taxonID	type	description	language	source
894E3E5CD562FF90F9AE929AB762FEE9.taxon	description	(Figs. 1 – 9, 16 – 18, 25 – 27, 34 – 35, 40 – 42, 49 – 50)	en	Roell, Talita, Brugnera, Ricardo, Lemaître, Valérie A. (2021): Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae). Zootaxa 4958 (1): 570-584, DOI: 10.11646/zootaxa.4958.1.33
894E3E5CD562FF90F9AE929AB762FEE9.taxon	description	Arma pallipes Dallas, syntype ♂ at NHMUK, labels: “ Venezuela / 47 26 ”; “ 29. ARMA CRASSIMARGO. ”; “ a ”; “ BRIT. MUS ”; “ Podisus sp. 1 - 198 ”; “ NHMUK 010938912 ”. (Figs 4 – 6) Podisus neniator Breddin, lectotype ♀ at SDEI, labels: “ Balzapamba (Ecuad.) R. Haensch S. ”; “ Podisus neniator Bredd. ”; “ coll. Breddin ”; “ Holo-typus ”; “ DEI EBRESWALDE [sic!] ”; “ DEI Hemimetabola # 100183 ”. (Figs 7 – 9) Non-types examined. COLOMBIA: Cundinamarca, Reserva Chicaque, Refugio, 04.6148667 ° N, 74.3112833 ° W, 2221 m, 24 - 28 - II- 2014, D. Forero col., 2 ♀ 1 ♂ (MPUJ); Choachi, La Victoria, N 4 ° 31, W 73 ° 55, 1927 m, 18 - VI- 2011, J. Pérez col., ♂ (UNAB); Boyacá, Buenavista, Patiño, Las Lomas, N 5 ° 3, W 73 ° 24, 1989 m, 15 - IV- 2004, W. Ávila col., ♀ (UNAB); Risaralda, La Suiza, 1900 m, VIII- 1992, Yaces col., ♀ (MPUJ).	en	Roell, Talita, Brugnera, Ricardo, Lemaître, Valérie A. (2021): Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae). Zootaxa 4958 (1): 570-584, DOI: 10.11646/zootaxa.4958.1.33
894E3E5CD562FF90F9AE929AB762FEE9.taxon	diagnosis	Diagnosis. Pronotum without spots, anterolateral pronotal margins calloused, crenulated, paler than pronotal disc; humeral angles angular, without a posterior tooth, laterally directed. Apex of scutellum concolourous with disc (Figs 1, 4, 7, 16). Apices of parameres obtuse (Figs 25 – 27, 34, par). Posterior margins of valvifers VIII strongly sinuous, forming a V-projection on posterior angles over valvifers IX (Fig. 49, vf 8).	en	Roell, Talita, Brugnera, Ricardo, Lemaître, Valérie A. (2021): Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae). Zootaxa 4958 (1): 570-584, DOI: 10.11646/zootaxa.4958.1.33
894E3E5CD562FF90F9AE929AB762FEE9.taxon	description	Redescription. Body length 9.86 – 11.29 mm (♀) and 10 mm (♂). Body castaneous to brown or rubrous, anterior half of lateral margins of pronotum forming a marginal wide carina, lighter than rest of pronotum. A thin central line mostly without punctures or lighter than rest of dorsum extends from anterior margin of pronotum to posterior margin of scutellum (Figs 1, 4, 7, 16, 18). Head subrectangular, mandibular plates uniformly punctured, clypeus as long as or slightly surpassing apex of mandibular plates; ocelli lying behind an imaginary line through posterior margins of compound eyes; antennal tubercles partially visible from above, proportion of antennomeres: I <II> III <IV> V (Figs 1, 4, 7, 16, 18); bucculae evanescent posteriorly, apex rounded; labium extending to posterior margin of metasternum, proportion of labiomeres: I <II> III> IV (Figs 2, 5, 8, 17). Pronotum hexagonal, uniformly punctured; cicatrices flat, almost indistinguishable; anterior and lateral margins concave in dorsal view, lateral margins slightly crenulated and lighter than disc on anterior half, anterior half of lateral margins preceded by a marginal sulcus; humeral angles angular, slightly produced, without a posterior tooth. Scutellum longer than wide, reaching an imaginary line connecting fifth connexives medially; frenal margins longer than post frenal margins. Coria longer than scutellum, each attaining abdominal segment IV, uniformly punctured; hemelytral membranes well surpassing apex of abdomen, with apex spotted in brown (Figs 1, 4, 7, 16). Pro-, meso- and metasterna covered by small thin setae; prosternum with a weak median carina; mesosternum carinated longitudinally; limit between mesosternum and metasternum tumid; metasternum flat. Pro-, meso-, and metapleura sparsely punctured, evaporatoria on posterior margin of each mesopleura; metapleural evaporatoria surrounding each ostiolar peritreme; each ostiolar peritreme discal-type, elongated, with a darker spot on apical angle; each ostiolar opening directed laterally (Figs 2, 5, 8, 17). Abdomen sparsely punctured, posterolateral angles of each connexivum projected as small spines, longer on seventh segment. Mesial tubercle projecting from sternite III not surpassing metacoxae (Figs 2, 5, 8, 17). Male. Without glandular patches on ventral surface of abdomen. Genitalia. Pygophore bowl-shaped, opened posteriorly, setose, setae denser on ventral rim (Figs 25 – 27); dorsal rim concave with 1 + 1 dorsal projections near segment X (Figs 25 – 27, dr, dp); ventral rim concave, with 1 + 1 ventral projections near posterolateral angles (Fig. 27, vr, vp, pa); surface between inferior and superior layers of ventral rim excavated (Figs 26 – 27, vr). Posterolateral angles acute, slightly surpassing seventh tergite (Fig. 5). Segment X tubular, ventrally directed. Head of parameres divided into two arms, each apex obtuse, well visible posteriorly (Figs 25 – 27, 34, par). Pseudoclasper subquadrate, outer margins sinuous, placed dorsolaterally to parameres, with globose projections and elevations (Figs 25 – 27, 35, pc, om). Phallus. Basal theca and thecal shield subequal in length, thecal shield slightly wider than basal theca (Figs 40 – 42, bt, ts), vesica golf club-shaped in lateral view (Fig. 41, v), apices of conjunctival lobes slightly sculptured (Fig. 40 – 42, cl). Female. Genitalia. Genital plates positioned ventroposteriorly (Fig. 17); valvifers VIII subtriangular, posterior margins setose, strongly sinuous, forming a V-projection on posterior angles over valvifers IX, sutural margins straight, juxtaposed (Fig. 49, vf 8). Laterotergites VIII triangular with spiracles on proximal angle (Fig. 49, la 8). Exposed portion of valvifers IX rectangular, wider than long (Fig. 49, vf 9). Exposed portion of laterotergites IX tongue-like, setose apically (Fig. 49, la 9), surpassing segment X and almost attaining mediotergite VIII. Segment X subquadrate (Fig. 49, X). Inner portion of valvifers IX projected in 1 + 1 straight elongated arms with acute apices (Fig. 50, vf 9). Valvulae IX with 1 + 1 small lunate secondary thickenings (Fig. 50, tva 9). Ring sclerites small, circular, placed posteriorly to thickening of vaginal intima (Fig. 50, rs, tvi); thickening of vaginal intima short, obtuse (Fig. 50, tvi); arcus placed right anteriorly of thickening of vaginal intima (Fig. 50, ar). Vesicular area almost four times length of pars intermedialis plus capsula seminalis (Fig. 50, va, pi, cs). Inner and median ducts of vesicular areas of uniform diameter, median duct slightly enlarged basally (Fig. 50, id, md). Pars intermedialis tubular, shorter and narrower than capsula seminalis; capsula seminalis oval, elongated, smooth (Fig. 50, pi, cs). Proximal and distal annular crests of similar width (Fig. 50, pa, da).	en	Roell, Talita, Brugnera, Ricardo, Lemaître, Valérie A. (2021): Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae). Zootaxa 4958 (1): 570-584, DOI: 10.11646/zootaxa.4958.1.33
894E3E5CD562FF90F9AE929AB762FEE9.taxon	distribution	Distribution: Venezuela, Colombia, Ecuador.	en	Roell, Talita, Brugnera, Ricardo, Lemaître, Valérie A. (2021): Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae). Zootaxa 4958 (1): 570-584, DOI: 10.11646/zootaxa.4958.1.33
894E3E5CD562FF90F9AE929AB762FEE9.taxon	discussion	Remarks: Dallas (1851) described A. ventralis and A. pallipes in the same work. He examined one or more females of A. ventralis and one or more males of A. pallipes. It is unclear how many specimens he had at the time of the description of each species as the lettering system he used is different from the one that had been agreed for the catalogues of zoological specimens in the collection of the British Museum, the one that, for instance, Francis Walker dutifully followed (Kondorosy et al. 2006; Roell et al. in prep.). After a morphological analysis of the typespecimens found by Roell et al. (in prep.), we can confirm our original hypothesis: Dallas’s syntype of A. pallipes is the male of Podisus ventralis. Arma submarginata Walker, 1867, synonymized under P. ventralis by Thomas (1992), was recently synonymized under Podisus distinctus (Stål, 1860) by Brugnera et al. (2020). Breddin (1904) described Podisus neniator from an undetermined number of female specimens (syntype (s )). Gaedike (1971), finding a unique female specimen in the collection of SDEI, labelled and listed it as the holotype. As syntypes from Breddin’s collection are still found scattered in many depositories and even still discovered in SDEI by experts of different hemipteran families (Kondorosy & Rédei 2015; Simões & Campos 2015; Rédei et al. 2016), we have accepted that Gaedike (1971) has designated the female specimen deposited at SDEI as lectotype by inference of holotype (ICZN 1999, Art. 74.6). Should further syntypes be found, they should be considered paralectotypes.	en	Roell, Talita, Brugnera, Ricardo, Lemaître, Valérie A. (2021): Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae). Zootaxa 4958 (1): 570-584, DOI: 10.11646/zootaxa.4958.1.33
894E3E5CD565FF9EF9AE9014B475F985.taxon	description	(Figs 10 – 12, 19 – 21, 28 – 30, 36 – 37, 43 – 45, 51 – 52, 55 – 57)	en	Roell, Talita, Brugnera, Ricardo, Lemaître, Valérie A. (2021): Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae). Zootaxa 4958 (1): 570-584, DOI: 10.11646/zootaxa.4958.1.33
894E3E5CD565FF9EF9AE9014B475F985.taxon	materials_examined	Holotype. Male. Labels: BRASIL, RS, São Domingos do Sul, Santa Gema, 28 ° 33 ’ 32 ’’ S, 51 ° 50 ’ 11 ’’ W. 14 - 16 vii 2017. Pano de batida. J. Avendaño, R. Brugnera & V. Castro col .. (MZSP) (Figs 10 – l 2) Paratypes. BRAZIL: Paraná, 1 ♀, Curitiba, 12 - III-year?, D. H. Habeck, col., (JEE); Santa Catarina, Seara [Nova Teutônia], 1 ♀, 27 ° 11 ’ S, 52 ° 23 ’ L, 06 - XIII- 1948, F. Plaumann col., (AMNH); São Lourenço, 20 mi. W, 28 - II- 1980, D. B. Thomas col., (DBTC); Rio Grande do Sul 10 ♂, 10 ♀, same data as holotype (UFRGS); 3 ♂, 3 ♀ same data as holotype (MZSP); ♂, Maquiné, Fepagro, I- 2006, (MCNZ); PARAGUAY: Alto Paraná, 3 ♀, 21 - 23 - X- 1989, G. Arriagada col., (JEE).	en	Roell, Talita, Brugnera, Ricardo, Lemaître, Valérie A. (2021): Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae). Zootaxa 4958 (1): 570-584, DOI: 10.11646/zootaxa.4958.1.33
894E3E5CD565FF9EF9AE9014B475F985.taxon	etymology	Etymology. We are pleased to dedicate this new species to Dr Jocelia Grazia (UFRGS) in recognition of her outstanding contributions to the knowledge of the Heteroptera, especially to that of Neotropical Pentatomoidea, the passion for which she has, over the years, graciously communicated to her many grateful students.	en	Roell, Talita, Brugnera, Ricardo, Lemaître, Valérie A. (2021): Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae). Zootaxa 4958 (1): 570-584, DOI: 10.11646/zootaxa.4958.1.33
894E3E5CD565FF9EF9AE9014B475F985.taxon	materials_examined	Type locality. Brazil	en	Roell, Talita, Brugnera, Ricardo, Lemaître, Valérie A. (2021): Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae). Zootaxa 4958 (1): 570-584, DOI: 10.11646/zootaxa.4958.1.33
894E3E5CD565FF9EF9AE9014B475F985.taxon	diagnosis	Diagnosis. Anterolateral margins of pronotum calloused, crenulated, paler than disc; humeral angles angular, without a posterior tooth, directed laterally. Apex of scutellum concolourous with disc (Figs 10, 19, 21). Apices of parameres obtuse (Figs 28 – 30, 36). Posterior margins of valvifers VIII slightly sinuous, posterior angles obtuse (Fig. 51).	en	Roell, Talita, Brugnera, Ricardo, Lemaître, Valérie A. (2021): Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae). Zootaxa 4958 (1): 570-584, DOI: 10.11646/zootaxa.4958.1.33
894E3E5CD565FF9EF9AE9014B475F985.taxon	description	Description. Body length 10.57 – 11.07 mm (♀) and 8.00 – 9.70 mm (♂). Body castaneous, anterior half of lateral margins of pronotum forming a marginal wide carina, lighter than rest of pronotum. A thin central line mostly without punctures or lighter than rest of dorsum extends from anterior margin of pronotum to posterior margin of scutellum (Figs 10, 19, 21, 55 – 57). Head subrectangular, mandibular plates uniformly punctured, clypeus as long as or slightly surpassing apex of mandibular plates; ocelli lying behind an imaginary line through posterior margins of compound eyes; antennal tubercles partially visible from above, proportion of antennomeres: I <II> III <IV> V (Figs 10, 19, 21); bucculae evanescent posteriorly, apex rounded; labium extending to posterior margin of metasternum, proportion of labiomeres: I <II> III> IV (Figs 11, 20). Pronotum hexagonal, uniformly punctured; cicatrices flat; anterior and lateral margins concave in dorsal view, lateral margins slightly crenulated and lighter than disc on anterior half, anterior half of lateral margins preceded by a marginal sulcus; humeral angles angular, slightly produced. Scutellum longer than wide, reaching an imaginary line connecting fifth connexives medially; frenal margins longer than post frenal margins. Coria longer than scutellum, each attaining abdominal segment IV, uniformly punctured; hemelytral membranes well surpassing apex of abdomen, with apex spotted in brown (Figs 10, 19, 21). Pro-, meso- and metasterna covered by small thin setae; prosternum with a weak median carina; mesosternum carinated longitudinally; limit between mesosternum and metasternum tumid; metasternum flat. Pro-, meso-, and metapleura sparsely punctured, evaporatoria on posterior margin of each mesopleura; metapleural evaporatoria surrounding each ostiolar peritreme; each ostiolar peritreme discal-type, elongated, with a darker spot on apical angle; each ostiolar opening directed laterally (Figs 11, 20). Abdomen sparsely punctured, posterolateral angles of each connexivum projected as small spines, longer on seventh segment. Mesial tubercle projecting from sternite III not surpassing metacoxae (Figs 11, 20). Male. Without glandular patches on ventral surface of abdomen. Genitalia. Pygophore bowl-shaped, opened posteriorly, setose, setae denser on ventral rim (Figs 28 – 30); dorsal rim concave with 1 + 1 dorsal projections near segment X (Fig 28, dr, dp) ventral rim concave, with 1 + 1 small ventral projections near posterolateral angles (Figs 29, 30, vr, vp, pa); surface between inferior and superior layers of ventral rim excavated (Fig. 29, 30, il, sl). Posterolateral angles acute (Figs 28 – 30, pa). Segment X tubular, ventrally directed. Head of parameres divided into two arms, each apex obtuse, well visible posteriorly (Figs 28 – 30, 36, par). Pseudoclasper C-shaped, outer margins sinuous, placed dorsolaterally to parameres, with globose projections and elevations (Figs 28 – 30, 37, pc). Phallus. Basal theca and thecal shield subequal in length, thecal shield slightly wider than basal theca (Figs 43 – 45), vesica golf club-shaped in lateral view (Fig. 44, v), apices of conjunctival lobes without sculptures (Figs 43 – 45, cl). Female. Genitalia. Genital plates positioned ventroposteriorly (Fig. 20); valvifers VIII subtriangular, posterior margins slightly sinuous, posterior angles obtuse, sutural margins straight, juxtaposed (Fig. 51, vf 8). Laterotergites VIII triangular with spiracles on proximal angle (Fig. 51, la 8). Exposed portion of valvifers IX rectangular, wider than long (Fig. 51, vf 9). Exposed portion of laterotergites IX tongue-like (Fig. 51, la 9), surpassing segment X, and almost attaining mediotergite VIII. Segment X subquadrate (Fig. 51, X). Inner portion of valvifers IX projected in 1 + 1 straight elongated arms with acute apices (Fig. 52, vf 9). Valvulae IX with 1 + 1 small lunate secondary thickenings (Fig. 52, tva 9). Ring sclerites inconspicuous, small, circular, placed posteriorly to thickening of vaginal intima (Fig. 52, rs, tvi); thickening of vaginal intima short, obtuse (Fig. 52, tvi); arcus placed anteriorly of thickening of vaginal intima (Fig. 52, ar). Vesicular area almost 2.5 times longer than length of pars intermedialis plus capsula seminalis (Fig. 52, va, pi, cs). Inner and median ducts of vesicular areas of uniform diameter (Fig. 52, id, md), outer duct constricted basally (Fig. 52, od). Pars intermedialis tubular, shorter and narrower than capsula seminalis; capsula seminalis oval, elongated, smooth (Fig. 52, pi, cs). Proximal and distal annular crests of similar width (Fig. 52, pa, da).	en	Roell, Talita, Brugnera, Ricardo, Lemaître, Valérie A. (2021): Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae). Zootaxa 4958 (1): 570-584, DOI: 10.11646/zootaxa.4958.1.33
894E3E5CD565FF9EF9AE9014B475F985.taxon	distribution	Distribution: Brazil (South), Paraguay.	en	Roell, Talita, Brugnera, Ricardo, Lemaître, Valérie A. (2021): Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae). Zootaxa 4958 (1): 570-584, DOI: 10.11646/zootaxa.4958.1.33
894E3E5CD565FF9EF9AE9014B475F985.taxon	discussion	Remarks. The external morphology of P. graziae resembles that of P. ventralis. Neither species have a paler spot on the post-frenal lobe of the scutellum (Figs 16, 19) and both present parameres with obtuse apices (Figs 34, 36). However, the female external and internal genitalia are very different; in P. ventralis, the posterior margins of valvifers VIII are strongly sinuous, forming a V-projection on the posterior angles over valvifers IX (Fig. 49), while in P. graziae these posterior margins are slightly sinuous, with obtuse posterior angles (Fig. 51); the vesicular area is longer in P. ventralis than in P. graziae (Figs 50, 52), and the outer duct of the vesicular area is constricted only in P. graziae (Fig. 52). Besides that, these two species are allopatric in the Neotropical region; P. ventralis can be found in the northern regions of South America (Venezuela, Colombia, Ecuador), while P. graziae is distributed in the southern regions (Brazil, Paraguay).	en	Roell, Talita, Brugnera, Ricardo, Lemaître, Valérie A. (2021): Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae). Zootaxa 4958 (1): 570-584, DOI: 10.11646/zootaxa.4958.1.33
894E3E5CD56BFF9AF9AE9177B4CAFE21.taxon	description	(Figs 13 – 15, 22 – 24, 31 – 33, 38 – 39, 46 – 48, 53 – 54, 58 – 60)	en	Roell, Talita, Brugnera, Ricardo, Lemaître, Valérie A. (2021): Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae). Zootaxa 4958 (1): 570-584, DOI: 10.11646/zootaxa.4958.1.33
894E3E5CD56BFF9AF9AE9177B4CAFE21.taxon	materials_examined	Holotype. Male. Labels: “ VENEZUELA: Mérida Las Cruces 9 July 1986 R. S. Miller colr. ”; “ B. M. 1991 - 55 ”; “ J. E. Eger Collection ”; “ NEOTYPE ARMA PALLIPES (DALLAS 1851) Desig. D. B. Thomas ”; “ NHMUK 010592332 ” (Figs 13 – 15). (NHMUK) Paratypes. PANAMA: Veraguas, Cerro Tute, 4 Km W Santa Fé, 680 m, 2 - XIII- 1995, C. W. & L. B. O’Brien col., ♀ (JEE); VENEZUELA: Lara, Humocaro Bajo, 10 - V- 1951, F. Fernandes, ♀ (MIZA). Carabobo, Barbula, 17 - IX- 1963, C. Gonzalez col., ♀ (MHNLS). Aragua, Guamitas (probably current La Guamita in the Henri Pittier National Park), 750 m, 24 - VIII- 62, F. Fernandes col., ♀ (MIZA); Rancho Grande (currently Henri Pittier National Park), 29 - VII- 1958, ♀ (MHNLS); 1100 m, 02 - I- 1966, F. Romero col., ♀ (MIZA); 1100 m, 6 - XII- 56, F. Fernandes & C. J. Rosales col., ♀ (MIZA); 1100 m, 12 - III- 1952, J. R. Reguena col., ♂ (MIZA); 1100 m, 25 - IV- 1966, J. & B. Bechine col., ♂ (MIZA); I- 1954, G. u. Helga Frey col., ♀ (EMG); El Limón, 450 m, 25 - V- 1996, F. Fernandes col., ♂ (MIZA); Barinas, El Junco, 1500 m, 9 - I- 1982, R. T. Schuh and B. M. Massie col., ♂ (AMNH). COLOMBIA: Boyacá, Santa Maria, sector La Almenara, 1123 m, 4.87, - 73.25, P. Erazo, C. Lesmes, L. López col., ♂ (MPUJ). Cundinamarca, Quipile, Vereda Arabia, Finca el Paraiso nº 12, 2026 m, 4.74, - 74.56, 15 - IX- 2012, A. Garcia col., ♂ (UNAB). Tolima, Roncesvalles, Vereda El Diamante, 05 - VI- 97, M. A. & D. R. col., ♀ (UNAB). ECUADOR: Imbabura, Paramba, 3500 m, IV- 1997, Rosenberg col., ♂ (NHMUK); Pichincha, Maquipucuna Biol. Reserve, 1350 m, 2 / 3 - VIII- 1998, W. Optiz col., ♂ (JEE).	en	Roell, Talita, Brugnera, Ricardo, Lemaître, Valérie A. (2021): Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae). Zootaxa 4958 (1): 570-584, DOI: 10.11646/zootaxa.4958.1.33
894E3E5CD56BFF9AF9AE9177B4CAFE21.taxon	etymology	Etymology. We are pleased to dedicate this species to Dr Donald B. Thomas (USDA-ARS) in recognition of his remarkable work on the World Asopinae. The publication of his catalogues (1992, 1994) has allowed further research on the group, including this paper in which we thank him for donating to NHMUK the male specimen we are designating above as holotype of this species.	en	Roell, Talita, Brugnera, Ricardo, Lemaître, Valérie A. (2021): Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae). Zootaxa 4958 (1): 570-584, DOI: 10.11646/zootaxa.4958.1.33
894E3E5CD56BFF9AF9AE9177B4CAFE21.taxon	materials_examined	Type locality. Venezuela	en	Roell, Talita, Brugnera, Ricardo, Lemaître, Valérie A. (2021): Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae). Zootaxa 4958 (1): 570-584, DOI: 10.11646/zootaxa.4958.1.33
894E3E5CD56BFF9AF9AE9177B4CAFE21.taxon	diagnosis	Diagnosis. Pronotum without spots, anterolateral pronotal margins not calloused, crenulated, paler than pronotal disc; humeral angles angular elongated, without a posterior tooth, laterally directed. Apex of scutellum paler than disc (Figs 13, 22, 58). Parameres divided into two arms, each apex acute (Fig. 38). Posterior margins of valvifers VIII slightly sinuous, posterior angles obtuse, lateral angles over laterotergites IX (Fig. 53).	en	Roell, Talita, Brugnera, Ricardo, Lemaître, Valérie A. (2021): Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae). Zootaxa 4958 (1): 570-584, DOI: 10.11646/zootaxa.4958.1.33
894E3E5CD56BFF9AF9AE9177B4CAFE21.taxon	description	Description. Body length. 10.30 – 11.08 mm (♀) and 8.50 – 9.40 mm (♂). Body castaneous to brown, anterior half of lateral margins of pronotum lighter than rest of pronotum, humeral angles slightly darker than disc, apex of scutellum pale, lighter than disc. A thin central line mostly without punctures or lighter than rest of dorsum extends from anterior margin of pronotum to posterior margin of scutellum (Figs. 13, 22). Head subrectangular, mandibular plates uniformly punctured, clypeus as long as or slightly surpassing apex of mandibular plates; ocelli lying behind an imaginary line through posterior margins of compound eyes; antennal tubercles partially visible from above, proportion of antennomeres: I <II> III <IV> V (Figs. 13, 22); bucculae evanescent posteriorly, apex rounded; labium extending to posterior margin of metasternum, proportion of labiomeres: I <II> III> IV (Figs. 14, 23). Pronotum hexagonal, uniformly punctured; cicatrices flat; anterior and lateral margins concave in dorsal view, lateral margins crenulated and lighter than disc on anterior half; humeral angles angular, slightly produced, darker than disc. Scutellum longer than wide, reaching an imaginary line connecting fifth connexives medially, apex pale, lighter than disc; frenal margins longer than post frenal margins. Coria longer than scutellum, each attaining abdominal segment IV, uniformly punctured; hemelytral membranes well surpassing apex of abdomen, with apex spotted in brown (Figs 13, 22, 24). Pro-, meso- and metasterna covered by small thin setae; prosternum with a weak median carina; mesosternum carinated longitudinally; metasternum flat. Pro-, meso-, and metapleura densely punctured, evaporatoria on posterior margin of each mesopleura; metapleural evaporatoria surrounding each ostiolar peritreme; each ostiolar peritreme discal-type, elongated, with a darker spot on apical angle; each ostiolar opening directed laterally (Fig. 14). Abdomen sparsely punctured, posterolateral angles of each connexivum projected as small spines, longer on seventh segment. Mesial tubercle projecting from sternite III not surpassing metacoxae (Figs. 14, 23). Male. Without glandular patches on ventral surface of abdomen. Genitalia. Pygophore bowl-shaped, opened posteriorly, setose, setae denser on ventral rim (Figs 31 – 33); dorsal rim concave, with 1 + 1 dorsal projections near segment X (Fig. 31, dr, dp); ventral rim concave, with 1 + 1 inconspicuous ventral projections near posterolateral angles (Figs 32 – 33, vr, vp, pa), surface between inferior and superior layers of ventral rim excavated (Figs 32 – 33, il, sl, vr). Posterolateral angles acute (Figs 31 – 33, pa), surpassing seventh tergite (Fig. 14). Segment X tubular, ventrally directed. Head of parameres divided into two distinct arms, each apex acute, well visible posteriorly (Figs. 31 – 33, 38, par). Pseudoclasper with transversal elevations, placed dorsolaterally to parameres (Figs. 31 – 33, 39, pc). Phallus. Basal theca and thecal shield subequal in length, thecal shield slightly wider than basal theca (Figs. 46 – 48, bt, ts); vesica with two elongated arms, golf club-shaped in lateral view (Fig. 47, v); apices of conjunctival lobes slightly sculptured (Fig. 46 – 48, cl). Female. Genitalia. Genital plates positioned ventroposteriorly (Fig. 23); valvifers VIII oval, posterior margins sinuous, posterior angles obtuse, sutural margins straight, juxtaposed, lateral angles over laterotergites IX (Fig. 53, vf 8). Laterotergites VIII triangular with spiracles on proximal angle (Fig. 53, la 8). Exposed portion of valvifers IX rectangular, wider than long (Fig. 53, vf 9). Exposed portion of laterotergites IX tongue-like (Fig. 53, la 9), surpassing segment X, and almost attaining mediotergite VIII. Segment X subrectangular (Fig. 53, X). Inner portion of valvifers IX projected in 1 + 1 straight elongated arms with acute apices (Fig. 54, vf 9). Valvulae IX with 1 + 1 small circular secondary thickenings. Ring sclerites inconspicuous, small, circular, placed posteriorly to thickening of vaginal intima; thickening of vaginal intima short, obtuse (Fig. 54, tvi); arcus placed right anteriorly of thickening of vaginal intima (Fig. 54, ar). Vesicular area almost three times the length of pars intermedialis plus capsula seminalis (Fig. 54); Inner and median ducts of vesicular areas of uniform diameter (Fig. 54, id, md), outer duct constricted basally (Fig. 54, od). Pars intermedialis tubular, shorter and narrower than capsula seminalis; capsula seminalis oval, elongated, smooth (Fig. 54, pi, cs). Proximal and distal annular crests of similar width (Fig. 54, pa, da).	en	Roell, Talita, Brugnera, Ricardo, Lemaître, Valérie A. (2021): Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae). Zootaxa 4958 (1): 570-584, DOI: 10.11646/zootaxa.4958.1.33
894E3E5CD56BFF9AF9AE9177B4CAFE21.taxon	distribution	Distribution: Panama, Venezuela, Colombia, Ecuador.	en	Roell, Talita, Brugnera, Ricardo, Lemaître, Valérie A. (2021): Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae). Zootaxa 4958 (1): 570-584, DOI: 10.11646/zootaxa.4958.1.33
894E3E5CD56BFF9AF9AE9177B4CAFE21.taxon	discussion	Remarks. Thomas (1992) examined the syntype of A. ventralis but was unable to find any type for A. pallipes and donated to NHMUK a male specimen that he designated as the neotype of this species. While cataloguing typespecimens of Asopinae, Roell et al. (in prep.) found an actual syntype of A. pallipes (Figs 4 – 6), the assessment of that specimen as a type is detailed in their catalogue; Thomas’s neotype is therefore hereby set aside (ICZN 1999: Art. 75.8) and we have described, in this work, a new species for this specimen, based also on the study of several other conspecific specimens. Podisus thomasi sp. n. can be differentiated from Podisus ventralis and Podisus graziae sp. n. by the following characteristics: apex of scutellum yellowish spotted (Figs. 13, 22, 58 – 60) [concolourous with the disc in P. ventralis and P. graziae], apices of parameres acute (Fig. 38) [obtuse in P. ventralis and P. graziae], and pseudoclasper subrectangular (Fig. 38) [suboval in P. ventralis and P. graziae]. P. thomasi is sympatric with P. ventralis. Both are distributed in the northern regions of South America.	en	Roell, Talita, Brugnera, Ricardo, Lemaître, Valérie A. (2021): Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae). Zootaxa 4958 (1): 570-584, DOI: 10.11646/zootaxa.4958.1.33
