taxonID	type	description	language	source
8B6287D79443FFFCFF66F3C6DB75FED3.taxon	description	(Figs 1 – 4, 6 A)	en	Melnitsky, Stanislav I., Ivanov, Vladimir D., Perkovsky, Evgeny E., Legalov, Andrei A. (2025): New Nyctiophylax Brauer, 1865 and Plectrocnemia Stephens, 1836 (Trichoptera: Polycentropodidae) from Eocene Rovno amber. Ecologica Montenegrina 83: 104-111, DOI: 10.37828/em.2025.83.11, URL: https://doi.org/10.37828/em.2025.83.11
8B6287D79443FFFCFF66F3C6DB75FED3.taxon	materials_examined	Type material: Holotype. Male. SIZK CC – 58, Stare Selo, Rovno amber, late Eocene. Syninclusions: SIZK CC – 58 – SIZK CC – 63: 2 Hymenoptera (one Megaspilidae, Lagynodes sp.), 9 Diptera Nematocera (2 Psychodidae, 6 Chironomidae), 2 Acari.	en	Melnitsky, Stanislav I., Ivanov, Vladimir D., Perkovsky, Evgeny E., Legalov, Andrei A. (2025): New Nyctiophylax Brauer, 1865 and Plectrocnemia Stephens, 1836 (Trichoptera: Polycentropodidae) from Eocene Rovno amber. Ecologica Montenegrina 83: 104-111, DOI: 10.37828/em.2025.83.11, URL: https://doi.org/10.37828/em.2025.83.11
8B6287D79443FFFCFF66F3C6DB75FED3.taxon	description	Description. Body length 3.6 mm; forewing length 4.6 mm. Head, antennae, legs, thorax and wings dark brown. Palps and abdomen brown. Wings with long brown hairs. Male genitalia. Ventral part of inferior appendages elongate with rounded and turned upwards apex. Dorsal part of inferior appendages makes clavate process directed upwards and covered with hairs. Segment X large, roof-shaped, rounded in dorsal view. Ventral membranous part of this segment well developed. Ventrolateral horn-shaped processes of segment X directed ventrocaudad and curved inwards, strong and long, longer than segment X. Preanal appendages (cerci) irregularly triangular with rounded apex. Aedeagus wide with two small ventral tubercles at the apex. Comparison. We placed this new species in the genus Nyctiophylax because of absence of the fork 1 on fore wing. Nonetheless, some structures of male genitalia in this specimen are similar to those typical for Plectrocnemia, with bilobed inferior appendages having smaller dorsal part. This composition of peculiar and presumably ancestral shape of the inferior appendages with a set of characters widespread among Nyctiophylax is the most characteristic for the new species. The probable closest species in the genus Nyctiophylax to the new species is N. hamatus Ulmer, 1912 from Baltic amber. These two species differ from each other by the structure of the inferior appendages and by the shape of ventrolateral projections of segment X. Nyctiophylax hamatus has ventrolateral processes of the same length as segment X; these processes curved downwards, not inwards as in the new species.	en	Melnitsky, Stanislav I., Ivanov, Vladimir D., Perkovsky, Evgeny E., Legalov, Andrei A. (2025): New Nyctiophylax Brauer, 1865 and Plectrocnemia Stephens, 1836 (Trichoptera: Polycentropodidae) from Eocene Rovno amber. Ecologica Montenegrina 83: 104-111, DOI: 10.37828/em.2025.83.11, URL: https://doi.org/10.37828/em.2025.83.11
8B6287D79443FFFCFF66F3C6DB75FED3.taxon	etymology	Etymology. The specific name ispokonvekov is indeclinable and must not agree in gender with its generic name; it derives from Russian and means “ since the beginning of time ”.	en	Melnitsky, Stanislav I., Ivanov, Vladimir D., Perkovsky, Evgeny E., Legalov, Andrei A. (2025): New Nyctiophylax Brauer, 1865 and Plectrocnemia Stephens, 1836 (Trichoptera: Polycentropodidae) from Eocene Rovno amber. Ecologica Montenegrina 83: 104-111, DOI: 10.37828/em.2025.83.11, URL: https://doi.org/10.37828/em.2025.83.11
8B6287D79443FFFCFF66F3C6DB75FED3.taxon	distribution	Distribution. Priabonian Rovno amber.	en	Melnitsky, Stanislav I., Ivanov, Vladimir D., Perkovsky, Evgeny E., Legalov, Andrei A. (2025): New Nyctiophylax Brauer, 1865 and Plectrocnemia Stephens, 1836 (Trichoptera: Polycentropodidae) from Eocene Rovno amber. Ecologica Montenegrina 83: 104-111, DOI: 10.37828/em.2025.83.11, URL: https://doi.org/10.37828/em.2025.83.11
8B6287D79443FFFCFF66F3C6DB75FED3.taxon	discussion	Remark. Big size of the piece and syninclused Psychodidae indicate that amber trap was situated on the base of tree trunk (Perkovsky et al. 2012).	en	Melnitsky, Stanislav I., Ivanov, Vladimir D., Perkovsky, Evgeny E., Legalov, Andrei A. (2025): New Nyctiophylax Brauer, 1865 and Plectrocnemia Stephens, 1836 (Trichoptera: Polycentropodidae) from Eocene Rovno amber. Ecologica Montenegrina 83: 104-111, DOI: 10.37828/em.2025.83.11, URL: https://doi.org/10.37828/em.2025.83.11
8B6287D79446FFFFFF66F6A9DCE3FAEF.taxon	description	(Figs 5, 6 B)	en	Melnitsky, Stanislav I., Ivanov, Vladimir D., Perkovsky, Evgeny E., Legalov, Andrei A. (2025): New Nyctiophylax Brauer, 1865 and Plectrocnemia Stephens, 1836 (Trichoptera: Polycentropodidae) from Eocene Rovno amber. Ecologica Montenegrina 83: 104-111, DOI: 10.37828/em.2025.83.11, URL: https://doi.org/10.37828/em.2025.83.11
8B6287D79446FFFFFF66F6A9DCE3FAEF.taxon	materials_examined	Type material: Holotype. Male. SIZK UA – 28504, Rovno amber, late Eocene. Syninclusion: Tipuloidea.	en	Melnitsky, Stanislav I., Ivanov, Vladimir D., Perkovsky, Evgeny E., Legalov, Andrei A. (2025): New Nyctiophylax Brauer, 1865 and Plectrocnemia Stephens, 1836 (Trichoptera: Polycentropodidae) from Eocene Rovno amber. Ecologica Montenegrina 83: 104-111, DOI: 10.37828/em.2025.83.11, URL: https://doi.org/10.37828/em.2025.83.11
8B6287D79446FFFFFF66F6A9DCE3FAEF.taxon	description	Description. Body length 5.7 mm; forewing length 6.8 mm. Head, antennae legs, abdomen and brown thorax, wings light brown. Head with light hairs. Male genitalia. Only ventral aspect of male genitalia is clearly visible. Inferior appendages short, trapezoid with a suture along distal margin delimiting an apical stripe. Aedeagus sclerotized, with apical oval sclerite provided with two pointed terminal lobes. Comparison. The new species has some similarity with Plectrocnemia scapulosa Mey, 1988 from Bitterfeld amber and differs from it the shape of the inferior appendages and apex of aedeagus: in P. scapulosa the inferior appendages looks more elongated in the ventral view, sharply tapering towards of apex, aedeagus gradually tapers towards the apex without subapical expansion.	en	Melnitsky, Stanislav I., Ivanov, Vladimir D., Perkovsky, Evgeny E., Legalov, Andrei A. (2025): New Nyctiophylax Brauer, 1865 and Plectrocnemia Stephens, 1836 (Trichoptera: Polycentropodidae) from Eocene Rovno amber. Ecologica Montenegrina 83: 104-111, DOI: 10.37828/em.2025.83.11, URL: https://doi.org/10.37828/em.2025.83.11
8B6287D79446FFFFFF66F6A9DCE3FAEF.taxon	etymology	Etymology. The specific name stremglav is indeclinable and must not agree in gender with its generic name; it derives from Russian and means “ headlong, reckless ”.	en	Melnitsky, Stanislav I., Ivanov, Vladimir D., Perkovsky, Evgeny E., Legalov, Andrei A. (2025): New Nyctiophylax Brauer, 1865 and Plectrocnemia Stephens, 1836 (Trichoptera: Polycentropodidae) from Eocene Rovno amber. Ecologica Montenegrina 83: 104-111, DOI: 10.37828/em.2025.83.11, URL: https://doi.org/10.37828/em.2025.83.11
8B6287D79446FFFFFF66F6A9DCE3FAEF.taxon	distribution	Distribution. Priabonian Rovno amber.	en	Melnitsky, Stanislav I., Ivanov, Vladimir D., Perkovsky, Evgeny E., Legalov, Andrei A. (2025): New Nyctiophylax Brauer, 1865 and Plectrocnemia Stephens, 1836 (Trichoptera: Polycentropodidae) from Eocene Rovno amber. Ecologica Montenegrina 83: 104-111, DOI: 10.37828/em.2025.83.11, URL: https://doi.org/10.37828/em.2025.83.11
8B6287D79446FFFFFF66F6A9DCE3FAEF.taxon	discussion	Discussion The second European trichopteran fauna studied in details after Baltic amber fauna was Bitterfeld amber fauna. The faunistic survey of the Bitterfeld yet not complete, although the preliminary results evidence the similarity of all European Trichoptera amber deposits (Wichard 2013 a). The new species described here, P. stremglav, has related species in Baltic (P. spinosa, with similar apical structures of aedeagus) and Bitterfeld (P. scapulosa) ambers, with closer similarity to the latter. The taxonomic discussion around the species described by Ulmer (1912) is difficult because a part of his material is lost, and his descriptions are based on syntypes to be revised and re-classified. For example, P. spinosa was described on 3 specimens, illustrated by 5 illustrations of 2 specimens, and these illustrations are markedly different in shape of inferior and preanal appendages. With new species genus Plectrocnemia represented in Rovno amber by nine species with seven endemic to Rovno amber, so now the number of endemic species already higher than in Bitterfeld amber. Reasons for relative rarity of Nyctiophylax in the non-Baltic fossil resins require additional investigations. This genus is abundantly represented by Nyctiophylax varians Ulmer, 1912 in the Baltic amber making 30 % of all specimens of the genus determined to the species level in the monograph of Ulmer (1912), while the same species occur in much lesser numbers, up to 5 specimens, also in the Rovno amber. Nevertheless, this species makes up to 55 % of all Nyctiophylax specimens found in the Rovno amber. The three genera (Plectrocnemia, Holocentropus, Nyctiophylax) comprise a half of all species in the Rovno amber. The genus is still not reported from Bitterfeld amber (Wichard 2013 a). The trichopteran fauna of Rovno amber turned to be better studied than the Bitterfeld fauna. There could be other species in Bitterfeld amber related to the Rovno ones. Until now, only endemic Bitterfeld species with one exception (Wichard 2013 b) were reported, although numerous Baltic amber species found in Bitterfeld amber (Wichard 2013 a) left unrevised. Unfortunately, only the first part of Wichard Eocene revision with new data on Bitterfeld fauna with the first species reported to Baltic and Bitterfeld amber, i. e., Electroadicella bitterfeldi Wichard, 2013 was published (Wichard 2013 b), and dominant Eocene annulipalpian Trichoptera were not included in it. We suppose that at least dominant Eocene species abundant in both Baltic and Rovno amber occurred in Bitterfeld amber as well. Since Bitterfeld amber probably originated on Russoscandia, its place of origin was at least partly in more western and southern regions than Baltic amber (Simutnik et al. 2025 and references therein). Further investigations might clarify the relationships of the European amber faunas. Acknowledgments Authors thanks Alexandr P. Rasnitsyn for Megaspilidae determination.	en	Melnitsky, Stanislav I., Ivanov, Vladimir D., Perkovsky, Evgeny E., Legalov, Andrei A. (2025): New Nyctiophylax Brauer, 1865 and Plectrocnemia Stephens, 1836 (Trichoptera: Polycentropodidae) from Eocene Rovno amber. Ecologica Montenegrina 83: 104-111, DOI: 10.37828/em.2025.83.11, URL: https://doi.org/10.37828/em.2025.83.11
8B6287D79446FFFFFF66F6A9DCE3FAEF.taxon	description	https: // doi. org / 10.37828 / em. 2024.72.14 Ivanov, V. D., Melnitsky, S. I. & Perkovsky, E. E. (2016) Caddisflies from Cenozoic resins of Europe. Paleontological Journal, 50 (5), 485 – 493. https: // doi. org / 10.1134 / S 0031030116050063 Lyubarsky, G. Yu., Legalov, A. A., Vasilenko, D. V. & Perkovsky, E. E. (2024 a) Discovery of Atomaria saxonica (Coleoptera: Cryptophagidae) in Rovno amber and Eocene silken fungus beetles diversity. Ecologica Montenegrina, 79, 133 – 144. https: // dx. doi. org / 10.37828 / em. 2024.79.13 Lyubarsky, G. Yu., Vasilenko, D. V., Legalov, A. A. & Perkovsky, E. E. (2024 b) New silken fungus beetle species (Coleoptera, Cucujiformia, Cryptophagidae) from Rovno amber. Ecologica Montenegrina, 73, 434 – 442. https: // dx. doi. org / 10.37828 / em. 2024.73.29 Lyubarsky, G. Yu., Vasilenko, D. V., Legalov, A. A. & Perkovsky, E. E. (2024 c) Two new silken fungus beetle species (Coleoptera, Cucujiformia, Cryptophagidae) from Rovno amber. Ecologica Montenegrina, 73, 90 – 99. https: // dx. doi. org / 10.37828 / em. 2024.73.9 Melnitsky, S. I. & Ivanov, V. D. (2010) New species of caddisfly (Insecta: Trichoptera) from the Rovno amber, Eocene of Ukraine. Paleontological Journal, 44 (3), 303 – 311. https: // doi. org / 10.1134 / S 003103011003010 X Melnitsky, S. I. & Ivanov, V. D. (2013) Seven new species of caddisflies (Insecta: Trichoptera) from the Rovno amber (Eocene of Ukraine). Paleontological Journal, 47 (3), 283 – 291. https: // doi. org / 10.1134 / S 0031030113030076 Melnitsky, S. I. & Ivanov, V. D. (2016 a) New species of caddisflies (Insecta: Trichoptera) from the Paleogene resins of Europe. Paleontological Journal, 50 (1), 69 – 72. https: // doi. org / 10.1134 / S 003103011601007 X Melnitsky, S. I. & Ivanov, V. D. (2016 b) New species of caddisflies (Insecta: Trichoptera) from the Rovno amber. Zoosymposia, 10, 278 – 291. https: // doi. org / 10.11646 / zoosymposia. 10.1.26 Melnitsky, S. I. & Ivanov, V. D. (2023 a) Plectrocnemia aristovi sp. nov., a new fossil species of Polycentropodidae (Insecta: Trichoptera) from Eocene Rovno amber. Palaeoentomology, 6 (2), 117 – 119. https: // doi. org / 10.11646 / palaeoentomology. 6.2.2. Melnitsky, S. I. & Ivanov, V. D. (2023 b) Plectrocnemia zolotuhini sp. nov., a new fossil species of Polycentropodidae (Insecta: Trichoptera) from Eocene Rovno amber. Euroasian Entomological Journal, 22 (3), 162 – 164. https: // doi. org / 10.15298 / euroasentj. 22.03.08. Melnitsky, S. I., Ivanov, V. D. & Perkovsky, E. E. (2021 a) A new species of Plectrocnemia (Trichoptera: Polycentropodidae) from Rovno amber. Zootaxa, 5006 (1), 106 – 109. https: // doi. org / 10.11646 / zootaxa. 5006.1.14 Melnitsky, S. I., Ivanov, V. D. & Perkovsky, E. E. (2021 b) A new species of the fossil genus Electrotrichia (Insecta: Trichoptera: Hydroptilidae) from Rovno amber (Zhytomyr region, Olevsk amber locality). Palaeoentomology, 4 (5), 421 – 424. https: // doi. org / 10.11646 / palaeoentomology. 4.5.4 Melnitsky, S. I., Ivanov, V. D. & Perkovsky, E. E. (2021 c) A new species of the genus Holocentropus (Trichoptera: Polycentropodidae) from Rovno amber. Zoosystematica Rossica, 30 (2), 298 – 302. https: // doi. org / 10.31610 / zsr / 2021.30.2.298 Melnitsky, S. I., Ivanov, V. D., Perkovsky, E. E. & Vasilenko, D. V. (2024 a) Electroadicella unipetra sp. nov., a new fossil species of Leptoceridae (Insecta: Trichoptera) from Eocene Rovno amber. Ecologica Montenegrina, 72, 11 – 18. https: // doi. org / 10.37828 / em. 2024.72.3 Melnitsky, S. I., Ivanov, V. D., Perkovsky, E. E. & Legalov, A. A. (2024 b) The new species and two new records of Polycentropodidae (Insecta: Trichoptera) from Eocene Rovno amber. Ecologica Montenegrina, 75, 1 – 11. https: // doi. org / 10.37828 / em. 2024.75.1 Melnitsky, S. I., Ivanov, V. D., Legalov, A. A. & Perkovsky, E. E. (2024 c) Palaeohelicopsyche netnetdaida sp. n., a first species of Helicopsychidae (Insecta: Trichoptera) from Eocene Rovno amber. Ecologica Montenegrina, 79, 118 – 124. https: // doi. org / 10.37828 / em. 2024.79.11 Melnitsky, S. I., Ivanov, V. D., Perkovsky, E. E., Vasilenko, D. V. & Legalov, A. A. (2024 d) The new species of Holocentropus, Plectrocnemia and Phryganea (Insecta: Trichoptera) from Eocene Rovno amber. Ecologica Montenegrina, 80, 1 – 12. https: // dx. doi. org / 10.37828 / em. 2024.80.1 Morse, J. C. (2025) Trichoptera World Checklist. Available online at http: // trichopt. app. clemson. edu / [Accessed 04 February 2025.] Perkovsky, E. E. (2017) Rovno amber caddisflies (Insecta, Trichoptera) from different localities, with information about three new sites. Vestnik zoologii, 51 (1), 15 – 22. https: // doi. org / 10.1515 / vzoo- 2017 - 0003 Perkovsky, E. E., Rasnitsyn, A. P., Vlaskin, A. P. & Rasnitsyn, S. P. (2012) Contribution to the study of the structure of amber forest communities based on analysis of syninclusions in the Rovno amber (Late Eocene of Ukraine). Paleontological Journal, 46 (3), 293 – 301. https: // doi. org / 10.1134 / S 0031030112030136 Rasnitsyn, A. P. & Quicke, D. (Eds.) (2002) History of Insects. Kluwer Academic Publishers. Dordrecht, Boston, London, 1 – 518. Simutnik, S. A., Pankowski, M. V. & Perkovsky, E. E. (2025) A new genus of Encyrtidae (Hymenoptera: Chalcidoidea), the first known hymenopteran in amber from Lower Lusatia (Germany). Palaeoentomology, 8 (1), 29 – 35. https: // doi. org / 10.11646 / palaeoentomology. 8.1.5 Ulmer, G. (1912) Die Trichopteren des Baltischen Bernsteins. Beiträge zur Naturkunde Preussens, 10, 1 – 380. Wichard, W. (2013 a) Wasserinsekten im Bitterfelder Bernstein. Exkursionsführer und Veröffentlichungen der Deutschen Gesellschaft für Geowissenschaften, 249, 45 – 53. Wichard, W. (2013 b) Overview and descriptions of Trichoptera in Baltic amber: Spicipalpia and Integripalpia. Remagen: Kessel, 1 – 230. Wichard, W. & Weitschat, W. (1996) Wasserinsekten im Bernstein: Eine Paläobiologische Studie. Entomologische Mitteilungen aus dem Löbbecke-Museum + Aquazoo, 4, 1 – 122.	en	Melnitsky, Stanislav I., Ivanov, Vladimir D., Perkovsky, Evgeny E., Legalov, Andrei A. (2025): New Nyctiophylax Brauer, 1865 and Plectrocnemia Stephens, 1836 (Trichoptera: Polycentropodidae) from Eocene Rovno amber. Ecologica Montenegrina 83: 104-111, DOI: 10.37828/em.2025.83.11, URL: https://doi.org/10.37828/em.2025.83.11
