taxonID	type	description	language	source
871187DDFF85FF83FF15F8F03778F96F.taxon	description	(Fig. 2) SPECIMEN EXAMINED. — Türkiye • Artvin province; Artvin, Irsa Plateau, on tree trunks (Abies nordmanniana (Stev.) Spach. subsp. nordmanniana); 2014 m alt.; 41 ° 51 ’ 54.7 ” N, 41 ° 15 ’ 47.7 ” E; 24. VIII. 2022; leg. N. Batan, H. Erata, N. Batan, H. Erata, KTUB [KTUB 1626]; duplicate in Erata 1215 (Herb. Erata).	en	Nevzat, Hüseyin Erata, Mevlüt, Nevzat Batan, Tülay, Mevlüt Alataş, Ezer, Tülay (2025): Three remarkable moss species new to Türkiye and Southwest Asia. Cryptogamie, Bryologie 20 (6): 51-59, DOI: 10.5252/cryptogamie-bryologie2025v46a6, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a6.pdf
871187DDFF85FF83FF15F8F03778F96F.taxon	biology_ecology	ECOLOGY. — Iwatsukiella leucotricha typically grows on tree trunks and fallen logs of Abies and Picea, and is occasionally on rock faces and in crevices (Dierssen 2001). In Türkiye, the specimens were collected from tree trunks of Abies nordmanniana (Stev.) Spach. Subsp. nordmanniana on the İrsa Plateau, Karçal Mountains, in the Artvin region. DISTRIBUTION. — Iwatsukiella leucotricha is distributed across Europe (notably in the European part of Russia); Asia (including Georgia, China, and Japan); and North America (in Canada, Alaska, and Oregon) (Ignatov et al. 2006; Hodgetts & Lockhart 2020; GBIF 2024: https: // www. gbif. org / species / 2681265). This record demonstrates that the Iwatsukiella leucotricha, previously known from North America and eastern Asia, has extended its distribution further within Asia, marking a new location. The majority of records for Iwatsukiella leucotricha are from Russia. In the European part of Russia, it has been documented in the Republic of Bashkortostan (Bashkiria) and the Perm Territory. In Asiatic Russia, occurrences span across numerous regions, including the Chukotka Autonomous Area, Khanty-Mansi Autonomous Area- Yugra, Republic of Sakha (Yakutia), Magadan Region, Irkutsk Region, Khabarovsk Territory, Kamchatka Territory, Commander Islands, Republic of Altai, Kemerovo Region, Republic of Buryatia, Trans-Baikal Territory, Amur Region, Primorye Territory, and Sakhalin Region (encompassing Sakhalin Island and the Kuril Islands). (Ignatov 1994; Ignatova et al. 1995, 1996, 2013; Ignatov et al. 2001; Czernyadjeva & Potemkin 2003; Zolotov & Baisheva 2003; Bakalın et al. 2009 a; b; Pisarenko 2014; Fedosov et al. 2016; Afonina et al. 2018; Pisarenko & Bakalin 2018; Dudov et al. 2018; Pisarenko et al. 2022). This new record represents the westernmost and southernmost points of Iwatsukiella leucotricha ’ s global distribution. Additionally, this locality establishes it as both the southernmost occurrence of the species worldwide and the closest point to Europe within its distribution range.	en	Nevzat, Hüseyin Erata, Mevlüt, Nevzat Batan, Tülay, Mevlüt Alataş, Ezer, Tülay (2025): Three remarkable moss species new to Türkiye and Southwest Asia. Cryptogamie, Bryologie 20 (6): 51-59, DOI: 10.5252/cryptogamie-bryologie2025v46a6, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a6.pdf
871187DDFF85FF83FF15F8F03778F96F.taxon	description	DESCRIPTION Plants small, slender 0.7 - 1.7 cm high, pale to yellowish-green. Stem creeping, densely pinnately branched. Brach 1 - 4 mm high and densely leaved. Leaves reflexed when moist and imbricate when dry. Leaves 1 - 2 mm tall and 0.2 - 1.3 mm wide, rounded or rounded ovate at base, and long, clear, hairpoint on the tip. Alar cells not clearly differentiated. Median leaf cells oblong or rhomboidal. Costa absent, or short and double. Iwatsukiella leucotricha bears resemblance to Habrodon perpusillus (De Not.) Lindb., yet it can be differentiated by its long, transparent hairpoint at the leaf tip and its undifferentiated alar cells. Iwatsukiella leucotricha is also morphologically close to Heterocladiella dimorpha (Brid.) Ignatov & Fedosov, from which it is distinguished by similar features: a long, clear hairpoint at the leaf tip and undifferentiated alar cells. Additionally, Heterocladiella dimorpha differs by its denticulate leaf margins and papillose cell surfaces, further setting it apart from Iwatsukiella leucotricha.	en	Nevzat, Hüseyin Erata, Mevlüt, Nevzat Batan, Tülay, Mevlüt Alataş, Ezer, Tülay (2025): Three remarkable moss species new to Türkiye and Southwest Asia. Cryptogamie, Bryologie 20 (6): 51-59, DOI: 10.5252/cryptogamie-bryologie2025v46a6, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a6.pdf
871187DDFF85FF83FF15F8F03778F96F.taxon	discussion	REMARKS Iwatsukiella leucotricha was most recently assessed for the IUCN Red List of Threatened Species in 2017, where it is categorized as Least Concern (LC) (Baisheva et al. 2019. This assessment, documented under the Europe region, is accessible in the IUCN Red List of Threatened Species (2019: e. T 87468107 A 87758253), with data available as of March 26, 2024.	en	Nevzat, Hüseyin Erata, Mevlüt, Nevzat Batan, Tülay, Mevlüt Alataş, Ezer, Tülay (2025): Three remarkable moss species new to Türkiye and Southwest Asia. Cryptogamie, Bryologie 20 (6): 51-59, DOI: 10.5252/cryptogamie-bryologie2025v46a6, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a6.pdf
871187DDFF84FF83FE81F8D33096F9F0.taxon	biology_ecology	ECOLOGY. — Andreaea alpestris typically grows on periodically wet rocks and lithosols often near snow beds. Also grows on rocks in subalpine and alpine habitats (Dierssen 2001; Smith 2004; Frey et al. 2006).	en	Nevzat, Hüseyin Erata, Mevlüt, Nevzat Batan, Tülay, Mevlüt Alataş, Ezer, Tülay (2025): Three remarkable moss species new to Türkiye and Southwest Asia. Cryptogamie, Bryologie 20 (6): 51-59, DOI: 10.5252/cryptogamie-bryologie2025v46a6, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a6.pdf
871187DDFF84FF83FE81F8D33096F9F0.taxon	distribution	DISTRIBUTION. — Andreaea alpestris has a broad distribution, found in Europe (Finland, Norway, Sweden, Great Britain, Andorra, Corsica, France, Italy, Madeira, Spain, Austria, Poland, Switzerland, Bulgaria, North Macedonia, Romania, Caucasus (in Europe); Artict Russia (NW Russia); Ukranie; Asia (China); America (Subartic America, EW America, Canada, Guatemala, Honduras, Costa Rica); (Murray 1988; Allen 1994; Smith 2004; Hodgetts & Lockhart 2020).	en	Nevzat, Hüseyin Erata, Mevlüt, Nevzat Batan, Tülay, Mevlüt Alataş, Ezer, Tülay (2025): Three remarkable moss species new to Türkiye and Southwest Asia. Cryptogamie, Bryologie 20 (6): 51-59, DOI: 10.5252/cryptogamie-bryologie2025v46a6, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a6.pdf
871187DDFF84FF83FE81F8D33096F9F0.taxon	description	DESCRIPTION Plants small, 1.0 - 3.4 cm high, reddish black, black or greenish brown cushions. Shoots 1.4 - 1.8 cm high. Leaves 0.4 - 0.8 mm long, usually straight, imbricate, lanceolate, widest above the base. Leaf margins entire, plane or slightly incurved. Costa absent; basal cells short rectangular. Upper cells of leaf almost quadrate. Papillae on the dorsal surface of the leaves low or absent. Perichaetial leaves differentiated, sheathing, convolute and 1.1 - 1 mm long and 0.5 - 0.6 mm wide (Murray 1988; Smith 2004; Frey et al. 2006). This species closely resembles Andreaea rupestris Hedw., but it differs by having straight leaves and low or absent papillae on the dorsal leaf surface. Additionally, A. alpestris is distinct from A. rupestris due to its smaller, imbricate, obtuse leaves and the non-porose nature of the upper leaf cells. Andreaea alpestris also bears similarities to Andreaea mutabilis Hook. f. & Wilson; however, it can be differentiated by its typically straight leaves and the low or absent papillae on the leaf surface (Murray 1988; Smith 2004; Frey et al. 2006; Zander 2007).	en	Nevzat, Hüseyin Erata, Mevlüt, Nevzat Batan, Tülay, Mevlüt Alataş, Ezer, Tülay (2025): Three remarkable moss species new to Türkiye and Southwest Asia. Cryptogamie, Bryologie 20 (6): 51-59, DOI: 10.5252/cryptogamie-bryologie2025v46a6, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a6.pdf
871187DDFF84FF83FE81F8D33096F9F0.taxon	discussion	REMARKS The species was last assessed for the IUCN Red List in 2017 and is currently classified as Data Deficient (DD) (Hallingbäck et al. 2019; IUCN Red List of Threatened Species, 2019: e. T 83659776 A 87832177, accessed on July 8, 2024).	en	Nevzat, Hüseyin Erata, Mevlüt, Nevzat Batan, Tülay, Mevlüt Alataş, Ezer, Tülay (2025): Three remarkable moss species new to Türkiye and Southwest Asia. Cryptogamie, Bryologie 20 (6): 51-59, DOI: 10.5252/cryptogamie-bryologie2025v46a6, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a6.pdf
871187DDFF84FF8CFC51F898347BF9CE.taxon	description	(Fig. 4) SPECIMEN EXAMINED. — Türkiye • Artvin province, Borçka district, Balcıköy Plateau, power plant surroundings on rock; 2095 - 2150 m alt.; 41 ° 56 ’ 36.2 ” N, 41 ° 20 ’ 35.7 ” E; 22. VI. 2022 • Borçka district, Beyazsu Plateau, on rock; 2356 m alt.; 41 ° 57 ’ 36.1 ” N, 41 ° 21 ’ 18.4 ” E; 23. VI. 2022; leg. N. Batan, H. Erata, det N. Batan, H. Erata, KTUB [KTUB 1628]; duplicate in Erata 1217 (Herb. Erata).	en	Nevzat, Hüseyin Erata, Mevlüt, Nevzat Batan, Tülay, Mevlüt Alataş, Ezer, Tülay (2025): Three remarkable moss species new to Türkiye and Southwest Asia. Cryptogamie, Bryologie 20 (6): 51-59, DOI: 10.5252/cryptogamie-bryologie2025v46a6, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a6.pdf
871187DDFF84FF8CFC51F898347BF9CE.taxon	biology_ecology	ECOLOGY. — Andreaea mutabilis typically grows on dry to wet, exposed acidic rocks, It is also found on periodically wet acid boulders crags or rubble, rarely on thin soil over rocks (Dierssen 2001; Smith 2004; Frey et al. 2006).	en	Nevzat, Hüseyin Erata, Mevlüt, Nevzat Batan, Tülay, Mevlüt Alataş, Ezer, Tülay (2025): Three remarkable moss species new to Türkiye and Southwest Asia. Cryptogamie, Bryologie 20 (6): 51-59, DOI: 10.5252/cryptogamie-bryologie2025v46a6, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a6.pdf
871187DDFF84FF8CFC51F898347BF9CE.taxon	distribution	DISTRIBUTION. — Andreaea mutabilis is distributed across various regions, including Europe (Fareo Islands, Great Britain, Andorra, France, Spain); Asia (Taiwan, China, and Malaysia); the Americas (North and South America); Oceania (New Guinea, Australia, Tasmania, and New Zealand). (Chiang 1998; Gao & Crosby 1999; Smith 2004; Hodgetts & Lockhart 2020).	en	Nevzat, Hüseyin Erata, Mevlüt, Nevzat Batan, Tülay, Mevlüt Alataş, Ezer, Tülay (2025): Three remarkable moss species new to Türkiye and Southwest Asia. Cryptogamie, Bryologie 20 (6): 51-59, DOI: 10.5252/cryptogamie-bryologie2025v46a6, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a6.pdf
871187DDFF84FF8CFC51F898347BF9CE.taxon	description	DESCRIPTION Plants small, 1.0 - 1.2 cm high, reddish to black cushions. Shoots 0.4 - 1 cm high. Leaves 0.4 - 0.7 mm long, erect-spreading and straight, lanceolate, widest above the base, acuminate to acute apex. Costa absent. Leaf margins entire, incurved. Basal laminal cells rectangular, basal marginal cells isodiametric, median laminal cells rounded-quadrate or rhomboid. Perichaetial leaves oblong-lingulate, acuminate (Murray 1988; Smith 2004; Frey et al. 2006). Andreaea mutabilis closely resembles Andreaea sinuosa B. M. Murray but can be distinguished by its quadrate basal laminal cells and non-sinuose basal cells. Additionally, A. mutabilis differs from Andreaea rupestris Hedw. by having isodiametric basal marginal cells, less pitted basal cell walls, and quadrate basal laminal cells, whereas A. rupestris has rectangular basal laminal cells and laminal papillae. Andreaea mutabilis also bears similarity to Andreaea alpestris, but differs in its minimally sheathing leaf bases, often displaying a patch of yellow cells at the base, with basal cells that are scarcely pitted. Its basal laminal cells are rectangular, and marginal cells are quadrate, measuring 0.8 - 1 mm in length and 0.3 - 0.4 mm in width (Chiang 1998; Murray 1988; Smith 2004; Frey et al. 2006; Zander 2007).	en	Nevzat, Hüseyin Erata, Mevlüt, Nevzat Batan, Tülay, Mevlüt Alataş, Ezer, Tülay (2025): Three remarkable moss species new to Türkiye and Southwest Asia. Cryptogamie, Bryologie 20 (6): 51-59, DOI: 10.5252/cryptogamie-bryologie2025v46a6, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a6.pdf
871187DDFF84FF8CFC51F898347BF9CE.taxon	discussion	REMARKS Andreaea mutabilis was last assessed for The IUCN Red List of Threatened Species in 2016, where it is classified as Least Concern (LC) (Hodgetts et al. 2019; The IUCN Red List of Threatened Species 2019: e. T 83659827 A 87792052 accessed on 11 July 2024).	en	Nevzat, Hüseyin Erata, Mevlüt, Nevzat Batan, Tülay, Mevlüt Alataş, Ezer, Tülay (2025): Three remarkable moss species new to Türkiye and Southwest Asia. Cryptogamie, Bryologie 20 (6): 51-59, DOI: 10.5252/cryptogamie-bryologie2025v46a6, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2025v46a6.pdf
