taxonID	type	description	language	source
896187BFF76FB615FE07C578FC31FF32.taxon	description	(Fig. 1)	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF76FB615FE07C578FC31FF32.taxon	materials_examined	Type material examined (ZIN). Holotype: m, ‘ Ugam Ranges / Baldarbek River / h = 1800 m / 14. vi. 1964 / leg. N. Skopin’ [in Cyrillics]. ALLOTYPE: 1 f, ‘ Karzhantau Range / Badam River / h = 1600 m / 18. v. 1938 / leg. N. Skopin’ [in Cyrillics]. Distribution. Kazakhstan: Ugam and Karzhantau ranges (SKOPIN 1977).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF76CB614FE92C11EFD7FFDCA.taxon	description	(Fig. 2) Allard, 1882: 135; Seidlitz, 1893: 294; Skopin, 1960: 50 (larva); Skopin, 1961: 189. Material examined (ZIN). 1 m, ‘ Southeast Kazakhstan / around Issyk Lake / vii. 1958 / leg. N. G. Skopin’ [in Cyrillics]; 1 m, ‘ Kazakhstan / southeast side of Ketmen Range / 20 km north of Sarynaz / h = 2000 / arid recess near crag / 3. vi. 1986. / leg. L. Egorov’ [in Cyrillics]; 1 f, ‘ Kazakhstan / Alma-Ata / Big Almaty Lake / 21. vii. 1958 / leg. N. G. Skopin’ [in Cyrillics]. Distribution. Kazakhstan, China (Gansu, Ningxia, Qinghai, Shaanxi, Xinjiang, Xizang provinces), Kyrgyzstan, Tajikistan, Afghanistan (LÖBL et al. 2008). Regional distribution. Southeast Kazakhstan and adjacent territories of Kyrgyzstan (Central Tian Shan) (SKOPIN 1961). Blaps caraboides intermittens Kaszab, 1962 (Fig. 3) Kaszab, 1962: 313; Skopin, 1961: 189. Type material examined (ZIN). Paratypes: 6 mm, ‘ Southeast Kazakhstan / Dzungarian Alatau / Malyy Baskan River / 10. vi. 1953 / leg. N. Balabas’ [in Cyrillics]. Material examined (ZIN). 2 mm, ‘ Southeast Kazakhstan / Dzungarian Alatau / Malyy Baskan River / 25 km east of Pokatilovka village / 18. vi. 1968 / leg. G. Bugaev’ [in Cyrillics] [45 ° 23 ’ 22.90 ” N, 80 ° 8 ’ 25.18 ” E]. Comments. The subspecies was formally described by Kaszab (1962), but a year before, SKOPIN (1961) already indicated, without giving a name, that Kaszab considered the populations of the Dzhungarian Alatau as a separate subspecies. Distribution. Kazakhstan: to the northeast of Ili River, spruce groves in Dzungarian Alatau (SKOPIN 1961, KASZAB 1962).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF76DB613FE27C270FE4CFCF1.taxon	description	(Figs 4, 21 A, B) Ménétriés, 1832: 199; Fischer von Waldheim, 1832: 192 (“ Blaps muricata ”); Seidlitz, 1893: 288; Skopin, 1968: 86; Medvedev & Nepesova, 1985: 119. Type material examined (ZIN). Lectotype of Blaps deplanata (designated by ABDURA- KHMANOV & NABOZHENKO 2011): m, ‘ Baku. // deplanata Menet. Baku // Lectotypus / Blaps deplanata Mén. 1832 / des. Abdurakhmanov & Nabozhenko’. Paralectotype: 1 f, ‘ Baku. // Paralectotypus / Blaps deplanata Mén. 1832 / des. Abdurakhmanov & Nabozhenko’. Lectotype of Blaps muricata (designated by ABDURAKHMANOV & NABOZHENKO 2011): m, ‘ Baku. // muricata Fisch. Baku // Lectotypus / Blaps deplanata Mén. 1832 / des. Abdurakhmanov & Nabozhenko’. Material examined (ZIN). 1 f, ‘ Kazakhstan / Barsa-Kelmes Island [former island of Aral Sea] / 13. xii. 1978 / leg. A. Konev’; 1 m, ‘ Uzbekistan / Between Bukhara and Kata Kurgan / v. [18] 84. / leg. Regel’; 3 mm, 2 ff, ‘ Uzbekistan / Juma – Samarkand / 12. vii. [18] 98 / leg. Retter’; 1 m, ‘ Uzbekistan / Around Samarkand / 1. iii. [18] 96 / leg. L. Barshevsky’ [in Cyrillics]; 11 mm, ‘ Uzbekistan / Around Samarkand / 3. iii. [18] 96 / leg. L. Barshevsky’ [in Cyrillics]; 1 m, 1 f, ‘ Uzbekistan / Around Samarkand / 13 - 15. iv. [18] 96 / leg. L. Barshevsky’ [in Cyrillics]; 1 m, ‘ Uzbekistan / Samarkand / 1898 / leg. Retter’; 1 f, ‘ Uzbekistan / Samarkand district / Samarkand / 7. v. 1904 / leg. G. Suvorov’ [in Cyrillics]; 1 m, ‘ Uzbekistan / Samarkand / 1892 / leg. О. Herz’; 1 m, ‘ Uzbekistan / Samarkand / 1892 / leg. Herz // coll. Sivers’. Distribution. Azerbaijan (Apsheron Peninsula), Iran, Kazakhstan, Turkmenistan, Uzbekistan (ABDURAKHMANOV & NABOZHENKO 2011, MEDVEDEV & NEPESOVA 1985). Regional distribution. The Barsakelmes Nature Reserve (a former island in the Aral sea), the outskirts of Northwestern Tian Shan (the Boroldaytau mountains) (SKOPIN 1968).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF768B611FE59C0ADFC4DFD5C.taxon	description	(Figs 5, 27 D) Seidlitz, 1893: 285; Skopin, 1960: 55 (larva); Skopin, 1961: 191; Skopin, 1968: 86. Material examined (ZIN). 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / Kurdayskaya station / h = 900 m / 07. viii. 1907 / leg. A. Jakobson’ [in Cyrillics]; 2 mm, 8 ff, ‘ Southeast Kazakhstan / Alma-Ata Region / Kurdayskiy pass / h = 1100 m / 16. v. 1907 / leg. A. Jakobson’ [in Cyrillics] [43 ° 16 ’ 23.25 ” N, 74 ° 49 ’ 37.78 ” E]; 1 m, 5 ff, ‘ Southeast Kazakhstan / Alma-Ata Region / Vernyi (Alma-Ata) suburbs / h = 900 m / 23 – 29. v. 1907 / leg. A. Jakobson’ [in Cyrillics] [43 ° 12 ’ 12.60 ” N, 76 ° 50 ’ 37.26 ” E]; 1 m, ‘ Southeast Kazakhstan / Alma-Ata Region / Kara-Bulak village / 24. vii. 1909 / leg. I. E. Boikov’ [in Cyrillics]. Distribution. Kazakhstan (LÖBL et al. 2008), “ Turkestan ” (SKOPIN 1961). Regional distribution. The plain near Trans-Ili Alatau Range (SKOPIN 1960), northern slopes and foothills of Trans-Ili Alatau Range (SKOPIN 1961), Zhetizhol Range, southeastern part of Karatau Ridge (SKOPIN 1968).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF768B610FE21C2E1FE28FEDB.taxon	description	(Figs 6, 27 E) Seidlitz, 1893: 305; Skopin, 1968: 86. Material examined (ZIN). 1 m, ‘ Kazakhstan / Around Kazalinsk / Dorandsh / 20. x. [18] 75 // 79221 ’ [in Cyrillics]; 1 m: ‘ Uzbekistan / Bukhara / Amu Darya / Kelif / 10. vi. 1904 / leg. Suvorov’ [in Cyrillics]; 1 f, ‘ Uzbekistan / Bishkent / Bukhara / 8. v. [18] 97 / leg. Kazankov’ [in Cyrillics]. Distribution. Kazakhstan, Uzbekistan, Turkmenistan, Tajikistan, Afghanistan (LÖBL et al. 2008). Regional distribution. Northern and northeastern parts of the Kyzylkum desert (SKOPIN 1968).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF769B61BFDE5C166FBB9FCCE.taxon	description	(Figs 7 – 11) Type material (ZIN). Holotype: m, ‘ South Kazakhstan / Akmechet-Aulie Cave / 100 km NW Bourny / 22. vi. 1964 / leg. V. Kusov // coll. N. Skopin’. Paratypes: 1 f, ‘ South Kazakhstan / Karatau Ridge / 30 km W Sholakkorgan / h = 915 m / 43 ° 46 ’ 27.11 ” N, 68 ° 48 ’ 50.20 ” E / 7. vi. 2015 / leg. A. Shapovalov’; 1 m: ‘ South Kazakhstan / Karatau Ridge / 15 km S Kozmoldak village / 43 ° 51 ’ 36.0 ’’ N, 068 ° 32 ’ 52.5 ’’ E / h = 931 m / 16. iv. 2014 / leg. A. V. Ivanov’. Description. Male. Body black, mat, slender. Anterior margin of epistoma weakly arcuately emarginate, straight in middle. Lateral margins of epistoma weakly rounded. Lateral margins of genae straight in anterior half, rounded at base. Lateral margins of head with distinct emargination between epistoma and genae. Head widest at level of posterior margin of eyes. Head 1.45 times as wide as interocular distance. Antennomeres 10 – 11 reaching base of elytra. Ratio of length / width of antennomeres 2 – 11 as 7 (9), 43 (10), 19 (10), 17 (10), 17 (11), 18 (14), 10 (9), 10 (9), 10 (9), 12 (10). Mentum transversely oval, convex in middle, its base straight. Punctation of head fine, sparse (puncture diameters on frons 5 times as wide as distance between punctures, puncture diameters on genae 2 times as wide as distance between punctures). Pronotum weakly transverse (1.15 times as wide as long), widest at middle, 1.72 times as wide as head. Ratio of pronotal width near anterior angles to widest part and width at base 5.3: 7.7: 7.2. Anterior margin of pronotum widely emarginate, lateral margins widely rounded, base straight. Lateral margins of pronotum weakly emarginate near base. Disc of pronotum weakly convex, lateral sides narrow flattened. Anterior angles obtuse, widely rounded, posterior angles right, narrowly rounded. Pronotum completely beaded, except for middle of apex and base. Pronotal punctation moderately coarse, dense (distance between punctures in middle subequal to one puncture diameter), punctation of lateral sides and near base sparser (puncture diameters 2 times as wide as distance between punctures). Prothoracic hypomera with small wrinkles and covered with small granules. Lateral margins of hypomera not excavate. Elytra weakly convex, elongate (1.93 times as long as wide together), flattened along suture, widest at middle, 3.42 times as long and 2.52 times as wide as pronotum, 2.52 times as wide as head. Caudal extension of elytra (mucro) distinct, 2.2 mm long; elytra 10 times as long as mucro. Elytral surface with fine microwrinkles and rasp-like punctation, obliterated from base to middle, and almost completely disappearing closer to elytral apex. Epipleura with fine wrinkles and sparse fine rasp-like punctures. Hair tuft between abdominal ventrites 1 and 2 present. Abdominal ventrite 1 without tubercle. Abdominal ventrites 1 – 3 with sparse rasp-like punctation and sparse granules, covered with short recumbent hairs. Abdominal ventrites 4 – 5 with moderately dense punctation, punctation at lateral sides of ventrite 4 forming wrinkles, ventrite 5 completely beaded, except for base and with unclear bead apically. Anterior margin of male inner sternite VIII emarginate; accessory gland of sternite VIII moderately long and thin. Rods of spiculum gastrale not merged at apex, forming long common stem. Aedeagus length 4.5 mm, width 0.9 mm. Aedeagus weakly C-curved. Lateral margins of parameres weakly rounded, widely emarginate in apical third, lateral sides of parameres with longitudinal impression in basal two-thirds, apex rounded. Parameres length 1.3 mm, width 0.9 mm. Legs long, slender. Ratio of lengths of femora, tibiae and tarsi of fore, middle and hind legs 8.4: 7.3: 5.3, 9.4: 7.9: 6.1, 11.6: 10.5: 7.8. All tarsomeres with one pair of setal brushes. apical piece, dorsal view; E = the same, ventral view; F = the same, lateral view. Scale bars: 100 µm for A – C, 30 µm for D, 200 µm for E – F Body length 29 – 30 mm, width 11 – 11.1 mm. Female. Shape of body and punctation similar to those of male. Head 1.41 times as wide as interocular distance. Antennomere 11 reaching base of pronotum. Ratio of pronotal width near anterior angles, in widest part and at base 4.9: 7.0: 6.9. Elytra elongate (1.76 times as long as wide together), 3.65 times as long and 1.55 times as wide as pronotum, 2.4 times as wide as head. Mucro short (1 mm). Ovipositor moderately long. Apical lobes straight in basal two-thirds, weakly arcuately emarginate at apical third. Apex of lobes acute. Ventral side of lobes with excavation near middle. Middle of ventral side of lobes with small deep longitudinal wrinkles. Ante- ca, r = reservoirs, g = accessory gland of spermatheca). Scale bars = 1 mm rior margin of proctiger widely emarginate. Basal duct of spermatheca between vagina and reservoirs long, gland of spermatheca short. Reservoirs of spermatheca separated from each other; 1 st reservoir sharply expanding from middle, wide in apical half; 2 nd reservoir almost spherical at apex, 1 st reservoir slightly larger than 2 nd. Base of spermathecal gland very thin. Stem of spiculum ventrale moderately long and wide. Body length 27.3 mm, width 11.1 mm. Etymology. The species epithet of this new species is dedicated to the late Czech entomologist, Ren é Fouque. Differential diagnosis. The body shape of the new species is similar to that in Blaps seriata Fischer von Waldheim, 1820 (Fig. 12), and differs from the latter in the following character states: antennomeres 10 – 11 of Blaps fouquei sp. n. reaching first quarter of elytra, those of B. seriata reaching only basal quarter of the pronotum; elytra of Blaps fouquei sp. n. flattened along the suture, elytra of B. seriata weakly convex; lateral sides of parameres of Blaps fouquei sp. n. weakly rounded in the basal two-thirds, apical third widely emarginate, lateral sides of parameres of B. seriata weakly rounded; reservoirs of the spermatheca in Blaps fouquei sp. n. are separated from each other, reservoirs of the spermatheca and valve of the accessory gland in B. seriata are closely located.	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF762B61AFE8AC4C5FC59FE23.taxon	description	(Fig. 13)	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF762B61AFE8AC4C5FC59FE23.taxon	materials_examined	Type material examined (ZIN). Holotype: m, ‘ Southeast Kazakhstan / Dzungarian Alatau / Altyn-Emel Range / 29. viii. 1962 / leg. N. G. Skopin’ [in Cyrillics]. Allotype: 1 f, ‘ Southeast Kazakhstan / upper terrace of Ili River / 24. viii. 1962 / leg. N. G. Skopin’ [in Cyrillics]. Paratypes: 1 m, 1 f, ‘ Southeast Kazakhstan / Dzungarian Alatau / Altyn-Emel Range / 30. viii. 1962 / leg. N. G. Skopin’ [in Cyrillics]; 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / Sary-Ozek suburbs / 04. vii. 1962 / leg. N. G. Skopin’ [in Cyrillics] [44 ° 21 ’ 7.37 ” N, 77 ° 59 ’ 5.12 ” E]; 1 f, ‘ Southeast Kazakhstan / Sarkand / 10. iv. 1965. / leg. N. Yerolskaya’ [in Cyrillics]. Material examined (ZIN). 2 ff, ‘ Southeast Kazakhstan / northern slope of Altyn- Emel Range / Kugaly suburbs / 16. ix. 1966 / leg. N. G. Skopin’ [in Cyrillics] [44 ° 25 ’ 56.56 ” N, 78 ° 42 ’ 26.28 ” E]; 1 f, ‘ Semirechye Region / env. of Araltobe / 14. ix. 1966 / leg. N. Skopin’ [in Cyrillics]; 1 m, ‘ Southeast Kazakhstan / env. of Tekeli / 22. v. 1967 / leg. V. Linsky’ [in Cyrillics]. Distribution. Kazakhstan (Almaty Region) (LÖBL et al. 2008). Regional distribution. Altyn-Emel Range (SKOPIN 1966, 1968).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF763B607FE78C1EEFBBCFCB4.taxon	description	(Figs 14, 27 B) Gebler, 1825: 47; Seidlitz, 1893: 287; Skopin, 1961: 189: Skopin, 1966: 336; Skopin, 1968: 85; Arnoldi & Medvedev, 1969: 403; Ren et al., 2016: 119. Material examined (ZIN). 1 m, ‘ Southeast Kazakhstan / Alma-Ata Region / bottom land of Bolshaya Almatinka River / h = 1000 m / 02. vi. 1949 / leg. N. G. Skopin’ [in Cyrillics]; 1 m, ‘ Southeast Kazakhstan / Alma-Ata Region / Ketmen Range / Podgornoye suburbs / 05. vii. 1960 / leg. N. G. Skopin’ [in Cyrillics] [44 ° 16 ’ 1.02 ” N, 79 ° 28 ’ 18.53 ” E]; 1 m, 1 f, ‘ Southeast Kazakhstan / South-Western Balkhash Lake region / Chu-Iliyskiye gory / 28. vii. 1965 / leg. V. Kombulin’ [in Cyrillics]; 1 m, ‘ Southeast Kazakhstan / Moyunkum Sands / Kos- Kuduk station / 12. x. 1949 / leg. A. Gyarynin’ [in Cyrillics] [44 ° 4 ’ 56.14 ” N, 77 ° 25 ’ 14.85 ” E]; 2 mm, ‘ Southeast Kazakhstan / Alma-Ata Region / Kurtogay natural boundary / 21. vii. 1959 / leg. N. G. Skopin’ [in Cyrillics] [43 ° 15 ’ 52.20 ” N, 78 ° 58 ’ 25.09 ” E]; 1 m, ‘ Southeast Kazakhstan / Sands along Khorgos River / 17. iv. 1965 / leg. N. G. Skopin’ [in Cyrillics]; 1 m, ‘ Southeast Kazakhstan / Alma-Ata Region / Alma-Ata / Zailiyskiy Alatau Range / 16. v. 1964 / leg. G. Kosolapova’ [in Cyrillics] [43 ° 9 ’ 43.70 ” N, 76 ° 53 ’ 16.83 ” E]; 2 mm, 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / Tekeli suburbs / 22. v. 1967 / leg. V. Linskiy’ [in Cyrillics] [44 ° 51 ’ 47.06 ” N, 78 ° 45 ’ 45.56 ” E]; 2 mm, 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / Dzungarian Alatau / Koktuma village / 26. vi. 1962 / leg. G. S. Medvedev’ [in Cyrillics] [45 ° 51 ’ 17.07 ” N, 81 ° 38 ’ 20.27 ” E]; 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / Karatal River / 16. vi. 1964 / leg. N. Gorbunov’ [in Cyrillics]; 1 m, ‘ Southeast Kazakhstan / Alma- Ata Region / lower reaches of Lepsy River / 16. vi. 1965 / leg. N. G. Skopin’ [in Cyrillics]; 1 m, ‘ Southeast Kazakhstan / Alma-Ata Region / Alma-Ata / 03. vi. 1957 / leg. N. G. Skopin’ [in Cyrillics] [43 ° 9 ’ 43.70 ” N, 76 ° 53 ’ 16.83 ” E]; 2 mm, ‘ Southeast Kazakhstan / Alma-Ata Region / Kokishbay village / 16. ix. 1948 / leg. D. Aleksandrov’ [in Cyrillics] [45 ° 2 ’ 19.10 ” N, 75 ° 26 ’ 55.43 ” E]; 1 m, ‘ Southeast Kazakhstan / Alma-Ata Region / Akkol village / 13. vii. 1969 / leg. N. G. Skopin’ [in Cyrillics] [45 ° 0 ’ 42.54 ” N, 75 ° 39 ’ 31.32 ” E]; 1 m, ‘ Southeast Kazakhstan / Alma-Ata Region / Dzungarian Alatau / 30 km south-west of Sarkand city / 15. vi. 1968 / leg. T. Bugaev’ [in Cyrillics] [45 ° 18 ’ 41.75 ” N, 79 ° 33 ’ 25.99 ” E]; 1 m, ‘ Southeast Kazakhstan / Alma- Ata Region / Araltobe suburbs / 14.09.1966 / leg. N. G. Skopin’ [in Cyrillics] [45 ° 2 ’ 10.45 ” N, 75 ° 27 ’ 11.49 ” E]; 1 m, 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / 40 km south of Khorgos village / 14. iv. 1965 / leg. N. G. Skopin’ [in Cyrillics] [43 ° 52 ’ 13.06 ” N, 80 ° 14 ’ 24.21 ” E]; 1 m, ‘ Southeast Kazakhstan / Jambyl Region / 120 km west of Burubayltal village / 15. ix. 1964 / leg. N. G. Skopin’ [in Cyrillics] [45 ° 15 ’ 52.41 ” N, 72 ° 29 ’ 3.35 ” E]; 1 m, ‘ Southeast Kazakhstan / Alma-Ata Region / Bakanas village / 09. vi. 1953 / leg. A. Savicheva’ [in Cyrillics] [44 ° 48 ’ 29.94 ” N, 76 ° 16 ’ 19.24 ” E]; 1 m, ‘ Southeast Kazakhstan / Alma-Ata Region / Sartaga [env. of Narynkol village] / 24. iv. 1964 / leg. N. G. Skopin’ [in Cyrillics] [42 ° 43 ’ 12.29 ” N, 80 ° 9 ’ 57.62 ” E]; 7 mm, 3 ff, ‘ Eastern Kazakhstan / Saykan Range / Kendyrlik (Saryterek) village / 02. vii. 1910 / leg. A. Jakobson’ [in Cyrillics] [47 ° 29 ’ 52.50 ” N, 85 ° 10 ’ 36.35 ” E]; 1 m, ‘ Uzbekistan / Malyy Chimgan Mount / 23. vi. 1936 / leg. N. G. Skopin’ [in Cyrillics] [41 ° 32 ’ 45.11 ” N, 70 ° 2 ’ 8.62 ” E]; 1 m, ‘ Kyrgyzstan / Kyrgyz Ridge / Shamsi ravine / viii. 1958 / leg. Pivovarov’ [in Cyrillics] [42 ° 39 ’ 17 ” N, 75 ° 23 ’ 20 ” E]. Comments. Specimens labelled as “ Blaps granulata mixta ” and “ Blaps granulata psammophila ” are deposited in ZIN and NMP collections (prepared by Skopin for description, but remained undescribed). The specimens have no distinct differences from the nominotypical subspecies and interpreted here as Blaps granulata granulata. Additional material. “ Blaps granulata mixta ” (unpublished name) (Figs 15 A – C). Material examined (ZIN). 3 mm, 3 ff, ‘ Southeast Kazakhstan / Alma-Ata Region / Jabyrtau mountains / 26. vi. 1966 / leg. N. G. Skopin’ [in Cyrillics] [42 ° 56 ’ 49 ” N, 79 ° 59 ’ 15 ” E]; 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / Narynkol village / 30. v. 1961 / leg. V. Chekalin’ [in Cyrillics] [42 ° 43 ’ 12.29 ” N, 80 ° 9 ’ 57.62 ” E]; 1 m, ‘ Southeast Kazakhstan / Alma-Ata Region / Terskey Alatau Range / Koksay River / 08. vi. 1957 / leg. L. Marinenko’ [in Cyrillics]; 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / Ketmen Range / 20. iii. 1960 / leg. V. Zadorin’ [in Cyrillics] [43 ° 20 ’ 35.53 ” N, 80 ° 40 ’ 42.01 ” E]; 1 m, ‘ Kyrgyzstan / North from Issyk-Kul lake / Chong-Örüktü village / 2. v. 1956 / leg. N. Skopin’ [in Cyrillics].	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF763B607FE78C1EEFBBCFCB4.taxon	materials_examined	“ Blaps granulata psammophila ” (unpublished name) (Figs 16 A – C). Material examined. 1 m, ‘ Thian-S. Musart’ (NMP); 1 f (NMP), ‘ Chan-Tengri. Mont. merid. ’ (NMP); 1 f, ‘ Turkestan, Reitter leg. ’ (NMP); 1 m, ‘ Southeast Kazakhstan / Panfilov [Zharkent city] / 12.05.1960 / leg. N. Skopin’ [in Cyrillics] (ZIN); 1 m, ‘ caudata Gebl. Turcom. ’ (NMP); 1 f, ‘ Southeast Kazakhstan / Panfilov [Zharkent city] / 12.05.1960 / leg. N. Skopin’ [in Cyrillics] (ZIN). Distribution. Central and Southeast Kazakhstan and adjacent areas of Kyrgyzstan (SKOPIN 1960, 1961; ARNOLDI & MEDVEDEV 1969), China (Gansu and Xinjiang provinces) (LÖBL et al. 2008, Ren et al. 2016). Regional distribution. Eastern part of Kyrgyz Range, the mountainous part of Talas River area, eastern part of Trans-Ili Alatau Range (SKOPIN 1968).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF77FB605FE6BC0ADFD9AFCA1.taxon	description	(Fig. 17) Ménétriés, 1849: 236 (“ Turcomanie ”); Skopin, 1966: 337; Skopin, 1968: 85. Material examined. 1 f, ‘ Turkestan // Blaps caudata / Turkestan / ii. 20 ’ (NMP); 1 m, ‘ Syr Daria / Bang-Haas’ (NMP); 1 m, ‘ Western Balkhash Lake region / 120 km W Baital village / 15. ix. 1964 / leg. N. Skopin’ [in Cyrillics] (ZIN); 1 f, ‘ Kyrgyz Range / Shamsi pass / viii. 1958 / leg. Pivovarov’ [in Cyrillics] (ZIN). Distribution. Kazakhstan (LÖBL et al. 2008). Regional distribution. Muyunkum Desert, to the west and south of Balkhash Lake (Baital), the lower course of the Ili River (SKOPIN 1966, 1968). Blaps granulata turcomana Fischer von Waldheim, 1843 (Fig. 18) Fischer von Waldheim, 1844: 88; Gebler, 1844: 103 (“ Blaps caudata Gebler, 1844 ”); Skopin, 1960: 50 (“ B. caudata ”) (larva); Skopin, 1961: 189 (“ B. caudata ”); Skopin, 1966: 337 (“ B. caudata ”); Skopin, 1968: 85 (“ B. caudata ”); Ren et al. 2016: 121. Material examined (ZIN). 1 m, 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / Bayserke village / 1907 / leg. V. Nedzeltsky’ [in Cyrillics] [43 ° 29 ’ 28.18 ” N, 77 ° 3 ’ 48.68 ” E]; 1 m, ‘ Southeast Kazakhstan / Alma-Ata Region / Alma-Ata / 25. iv. 1950 / leg. N. G. Skopin’ [in Cyrillics] [43 ° 12 ’ 12.60 ” N, 76 ° 50 ’ 37.26 ” E]; 5 mm, 3 ff, ‘ Southeast Kazakhstan / Alma-Ata Region / Anrahai Mountains / canyon of Kopaly River / 14. v. 1909 / leg. Nedovpukov’ [in Cyrillics] [44 ° 1 ’ 11.62 ” N, 75 ° 5 ’ 54.61 ” E]; 1 m, 1 f, ‘ Southeast Kazakhstan / Semirechye / Kurdayskiy pass / h = 1100 m / 16. v. 1907 / leg. A. Jakobson’ [in Cyrillics] [43 ° 16 ’ 23.25 ” N, 74 ° 49 ’ 37.78 ” E]; 4 mm, 3 ff, ‘ Southeast Kazakhstan / Alma-Ata Region / Verniy (Alma-Ata) / 25. vii. 1909 / leg. Nedovpuk [ov] ’ [in Cyrillics] [43 ° 12 ’ 12.60 ” N, 76 ° 50 ’ 37.26 ” E]; 3 mm, 5 ff, ‘ Southeast Kazakhstan / Alma-Ata Region / Jambyl region / Korday village / 07. vii. 1906 / leg. N. Radkevich’ [in Cyrillics] [43 ° 2 ’ 14.36 ” N, 74 ° 42 ’ 50.64 ” E]; 4 mm, 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / Alma-Ata city suburbs / 19. vi. 1909 / leg. Nedovpuk [ov] ’ [in Cyrillics] [43 ° 12 ’ 12.60 ” N, 76 ° 50 ’ 37.26 ” E]; 2 mm, 3 ff, ‘ Southeast Kazakhstan / Alma-Ata Region / Anrahai Mountains / Almalysai River / 10 – 14. v. 1909 / leg. Nedovpukov’ [in Cyrillics] [44 ° 0 ’ 6.33 ” N, 75 ° 48 ’ 49.63 ” E]; 2 mm, 1 f, ‘ Kyrgyzstan / Jalal-Abad Region / Kara-Kul city / 27. vi. 1913 / leg. Chernivin’ [in Cyrillics] [41 ° 37 ’ 47.53 ” N, 72 ° 40 ’ 14.68 ” E]; 1 m, ‘ Kyrgyzstan / Naryn Region / Kokomeren River / 18 – 19. v. 1914 / leg. Mikhalevskaya’ [in Cyrillics]. Distribution. Kazakhstan, Uzbekistan, Kyrgyzstan, China (Nei Mongol and Xinjiang provinces) (LÖBL et al. 2008, REN et al. 2016). Regional distribution. The distribution overlaps with that of the nominotypical subspecies (SKOPIN 1968).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF77AB603FE74C0ADFE65FD6C.taxon	description	(Figs 19, 27 A)	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF77AB603FE74C0ADFE65FD6C.taxon	materials_examined	Type material examined (ZIN). Holotype (m), allotype (m) and paratypes (2 ff): ‘ Eastern Kazakhstan / east coast of Lake Alakol / 20 km north of Uzynbulak village / 06. vii. 1965 / leg. N. G. Skopin’ [in Cyrillics] [45 ° 54 ’ 44.73 ” N, 82 ° 10 ’ 50.51 ” E]. Comments. Skopin (1966) described this species (after study of six specimens available to him), but did not indicate the number of males and females in the description. There are four specimens in the collection of ZIN: the holotype, the allotype (male!) and 2 paratypes. Distribution. Kazakhstan: the area surrounding Alakol Lake and Zhalanashkol Lake (SKOPIN 1966).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF77AB602FEADC2D1FD88FD3D.taxon	description	(Figs 20, 21 C, D) Fischer von Waldheim, 1844: 71 (Desertis Kirgisorum); Skopin, 1960: 57 (larva); Skopin, 1961: 192; Skopin, 1968: 86; Arnoldi & Medvedev, 1969: 403; Medvedev & Nepesova, 1985: 119 (Blaps scutellata); Chigray et al., 2016: 9. Material examined (ZIN). 2 mm, ‘ Southeast Kazakhstan / Jambyl Region / Moyynkum village / 20. vii. 1960 / leg. L. Serkova’ [in Cyrillics] [44 ° 15 ’ 51.05 ” N, 72 ° 55 ’ 15.69 ” E]; 1 m, ‘ Southeast Kazakhstan / Jambyl Region / Muyunkum village / 22. iv. 1954 / leg. L. Serkova’ [in Cyrillics] [44 ° 15 ’ 51.05 ” N, 72 ° 55 ’ 15.69 ” E]; 1 m, 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / 40 km south-west of Zharkent city / 26. iv. 1965 / leg. N. G. Skopin’ [in Cyrillics] [43 ° 54 ’ 33.36 ” N, 79 ° 38 ’ 30.23 ” E]; 1 m, ‘ Southeast Kazakhstan / Alma-Ata Region / Sasykbulak spring / Ulken-Bugutty mountains / 29. vi. 1960 / leg. N. Skopin’ [in Cyrillics] [43 ° 32 ’ 25.36 ” N, 79 ° 4 ’ 15.09 ” E]; 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / Kokishbay village / lower course of Ili River / Rondy duct / 16. ix. 1948 / leg. D. Aleksandrov’ [in Cyrillics] [44 ° 58 ’ 42.14 ” N, 75 ° 33 ’ 42.53 ” E]; 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / Saryyesik- Atyrau sands / 12. x. 1953 / leg. V. Chekalin’ [in Cyrillics] [45 ° 30 ’ 44.27 ” N, 77 ° 1 ’ 55.41 ” E]; 1 f, Southeast Kazakhstan / Alma-Ata Region / Sarybulak village / 06. vii. 1907 / leg. A. Jakobson’ [in Cyrillics] [43 ° 14 ’ 57.69 ” N, 74 ° 17 ’ 35.90 ” E]; 1 m, ‘ Southeast Kazakhstan / Semirechye Region / Dzharkentskiy uyezd / Ili River / 18. iv. 1906 / leg. V. Ryunbeyl’ [in Cyrillics]; 1 m, 3 ff, ‘ Semirechye province. / Galford Mountain pass’ [in Cyrillics]. Comments. ARNOLDI and MEDVEDEV (1969) distinguished two species: Blaps holconota and Blaps scutellata Fischer von Waldheim, 1844. These names are listed in the catalogue of LÖBL et al. (2008) as concerning different species, but the authors of this paper use the name Blaps holconota because didn’t find distinct differences between these two taxa (CHIGRAY et al. 2016: 9). Distribution. Northern and Southern Kazakhstan, Uzbekistan, Turkmenistan, Afghanistan (SKOPIN 1961, ARNOLDI & MEDVEDEV 1969, LÖBL et al. 2008). Regional distribution. Desert areas of South Kazakhstan (SKOPIN 1961, ARNOLDI & MEDVEDEV 1969).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF77BB602FE3AC301FE0FFB50.taxon	materials_examined	Material examined (ZIN). 1 m, ‘ Kazakhstan / middle of Syr Darya River / Kyzylorda / 30. v. 1933 / leg. N. G. Skopin’ [in Cyrillics]; 2 mm, ‘ South Kazakhstan / left bank of Syr Darya River / Tugay / between Shardara and Kok-su / 11. v. 1964 / leg. N. G. Skopin’ [in Cyrillics]. Distribution. Kazakhstan, Uzbekistan (from Namangan to Amu Darya River delta), Turkmenistan (MEDVEDEV & NEPESOVA 1985 (Amu Darya River delta); LÖBL et al. 2008). Regional distribution. The species is distributed in Syr Darya River valley (SKOPIN 1968).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF77BB602FE51C4FEFDC3FA14.taxon	materials_examined	Type material examined (ZIN). Holotype and paratypes from both localities were studied (MEDVEDEV 2004). Distribution. Kazakhstan: Atyrau Region (Kulsary), North Aral Sea coast (Butakov gulf) (MEDVEDEV 2004).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF778B601FE95C0ADFEA6FD11.taxon	materials_examined	Type material examined (ZIN). LECTOTYPE of Blaps robusta (designated by ABDURA- KHMANOV & NABOZHENKO 2011): f, ‘ Lehmann // Blaps robusta Motsch // Lectotypus / Blaps robusta / des. Abdurakhmanov & Nabozhenko’. LECTOTYPE of Blaps anthracina (designated by ABDURAKHMANOV & NABOZHENKO 2011): f, ‘ Caucas. // anthracina Fald. Caucas. // Lectotypus / Blaps anthracina / des. Abdurakhmanov & Nabozhenko’. Material examined (ZIN). More than 60 specimens from Western Kazakhstan Region. Distribution. Europe (everywhere, north to Scandinavia and the European part of Russia), North Africa, the Middle East, North Caucasus, Western Siberia (ABDURAKHMANOV & NABOZHENKO 2011), Western (CHIGRAY et al. 2016), North and NE Kazakhstan (SKOPIN 1960), China: Xijiang (REN at al. 2016). Regional distribution. Karaganda Region (SKOPIN 1960, ARNOLDI & MED- VEDEV 1969).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF778B600FE9AC33DFCF6FDB9.taxon	description	(Figs 22, 36 A) Motschulsky, 1845: 65 (“ Blaps gigantea ”); Seidlitz, 1893: 268 (“ B. gigantea ”); BOGATCHEV, 1947: 515 (B. motschulskiana, new name for B. gigantea Motschulsky, 1845 nec B. gigantea L. Petagna, 1819); Skopin, 1968: 87 (“ Lithoblaps gigantea ”). Type material examined (ZIN). Lectotype of Blaps gigantea Motschulsky, 1845 is designated here: m, ‘ Blaps gigantea Motschulsky / Tekke. // Lectotypus / Blaps gigantea Motschulsky, 1845 / des. I. A. Chigray’. Material examined (ZIN). 1 m, ‘ Turkmenistan / Karahan / 15. v. 1976 / leg. G. Medvedev’ [in Cyrillics]; 1 f, ‘ Turkestan’; 1 f ‘ Turkmenistan / Trans-Caspian Region / Ashgabat / 3. v. 1889 / leg. A. Semenov’ [in Cyrillics]; 1 m ‘ Turkmenistan / Trans-Caspian Region / Ashgabat / 2. v. 1889 / leg. A. Semenov’ [in Cyrillics]; 2 ff ‘ Trans-Caspi G. / Turcmenien / leg. E. König’; 1 f ‘ Turkmenistan / Ashgabat / 1986 / leg. Varentsev’ [in Cyrillics]. Distribution. Kazakhstan (SKOPIN 1968), Turkmenistan, Afghanistan (LÖBL et al. 2008). Regional distribution. Southeastern parts of Kyzylkum desert and Syr Darya River near the border of Uzbekistan (SKOPIN 1968).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF779B60FFE6AC285FBBEFDE1.taxon	description	(Fig. 23) Fischer von Waldheim 1844: 96; Seidlitz, 1893: 280; Skopin, 1961: 190. Material examined (ZIN). 1 m, 1 f, ‘ Central Kazakhstan / Karaganda Region / 30 km west of Gulshad village [46 ° 37 ’ 27.91 ” N, 73 ° 57 ’ 55.11 ” E] / 17. ix. 1964 / leg. N. G. Skopin’ [in Cyrillics]. Comments. SKOPIN (1973) discussed the taxonomic position of Blaps nitida Fischer von Waldheim, 1844 and the very variable B. lethifera Marsham, 1802. The first taxon is listed as a subspecies of B. lethifera in the card index and the collection of ZIN, but as a valid species in the Catalogue of Palaearctic Coleoptera (LÖBL et al. 2008). The status of B. nitida can be clarified after the study of type specimens. In the subsequent key, B. nitida and B. lethifera are given in the same key couplet. ARNOLDI and MEDVEDEV (1969) did not mention B. nitida for Central Kazakhstan. Distribution. Kazakhstan, Russia (Western Siberia) (LÖBL et al. 2008). Regional distribution. Central and Southeast Kazakhstan (SKOPIN 1961).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF776B60FFEBEC22DFECAFB2A.taxon	materials_examined	Material examined. 1 f, ‘ Kazakhstan / Aral Sea / Barsa-Kelmes Island [a former island of Aral Sea] / 20. vi. 1940 / leg. Zvorygin’ [in Cyrillics]. Additional material examined. More than 50 specimens from Western Kazakhstan. Comments. The records for Karaganda and Jambyl regions (ARNOLDI & MEDVEDEV 1969) are erroneous and probably belong to the subspecies Blaps parvicollis quadricollis Ballion, 1878. Differentiation of the subspecies by females is difficult. The female studied from Barsakelmes Reserve (the western part of Southern Kazakhstan) has the body shape and the structure of genitalia similar to those of the nominotypical subspecies from Western Kazakhstan. The authors of this paper have not seen any male of the nominotypical subspecies from South Kazakhstan. Distribution. South of European part of Russia, Azerbaijan (Apsheron), Kazakhstan (ABDURAKHMANOV & NABOZHENKO 2011) Regional distribution. The Barsakelmes Nature Reserve (a former island in the Aral Sea).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF776B60CFE87C517FD93FDE1.taxon	description	(Fig. 24) Ballion, 1878: 304; Skopin, 1973: 868 (B. parvicollis, subsp.).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF776B60CFE87C517FD93FDE1.taxon	materials_examined	Type material examined (NHM). LECTOTYPE of Blaps subcordata (designated by Skopin): m, ‘ Haberhauer / Turkestan / 89 // Lectotypus / Blaps subcordata Soll. / N. Skopin design. / 1978 // Blaps parvicollis quadricollis Ball. / N. Skopin det. / 1978 ’. PARALECTOTYPES: 1 m, 1 f, ‘ Haberhauer / Turkestan / 89 ’. Material examined. ZIN. 1 m, 2 ff, ‘ Southeast Kazakhstan / Alma-Ata Region / Anrahai Mountains / canyon of Kopaly River / 16 – 19. v. 1909 / leg. Nedovpuk [ov] ’ [in Cyrillics] [44 ° 1 ’ 11.62 ” N, 75 ° 5 ’ 54.61 ” E]; 2 mm, 2 ff, ‘ Southeast Kazakhstan / Alma-Ata Region / canyon of Kopaly River / 19. v. 1909 / leg. Nedovpuk [ov] ’ [in Cyrillics] [44 ° 1 ’ 11.62 ” N, 75 ° 5 ’ 54.61 ” E]; 2 ff, ‘ Southeast Kazakhstan / Jambyl region / Akyr-Tyube village / 25. v. 1931 / leg. Veltishchev’ [in Cyrillics] [42 ° 59 ’ 31.57 ” N, 72 ° 6 ’ 5.64 ” E]; 2 mm, ‘ Southeast Kazakhstan / Alma-Ata Region / Ili-Karatal along Turkestan-Siberian Railway / 05. vi. 1930 / leg. V. Kizeritskiy’ [in Cyrillics]; 1 m, 1 f, ‘ Southeast Kazakhstan / Semirechye Region / Kunkuzskaya Upland (Altyn-Emel Range) to north of Aladan / iv. 1879 / leg. E. Regel’; 1 m, 1 f, ‘ Southeast Kazakhstan / Semirechie / upper reaches of River Bota-Boruly / 20. v. 1909 / leg. Nedovpuk [ov] ’ [in Cyrillics]; 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / Alma-Ata city suburbs / 03. v. 1908 / leg. Nedzvetszkiy’ [in Cyrillics] [43 ° 12 ’ 12.60 ” N, 76 ° 50 ’ 37.26 ” E]; 1 m, 2 ff, ‘ Southeast Kazakhstan / env. of Jambyl / Merke village / iv. 1910 / leg. E. Fischer’ [42 ° 52 ’ 5.52 ” N, 73 ° 11 ’ 46.21 ” E]; 1 m, ‘ Southeast Kazakhstan / Alma-Ata Region / Uzun- Kargaly River / 15. iv. 1909 / leg. Nedzvetszkiy’ [in Cyrillics] [43 ° 6 ’ 51.03 ” N, 76 ° 21 ’ 34.08 ” E]. Comments. SKOPIN (1960) suggested that Blaps parvicollis subcordata Seidlitz, 1893 could be the eastern subspecies of B. parvicollis, and indicated that the larvae of both subspecies are very similar in a structure. Later, SKOPIN (1973) synonymized B. parvicollis subcordata with B. parvicollis quadricollis Ballion, 1878 and pointed to the error of SEMENOV TIAN­SHANSKY and BOGATCHEV (1936), who noted that B. quadricollis is indistinguishable from B. pterosticha Fischer von Waldheim, 1844. LÖBL et al. (2008) erroneously listed Blaps parvicollis subcordata as a separate species and B. parvicollis quadricollis as a junior synonym of B. pterosticha. Blaps parvicollis quadricollis differs from B. pterosticha in smaller body size, coarser pronotal bead, male metatibia with thickening in apical half and subglobular reservoirs of spermatheca. Based on the more elongated pronotum and elytra, coarser and thicker pronotal bead and male metatibia with thickening in the apical half (Figs 25 A, B), structures of male and female genitalia and also presence of populations with intermediate characters in intergradation zones, it is thought that B. p. quadricollis is a subspecies of B. parvicollis. Separation of females is difficult, as the body shape (in particular structure of pronotum and elytra) and the structure of spermatheca are rather variable. Some authors (MEDVEDEV 2001, CHIGRAY et al. 2016) illustrated female genital ducts of B. p. parvicollis. Recent studies revealed that the genital ducts of each of both subspecies of B. parvicollis in Southern Kazakhstan are represented by three types: 1) bases of both reservoirs of the spermatheca are joined in a common duct at a distance from the basal duct (Fig. 25 C); 2) bases of reservoirs of the spermatheca are joined and do not form a common tube before the basal duct (Fig. 25 D); 3) bases of reservoirs of the spermatheca are separately joined with the basal duct (Fig. 25 E). Distribution. The subspecies is widely distributed in the southern zone of Kazakhstan (SKOPIN 1968).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF775B60CFE23C22DFD90FA84.taxon	materials_examined	Type material examined (ZIN). Lectotype of Blaps pruinosa (designated by ABDURA- KHMANOV & NABOZHENKO 2011): m, ‘ Kirgis Step // Lehmann // Blaps pruinosa Eversm. Kirgs. Desert // golden square // Lectotypus / Blaps pruinosa Eversmann / des. Abdurakhmanov & Nabozhenko’. Lectotype of Blaps rorulenta Motschulsky, 1845 (designated by ABDURA- KHMANOV & NABOZHENKO 2011): f, ‘ rorulenta // Rhisoblaps rorulenta Motch. Songoria // c. Motschulsky // golden square // Lectotypus / Blaps rorulenta 1845 / des. Abdurakhmanov & Nabozhenko’. Material examined (ZIN). 1 m, 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / 40 km south-west of Zharkent village / 26. iv. 1965 / leg. N. G. Skopin’ [in Cyrillics] [43 ° 59 ’ 24.43 ” N, 79 ° 34 ’ 3.92 ” E]. Additional material examined (ZIN). More than 50 specimens from the Caspian Depression. Distribution. Southeast of the European part of Russia, Kazakhstan (from the Caspian sea to southern foothills of Tarbagatay Mts.), Uzbekistan, Turkmenistan, Tajikistan (MEDVEDEV & NEPESOVA 1985, ABDURAKHMANOV & NABOZHENKO 2011). Regional distribution. The species is widely distributed in the central (ARNOLDI & MEDVEDEV 1969) and southern zones of Kazakhstan to eastern Balkhash Lake (SKOPIN 1961).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF775B60AFE88C58AFD4BFC33.taxon	description	(Figs 26, 27 C) Fischer von Waldheim, 1844: 93; Seidlitz, 1893: 286 (“ Turkestan ”); Skopin, 1960: 55 (larva); Skopin, 1961: 191; Skopin, 1968: 86; Ren et al., 2016: 153. Material examined (ZIN). 1 m, 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / Konyrolen village / 16. vii. 1957 / leg. N. G. Skopin’ [in Cyrillics] [44 ° 16 ’ 36.38 ” N, 79 ° 18 ’ 5.33 ” E]; 1 m, 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / 40 km south-west of Zharket village / 26. iv. 1965 / leg. N. G. Skopin’ [in Cyrillics] [43 ° 59 ’ 24.43 ” N, 79 ° 34 ’ 3.92 ” E]; 1 m, ‘ Southeast Kazakhstan / Alma-Ata Region / 30 km north of Taldykorgan city / 07. ix. 1962 / leg. N. G. Skopin’ [in Cyrillics] [45 ° 18 ’ 9.84 ” N, 78 ° 21 ’ 9.40 ” E]; 1 m, ‘ Southeast Kazakhstan / Alma-Ata Region / Kurtogay natural boundary / 15. vii. 1959 / leg. N. G. Skopin’ [in Cyrillics] [43 ° 15 ’ 52.20 ” N, 78 ° 58 ’ 25.09 ” E]; 2 f, ‘ Southeast Kazakhstan / Alma-Ata Region / 30 km south of Dubun village / 10. vi. 1969 / leg. N. G. Skopin’ [in Cyrillics] [43 ° 26 ’ 5.87 ” N, 80 ° 11 ’ 13.00 ” E]; 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / Kurtogay natural boundary / 12. v. 1961 / leg. N. G. Skopin’ [in Cyrillics] [43 ° 15 ’ 52.20 ” N, 78 ° 58 ’ 25.09 ” E]; 1 m, 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / 40 km north of Zharkent village / 14. v. 1961 / leg. N. G. Skopin’ [in Cyrillics] [44 ° 31 ’ 44.58 ” N, 79 ° 59 ’ 36.85 ” E]; 1 m, 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / Saryzhaz village / 25. vi. 1966 / leg. N. G. Skopin’ [in Cyrillics] [42 ° 54 ’ 31.02 ” N, 79 ° 35 ’ 49.30 ” E]; 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / Kapchagay city / 29. iii. 1962 / leg. N. G. Skopin’ [in Cyrillics] [43 ° 51 ’ 40 ” N, 77 ° 2 ’ 45.73 ” E]; 1 m, 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / Zharkent village / 12. v. 1961 / leg. N. G. Skopin’ [in Cyrillics] [44 ° 9 ’ 24.73 ” N, 79 ° 59 ’ 53.04 ” E]. Comments. SKOPIN (1961) showed that Blaps pterosticha and B. tenuicauda Seidlitz, 1893 are morphologically similar species and their lifestyles are the same; he suggested that B. tenuicauda is only a form of B. pterosticha. Recent studies confirmed this opinion based on the absence of any significant external and internal structural differences between the specimens formerly considered as these two “ taxa ”, including those provided by Skopin with the label “ Homotypus ”. The validity of these taxa needs a confirmation by reexamination of the type specimens of both “ species ”. In the subsequent key B. pterosticha and B. tenuicauda are given in the same couplet. Distribution. Kazakhstan, Kyrgyzstan, China (Xizang province), Mongolia (LÖBL et al. 2008, REN et al. 2016), Afghanistan (KASZAB 1970) Regional distribution. North of the Muyunkum Desert, Middle Irtysh region, Balkhash Lake region (SKOPIN 1961).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF773B634FDE4C41EFD10FEDB.taxon	description	(Figs 28 – 33, 37 C) Type material. Holotype (m) and paratypes (1 m, 2 ff): ‘ Southeast Kazakhstan / Ketmen Range / 7 km SW Aktam village / 43 ° 23 ’ 21.6 ’’ N, 79 ° 52 ’ 48.3 ’’ E / h = 1376 m / 30. v. 2015 / leg. A. V. Ivanov’ (ZIN); Paratypes: 1 m, 1 f: ‘ Southeast Kazakhstan / Sugates Mountains / 4. ix. 1968 / leg. N. G. Skopin’ (ZIN); 1 m, 3 ff: ‘ Southeast Kazakhstan / Ketmen Range / 4 km N Talas village / 43 ° 10 ’ 07.2 ’’ N, 79 ° 47 ’ 38.6 ’’ E / h = 2185 m / 24. iv. 2018 / leg. A. V. Ivanov’ (ZIN); 2 mm: ‘ Southeast Kazakhstan / Ketmen Range / 4 km N Talas village / 43 ° 10 ’ 07.2 ’’ N, 79 ° 47 ’ 38.6 ’’ E / h = 2185 m / 09. iv. 2019 / leg. A. V. Ivanov’ (IEPaAY, ZIN); 1 m: ‘ Southeast Kazakhstan / 14 km NW Aktogay village / 43 ° 17 ’ 43.9 ’’ N, 78 ° 59 ’ 27.7 ’’ E / h = 1006 m / 24. v. 2016 / leg. A. V. Ivanov’ (ZIN). Description. Male. Body black, mat, slender. Anterior margin of epistoma weakly emarginate, straight in middle. Lateral margins of epistoma straight. Lateral margins of genae straight in anterior half, rounded at base. Lateral margins of head with indistinct emargination between epistoma and genae. Head widest at level of eyes and temples. Head 1.41 times as wide as interocular distance. Antennomeres 10 – 11 reaching base of pronotum when directed backwards. Ratio of length / width of antennomeres 2 – 11 as 6 (8), 31 (9), 11 (9), 11 (9), 11 (9), 15 (11), 10 (8), 10 (9), 10 (10), 14 (10). Mentum hexagonal, with weak outer angles. Punctation of head sparse, dense (distance between punctures subequal to or smaller than puncture diameter), bottom of punctures with one microgranule. Pronotum transverse (1.32 times as wide as long), widest at middle, 1.94 times as wide as head. Ratio of pronotal width near anterior angles to widest part and that at base 4.1: 7.8: 6.2. Disc of pronotum weakly convex, narrow flattened along lateral sides and base. Anterior margin of pronotum widely emarginate, lateral margins widely rounded in anterior third and weakly rounded in posterior half, base of straight. Disk completely beaded except for apical middle. Anterior angles widely rounded, posterior angles narrowly rounded and right. Punctures of pronotum similar to those on head, strongly dense, puncture subcontiguous, sparser in middle (distance between punctures smaller than puncture diameter). Prothoracic hypomera with small wrinkles and covered with sparse small granules. Hypomera along lateral margins narrowly excavate. Elytra weakly convex, elongate (2.1 times as long as wide together), widest at middle, 3.86 times as long and 1.37 times as wide as pronotum, 2.39 times as wide as head. Caudal extension of elytra (mucro) distinct, 5 mm long; elytra 4 times as long as mucro. Elytra with rasp-like punctures, punctation at sides becoming denser. Epipleura with fine wrinkles and sparse fine rasp-like punctation. Hair tuft between abdominal ventrites 1 and 2 present. Abdominal ventrites 1 – 3 with large transverse wrinkles in middle, ventrites 4 – 5 without wrinkles. Abdominal ventrites 1 – 3 covered with sparse small granules and rasp-shaped punctures, ventrite 4 with only rasp-like punctures, ventrite 5 with simple punctures, completely beaded except for base. Legs slender. Ratio of lengths of femora, tibiae and tarsi of fore, middle and hind legs 6.5: 5.5: 3.2; 7.2: 6.3: 4.1; 9.1: 8.2: 4.7. All tarsomeres with bifurcated setal brush. Anterior margin of male inner sternite VIII weakly emarginate, straight in middle, accessory gland of sternite VIII moderately long and thin, middle of sternite without hairs. Rods of spiculum gastrale merged at apex, forming long common stem, lobes of spiculum gastrale elongated. Aedeagus length 4.3 – 4.4 mm, width 0.8 – 0.9 mm. Aedeagus weakly Cshaped. Basal third of lateral margins of parameres almost straight, middle widely round- ed, apical third strongly acuminate at apex. Parameres length 1.4 mm, width 0.5 mm. Sides of parameres with two longitudinal impressions: one longer beginning from base and another shorter and disposed at middles. Body length 27 – 28 mm, width 9.7 – 9.9 mm. Female. Shape of body and punctation similar to those of male. Head 1.26 times as wide as interocular distance. Antennomeres 10 – 11 reaching base of pronotum when directed backwards. Ratio of pronotal width near anterior angles, in widest part and at base 3.8: 6.9: 6.4. Elytra elongate (1.72 times as long as wide together), 3.16 times as long and 1.38 times as wide as pronotum, 2.7 times as wide as head. Mucro short (1.6 mm). Ovipositor moderately long. Apical lobes straight in basal thirds, weakly arcuately rounded in middle, apical third weakly arcuately emarginate. Apex of lobes acute. Anterior margin of proctiger with deep narrow emarginated in middle. Basal duct of spermatheca between vagina and reservoirs long, gland of spermatheca short. Bases of reservoirs of spermatheca thin, their apical half wider, 1 st reservoir twice larger than 2 nd. Stem of spiculum ventrale moderately long and thin. Body length 22 – 24 mm, width 9 – 9.2 mm. apical piece, dorsal view; E = the same, ventral view; F = the same, lateral view. Scale bars: 30 µm for D, E, 100 µm for B, F, 200 µm for C, 300 µm for A Etymology. The species is named after the late Nikolay Georgievich Skopin famous specialist on Central Asian Tenebrionidae. Differential diagnosis. The body shape and punctation of this new species is similar to B. granulata, B. granulipennis and B. tsharynensis. Blaps skopini sp. n. differs from: − B. granulata in the more elongate pronotum (pronotum of B. granulata 1.4 times as wide as long); longer mucro of males (male elytra of B. granulata 5.5 times as long as mucro); in contrast to those of Blaps skopini sp. n., the sides of parameres of B. granulata are smooth, without longitudinal impression, slightly and gradually arched to apex (viewing from above), apex of the latter is straight (Fig. 28 B); − B. granulipennis in the coarser and denser punctation of the pronotum (punctures in Blaps skopini sp. n. are subcontiguous, while in B. granulipennis are subcontiguous everywhere except for middle); longer mucro of males and females (elytra of B. granulipennis 7.9 times as long as mucro); sides of parameres of B. granulipennis are widely rounded to the apex, with obtuse distinct emargination near the apex; lateral sides of apex of parameres are straight and parallel relative to each other (Fig. 27 A); − B. tsharynensis in the coarser and denser punctation of the pronotum (punctures in Blaps skopini sp. n. are subcontiguous, while they in B. tsharynensis are not contiguous); longer mucro of males and females (elytra of B. tsharynensis 10.5 times as long as mucro); larger and more distinct hair tuft between abdominal ventrites 1 and 2 (Figs 35 B); parameres of Blaps skopini are elongated (2.48 times as long as wide), while parameres of B. tsharynensis are shorter (1.8 times as long as wide); joined apex of parameres in Blaps skopini sp. n. is thin and long (Fig. 30), while ca, r = reservoirs, g = accessory gland of spermatheca). Scale bars = 1 mm that in B. tsharynensis is thin, but short; reservoirs of spermatheca of Blaps skopini sp. n. are thin and fusiform, while reservoirs of spermatheca in B. tsharynensis are elliptic; stem of spiculum ventrale of the new species is thinner than that in B. tsharynensis.	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF74DB634FE28C167FC9AFCFC.taxon	description	(Fig. 33) Seidlitz 1893: 307; Skopin, 1960: 56 (larva); Skopin, 1961: 191; Skopin, 1968: 86. Material examined (ZIN). 1 f, ‘ South Kazakhstan / South-Kazakhstan Region / Sharapkhana village / 06. v. 1964 / leg. N. G. Skopin’ [in Cyrillics] [41 ° 51 ’ 52.60 ” N, 69 ° 26 ’ 43.72 ” E]; 1 m, ‘ South Kazakhstan / South-Kazakhstan Region / Arys city / 14. v. 1967 / leg. N. G. Skopin’ [in Cyrillics] [42 ° 26 ’ 42.32 ” N, 68 ° 51 ’ 31.23 ” E]. Distribution. Kazakhstan (LÖBL et al. 2008), Uzbekistan (SKOPIN 1961). Regional distribution. South Kazakhstan, hilly steppe between Tashkent and Shymkent cities (SKOPIN 1961); Syrdarya River region (SKOPIN 1968). Comments. See comments at Blaps pterosticha.	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF74AB633FED8C0ADFDAFFCF0.taxon	description	(Fig. 34) Fischer von Waldheim, 1844: 105; Seidlitz 1893: 285 (“ Turkestan ”); Skopin, 1960: 55 (larva); Skopin, 1961: 191; Skopin, 1968: 86; Arnoldi & Medvedev, 1969: 404; Ren et al. 2000; 27; Ren et al., 2016: 171. Material examined (ZIN). 1 m, ‘ Southeast Kazakhstan / Alma-Ata Region / canyon of Kopaly River / 19. v. 1909 / leg. Nedovpuk [ov] ’ [in Cyrillics] [44 ° 1 ’ 11.62 ” N, 75 ° 5 ’ 54.61 ” E]; 1 m, ‘ Southeast Kazakhstan / env. of Jambyl / Ulanbel village / 02. v. 1960 / leg. L. Serkova’ [in Cyrillics] [44 ° 49 ’ 34.59 ” N, 71 ° 8 ’ 17.27 ” E]; 1 m, ‘ Southeast Kazakhstan / Jambyl / Khantau mountains / 25. vi. 1969 / leg. K. Nurpeisov’ [in Cyrillics] [44 ° 13 ’ 28.19 ” N, 73 ° 55 ’ 28.64 ” E]; 1 f, ‘ Kyrgyzstan / Bishkek city / 15. v. 1907 / leg. A. Jakobson’ [in Cyrillics] [42 ° 53 ’ 10.32 ” N, 74 ° 32 ’ 5.29 ” E]; 1 m, ‘ Kyrgyzstan / Chu River / Boom canyon / 30. iv. 1956 / leg. N. G. Skopin’ [in Cyrillics] [42 ° 40 ’ 30.86 ” N, 75 ° 53 ’ 35.95 ” E]. Distribution. Kazakhstan (ARNOLDI & MEDVEDEV 1969), China (LÖBL et al. 2008), Kyrgyzstan (SKOPIN 1961, 1968). Regional distribution. Between Talas and the Chu rivers (SKOPIN 1961); eastern part of the Karatau Range, the Muyunkum Desert and valley area near the Chu River (SKOPIN 1968).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF74BB631FE95C0ADFEB1FB57.taxon	description	(Figs 35, 36 B)	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF74BB631FE95C0ADFEB1FB57.taxon	materials_examined	Type material examined (ZIN). Holotype: m, ‘ Southeast Kazakhstan / Alma-Ata Region / Kurtogay natural boundary / 21. vii. 1959 / leg. N. G. Skopin’ [in Cyrillics] [43 ° 15 ’ 52.20 ” N, 78 ° 58 ’ 25.09 ” E]. Allotype: f, ‘ Southeast Kazakhstan / Alma-Ata Region / Kurtogay natural boundary / 21. vii. 1959 / leg. N. G. Skopin’ [in Cyrillics] [43 ° 15 ’ 52.20 ” N, 78 ° 58 ’ 25.09 ” E]. Paratypes: 1 m, ‘ Southeast Kazakhstan / Alma-Ata Region / middle course of Charyn River / 23. iv. 1960 / leg. N. G. Skopin’ [in Cyrillics]; 1 m, ‘ Southeast Kazakhstan / Alma-Ata Region / Kurtogay natural boundary / 12. v. 1961 / leg. N. G. Skopin’ [in Cyrillics] [43 ° 15 ’ 52.20 ” N, 78 ° 58 ’ 25.09 ” E]; 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / Kurtogay natural boundary / 05 – 19. vii. 1959 / leg. N. G. Skopin’ [in Cyrillics] [43 ° 15 ’ 52.20 ” N, 78 ° 58 ’ 25.09 ” E]. Material examined (ZIN). 1 m, 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / Lavar village suburbs / 04. vi. 1966 / leg. N. G. Skopin’ [in Cyrillics] [43 ° 34 ’ 4.62 ” N, 78 ° 5 ’ 22.28 ” E]; 2 mm, 2 ff, ‘ Southeast Kazakhstan / Alma-Ata Region / 40 km south-west of Dzharkent city / 26. iv. 1965 / leg. N. G. Skopin’ [in Cyrillics] [43 ° 55 ’ 58.35 ” N, 79 ° 35 ’ 39.24 ” E]; 1 m, 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / 60 km west of Gulshat village / 17. ix. 1964 / leg. N. G. Skopin’ [in Cyrillics] [46 ° 36 ’ 57.83 ” N, 73 ° 34 ’ 7.28 ” E]; 1 m, 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / lower reaches of Karatal River / 28. v. 1965 / leg. N. G. Skopin’ [in Cyrillics] [46 ° 8 ’ 18.41 ” N, 77 ° 9 ’ 44.45 ” E]. Distribution. Kazakhstan (LÖBL et al. 2008). Regional distribution. The Kurtogay canyon, Ili River region (near the spring of Ayak-Kalkan) (SKOPIN 1961).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF74BB631FE95C0ADFEB1FB57.taxon	materials_examined	Type material examined (ZIN). Holotype: m, ‘ Kazakhstan / North of Balkhash Lake / 60 КМ W Gulshat village / 17. ix. 1964 / leg. N. G. Skopin’ [in Cyrillics]. Paratypes: 1 m, 1 f, ‘ Central Kazakhstan / Karaganda Region / 40 km south of Sayak village / 21. ix. 1964 / leg. N. G. Skopin’ [in Cyrillics] [46 ° 38 ’ 59.60 ” N, 77 ° 18 ’ 17.37 ” E]; 1 m, ‘ South Kazakhstan / right bank of Chu River / 13. ix. 1964 / leg. N. G. Skopin’ [in Cyrillics]. Material examined (ZIN). 5 m, ‘ Southeast Kazakhstan / Alma-Ata Region / low- er reaches of Karatal River / 28. v. 1965 / leg. N. G. Skopin’ [in Cyrillics] [46 ° 8 ’ 18.41 ” N, 77 ° 9 ’ 44.45 ” E]; 1 f, ‘ Southeast Kazakhstan / Alma-Ata Region / 30 km west of Gulshat village / 17. ix. 1964 / leg. N. G. Skopin’ [in Cyrillics] [46 ° 36 ’ 59.25 ” N, 73 ° 56 ’ 35.18 ” E]. Distribution. Central and Southeast Kazakhstan (Balkhash Lake region) (SKOPIN 1966).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF748B631FE6DC4FAFD75F9D4.taxon	description	(Figs 27 F, 38) Seidlitz, 1893: 305 Material examined (ZIN). 1 m, 1 f, ‘ Kazakhstan / Dzhambul [Jambyl] // Blaps turcomanorum ’. Distribution. Kazakhstan (first record for the country), Uzbekistan (LÖBL et al. 2008). Regional distribution. Southeast Kazakhstan.	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF746B63EFE2BC0ADFEE3FEDB.taxon	description	(Fig. 39) Seidlitz, 1893: 307; Skopin, 1961: 190; Ren et al. 2000; 27; Ren et al., 2016: 178. Material examined. NHM. 1 m, ‘ Haberhauer / Turkestan / 89 // Blaps akinina All // sagitta // Cotypus / Blaps sagitta Seidl. / N. Skopin design. / 1978 ’. ZIN. 1 m, 4 ff, ‘ Naryn mountains / Semirechye Region / 6. vi. 1905. / leg. Nezhivov’ [in Cyrillics]; 1 m, ‘ Naryn mountains / Semirechye Region / 8. vi. 1905. / leg. Nezhivov’ [in Cyrillics]; 2 ff, ‘ Naryn mountains / Semirechye Region / 11. vi. 1905. / leg. Nezhivov’ [in Cyrillics]; 1 m, Przhevalsk [Karakol] / 28. vi. 04 / leg. Nezhivov’ [in Cyrillics]; 2 ff, ‘ Naryn mountains / 9. vii. 1904 / leg. Nezhivov’ [in Cyrillics]; 3 ff, ‘ Naryn town / 10. vii. 1904 // leg. Herz’; 1 f, ‘ Naryn town / 11. vii. 1904 ’; 1 f, ‘ Naryn town / 20. vi. 04 / leg. Nezhivov’ [in Cyrillics]; 3 mm, 1 f, ‘ Southeast Kazakhstan / Dzungarian Alatau / 1. vi. 1968 / leg. N. G. Skopin’ [in Cyrillics]; 1 m, 1 f, ‘ Dzungarian Alatau / 5 km N Bien River / 22. v. 1968 / leg. N. G. Skopin’ [in Cyrillics]. Comments. A population of B. virgo with short mucro (1 – 1,5 mm) occurs near Bien River (Alma-Ata Region). Distribution. Kazakhstan, Kyrgyzstan, China (Xinjiang Region) (LÖBL et al. 2008, REN et al. 2016). Regional distribution. Northern slopes of Terskey Alatau Range (Narynkol Region).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF747B63EFE95C20BFC1BFBA1.taxon	materials_examined	Type material examined (ZIN). Holotype of Blaps convexicollis Motschulsky, 1845: m, ‘ Mt. Altai // Blaps convexicollis m. Altai’. Lectotype of Blaps confusa Ménétriés, 1832 designated here: m, ‘ Caucas. // confusa Ménét. Caucas. // Blaps longicollis // Lectotypus / Blaps confusa Ménétriés, 1832 / des. I. A. Chigray’. Material examined (ZIN). 2 ff, ‘ Kazakhstan / Zaisan district / 5. vii. [19] 26 / leg. Filatova’ [in Cyrillics]; 1 m, ‘ Kazakhstan / Zaisan / Neygy Kuma / into Achnatherum sp. / 10. vi. 1958 / leg. N. G. Skopin’ [in Cyrillics]; 1 f, ‘ Kazakhstan / Zaisan hollow / 10. vi. 1958 / leg. N. G. Skopin’; 1 f, ‘ Semirechye Region / Alatau / Kurdayskiy pass / 16. vii. 1906 / leg. N. Radkevich’ [in Cyrillics]. Distribution. Part of Central, Southern and Eastern Europe, the European part of Russia, the Great Caucasus, Southern Siberia, North Kazakhstan (ARNOLDI & MEDVEDEV 1969, ABDURAKHMANOV & NABOZHENKO 2011). Regional distribution. Karaganda Region (SKOPIN 1960).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF747B63EFE7EC167FCC3FE39.taxon	materials_examined	Type species: Blaps confusa Ménétriés, 1832 (= Blaps halophila Fischer von Waldheim, 1820) by the subsequent designation (MEDVEDEV & IWAN 2006).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF747B63CFE86C46DFC41FC84.taxon	materials_examined	Type species Blaps allardiana Reitter, 1889 by the original designation (SKOPIN & KASZAB 1978). Comments. MEDVEDEV (2001) interpreted the genus Ablapsis Reitter, 1887 as a subgenus of Blaps and some species of Prosoblapsia (Blaps (Ablapsis) compressipes Reitter, 1887, B. (A.) berezowskii G. S. Medvedev, 1998, B. (A.) lucidula G. S. Medvedev, 1998, B. (A.) znoikoi SEMENOV & BOGATCHEV, 1936 and B. (A.) gentilis gentilis Fairmaire, 1887) were included by him in the subgenus Ablapsis. However, further specialists have not accepted this opinion (LÖBL et al., 2008, REN et al., 2016). The taxonomic position and composition of Ablapsis and Prosoblapsia will be discussed in further works.	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF747B63CFE86C46DFC41FC84.taxon	description	Ballion, 1878: 301; Seidlitz, 1893: 296 (“ Turkestan ”); Skopin, 1960: 52 (larva); Skopin, 1961: 190; Skopin, 1968: 84; Skopin, 1977: 149; Ren et al., 2016: 196. Material examined (ZIN). 1 m, 1 f, ‘ Tian Shan, Musart // coll. N. Skopin’ [in Cyrillics]; 1 m, ‘ Ost – Turkestan / Acsu River / 1067 m. / v. 1903 / leg. Hauser // coll. N. Skopin’ [in Cyrillics]; 1 m, 1 f, ‘ Uzbekistan / Tian Shan / Przhevalsk [Karakol] / 3300 m. / vi. 77 ’ [in Cyrillics]; 1 m, ‘ Cat River [tributary of Ili River] / Dzungarian Region / leg. Regel’; 1 m, 1 f, ‘ Env. of Kuldzha / 22. iv. 79 / leg. A. Regel // Topotype // Homotype Blaps rudesculpta / det. N. Skopin’. Comments. Blaps transversimsulcata rudesculpta Semenov Tian-Shansky et Bogatchev, 1936 is a younger synonym of B. t. transversimsulcata (SKOPIN 1977), not of Blaps maeander Kraatz, 1885 (mistake in LÖBL et al. 2008). Distribution. Kazakhstan, China (Xinjiang Province) (LÖBL et al. 2008, REN et al. 2016). Regional distribution. Talas Ala-Too and Ugam Ranges (SKOPIN 1968).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
896187BFF747B63CFE86C46DFC41FC84.taxon	materials_examined	Type material examined (ZIN). Holotype: m, ‘ Southeast Kazakhstan / Alma-Ata Region / Alma-Ata city suburbs / 19. vi. 1957 / leg. N. G. Skopin’ [in Cyrillics] [43 ° 12 ’ 12.60 ” N, 76 ° 50 ’ 37.26 ” E]. Paratypes: 1 m, ‘ Southeast Kazakhstan / Alma-Ata Region / Alma-Ata city / 24. vi. 1952 / leg. N. G. Skopin’ [in Cyrillics] [43 ° 12 ’ 12.60 ” N, 76 ° 50 ’ 37.26 ” E]; 1 m, ‘ Southeast Kazakhstan / Alma-Ata Region / Zailiyskiy Alatau / Khargaly canyon / 12. vii. 1969 / leg. K. Nurpeisov’ [in Cyrillics] [43 ° 7 ’ 47.74 ” N, 76 ° 22 ’ 30.08 ” E]; 1 m, ‘ Southeast Kazakhstan / Alma-Ata Region / Alma-Ata city suburbs / 13. vii. 1953 / leg. N. G. Skopin’ [in Cyrillics] [43 ° 12 ’ 12.60 ” N, 76 ° 50 ’ 37.26 ” E]; 1 m, ‘ Southeast Kazakhstan / Alma-Ata Region / Alma-Ata city suburbs / 23. vi. 1957 / leg. N. G. Skopin’ [in Cyrillics] [43 ° 12 ’ 12.60 ” N, 76 ° 50 ’ 37.26 ” E]; 1 m, ‘ Kazakhstan / Alma-Ata / h = 1400 m / 31. v. 1950 / leg. N. G. Skopin’; 1 m, ‘ f, ‘ Kyrgyzstan / Bishkek city / 2. vi. 1976 / leg. Yanushev’ [in Cyrillics]. Material examined (ZIN). 1 m, 6 ff, ‘ Eastern Kazakhstan / Alma-Ata Region / Zailiyskiy Alatau Range / gorge of Uzun-Kargaly River [N 43 ° 4 ’ 40.23 ”, E 76 ° 24 ’ 9.25 ”] / 31.05 – 08. vi. 1907 / leg. A. Jakobson’ [in Cyrillics]. Distribution. Northern, North-East and Central Tian Shan (Kazakhstan, Kyrgyzstan) (SKOPIN 1977). Regional distribution. Southeast Kazakhstan (SKOPIN 1977).	en	Chigray, Ivan, Ivanov, Alexander (2020): A Review Of The Genus Blaps (Coleoptera: Tenebrionidae) Of Central And South Kazakhstan With Description Of Two New Species. Acta Zoologica Academiae Scientiarum Hungaricae 66 (2): 111-162, DOI: 10.17109/AZH.66.2.111.2020, URL: http://dx.doi.org/10.17109/azh.66.2.111.2020
