identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
891387A9FFC5FFCE13A6FBF341D97CB1.text	891387A9FFC5FFCE13A6FBF341D97CB1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta flisa	<div><p>1 Catasticta flisa (Herrich-Schäffer, [1858]) (TL: Mexico)</p><p>(fig. 3: a-i; fig. 12)</p><p>Catasticta flisa is the most common and widespread species of the genus, occurring in a wide variety of habitats and distributed over more than 50% of the known range of the entire genus. Two subspecies were previously known for the country: Catasticta flisa briseis Eitschberger &amp; Racheli, 1998 (TL: Venezuela, Aragua, Maracay), distributed from 1000 to 2200 meters in the Cordillera de la Costa, from Yaracuy to Miranda, and Catasticta flisa dilutior Avinoff, 1926 (TL: Colombia, Chaochi [= Choachi]) (= C. hermione Reissinger, 1972), which is quite common from Trujillo south-westwards to Táchira.</p><p>The species was recently collected for the first time in the Serranía del Turimiquire (Sucre and Anzoátegui), where it is quite scarce. Although only a few specimens were available, we describe a new subspecies to highlight their peculiar pattern and isolated distribution.</p></div>	https://treatment.plazi.org/id/891387A9FFC5FFCE13A6FBF341D97CB1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFC5FFCF13A6FA0E46F278DC.text	891387A9FFC5FFCF13A6FA0E46F278DC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta flisa subsp. viloriai	<div><p>Catasticta flisa viloriai ssp. nov.</p><p>Holotype: male, Cerro Quiriquire (S. Antonio), Estado Monagas, Venezuela, 1500m, 14-VIII-2005, col. Costa, in MIZA (fig. 3: g-h)</p><p>Description: FW length: 27 mm.</p><p>Dorsal surface: blackish ground-color with white marking. A series of evident premarginal white dots on either FWs and HWs, the apical ones of FWs clearly elongated; submarginal white spots of HWs elongated. Ventral surface: FWs with white discal spots highly suffused by dark scales; premarginal yellow spots very elongated, nearly merged with submarginal ones; HWs with submarginal yellow interneural spots elongated.</p><p>Diagnostic characters: the new subspecies is characterized by strongly elongated white submarginal spots of either dorsal and ventral FWs and HWs, thus permitting an immediate visual recognition, and approaching ssp. duna Eitschberger &amp; Racheli, 1998 from North-Western Ecuador. In nearby ssp. briseis, corresponding spots are always small and rounded.</p><p>Paratypes (4 males, 1 female): 1 male, same data as HT, but 22-VII-2005; 1 female, same data as HT, but 1200 m., 31-III-2005, all in MCCV; 1 male, same data as HT, but 20-VII-2005, DNA sample n° 146; 1 male, same data as HT, but 14-VIII-2005, all in MBLI; 1 male, same data as HT, but 21-VII-2005, in FRSV. Distribution: the new subspecies is restricted to Serranía del Turimiquire, where it is quite scarce and localized. Apparently C. flisa is the only known species of Catasticta in the area.</p><p>Etymology: a noun in the genitive case after Angel Viloria (Maracaibo, Venezuela) for his rediscovery of the interesting butterfly fauna of Serranía del Turimiquire (Viloria &amp; Camacho, 1999).</p></div>	https://treatment.plazi.org/id/891387A9FFC5FFCF13A6FA0E46F278DC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFC4FFCF13A6FE0446BF7AC2.text	891387A9FFC4FFCF13A6FE0446BF7AC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta sisamnus (Fabricius 1793) Fabricius 1793	<div><p>2 Catasticta sisamnus (Fabricius, 1793) (TL: Not stated. [Panama])</p><p>(fig.4: a-f; fig. 13)</p><p>Catasticta sisamnus is known from the Cordillera de Mérida, where Catasticta sisamnus merida Eitschberger &amp; Racheli, 1998 (TL: Venezuela, env. de Mérida) was described. The species is widespread, usually uncommon and encountered as single specimens, although on some occasions it has been reported in numbers. The elevational range is 900 to 2000 meters.</p><p>Even though Catasticta sisamnus is not yet reported from El Tamá, we think it likely that the species occurs there, since several specimens have been collected near S. Cristóbal, between 900 and 1500 meters.</p><p>Specimens recently collected by Steve Fratello on Mt. Ayanganna (Guyana) represent a new subspecies, being both differentiable and geographically distinct. This population does not fall within the political boundaries of Venezuela, but its occurrence on the tepuis of eastern Venezuela is highly probable.</p></div>	https://treatment.plazi.org/id/891387A9FFC4FFCF13A6FE0446BF7AC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFC4FFCF13A6FC1C43D57E03.text	891387A9FFC4FFCF13A6FC1C43D57E03.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta sisamnus subsp. ayanganna	<div><p>Catasticta sisamnus ayanganna ssp. nov.</p><p>Holotype: male, Guyana: Region 7, Mt. Ayanganna, Koatse R., 2500’–3300’, 05°26.0’N 60°00.4’W, 2–25 Apr 1999, leg. S. Fratello, R. Hanner, W. Prince, R. Williams/ Bar code, USNM ENT 0 0 232165, in USNM, will be deposited in CSBD (fig. 4: d-e)</p><p>Description: FW length: 28 mm.</p><p>Dorsal surface: black ground color with white yellowish markings. FW discal cell with a small spot; white yellowish discal band composed of three spots separated from each other by black veins, that in cell M3-CuA1 being small and oval. Thick and elongated submarginal spots in cells R4/R5-M1, M1-M2, M2-M3, M3-CuA1 and CuA1-CuA2. White yellowish discal band of HWs extended to the end of discal cell; no submarginal spots, but two faintly scaled in cells M1-M2 and M2-M3.</p><p>Ventral surface: black ground color with deep yellowish markings. FW as on dorsal surface, but with a series of premarginal elongated interneural spots. HW as on dorsal surface, but with complete series of submarginal interneural elongated spots and premarginal dots.</p><p>Diagnostic characters: a distinctive subspecies, with reduced clear markings. Its pattern approaches ssp. bythina Röber, 1924 from the Colombian Cordillera Oriental.</p><p>Paratypes (8 males, 1 female): 5 males, same data as HT, but Bar Code 0 0 232158, 0 0 232159, 0 0 232160, 0 0 232161 and 00232162; 1 female, same data as HT, but Bar Code 0 0 232167, in USNM; 3 males, same data as HT, but Bar Code 0 0 232163, 0 0 232164 and 0 0 232166, in USNM, will be deposited in CSBD.</p><p>Distribution: Known only from the type locality, where it is localized but not scarce. It should be looked for on other mountains of the Guyana Shield, especially near the border with Guyana.</p><p>Etymology: this subspecies is named after its type locality, Mt. Ayanganna, the highest mountain situated entirely within Guyana.</p></div>	https://treatment.plazi.org/id/891387A9FFC4FFCF13A6FC1C43D57E03	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFC3FFC813A6FF66436C7892.text	891387A9FFC3FFC813A6FF66436C7892.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta prioneris (Hopffer 1874) Hopffer 1874	<div><p>3 Catasticta prioneris (Hopffer, 1874) (TL: Peru Chanchamayo)</p><p>(fig. 4: g-l, fig. 14)</p><p>Catasticta prioneris araguana Eitschberger &amp; Racheli, 1998 (TL: Venezuela, Aragua, Maracay) is seldom encountered, usually as single individuals, in the Cordillera de la Costa at 1400–2200 meters. Recently collected specimens from El Tamá and the south-eastern Cordillera de Mérida differ in wing pattern and distribution from nearby ssp. araguana, ssp. estancia Eitschberger &amp; Racheli, 1998 (TL: Ecuador, Tungurahua, Río Estancias) and ssp. nigrilineata Bollino, Rodríguez &amp; Constantino, 2004 (TL: Colombia, Santander, San José de Suaita).</p></div>	https://treatment.plazi.org/id/891387A9FFC3FFC813A6FF66436C7892	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFC3FFC913A6FDBB40877B14.text	891387A9FFC3FFC913A6FDBB40877B14.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta prioneris subsp. parrishi	<div><p>Catasticta prioneris parrishi ssp. nov.</p><p>Holotype: male, Matamula, Tamá, 1900 m., Estado Táchira, Venezuela, 8-VII-2002, col. Costa, in MIZA (fig. 4: j-k)</p><p>Description: FW length: 26 mm.</p><p>Dorsal surface: FWs with black marginal band as wide as in araguana, and white spots in cells M3-CuA1, CuA1-CuA2 and CuA2-2A merged with submarginal white spots; white discal band occupying outer half of the cell; dark basal area reduced to the base of the wing; HWs with black marginal band thick, nearly absent in anal cells, four white marginal interneural triangular spots</p><p>Ventral surface: FWs with white discal band very reduced, composed of separated from each other and marginally dark scaled spots in cells M3-CuA1, CuA1-CuA2 and CuA2-2A; HWs with white discal band highly reduced, nearly only represented by yellow interneural bands.</p><p>Allotype: female, vía Chorro del Indio, 1500 m., Estado Táchira, IV-1994, col. F. Rey, in MIZA (fig. 4: l)</p><p>Description: FW length: 24 mm.</p><p>Dorsal surface: similar to araguana, but with larger submarginal white spots on FWs; HWs with black marginal band larger than in male; ventral surface like in male, but white discal band of HWs with more white.</p><p>Ventral surface: as in male.</p><p>Diagnostic characters: the new subspecies has white marking extended on dorsal surface, thus approaching ssp. estancia and ssp. araguana, but with a peculiar dark ventral surface highly reminiscent of ssp. hegemon Godman &amp; Salvin, 1889 from Costa Rica, Panama and the northern part of Central Colombian Cordillera.</p><p>Paratypes (30 males, 10 females): 1 male, Delicias, Venezuela, Táchira, 7-X-1982, col. G. Borges; 1 male, Venezuela, Táchira, Parque Nacional El Tamá, 27-VII-1980, col. R. Manrique; 1 female, same data, but 23- VII-80; 1 female, Venezuela, Táchira, Carret. Bramón Delicias, 1860 m., 9-11-XII-1980, J. Clavijo A. Chacón Ayala, all in MIZA; 1 male, Venezuela, Táchira, El Reposo (P.N. El Tamá), ~ 7°32’N 72°24’W, m. 2100 – 6.VII.2002, lg. Mauro Costa; 1 male, Venezuela, Táchira, Mata Mula (P.N. El Tamá), ~ 7°38’N 72°26’W, m. 1900-2000 – 7.VII.2002, lg. Mauro Costa; 1 male, 1 female, Venezuela – Táchira, El Chimborazo, ~ 7°43’N 72°12’W, m. 1300 - XII.1988, lg. local collectors, all in MBLI; 1 male, Matamula, Tamá, 1900 m., Estado Táchira, Venezuela, col. Costa, 8-VII-2002; 1 male, 1 female, same data, but 3-I-2002; 1 male, vía Chorro del Indio, 1500 m., Estado Táchira, Venezuela, I-1993, col. Costa; 1 male, vía Delicias, 1900 m., Estado Táchira, Venezuela, 19-I-1993, col. Costa; 1 male, S. Isidro, 1500 m, Estado Barinas, Venezuela, 02-I-2006, col. Costa; 1 male, S. Isidro, 1500 m, Estado Barinas, Venezuela, 04-I-2007, col. Costa, all in MCCV; 2 males, vía Delicias, 1950 m., III-83, C.f.R., Estado Táchira; 1 male, vía Delicias, 1700 m., V-91, C.f.R., Estado Táchira; 2 males, same data, but IV-90; 2 males, same data, but III-90; 1 male, same data, but V-87; 1 female, sama data, but VI-89; 3 males, 1 female, vía Delicias, 1800 m., IV-87, C.f.R., Estado Táchira; 1 male, same data, but VI- 87; 1 male, 1 female, vía Delicias, 1900 m., XII-84, C.f.R., Estado Táchira; 2 males, same data, but V-1986; 1 male, Barinitas, 1400 m., VIII-92, C.f.R., Estado Barinas; 1 female, same data, but II-89, all in FRMV; 1 male, Palo Grande, 1400 m., Municipio Lobatera, Edo. Táchira, VZLA.; 1 female, IX-1990, Hacienda Pánaga, 800 msnm., Mpio San Cristóbal, Táchira, VZLA, all in FRCV; 1 male, Chorro del Indio, 1300 m., Táchira, Venezuela, 10-XII-88, in DSBV; 1 female, El Paramito (Escuque), 1600m (Trujillo), 6-11 Octubre 2001, in PBPF; 1 male, estado Trujillo, arriba de Escuque, El Paramito (la cumbre), 1600 m, 15. IV.2006, T. Pyrcz leg., in MZUJ.</p><p>Distribution: C. prioneris parrishi ssp. nov. is known from the Eastern side of El Tamá and the Cordillera de Mérida north to Trujillo. It is generally scarce, and additional field work will be necessary to establish its northern limit.</p><p>Etymology: a noun in the genitive case after Chris Parrish (Sewanee, USA), for his contribution to a better knowledge of the ornithological fauna of Venezuela.</p></div>	https://treatment.plazi.org/id/891387A9FFC3FFC913A6FDBB40877B14	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFC2FFC913A6FBBB43047F82.text	891387A9FFC2FFC913A6FBBB43047F82.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta hebra (Lucas 1852) Lucas 1852	<div><p>5 Catasticta hebra (Lucas, 1852) (TL: Colombia)</p><p>(fig. 5: d-l; fig. 16)</p><p>Catasticta hebra is a common and widespread species. Three subspecies are present in Venezuela: Catasticta hebra potamea (C. &amp; R. Felder, 1861) (TL: Venezuela, [Aragua, Colonia] Tovar) (= angulifera Avinoff, 1926, TL.: Venezuela, Las Quiguas), Catasticta hebra crowleyi Butler, 1901 (TL: Venezuela, (Mérida), Monte Sierra and Culata), and Catasticta hebra apollinari Fassl, 1915 (TL: Colombia, [Cundinamarca], Río Negro).</p><p>Catasticta hebra potamea would occur from the Sierra Nevada de Santa Marta (Colombia), eastward to Miranda and Guárico, along the Venezuelan Cordillera de La Costa and Cordillera del Interior. It flies between 1200 and 2000 meters. Very few specimens from the Sierra Nevada de Santa Marta (Colombia) are known to us and specimens from Sierra de Perijá (Zulia) were not available, thus we can not assert all populations belong to the same subspecies. Specimens from Sierra de Aroa (NE of Barquisimeto) are transitional in pattern between ssp. potamea and crowleyi.</p><p>In the Cordillera de Mérida, Catasticta hebra crowleyi is also common and widespread, flying from 1200 to 2200 meters. Táchira populations east of San Cristóbal Valley are transitional in pattern between ssp. crowleyi and apollinari from El Tamá.</p><p>Catasticta hebra apollinari is restricted to El Tamá in Venezuela, being mostly a species of the Colombian Cordillera Oriental.</p></div>	https://treatment.plazi.org/id/891387A9FFC2FFC913A6FBBB43047F82	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFC2FFC913A6FD4343E07A82.text	891387A9FFC2FFC913A6FD4343E07A82.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta notha (Doubleday 1847) Doubleday 1847	<div><p>4 Catasticta notha (Doubleday, 1847) (TL: Venezuela)</p><p>(= notha (Lucas, 1852), TL: Venezuela) (fig. 5: a-c; fig. 15)</p><p>Catasticta notha was described by Doubleday from Venezuela without additional data. The only specimens claimed to be from Venezuela are the types of notha Doubleday and notha Lucas (the latter a junior primary homonym and junior synonym). However, it occurs in Santander and Cundinamarca (Cordillera Oriental, Colombia) and may also occur in the El Tamá area. For this reason, we provisionally retain it as a member of the Venezuelan fauna.</p></div>	https://treatment.plazi.org/id/891387A9FFC2FFC913A6FD4343E07A82	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFC2FFCA13A6F8BB40867885.text	891387A9FFC2FFCA13A6F8BB40867885.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta philais	<div><p>6 Catasticta philais (C. &amp; R. Felder, 1865) (TL: Colombia, Bogotá)</p><p>(fig. 6: a-b; fig. 17)</p><p>The nominotypical subspecies is widespread along the northern part of the Colombian Cordillera Oriental, but is never common. Catasticta philais borgesi Eitschberger &amp; Racheli, 1998 (TL: Venezuela, Táchira) is the Venezuelan representative of the species, and is restricted to El Tamá, where it flies in sympatry with C. philoscia . These species can hardly be distinguished in the field, but the following differences, mostly ventral wing pattern differences (fig. 1), are consistent.</p><p>A) FW yellow interneural spot in cell R4/R5-M1 large in philoscia, a small dot in philais . B) HW whitish area at base of cells M1-M2 and CuA1-CuA2) large in philoscia, small in philais . C) HW yellow interneural spot in cell CuA1-CuA2 (and usually also other yellow interneural spots of</p><p>VHW) long in philoscia, a small dot in philais .</p><p>No females of ssp. philais or borgesi are known.</p></div>	https://treatment.plazi.org/id/891387A9FFC2FFCA13A6F8BB40867885	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFC1FFCA13A6FDA941167F7B.text	891387A9FFC1FFCA13A6FDA941167F7B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta philoscia	<div><p>7 Catasticta philoscia (C. &amp; R. Felder, 1861) (TL: Venezuela, [Aragua, Colonia] Tovar)</p><p>(= Catasticta subtoca Röber, 1924, TL: Venezuela, Mérida) (fig. 6: c-h; fig. 18)</p><p>Catasticta philoscia was described by C. &amp; R. Felder (1861) from male and female specimens collected by Moritz in Colonia Tovar. Subsequently, Röber (1924) described Catasticta subtoca from a single female from Mérida, apparently not realizing that his specimen matched the description of philoscia . Lamas (1993) synonymized subtoca Röber with philoscia philoscia .</p><p>Colonia Tovar is a problematical type locality for Catasticta philoscia . Although Colonia Tovar is a wellknown butterfly collecting locality in Venezuela, no specimens of C. philoscia other than the type series are known from there. We do not know if the type specimens were mislabeled or if there was local extinction of this species. Moritz collected in Colonia Tovar and Mérida (Viloria et al., 2001), and no specimens of this usually common species are known from the Cordillera de La Costa and the North-Eastern part of Cordillera de Mérida. On the other hand, populations of Catasticta seitzi have become noticeably scarcer in the Colonia Tovar area in recent decades. We provisionally retain Colonia Tovar as the type locality of C. philoscia . The current distribution of Catasticta philoscia in Venezuela is the Central and South-western portions of the Cordillera de Mérida from 1900 to 2400 meters.</p><p>Females of Catasticta philoscia are extremely rare. A few females from El Tamá have dorsal brownish bands and spots on the FW and HW in contrast to nominotypical females (and one female from Cundinamarca, Colombia), which have the corresponding bands and spots white. Even though various Catasticta species have recently been found to have dimorphic females (Bollino, Greeney &amp; Vitale, 2002; Bollino, Boyer &amp; Vitale, 2003; Bollino &amp; Boyer, in prep., and cf. Catasticta gelba, C. troezene, C. seitzi here below), no species related to philoscia is dimorphic. Since the dark females of C. philoscia philoscia appear to form a geographical entity, we describe these populations as a new subspecies.</p></div>	https://treatment.plazi.org/id/891387A9FFC1FFCA13A6FDA941167F7B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFC1FFCB13A6F9CA40407FB2.text	891387A9FFC1FFCB13A6F9CA40407FB2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta philoscia subsp. reyi	<div><p>Catasticta philoscia reyi ssp. nov.</p><p>Holotype: female, Fundo Piedras Blancas—San Vicente de la Revancha, 2350 m, Estado Táchira, Venezuela, 22-XII-1997, Col. F. Rey, in MIZA. (fig. 6: f-g)</p><p>Description: FW length: 22 mm.</p><p>Dorsal surface: ground color chocolate brown with complete series of lighter markings. FW with discal band, submarginal and premarginal spots whitish and suffused of dark scales. HW with discal band, submarginal and premarginal spots light brown. Submarginal spots are arrow-shaped, those in cells M3-CuA1 and CuA1-CuA2 with an evident indentation at outer edge.</p><p>Ventral surface: as in nominotypical subspecies, but discal band of FW with spots smaller.</p><p>Diagnostic characters: males and females distinguished by the shape of DHW submarginal spots in cells M3-CuA1 and CuA1-CuA2: in C. philoscia reyi such spots are nearly always arrow-shaped with an evident indentation at outer edge, while in nominotypical specimens these spots are usually triangular, without any indentation at outer edge.</p><p>Paratypes (119 males, 17 females): 2 males, Venezuela, Táchira, Las Copas, 2-11-XI-1999, J. DeMarmels, A. Chacón; 1 male, Venezuela, Táchira, La Pesa, San Vicente de la Revancha, 2350 m, 13-19-I-1999, J. DeMarmels, A. Chacón; 1 male, Delicias, Venezuela, Táchira, 1900 m, 13-VI-1987, Col. G. Borges; 5 males, vía Delicias, Venezuela, Táchira, III-1983, Col. G. Vivas; 2 males, Venezuela, Táchira, Parque Nacional El Tamá, 23-VII-80, Col. G. Manrique; 1 male, same data, but 27-VII-80; 1 male, Delicias, Venezuela, Táchira, 13-VI-1982, Col. G. Borges; 1 male, vía Delicias, La Peña, Venezuela, Táchira, 13-VI-1982, Cols. Borges Romero, all in MIZA; 3 males, Venezuela, Táchira, El Reposo (P.N. El Tamá), ~ 7°32’N 72°24’W, m. 2100 – 4-5.I. 2001, lg. Mauro Costa; 1 male, Venezuela, Táchira, Betania, mt. 2250, 9-8-[19]87, lg. F. Rey; 1 male, Venezuela, Táchira, Junín, Fundo Piedras Blancas, ~ 7°32’N 72°23’W, m. 2300-2400, I.2004 (local collectors); 2 males, 1 female, idem, but m. 2500; 2 males, idem, but m. 2700; 2 males, idem, but m. 2800; 2 males, idem, but m. 2900; 3 males, Venezuela, Táchira, Junin, Taburete, Páramo La Revancha, ~ 7° 32’N 72° 23’W, m. 2800, I-II.2004, (local collectors); 17 males, 3 females, idem, but III.2004; 1 female, Venezuela, Táchira, Fundo Piedras Blancas, ~ 7°32’N 72°23’W, m. 2300–2500, I.2004 (local collectors); 3 females, Venezuela, Edo. Táchira, Fundo Piedras Blancas, Páramo La Revancha, ~ 7° 30’N 72° 21’W, m. 2350, VIII.1983, (local collectors), all in MBLI; 11 males, La Revancha, 2800 m, Estado Táchira, Venezuela, II-2004; 3 females, La Revancha, 2300 m, Estado Táchira, Venezuela, II-2004; 2 females, La Revancha, 2700 m, XII-1997; 3 males, El Reposo, 2100 m, Estado Táchira, Venezuela, 4-I-2001, Col. Costa, all in MCCV; 1 male, 1 female, 12-IV- 1995, F.P.B., 2.300 msnm, Táchira, VZLA; 1 male, II-2004, La Revancha, Tamá, 2.800 msnm, Táchira, VZLA; 2 males, same data, but III-2004; 1 female, III-2004, La Revancha, Tamá, Táchira, VZLA, all in FRSV; 3 males, Vía Delicia, 1900 m, XI-82, C.f.R.; 1 male, same data, but III-83; 2 males, same data, but XI- 83; 5 males, same data, but VI-84; 12 males, same data, but XII-84; 7 males, same data, but V-86; 14 males, same data, but V-86; 3 males, same data, but VI-86; 3 males, same data, but I-87, all in FRMV; 1 male, El Reposo, 2100 m, Táchira, Venezuela, 26-XII-1998, Col. F. Rey; 1 male, same data, but 7-XII-2002; 1 female, same data, but 02-I-2002, all in DSBV; 1 male, La Revancha, 2800 m, Estado Táchira, Venezuela, II-2004, Col. F. Rey; 1 male, Betania, 2250 m, Estado Táchira, Venezuela, 10-VIII-1987 Col. F. Rey; 1 female, La Revancha, 2300 m, Estado Táchira, Venezuela, II-2004, Col. F. Rey; 1 male, El Reposo, E of Delicias, PN El Tamá, SE Táchira State, Western Venezuela, 07°33'N, 072°25'W, 2080m., Coll. Neild; 2 males, Mata Mula area, Bramón–Delicias, PN El Tamá, SE Táchira State, W. Venezuela, Coll. Neild, 1900 m, V-86, Leg. C.F. Romero; 2 males, Mata Mula area, Bramón–Delicias, PN El Tamá, SE Táchira State, W. Venezuela. Coll. Neild, 1900 m, II-84, Leg: C.F. Romero; 2 males, Mata Mula area, Bramón–Delicias, PN El Tamá, SE Táchira State, W. Venezuela. Coll. Neild 1900 m, VI-84, Leg: C.F. Romero, all in ANLE.</p><p>Distribution: Restricted to the eastern side of El Tamá (Táchira), at elevations between 2100 and 2900 meters. No specimens from the Colombian side of this range are known.</p><p>Etymology: a noun in the genitive case, in honour of Fernando Rey (San Cristóbal, Táchira, Venezuela), friend and colleague, for his contribution to the knowledge of Venezuelan Catasticta, skillful observations and precious notes about their ethology.</p></div>	https://treatment.plazi.org/id/891387A9FFC1FFCB13A6F9CA40407FB2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFC0FFC413A6F8D147067BEC.text	891387A9FFC0FFC413A6F8D147067BEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta troezene	<div><p>8 Catasticta troezene (C. &amp; R. Felder, 1865) (TL: Colombia, Bogotá)</p><p>(fig. 7: a-d; fig. 19)</p><p>Catasticta troezene was described from male and females, but only two females were found in the type series at the BMNH. Lamas (1993) selected one of these as a lectotype, but erroneously wrote that it was a male (Abadjiev 2006). Felder (1865), in the same paper, described Euterpe zancle from two or more females, but the specimen labelled as " type " appears to be a male. Both taxa were described from " Nova Granada: Bogotá (Lindig)". Examination of males and females recently collected along the eastern side of the Colombian Cordillera Oriental confirms that Euterpe zancle C. &amp; R. Felder 1865 is a synonym of Euterpe troezene C. &amp; R. Felder 1865, as previously stated by various authors (Reissinger, 1972; Eitschberger &amp; Racheli, 1998; Lamas, 2004). Two additional taxa, i.e. latiplaga Röber 1924 (TL: Colombia, E-Cordilliere, Pacho) and lactea Avinoff 1926 (TL: Colombia, Choachi), are also synonyms of Catasticta troezene troezene C. &amp; R. Felder 1865. The lectotype of Catasticta troezene troezenides Röber, 1908 (TL: Colombia) selected by Lamas (1993) corresponds to Colombian populations of Catasticta troezene distributed along Magdalena Valley on both the western side of the Cordillera Oriental and the eastern side of the Cordillera Central (Bollino &amp; Rodríguez, 2004). It does not occur in Venezuela.</p><p>Catasticta gelba Brown &amp; Gabriel, 1939 has been considered to be the “Venezuelan subspecies” of Catasticta troezene (Eitschberger &amp; Racheli, 1998; Lamas, 2004). Extensive material recently collected in Táchira reveals that C. gelba is sympatric and synchronic with a new subspecies of C. troezene, which we describe.</p></div>	https://treatment.plazi.org/id/891387A9FFC0FFC413A6F8D147067BEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFCFFFC413A6FD1446AC7F96.text	891387A9FFCFFFC413A6FD1446AC7F96.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta troezene subsp. miza	<div><p>Catasticta troezene miza ssp. nov.</p><p>Holotype: male, vía Delicias 1700 m, Estado Táchira, Venezuela 2-I-2002 Col. Costa, in MIZA (fig. 7: a-b)</p><p>Description: FW length: 25 mm.</p><p>Dorsal surface: ground color brownish black. FW with discal band and submarginal spots of whitish color, with strong suffusion of dark scales. HW with discal band of the same color, the anal portion being the lighter in color. Five small triangular submarginal spots strongly suffused by dark scales.</p><p>Ventral surface: as in nominotypical subspecies.</p><p>Diagnostic characters: the new subspecies is characterized in both sexes by having DFW discal bands and submarginal spots with a strong suffusion of dark scales, and DHW submarginal spots small and with black suffusion. DHW submarginal spot in cell CuA1-CuA2 is always separated by the discal band. In C. troezene troezene males have corresponding bands and spots brownish yellow in color, with DHW submarginal spot in cell CuA1-CuA2 always merged with discal band; females with light yellow markings and DHW submarginal spots very large.</p><p>Paratypes (8 males, 2 females): 2 males, Venezuela – Táchira, Junín, Fundo Piedras Blancas, ~ 7°32’N 72°23’W, m. 2300–2400 - I.2004, lg. local collectors, in MBLI; 1 male, vía Delicias 1700 m Estado Táchira Venezuela 15-X-1990 Col. Costa; 2 males, La Revancha 2300 m Estado Táchira Venezuela 20-III-2004 Col. F. Rey; 1 female, same data but 10-XII-1997; all in MCCV; 1 male, vía Delicias 1800 m (Matamula) IV-87 C.F. Romero, Estado Táchira, in FRMV; 1 male, 10-IV-1996 Fundo Piedra Blanca (F.P.B.) San Vicente de la Revancha 2.300 msnm., Táchira, Vzla.; 1 male, Matamula, vía Delicias, Municipio Junín, 1.900 msnm. Táchira, Vzla; 1 female, 26-IV-96 Matamula, vía Delicias, Municipio Junín, 1.900 m, Táchira, Venezuela, all in FRSV.</p><p>Distribution: The new subspecies occurs only on the Venezuelan side of El Tamá, where it is rare, although it may also occur on nearby Colombian slopes of the same range.</p><p>Etymology: a noun in the nominative singular standing in apposition after MIZA (Museo del Instituto de Zoología Agrícola, Maracay, Venezuela) for the great cooperation we received from its staff.</p></div>	https://treatment.plazi.org/id/891387A9FFCFFFC413A6FD1446AC7F96	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFCFFFC513A6F8B447E97B14.text	891387A9FFCFFFC513A6F8B447E97B14.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta gelba Brown & Gabriel 1939	<div><p>9 Catasticta gelba Brown &amp; Gabriel, 1939 stat. rev. (TL.: Venezuela, Mérida)</p><p>(fig. 7: e-h; fig. 20)</p><p>Catasticta gelba Brown &amp; Gabriel, 1939 has been treated as a subspecies of troezene (Eitschberger &amp; Racheli 1998, Lamas 2004). As previously mentioned, we found it sympatric and synchronic with Catasticta troezene miza in Táchira, for which reason we recognize Catasticta gelba Brown &amp; Gabriel, 1939 as a distinct species, stat. rev. The best characters differentiating the two species are on VHW, with the dark discal band of gelba being significantly wider, “entering” the outer portion of the cell, while in troezene it is thin, never “entering” the cell (fig.2).</p><p>In females, the DHW marginal black band of gelba is large and touching the cell, with arrow-shaped white marks, while in troezene miza the band is thinner, not touching the cell, and with chevron-shaped marks suffused with black scales.</p><p>Females of both species are dimorphic: all examined gelba females are white, except one which is yellow, while of two known troezene miza females, one is yellow and the other yellowish-white.</p><p>Catasticta gelba is generally common in Mérida and the eastern side of El Tamá. No specimens are yet known from nearby Colombia. Populations on opposite sides of the Táchira depression show minor variation.</p></div>	https://treatment.plazi.org/id/891387A9FFCFFFC513A6F8B447E97B14	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFCEFFC513A6FD3641587D3A.text	891387A9FFCEFFC513A6FD3641587D3A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta seitzi Lathy & Rosenberg 1912	<div><p>10 Catasticta seitzi Lathy &amp; Rosenberg, 1912 (TL: W-Colombia, Rio Cauca, Guabinas)</p><p>(fig. 8: a-d; fig. 21)</p><p>Catasticta seitzi had not previously been reported from Venezuela, so we were perplexed at finding Venezuelan specimens of C. seitzi collected on the Cordillera de La Costa (Aragua and Distrito Federal, now Distrito Capital) and represented in most public and private Venezuelan collections. The peculiar wing pattern of these specimens is reminiscent of nominotypical C. seitzi from the Colombian Cordillera Oriental, but differs consistently in some characters on the ventral surfaces of both wings. Although differences in ventral wing patterns often differentiate species of Catasticta, we opt for the more conservative strategy of describing the Venezuelan population of C. seitzi as a new subspecies.</p></div>	https://treatment.plazi.org/id/891387A9FFCEFFC513A6FD3641587D3A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFCEFFC613A6FB8B477E7AFC.text	891387A9FFCEFFC613A6FB8B477E7AFC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta seitzi subsp. clarita	<div><p>Catasticta seitzi clarita ssp. nov.</p><p>Holotype: male, Cumbre el Avila, DF, Venezuela, 2100 m., 4.III. [19]61, Rodolfo Feige col., in MIZA (fig. 8: a-b)</p><p>Description: FW length: 23 mm.</p><p>Dorsal surface: dark brown with brownish yellow markings. FW with one small spot at apex of the cell, and a complete series of discal, submarginal and premarginal spots. HW with discal band and submarginal spots of the same color as in FW.</p><p>Ventral surface: similar to nominotypical subspecies, but with FW premarginal spot in cell R3-R4/R5 inwardly incompletely bordered by black line ( C. seitzi seitzi with spot totally bordered by black line); FW submarginal spots outwardly indented ( C. seitzi seitzi with round spots); HW yellow portion of submarginal interneural stripes as long as the brown portion of same stripes (in nominotypical specimens such stripes are mostly brown, with a basal yellow small dot).</p><p>Allotype: female: kilómetro 23, El Junquito, Dist. Federal, 1900 m., 14.IV.1983, [lg.] Alcides Huerta, in MIZA (fig. 8: c)</p><p>Description: FW length: 22 mm.</p><p>Dorsal surface: blackish brown with white markings. FW with one small spot at apex of the cell, and complete series of discal submarginal spots faintly marked. HW with discal band and premarginal spots of the same color as in FW; submarginal spots nearly obsolete.</p><p>Ventral surface: as in male.</p><p>Paratypes (12 males, 9 females): 1 male, Caracas, Avila, 6.5.1959 (ex col. Feige); 1 male, Caracas, Avila 1.4.1961 (ex col. Feige); 1 male, Caracas, Avila, 2100 m., 8.IV. [19]61; 1 female, El Junquito, D.F., Venezuela, 2000 m., 5.VII.1950, [lg.] F. Fernández Yépez; all in MIZA; 1 female, Venezuela – Aragua, Colonia Tovar, ~ 10°25’N 67°17’W, m. 2000 – 30.I.2005, Lg. Mauro Costa, in MBLI; 1 male, 1 female, Agua Fría - El Jarillo, 1800 m, Estado Miranda Venezuela 30-VI-1985 Col. Costa; 1 male, same data but 24-VI-1985; 1 female, Colonia Tovar 2150 m Estado Aragua Venezuela 7-III-2004 Col. Costa; 1 female, Colonia Tovar 2000 m Estado Aragua Venezuela 7-XI-2004; all in MCCV; 1 male, kilómetro 23, El Junquito, D. F., 1830 m., 30.V.1985, (16:30 hr) [lg.] J.B. Rodríguez; 1 male, via El Junquito, Colonia Tovar, Dist. Federal, 11.VI.1977, (9:00 hr) [lg.] J.B. Rodríguez; 1 female, vía El Capachal, Colonia Tovar, Estado Aragua, 2.I.1981, (14:00 hr) [lg.] J.B. Rodríguez, all in coll. BR; 1 male, Colonia Tovar, I-1987 C.f.R. 1550 m., Estado Aragua.; 1 female, Colonia Tovar, 2100 m. C.f.R. Estado Aragua. IV-1982; 1 female, Colonia Tovar, 1550 m. C.f.R. Estado Aragua. I-1981; 1 female, Colonia Tovar, 1700 m. C.f.R. Estado Aragua. VI-1987, all in FRMV; 1 male, Colonia Tovar, 2070 m., Edo. Aragua, Venezuela, 07-VII-2002, Col. María G. Alvarez, in DSBV; 1 male, Venezuela, Caracas, Avila, 25.IV. [19]61, leg. Feige, in ZSMC; 2 males, Colonia Tovar, sector los Colonos, 1900/ 2000m (Aragua), 30/3/2006, in PBPF.</p><p>Distribution: As far as we know, the species has been collected recently only in the vicinity of Colonia Tovar (Aragua), while in the ’60s and ’70s it was observed also on Pico El Avila at 1800–2100 m. Over the last few decades, the species has become scarcer for unknown reasons. Curiously, females are encountered as commonly as males, which is unusual in Catasticta .</p><p>Etymology: a noun in apposition in the nominative case after Mrs. Clara (Clarita) Baldi, wife and mentor of the junior author, for her enthusiastic contribution to our knowledge of Venezuelan butterflies.</p></div>	https://treatment.plazi.org/id/891387A9FFCEFFC613A6FB8B477E7AFC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFCDFFC613A6FC2E46F87C4A.text	891387A9FFCDFFC613A6FC2E46F87C4A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta philone	<div><p>11 Catasticta philone (C. &amp; R. Felder, 1865) (TL: Colombia, Bogotá; Ecuador)</p><p>(fig. 8: e-f; fig. 22)</p><p>The species is widespread and common on both sides of the Cordillera Oriental in Colombia (Bollino &amp; Rodriguez, 2004), but it has not been previously reported from Venezuela. Two males of this species were recently collected at Fundo Piedras Blancas and Páramo La Revancha (Parque Nacional El Tamá, Táchira) between 2300 and 2800 meters.</p><p>Both specimens belong to the nominotypical species. Some characters suggest that they represent a different subspecies, but additional material would be needed to confirm this possibility.</p></div>	https://treatment.plazi.org/id/891387A9FFCDFFC613A6FC2E46F87C4A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFCDFFC613A6FA6346CF7E74.text	891387A9FFCDFFC613A6FA6346CF7E74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta revancha Rey & Pyrcz 1996	<div><p>12 Catasticta revancha Rey &amp; Pyrcz, 1996 (TL: Fundo Piedra Blanca, San Vicente de la Revancha, Parque Nacional El Tamá, Táchira, Venezuela, 2350 m.)</p><p>(fig. 8: g-i; fig. 23)</p><p>This species is apparently endemic to the Venezuelan side of El Tamá, but sampling from the Colombian side of the same massif is lacking. Information about its ecology, ethology, and resemblance to Leptophobia eleone (Doubleday, 1847) were provided by Rey &amp; Pyrcz (1996). Those authors considered it to be a close relative of Catasticta poujadei Dognin from southern Ecuador and northern and central Peru because of similarities in its wing pattern. However, phylogeny of Andean Pieridae is yet poorly studied, and the striking similarity convergence in pattern between C. revancha and C. poujadei may be fortuitous.</p></div>	https://treatment.plazi.org/id/891387A9FFCDFFC613A6FA6346CF7E74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFCDFFC713A6F89643627F04.text	891387A9FFCDFFC713A6F89643627F04.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta chrysolopha (Kollar 1850) Kollar 1850	<div><p>13 Catasticta chrysolopha (Kollar, 1850) (TL: [Colombia, Cundinamarca], “ Venezuela ”)</p><p>(fig. 9: a-i; fig. 24)</p><p>Catasticta chrysolopha was described by Kollar from Venezuela based on material collected by Prince Maximilian Sulkowsky in 1843 in both Colombia and Venezuela. Subsequently, Reissinger (1972) described Catasticta spectrum (TL: Venezuela, Mérida, Rio Albarragas), which is currently considered to be a subspecies of C. chrysolopha from the Cordillera de Mérida (Lamas, 2004). Lamas (1995) emphasized that C. chrysolopha chrysolopha should be restricted to Colombia because “localities of many butterflies collected by Sulkowsky are ambiguous or erroneous.” The origin of the type specimen was also discussed by Wojtusiak (1998) who arrived at the same conclusion as Lamas (1995), that the type specimen of Catasticta chrysolopha was collected somewhere in Colombia. Neild (pers. comm.) agrees with both Lamas and Wojtusiak about the Colombian origin of the type material because “almost every taxon described before about 1950 (probably even up to 1975) from the Venezuelan Andes was collected in the Mérida range (I have seen none which could have been from the Serrania de El Tamá), so it is almost inconceivable that a name from the 19th century was based on an El Tamá specimen, even if it does fly there!”. Careful study of the type specimen by both us and Wojtusiak confirms that it matches Colombian specimens, so that the type locality of the species can be restricted to Cundinamarca Department (Colombia), and its distribution consequently encompasses only the Central and Oriental Colombian Cordilleras northward to the Venezuelan side of Serranía de El Tamá.</p><p>Authors treating Catasticta spectrum offered a variety of opinions on its taxonomic status. Reissinger (1972) described it as a species, but (unpublished manuscript) subsequently suggested that it be considered a subspecies of Catasticta chrysolopha . Wojtusiak (1998) also considered it to be a valid species due to its distinct dorsal pattern and ecology, and Eitschberger &amp; Racheli (1998) synonymized it with Catasticta chrysolopha chrysolopha . Finally, Lamas (2004) proposed a new combination, considering it to be a distinct subspecies of chrysolopha, but without providing any additional details.</p><p>We assign specimens from El Tamá to the nominotypical subspecies, but at the same time recognize that long series of specimens from both El Tamá and the Cordillera de Mérida demonstrate stable morphological differences. Thus, we reject the treatment of C. chrysolopha chrysolopha and C. spectrum as synonyms, and accept the distinctiveness of the populations from the Cordillera de Mérida. Furthermore, we have encountered specimens from Yacambú, northeast of the Cordillera de Mérida, which clearly represent an additional taxon in this group. Interspecific differences in the genus Catasticta are generally sustained by clear and discrete differences on the VHW, but in these cases neither Wojtusiak nor we have found them. Genitalic differences between both taxa were reported by our Polish colleague, but we find no evidence in the literature of any species-specific differences in the various taxa of the genus Catasticta, apart from a single vague reference (Eitschberger &amp; Racheli, 1998: 12). Moreover, dissections of eight specimens (four from El Tamá and four from Mérida) did not indicate any differences, except for some individual variability in the shape of the harpe and saccus. Recognizing three different taxa in this group of Venezuelan Catasticta broaches the question of the nature of their relationships, and the problem of reflecting those relationships in an appropriate taxonomy. Accepting Wojtusiak’s thesis that the populations in the Cordillera de Merida represent a distinct species of butterfly would seem to force an analogous treatment for the populations in Yacambú. The degree of morphological differentiation present in this complex, and the clear biogeographic relationships, are expressed by considering all three distinct Venezuelan populations as comprising subspecies of the same specific entity.</p></div>	https://treatment.plazi.org/id/891387A9FFCDFFC713A6F89643627F04	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFCCFFC713A6F88946F27EAA.text	891387A9FFCCFFC713A6F88946F27EAA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta chrysolopha subsp. adamsi (Eitschberger & Racheli 1998) Eitschberger & Racheli 1998	<div><p>Catasticta chrysolopha adamsi (Eitschberger &amp; Racheli, 1998) (TL: North Colombia, Sierra Nevada de Santa</p><p>Marta)</p><p>= Catasticta abdita Wojtusiak, 1998 (TL: North Colombia, Sierra Nevada de Santa Marta)</p></div>	https://treatment.plazi.org/id/891387A9FFCCFFC713A6F88946F27EAA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFCCFFC713A6F94141687E77.text	891387A9FFCCFFC713A6F94141687E77.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta chrysolopha subsp. chrysolopha (Kollar 1850) Kollar 1850	<div><p>Catasticta chrysolopha chrysolopha (Kollar, 1850) (TL: [Colombia, Cundinamarca], “ Venezuela ”)</p><p>= Archonias xeque Mengel, 1899 (TL: Colombia, [Huila])</p><p>= Archonias chrysolophana Mengel, 1899, misspelling</p></div>	https://treatment.plazi.org/id/891387A9FFCCFFC713A6F94141687E77	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFCBFFC013A6FB42417F7CC6.text	891387A9FFCBFFC013A6FB42417F7CC6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta apaturina Butler 1901	<div><p>14 Catasticta apaturina Butler, 1901 (TL: Ecuador, [Cotopaxi] Angamarca)</p><p>(fig. 10: a-c; fig. 25)</p><p>Prior to the present study, Catasticta apaturina was not reported from Venezuela. However, we obtained a few specimens from El Tamá at 2300–2800 m elevation, where they are quite scarce. These specimens belong to Catasticta apaturina subturina Reissinger, 1972 (TL: Colombia, Muzo) from the Central and Eastern Colombian Cordilleras (Bollino &amp; Rodríguez, 2004).</p></div>	https://treatment.plazi.org/id/891387A9FFCBFFC013A6FB42417F7CC6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFCBFFC013A6FF66400A7D2B.text	891387A9FFCBFFC013A6FF66400A7D2B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta chrysolopha subsp. beatrizae	<div><p>Catasticta chrysolopha beatrizae ssp. nov.</p><p>Holotype: male, Vía Sanare – Caspo, 1900 m., Edo. Lara, Venezuela, 15-II-2004, Col. Juan C. De Sousa C., presently in DSBV, will be deposited in MIZA (fig. 9: g-h)</p><p>Description: FW length: 22 mm.</p><p>Dorsal surface: black ground color. FW discal band with four spots: the anal one whitish with dark suffusion, those in cells CuA1-CuA2 and CuA2-2A whitish with strong orange suffusion, that in cell M3-CuA1 faintly marked. Submarginal spots nearly obsolete. HW with discal band as in other subspecies, but costal white spots dashed by dark scales. Submarginal spots nearly obsolete.</p><p>Ventral surface: FW with greyish discal band; premarginal series uncomplete, composed by four small spots in cells R3-R4/R5, R4/R5-M1, M1-M2 and M2-M3; submarginal spots small and subtriangular, those in cells R4/R5-M1, M1-M2 and M2-M3 mostly yellow, the remnants greyish. HW the same as in other subspecies, but postdiscal black band heavily marked and large; submarginal portion of cells greyish.</p><p>Diagnostic characters: the new subspecies is characterized by its large size and melanistic appearance of both wing surfaces. Orange suffusion of FW discal spots in cells CuA1-CuA2 and CuA2-2A was not observed in long series of other subspecies.</p><p>Paratypes (6 males, 1 female): 2 males, same data as holotype; 1 male, same data as holotype, but 23-I- 2000, all in DSBV; 1 male, same data as holotype, in MCCV; 1 male, same data as holotype, in FRSV; 1 male, same data as holotype, in MBLI; 1 female, Vía Sanare–Caspo, 1900 m., Edo. Lara, Venezuela, 26-XII-1999, Col. Andrés M. Orellana, in AOMV</p><p>Distribution: To the best of our knowledge, the new subspecies is restricted to the vicinity of its type locality.</p><p>Etymology: a noun in the genitive case after Mrs. Beatriz Coelho De Sousa, mother of Juan Carlos De Sousa who collected most of type series.</p></div>	https://treatment.plazi.org/id/891387A9FFCBFFC013A6FF66400A7D2B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFCBFFC113A6F8B14627782C.text	891387A9FFCBFFC113A6F8B14627782C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta duida Brown 1932	<div><p>16 Catasticta duida Brown, 1932 (TL: Venezuela, Mt. Duida)</p><p>(fig. 10: g-h; fig. 27)</p><p>Catasticta duida was described from Mt. [Cerro] Duida (Amazonas, ~ 3°21’N, 65°40’W) on the basis of 7 males and 2 females, plus 1 male from Mt. Roraima (Brazil). Recently we examined 6 males in MIZA that were collected on Cerro Yutajé (Amazonas: 5°45’N, 66°6’W), Kukenán Tepui (Bolívar: 5°19’N, 60°49’W) and Kamarkabarac Tepui (Bolívar: 5°59’N, 62°01’W). These data suggest that C. duida likely occurs on most Venezuelan tepuis higher than 1700–1800 m. In spite of the isolation of these populations from each other, there is no evidence of subspecific differentiation. Catasticta duida was filmed by a Japanese group on top of Cerro de La Neblina (Amazonas: ~ 0°48’N, 66°01’W) (A. Orellana, pers. comm.).</p></div>	https://treatment.plazi.org/id/891387A9FFCBFFC113A6F8B14627782C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFCBFFC013A6F9E143577E6E.text	891387A9FFCBFFC013A6F9E143577E6E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta tomyris	<div><p>15 Catasticta tomyris (C. &amp; R. Felder, 1865) (TL: Nova Granada, Bogotá)</p><p>(fig. 10: d-f; fig. 26)</p><p>Catasticta tomyris subtamina (Eitschberger &amp; Racheli, 1998) (TL: Venezuela, Mérida, Tal des Rio Albarragas) is apparently the only Venezuelan representative of this species. It is distributed along the Cordillera de Mérida from 1800 to 2500 m, southwestward to the Serranía de El Tamá. Specimens are not commonly encountered.</p></div>	https://treatment.plazi.org/id/891387A9FFCBFFC013A6F9E143577E6E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFCAFFC113A6FD2443F27AA4.text	891387A9FFCAFFC113A6FD2443F27AA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta cinerea Butler 1897	<div><p>18 Catasticta cinerea Butler, 1897 (TL: [S. Peru] “ Ecuador ”)</p><p>(fig. 11: figs. a-c; fig. 29)</p><p>Catasticta cinerea is widespread, but not rare, along the Central and Oriental Colombian Cordilleras, and belongs to C. cinerea . suprema Fassl, 1915 (TL: Colombia, Central-Cordilliere, Quindiu Pass). Specimens recently collected in El Tamá (Táchira), at elevations ranging from 2500 to 2900 m have wing patterns without much variability.</p></div>	https://treatment.plazi.org/id/891387A9FFCAFFC113A6FD2443F27AA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFCAFFC113A6FE5440AB7BFC.text	891387A9FFCAFFC113A6FE5440AB7BFC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta semiramis (Lucas 1852) Lucas 1852	<div><p>17 Catasticta semiramis (Lucas, 1852) TL: Colombia, [Cundinamarca])</p><p>(fig. 10: i-k; fig. 28)</p><p>Catasticta semiramis (Lucas, 1852) was described from Venezuela as ssp. costarum Bollino &amp; Lamas 2004. The species occurs from 2700 to 3200 m, on both the Colombian and Venezuelan sides of El Tamá (Táchira). Individuals are highly localized, though not rare.</p></div>	https://treatment.plazi.org/id/891387A9FFCAFFC113A6FE5440AB7BFC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFCAFFC313A6FBDC40057BC4.text	891387A9FFCAFFC313A6FBDC40057BC4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta rochereaui Le Cerf 1924	<div><p>19 Catasticta rochereaui Le Cerf, 1924 stat rev. (TL: Colombia, Pamplona)</p><p>(fig. 11: d-f; fig. 30)</p><p>In collections we found one female (collected at El Azul, vía El Reposo, El Tamá, Táchira, at an elevation of 2200 m.) that we preliminarily identified as Catasticta cinerea rochereaui Le Cerf, and one male (collected at Mata Mula, El Tamá, Táchira, at an elevation of 2000 m.) that appeared to be Catasticta cinerea hollandi Avinoff. Field work yielded another male and two females, which were practically identical to the previously known material. These specimens were found in the same area, at elevations ranging from 2300 to 2600 m, and are partially sympatric with Catasticta cinerea suprema . The similarity of the ventral wing pattern of these specimens, which is different from the wing pattern of the sympatric C. cinerea, led us to the conclusion that those males and females belong to the same taxon, which is distinct from C. cinerea .</p><p>Catasticta rochereaui was described from a single female, and the male has been unknown. Similarly, Catasticta hollandi was known only from males. We recognize Catasticta rochereaui Le Cerf stat. rev. as a distinct species and transfer the taxon hollandi Avinoff, 1926 (TL: Colombia, Santander, Peña Blanca) to the Catasticta rochereaui species group.</p><p>Since Venezuelan males of C. rochereaui differ from the holotype of hollandi, we tried to ascertain its type locality before deciding its subspecific status. Avinoff (1926) provided only the data present on the label: Peña Blanca, Santander, Colombia, Feb. 1917. The type specimen of C. hollandi was collected by Melbourne Armstrong Carriker during his Colombian Santander-Casanare ornithological expedition (1916–1917). Carriker was an American ornithologist who lived for a long time in Colombia selling bird skins to the Carnegie Museum (where the holotype of C. hollandi is deposited), and his biography (Carriker, 2001) was written by his eldest son. We asked the ornithological staff of the Carnegie Museum if they might be able to supply any further information on the possible location of the collecting site Peña Blanca. Robin K. Panza (Section of Birds, Carnegie Museum of Natural History, Pittsburgh, USA) kindly responded with the following notes: In the "Ornithological Gazetteer of Colombia " (2nd. edition, 1997, by Raymond A. Paynter, Jr.), this locality (Peña Blanca) is given as:</p><p>ca. 06:33/72:30 (de Schauensee, 1948a:319)</p><p>2,800 m. on western slope of Eastern Andes, a valley SW of Chiscas [06:33/72:29], northern Boyacá, near border of Santander (de Schauensee, 1948a:319); not shown on our maps; Carriker, 24–28 Feb., 1–3, 5 –6 Mar. 1917 (WFVZ, as Peña Blanca, Santander; Todd, 1919:113, as Peña Blanca, Santander; CM, as Peña Blanca, Santander); see El Cardón for discussion of probable error in coordinates for Peña Blanca.</p><p>The entry for El Cardón, Santander is:</p><p>3,300 m pass above Peña Blanca [ca. 06:33/72:30 (de Schauensee 1948a:319)], Carriker, 5 Mar 1917 (de Schauensee, 1948a:291; CM, as "El Cardón, Boyacá); there are two places with the same name in the general area, vsz. at ca. 06:08/72:32, on the western slope in Boyacá (Atlas, 1977) and the one on the eastern slope in Casanare, on the Rio Chinivaque [ca. 06:09/72:20 (Atlas, 1977)], 8 km NNW of La Salina [06:06/72:20 (USBGN)] (Atlas, 1977); however, both Todd (1942, Ann. Carnegie Mus., 29:366) and de Schauensee closely associate Peña Blanca with El Cardón and the coordinates given by de Schauensee for Peña Blanca indicate it is quite some distance from either of these sites; the locality may actually be Alto Cardón [coordinates?], SE of Carcasí [06:42/72:35 (USBGN)], Santander (Dicc Geog.)</p><p>Todd was the curator here when we received specimens from Carriker, and he consulted with Carriker about his collecting localities.</p><p>Here are the collecting localities we have from Carriker for February and March of 1917:</p><p>FEB 07–17 LA PICA</p><p>FEB 24 - MAR 0 3 PEÑA BLANCA</p><p>MAR 0 5 EL CARDÓN</p><p>MAR 05–06 PEÑA BLANCA</p><p>MAR 13–20 LAGUNILLAS</p><p>MAR 24 EL COCUY (7 KM S, 10 KM E; PEÑA NEGRA) MAR 26–30 RIO NEGRO</p><p>MAR 31 CHINIVAQUE</p><p>MAR 31 RIO NEGRO</p><p>Carriker was very careful about the accuracy of his data. Therefore, Peña Blanca and El Cardón must be near each other, probably on or very close to the Santander/Boyacá border, as he collected in both locations on 5 March.</p><p>Peña Blanca is located on the southwestern side of Sierra Nevada del Cocuy. Due to the different geographical distributions of C. rochereaui and C. hollandi, which is correlated with differences in the width of the white discal band of DHW in males, we retain the latter as a distinct subspecies. Furthermore, we consider Catasticta cinerea laurentina (Eitschberger &amp; Racheli, 1998) as conspecific with rochereaui (character evidence below), and summarize our conclusions as follows:</p><p>Catasticta rochereaui rochereaui Le Cerf, 1924 stat. rev. (TL: Colombia, Pamplona [Norde de Santander])</p><p>Distribution: both Colombian and Venezuelan sides of Serranía de El Tamá.</p><p>Catasticta rochereaui hollandi Avinoff, 1926 comb. nov. (TL: Peña Blanca, “Santander” [Boyacá], Colombia)</p><p>Distribution: Sierra Nevada del Cocuy (Boyacá, Colombia)</p><p>Catasticta rochereaui laurentina (Eitschberger &amp; Racheli, 1998) comb. nov. (TL: Colombia, Sierra Nevada de Santa Marta, San Lorenzo)</p><p>Distribution: As far as we know, this subspecies is restricted to the Sierra Nevada de Santa Marta (Colombia), but its occurrence in the Sierra de Perijá cannot be discarded a priori. Wojtusiak &amp; Rey (1999) point out that “most cloud forest and Páramo species found in the Tamá have their relatives in the eastern Cordillera [of Colombia], particularly the Cocuy area, and in the Perijá range”.</p><p>Males of Catasticta rochereaui can be differentiated from males of C. cinerea by the white discal bands on both DFW and DHW. Females of C. rochereaui, apart from being much larger, can be easily differentiated from females of C. cinerea by the large yellow anal area on DHW, while in C. cinerea the same area is dark brown with, at most, a few yellow scales.</p></div>	https://treatment.plazi.org/id/891387A9FFCAFFC313A6FBDC40057BC4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFC8FFC313A6FB0647517FBA.text	891387A9FFC8FFC313A6FB0647517FBA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta tricolor Butler 1897	<div><p>21 Catasticta tricolor Butler, 1897 (TL: Ecuador)</p><p>(fig. 11: j-l; fig. 32)</p><p>Catasticta tricolor tomasi Wojtusiak &amp; Rey 1999 (= tricolor viridia Wojtusiak &amp; Rey 1999 nomen nudum) was described from Páramo El Tamá, Táchira. The type specimens were collected at the same time and place as specimens of Catasticta uricoecheae inopa and remain the only known ones from the Venezuelan side of the type locality despite subsequent field work. A male belonging to the same subspecies was collected on the Colombian side of El Tamá, and it is presently deposited in JLBC (Bollino &amp; Rodríguez, 2004). Another male of this species was collected by Viloria at El Peñasco, Cerro Tetari, Sierra de Perijá (Zulia), and deposited in UZMC (plate 9: fig. l). Its wing patterns seems to be intermediate between it and ssp. rodriguezi (Eitschberger &amp; Racheli) (TL: Colombia, Antioquia, Páramo de Belmira). We provisionally consider it as belonging to the Venezuelan subspecies, pending additional material from this poorly sampled area.</p></div>	https://treatment.plazi.org/id/891387A9FFC8FFC313A6FB0647517FBA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFC8FFC313A6FCE641307DEA.text	891387A9FFC8FFC313A6FCE641307DEA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta uricoecheae	<div><p>20 Catasticta uricoecheae (C. &amp; R. Felder, 1861) (TL: Colombia, Muzo)</p><p>(fig. 11: g-i; fig. 31)</p><p>Catasticta uricoecheae was reported for the first time from Venezuela by Wojtusiak &amp; Rey (1999), when they described the subspecies inopa from Páramo El Tamá, Táchira. This subspecies is confined to a narrow altitudinal range, 2800 to 3100 m., but it is not rare. It is sympatric and synchronic with Catasticta tricolor, but the latter is much rarer. The same uricoecheae subspecies is present also in the Colombian side of El Tamá (Bollino &amp; Rodríguez, 2004). We illustrate the female of this subspecies for the first time (plate 7: fig. i).</p><p>Catasticta uricoecheae can be separated from C. tricolor by its DFW whitish-grey discal markings. The corresponding markings in C. tricolor are yellowish green.</p></div>	https://treatment.plazi.org/id/891387A9FFC8FFC313A6FCE641307DEA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFC8FFDC13A6F87E46F2787C.text	891387A9FFC8FFDC13A6F87E46F2787C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta radiata (Kollar 1850) Kollar 1850	<div><p>Catasticta radiata (Kollar, 1850) (TL: [Colombia, Cundinamarca] “ Venezuela ”)</p><p>(= Catasticta hanna Reissinger, 1970, TL: Colombia, Muzo)</p><p>The species was described by Kollar based on Sulkowsky’s material collected in Venezuela. We have already mentioned the sometimes vague or erroneous localities of this collector (cf. C. chrysolopha). Lamas (1995) noted that the species is not present in Venezuela, and we have found no records for it.</p></div>	https://treatment.plazi.org/id/891387A9FFC8FFDC13A6F87E46F2787C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
891387A9FFD7FFDC13A6FEAE41CC7A7A.text	891387A9FFD7FFDC13A6FEAE41CC7A7A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catasticta truncata Lathy & Rosenberg 1912	<div><p>Catasticta truncata Lathy &amp; Rosenberg, 1912 (TL: [Ecuador, Loja] “ Venezuela, Mérida”)</p><p>(= xanthotaenia Röber, 1924, TL: Ecuador)</p><p>This species was described from at least one male labeled “Mérida”, and Abadjiev (2005) designated a lectotype. Röber (1924) described Catasticta xanthotaenia from Ecuador, and Lamas (1993) considered it to be a subspecies of truncata . Subsequent authors considered both taxa as subspecies of truncata (Reissinger, 1972; Lamas, 2004), or considered xanthotaenia to be a synonym of truncata (Eitschberger &amp; Racheli, 1998) . No one noted that the type locality of truncata might be wrong. Catasticta truncata is a highly localized and specialized species, restricted to dry areas of southern Ecuador and northwestern Peru (Lamas, pers. comm.). As no Venezuelan specimens of the species are known except for the questionably labeled lectotype, we change the type locality of the species to southern Ecuador (Loja Province).</p></div>	https://treatment.plazi.org/id/891387A9FFD7FFDC13A6FEAE41CC7A7A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bollino, Maurizio;Costa, Mauro	Bollino, Maurizio, Costa, Mauro (2007): An illustrated annotated check-list of the species of Catasticta (s. l.) Butler (Lepidoptera: Pieridae) of Venezuela. Zootaxa 1469: 1-42, DOI: 10.5281/zenodo.176651
