starts). A articulations, A 2 between gg-m and pl, A 3 between gg-m and basal sclerotisation of v 1, A 5 between pt 8,9 and vlf. (See Supplement 1 and Supplement 3 Fig. S 2 A, B, C for additional explanations and more complete labeling using different terminologies.)	Figure 20. Terminalia with genitalia of females of Attaphila species, ventral view, posterior end on top. A, B: A. paucisetosa (PT Bo 1226, PT Bo 1255), A with, B without laterosternal shelf area. C, D: A. schuppi (ST Bo 1234, ST Bo 1237), C with, D without laterosternal shelf area. - Abbreviations: aa anterior arch; ca central apodeme; ce cercus; gcf fold dividing genital chamber (posterior edge labeled); gcp pouch of genital chamber (one-sided); gg-m mesal gonangulum sclerite; IC intercalary sclerite; ls laterosternal-shelf sclerite (with central part c, arm part a, wing part w, posterolateral extremity p); lst laterosternal-shelf tube; pl posterior lobe; PP paraproct; pt 8,9 extension of paratergite 9; sp spermathecal plate; T 9, T 10 abdominal tergites 9 and 10; T 10 - p posterior border of tergite 10; v 1, v 3 1 st and 3 rd valves of ovipositor; vlf valvifer of segment 8 (arrow pointing to zone where median widening starts). A articulations, A 2 between gg-m and pl, A 3 between gg-m and basal sclerotisation of v 1, A 5 between pt 8,9 and vlf. (See Supplement 1 and Supplement 3 Fig. S 2 A, B, C for additional explanations and more complete labeling using different terminologies.)	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
31348359E8745C70B5E357A00B36B7DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563290	https://doi.org/10.3897/asp.79.e67569.figure21	Figure 21. A-C: Terminalia with genitalia of females of Attaphila species, ventral view, posterior end on top. A: A. multisetosa (HT Bo 1270), without laterosternal shelf area, cerci damaged; inset showing median part of genital chamber dorsal wall. B: A. sinuosocarinata (HT Bo 1273), without laterosternal shelf area; inset showing median part of genital chamber dorsal wall. C: A. sexdentis (HT Bo 1233), with laterosternal shelf area. - D: Female of A. paucisetosa carrying an ootheca; length of animal ca. 3 mm. - Abbreviations: aa anterior arch; ca central apodeme; ce cercus; gcf fold dividing genital chamber (posterior edge labeled); gcp pouch of genital chamber (one-sided); gg-m mesal gonangulum sclerite; IC intercalary sclerite; ls laterosternal-shelf sclerite (with posterolateral extremity p); lst laterosternal-shelf tube; otkl ootheca keel; pl posterior lobe; PP paraproct; pt 8,9 extension of paratergite 9; sp spermathecal plate; T 9, T 10 abdominal tergites 9 and 10; T 10 - p posterior border of tergite 10; vlf valvifer of segment 8; A articulations, A 2 between gg-m and pl, A 3 between gg-m and basal sclerotisation of v 1, A 5 between pt 8,9 and vlf. Arrow in A, B showing mesal border of putative weak sclerotisation in dorsal wall of genital chamber. (See Supplement 1 and Supplement 3 Fig. S 2 A, B, C for additional explanations and more complete labeling using different terminologies.)	Figure 21. A-C: Terminalia with genitalia of females of Attaphila species, ventral view, posterior end on top. A: A. multisetosa (HT Bo 1270), without laterosternal shelf area, cerci damaged; inset showing median part of genital chamber dorsal wall. B: A. sinuosocarinata (HT Bo 1273), without laterosternal shelf area; inset showing median part of genital chamber dorsal wall. C: A. sexdentis (HT Bo 1233), with laterosternal shelf area. - D: Female of A. paucisetosa carrying an ootheca; length of animal ca. 3 mm. - Abbreviations: aa anterior arch; ca central apodeme; ce cercus; gcf fold dividing genital chamber (posterior edge labeled); gcp pouch of genital chamber (one-sided); gg-m mesal gonangulum sclerite; IC intercalary sclerite; ls laterosternal-shelf sclerite (with posterolateral extremity p); lst laterosternal-shelf tube; otkl ootheca keel; pl posterior lobe; PP paraproct; pt 8,9 extension of paratergite 9; sp spermathecal plate; T 9, T 10 abdominal tergites 9 and 10; T 10 - p posterior border of tergite 10; vlf valvifer of segment 8; A articulations, A 2 between gg-m and pl, A 3 between gg-m and basal sclerotisation of v 1, A 5 between pt 8,9 and vlf. Arrow in A, B showing mesal border of putative weak sclerotisation in dorsal wall of genital chamber. (See Supplement 1 and Supplement 3 Fig. S 2 A, B, C for additional explanations and more complete labeling using different terminologies.)	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
31348359E8745C70B5E357A00B36B7DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563291	https://doi.org/10.3897/asp.79.e67569.figure22	Figure 22. Laterosternal shelf area of females of Attaphila species, ventral view, posteriorly on top. A, B: A. aptera (Bo 1225, Bo 1257), in A left tube distorted (ventral part of tube squeezed laterally, as indicated by grey arrow). C, D: A. bergi (Bo 1282, Bo 1239). E, F: A. fungicola (Bo 1264, Bo 1447). G: A. multisetosa (HT Bo 1270). H: A. paucisetosa (PT Bo 1255). I: A. sexdentis (HT Bo 1233). J: A. schuppi (ST Bo 1237). K, L: A. sinuosocarinata (HT Bo 1273, PT Bo 1288). - Abbreviations: isf intersternal folds (of floor of vestibulum); ls laterosternal-shelf sclerite (with central part c, arm part a, wing part w, posterolateral extremity p); lst laterosternal-shelf tube; vfl floor of vestibulum. (See Supplement 3 Fig. S 2 D, E for more complete labeling using different terminologies.) - Arrows: Black arrows in A, D, I: twisting of sclerite at transition between ls-w and ls-p. Grey arrow in A indicates direction of squeezing of tube lst. Blue arrowheads: anterior extremity of wing part ls-w on mesal border of tube lst, ls-w either restricted to posterior half of tubes lst (e. g. K) or reaching far into anterior half (e. g. D). Red arrowheads: area where the posterior margin of ls-a bends dorsally, and where a bend or kink is often present in (inner) lateral border of tube lst (compare white arrowheads in Fig. 23 C, D, H, I, J). Black arrowheads: membranous inner lateral border of tube lst posterior to the bend or kink (compare Fig. 23 C, D, H, I and black arrowheads in sections beside D). - Inserts in B, D, E, K showing right (of picture and animal) tube lst with posterior margin of ls-a (red line; compare Fig. 23 C-E) and anterior margin of ls-w (blue line; compare Fig. 23 F, G).	Figure 22. Laterosternal shelf area of females of Attaphila species, ventral view, posteriorly on top. A, B: A. aptera (Bo 1225, Bo 1257), in A left tube distorted (ventral part of tube squeezed laterally, as indicated by grey arrow). C, D: A. bergi (Bo 1282, Bo 1239). E, F: A. fungicola (Bo 1264, Bo 1447). G: A. multisetosa (HT Bo 1270). H: A. paucisetosa (PT Bo 1255). I: A. sexdentis (HT Bo 1233). J: A. schuppi (ST Bo 1237). K, L: A. sinuosocarinata (HT Bo 1273, PT Bo 1288). - Abbreviations: isf intersternal folds (of floor of vestibulum); ls laterosternal-shelf sclerite (with central part c, arm part a, wing part w, posterolateral extremity p); lst laterosternal-shelf tube; vfl floor of vestibulum. (See Supplement 3 Fig. S 2 D, E for more complete labeling using different terminologies.) - Arrows: Black arrows in A, D, I: twisting of sclerite at transition between ls-w and ls-p. Grey arrow in A indicates direction of squeezing of tube lst. Blue arrowheads: anterior extremity of wing part ls-w on mesal border of tube lst, ls-w either restricted to posterior half of tubes lst (e. g. K) or reaching far into anterior half (e. g. D). Red arrowheads: area where the posterior margin of ls-a bends dorsally, and where a bend or kink is often present in (inner) lateral border of tube lst (compare white arrowheads in Fig. 23 C, D, H, I, J). Black arrowheads: membranous inner lateral border of tube lst posterior to the bend or kink (compare Fig. 23 C, D, H, I and black arrowheads in sections beside D). - Inserts in B, D, E, K showing right (of picture and animal) tube lst with posterior margin of ls-a (red line; compare Fig. 23 C-E) and anterior margin of ls-w (blue line; compare Fig. 23 F, G).	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
31348359E8745C70B5E357A00B36B7DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563294	https://doi.org/10.3897/asp.79.e67569.figure25	Figure 25. Subgenital plate (sternite S 9) and phallomeres of males of Attaphila species, dorsal view, anteriorly on top. A, B: A. paucisetosa, subgenital plate with phallomeres (PT Bo 1254, Bo 1444), hook and right phallomere removed in B. C: A. flava, subgenital plate with phallomeres (HT Bo 1280). D, E: A. fungicola. D showing subgenital plate with phallomeres (LT Bo 1265), hook not seen in profile; E showing isolated hook seen in profile (Bo 1229). - Abbreviations: T 9 p ventrally bent lateral (paratergal) part of tergite 9; otherwise as for Fig. 24.	Figure 25. Subgenital plate (sternite S 9) and phallomeres of males of Attaphila species, dorsal view, anteriorly on top. A, B: A. paucisetosa, subgenital plate with phallomeres (PT Bo 1254, Bo 1444), hook and right phallomere removed in B. C: A. flava, subgenital plate with phallomeres (HT Bo 1280). D, E: A. fungicola. D showing subgenital plate with phallomeres (LT Bo 1265), hook not seen in profile; E showing isolated hook seen in profile (Bo 1229). - Abbreviations: T 9 p ventrally bent lateral (paratergal) part of tergite 9; otherwise as for Fig. 24.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
31348359E8745C70B5E357A00B36B7DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563295	https://doi.org/10.3897/asp.79.e67569.figure26	Figure 26. Subgenital plate (sternite S 9) and phallomeres of males of Attaphila species, phase contrast images of the same objects as shown in Fig. 25 A, C, D, anteriorly on top. A: A. flava (HT Bo 1280). B: A. fungicola (LT Bo 1265). C: A. paucisetosa (PT Bo 1254). - Abbreviations: A 1 articulation between S 9 and pt 9; cs cleft sclerite of right phallomere; ea endophallic apodeme; h hook; psa process; pt 9 extension of paratergite 9 contacting sternite 9; R 3 anterior principal sclerite of right phallomere; sll left stylus; sta 9 anterior apodeme of subgenital plate S 9; vf 9 subgenital lobe; vi virga. Arrow: in A indicating that the distal part of the subgenital lobe is broken off.	Figure 26. Subgenital plate (sternite S 9) and phallomeres of males of Attaphila species, phase contrast images of the same objects as shown in Fig. 25 A, C, D, anteriorly on top. A: A. flava (HT Bo 1280). B: A. fungicola (LT Bo 1265). C: A. paucisetosa (PT Bo 1254). - Abbreviations: A 1 articulation between S 9 and pt 9; cs cleft sclerite of right phallomere; ea endophallic apodeme; h hook; psa process; pt 9 extension of paratergite 9 contacting sternite 9; R 3 anterior principal sclerite of right phallomere; sll left stylus; sta 9 anterior apodeme of subgenital plate S 9; vf 9 subgenital lobe; vi virga. Arrow: in A indicating that the distal part of the subgenital lobe is broken off.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
31348359E8745C70B5E357A00B36B7DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563280	https://doi.org/10.3897/asp.79.e67569.figure11	Figure 11. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. schuppi (ST Bo 1237). A: T 2 - 5; B: T 6,7. C, D: A. sexdentis (HT Bo 1233). C: T 1 - 5; D: T 6,7. - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to median and lateral excurvations of transversal ridges to the anterior.	Figure 11. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. schuppi (ST Bo 1237). A: T 2 - 5; B: T 6,7. C, D: A. sexdentis (HT Bo 1233). C: T 1 - 5; D: T 6,7. - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to median and lateral excurvations of transversal ridges to the anterior.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
BA4D3E16229A56FA8322212D7AC846F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563280	https://doi.org/10.3897/asp.79.e67569.figure11	Figure 11. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. schuppi (ST Bo 1237). A: T 2 - 5; B: T 6,7. C, D: A. sexdentis (HT Bo 1233). C: T 1 - 5; D: T 6,7. - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to median and lateral excurvations of transversal ridges to the anterior.	Figure 11. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. schuppi (ST Bo 1237). A: T 2 - 5; B: T 6,7. C, D: A. sexdentis (HT Bo 1233). C: T 1 - 5; D: T 6,7. - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to median and lateral excurvations of transversal ridges to the anterior.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
BA4D3E16229A56FA8322212D7AC846F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563281	https://doi.org/10.3897/asp.79.e67569.figure12	Figure 12. Abdominal tergites T 6,7 of females of Attaphila species. A: A. aptera (Bo 1257). B: A. bergi (Bo 1282). C: A. fungicola (Bo 1264). D: A. multisetosa (HT Bo 1270). E: A. paucisetosa (PT Bo 1255). F: A. schuppi (ST Bo 1237), transversal ridge tr 7 almost completely missing, only tiny remnants present near the associated bristles (see Fig. 14 F). - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 12. Abdominal tergites T 6,7 of females of Attaphila species. A: A. aptera (Bo 1257). B: A. bergi (Bo 1282). C: A. fungicola (Bo 1264). D: A. multisetosa (HT Bo 1270). E: A. paucisetosa (PT Bo 1255). F: A. schuppi (ST Bo 1237), transversal ridge tr 7 almost completely missing, only tiny remnants present near the associated bristles (see Fig. 14 F). - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
BA4D3E16229A56FA8322212D7AC846F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563283	https://doi.org/10.3897/asp.79.e67569.figure14	Figure 14. Size and distribution of bristles in the median part of tergites T 6,7 in females (A-H) and males (I) of Attaphila species, phase contrast images. A: A. aptera, female (Bo 1257). B: A. bergi, female (Bo 1282). C: A. fungicola, female (Bo 1264). D: A. multisetosa, female (Bo 1270). E: A. paucisetosa, female (PT Bo 1255). F: A. schuppi, female (ST Bo 1237), with only short remnants of transversal ridge tr 7. G: A. sexdentis, female (Bo 1233). H: A. sinuosocarinata, female (HT Bo 1273). I: A. aptera, male (Bo 1256). - Abbreviations: T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 14. Size and distribution of bristles in the median part of tergites T 6,7 in females (A-H) and males (I) of Attaphila species, phase contrast images. A: A. aptera, female (Bo 1257). B: A. bergi, female (Bo 1282). C: A. fungicola, female (Bo 1264). D: A. multisetosa, female (Bo 1270). E: A. paucisetosa, female (PT Bo 1255). F: A. schuppi, female (ST Bo 1237), with only short remnants of transversal ridge tr 7. G: A. sexdentis, female (Bo 1233). H: A. sinuosocarinata, female (HT Bo 1273). I: A. aptera, male (Bo 1256). - Abbreviations: T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
BA4D3E16229A56FA8322212D7AC846F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563285	https://doi.org/10.3897/asp.79.e67569.figure16	Figure 16. Subgenital plate (sternite S 7) of females of Attaphila species. A, B: A. bergi (Bo 1275, Bo 1282). C, D: A. fungicola (Bo 1264, Bo 1228). E, F: A. paucisetosa (Cb 2 / 2, PT Bo 1255). G, H: A. schuppi (ST Bo 1237, ST Bo 1234). - Abbreviations: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars); vf 7 ventral fold of segment 7 = subgenital lobe (across entire width of S 7; approximate longitudinal extension given by double-headed arrow).	Figure 16. Subgenital plate (sternite S 7) of females of Attaphila species. A, B: A. bergi (Bo 1275, Bo 1282). C, D: A. fungicola (Bo 1264, Bo 1228). E, F: A. paucisetosa (Cb 2 / 2, PT Bo 1255). G, H: A. schuppi (ST Bo 1237, ST Bo 1234). - Abbreviations: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars); vf 7 ventral fold of segment 7 = subgenital lobe (across entire width of S 7; approximate longitudinal extension given by double-headed arrow).	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
BA4D3E16229A56FA8322212D7AC846F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563287	https://doi.org/10.3897/asp.79.e67569.figure18	Figure 18. Anterior half of subgenital plate (sternite S 7) of females of Attaphila species, shape of sternal transversal ridge, phase contrast images (same objects as shown in Figs 16, 17, except for Bo 1236 in F). A, B: A. aptera (Bo 1225, Bo 1253). C, D: A. bergi (Bo 1275, Bo 1282). E, F: A. fungicola (Bo 1264, Bo 1236). G: A. multisetosa (HT Bo 1270). H, I: A. paucisetosa (Cb 2 / 2, PT Bo 1255). J, K: A. schuppi (ST Bo 1234, ST Bo 1237). L: A. sexdentis (HT Bo 1233). M, N: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). - Abbreviations and symbols: sr 7 transversal ridge of subgenital plate S 7; short interruptions of this ridge indicated by arrows, larger gaps delimited by double bars.	Figure 18. Anterior half of subgenital plate (sternite S 7) of females of Attaphila species, shape of sternal transversal ridge, phase contrast images (same objects as shown in Figs 16, 17, except for Bo 1236 in F). A, B: A. aptera (Bo 1225, Bo 1253). C, D: A. bergi (Bo 1275, Bo 1282). E, F: A. fungicola (Bo 1264, Bo 1236). G: A. multisetosa (HT Bo 1270). H, I: A. paucisetosa (Cb 2 / 2, PT Bo 1255). J, K: A. schuppi (ST Bo 1234, ST Bo 1237). L: A. sexdentis (HT Bo 1233). M, N: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). - Abbreviations and symbols: sr 7 transversal ridge of subgenital plate S 7; short interruptions of this ridge indicated by arrows, larger gaps delimited by double bars.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
BA4D3E16229A56FA8322212D7AC846F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563289	https://doi.org/10.3897/asp.79.e67569.figure20	Figure 20. Terminalia with genitalia of females of Attaphila species, ventral view, posterior end on top. A, B: A. paucisetosa (PT Bo 1226, PT Bo 1255), A with, B without laterosternal shelf area. C, D: A. schuppi (ST Bo 1234, ST Bo 1237), C with, D without laterosternal shelf area. - Abbreviations: aa anterior arch; ca central apodeme; ce cercus; gcf fold dividing genital chamber (posterior edge labeled); gcp pouch of genital chamber (one-sided); gg-m mesal gonangulum sclerite; IC intercalary sclerite; ls laterosternal-shelf sclerite (with central part c, arm part a, wing part w, posterolateral extremity p); lst laterosternal-shelf tube; pl posterior lobe; PP paraproct; pt 8,9 extension of paratergite 9; sp spermathecal plate; T 9, T 10 abdominal tergites 9 and 10; T 10 - p posterior border of tergite 10; v 1, v 3 1 st and 3 rd valves of ovipositor; vlf valvifer of segment 8 (arrow pointing to zone where median widening starts). A articulations, A 2 between gg-m and pl, A 3 between gg-m and basal sclerotisation of v 1, A 5 between pt 8,9 and vlf. (See Supplement 1 and Supplement 3 Fig. S 2 A, B, C for additional explanations and more complete labeling using different terminologies.)	Figure 20. Terminalia with genitalia of females of Attaphila species, ventral view, posterior end on top. A, B: A. paucisetosa (PT Bo 1226, PT Bo 1255), A with, B without laterosternal shelf area. C, D: A. schuppi (ST Bo 1234, ST Bo 1237), C with, D without laterosternal shelf area. - Abbreviations: aa anterior arch; ca central apodeme; ce cercus; gcf fold dividing genital chamber (posterior edge labeled); gcp pouch of genital chamber (one-sided); gg-m mesal gonangulum sclerite; IC intercalary sclerite; ls laterosternal-shelf sclerite (with central part c, arm part a, wing part w, posterolateral extremity p); lst laterosternal-shelf tube; pl posterior lobe; PP paraproct; pt 8,9 extension of paratergite 9; sp spermathecal plate; T 9, T 10 abdominal tergites 9 and 10; T 10 - p posterior border of tergite 10; v 1, v 3 1 st and 3 rd valves of ovipositor; vlf valvifer of segment 8 (arrow pointing to zone where median widening starts). A articulations, A 2 between gg-m and pl, A 3 between gg-m and basal sclerotisation of v 1, A 5 between pt 8,9 and vlf. (See Supplement 1 and Supplement 3 Fig. S 2 A, B, C for additional explanations and more complete labeling using different terminologies.)	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
BA4D3E16229A56FA8322212D7AC846F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563291	https://doi.org/10.3897/asp.79.e67569.figure22	Figure 22. Laterosternal shelf area of females of Attaphila species, ventral view, posteriorly on top. A, B: A. aptera (Bo 1225, Bo 1257), in A left tube distorted (ventral part of tube squeezed laterally, as indicated by grey arrow). C, D: A. bergi (Bo 1282, Bo 1239). E, F: A. fungicola (Bo 1264, Bo 1447). G: A. multisetosa (HT Bo 1270). H: A. paucisetosa (PT Bo 1255). I: A. sexdentis (HT Bo 1233). J: A. schuppi (ST Bo 1237). K, L: A. sinuosocarinata (HT Bo 1273, PT Bo 1288). - Abbreviations: isf intersternal folds (of floor of vestibulum); ls laterosternal-shelf sclerite (with central part c, arm part a, wing part w, posterolateral extremity p); lst laterosternal-shelf tube; vfl floor of vestibulum. (See Supplement 3 Fig. S 2 D, E for more complete labeling using different terminologies.) - Arrows: Black arrows in A, D, I: twisting of sclerite at transition between ls-w and ls-p. Grey arrow in A indicates direction of squeezing of tube lst. Blue arrowheads: anterior extremity of wing part ls-w on mesal border of tube lst, ls-w either restricted to posterior half of tubes lst (e. g. K) or reaching far into anterior half (e. g. D). Red arrowheads: area where the posterior margin of ls-a bends dorsally, and where a bend or kink is often present in (inner) lateral border of tube lst (compare white arrowheads in Fig. 23 C, D, H, I, J). Black arrowheads: membranous inner lateral border of tube lst posterior to the bend or kink (compare Fig. 23 C, D, H, I and black arrowheads in sections beside D). - Inserts in B, D, E, K showing right (of picture and animal) tube lst with posterior margin of ls-a (red line; compare Fig. 23 C-E) and anterior margin of ls-w (blue line; compare Fig. 23 F, G).	Figure 22. Laterosternal shelf area of females of Attaphila species, ventral view, posteriorly on top. A, B: A. aptera (Bo 1225, Bo 1257), in A left tube distorted (ventral part of tube squeezed laterally, as indicated by grey arrow). C, D: A. bergi (Bo 1282, Bo 1239). E, F: A. fungicola (Bo 1264, Bo 1447). G: A. multisetosa (HT Bo 1270). H: A. paucisetosa (PT Bo 1255). I: A. sexdentis (HT Bo 1233). J: A. schuppi (ST Bo 1237). K, L: A. sinuosocarinata (HT Bo 1273, PT Bo 1288). - Abbreviations: isf intersternal folds (of floor of vestibulum); ls laterosternal-shelf sclerite (with central part c, arm part a, wing part w, posterolateral extremity p); lst laterosternal-shelf tube; vfl floor of vestibulum. (See Supplement 3 Fig. S 2 D, E for more complete labeling using different terminologies.) - Arrows: Black arrows in A, D, I: twisting of sclerite at transition between ls-w and ls-p. Grey arrow in A indicates direction of squeezing of tube lst. Blue arrowheads: anterior extremity of wing part ls-w on mesal border of tube lst, ls-w either restricted to posterior half of tubes lst (e. g. K) or reaching far into anterior half (e. g. D). Red arrowheads: area where the posterior margin of ls-a bends dorsally, and where a bend or kink is often present in (inner) lateral border of tube lst (compare white arrowheads in Fig. 23 C, D, H, I, J). Black arrowheads: membranous inner lateral border of tube lst posterior to the bend or kink (compare Fig. 23 C, D, H, I and black arrowheads in sections beside D). - Inserts in B, D, E, K showing right (of picture and animal) tube lst with posterior margin of ls-a (red line; compare Fig. 23 C-E) and anterior margin of ls-w (blue line; compare Fig. 23 F, G).	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
BA4D3E16229A56FA8322212D7AC846F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563279	https://doi.org/10.3897/asp.79.e67569.figure10	Figure 10. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. multisetosa (HT Bo 1270). A: T 2 - 5; B: T 6,7. C - E: A. sinuosocarinata. C: T 1 - 5 (HT Bo 1273); D: T 6,7 (HT Bo 1273); E: T 6 (PT Bo 1288). - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to weak median and lateral excurvations of transversal ridges to the anterior.	Figure 10. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. multisetosa (HT Bo 1270). A: T 2 - 5; B: T 6,7. C - E: A. sinuosocarinata. C: T 1 - 5 (HT Bo 1273); D: T 6,7 (HT Bo 1273); E: T 6 (PT Bo 1288). - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to weak median and lateral excurvations of transversal ridges to the anterior.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
7A973A062FFE514487AED823723C301D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563280	https://doi.org/10.3897/asp.79.e67569.figure11	Figure 11. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. schuppi (ST Bo 1237). A: T 2 - 5; B: T 6,7. C, D: A. sexdentis (HT Bo 1233). C: T 1 - 5; D: T 6,7. - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to median and lateral excurvations of transversal ridges to the anterior.	Figure 11. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. schuppi (ST Bo 1237). A: T 2 - 5; B: T 6,7. C, D: A. sexdentis (HT Bo 1233). C: T 1 - 5; D: T 6,7. - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to median and lateral excurvations of transversal ridges to the anterior.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
7A973A062FFE514487AED823723C301D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563282	https://doi.org/10.3897/asp.79.e67569.figure13	Figure 13. Abdominal tergites T 6,7 of females (A, B) and males (C - F) of Attaphila species. A: A. sexdentis, female (Bo 1233). B: A. sinuosocarinata, female (HT Bo 1273). C: A. aptera, male (Bo 1256). D: A. bergi, male (Bo 1274), transversal ridge tr 7 weakly developed, sublaterally with a large gap. E: A. paucisetosa, male (Bo 1240), transversal ridge tr 7 missing. F: A. fungicola, male (Bo 1229), transversal ridge tr 7 missing. - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 13. Abdominal tergites T 6,7 of females (A, B) and males (C - F) of Attaphila species. A: A. sexdentis, female (Bo 1233). B: A. sinuosocarinata, female (HT Bo 1273). C: A. aptera, male (Bo 1256). D: A. bergi, male (Bo 1274), transversal ridge tr 7 weakly developed, sublaterally with a large gap. E: A. paucisetosa, male (Bo 1240), transversal ridge tr 7 missing. F: A. fungicola, male (Bo 1229), transversal ridge tr 7 missing. - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
7A973A062FFE514487AED823723C301D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563283	https://doi.org/10.3897/asp.79.e67569.figure14	Figure 14. Size and distribution of bristles in the median part of tergites T 6,7 in females (A-H) and males (I) of Attaphila species, phase contrast images. A: A. aptera, female (Bo 1257). B: A. bergi, female (Bo 1282). C: A. fungicola, female (Bo 1264). D: A. multisetosa, female (Bo 1270). E: A. paucisetosa, female (PT Bo 1255). F: A. schuppi, female (ST Bo 1237), with only short remnants of transversal ridge tr 7. G: A. sexdentis, female (Bo 1233). H: A. sinuosocarinata, female (HT Bo 1273). I: A. aptera, male (Bo 1256). - Abbreviations: T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 14. Size and distribution of bristles in the median part of tergites T 6,7 in females (A-H) and males (I) of Attaphila species, phase contrast images. A: A. aptera, female (Bo 1257). B: A. bergi, female (Bo 1282). C: A. fungicola, female (Bo 1264). D: A. multisetosa, female (Bo 1270). E: A. paucisetosa, female (PT Bo 1255). F: A. schuppi, female (ST Bo 1237), with only short remnants of transversal ridge tr 7. G: A. sexdentis, female (Bo 1233). H: A. sinuosocarinata, female (HT Bo 1273). I: A. aptera, male (Bo 1256). - Abbreviations: T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
7A973A062FFE514487AED823723C301D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563286	https://doi.org/10.3897/asp.79.e67569.figure17	Figure 17. Subgenital plate (sternite S 7) of females (A - F) and larval females (G, H) of Attaphila species. A, B: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). C, D: A. aptera (Bo 1225, Bo 1253). E: A. multisetosa (HT Bo 1270). F: A. sexdentis (HT Bo 1233). G: A. aptera, larval female (Bo 1289). H: A. bergi, larval female (Bo 1231). - Abbreviations and arrows: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars). Arrows in B, C, F pointing to lateral borders of S 7, which are parallel or slightly divergent (to the anterior) in A-E, but convergent in F.	Figure 17. Subgenital plate (sternite S 7) of females (A - F) and larval females (G, H) of Attaphila species. A, B: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). C, D: A. aptera (Bo 1225, Bo 1253). E: A. multisetosa (HT Bo 1270). F: A. sexdentis (HT Bo 1233). G: A. aptera, larval female (Bo 1289). H: A. bergi, larval female (Bo 1231). - Abbreviations and arrows: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars). Arrows in B, C, F pointing to lateral borders of S 7, which are parallel or slightly divergent (to the anterior) in A-E, but convergent in F.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
7A973A062FFE514487AED823723C301D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563287	https://doi.org/10.3897/asp.79.e67569.figure18	Figure 18. Anterior half of subgenital plate (sternite S 7) of females of Attaphila species, shape of sternal transversal ridge, phase contrast images (same objects as shown in Figs 16, 17, except for Bo 1236 in F). A, B: A. aptera (Bo 1225, Bo 1253). C, D: A. bergi (Bo 1275, Bo 1282). E, F: A. fungicola (Bo 1264, Bo 1236). G: A. multisetosa (HT Bo 1270). H, I: A. paucisetosa (Cb 2 / 2, PT Bo 1255). J, K: A. schuppi (ST Bo 1234, ST Bo 1237). L: A. sexdentis (HT Bo 1233). M, N: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). - Abbreviations and symbols: sr 7 transversal ridge of subgenital plate S 7; short interruptions of this ridge indicated by arrows, larger gaps delimited by double bars.	Figure 18. Anterior half of subgenital plate (sternite S 7) of females of Attaphila species, shape of sternal transversal ridge, phase contrast images (same objects as shown in Figs 16, 17, except for Bo 1236 in F). A, B: A. aptera (Bo 1225, Bo 1253). C, D: A. bergi (Bo 1275, Bo 1282). E, F: A. fungicola (Bo 1264, Bo 1236). G: A. multisetosa (HT Bo 1270). H, I: A. paucisetosa (Cb 2 / 2, PT Bo 1255). J, K: A. schuppi (ST Bo 1234, ST Bo 1237). L: A. sexdentis (HT Bo 1233). M, N: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). - Abbreviations and symbols: sr 7 transversal ridge of subgenital plate S 7; short interruptions of this ridge indicated by arrows, larger gaps delimited by double bars.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
7A973A062FFE514487AED823723C301D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563290	https://doi.org/10.3897/asp.79.e67569.figure21	Figure 21. A-C: Terminalia with genitalia of females of Attaphila species, ventral view, posterior end on top. A: A. multisetosa (HT Bo 1270), without laterosternal shelf area, cerci damaged; inset showing median part of genital chamber dorsal wall. B: A. sinuosocarinata (HT Bo 1273), without laterosternal shelf area; inset showing median part of genital chamber dorsal wall. C: A. sexdentis (HT Bo 1233), with laterosternal shelf area. - D: Female of A. paucisetosa carrying an ootheca; length of animal ca. 3 mm. - Abbreviations: aa anterior arch; ca central apodeme; ce cercus; gcf fold dividing genital chamber (posterior edge labeled); gcp pouch of genital chamber (one-sided); gg-m mesal gonangulum sclerite; IC intercalary sclerite; ls laterosternal-shelf sclerite (with posterolateral extremity p); lst laterosternal-shelf tube; otkl ootheca keel; pl posterior lobe; PP paraproct; pt 8,9 extension of paratergite 9; sp spermathecal plate; T 9, T 10 abdominal tergites 9 and 10; T 10 - p posterior border of tergite 10; vlf valvifer of segment 8; A articulations, A 2 between gg-m and pl, A 3 between gg-m and basal sclerotisation of v 1, A 5 between pt 8,9 and vlf. Arrow in A, B showing mesal border of putative weak sclerotisation in dorsal wall of genital chamber. (See Supplement 1 and Supplement 3 Fig. S 2 A, B, C for additional explanations and more complete labeling using different terminologies.)	Figure 21. A-C: Terminalia with genitalia of females of Attaphila species, ventral view, posterior end on top. A: A. multisetosa (HT Bo 1270), without laterosternal shelf area, cerci damaged; inset showing median part of genital chamber dorsal wall. B: A. sinuosocarinata (HT Bo 1273), without laterosternal shelf area; inset showing median part of genital chamber dorsal wall. C: A. sexdentis (HT Bo 1233), with laterosternal shelf area. - D: Female of A. paucisetosa carrying an ootheca; length of animal ca. 3 mm. - Abbreviations: aa anterior arch; ca central apodeme; ce cercus; gcf fold dividing genital chamber (posterior edge labeled); gcp pouch of genital chamber (one-sided); gg-m mesal gonangulum sclerite; IC intercalary sclerite; ls laterosternal-shelf sclerite (with posterolateral extremity p); lst laterosternal-shelf tube; otkl ootheca keel; pl posterior lobe; PP paraproct; pt 8,9 extension of paratergite 9; sp spermathecal plate; T 9, T 10 abdominal tergites 9 and 10; T 10 - p posterior border of tergite 10; vlf valvifer of segment 8; A articulations, A 2 between gg-m and pl, A 3 between gg-m and basal sclerotisation of v 1, A 5 between pt 8,9 and vlf. Arrow in A, B showing mesal border of putative weak sclerotisation in dorsal wall of genital chamber. (See Supplement 1 and Supplement 3 Fig. S 2 A, B, C for additional explanations and more complete labeling using different terminologies.)	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
7A973A062FFE514487AED823723C301D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563291	https://doi.org/10.3897/asp.79.e67569.figure22	Figure 22. Laterosternal shelf area of females of Attaphila species, ventral view, posteriorly on top. A, B: A. aptera (Bo 1225, Bo 1257), in A left tube distorted (ventral part of tube squeezed laterally, as indicated by grey arrow). C, D: A. bergi (Bo 1282, Bo 1239). E, F: A. fungicola (Bo 1264, Bo 1447). G: A. multisetosa (HT Bo 1270). H: A. paucisetosa (PT Bo 1255). I: A. sexdentis (HT Bo 1233). J: A. schuppi (ST Bo 1237). K, L: A. sinuosocarinata (HT Bo 1273, PT Bo 1288). - Abbreviations: isf intersternal folds (of floor of vestibulum); ls laterosternal-shelf sclerite (with central part c, arm part a, wing part w, posterolateral extremity p); lst laterosternal-shelf tube; vfl floor of vestibulum. (See Supplement 3 Fig. S 2 D, E for more complete labeling using different terminologies.) - Arrows: Black arrows in A, D, I: twisting of sclerite at transition between ls-w and ls-p. Grey arrow in A indicates direction of squeezing of tube lst. Blue arrowheads: anterior extremity of wing part ls-w on mesal border of tube lst, ls-w either restricted to posterior half of tubes lst (e. g. K) or reaching far into anterior half (e. g. D). Red arrowheads: area where the posterior margin of ls-a bends dorsally, and where a bend or kink is often present in (inner) lateral border of tube lst (compare white arrowheads in Fig. 23 C, D, H, I, J). Black arrowheads: membranous inner lateral border of tube lst posterior to the bend or kink (compare Fig. 23 C, D, H, I and black arrowheads in sections beside D). - Inserts in B, D, E, K showing right (of picture and animal) tube lst with posterior margin of ls-a (red line; compare Fig. 23 C-E) and anterior margin of ls-w (blue line; compare Fig. 23 F, G).	Figure 22. Laterosternal shelf area of females of Attaphila species, ventral view, posteriorly on top. A, B: A. aptera (Bo 1225, Bo 1257), in A left tube distorted (ventral part of tube squeezed laterally, as indicated by grey arrow). C, D: A. bergi (Bo 1282, Bo 1239). E, F: A. fungicola (Bo 1264, Bo 1447). G: A. multisetosa (HT Bo 1270). H: A. paucisetosa (PT Bo 1255). I: A. sexdentis (HT Bo 1233). J: A. schuppi (ST Bo 1237). K, L: A. sinuosocarinata (HT Bo 1273, PT Bo 1288). - Abbreviations: isf intersternal folds (of floor of vestibulum); ls laterosternal-shelf sclerite (with central part c, arm part a, wing part w, posterolateral extremity p); lst laterosternal-shelf tube; vfl floor of vestibulum. (See Supplement 3 Fig. S 2 D, E for more complete labeling using different terminologies.) - Arrows: Black arrows in A, D, I: twisting of sclerite at transition between ls-w and ls-p. Grey arrow in A indicates direction of squeezing of tube lst. Blue arrowheads: anterior extremity of wing part ls-w on mesal border of tube lst, ls-w either restricted to posterior half of tubes lst (e. g. K) or reaching far into anterior half (e. g. D). Red arrowheads: area where the posterior margin of ls-a bends dorsally, and where a bend or kink is often present in (inner) lateral border of tube lst (compare white arrowheads in Fig. 23 C, D, H, I, J). Black arrowheads: membranous inner lateral border of tube lst posterior to the bend or kink (compare Fig. 23 C, D, H, I and black arrowheads in sections beside D). - Inserts in B, D, E, K showing right (of picture and animal) tube lst with posterior margin of ls-a (red line; compare Fig. 23 C-E) and anterior margin of ls-w (blue line; compare Fig. 23 F, G).	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
60B57AB4EDE85E7E9DFC9726AC297AEB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563279	https://doi.org/10.3897/asp.79.e67569.figure10	Figure 10. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. multisetosa (HT Bo 1270). A: T 2 - 5; B: T 6,7. C - E: A. sinuosocarinata. C: T 1 - 5 (HT Bo 1273); D: T 6,7 (HT Bo 1273); E: T 6 (PT Bo 1288). - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to weak median and lateral excurvations of transversal ridges to the anterior.	Figure 10. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. multisetosa (HT Bo 1270). A: T 2 - 5; B: T 6,7. C - E: A. sinuosocarinata. C: T 1 - 5 (HT Bo 1273); D: T 6,7 (HT Bo 1273); E: T 6 (PT Bo 1288). - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to weak median and lateral excurvations of transversal ridges to the anterior.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
60B57AB4EDE85E7E9DFC9726AC297AEB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563282	https://doi.org/10.3897/asp.79.e67569.figure13	Figure 13. Abdominal tergites T 6,7 of females (A, B) and males (C - F) of Attaphila species. A: A. sexdentis, female (Bo 1233). B: A. sinuosocarinata, female (HT Bo 1273). C: A. aptera, male (Bo 1256). D: A. bergi, male (Bo 1274), transversal ridge tr 7 weakly developed, sublaterally with a large gap. E: A. paucisetosa, male (Bo 1240), transversal ridge tr 7 missing. F: A. fungicola, male (Bo 1229), transversal ridge tr 7 missing. - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 13. Abdominal tergites T 6,7 of females (A, B) and males (C - F) of Attaphila species. A: A. sexdentis, female (Bo 1233). B: A. sinuosocarinata, female (HT Bo 1273). C: A. aptera, male (Bo 1256). D: A. bergi, male (Bo 1274), transversal ridge tr 7 weakly developed, sublaterally with a large gap. E: A. paucisetosa, male (Bo 1240), transversal ridge tr 7 missing. F: A. fungicola, male (Bo 1229), transversal ridge tr 7 missing. - Abbreviations: T 6, T 7 tergites T 6, T 7; T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
60B57AB4EDE85E7E9DFC9726AC297AEB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563283	https://doi.org/10.3897/asp.79.e67569.figure14	Figure 14. Size and distribution of bristles in the median part of tergites T 6,7 in females (A-H) and males (I) of Attaphila species, phase contrast images. A: A. aptera, female (Bo 1257). B: A. bergi, female (Bo 1282). C: A. fungicola, female (Bo 1264). D: A. multisetosa, female (Bo 1270). E: A. paucisetosa, female (PT Bo 1255). F: A. schuppi, female (ST Bo 1237), with only short remnants of transversal ridge tr 7. G: A. sexdentis, female (Bo 1233). H: A. sinuosocarinata, female (HT Bo 1273). I: A. aptera, male (Bo 1256). - Abbreviations: T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	Figure 14. Size and distribution of bristles in the median part of tergites T 6,7 in females (A-H) and males (I) of Attaphila species, phase contrast images. A: A. aptera, female (Bo 1257). B: A. bergi, female (Bo 1282). C: A. fungicola, female (Bo 1264). D: A. multisetosa, female (Bo 1270). E: A. paucisetosa, female (PT Bo 1255). F: A. schuppi, female (ST Bo 1237), with only short remnants of transversal ridge tr 7. G: A. sexdentis, female (Bo 1233). H: A. sinuosocarinata, female (HT Bo 1273). I: A. aptera, male (Bo 1256). - Abbreviations: T 6 - p, T 7 - p posterior borders of tergites T 6, T 7; tr 6, tr 7 transversal ridges of tergites T 6, T 7.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
60B57AB4EDE85E7E9DFC9726AC297AEB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563286	https://doi.org/10.3897/asp.79.e67569.figure17	Figure 17. Subgenital plate (sternite S 7) of females (A - F) and larval females (G, H) of Attaphila species. A, B: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). C, D: A. aptera (Bo 1225, Bo 1253). E: A. multisetosa (HT Bo 1270). F: A. sexdentis (HT Bo 1233). G: A. aptera, larval female (Bo 1289). H: A. bergi, larval female (Bo 1231). - Abbreviations and arrows: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars). Arrows in B, C, F pointing to lateral borders of S 7, which are parallel or slightly divergent (to the anterior) in A-E, but convergent in F.	Figure 17. Subgenital plate (sternite S 7) of females (A - F) and larval females (G, H) of Attaphila species. A, B: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). C, D: A. aptera (Bo 1225, Bo 1253). E: A. multisetosa (HT Bo 1270). F: A. sexdentis (HT Bo 1233). G: A. aptera, larval female (Bo 1289). H: A. bergi, larval female (Bo 1231). - Abbreviations and arrows: S 7 - p posterior border of subgenital plate S 7; sr 7 - l, sr 7 - m lateral, median part of transversal ridge of subgenital plate S 7 (borders indicated by double bars). Arrows in B, C, F pointing to lateral borders of S 7, which are parallel or slightly divergent (to the anterior) in A-E, but convergent in F.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
60B57AB4EDE85E7E9DFC9726AC297AEB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563287	https://doi.org/10.3897/asp.79.e67569.figure18	Figure 18. Anterior half of subgenital plate (sternite S 7) of females of Attaphila species, shape of sternal transversal ridge, phase contrast images (same objects as shown in Figs 16, 17, except for Bo 1236 in F). A, B: A. aptera (Bo 1225, Bo 1253). C, D: A. bergi (Bo 1275, Bo 1282). E, F: A. fungicola (Bo 1264, Bo 1236). G: A. multisetosa (HT Bo 1270). H, I: A. paucisetosa (Cb 2 / 2, PT Bo 1255). J, K: A. schuppi (ST Bo 1234, ST Bo 1237). L: A. sexdentis (HT Bo 1233). M, N: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). - Abbreviations and symbols: sr 7 transversal ridge of subgenital plate S 7; short interruptions of this ridge indicated by arrows, larger gaps delimited by double bars.	Figure 18. Anterior half of subgenital plate (sternite S 7) of females of Attaphila species, shape of sternal transversal ridge, phase contrast images (same objects as shown in Figs 16, 17, except for Bo 1236 in F). A, B: A. aptera (Bo 1225, Bo 1253). C, D: A. bergi (Bo 1275, Bo 1282). E, F: A. fungicola (Bo 1264, Bo 1236). G: A. multisetosa (HT Bo 1270). H, I: A. paucisetosa (Cb 2 / 2, PT Bo 1255). J, K: A. schuppi (ST Bo 1234, ST Bo 1237). L: A. sexdentis (HT Bo 1233). M, N: A. sinuosocarinata (PT Bo 1287, HT Bo 1273). - Abbreviations and symbols: sr 7 transversal ridge of subgenital plate S 7; short interruptions of this ridge indicated by arrows, larger gaps delimited by double bars.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
60B57AB4EDE85E7E9DFC9726AC297AEB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563290	https://doi.org/10.3897/asp.79.e67569.figure21	Figure 21. A-C: Terminalia with genitalia of females of Attaphila species, ventral view, posterior end on top. A: A. multisetosa (HT Bo 1270), without laterosternal shelf area, cerci damaged; inset showing median part of genital chamber dorsal wall. B: A. sinuosocarinata (HT Bo 1273), without laterosternal shelf area; inset showing median part of genital chamber dorsal wall. C: A. sexdentis (HT Bo 1233), with laterosternal shelf area. - D: Female of A. paucisetosa carrying an ootheca; length of animal ca. 3 mm. - Abbreviations: aa anterior arch; ca central apodeme; ce cercus; gcf fold dividing genital chamber (posterior edge labeled); gcp pouch of genital chamber (one-sided); gg-m mesal gonangulum sclerite; IC intercalary sclerite; ls laterosternal-shelf sclerite (with posterolateral extremity p); lst laterosternal-shelf tube; otkl ootheca keel; pl posterior lobe; PP paraproct; pt 8,9 extension of paratergite 9; sp spermathecal plate; T 9, T 10 abdominal tergites 9 and 10; T 10 - p posterior border of tergite 10; vlf valvifer of segment 8; A articulations, A 2 between gg-m and pl, A 3 between gg-m and basal sclerotisation of v 1, A 5 between pt 8,9 and vlf. Arrow in A, B showing mesal border of putative weak sclerotisation in dorsal wall of genital chamber. (See Supplement 1 and Supplement 3 Fig. S 2 A, B, C for additional explanations and more complete labeling using different terminologies.)	Figure 21. A-C: Terminalia with genitalia of females of Attaphila species, ventral view, posterior end on top. A: A. multisetosa (HT Bo 1270), without laterosternal shelf area, cerci damaged; inset showing median part of genital chamber dorsal wall. B: A. sinuosocarinata (HT Bo 1273), without laterosternal shelf area; inset showing median part of genital chamber dorsal wall. C: A. sexdentis (HT Bo 1233), with laterosternal shelf area. - D: Female of A. paucisetosa carrying an ootheca; length of animal ca. 3 mm. - Abbreviations: aa anterior arch; ca central apodeme; ce cercus; gcf fold dividing genital chamber (posterior edge labeled); gcp pouch of genital chamber (one-sided); gg-m mesal gonangulum sclerite; IC intercalary sclerite; ls laterosternal-shelf sclerite (with posterolateral extremity p); lst laterosternal-shelf tube; otkl ootheca keel; pl posterior lobe; PP paraproct; pt 8,9 extension of paratergite 9; sp spermathecal plate; T 9, T 10 abdominal tergites 9 and 10; T 10 - p posterior border of tergite 10; vlf valvifer of segment 8; A articulations, A 2 between gg-m and pl, A 3 between gg-m and basal sclerotisation of v 1, A 5 between pt 8,9 and vlf. Arrow in A, B showing mesal border of putative weak sclerotisation in dorsal wall of genital chamber. (See Supplement 1 and Supplement 3 Fig. S 2 A, B, C for additional explanations and more complete labeling using different terminologies.)	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
60B57AB4EDE85E7E9DFC9726AC297AEB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563291	https://doi.org/10.3897/asp.79.e67569.figure22	Figure 22. Laterosternal shelf area of females of Attaphila species, ventral view, posteriorly on top. A, B: A. aptera (Bo 1225, Bo 1257), in A left tube distorted (ventral part of tube squeezed laterally, as indicated by grey arrow). C, D: A. bergi (Bo 1282, Bo 1239). E, F: A. fungicola (Bo 1264, Bo 1447). G: A. multisetosa (HT Bo 1270). H: A. paucisetosa (PT Bo 1255). I: A. sexdentis (HT Bo 1233). J: A. schuppi (ST Bo 1237). K, L: A. sinuosocarinata (HT Bo 1273, PT Bo 1288). - Abbreviations: isf intersternal folds (of floor of vestibulum); ls laterosternal-shelf sclerite (with central part c, arm part a, wing part w, posterolateral extremity p); lst laterosternal-shelf tube; vfl floor of vestibulum. (See Supplement 3 Fig. S 2 D, E for more complete labeling using different terminologies.) - Arrows: Black arrows in A, D, I: twisting of sclerite at transition between ls-w and ls-p. Grey arrow in A indicates direction of squeezing of tube lst. Blue arrowheads: anterior extremity of wing part ls-w on mesal border of tube lst, ls-w either restricted to posterior half of tubes lst (e. g. K) or reaching far into anterior half (e. g. D). Red arrowheads: area where the posterior margin of ls-a bends dorsally, and where a bend or kink is often present in (inner) lateral border of tube lst (compare white arrowheads in Fig. 23 C, D, H, I, J). Black arrowheads: membranous inner lateral border of tube lst posterior to the bend or kink (compare Fig. 23 C, D, H, I and black arrowheads in sections beside D). - Inserts in B, D, E, K showing right (of picture and animal) tube lst with posterior margin of ls-a (red line; compare Fig. 23 C-E) and anterior margin of ls-w (blue line; compare Fig. 23 F, G).	Figure 22. Laterosternal shelf area of females of Attaphila species, ventral view, posteriorly on top. A, B: A. aptera (Bo 1225, Bo 1257), in A left tube distorted (ventral part of tube squeezed laterally, as indicated by grey arrow). C, D: A. bergi (Bo 1282, Bo 1239). E, F: A. fungicola (Bo 1264, Bo 1447). G: A. multisetosa (HT Bo 1270). H: A. paucisetosa (PT Bo 1255). I: A. sexdentis (HT Bo 1233). J: A. schuppi (ST Bo 1237). K, L: A. sinuosocarinata (HT Bo 1273, PT Bo 1288). - Abbreviations: isf intersternal folds (of floor of vestibulum); ls laterosternal-shelf sclerite (with central part c, arm part a, wing part w, posterolateral extremity p); lst laterosternal-shelf tube; vfl floor of vestibulum. (See Supplement 3 Fig. S 2 D, E for more complete labeling using different terminologies.) - Arrows: Black arrows in A, D, I: twisting of sclerite at transition between ls-w and ls-p. Grey arrow in A indicates direction of squeezing of tube lst. Blue arrowheads: anterior extremity of wing part ls-w on mesal border of tube lst, ls-w either restricted to posterior half of tubes lst (e. g. K) or reaching far into anterior half (e. g. D). Red arrowheads: area where the posterior margin of ls-a bends dorsally, and where a bend or kink is often present in (inner) lateral border of tube lst (compare white arrowheads in Fig. 23 C, D, H, I, J). Black arrowheads: membranous inner lateral border of tube lst posterior to the bend or kink (compare Fig. 23 C, D, H, I and black arrowheads in sections beside D). - Inserts in B, D, E, K showing right (of picture and animal) tube lst with posterior margin of ls-a (red line; compare Fig. 23 C-E) and anterior margin of ls-w (blue line; compare Fig. 23 F, G).	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
60B57AB4EDE85E7E9DFC9726AC297AEB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/563280	https://doi.org/10.3897/asp.79.e67569.figure11	Figure 11. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. schuppi (ST Bo 1237). A: T 2 - 5; B: T 6,7. C, D: A. sexdentis (HT Bo 1233). C: T 1 - 5; D: T 6,7. - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to median and lateral excurvations of transversal ridges to the anterior.	Figure 11. Abdominal tergites of Attaphila species only known from the female sex. A, B: A. schuppi (ST Bo 1237). A: T 2 - 5; B: T 6,7. C, D: A. sexdentis (HT Bo 1233). C: T 1 - 5; D: T 6,7. - Abbreviations: T n tergite (numbered); T n - a anterior border of tergite (numbered); T n - p posterior border of tergite (numbered); tr n transversal ridge of tergite (numbered). Grey arrows pointing to median and lateral excurvations of transversal ridges to the anterior.	2021-07-05	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter		Zenodo	biologists	Bohn, Horst;Nehring, Volker;G., Jonathan Rodriguez;Klass, Klaus-Dieter			
