taxonID	type	description	language	source
7F3D87954A1CFFAEFF5AFF5C6D7F92E5.taxon	description	Description. Body elongate, gourd-shaped, medium to large sized (ca. 12.0 – 27.0 mm long). Head hypognathus, narrower than pronotum, and widest across middle of eyes; without visible membrane between labrum and clypeus; clypeus transverse, nearly hexagonal, with straight to emarginate anterior margin, usually with transverse clypeal impression (Fig. 1 A; cti) before frontoclypeal suture, frontoclypeal suture well developed, broadly U-shaped; genae strongly raised, depressed before eyes, roundly produced anterolaterally; eyes transversely reniform, strongly protruding; inner ocular sulci weakly depressed to shallowly grooved; frons broad, smooth or faintly wrinkled, sometimes with paired frontal impressions (Fig. 1 A; fi), paired vertexal lateral impressions (Fig. 1 A; vli) and vertexal median impression (Fig. 1 A; vmi). Labrum (Fig. 1 C) transverse, elongate oval, anterior margin emarginate, bearing serried setae; Mandibles (Figs. 1 D, E) truncate, apex bidentate in most species, prosthecae (Fig. 1 D, E; pst) broad, membranous. Maxilla (Fig. 1 G) with galea (Fig. 1 G; gal) long and straight, with long and dense bristles; lacinia (Fig. 1 G; lac) straight, well-developed, clearly separated from mediostipes (Fig. 1 G; mst), with dense long bristles along inner margin, basistipes (Fig. 1 G; bst) sometimes depressed in middle. Gula with posterior tentorial pits marked (Fig. 16 F; ptp). Labium (Fig. 1 F) with submentum fused with hypostoma, lateral suture of submentum faintly presented; mentum (Fig. 1 F; mn) with medial surface with several large pores bearing long setae, gradually rising anteriorly, depressed along both sides; prementum (Fig. 1 F; pmn) strongly emarginate; glossa (Fig. 1 F; gl) flabellate, emarginate, with a pair of lobes densely pubescent anteriorly; labial palpi short; with palpomere II and palpomere III (Fig. 1 F; lp 2, lp 3) each dilated distally. Antennae slender, antennomeres elongate, and gradually thickening distally. Antennomere XI oblong-oval or fusiform, antennomeres V – XI pubescent, last five antennomeres with stellate sensoriae. Pronotum with marginal borders around entire outline, anterior marginal border occasionally obscured in middle; posterior portions of lateral marginal borders obscured in dorsal view; pronotal disc convex, usually with a pair of impressions in middle. Scutellum (Fig. 2 D), triangular or semicircular. Metanotum (Fig. 2 F; mtn) strongly shortened. Elytra fused, oval to fusiform, convex above; humeri straight, neither angulate nor swelled; disc with nine grooved or furrowed striae (atavus - and metallicu s-group); or with striae interrupted, adjacent segments of striae oppositely curved, connected, forming rows of encircled areas (aenescens - and elegantulu s-group); or with irregularly scattered strial punctures (most species of jendeki - group), or with irregularly distributed tubercles (chongli - and jendeki - group); right elytron overlapping the left one at apex; epipleura oblique inwards, gradually narrowing apically. Hind wings absent. Prosternum laterally fused with hypomera, pronotosternal sutures fine, prosternal process (Fig. 2 A; psp) dilated in ventral view, produced posteriorly, more or less declivous terminally. Mesoventrite with anterior ridges weakly and obtusely angulate anteriorly, anterior part with distinct shiny carina, sparsely pubescent. Metaventrite narrow, finely depressed along the midline, forming a suture; anterior metaventral process (Fig. 2 E; amtp) strongly protruded anteriorly, feebly convex and depressed at base, forming a wide V-shaped suture. Abdomen more or less depressed in male, flattened or weekly convex in female. Intercoxal process of sternite III broad, quadrate, anterior margin weakly convex, wrinkled along anterior margin; sternites IV – VI convex in middle, and depressed in each side; sternites VI and VII usually sulcate along both sides, sternite VII with a pair of shallow impressions in middle; in some species, male sternites III and IV depressed in posterior middle, whose central part weakly convex; membranes visible between last three sternites. Defensive reservoirs elongate, transverse rings indistinct (Fig. 2 I). Legs with femora and tibiae long and slender. Procoxae (Fig. 2 C) projecting below prosternum. Trochanters glabrous, without long setae. Profemora ventrally possessing one or two small impressions near each base; inner margins of tibiae pubescent along apical portions, in male pubescence denser and longer, forming ‘ hair brush’, outer margin of pro- and mesotibiae often depressed near apices. Male pro- and mesotibiae more curved than those in female. Male pro- and mesotarsi slightly widened, all tarsomeres slanted. Aedeagus, strongly sclerotized, curved in lateral view, parameres constricted before dilated apex, apex ovate to flabellate. Median lobe (Fig. 2 J – K; ml), strongly sclerotized, hamular, erected when mating. Sternite VIII strongly sclerotized and emarginated in male, with a pair of knife shaped or hooked apical lobes (Fig. 17 Ld, Ll; al) projecting at both sides of the apical notch. Female genitalia with internal female reproductive tract consisting of slender vagina, opening into enlarged bursa copulatix (Fig. 2 O, bc), oviduct (Fig. 2 O, od) produced from vagina before bursa, slender spermathecal accessory gland (Fig. 2 O, sag) attached at apex of bursa. Ovipositor (Fig. 2 O, ov) strongly sclerotized, blade-like, coxites fused, curved strongly dorsad, gonostyles strongly reduced, extremely short. Spiculum ventrale (Fig. 2 N) with short trunk, weakly sclerotized. Sexual dimorphism. Male: body slender, less markedly constricted between pronotum and elytra, less convex dorsally; abdomen depressed, sometimes with impressions on posterior middle of sternites III and IV; legs more elongate, tibiae more curved, modified in some species, with protrusions on inner margins of pro- and mesotibiae; pro- and mesotarsi more widened. Female: body wider or stouter, more markedly constricted between pronotum and elytra, more convex dorsally; abdomen flat, without impressions on posterior middle of sternites III and IV, legs less elongate in most species, tibiae less curved, without modifications; pro- and mesotarsi less widened.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A1CFFAEFF5AFF5C6D7F92E5.taxon	distribution	Distribution (Map 1). China, Myanmar, Thailand, Vietnam.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A1CFFAEFF5AFF5C6D7F92E5.taxon	discussion	Comments. Inclusion of Morphostenophanes in the tribe Cnodalonini is supported by the structure of defensive reservoirs and presence of stellate sensoriae on antennomeres VII – XI. According to Bouchard & Yeates (2001, as Coelometopini), ovipositors are especially useful in determining close relationship within Cnodalonini. In the Oriental fauna, Ainu Lewis, 1894 (ovipositor shown in Yuan et al. 2018, Fig. 4 L, M), Lycidioides Ando, 2003 (ovipositor shown in Masumoto & Akita 2007, Fig. 12), Pseudonautes Fairmaire, 1893, Scotaeus Hope, 1836 and Thesilea Haag-Rutenberg, 1878 (Masumoto, 1981) have similar strongly sclerotized, blade like ovipositors. However, these genera are all winged and with general appearances being very different from Morphostenophanes. The Palaearctic apterous genus Stenophanes Solsky, 1876 shares the similar elongate habitus and unclavate femora with Morphostenophanes, however, it has weakly sclerotized ovipositor with segmented coxites (according to the observation of Stenophanes mesostena Solsky, 1871 specimens in CZDY). In fact, Hegemona Laporte, 1840 and Saziches Champion, 1886 both from Central America seems really closest related to Morphostenophanes, they share similar appearance, fused elytra, absent hindwings, similar internal female reproductive tracts and sclerotized ovipositor with fused gonocoxites (Doyen 1987), but they distinctly differ from Morphostenophanes in having non overlapping elytral apex, and different attachment point of spermathecal accessory gland. However, as the cnodalonine fauna is far from being clearly understood, it is too early to clarify the actual relationship between Morphostenophanes and other members within this tribe. Morphostenophanes can be readily distinguished from other Oriental cnodalonine genera by the following characters: body medium-sized to large, elongate, gourd-shaped; clypeolabral membrane invisible; antennae slen- der, apical segments unclubbed, with antennomeres III – XI each elongate; pronotum with complete lateral marginal borders, without an indentation at each side before base; elytra fused, with right elytron overlapping the left one at apex. Hind wings absent. Procoxae each well projecting below prosternum. Femora not clavate. Male pro- and mesotarsi slightly widened. Parameres fused, slender, with dilated apex. Sternite VIII strongly sclerotized, with a pair of apical lobes projecting at both sides of the apical notch. Spermathecal accessory gland attached at apex of bursa. Ovipositor with gonocoxites fused, modified as a stout, sclerotized, laterally compressed, blade-like structure. After cleaning several fresh specimens of Morphostenophanes linglong Zhou, new species and M. furvus Zhou, new species in warm water, a coagulated secretion was found at the apex of antennomere XI (Fig. 1 B). In a female specimen of Stenophanes mesostena the author also found the similar phenomenon. The antennal apices of these specimens were also observed under a 15 x magnification, but no glands could be seen. The author speculates that the secretion could be some kind of pheromone related to mating. Electron microscopy is needed for a further study of their antennal surface, observation of more living individuals are needed to answer the function of the secretion, and further detailed study of all cnodalonine genera is needed to answer whether this phenomenon is common within the tribe. In the present work, six species groups of the genus Morphostenophanes are proposed according to different morphological characters and geographical distribution.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A10FFAAFF5AFF196CF890A4.taxon	description	(Figs. 3 A – K; 4 A – D; 16 A, F; 17 O, T; 18 A, L – N; 19 P; 20 A, L)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A10FFAAFF5AFF196CF890A4.taxon	materials_examined	Type material examined. Lectotype designated here: Yunnan [handwriting on pale label] // V. subparallelus Pic [handwriting on pale yellow label] // coll Pic [handwriting on pale yellow label] // Morphostenophanes aenescens Pic det. dr. Kaszab [handwriting of the first two lines on pale label] // MUSÉUM PARIS 1958 coll. M. PIC // SYN- TYPE [red label] // SYNTYPE Morphostenophanes aenescens var. subparallelus Pic, 1925 // MNHN EC 9694 // LECTOTYPE, Morphostenophanes aenescens Pic, 1925, ♂, des. De-Yao Zhou 2019 [printed on white label] (male, MNHN, Fig. 3 A – C). Paralectotype labelled: Yunnan [handwriting on pale label] // Morphostenophanes aenescens Pic [handwriting on pale label] // MUSÉUM PARIS 1958 coll. M. PIC // SYNTYPE [red label] // SYNTYPE Morphostenophanes aenescens Pic, 1925 // MNHN EC 9693 // PARALECTOTYPE, Morphostenophanes aenescens Pic, 1925, ♂, des. De-Yao Zhou 2019 [printed on white label] (1 female, MNHN, Fig. 3 G, H). Both types examined through ten photographs taken by Antoine Mantilleri.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A10FFAAFF5AFF196CF890A4.taxon	discussion	Comments on type material. This species was originally described from an unspecified number of specimens of Pic’s collection, which is now deposited in MNHN. However, characteristic descriptions of both sexes and two varieties, i. e. Morphostenophanes aenescens var. diversus, and M. aenescens var. subparallelus, were presented in the original description (Pic, 1925); therefore at least four specimens including both sexes were examined by Pic. It is difficult to determine the exact number of the specimens in the type series because of the mess of Pic’s collection. Kaszab (1941) placed both varieties as junior synonyms of Morphostenophanes aenescens. Here the author examined one female syntype of M. aenescens, and one male syntype of M. aenescens var. subparallelus with a handwritten identification label of Kaszab. Additional material examined (5 ♂♂, 2 ♀♀). CHINA: Yunnan: 1 ♂ (CZDY, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C), Qujin City, Xishan District, Mount Cui, 25 ° 32 ’ 57.96 ’’ N, 103 ° 42 ’ 04.92 ’’ E, ca. 2000 m, 2015. ix. 20, Ye-Hui Fang; 1 ♂ (CZDY, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C), Qujin City, Zhanyi District, Zhujiangyuan, ca. 2100 m, 2018. viii, Zheng Zhou; 1 ♀ (CJQY), Qujin City, Zhanyi District, Longxijian, 25 ° 54 ’ 18 ’’ N, 103 ° 55 ’ 52.18 ’’ E, 2222 m, 2019. vii. 30, Jia-Mian Liang; 1 ♂ (CZRZ, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C in CZDY), Qujin City, Fuyuan County, Fa’ao Village, Hejiafangzi, 2018. xii, native collector; 2 ♂♂ (ERPC), 1 ♀ (CZDY), Mianmianshan, 20 km W Lugu Lake, 20 ° 42 ’ N, 100 ° 34 ’ E, 2005. viii, V. Siniaev.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A10FFAAFF5AFF196CF890A4.taxon	diagnosis	Diagnosis. Medium-sized, moderately elongate and convex, color dark grayish blue, somewhat purplish. Elytra with densely and irregularly scattered round or oval segments of encircled striae, each encircled area moderately convex, areas around them with pinkish luster (Fig. 3 E, J). Head and pronotum densely scattered large and deep punctures, some connected with each other. Elytra widest in apical 2 / 5. Protibiae nearly straight.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A10FFAAFF5AFF196CF890A4.taxon	description	Redescription. Male (Fig. 4 A, C). Dark greyish blue, antennae, mouthparts, and claws dark brown, areas around encircled segments of elytral striae pinkish, all legs and abdomen purplish, shagreened. Body moderately elongate, length 18.6 – 20.4 mm, width 6.7 – 7.4 mm, strongly convex, noticeably constricted between pronotum and elytra. Head (Fig. 16 A, F) quadrate, finely microsculptured, with densely scattered large and deep punctures, some connected with each other; genal areas before eyes impunctate; with outer margin distinctly notched between genae and clypeus; clypeus transversely hexagonal, gently bent downwards in front, anterior margin weakly emarginate, frontoclypeal suture deeply grooved, widely U-shaped; genae strongly raised, depressed before eyes, strongly and roundly produced anterolaterally; frons broad, anterior part gradually sloping forwards, vertexal lateral impressions and vertexal median impression vaguely presented; eyes transversely reniform, strongly convex laterally; inner ocular sulci shallowly grooved along inner margins, becoming broader posteriorly; tempora moderately convex, coarsely punctate. OI = 58.5 – 60.9. Antennae (Fig. 18 A) slender, reaching basal fifth of elytra, with antennomeres weakly thickened to apices; relative lengths of antennomeres: 0.60: 0.26: 0.90: 0.82: 0.87: 0.85: 0.83: 0.82: 0.80: 0.73: 0.87. Mentum (Fig. 16 F) transversely quadrate, lateral margins nearly straight; medial surface coarsely punctate, with several large pores with long setae, gradually rising anteriorly, depressed along both sides. Pronotum (Fig. 17 O) barrel-shaped, PW / PL = 1.14 – 1.17, widest slightly anterior to midpoint, anterior margin bisinuate, anterior marginal border marked; lateral margins weakly rounded, more or less constricted in posterior sixth, lateral marginal borders thin, visible in dorsal view slightly exceeding anterior half, or between anterior fourth and the half, somewhat sinuate; posterior margin nearly straight, more or less emarginate, posterior marginal border marked; anterior angles rounded, more or less bent ventrally; posterior angles obtuse; disc strongly convex, shagreened, with densely scattered large and deep punctures, some connected with each other, presenting a longitude impression along midline, with a pair of vague impressions on both sides. Scutellum triangular, coarsely punctate. Elytra elongate oval, widest in apical 2 / 5, EL / EW = 1.80 – 1.90; strongly convex, highest near middle; with densely and irregularly scattered round or oval segments of striae, each encircled area moderately convex; intervals slightly wrinkled, finely punctate. Prosternum (Fig. 17 T) shagreened, finely and sparsely punctate; prosternal process declivous, truncate at apex; hypomeron strongly rugulose, shagreened. Metasternum weakly wrinkled. Abdomen (Fig. 19 P) depressed, surface somewhat rough and weakly wrinkled, densely and markedly punctate; sternites III and IV each depressed in posterior middle, with central portion of each impressions weakly convex. Legs slender. Protibiae (Fig. 18 L) nearly straight, slightly recurved behind apical half, apical 3 / 5 of inner margins emarginated and pubescent; mesotibiae (Fig. 18 M) nearly straight, weakly curved in apical seventh, apical half of inner margins weakly pubescent; metatibiae (Fig. 18 N) nearly straight, apical 3 / 5 of inner margins weakly pubescent, outer margins depressed before apices. Aedeagus (Fig. 20 A) elongate, curved in lateral view; parameres slender, 0.23 as long as total length, recurved in middle in lateral view, with ovoid apex, slightly bent ventrally in lateral view. Sternite VIII (Fig. 20 P) with apical lobes broadly hooked in lateral view. Female (Fig. 4 B, D). Stouter than male, length 21.5 mm. IE / TD = 58.6, PW / PL = 1.21; elytra more convex, highest behind middle, EL / EW = 1.74; abdomen straight in lateral view; without impressions on sternites. Ovipositor shortened, abruptly narrowing terminally from apical third. Variability. Only one male from Xishan District, Qujin city has the pronotum distinctly constricted before the base (as shown in Fig. 17 O-XsD), as shown in the lectotype. Species from Fuyuan County, Qujin City have pronotal anterior angles distinctely bent ventrally, lateral margin markedly projecting laterally at widest point, lateral marginal borders slightly sinuate (as shown in Fig. 17 O-FyC). Comparative notes. Morphostenophanes aenescens most resembles M. yunnanus. It can be readily distinguished from M. yunnanus by its greyish blue body color (grey-black in the latter), head and pronotum with large and deep punctures (those in the latter are much smaller and shallower), stouter elytra than the latter, and nearly straight male protibiae (curved at apical third in the latter).	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A10FFAAFF5AFF196CF890A4.taxon	discussion	Comments. Morphostenophanes aenescens is the type species of Morphostenophanes. Pic (1925) gave a very brief description of this species, with an imprecise locality information of ‘ Yunnan’. According to the original description, M. aenescens is shining and glossy, black or pitch-black, dorsum bronze (‘ Nitidus, glaber, nigro aut piceo metallicus, supra aeneus’). Within the description, Pic described two varieties, i. e. M. aenescens var. diversus, and M. aenescens var. subparallelus. The former is somewhat greenish (‘ aliquot viridescens’), and the latter has the habitus more parallel, impressions on elytra more coarsely punctate (‘ presente une forme plus étroite, presque parallele et les elytres ont quelques gros points nets dans leurs impressions’). Later, Kaszab (1941) synonymized these two variants. In his key, Kaszab described this species as black, with very weak metallic luster (‘ Schwarz, mit sehr schwachem Metallschimmer’). The identification label hand-written by Kaszab (shown in Fig. 3 F) also attests that he did examine the type series. In Kaszab’s work (1941), only one indistinct photo of M. aenescens was displayed. Subsequent works (Masumoto & Bečvář 2008; Gao & Ren 2009) also presented photos of male M. aenescens, but they are different from each other. Though the newly designated lectotype and paralectotype were seriously damaged, some important features were visible. Their heads and pronota are covered with large deep punctures, some connected with each other; prothorax of male constricted before base; and elytra possess rows of small encircled segments of striae, each with pinkish luster. The author found three males and two female specimens possessing these characteristics, thus allowing for a better characterization and more accurate location of this species. The author also found many similar specimens from Dali Prefecture and neighboring areas, distinguished from M. aenescens by darker body color, more glossy head and pronotum, strongly curved protibiae and sinuous metatibiae (nearly straight in M. aenescens), elytral impressions without pinkish luster. Thus these specimens are identified as new and name as M. yunnanus. Based on the revised diagnostic characters and known locality of M. aenescens, specimens examined by Masumoto & Bečvář (2008) and the record reported by Gao & Ren (2009) from Hetaoping, Binchuan County are considered as misidentifications of M. yunnanus; another pair of specimens cited by Gao & Ren (2009) from Longxin, Longling County may belongs to M. gaoligongensis, which belongs to the elegantulus - group. Pinkish luster on elytral impressions is a common character in M. cuproviridis, sometimes observed in M. aenescens yelang, and vaguely shown in M. papillatus, and on some individuals of M. iridescens. These species together with M. aenescens are successively distributed from east Yunnan to east Guizhou (Map 1). This character is also faintly present on the smallest individuals of M. yunnanus, which occurrs in midwestern Yunnan. Therefore, such character may reveals closer relations of these species within aenescens - group, and shows a gradual spreading and speciation scene from west to east.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A10FFAAFF5AFF196CF890A4.taxon	distribution	Distribution. (Map 1) CHINA: Yunnan.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A15FFABFF5AFABC6F2694B5.taxon	description	(Figs. 5 A – D; 16 M, Q; 17 L, Q; 18 H; 19 G – I, W; 20 H, S)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A15FFABFF5AFABC6F2694B5.taxon	materials_examined	Type materials. CHINA: Yunnan: ♂ (Holotype, SNUC), Qujing City, Shizong County, 1865 m, 2018. vii. 30, Ke Tang. Paratypes (2 ♀♀): CHINA: Yunnan: 1 ♀ (SNUC, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C in CZDY), Qujing City, Shizong County, 1865 m, 2018. vii. 30, Ke Tang; 1 ♀ (RBINS), Kouang Si Hien [Guangxi County = Luxi County], alt. 2100 m, Steihmetz. Diagnostic description. Male (Fig. 5 A, C). Color lighter, greyish-green, with coppery luster, pronotum faintly presented an annular greenish area. Body elongate, length 20.9 mm, width 7.4 mm, less convex. Head (Fig. 16 M, Q) with OI of 56.7. Antennae (Fig. 18 H) more slender, relative lengths of antennomeres: 0.60: 0.31: 1.02: 0.92: 0.94: 0.99: 1.04: 1.00: 0.97: 0.87: 1.09. Pronotum (Fig. 17 L) barrel-shaped, PW / PL = 1.12, disc moderately convex, shagreened, with densely scattered large and deep punctures, with a pair of vague impressions on both sides. Elytra more elongate and flattened, widest near middle, EL / EW = 1.89. Aedeagus (Fig. 20 H) more elongate, 0.29 as long as the total length, apicale of parameres wider. Female (Fig. 5 B, D). Stouter than male, length 19.2 mm. OI = 56.9; PW / PL = 1.28; disc of elytra more flattend, dorsum nearly straight, gradually rising posteriorly, highest in apical third; EL / EW = 1.58. Comparative notes. This subspecies can be distinguished from the nominate subspecies mainly by its lighter and somewhat purplish body color (dark greyish blue in the latter), distinctly more slender antenna and more elongate legs, sometimes observed faint annular greenish area on pronotum (absent in the latter), and less convex elytra.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A15FFABFF5AFABC6F2694B5.taxon	distribution	Distribution. (Map 1). CHINA: Yunnan.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A15FFABFF5AFABC6F2694B5.taxon	etymology	Etymology. The new subspecies is named from ‘ Yelang’, an ancient country located around the type locality in the Han dynasty.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A14FFA8FF5AFA626CEC9775.taxon	description	(Figs. 6 A – D; 16 B, G; 17 A, F; 18 B, O – Q; 19 Q; 20 B, M)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A14FFA8FF5AFA626CEC9775.taxon	materials_examined	Material examined. (8 ♂♂, 6 ♀♀). CHINA: Guizhou: 1 ♀ (CZDY, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C), Leishan County, Mount Leigong, 1610 m, 2018. vii. 9, Tian-Yong Yang; 2 ♂♂ (CZDY), Leishan County, Mount Leigong, 2015. vii. 12, Bo-Yan Li; 2 ♀♀ (CZDY), Leishan County, Mount Leigong, light trap, 2018. v. 25, Bo-Yan Li; 1 ♂ (MYNU), Leishan County, Mount Leigong, 2016. viii. 12, collector unknown; 1 ♂ (CZDY), Leishan County, Lianhuaping, 1572 m, 26 ° 22 ’ 25.18 ’’ N, 108 ° 11 ’ 55.72 ’’ E, 2017. vii. 22, Min-Zhi Zhao; 1 ♂ (MYNU), Leishan County, Mount Leigong, Lianhuaping, 1000 m, Hao Xu & Jian-Yue Qiu; 1 ♂, 2 ♀♀ (MYNU), Leishan County, Fangxiang Country, Queniao Village, 1600 m, 2013. vii. 28, Hao Xu & Jian-Yue Qiu; 1 ♂ (CQL), Leishan County, Mount Leigong, 1600 m, 2017. vii. 30. Zi-Hao Yang; 1 ♂ (CZDY, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C), Guiyang City, Lindai Village, 26 ° 39 ’ 36.55 ’’ N, 106 ° 22 ’ 20.88 ’’ E, 2018. viii, Min-Zhi Zhao; 1 ♀ (GFGY), Kaiyang County, 1978. xii. 9, Wang.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A14FFA8FF5AFA626CEC9775.taxon	description	Description of female. Female (Fig. 6 B, D). Stouter than male, length 17.8 – 19.3 mm. Distance between eyes longer than male, OI = 53.1 – 56.0. PW / PL = 1.14 – 1.21. Elytra more convex, highest near middle, EL / EW = 1.59 – 1.69; abdomen straight in lateral view; without impressions on sternites. Ovipositor shortened, abruptly narrowing terminally from apical third. Variability. Females from the type locality, Leigong Mountain, have shortened elytra with aspect ratio ranging from 1.59 – 1.62, but the only female from Kaiyang County City shows longer elytra with aspect ratio of 1.69. Comparative notes. Morphostenophanes cuproviridis was described in details by Gao & Ren (2009). It is the most brightly-coloured species of the genus, with green shining body, slightly bronze; pronotum colored pink, with an annular green band; area along elytral suture, areas of encircled segments of striae and epipleura, middle parts of pro-, meso- and metafemora, both sides of abdomen with pinkish luster. Morphostenophanes cuproviridis is similar to M. iridescens, which has almost the same bright-coloured body as M. cuproviridis. The two species seems to be closely related, but M. cuproviridis can be clearly distinguished from the new species by more shiny body surface, male pronotum distinctly constricted before the base, less convex elytra in male, more distinct pinkish luster on elytral impressions, less widened male protarsi, different aedeagus, and shorter female elytra (at least for those from the type locality). M. cuproviridis also resembles M. aenescens yelang by sharing extremely similar male habitus, but can be distinguished from the latter by different body color, genae more strongly protruding, much longer antennae, less coarsely punctate pronotum and head, much curved male protibiae (almost straight in the latter), less rounded sternite VII in male, and shorter female elytra.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A14FFA8FF5AFA626CEC9775.taxon	distribution	Distribution. (Map 1) CHINA: Guizhou.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A16FFB6FF5AFF196E8595C4.taxon	description	(Figs. 2 O; 7 A – D; 16 C, H, V; 17 B, G; 18 C, R – T; 19 R; 20 C, N)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A16FFB6FF5AFF196E8595C4.taxon	materials_examined	Type materials. CHINA: Guangxi: ♂ (Holotype, SNUC), Mount Mao’er, 2000 m, 2012. vii. 22, Wen-Xuan Bi. Paratypes (5 ♀♀): CHINA: Guangxi: 1 ♀ (SNUC), 1 ♀ (CZBS, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C in CZDY), Mount Mao’er, 2000 m, 2018. viii, Bi-Shen Zhan; 1 ♀ (CBWX), Mount Mao’er, 1577 m, 2019. vii. 8, Y. - Q. Lu; 2 ♀♀ (CBWX), Guilin City, Lingui District, Huangsha Country, 1350 m, 2019. vi. 4, Y. - Q. Lu.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A16FFB6FF5AFF196E8595C4.taxon	diagnosis	Diagnosis. Medium-sized, slender, moderately convex, bronze and slightly greenish, with metallic luster. Elytra widest slightly behind middle, with densely and irregularly scattered round or oval segments of striae, each encircled area moderately convex. Male pro- and mesotibiae evenly curved, metatibiae straight.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A16FFB6FF5AFF196E8595C4.taxon	description	Description. Male (Fig. 7 A, C). Greenish, bearing bronze luster, antennae, mouth parts, and tarsi dark brown. Body elongate, length 18.1 mm, width 6.1 mm, moderately convex, noticeably constricted between pronotum and elytra. Head (Fig. 16 C, H) quadrate, densely scattered with large and shallow punctures; with outer margin deeply notched between genae and clypeus; clypeus transversely heptagonal, gently bent downwards in front, anterior margin rounded, nearly straight in middle, clypeal transverse impression vague, frontoclypeal suture deeply furrowed, widely U-shaped; genae strongly raised, depressed before eyes, roundly produced anterolaterally; frons broad, anterior part gradually sloping forwards, vertexal lateral impressions triangular, each furrowed at deepest point; eyes transversely reniform, strongly convex laterally; inner ocular sulci finely depressed along inner margins; tempora weakly convex, coarsely punctate. OI = 54.0. Antennae (Fig. 18 C) slender, reaching basal fifth of elytra, with antennomeres weakly thickened to apices; relative lengths of antennomeres: 0.51: 0.26: 0.82: 0.82: 0.82: 0.89: 0.89: 0.85: 0.85: 0.85: 0.99. Mentum (Fig. 16 H) quadrate, lateral margins nearly straight; medial surface sparsely and finely punctate, with several large pores with long setae, gradually rising anteriorly, depressed along both sides. Pronotum (Fig. 17 B) quadrate, PW / PL = 1.16, widest in anterior third, anterior margin emarginate, anterior marginal border strongly marked; lateral margins nearly straight, lateral marginal borders thin, visible in dorsal view along anterior half; posterior margin feebly rounded, emarginate, posterior marginal border marked; anterior angles rounded; posterior angles obtuse; disc strongly convex, with a pair of shallow transverse impressions near basal third, shagreened, densely scattered with markedly punctaures. Scutellum triangular, glossy. Elytra oblong, widest slightly behind middle, EL / EW = 1.98; moderately convex, highest slightly before middle; with densely and irregularly scattered round or oval segments of encircled striae, each encircled area moderately convex; intervals sparsely and finely punctate. Prosternum (Fig. 17 G) shagreened, finely and sparsely punctate; prosternal process straight produced posteriorly; hypomeron rugulose, shagreened. Metasternum weakly wrinkled. Abdomen (Fig. 19 R) depressed, surface smooth, somewhat wrinkled along anterior margin and sides of each sternite, densely and markedly punctate; sternites III and IV each feebly depressed in posterior middle, with central portion of each impressions weakly convex. Legs slender. Protibiae (Fig. 18 R) evenly curved, apical 3 / 5 of inner margins pubescent; mesotibiae (Fig. 18 S) evenly curved, apical 3 / 5 of inner margins pubescent; metatibiae (Fig. 18 T) straight, apical 3 / 5 of inner margins weakly pubescent, outer margins depressed in apical fifth. Aedeagus (Fig. 20 C) elongate, weakly curved in lateral view; parameres slender, 0.22 as long as total length, with flabellate apex. Sternite VIII (Fig. 20 N) with apical lobes each produced posteriorly, superior margin bent to exterior margin, roundly bent to inferior margin. Female (Fig. 7 B, D). Stouter than male, length 20.9 – 25.2 mm. OI = 48.4 – 56.3, PW / PL = 1.15; elytra more convex and widened, EL / EW = 1.63, dorsum only weakly rounded, highest in middle, elytral impressions lightly pinkish; abdomen straight in lateral view; without impressions on sternites. Ovipositor (Fig. 2 O) shortened, abruptly narrowing terminally from apical third. Variability. The holotype male and one female have mesosternum with posterior margins of lateral portions roundly protruding (Fig. 16 V), while the rest of female paratypes have straight one as in any other species (Fig. 16 U). Comparative notes. Morphostenophanes curvitibialis resembles M. luoxiaoshanus, but can be distinguished by its slightly more slender body, quadrate pronotal outline (barrel-shaped in the latter), evenly curved male pro- and mesotibiae (weakly curved at apical fourth of the latter), and pinkish luster of elytral encircled areas faintly presented, and more distinct in female (completely missing in the latter).	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A16FFB6FF5AFF196E8595C4.taxon	discussion	Comments. Type series of this species were all from the northern end of Yuechengling Mountains, which run through north Guangxi and south Hunan. Therefore, this species may be found in Hunan in the future.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A16FFB6FF5AFF196E8595C4.taxon	distribution	Distribution. (Map 1) CHINA: Guangxi.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A16FFB6FF5AFF196E8595C4.taxon	etymology	Etymology. The new specific epithet refers to the evenly curved pro- and mesotibiae of the male of the new species.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A09FFB7FF5AF91F6F0993D8.taxon	description	(Figs. 8 A – D; 16 D, I; 17 C, H; 18 D, U – W; 19 S; 20 D, O)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A09FFB7FF5AF91F6F0993D8.taxon	materials_examined	Type materials. CHINA: Yunnan: ♂ (Holotype, SNUC), Wenshan Prefecture, Maguan County, Bazhai Town, 1700 – 1800 m, 2018. xi. 27, native collector. Paratypes (10 ♂♂, 9 ♀♀): CHINA: Yunnan: 1 ♀ (SNUC), 2 ♂♂, 2 ♀♀ (MHBU), 3 ♂♂, 3 ♀♀ (CZDY, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C), 5 ♂♂, 3 ♀♀ (CZMZ), Wenshan Prefecture, Maguan County, Bazhai Town, 1700 – 1800 m, 2018. xi. 27, native collector.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A09FFB7FF5AF91F6F0993D8.taxon	diagnosis	Diagnosis: Large, elongate, moderately convex, greenish, pronotum with annular pinkish band; anterior of clypeus and posterior of head, elytral epipleura and intervals, middle parts of pro-, meso- and metafemora, middle part of abdomen with pinkish luster. Male elytra widest in apical 2 / 5, dorsum flattened, with densely and irregularly scattered round or oval segments of striae, each encircled area moderately convex. Male protarsi strongly widened, protibiae with dorsal side of inner margin more concave than ventral one.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A09FFB7FF5AF91F6F0993D8.taxon	description	Description: Male (Fig. 8 A, C). Greenish, with bronze luster, antennae, mouthparts, and tarsi dark brown, pronotum with annular pinkish band; anterior of clypeus and posterior of head, elytral epipleura and intervals, middle parts of pro-, meso- and metafemora, middle part of abdomen with pinkish luster; elytral encircled areas purplish, or pinkish in few small individuals. Body elongate, length 21.9 – 26.0 mm, width 7.6 – 8.5 mm, moderately convex dorsad, noticeably constricted between pronotum and elytra. Head (Fig. 16 D, I) transversely quadrate, densely and markedly punctate; with outer margin weakly notched between genae and clypeus; clypeus transversely hexagonal, gently bent downwards in front, anterior margin weakly emarginate, clypeal transverse impression marked to absent, frontoclypeal suture deeply furrowed, widely U-shaped; genae strongly raised, depressed before eyes, roundly produced anterolaterally; frons broad, anterior part gradually sloping forwards, frontal impressions each transversely marked, somewhat shallowly extending backwards, connecting shallow vertexal lateral impression, vertexal median impression feebly presented; eyes transversely reniform, strongly convex laterally; inner ocular sulci grooved along inner margins; tempora weakly convex, coarsely punctate. OI = 48.2 – 50.5. Antennae (Fig. 18 D) slender, reaching basal fourth of elytra, with antennomeres weakly thickened to apices; relative lengths of antennomeres: 0.77: 0.37: 1.33: 1.19: 1.22: 1.22: 1.19: 1.14: 1.11: 1.04: 1.11. Mentum (Fig. 16 I) quadrate, lateral margins straight; medial surface sparsely and coarsely punctate, with several large pores with long setae, gradually rising anteriorly, depressed along both sides. Pronotum (Fig. 17 C) barrel-shaped, PW / PL = 1.08 – 1.12, widest in anterior third, anterior margin nearly straight, projecting in middle, anterior marginal border marked; lateral margins weakly rounded, slightly more widened at widest point, lateral marginal borders thin, visible in dorsal view slightly before the middle; posterior margin nearly straight, weakly emarginate, posterior marginal border marked; anterior and posterior angles obtuse; disc strongly convex, a pair of impressions vague to absent in middle, shagreened, densely and finely punctate. Scutellum triangular, glossy. Elytra oblong, widest in apical 2 / 5, EL / EW = 1.96 – 1.97; moderately convex, dorsum nearly straight, with densely and irregularly scattered round or oval segments of striae, each encircled area moderately convex; intervals sparsely and finely punctate. Prosternum (Fig. 17 H) shagreened, sparsely and finely punctate; prosternal process declivous, truncate at apex; hypomeron strongly rugulose, shagreened. Metasternum weakly wrinkled. Abdomen (Fig. 19 S) depressed, surface somewhat rough, sternite III notably wrinkled, densely and finely punctate; sternites III and IV each shallowly depressed in posterior middle, with central portion of each impressions weakly convex. Legs slender. Protarsi strongly widened. Protibiae (Fig. 18 U) curved in apical third, apical 3 / 5 of inner margins emarginated, each with dorsal side of inner margin more concave than ventral one, pubescent; mesotibiae (Fig. 18 V) curved in apical third, apical 3 / 5 of inner margins pubescent; metatibiae (Fig. 18 W) sinuous, apical 3 / 5 of inner margins feebly pubescent, outer margins depressed before apices. Aedeagus (Fig. 20 D) elongate, weakly curved in lateral view; parameres slender, 0.22 as long as total length, with widely rhomboid apex. Sternite VIII (Fig. 20 O) with apical lobes nearly rounded, each inferior margin projecting, forming a hook. Female (Fig. 8 B, D). Stouter than male, length 21.0 – 23.7 mm. OI = 53.5, PW / PL = 1.15 – 1.19; elytra more widened and convex, EL / EW = 1.72 – 1.74, abdomen straight in lateral view; without impressions on sternites. Ovipositor shortened, abruptly narrowing terminally from apical third. Comparative notes. Morphostenophanes iridescens resembles M. cuproviridis. They share similar bright-colored body and similar male protibiae. M. iridescens can be easily distinguished from M. cuproviridis by its strongly convex pronotum colored green, with a pinkish annular band, lateral margin of elytra not constricted before base, moderately convex elytral widest in apical 2 / 5, elytral encircled areas purplish in most individuals, strongly widened male protarsi; and more elongate female elytra with the aspect ratio greater than 1.7, compared to that of the latter less than 1.7.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A09FFB7FF5AF91F6F0993D8.taxon	distribution	Distribution. (Map 1) CHINA: Yunnan.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A09FFB7FF5AF91F6F0993D8.taxon	etymology	Etymology. The new species is named from the Latin epithet ‘ iridescens’ (colored as rainbow) referring to its bright-colored body.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A0BFFB5FF5AFA216B06933D.taxon	description	(Figs. 9 A – D; 16 E, J, U; 17 D, I; 18 E, X – Z; 19 T; 20 E, P)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A0BFFB5FF5AFA216B06933D.taxon	materials_examined	Type materials. CHINA: Hunan: ♂ (Holotype, SNUC), Yueyang City, Pingjiang County, Mount Fushou, 28 ° 28 ’ N, 113 ° 46 ’ E, 800 – 1079 m, 2016. ix. 10 – 17, Ri-Xin Jiang & De-Yao Zhou. Paratypes (81 ♂♂, 30 ♀♀): CHINA: Hunan: 1 ♀ (SNUC), 61 ♂♂, 15 ♀♀ (CZDY, one male was decomposed, a male and a female were preserved in 99.7 % etha- nol at – 18 ° C), 5 ♂♂, 5 ♀♀ (MHBU), 2 ♂♂, 2 ♀♀ (CJRX), 1 ♂, 1 ♀ (CZX), Yueyang City, Pingjiang County, Mount Fushou, 28 ° 28 ’ N, 113 ° 46 ’ E, 800 – 1079 m, 2016. ix. 10 – 17, Ri-Xin Jiang & De-Yao Zhou; 1 ♂, (CJRX) Liuyang City, Mount Dawei, 2016. vi. 4, Ri-Xin Jiang; 1 ♂ (CZJZ) Liuyang City, Mount Dawei, 1300 m, 2019. v. 23, Jia-Zhi Zhang; 1 ♀ (MYNU), Liuyang City, Mount Dawei, 1400 m, 2018. viii. 8 – 9, Hao Xu & Jian-Yue Qiu; 1 ♂, 1 ♀ (CZDY), Pingji- ang County, Mount Mufushan, 1300 m, 2016. viii, Jiang Zhu leg.; 1 ♀ (MYNU), Pingjiang County, Mount Mufu, 2013. late vii, Cong-Chao Dai leg; 2 ♂♂, 2 ♀♀ (CZDY, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C), Yanlin Country, Taoyuandong Nature Reserve, Nanfengmian, 26 ° 17 ’ 59.29 ’’ N, 113 ° 59 ’ 21.85 ’’ E, 1665 m, 2016. vi., De-Yao Zhou & Yi-Xiao Liu. Hubei: 5 ♂♂ (CZDY), Xianning City, Tongshan County, Mount Jiugong, 29 ° 24 ’ 13 ’’ N, 114 ° 40 ’ 10 ’’ E, 1200 – 1250 m, 30. vii. 2016, Yi-Xiao Liu; 1 ♂, 1 ♀ (MYNU), Mount Jiugong, 2010. viii, Wei-Cheng Huang. Jiangxi: 1 ♂ (CZDY), Mount Jinggang, Huangyangjie, 26 ° 37 ’ 21.82 ’’ N, 114 ° 7 ’ 2.11 ’’ E, 1253 m, 2017. vii. 23, Kang Shuang.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A0BFFB5FF5AFA216B06933D.taxon	diagnosis	Diagnosis. Medium sized, elongate, strongly convex species, color slightly coppery greenish, with metallic luster. Elytra widest slightly behind middle, with densely and irregularly scattered round or oval segments of striae, each encircled area moderately convex. Male pro- and mesotibiae curved in apical fourth, metatibiae straight.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A0BFFB5FF5AFA216B06933D.taxon	description	Description. Male (Fig. 9 A, C). Color copper, slightly greenish, with metallic luster, all legs dark green, somewhat purplish, antennae and mouthparts dark brown. Body elongate, length 17.5 – 22.5 mm, width 6.2 – 7.6 mm, moderately convex, distinctly constricted between pronotum and elytra. Head (Fig. 16 E, J) quadrate, finely microsculptured, densely and coarsely punctate; with outer margin deeply notched between genae and clypeus; clypeus transversely hexagonal, gently bent downwards in front, anterior margin nearly straight, emarginate in middle, clypeal transverse impression marked to absent, frontoclypeal suture grooved, widely U-shaped; genae strongly raised, depressed before eyes, roundly produced anterolaterally; frons broad, anterior part gradually sloping forwards, frontal impressions and vertexal lateral impressions marked, vertexal median impression vague; eyes transversely reniform, strongly convex laterally; inner ocular sulci grooved along inner margins; tempora weakly convex, coarsely punctate. OI = 47.9 – 51.7. Antennae (Fig. 18 E) slender, reaching basal 2 / 7 of elytra, with antennomeres weakly thickened to apices; relative lengths of antennomeres: 0.70: 0.27: 1.19: 1.14: 1.19: 1.22: 1.16: 1.12: 1.05: 0.94: 1.07. Mentum (Fig. 8 J) inversely trapezoidal, lateral margins roundly curved; medial surface finely punctate, somewhat rough, with several large pores with long setae, gradually rising anteriorly, depressed along both sides. Pronotum (Fig. 17 D) barrel-shaped, PW / PL = 1.03 – 1.07, widest near the middle, anterior margin bisinuate, anterior marginal border marked; lateral margins rounded, lateral marginal borders thin, visible in dorsal view along anterior half; posterior margin nearly straight, emarginate, posterior marginal border marked; anterior angles rounded, slightly projecting anteriorly; posterior angles obtuse; disc strongly convex, a pair of rounded impressions vague to absent in middle, shagreened, with densely scattered marked punctures. Scutellum triangular, smooth. Elytra oblong, widest slightly behind middle, EL / EW = 1.76 – 1.97; strongly convex, highest near middle; with densely and irregularly scattered round or oval segments of striae, each encircled area moderately convex; intervals glossy, sparsely and finely punctate. Prosternum (Fig. 17 I) shagreened, finely and sparsely punctate; prosternal process weakly declivous, apex truncate; hypomeron rugulose, shagreened. Metasternum weakly wrinkled. Abdomen (Fig. 19 T) depressed, surface smooth, somewhat wrinkled along anterior margin and sides of each sternite, densely and markedly punctate; sternites III and IV each depressed in posterior middle, with central portion of each impressions weakly convex. Legs slender. Protibiae (Fig. 18 X) curved in apical fourth, apical 3 / 5 of inner margins pubescent; mesotibiae (Fig. 18 Y) curved in apical fourth, apical 3 / 5 of inner margins pubescent; metatibiae (Fig. 18 Z) straight, apical 3 / 5 of inner margins weakly pubescent, outer margins depressed in apical fifth. Aedeagus (Fig. 20 E) elongate, weakly curved in lateral view; parameres slender, 0.22 as long as total length, with flabellate apex. Sternite VIII (Fig. 20 P) with apical lobes each produced posteriorly, superior margin bent to exterior margin, roundly bent to the inferior margin. Female (Fig. 9 B, D). Stouter than male, length 16.9 – 23.8 mm. Space between eyes wider than on male, OI = 57.3 – 59.0; PW / PL = 1.07 – 1.10; elytra extremely convex, dorsum slightly rounded, highest in middle, EL / EW = 1.56 – 1.64; abdomen straight in lateral view; without impressions on sternites. Ovipositor shortened, abruptly narrowing terminally from apical third. Comparative notes. Morphostenophanes luoxiaoshanus resembles M. curvitibialis, and can be distinguished by having pronotal lateral margins roundly curved (almost straight in the latter), male pro- and mesotibiae curved in apical fourth (evenly curved in the latter), and posterior margins of lateral portions of mesosternum straight (rounded in the latter).	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A0BFFB5FF5AFA216B06933D.taxon	discussion	Comments. Morphostenophanes luoxiaoshanus is the easternmost known species. All specimens of the type series are from Luoxiaoshan Mountains, a north-south mountain range located along the boundary between Hunan and Jiangxi, which consists of a series of mountains over 1000 meters, and separated by lower altitudes. However, there is little difference between the populations from each mountain.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A0BFFB5FF5AFA216B06933D.taxon	distribution	Distribution. (Map 1) CHINA: Hunan, Hubei, Jiangxi.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A0BFFB5FF5AFA216B06933D.taxon	etymology	Etymology. The new species is named after the Luoxiaoshan Mountains, where the type series came from.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A0DFFB3FF5AFAEE6E83929B.taxon	description	(Figs. 10 A – D; 16 K, O; 17 E, J; 18 F; 19 A – C, U; 20 F, Q)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A0DFFB3FF5AFAEE6E83929B.taxon	materials_examined	Material examined (7 ♂♂, 9 ♀♀). CHINA: Chongqing: 1 ♀ (CZDY), Mount Simian Nature Reserve, 28 ° 35 ’ 13.87 ” N, 106 ° 22 ’ 15.79 ” E, 1033 m, 2018. viii. 15, De-Yao Zhou; 1 ♀ (CQL), Simianshan Nature Reserve, Feilong Temple, 2017. vii. 7 – 11, Yi Zhang; 1 ♀ (CBNX), Simianshan Nature Reserve, 1000 m, 2017. v. 1, Qiao-Qiao Liu; 1 ♀ (MYNU), Simianshan Nature Reserve, 2016. vi. 3, Hong-Mei Lou; 1 ♀ (MYNU), Simianshan Nature Re- serve, Feilong Temple, 1000 m, 2014. iv. 26, Hao Xu; 1 ♀ (MYNU), Simianshan Nature Reserve, Dawopu, 2013. vii. 24, Hao Xu & Jian-Yue Qiu; 1 ♂ (CBWX), Simianshan Nature Reserve, Dawopu, 2008. ix. 8, Xiao-Dong Yang; 2 ♂♂, 1 ♀ (MYNU), Simianshan Nature Reserve, Dawopu, 2014. vii. 28, Hao Xu & Jian-Yue Qiu. Guizhou: 1 ♂ (CZDY), Xishui Nature Reserve, Sanchahe, 890 m, pupa in rotten word 2015. vii. 12, adult merged 2015. viii, De- Yao Zhou; 1 ♀ (CLRY), Zunyi City, Mount Jinding, 1100 m, without date, Run-Yu Li; 1 ♂, 1 ♀ (MYNU), Zunyi City, Dabanshui forest park, 2100 m, Run-Yu Li. Sichuan: 1 ♂ (MYNU), Luzhou City, Gulin County, Honglonghu Park, 2012. viii. 18, Lu Qiu. Yunnan: 1 ♂ (CZDY), Shaotong City, Weixing County, back hill near Ba’erlin, ca. 1350 m, 2018. ix, native collector. Comparative notes. Morphostenophanes papillatus is distinguished from congeners by the rows of strongly inflated tubercles on the elytra. It is also distinguishable by its extremely convex body and nearly straight pro- and mesotibiae. Alhough M. papillatus has exaggerated elytral tubercles different from other congeners, other character states (cephalic impressions, elytra widest behind middle, with encircled and merely interrupted segments of elytral striae, impressions on sternites III and IV) undoubtedly place it within the aenescens - group. Similar bronze body color suggest possible close relationship to M. luoxiaoshanus and M. curvitibialis.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A0DFFB3FF5AFAEE6E83929B.taxon	discussion	Comments. Until now, all known specimens of this species were from Dalou Mountain, except one male examined by Gao & Ren (2009), which is from Fanjing Mountain, a part of Wuling Mountains. This specimen has less convex elytral tubercles, and narrower pronotum, which is different from those from Dalou Mountains. Therefore it is considered here as an uncertain identification. More specimens from Wuling Mountains are required for a reliable identification.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A0DFFB3FF5AFAEE6E83929B.taxon	distribution	Distribution. (Map 1) CHINA: Chongqing, Guizhou, Sichuan (new province record), Yunnan (new province record).	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A0CFFB1FF5AF8A56F119314.taxon	description	(Figs. 11 A – D; 17 K, P; 18 G; 19 D – F, V; 20 G, R, V)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A0CFFB1FF5AF8A56F119314.taxon	materials_examined	Type materials. CHINA: Yunnan: ♂ (Holotype, SNUC), Pingbian County, Mount Dawei, near the hotel on the peak, ca. 2093 m, 2018. v. 26, Zhi-Wei Dong & Lu Qiu. Paratypes (8 ♂♂, 14 ♀♀): CHINA: Yunnan: 1 ♀ (SNUC), 1 ♂, 2 ♀♀ (CZDY, a sample of eggs was preserved in 99.7 % ethanol at – 18 ° C), 2 ♀♀ (CQL), Pingbian County, Mount Dawei, near the hotel on the peak, ca. 2093 m, 2018. v. 26 – 27, Zhi-Wei Dong & Lu Qiu; 1 ♀ (CQL), Pingbian County, Mount Dawei, at the top of the mountain, 2016. v. 16, Lu Qiu; 1 ♂ (CZDY), 1 ♀ (CQL), Pingbian County, Mount Dawei, 2016. v. 18, Lu Qiu; 1 ♀ (CJM), Pingbian County, Mount Dawei, 2017. vii. 24 – 28, Ming Jin; 1 ♂ (CYM), 3 ♀♀ (CZDY), 2 ♀♀ (MHBU), Pingbian County, Mount Dawei, near 22 ° 54 ’ 21.21 ” N, 103 ° 41 ’ 48.73 ” E, 1700 – 2100 m, 2015. vi. 2, Tian-Long He; 1 ♂ (CBWX), Pingbian County, Mount Dawei, 2000 – 2200 m, 2009. v. 20 – 21, Wen-Xuan Bi; 1 ♂ (MHBU), 2 ♂♂, 1 ♀ (MYNU), Pingbian County, Mount Dawei, Dajianshan, 2100 m, 2015. v. 20, Jian-Yue Qiu; 1 ♂ (MYNU), Pingbian County, Mount Dawei, Dajianshan, 2100 m, 2018. v. 25 – 27, Hao Xu & Jian-Yue Qiu.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A0CFFB1FF5AF8A56F119314.taxon	diagnosis	Diagnosis. Large sized, elongate, strongly convex, green colored, anterior part of head, and all legs purple, pronotum, middle part of elytra and abdomen with purplish luster. Tempora strongly convex, separated from head by sulci. Elytra moderately convex, widest in apical 2 / 5, with densely and irregularly scattered round or oval encircled striae, each encircled area moderately convex. Male protarsi extremely narrow, pro- and mesotibiae nearly straight. Female possessing relative long legs. Ovipositor elongate, acute at apex.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A0CFFB1FF5AF8A56F119314.taxon	description	Description. Male (Fig. 11 A, C). Light to dark green, antennae, mouthparts, and tarsi dark brown, anterior part of head, and all legs purple, pronotum, middle part of elytra and abdomen with purplish luster. Body elongate, length 23.6 – 26.1 mm, width 7.9 – 8.9 mm, strongly convex, noticeably constricted between pronotum and elytra. Head (Fig. 16 L, P) transversely quadrate, densely and markedly punctate; with outer margin deeply notched between genae and clypeus; clypeus transversely hexagonal, gently bent downwards in front, anterior margin emarginate, clypeal transverse impression marked; frontoclypeal suture furrowed, widely U-shaped; genae strongly raised, depressed before eyes, roundly produced anterolaterally; frons broad, strongly wrinkled, frontal impressions vague, vertexal median impressions broadly presented; eyes transversely reniform, strongly convex laterally; inner ocular sulci grooved along inner margins, extending posteriorly and ventrally, encircled and separated tempora; tempora strongly convex, finely punctate. OI = 54.0 – 54.8. Antennae (Fig. 18 G) slender, reaching basal third of elytra, with antennomeres weakly thickened to apices; relative lengths of antennomeres: 0.85: 0.37: 1.39: 1.38: 1.45: 1.45: 1.34: 1.24: 1.21: 1.14: 1.34. Mentum (Fig. 16 P) inversely trapezoidal, lateral margins straight; medial surface sparsely and coarsely punctate, with several large pores with long setae, gradually rising anteriorly, depressed along both sides. Pronotum (Fig. 17 K) quadrate, PW / PL = 1.04 – 1.10, widest in anterior third, anterior margin nearly straight, anterior marginal border marked; lateral margins nearly straight, slightly widened at widest point, lateral marginal borders extremely thin, faintly presented; posterior margin weakly rounded, emarginate, posterior marginal border markedly presented; anterior angles rounded; posterior angles obtuse; disc strongly convex, a pair of impressions vague to absent in middle, shagreened, and densely markedly punctates. Scutellum triangular, glossy. Elytra oblong, widest in apical 2 / 5, EL / EW = 1.96 – 1.97; strongly convex, highest in middle; with densely and irregularly scattered round or oval striae, each encircled area moderately convex; intervals sparsely and finely punctate. Prosternum (Fig. 17 P) shagreened, finely and sparsely punctate; prosternal process declivous, pointed at apex; hypomeron strongly rugulose, shagreened. Metasternum glossy, metaventral anterior process transversely aciculate. Abdomen (Fig. 19 V) depressed, surface somewhat rough, wrinked along anterior margins and sides of each sternite, densely and finely punctate; sternites III and IV each shallowly depressed in posterior middle, with central portion of each impressions weakly convex. Legs slender. Protibiae (Fig. 19 D) nearly straight, apical half of inner margins pubescent; mesotibiae (Fig. 19 E) nearly straight, weakly curved in apical fourth, apical half of inner margins pubescent; metatibiae (Fig. 19 F) sinuous, apical 3 / 5 of inner margins feebly pubescent, outer margins depressed before apices. Aedeagus (Fig. 20 G) elongate, weakly curved in lateral view; parameres slender, 0.22 as long as total length, with rounded apex. Sternite VIII (Fig. 20 R) with apical lobes broadly hooked. Female (Fig. 11 B, D). Wider than male, length 21.4 – 24.4 mm. Elytra more convex, highest in middle; abdomen straight in lateral view; without impressions on sternites. OI = 42.4 – 46.7, PW / PL = 1.15 – 1.18, EL / EW = 1.66 – 1.67. Ovipositor (Fig. 20 V) elongate, gradually narrowing apically, with acute weakly beveled. Variability. Several individuals have antennomere XI strongly narrowing from middle towards the apex (Fig. 18 Gv). Comparative notes. Morphostenophanes purpurascens is assigned to the aenescens - group due to its slender habitus; elongate elytra widest in apical 2 / 5, with densely and irregularly scattered encircled segments of striae; and male sternites III and IV each depressed posterior of the middle. However, M. purpurascens is unique within aenescens - group. Compared with the stout females of other congeners, the female of M. purpurascens is much more slender with long legs; thus appearing like a male with wider and more convex elytra. The more acute ovipositor is similar to those of elegantulus and atavus - group. M. purpurascens can also be easily distinguished from congeners by its purplish body color, strongly convex tempora separated by sulci, and straight pro- and mesotibiae.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A0CFFB1FF5AF8A56F119314.taxon	discussion	Comments. Localities of M. purpurascens and M. iridescens are only 39 kilometers apart. However, the two species are very different in body colors, habitus, and many detailed characters, which suggests they may have different last common ancestors. M. iridescens is clearly related to M. cuproviridis and M. aenescens yelang (see comparative note of M. iridescens and comments of M. aenescens). M. purpurascens has monotonous colors, lacks pinkish luster on the pronotum, elytra striae and legs, and unique acute ovipositor, which is typical in the atavus - and elegantulus - group, and could be a primitive feature, which indicates it may have developed from an earlier ancestor.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A0CFFB1FF5AF8A56F119314.taxon	distribution	Distribution. (Map 1) CHINA: Yunnan.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A0CFFB1FF5AF8A56F119314.taxon	etymology	Etymology. The new species is named from the Latin epithet ‘ purpurascens’ (purple colored) referring to its purple body color.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A01FFBAFF5AFF196A6894B4.taxon	description	(Figs. 12 A – D; 13; 16 S, T; 17 U; 18 J; 19 J – L, X; 20 I, T, W)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A01FFBAFF5AFF196A6894B4.taxon	materials_examined	Type materials. CHINA: Yunnan: ♂ (Holotype, SNUC), Dali Prefecture, Weishan County, Mount Weibao, 2450 – 2580 m, 2017. vii. 7, Wen-Xuan Bi. Paratypes (65 ♂♂, 46 ♀♀): CHINA: Yunnan: 1 ♀ (SNUC), 1 ♂, 1 ♀ (MHBU), 10 ♂♂, 6 ♀♀ (CBWX, three samples each including eggs and muscle tissue were preserved in 99.7 % ethanol at – 18 ° C in CZDY), Dali Prefecture, Weishan County, Mount Weibao, 2450 – 2580 m, 2017. vii. 7, Wen-Xuan Bi; 1 ♂, 1 ♀ (CZDY), 6 ♂♂, 4 ♀♀ (CBWX), Dali Prefecture, Weishan County, Mount Weibao, 2400 – 2500 m, 2015. viii. 16 – 18, Wen-Xuan Bi; 2 ♂♂, 2 ♀♀ (CZDY, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C), 1 ♂♂, 2 ♀♀ (CDZW), 2 ♂♂, 2 ♀♀ (KIZ) Dali Prefecture, Weishan County, Ma’anshan Country, 25 ° 19 ’ 59.67 ” N, 100 ° 3 ’ 13.91 ” E, 2019. I, Zhi-Wei Dong; 1 ♂ (CZDY, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C), 1 ♀ (CZJZ), Dali Prefecture, Mount Cang, ca. 2100 m, 2018. vii. 20, Jia-Zhi Zhang; 13 ♂♂, 6 ♀♀ (MYNU), Dali Prefecture, Mount Cang, Jiangjundong, 2015. vii. 27, Xin-Ran Li & Zhi-Wei Qiu; 2 ♂♂, 2 ♀♀ (MYNU), Dali Prefecture, Mount Cang, Malong Peak, 2500 m, 2011. ii. 17, Yun-Chun Li & Yong-Sheng Pan; 1 ♂, 1 ♀ (CZDY), Dali Prefecture, Binchuan County, Jizushan Nature Reserve, 25 ° 57 ’ 37.75 ’’ N, 100 ° 23 ’ 13.38 ’’ E, 2297 m, 2010. vii. 12, Xiao-Bin Song; 1 ♂, 1 ♀ (MHBU), 5 ♂♂, 4 ♀♀ (CZDY, a sample of eggs tissue was preserved in 99.7 % ethanol at – 18 ° C) Dali Prefecture, Binchuan County, Jizushan Nature Reserve, 2.5 km WS. Jizushan Town, 2400 – 2600 m, 2019. vi. 7, native collector; 4 ♂♂, 2 ♀♀ (MYNU), Dali Prefecture, Binchuan County, Jizushan Nature Reserve, 2350 m, 2016. ii. 20, Hao Xu & Jian-Yue Qiu; 1 ♂ (CZDY, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C), Dali City, Binchuan County, nr. Shifenshuiliangzi, 2018. iv. 16, native collector. 1 ♀ (CZDY), Dali City, Binchuan County, 2 km N Muzhulong, 2019. iii, native collector. 2 ♂♂, 2 ♀♀ (CBWX, a sample of eggs was preserved in 99.7 % ethanol at – 18 ° C in CZDY), Mount Wuliang, near De’an Village, Sheyaoqing, 2300 – 2450 m, 2017. vii. 10 – 11, Wen-Xuan Bi; 2 ♂♂ (CZDY, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C), Lincang City, Yun County, Manwan Town, Shuibatou Village, 24 ° 41 ’ 43 ’’ N, 100 ° 21 ’ 10 ’’ E, 1942 m, 2016. ii. 10, Zi-Chun Xiong; 1 ♂ (CZDY), Lincang City, Yun County, Manwan Town, Shuibatou Village, 2200 m, 2017. viii, Zi- Chun Xiong; 1 ♂, 1 ♀ (CZDY), Lincang City, Manwan Town, Shuibatou Village, 2200 m, 2018. x, Zi-Chun Xiong; 2 ♂♂, 2 ♀♀ (MYNU), Lincang City, Manwan Town, 2016. iii. 8, Zi-Chun Xiong; 1 ♂, 1 ♀ (CYM), Lijiang City, Qina Town, Dongfeng Country, 2017. viii. 9, Zhen-Lian Ya; 1 ♂ (CZDY), Lijiang City, Yongsheng County, Ludila Town, nr. Zhiduping Village, 2018. viii, native collector; 1 ♂, 1 ♀ (MYNU), 1 ♂ (CZDY), 1 ♂, 2 ♀♀ (CQL), Xinping County, Mount Ailao, Laoniuchang, 23 ° 58 ’ 27 ” N, 101 ° 33 ’ 18 ” E, 1987 m, in pine forest, 2018. v. 24, Lu Qiu & Zhi-Wei Dong; 1 ♂ (CQL), Xinping County, Mount Ailao, Gasa Town, Yaonan Village, 2016. v. 13, Lu Qiu.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A01FFBAFF5AFF196A6894B4.taxon	diagnosis	Diagnosis. Medium sized, elongate, moderately convex, uniformly grey-black, elytral encircled segments of striae usually without pinkish luster, in very few individuals with extremely indistinct pinkish luster. Elytra widest at apical 2 / 5 in male, with densely and irregularly scattered round or oval segments of striae, each encircled area moderately convex. Male protibiae curved in apical third, each with dorsal side of inner margin more concave than ventral one.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A01FFBAFF5AFF196A6894B4.taxon	description	Description. Male (Figs. 12 A, B, D; 13). Grey-black, shagreened, mouth parts, and tarsi dark brown. Body elongate, length 17.8 – 22.3 mm, width 6.0 – 7.2 mm, moderately convex dorsad, noticeably constricted between pronotum and elytra. Head (Fig. 16 S, T) transversely quadrate, finely microsculptured, with densely scattered marked punctures; with outer margin moderately notched between genae and clypeus; clypeus transversely hexagonal, gently bent downwards in front, anterior margin nearly straight, emarginate, clypeal transverse impression marked to absent, frontoclypeal suture deeply grooved, widely U-shaped; genae strongly raised, depressed before eyes, roundly produced anterolaterally; frons broad, anterior part gradually sloping forwards, frontal impressions and vertexal lateral impressions vague to marked, frontal impressions sometimes connected in middle, vertexal median impression shallow, vaguely presented; eyes transversely reniform, strongly convex laterally; inner ocular sulci sharply grooved along inner margins; tempora weakly convex, coarsely punctate. Distance between eyes about 2 – 2.5 times as transverse diameter of an eye. Antennae (Fig. 18 J) slender, reaching basal 2 / 9 to 2 / 7 of elytra, with antennomeres weakly thickened to each apices. Mentum (Fig. 16 T) quadrate, lateral margins nearly straight; medial surface sparsely and coarsely punctate, with several large pores with long setae, gradually rising anteriorly, depressed along both sides. Pronotum (Fig. 17 U) barrel-shaped, slightly wider than long, widest at or anterior to the midpoint, anterior margin nearly straight, more or less projecting in middle, anterior marginal border marked; lateral margins rounded, more or less widened at widest point, lateral marginal borders thin, visible in dorsal view slightly behind anterior half; posterior margin nearly rounded, emarginate, posterior marginal border; anterior angles rounded or obtuse; posterior angles obtuse or nearly vertical; disc moderately convex, a pair of impressions vague to absent in middle, a stria connecting small strial punctures presented before the posterior marginal border, shagreened, sparsely and finely to coarsely punctate. Scutellum triangular, glossy. Elytra oblong, widest near apical 2 / 5, the length about two times the width; moderately convex, with densely and irregularly scattered round or oval segments of striae, each encircled area moderately convex; intervals more or less wrinkled, densely and finely punctate. Prosternum (Fig. 17 U-CDls-v) shagreened, finely and sparsely punctate; prosternal process declivous, pointed at apex; hypomeron strongly rugulose, shagreened. Metasternum transversely wrinkled. Abdomen (Fig. 19 X) depressed, surface somewhat rough and wrinkled, densely and finely punctate; sternites III and IV each shallowly depressed in posterior middle, with central portion of each impressions weakly convex. Legs slender. Protibiae (Fig. 19 J) curved in apical third, apical 3 / 5 of inner margins concave, each with dorsal side of inner margin more concave than ventral one, pubescent; mesotibiae (Fig. 19 H) curved in apical third, apical half of inner margins pubescent; metatibiae (Fig. 19 L) more or less sinuous, apical 3 / 5 of inner margins weakly pubescent, outer margins depressed before apices. Aedeagus (Fig. 20 I) elongate, curved in lateral view; parameres slender, about 0.21 as long as total length, with flabellate apex. Sternite VIII (Fig. 20 T) with apical lobes nearly rounded, each inferior margin projecting, forming a hook. Female. Wider than male, length 17.9 – 22.2 mm. OI = 55.6 – 59.2. Elytra more convex, elytra much wider, widest in middle; abdomen straight in lateral view; without impressions on sternites. Ovipositor (Fig. 20 W) shortened, abruptly narrowing terminally from apical third. Variability. This species is widely distributed from central to eastern Yunnan with distinct geographical variation, which can be divided into seven populations. All comparisons are based on the general properties of specimens from the locality of the holotype, i. e., Weibao Mountain. Ranges of all seven populations are shown in Map 2. The populations from Weibao Mountain, Cang Mountain, and Ma’an Mountain are extremely similar, and are defined here as a Central Dali population (CDl in Figs. 13, 16, 17, 18). They are characterized by having clypeus weakly protruding anteriorly, with anterior margin gently rounded and weakly emarginate in middle; antennae reaching basal fourth of elytra; relative lengths of antennomeres: 0.73: 0.34: 1.24: 1.12: 1.16: 1.19: 1.12: 1.02: 1.00: 0.95: 1.12; male OI = 49.4; pronotum barrel-shaped in most individuals, sometimes variable in shape, with anterior angles more strongly projecting anteriorly with lateral margins buckled in middle, PW / PL = 1.03 – 1.12, disc with densely scattered shallow but conspicuously marked punctures, punctures more or less connected with each other on middle of disc, pronotal anterior angles only weakly protruded, feebly rounded, but in few small individuals more strongly protruding anteriorly, obtusely angled. Female OI = 55.5. Pronotum moderately convex, PW / PL = 1.12 – 1.21. Elytra wider, EL / EW = 1.74 – 1.79, dorsum nearly straight, gradually risen posteriorly, highest slightly behind basal half. The population from Jizu Mountain (JzM in Figs. 13, 16, 17, 18) has darker pitch-black body color, head and pronotum more coarsely punctate, punctures on all legs also larger and deeper; head notably narrower, with eyes narrower and less protruding laterally; antennae slightly shorter, with antennomeres III, IV, X and XI notably shorter; head notably narrower; elytral intervals more strongly wrinkled. The population from Shifenshuiliangzi and adjacent mountains on the east side of Binchuan County is defined here as Eastern Binchuan population (EBc in Figs. 13, 16, 17, 18), which resembles the type population, but with widier head, narrower eyes less protruding laterally, distance between eyes extremely longer than transverse diameter of an eye, OI = 55.3 in male, and 2.43 in female. Antennae distinctly shorter, with antennomeres III and IV notably shortened, relative lengths of antennomeres: 0.65: 0.27: 1.05: 0.95: 0.97: 0.99: 0.97: 0.95: 0.90: 0.83: 0.99. Pronotum densely and markedly punctate, with punctures merely connected with each other on median area of disc. The population from Yongsheng County, Lijiang City (YsC in Figs. 13, 16, 17, 18) is with darker pitch-black body color, clypeus more protruding anteriorly, with anterior margin more distinctly emarginate; antennae distinctly short, only reaching basal sixth of elytra, relative lengths of antennomeres: 0.73: 0.32: 1.00: 1.00: 1.16: 1.07: 1.07: 1.02: 1.00: 0.92: 0.95; OI = 53.9 in male, and 2.59 in female; pronotum straight or only weakly projecting in middle, anterior angles obtuse; elytral encircled segments of striae with distinctly connected punctures; intervals more wrinkled. One male has genae more strongly and roundly protruding laterally, and only weakly curved, shortened aedeagus. The population from Manwan Town, Lincang City (MwT in Figs. 13, 16, 17, 18) has clypeus more protruding anteriorly, with anterior margin more distinctly emarginate; antennae elongate, reaching basal fourth of elytra or slightly longer; relative lengths of antennomeres: 0.75: 0.34: 1.36: 1.28: 1.33: 1.36: 1.29: 1.19: 1.16: 1.11: 1.28; OI = 48.8 – 50.2 in male, and 2.57 in female; head and pronotum more glabrous, sparsely and finely punctate, punctures more distinct in small individuals; lateral margins evenly rounded. All legs with smaller and shallower punctures. The population from Wuliang Mountain (WlM in Figs. 13, 16, 17, 18) has clypeus more protruding anteriorly, with anterior margin more distinctly emarginate; antennae slightly longer, reaching basal 2 / 7 of elytra; relative lengths of antennomeres: 0.70: 0.38: 1.28: 1.16: 1.28: 1.26: 1.19: 1.16: 1.11: 1.00: 1.05; OI = 48.7 – 51.9 in male, 2.34 in female; genae less projecting anterolaterally; pronotum hexagonal, lateral margins buckled in middle, anterior angels bent ventrally, posterior angles vertical, somewhat projecting laterally; disc densely scattered shallow punctures. The population from Ailao Mountain (AlM in Figs. 13, 16, 17, 18) has clypeus more protruding anteriorly, with anterior margin more distinctly emarginate; antennae slightly shorter, reaching basal fifth of elytra, with each antennomeres notably shorter, relative lengths of antennomeres: 0.77: 0.42: 1.31: 1.26: 1.29: 1.26: 1.19: 1.11: 1.07: 0.99: 1.09; OI = 50, and 2.16 in female; pronotum densely and markedly punctate, posterior angles distinctly projecting laterally, more notebly constricted behind widest point in female; male possessing slightly wider protarsi; impressions of sternites III and IV with central swelling portions ridged.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A01FFBAFF5AFF196A6894B4.taxon	distribution	Distribution. (Map 1, 2) CHINA: Yunnan. Comparative notes. Morphostenophanes yunnanus resembles M. aenescens, see comparative notes of the latter.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A01FFBAFF5AFF196A6894B4.taxon	discussion	Comments Morphostenophanes yunnanus is here defined to include a series of populations from several adjacent localities in central and eastern Yunnan. They are very similar in general appearance, including dark-grey body color, slender and moderately convex habitus, densely scattered elytral encircled segments of striae without pinkish luster (in most cases), male protibiae curved in apical third, with dorsal side of inner margin more concave than ventral one, and almost the same genital structures; but each slightly differs from each other in certain characters, e. g. length of antennae, cephalic and pronotal punctation. Comparison of any two populations may result in a subspecifice or even species level classification (e. g. specimens from Manwan Town and Jizu Mountain seems very different, with the former possessing wider head and pronotum, sparsely and finely punctate, antennae reaching basal fourth of elytra, while the latter possessing narrower head and pronotum, densely and coarsely punctate, antennae only reaching basal fifth of elytra). However, a comprehensive comparison of all the seven populations shows relatively continuous variability: individuals from south-central populations (Lincang, Wuliang Mountain, Ailao Mountain) bear less obvious cephalic and pronotal punctations, which is more pronounced in northeastern populations (Central Dali, Jizu Mountain, Eastern Binchuan, Lijiang City); length of antennae also successively becoming shorter from the west to the east. In the present study, thirty specimens from Weibao Mountain were examined by the author, which showed considerable intrapopulation variations, especially in pronotal shapes. The pronotum of most individuals is barrelshaped with lateral margins roundly curved, inclined inwards anteriorly, and somewhat constricted before the base, but in few individuals, the lateral margins are more or less projecting at the widest point, which is similar to those of Wuliang Mountain population. Specimens from other localities were sparser but it is reasonable to believe that such intrapopulation variations also exist in those populations. Specimen data and field observations show that M. yunnanus lives at elevations of 1900 to 2600 meters in central and eastern Yunnan. Such mountainous regions are spread all over this region and connected with each other. Similar general appearance among the above-mentioned seven populations suggest that all these populations spread to these mountains within a relatively short period of time, or relatively high rate of interpopulation gene flow were maintained. Therefore, it is inadequate to classify any of these populations as distinct subspecies or species. A mo- lecular analysis of with more specimens from this whole region is required to reveal exact relations among these populations.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A01FFBAFF5AFF196A6894B4.taxon	etymology	Etymology. The new species is named after the province of its type locality.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A05FFB8FF5AFDE26AB59498.taxon	description	(Figs. 14 A – D; 16 N, R; 17 M, R; 18 I; 19 M – O, Y; 20 J, U)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A05FFB8FF5AFDE26AB59498.taxon	materials_examined	Type materials. CHINA: Yunnan: ♂ (Holotype, SNUC), Pu’er City, Zhengyuan County, Heping Country, 23 ° 56 ’ 13.73 ” N, 101 ° 29 ’ 4.98 ” E, 2299 m, 2011. ix. 26, Guo-Lin Li. Paratypes (2 ♂♂, 4 ♀♀): CHINA: Yunnan: 1 ♀ (SNUC), 1 ♂, 1 ♀ (MHBU), 1 ♂, 1 ♀ (CZDY, two samples each including muscle tissue and eggs were preserved in 99.7 % ethanol at – 18 ° C), Pu’er City, Zhengyuan County, Heping Country, 23 ° 56 ’ 13.73 ” N, 101 ° 29 ’ 4.98 ” E, 2299 m, 2011. ix. 26, Guo-Lin Li; 1 ♀ (CBWX), Pu’er City, Mengda Town, Baiyanzi, 1700 m, 2019. viii. 29, Y. - H. Li.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A05FFB8FF5AFDE26AB59498.taxon	diagnosis	Diagnosis. Medium-sized, coppery colored, elongate and strongly convex species. Head and pronotum with extremely coarse punctation on the head and pronotum. Pronotal disc with a pair of deep impressions in middle. Elytra ovate, short strial punctures irregularly scattered and interconnected, forming a network, intervals uneven in sizes and shapes, some strongly swelling, forming tubercles.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A05FFB8FF5AFDE26AB59498.taxon	description	Description. Male (Fig. 14 A, C). Color coppery, hypomeron and prosternum dark green, antennae, mouthparts and tarsi dark brown, claws reddish brown. Body elongate, length 18.9 – 19.9 mm, width 6.7 mm, strongly convex, noticeably constricted between pronotum and elytra. Head (Fig. 16 N, R) transverse, subquadrate, coarsely microsculptured, scattered with large and deep punctures; outer margin strongly notched between genae and clypeus; clypeus transversely hexagonal, gently bent downwards in front, with anterior margin nearly straight, weakly emarginate in middle, clypeal transverse impression marked; frontoclypeal suture deeply grooved, becoming shallower anterolaterally, widely U-shaped; genae strongly raised, depressed before eyes, strongly and roundly produced anterolaterally; frons broad, gradually sloping forwards, vertexal lateral impressions shallow and broad, vertexal median impression marked, becoming deeper posteriorly; eyes transversely reniform, strongly convex laterally; inner ocular sulci deeply grooved along inner margins, becoming broader posteriorly; tempora convex, coarsely punctate. OI = 52.3 – 53.4. Antennae (Fig. 18 I) slender, reaching basal 2 / 7 of elytra, with antennomeres weakly thickened to apices; relative lengths of antennomeres: 0.58: 0.49: 1.05: 1.02: 1.12: 1.11: 1.12: 1.00: 1.02: 1.00: 1.14. Mentum (Fig. 16 R) quadrate, lateral margins straight or weakly rounded; medial surface sparsely and coarsely punctate, with several large pores with long setae; gradually rising anteriorly, depressed along both sides. Pronotum (Fig. 17 M) barrel-shaped, PW / PL = 1.05 – 1.08, finely microsculptured, widest in middle, anteri- or margin emarginate, projecting in middle, with thin marginal border, absent on middle; lateral margins weakly rounded, lateral marginal borders thin, visible in dorsal view slightly exceeding anterior half; posterior margin weakly rounded, slightly emarginate, posterior marginal border distinct; anterior angles rounded; posterior angles obtuse; disc moderately convex, with scattered large and deep punctures, with a pair of rounded impressions in middle, furrowed behind each impression. Scutellum widely triangular, glossy, sparsely and finely punctate. Elytra ovate, widest near basal 3 / 5, EL / EW = 1.89; moderately convex, highest near middle; short strial punctures irregularly scattered and interconnected, forming a network, intervals uneven in size and shape, some strongly swelling, forming tubercles. Prosternum (Fig. 17 R) rugulose, finely and sparsely punctate; prosternal process weakly declivous, apex truncate; hypomeron rugulose, finely microsculptured, very sparsely and finely punctate. Metasternum finely wrinkled. Abdomen (Fig. 19 Y) depressed, surface weakly rough, densely and finely punctate, sternite III more or less convex between metacoxae; sternite IV broadly depressed in apical middle, with the central part convex. Legs slender. Protibiae (Fig. 19 M) nearly straight, curved in apical fourth, thickening terminally, apical 3 / 5 of inner margins emarginate and pubescent; mesotibiae (Fig. 19 N) curved in apical fourth, apical 3 / 5 of inner margins pubescent; metatibiae (Fig. 19 O) nearly straight, apical 3 / 5 of inner margins emarginate, weakly pubescent. Aedeagus (Fig. 20 J) elongate, weakly curved in lateral view; parameres slender, 0.27 as long as total length, flattened terminally, with broadly widened and flabellate apex. Apical lobes of sternite VIII (Fig. 20 K) hooked. Female (Fig. 14 B, D). Stouter than male, length 19.1 – 21.2 mm; OI = 56.5, PW / PL = 1.12; elytra stouter, EL / EW = 1.59 – 1.69; abdomen straight in lateral view; without impressions on sternites. Ovipositor shortened, abruptly narrowing terminally from apical third. Comparative notes. Morphostenophanes chongli resembles M. tuberculatus. Both species have elytra scattered with unevenly sized tubercles. However, M. chongli has intermittent distribution of elytral tubercles (continuous in the latter), with dark striae around each tubercle (without in the latter). Additionally, M. chongli can be distinguished from the latter by its coarsely punctate head and pronotum (fine in the latter), larger body size, and less widened paramere apex.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A05FFB8FF5AFDE26AB59498.taxon	discussion	Comments. Similarly oblong elytra, impressions on head and abdomen, weak pubescence along inner margins of metatibiae and shortened ovipositor of Morphostenophanes chongli show its possible relation with aenescens - group, but none of the species in the aenescens - group have elytra sculpture similar to that of M. chongli. A future molecular study is required to verify the exact status of M. chongli within the genus.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A05FFB8FF5AFDE26AB59498.taxon	distribution	Distribution. (Map 1) CHINA: Yunnan.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A05FFB8FF5AFDE26AB59498.taxon	etymology	Etymology. The new species is named from ‘ Chongli’, a god of fire in Chinese mythology	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A07FF87FF5AFEA96F4E93E6.taxon	description	(Figs. 15 A – D; 17 N, S; 18 K; 20 K)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A07FF87FF5AFEA96F4E93E6.taxon	materials_examined	Type materials. CHINA: Yunnan: ♂ (Holotype, SNUC), Honghe Prefecture, Lüchun County, Huanglianshan Nature Reserve, 2018. v. 20, Chao-Ming Chen. Paratypes (5 ♂♂, 11 ♀♀): CHINA: Yunnan: 1 ♀ (SNUC), 1 ♀ (CZBS), Honghe Prefecture, Lüchun County, Huanglianshan Nature Reserve, 2018. v. 20, Chao-Ming Chen; 2 ♀♀ (CZDY, two samples each including muscle tissue and eggs were preserved in 99.7 % ethanol at – 18 ° C), Huanglianshan Nature Reserve, Yakou station, 1900 m, light trap, 2018. vi. 13, Hao Huang; 1 ♂, 1 ♀ (CZDY), 3 ♂♂, 5 ♀♀ (CBWX) Honghe Prefecture, Lüchun County, Huanglianshan Nature Reserve, 2019. viii. 26 – 28, Wen-Xuan Bi; 1 ♀ (CWC), Honghe Prefecture, Lüchun County, Huanglianshan Nature Reserve, 1800 – 1900 m, 2019. viii. 27, Chao Wu. VIET- NAM: 1 ♂ (CZJZ) Mount Sapa, 2019. vi, native collector. Diagnostic description. Male (Fig. 15 A, C). Color coppery, prosternum dark green, antennae, mouthparts and tarsi dark brown, claws reddish brown. Body elongate, length 18.2 – 22.2 mm, width 6.5 – 7.6 mm, strongly convex dorsally. Cephalic punctures shallower, OI = 48.5 – 50.9. Antennae (Fig. 18 K) much longer, reaching basal third of elytra; antennomeres more elongate, relative lengths of antennomeres: 0.75: 0.36: 1.21: 1.05: 1.28: 1.33: 1.29: 1.22: 1.22: 1.17: 1.36. Pronotum (Fig. 17 N) quadrate, PW / PL = 1.04, widest in anterior 2 / 7, anterior margin projecting in middle, with thin marginal border, becoming thinner towards middle; lateral margins nearly straight, lateral marginal borders thin, visible in dorsal view along anterior third; posterior margin feebly emarginate, posterior marginal border marked; anterior angles rounded, weakly projecting forwards; posterior angles obtuse; disc moderately convex, with shallower punctures, presenting a pair of rounded impressions in anterior third, and a pair of shallower impressions near anterior fifth. Elytra ovate, EL / EW = 1.80 – 1.88. Aedeagus (Fig. 20 K) elongate, basal piece longer, and much strongly curved in lateral view; parameres 0.23 as long as total length. Apical lobes of sternite VIII hooked. Female (Fig. 15 B, D). Stouter than male, length 20.1 – 23.3 mm; elytra more convex; pronotum wider; abdomen straight in lateral view; without impressions on sternites. OI = 54.5, PW / PL = 1.09 – 1.18, EL / EW = 1.60 – 1.63. Comparative notes. Morphostenophanes chongli glaber can be easily distinguished from the nominate subspecies by lighter body color, more slender antennae, shallower punctures on head and pronotum, a pair of extra shallow impressions on anterior fifth of pronotum, and much strongly curved aedeagus.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A07FF87FF5AFEA96F4E93E6.taxon	discussion	Comments. Morphostenophanes chongli glaber occurs in Huanglian Mountain at an altitude near 1900 meters, while the nominate subspecies distributed in Ailao Mountains at an altitude over 2300 meters. The two mountains are separated by several low altitude valleys, which must have also isolated these two populations, and caused differentiation.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A07FF87FF5AFEA96F4E93E6.taxon	distribution	Distribution. (Map 1) CHINA: Yunnan; North VIETNAM.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A07FF87FF5AFEA96F4E93E6.taxon	etymology	Etymology. The new subspecies is named from the Latin epithet ‘ glaber’, referring its relatively smooth head and pronotum.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A3DFF83FF5AFE296F079238.taxon	description	(Figs. 21 A – D; 23 A, E, C, G, I, K – M; 24 A, C, F)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A3DFF83FF5AFE296F079238.taxon	materials_examined	Type materials. CHINA: Yunnan: ♂ (Holotype, SNUC), Lincang City, Mount Dachao, 24 ° 12 ’ 18.14 ” N, 100 ° 19 ’ 14.25 ” E, 2018. xii., Zi-Chun Xiong. Paratypes (3 ♂♂, 4 ♀♀): CHINA: Yunnan: 1 ♀ (SNUC), 1 ♀ (MHBU), 2 ♀♀ (CZDY, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C), Lincang City, Mount Dachao, 24 ° 12 ’ 18.14 ” N, 100 ° 19 ’ 14.25 ” E, 2018. xi. 10, Zi-Chun Xiong; 1 ♂ (MHBU), 2 ♂♂ (CZDY), Lincang City, Mount Dachao, 24 ° 12 ’ 18.14 ” N, 100 ° 19 ’ 14.25 ” E, 2018. xii, Zi-Chun Xiong.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A3DFF83FF5AFE296F079238.taxon	diagnosis	Diagnosis. Small, bronze, extremely glossy and strongly shiny, short and strongly convex species. Elytral with rows of tubercles of variable sizes, each with purple central part. Superior margin of apical lobes of sternite VIII with shallow tick-shaped notch, dorsum of parameres strongly ridged.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A3DFF83FF5AFE296F079238.taxon	description	Description. Male (Fig. 21 A, C). Bronze and strongly shiny, antennomeres VII – XI and mouthparts brownish; elytra slightly greenish, elytral sutural band and central parts of elytral tubercles purplish; claws reddish brown. Body elongate, length 14.0 – 14.8 mm, width 5.2 – 5.5 mm, strongly convex, noticeably constricted between pronotum and elytra. Head (Fig. 23 A, E) transversely subquadrate, sparsely and finely punctate, outer margin strongly notched between genae and clypeus; clypeus transversely hexagonal, slightly convex in middle, gently bent downwards in front, with anterior margin nearly straight, weakly emarginate in middle, convex in middle before frontoclypeal suture; frontoclypeal suture moderately depressed; genae strongly raised, depressed before eyes, strongly and roundly produced anterolaterally; frons broad, gradually sloping forwards, vertexal lateral impressions vague; eyes transversely reniform, strongly convex laterally; inner ocular sulci shallowly grooved along inner margin; tempora moderately convex, finely punctate. OI = 53.4 – 55.5. Antennae (Fig. 23 I) slender, extremely long, exceeding basal third of elytra, with antennomeres weakly thickened to each apices; relative lengths of antennomeres: 0.54: 0.27: 0.83: 0.78: 0.78: 0.80: 0.85: 0.85: 0.85: 0.82: 0.99. Mentum (Fig. 23 E) quadrate, lateral margins nearly straight, finely punctate, with several large pores bearing long setae, gradually rising from base to apex, both sides of posterior half depressed. Pronotum (Fig. 23 C) quadrate, PW / PL = 1.10 – 1.13, widest slightly anterior to the midpoint, anterior margin projecting in middle, anterior marginal border faintly presented, missing in middle; lateral margins weakly curved, lateral marginal borders visible in dorsal view along anterior half; posterior margin nearly straight, posterior marginal border marked; anterior angles rounded; posterior angles obtuse; disc moderately convex, smooth, sparsely scattered with small but sharply marked punctures, pair of impressions marked or missing slightly before the middle. Scutellum widely triangular, extremely glossy. Elytra fusiform, widest in apical 2 / 5, EL / EW = 1.73 – 1.83; strongly convex, highest in basal third, weakly ridged in apical portion along suture; striae twisted, intervals intermittently dilated and convex, forming rounded or ovate tubercles, glossy, sparsely and finely punctate. Prosternum (Fig. 23 G) weakly rugulose, finely and sparsely punctate; prosternal process declivous, truncate at apex; hypomeron weakly rugulose. Metasternum glossy, weakly wrinkled. Abdomen depressed, surface smooth, somewhat wrinked along anterior margins and sides of each sternite, densely and finely punctate; sternite III with a short median border on anterior part between metacoxae; sternite III and IV weakly sulcate along both sides. Legs slender. Protibiae (Fig. 23 K) moderately curved at apical third, apical half of inner margins sparsely pubescent; mesotibiae (Fig. 23 L) weakly curved, apical 2 / 5 of inner margins pubescent; metatibiae (Fig. 23 M) nearly straight, more than half of apical inner margins pubescent, outer margins depressed before apices. Aedeagus (Fig. 24 A) elongate, curved in lateral view, with pair of large pits on apical flank of basal piece; parameres slender, dorsum strongly ridged along midline, 0.26 as long as total length, with broadly widened and flabellate apex, apical margin weakly thickened. Sternite VIII (Fig. 24 C) with apical lobes strongly oppositely curved in dorsal view, each superior margin with a shallow tick-shaped notch in lateral view. Female (Fig. 21 B, D) slightly larger than male, stouter, length 15.2 – 16.6 mm. More constricted between pronotum and elytra; OI = 55.8, PW / PL = 1.15 – 1.17; elytra more convex, EL / EW = 1.50 – 1.60; abdomen straight in lateral view. Ovipositor (Fig. 24 F) shortened, abruptly narrowing terminally from apical third, apex beveled. Comparative notes. Morphostenophanes linglong is closely related to M. metallicus. Both species have small and glossy body with reflective luster, slender antennae and legs, similar pronotal shape, fusiform elytra, similar sternite VIII and short ovipositor. The elytra of M. linglong bear intervals intermittently convex, forming purple tubercles. This character state is shared with M. bannaensis, but not seen in M. metallicus. However, M. bannaensis is clearly related to species within atavus - group, which contains other species with striate elytra.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A3DFF83FF5AFE296F079238.taxon	distribution	Distribution. (Map 1) CHINA: Yunnan. Etymolo gy. The specific epithet ‘ linglong [Ḣẇ] ’ means small and elegant in Chinese, in reference to its small and gorgeous body.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A3CFF8EFF5AF9496EA096C8.taxon	description	(Figs. 22 A – D; 23 B, F, D, H, J, N – P; 24 B, D, E)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A3CFF8EFF5AF9496EA096C8.taxon	materials_examined	Type materials. CHINA: Yunnan: ♂ (Holotype, SNUC), Dehong Prefecture, Longchuan County, 2000 m, 2017. vi. 16 – 28, Wen-Xuan Bi. Paratypes (6 ♂♂, 14 ♀♀): CHINA: Yunnan: 1 ♀ (SNUC), 1 ♂, 1 ♀ (CZDY), 1 ♂, 1 ♀ (CBWX), Dehong Prefecture, Longchuan County ,, 2000 m, 2017. vi. 16 – 28, Wen-Xuan Bi; 1 ♂, 3 ♀♀ (CBWX), Tengchong City, Houqiao Town, Heinitang Village, 2000 m, 2017. ix. 7 – 10, Xiao-Dong Yang; 1 ♂, 1 ♀ (CZDY), 1 ♂, 1 ♀ (MHBU), 1 ♂, 2 ♀♀ (CBWX), Tengchong City, Houqiao Town, Heinitang Village, 1850 – 1950 m, 2018. ix. 4 – 6, Wen-Xuan Bi; 2 ♀♀ (CBWX), Tengchong City, Houqiao Town, Heinitang Village, 1850 – 1950 m, 2018. ix. 11 – 13, Xiao-Dong Yang; 1 ♀ (MYNU), Tengchong City, Mingguang Country, Zizhi Village, 2200 m, 2016. ii. 14, Hao Xu & Jian-Yue Qiu; 1 ♀ (CJQY), Yingjiang County, Xima Town, 5 km N Baobian Village, 1870 m, 2018. xii, native collector.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A3CFF8EFF5AF9496EA096C8.taxon	diagnosis	Diagnosis. Small to medium sized, bronze and strongly shiny, extremely smooth, shortened and strongly convex species. Elytral striae grooved, intervals slightly ridged. Antennae and legs slender. Apical lobes of sternite VIII oppositely curved in dorsal view, spatular in lateral view, with each superior margin presenting a tick-shaped notch.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A3CFF8EFF5AF9496EA096C8.taxon	description	Description. Male (Fig. 22 A – C). Bronze and strongly shiny, antennae segments VII – XI and mouthparts brownish; elytra slightly greenish; tarsus claws reddish brown. Body elongate, length 15.3 – 18.1 mm, width 5.6 – 6.8 mm, strongly convex, noticeably constricted between pronotum and elytra. Head (Fig. 23 B, F) transversely subquadrate, sparsely and finely punctate, with outer margin strongly notched between genae and clypeus; clypeus transversely heptagonal, slightly convex in middle, gently bent downwards in front, anterior margin nearly straight, weakly emarginate at middle, frontoclypeal suture deeply grooved, arcuate; genae strongly raised, depressed before eyes, strongly and roundly produced anterolaterally; frons broad, gradually sloping forwards, vertexal lateral impressions vague; eyes transversely reniform, strongly convex laterally; inner ocular sulci shallowly grooved along inner margins, becoming broader posteriorly; tempora moderately convex, finely punctate. OI = 57.7 – 60.5. Antennae (Fig. 23 J) slender, reaching basal third of elytra, with antennomeres weakly thickened forwards, and distinctly dilated at apices; relative lengths of antennomeres: 0.68: 0.34: 1.11: 1.07: 1.17: 1.16: 1.19: 1.11: 1.11: 0.97: 1.11. Mentum (Fig. 23 F) quadrate, lateral margin slightly rounded; medial surface sparsely and coarsely punctate, with several large pores bearing long setae, gradually rising anteriorly, both sides of posterior half depressed. Pronotum (Fig. 23 D) quadrate, PW / PL = 1.06 – 1.14, widest at anterior angles, anterior margin nearly straight, anterior marginal border marked; lateral margins nearly straight, lateral marginal borders thin, visible in dorsal view along anterior third; posterior margin emarginate, posterior marginal border marked; anterior and posterior angles rounded; disc moderately convex, extremely smooth, very finely and sparsely punctate, a pair of impressions marked to entirely missing slightly beforemiddle. Scutellum widely triangular, extremely glossy. Elytra fusiform, widest in middle, EL / EW = 1.68 – 1.71; strongly convex, highest slightly before middle, ridged in apical portion along suture; with 9 grooved striae, 9 th stria branching at apical fourth of elytra, 1 st and 9 th, 2 nd and upper branch of 9 th, 3 rd and 8 th, 4 th and 5 th, and 6 th and 7 th converingat ends, 2 nd and 3 rd converging at fore-ends; intervals weakly ridged, glabrous, sparsely and finely punctate. Prosternum (Fig. 23 H) weakly rugulose, finely and sparsely punctate; prosternal process strongly declivous; hypomeron rugulose. Metasternum glossy, weakly wrinkled. Abdomen depressed, surface smooth, somewhat wrinked along anterior margins and sides of each sternite, densely and finely punctate; sternite III with a short median border on anterior part between metacoxae; sternite III and IV weakly sulcate along both sides. Legs slender. Protibiae (Fig. 23 N) moderately curved in apical third, apical half of inner margins pubescent; mesotibiae (Fig. 23 O) weakly curved, apical half of inner margins pubescent; metatibiae (Fig. 23 P) sinuous, more than half of inner margins pubescent, outer margins weakly depressed before apices. Aedeagus (Fig. 24 B) elongate, curved in lateral view; parameres slender, dorsal margin depressed slightly before apical half in lateral view, dorsum ridged along midline, 0.21 as long as total length, with broadly widened and flabellate apex. Sternite VIII (Fig. 24 D) with apical lobes strongly oppositely curved in dorsal view, each superior margin with tick-shaped notch in lateral view. Female (Figs. 22 B, D). Stouter than male, length 14.6 – 18.5 mm. Distance between eyes slightly longer, OI = 61.9 – 62.5; PW / PL = 1.16 – 1.18, more constricted between pronotum and elytra; elytra more convex, EL / EW = 1.51 – 1.63; abdomen straight in lateral view. Ovipositor (Fig. 24 E) shortened, abruptly narrowing terminally from apical third.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A3CFF8EFF5AF9496EA096C8.taxon	discussion	Comments. Morphostenophanes metallicus is distinctive for its extremely glossy body with mirror-like reflective luster. Additionally, the grooved elytral striae and ridged intervals are not present in other congeners. Striate elytra are common within the atavus - group. However, elytral striae of the atavus - group are furrowed with domed intervals. The small to medium body size, widely flabellate apex of parameres, strongly crooked apical lobes of sternite VIII with tick shaped notch in each superior margin, and shortened ovipositor clearly separate this species from those of atavus - group. Within the entire genus, M. metallicus is closely related to M. linglong, the latter resembling a miniature M. metallicus, but with enlarged and strongly raised tubercles unevenly distributed along each elytral intervals.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A3CFF8EFF5AF9496EA096C8.taxon	distribution	Distribution. (Map 1) CHINA: Yunnan.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A3CFF8EFF5AF9496EA096C8.taxon	etymology	Etymology. The new species is named from the Latin epithet ‘ metallicus’ referring its strongly glossy and shiny body.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A30FF8DFF5AFF196CF896BC.taxon	description	(Figs. 25 A – G; 31 A, E, K, P; 32 A, G – I; 33 A, G)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A30FF8DFF5AFF196CF896BC.taxon	materials_examined	Type material examined. Holotype of Promorphostenophanes atavus labelled: ‘ Yunnan Tientshun [handwritten in pink label] // Holotypus 1960 Promorphostenophanes atavus Kaszab [handwritten on a pink label with red border] ’ (female, HMHM, Fig. 25 C – G). Examined through five photographs taken by Tamás Németh (HNHM) and sent by Ottó Merkl. Additional material examined (1 ♂). CHINA: Yunnan: 1 ♂ (MYNU), east slope of Mount Gaoligong, 25 ° 17 ’ 48.33 ” N, 98 ° 45 ’ 52.98 ” E, 2402 m, 2014. iv. 11, at night, Xuan-Kong Jiang & Tian Lu.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A30FF8DFF5AFF196CF896BC.taxon	diagnosis	Diagnosis. Body large and elongate, black. Antennae short, reaching basal 2 / 7 of elytra; Pronotum widest in anterior fourth; elytra striate, with aspect ratio exceeding 1.8 in male, and 1.7 in female; apical spurs of mesotibiae exposed and visible in ventral view; metafemora reaching anterior half of sternite VII; metatibiae nearly straight; apical lobes of sternite VIII narrowly hooked in lateral view.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A30FF8DFF5AFF196CF896BC.taxon	description	Description of male. Male (Fig. 25 A, B). Color black, antennae, all tibiae, mouthparts, and claws reddish brown, shagreened. Body elongate, length 23.7 mm, width 8.4 mm, strongly convex, noticeably constricted between pronotum and elytra. Head (Fig. 31 A, F) transversely quadrate, sparsely and finely punctate, with outer margin distinctly notched between genae and clypeus; clypeus transversely hexagonal, slightly convex in middle, gently bent downwards in front, anterior margin nearly straight, weakly emarginate; frontoclypeal suture finely depressed, widely U-shaped, weakly protruding backwards in middle; genae strongly raised, depressed before eyes, strongly and roundly produced anterolaterally; frons broad, anterior part gradually sloping forwards, flattened in middle; eyes transversely reniform, strongly convex laterally; inner ocular sulci finely depressed, moderately grooved along inner margins, becoming broader posteriorly; tempora moderately convex, more coarsely punctate than frons. OI = 46.4. Antennae (Fig. 32 A) slender, reaching basal 2 / 7 of elytra, with antennomeres weakly thickened to apices; relative lengths of antennomeres: 0.71: 0.34: 1.19: 1.14: 1.29: 1.31: 1.29: 1.16: 1.14: 1.04: 1.19. Mentum (Fig. 31 F) inversely trapezoidal, lateral margins weakly rounded; medial surface finely punctate, with several large pores bearing long setae, gradually rising from base to apex, depressed along both sides. Pronotum (Fig. 31 K) inversely trapezoidal, PW / PL = 1.10, widest in anterior fourth, anterior margin nearly straight, anterior marginal border fine, interrupted in middle; lateral margins weakly rounded, lateral marginal borders thin, visible in dorsal view slightly before anterior half; posterior margin weakly rounded, emarginate in middle, posterior marginal border marked; anterior angles rounded; posterior angles obtuse; disc strongly convex, shagreened, finely and sparsely punctate, with a pair of vague impressions on middle. Scutellum widely triangular, glossy, finely punctate. Elytra elongate oval, widest near middle, EL / EW = 1.84; strongly convex, highest in basal 2 / 5; with 9 continuous furrowed striae; 9 th stria branching before apex, 1 st and 9 th, 2 nd and 7 th, 3 rd and 6 th, and 4 th and 5 th, converging at ends, 2 nd and 3 rd, and 5 th and 6 th converging at anterior ends; intervals strongly convex, shagreened, sparsely and finely punctate. Prosternum (Fig. 31 P) shagreened, finely and sparsely punctate; prosternal process declivous, truncate at apex; hypomeron weakly rugulose, shagreened. Metasternum glossy, metaventral anterior process weakly wrinkled. Abdomen depressed, somewhat rough and finely punctate, sternites III and IV sulcate on both sides, sternite V depressed laterally. Legs slender. Protibiae (Fig. 32 G) weakly curved, apical 3 / 5 of inner margins pubescent; mesotibiae (Fig. 32 H) weakly curved near apical third, apical 3 / 5 of inner margins pubescent, ventral apical tuft interrupted, mesotibial spur uncovered (marked in Fig. 32 H); metatibiae (Fig. 32 I) straight, apical 3 / 5 of inner margins pubescent, outer margins depressed before apices. Aedeagus (Fig. 33 A) elongate, curved in lateral view; parameres slender, 0.22 as long as total length, with ovoid apex. Sternite VIII (Fig. 33 G) with apical lobes narrowly hooked in lateral view. Comparative notes. Morphostenophanes atavus most resembles M. brevigaster. They share similar appearance and indistinguishable aedeagi. However, M. atavus can be easily distinguished from M. brevigaster by its shorter antennae which reaches 2 / 7 of elytra, compared to those of M. brevigaster reaching third of elytra; each antennomere less sharply dilated towards apex; pronotum widest in apical fourth, compared to that of M. brevigaster widest near the middle; elytra more elongate, highest posterior to the midpoint, compared to those of M. brevigaster highest anterior to the midpoint; elytral striae less depressed than those of the latter; all legs shorter, metafemora reaching anterior half of sternite VII, compared to those M. brevigaster reaching apex of sternite VII; protibiae in male evenly curved, compared to that of M. brevigaster curved at apical third; mesotibiae with apical spur exposed and visible in ventral view, compared to those of M. brevigaster covered by tuft and obscured; metatibiae slightly more straight; apical lobes of male sternite VIII narrowly hooked, compared to those of M. brevigaster broadly hooked.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A30FF8DFF5AFF196CF896BC.taxon	discussion	Comments. Morphostenophanes atavus was described from a single female from ‘ Yunnan’ without detailed location (‘ ohne nähere Angabe des Fundortes’ in original description). The male was then illustrated by Masumoto & Bečvář (2008), identified as M. atavus, however, no description or locality was given. Later, Gao & Ren (2009) provided detiled locality for this species, as well as a photograph of the female. In the present study, the author examined the holotype of M. atavus through photos. Specimens with similar appearances from three localities were studied, with one male from the eastern slope of Gaoligong Mountains, and two series containing both males and females from Yingjiang County and an adjacent area and Lincang City. According to the original description, the elytral aspect ratio of the M. atavus holotype female is 1.7, which is different from those from around Yingjiang County with an average ratio of 1.59 (n = 10) and maximum of 1.65. Moreover, the elytral striae of the M. atavus holotype are distinctly less depressed than those of the latter. The elytral aspect ratio of those from Lincang ranges from 1.7 to over 1.9, which matches that of the M. atavus holotype, but they are distinctly different by the latter having much more convex pronotum and elytra. The elytral aspect ratio of the male from the western slope of Gaoligong Mountains is 1.84, which is comprable to the M. atavus female. Moreover, this male has antennomeres less thickened anteriorly, elytral intervals less convex, and ventral apical tuft is interrupted, which are identical with those of the female holotype. Therefore, this male from the Gaoligong Mountains is identified as M. atavus. The female holotype possesses bulges at each side of the base of the pronotum, which are indistinct in the male. This is probably due to interspecific variation. Specimens from Yingjiang County and Lincang City are identified as belonging to M. brevigaster and M. lincangensis. On the label of M. atavus holotype, the author found ‘ Tientshun’ after the locality ‘ Yunnan’, which refers to ‘ Tengchong’, a city reaching the western slope of the Gaoligong Mountains. Such detailed locality was not mentioned in the original publication. The type locality associated with data of here examined male suggest a potential distribution of M. atavus along the southern Gaoligong Mountains, spreading over both slopes of mountains. The male specimen in Masumoto & Bečvář’s photograph (2008, Fig. 4) was in dorsal view, so the author measured its elytral aspect ratio, the result is 1.86. Moreover, its pronotum is constricted before the base, and its protibiae are only slightly curved, based on the revised diagnostic characters, it is undoubtedly a true M. atavus. Based on the known distribution, the female specimen mentioned by Gao & Ren (2009) from Yingjiang county is considered a misidentified of M. brevigaster.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A30FF8DFF5AFF196CF896BC.taxon	distribution	Distribution. (Map 1) CHINA: Yunnan.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A35FF8AFF5AFF196B4693D7.taxon	description	(Figs. 26 A – C; 31 B, G, L, Q; 32 B, J – L; 33 B, H)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A35FF8AFF5AFF196B4693D7.taxon	materials_examined	Type materials. CHINA: Yunnan: ♂ (Holotype, MYNU), Jinghong City, Mengyang Town, Yexianggu, 2016. xi. 12, native collector.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A35FF8AFF5AFF196B4693D7.taxon	diagnosis	Diagnosis. Body large, greenish, elongate and strongly convex. Antennae and legs reddish. Pronotum relatively narrow, slightly constricted before base. Elytra with rows of tubercles of varying sizes, each with purplish central portions. Metatibiae strongly curved. Aedeagus with narrowed and fusiform apex of parameres. Apical lobes of sternite VIII spatulate.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A35FF8AFF5AFF196B4693D7.taxon	description	Description. Male (Fig. 26 A – C). Greenish, antennae, mouthparts, and legs reddish brown; central parts of elytral tubercles purplish. Body elongate, length 25.1 mm, width 9.0 mm, strongly convex dorsad, noticeably constricted between pronotum and elytra. Head (Fig. 31 B, G) transversely subquadrate, densely and finely punctate, with outer margin strongly notched between genae and clypeus; clypeus transversely hexagonal, slightly convex in middle, gently bent downwards in front, with anterior margin nearly straight, weakly emarginate in middle, clypeal transverse impression marked; frontoclypeal suture deeply grooved, more deeply concaved in middle, becoming weaker laterally, widely U-shaped; genae strongly raised, depressed before eyes, strongly and roundly produced anterolaterally; frons broad, gradually sloping forwards; eyes transversely reniform, strongly convex laterally; inner ocular sulci shallowly grooved along inner margins, extending posterolaterally; tempora moderately convex, coarsely punctate. OI = 46.5. Antennae (Fig. 32 B) slender, reaching basal fourth of elytra, with antennomeres weakly thickened to apices; relative lengths of antennomeres: 0.83: 0.34: 1.29: 1.22: 1.38: 1.38: 1.36: 1.33: 1.22: 1.17: 1.21. Mentum (Fig. 31 G) quadrate, lateral margins straight; medial surface coarsely punctate, with several large pores bearing long setae, gradually rising from basal to apex, both sides of posterior half depressed. Pronotum (Fig. 31 L) quadrate, PW / PL = 1.00, widest in anterior third, anterior margin projecting anteriorly, marginal border marked; lateral margins rounded in anterior third, becoming straight posteriorly, lateral marginal borders thin, visible in dorsal view along anterior half; posterior margin rounded, weakly emarginate in middle, posterior marginal border markedly presented; anterior angles rounded; posterior angles obtuse, slightly projecting laterally; disc moderately convex, shagreened, sparsely and finely punctate. Scutellum widely triangular, extremely glossy. Elytra fusiform, widest in middle, EL / EW = 1.86; strongly convex, highest behind apical third; striae furrowed, sinuate; intervals intermittently expanded and convex, forming rounded or ovate tubercles, shagreened and weakly wrinkled, sparsely and finely punctate. Prosternum (Fig. 31 Q) weakly rugulose, finely and sparsely punctate; prosternal process strongly declivous; hypomeron rugulose, finely microsculptured. Metasternum densely and finely punctate, metaventral anterior process transversely wrinkled. Abdomen depressed, surface somewhat rough, shagreened, densely and finely punctate, with sternites III and IV sulcate at both sides. Legs slender. Protibiae (Fig. 32 J) moderately curved in apical fourth, apical 2 / 3 of inner margin sparsely pubescent; mesotibiae (Fig. 32 K) weakly curved in apical fourth, apical half of inner margin emarginate and pubescent; metatibiae (Fig. 32 L) strongly recurved, 2 / 3 of apical inner margin densely pubescent. Aedeagus (Fig. 33 B) elongate, curved in lateral view; parameres slender, 0.22 as long as total length, with narrowed and fusiform apex. Sternite VIII (Fig. 33 H) with spatulater apical lobes strongly produced backwards, constricted at each base. Female: Unknown. Comparative notes. Morphostenophanes bannaensis is the only species within the atavus - group possessing rows of purplish tubercles along eltra intervals. Such character state is also seen in M. linglong. However, M. linglong is clearly related to M. metallicus (see comparative notes of M. linglong). Large and elongate habitus, strongly curved metatibiae, narrowed apex of parameres, and similar apical lobes of sternite VIII suggest close relation between Morphostenophanes bannaensis, M. birmanicus, M. lincangensis and M. vietnamicus.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A35FF8AFF5AFF196B4693D7.taxon	distribution	Distribution. (Map 1) CHINA: Yunnan.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A35FF8AFF5AFF196B4693D7.taxon	etymology	Etymology. The new species is named after a simplified spelling of its type locality, Xishuangbanna.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A34FF88FF5AFF196ABA95C4.taxon	description	(Figs. 27 A – E; 31 C, H, M, R; 32 C, M – O; 33 C, I)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A34FF88FF5AFF196ABA95C4.taxon	materials_examined	Type material examined. Paratype of Promorphostenophanes koyamai labelled: THAILAND CHiang Mai Fang, 24. VIII. 1989 leg. K. Masumoto [handwritten in white label] // Paratype Promorphostenophanes komiyai [sic!] MA- SUMOTO [handwritten in pink label with underlines] (male, HNHM, examined through five photographs taken by Tamás Németh, sent by Ottó Merkl, Fig. 27 D). Additional material examined. CHINA: Yunnan: 1 ♂ (MYNU), Pu’er City, Lancang County, Zhutang Coun- try, Cizhuhe Village, Xiaolushan, 22 ° 45 ’ 31.78 ” N, 99 ° 42 ’ 23.72 ” E, 2180 m, 2017. i. 30, Hao Xu & Jian-Yue Qiu; 1 ♂ (MYNU, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C in CZDY), Pu’er City, Lancang County, Zhutang Country, Cizhuhe Village, Xiaolushan, 22 ° 45 ’ 31.78 ” N, 99 ° 42 ’ 23.72 ” E, 2180 m, larva 2017. i. 30, adult emerged 2017. v, Hao Xu & Jian-Yue Qiu. Comparative notes. M or phostenophanes birmanicus is closely related to M. lincangensis, M. bannaensis and M. vietnamicus, but can be easily distinguished from the related species by its unmodified pro- and mesotibiae, elytra with bronze metallic luster, elytral intervals evenly convex, elytral apex distinctly furcate.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A34FF88FF5AFF196ABA95C4.taxon	discussion	Comments. This species was originally treated as a subspecies of Promorphostenophanes atavus by Kaszab (1980) with a very brief description. Later, it was elevated to species rank (Masumoto & Bečvář 2008). In 1990, Ma- sumoto described a new species from north Thailand named P. koyamai, which was later placed as a junior synonym of M. birmanicus by Masumoto & Bečvář (2008). Thanks to this little ‘ episode’, M. birmanicus was redescribed in detail, and compared with M. atavus. In the present study, the author examined one male paratype of M. koyamai through photos taken from dorsal, lateral and ventral views. M. birmanicus is herewith recorded from China as a northernmost part of distribution. It was observed by the author that specimens from China and Thailand are almost identical.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A34FF88FF5AFF196ABA95C4.taxon	distribution	Distribution. (Map 1) CHINA: Yunnan (new country record), MYANMAR, THAILAND.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A37FF89FF5AFE5D6F6B93B7.taxon	description	(Figs. 28 A – D; 31 D, I, N, S; 32 D, P – R; 33 D, J)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A37FF89FF5AFE5D6F6B93B7.taxon	materials_examined	Type materials. CHINA: Yunnan: ♂ (Holotype, SNUC), Yingjiang County, Taiping Town, Shangbangwa, ca. 800 m, 2019. i, native collector. Paratypes (16 ♂♂, 19 ♀♀): CHINA: Yunnan: 1 ♀ (SNUC), 1 ♂ (MHBU), 1 ♂ (CJQY), Yingjiang County, Taiping Town, Mangyun, late vi, native collector; 1 ♀ (CJQY), 1 ♀ (CZDY, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C), Yingjiang County, Taiping Town, 2019. ii, native collector; 3 ♂ (CZDY), 2 ♂♂ (CQL), 1 ♀ (CJQY), Yinjiang County, Nongzhang Town, ca. 900 m, 2019. xii – 2020. i. 11, native col- lector; 1 ♂ (CZDY), Yingjiang County, Xima Town, Hulukou, 1200 m, 2018. vi – vii, Wei-Zong Yang; 1 ♂ (CJQY), Yingjiang County, Xima Town, Mountain near Menglai River, 1800 m, 2019, late vi, native collector; 6 ♀♀ (CBWX), Dehong Prefecture, Longchuan County, 2000 m, 2017. vi. 15 – 28, Wen-Xuan Bi; 1 ♀ (CBWX), Dehong Prefecture, Longchuan County, 2000 – 2080 m, 2018. vi. 4 – 8, Wen-Xuan Bi; 1 ♂ (CBWX), Tengchong City, Houqiao Town, Heinitang Village, 2000 m, 2017. ix. 7 – 10, Xiao-Dong Yang; 1 ♂, 2 ♀♀ (CZDY), 1 ♂, 2 ♀♀ (CBWX), Tengchong City, Houqiao Town, Heinitang Village, 1850 – 1950 m, 2018. ix. 4 – 6, Wen-Xuan Bi; 1 ♀ (CBWX), Tengchong City, Houqiao Town, Heinitang Village, 1850 m, 2018. ix. 13 as larva, adult emerged 2018. xii. 21, Wen-Xuan Bi; 1 ♀ (CBWX), Lushui County, Pianma Town, Gangfang, 2050 m, 2015. vi. 10, Wen-Xuan Bi; 2 ♂♂, 1 ♀ (CZDY), Tengchong County, Diantan Town, Mount Yunfeng, 25 ° 22 ’ 44.26 ” N, 98 ° 24 ’ 32.40 ” E, 2200 m, 2016. vii. 1, Zi-Chun Xiong; 1 ♂ (CYM), Yingjiang County, Taiping Town, Shangbangwa, ca. 800 m, 2019. i, native collector; 1 ♀ (CYM), Tengchong County, Diantan Town, Dadongshanhe, 25 ° 32 ’ 10.38 ” N, 98 ° 22 ’ 57.28 ” E, 2000 m, 2016. vii. 4, Mao Ye & Zi-Chun Xiong; 1 ♂ (CYM), Yingjiang County, Sudian Country, 2000 m, 2018. viii. 13, Ye Mao.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A37FF89FF5AFE5D6F6B93B7.taxon	diagnosis	Diagnosis. Body large and stout, grayish black. Antennae long, reaching apical third of elytra; Pronotum widest in middle; elytra striate, with ratio of length to width about 1.6 in male, and about 1.5 in female; all legs slender, metafemora reaching apex of sternite VII; metatibiae weakly sinuous; apical lobes of sternite VIII broadly hooked in lateral view.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A37FF89FF5AFE5D6F6B93B7.taxon	description	Description of male. Male (Fig. 28 A, C). Color greyish- black, antennae, claws reddish brown, shagreened. Body stout, length 20.0 – 24.5 mm, width 8.2 – 9.7 mm, strongly convex, noticeably constricted between pronotum and elytra. Head (Fig. 31 D, I) transversely subquadrate, sparsely and finely punctate, with outer margin distinctly notched between genae and clypeus; clypeus transversely hexagonal, slightly convex in middle, gently bent downwards in front, anterior margin bisinuate, clypeal transverse impression short, marked to absent; frontoclypeal suture deeply grooved, becoming weaker laterally, widely U-shaped; genae strongly raised, depressed before eyes, strongly and roundly produced anterolaterally; frons broad, anterior part gradually sloping forwards, vertexal lateral impressions shallow to absent; eyes transversely reniform, strongly convex laterally; inner ocular sulci shallowly depressed along inner margins, becoming broader posteriorly; tempora moderately convex. OI = 49.1 – 50. Antennae (Fig. 32 D) slender, reaching basal third of elytra, with antennomeres weakly thickened forwards, apices dilated; relative lengths of antennomeres: 0.77: 0.34: 1.28: 1.26: 1.38: 1.41: 1.34: 1.24: 1.21: 1.16: 1.24. Mentum (Fig. 31 I) inversely trapezoidal, lateral margins weakly rounded; medial surface finely punctate, with several large pores bearing long setae, gradually rising from base to apex, depressed along both sides. Pronotum (Fig. 31 N) quadrate, PW / PL = 1.12 – 1.14, widest slightlyanterior to the midpoint, anterior margin weakly rounded, anterior marginal border thin; lateral margins weakly rounded, lateral marginal borders visible in dorsal view along anterior half; posterior margin weakly rounded, emarginate in middle, posterior marginal border marked; anterior angles rounded, posterior angles obtuse; disc strongly convex, shagreened, finely and sparsely punctate. Scutellum widely triangular, glossy, finely punctate. Elytra fusiform, widest near middle, EL / EW = 1.61 – 1.64; strongly convex, highest in apical 2 / 5; with 9 continuous furrowed striae, 8 th ending at apical seventh of elytra, 1 st and 9 th, 2 th and 7 th convergingat ends, 4 th, 5 th successively joined 6 th at each end, 1 st and 4 th, 2 nd and 3 rd, 5 th and 6 th, and 7 th and 8 th convergingat fore-ends; intervals strongly convex, shagreened, sparsely and finely punctate. Prosternum (Fig. 31 S) shagreened, finely and sparsely punctate; prosternal process declivous, pointed at apex; hypomeron weakly rugulose, shagreened. Metasternum smooth, metaventral anterior process weakly wrinkled. Abdomen depressed, somewhat rough and finely punctate, sternites III and IV sulcate on both sides, sternite V depressed laterally. Legs slender. Protibiae (Fig. 32 P) weakly curved in apical third, apical 3 / 5 of inner margins pubescent; mesotibiae (Fig. 32 Q) weakly curved in apical third, apical half of inner margins pubescent; metatibiae (Fig. 32 R) weakly sinuous, apical 3 / 5 of inner margins pubescent, outer margins depressed before apices. Aedeagus (Fig. 33 D) elongate, curved in lateral view; parameres slender, 0.21 as long as total length, with ovoid apex. Sternite VIII (Fig. 33 J) with apical lobes broadly hooked in lateral view. Female (Fig. 28 B, D). Stouter than male, length 19.8 – 27.8 mm. OI = 49.0, PW / PL = 1.14 – 1.18; elytra more convex, highest in middle, EL / EW = 1.55 – 1.65; abdomen straight in lateral view. Ovipositor elongated, gradually narrowing posteriorly, apex acute. Comparative notes. M or phostenophanes brevigaster is similar to M. atavus. Detailed comparison and diagnosis is provided in the comparative notes of M. atavus.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A37FF89FF5AFE5D6F6B93B7.taxon	distribution	Distribution. (Map 1) CHINA: Yunnan.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A37FF89FF5AFE5D6F6B93B7.taxon	etymology	Etymology. The new species is named from a combination of the Latin stem, ‘ brevi’ and ‘ gaster’ referring to its short elytra.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A29FF97FF5AFF196A7E9670.taxon	description	(Figs. 29 A – D; 31 E, J, O, T, Y; 32 E, S – U; 33 E, K)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A29FF97FF5AFF196A7E9670.taxon	materials_examined	Type materials. CHINA: Yunnan: ♂ (Holotype, SNUC), Lincang City, Gengma County, Nangunhe Nature Reserve, 23 ° 38 ’ 38.83 ” N, 99 ° 23 ’ 13.84 ” E, 2006 m, 2017. vi. 26, Zi-Chun Xiong. Paratypes (12 ♂♂, 7 ♀♀): CHINA: Yunnan: 1 ♀ (SNUC), 1 ♀ (MHBU), 1 ♂, 1 ♀ (CYM), 3 ♂♂, 2 ♀♀ (CZDY, a female was preserved in 99.7 % ethanol at – 18 ° C), Lincang City, Gengma County, Nangunhe Nature Reserve, 2000 m, 2018. viii. 2, Zi-Chun Xiong; 4 ♂♂ (CZDY), Lincang City, Gengma County, Nangunhe Nature Reserve, 23 ° 38 ’ 38.83 ” N, 99 ° 23 ’ 13.84 ” E, 2006 m, 2017. vi. 26, Zi-Chun Xiong; 1 ♂, 1 ♀ (CZDY), Lincang City, Daxueshan Nature Reserve, 23 ° 59 ’ 28.40 ” N, 100 ° 18 ’ 46.21 ” E, 2018. xi. 7, Zi-Chun Xiong; 1 ♂ (CYM), Yongde County, Daxueshan Nature Reserve, near Yinchangjie Reservoir, 24 ° 11 ’ 53.58 ” N, 99 ° 39 ’ 14.17 ” E, 2447 m, in a hollow of Saurauia tree, 2015. vi. 18, Mao Ye; 1 ♂ (CBWX), Yongde County, Yalian Country, Damaidi Village, 2012. v. 6, Xiao-Dong Yang; 1 ♂, 1 ♀ (HNHM, identified by Ottó Merkl), Yunnan Province, SSE Shuangjiang Town, 23 ° 25 ’ 01 ’’ N / 99 ° 57 ’ 01 ’’ E, 23 ° 25 ’ 10 ’’ N / 99 ° 57 ’ 24 ’’ E, 2425 – 2570 m, 2011. vi. 20, Belousov, Kabak & Korolev.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A29FF97FF5AFF196A7E9670.taxon	diagnosis	Diagnosis. Body large and elongate, greyish-black, basal 3 / 4 of femora reddish. Antennae long, reaching apical 2 / 7 of elytra; elytra striate, with aspect ratio over 1.9 in male, and over 1.7 in female; protibiae thickened, strongly curved in apical third, metatibiae strongly recurved; leaf shaped apical lobes of sternite VIII.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A29FF97FF5AFF196A7E9670.taxon	description	Description of male. Male (Fig. 29 A, C). Color greyish-black, antennae, mouthparts, basal 3 / 4 of femora, apical portions of tibiae, and tarsi reddish brown, shagreened. Body slender, length 22.3 – 26.3 mm, width 7.6 – 8.6 mm, moderately convex, noticeably constricted between pronotum and elytra. Head (Fig. 31 E, J) transversely subquadrate, densely and markedly punctate, with outer margin strongly notched between genae and clypeus; clypeus transversely hexagonal, gently bent downwards in front, with anterior margin straight, clypeal transverse impression short, marked to absent; frontoclypeal suture deeply grooved, becoming weaker laterally, widely U-shaped; genae strongly raised, depressed before eyes, strongly and roundly produced anterolaterally; frons broad, anterior part gradually sloping forwards, vertexal lateral impressions narrow, distinct to vague, extending backward and curved oppositely, forming a U-shaped impressions; eyes transversely reniform, strongly convex laterally; inner ocular sulci shallowly depressed along inner margins, becoming broader posteriad; tempora slightly convex, punctures as those on frons. OI = 44.0 – 49.5. Antennae (Fig. 32 E) slender, reaching basal 2 / 7 of elytra, with antennomeres weakly thickened to apices; relative lengths of antennomeres: 0.92: 0.41: 1.53: 1.36: 1.53: 1.48: 1.36: 1.34: 1.26: 1.14: 1.22. Mentum (Fig. 31 J) inversely trapezoidal, lateral margins straight; medial surface coarsely punctate, with several large pores bearing long setae; gradually rising from base to apex, depressed along each side. Pronotum (Fig. 31 O) quadrate, PW / PL = 0.93 – 1.02, widest slightly before anterior third, anterior margin weakly rounded, anterior marginal border thin, vague; lateral margins weakly rounded, lateral marginal borders thin, visible in dorsal view along anterior half; posterior margin weakly rounded, emarginate in middle, posterior marginal border marked; anterior angles rounded, posterior angles obtuse; disc moderately convex, shagreened, finely and sparsely punctate. Scutellum widely triangular, glossy, finely punctate. Elytra elongate ovate, widest near middle, EL / EW = 1.90 – 1.99; strongly convex, highest near middle; with 9 continuous furrowed striae, 1 st branching, and 8 th ending in apical seventh of elytra; 1 st, 2 th, 3 rd, 7 th, and 8 th converged at ends, 4 th, 5 th, and 6 th converging in apical seventh, 2 nd and 3 rd converging at fore-ends; intervals strongly convex, glossy, coarsely wrinkled on apical seventh, sparsely and finely punctate. Prosternum (Fig. 31 T) shagreened, finely and sparsely punctate; prosternal process almost vertically declivous behind coxae, pointed at apex; hypomeron moderately rugulose, shagreened. Metasternum smooth, metaventral anterior process finely wrinkled. Abdomen depressed, somewhat wrinkled, densely and finely punctate, sternites III and IV sulcate on sides. Legs slender. Protibiae (Fig. 32 S) thick, strongly curved in apical third, more than apical half of inner margins emarginate and pubescent; mesotibiae (Fig. 32 T) weakly curved in apical fourth, apical half of inner margins emarginate and pubescent; metatibiae (Fig. 32 U) strongly curved in middle, apical 3 / 5 of inner margins pubescent, outer margins depressed before apices. Aedeagus (Fig. 33 E) elongated, curved in lateral view; parameres slender, 0.21 as long as the total length, with ovoid apex. Sternite VIII (Fig. 33 K) strongly sclerotized, with leaf shaped apical lobes in lateral view, constricted at each base, produced posteriorly, apex pointed. Female (Fig. 29 B, D). Wider than male, length 24.3 – 26.5 mm. OI = 44.6 – 47.1, PW / PL = 1.08 – 1.10. Elytra more convex, highest in middle, EL / EW = 1.68 – 1.90; abdomen straight in lateral view. Ovipositor (Fig. 31 Y) elongate, sclerotized and acinaciform, apex of gonocoxite acute. Comparative notes. Morphostenophanes lincangensis looks like a longer M. atavus. Its strongly elongate and less convex elytra, thicker tibiae, curved metatibiae, and shape of apical lobes of sternite VIII are shared with M. birmanicus, M. bannaensis and M. vietnamicus. M. lincangensis can be distinguished from M. birmanicus by the latter possessing more elongate antennae reaching apical third of elytra, greenish colored pronotum widest in anterior 2 / 5, much wider elytra with bronze metallic luster. M. lincangensis can be distinguished from M. bannaensis by the latter having tubercules of different sizes along each elytral interval. M. lincangensis can be distinguished from M. vietnamicus by the latter having bronzy color, male pro- and mesotibiae bugled in apical 2 / 5.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A29FF97FF5AFF196A7E9670.taxon	distribution	Distribution. (Map 1) CHINA: Yunnan.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A29FF97FF5AFF196A7E9670.taxon	etymology	Etymology. The new species is named after its type locality, Lincang City.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A28FF92FF5AF9606CD493CD.taxon	description	(Figs. 30 A – D; 31 U – X, Z; 32 F, V – X; 33 F, L)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A28FF92FF5AF9606CD493CD.taxon	materials_examined	Materials examined. (8 ♂♂, 1 ♀). VIETNAM: 1 ♂, 1 ♀ (MYNU, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C in CZDY), Lào Cai Province, Mount Sapa, 1600 m, native collector; 2 ♂ ♂ (CZDY), Yên Bái Province, Mù Cang Ch ải District, 1700 m, 2016. v – vii, native collector; 1 ♂ (HNHM, identified by Ottó Merkl), Lào Cai Province, 16 km W of Sa Pa, 1800 m, first Frontier base camp, 1998. iii. 18, under bark, L. Peregovits & T. Vásárhelyi; 4 ♂♂ (HNHM, identified by Ottó Merkl), Yên Bái Province, Mù Cang Ch ải District, Che Tao commune, Mù Cang Ch ải Species and Habitats Conservation Area, around Cong Troi (Gate to Heaven) Pass, 21.7686 ° N, 104.0442 ° E, 1940 m, upper montane evergreen forest, hand-collected from dead trees at night, 2019. ix. 24 – 29, Ottó Merkl & Phu Pham Van.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A28FF92FF5AF9606CD493CD.taxon	diagnosis	Diagnosis. Body large and elongate, aeneous, with metallic luster. Elytra striate, with aspect ratio of 1.8 in male, and of 1.7 in female; protibiae and mesotibiae each bulged in apical 2 / 5.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A28FF92FF5AF9606CD493CD.taxon	description	Redescription. Male (Fig. 30 A, C), aeneous, with metallic luster. Body elongate, length 22.3 – 23.9 mm, width 8.1 – 8.7 mm, strongly convex dorsad, noticeably constricted between pronotum and elytra. Head (Fig. 31 U, W) transversely subquadrate, densely and markedly punctate, with outer margin strongly notched between genae and clypeus; clypeus transversely hexagonal, gently bent downwards in front, convex before middle of the frontoclypeal suture, with anterior margin straight; frontoclypeal suture furrowed, becoming weaker laterad, widely U-shaped; genae strongly raised, depressed before eyes, strongly and roundly produced anterolaterally; frons broad, anterior part gradually sloping forwards; eyes transversely reniform, strongly convex laterally; inner ocular sulci shallowly grooved along inner margins; tempora slightly convex, punctures as those on frons. OI = 48.7. Antennae (Fig. 32 F) slender, reaching basal 2 / 7 of elytra, with antennomeres weakly thickened to apices; relative lengths of antennomeres: 0.77: 0.37: 1.28: 1.21: 1.34: 1.38: 1.33: 1.22: 1.21: 1.14: 1.33. Mentum (Fig. 31 W) rounded, medial surface weakly wrinked, coarsely punctate, with several large pores bearing long setae; gradually rising from base to apex, depressed along each side. Pronotum (Fig. 31 V) quadrate, PW / PL = 1.13, widest slightly before anterior third, anterior margin strongly projecting in middle, with marked marginal border; lateral margins nearly straight, roundly projecting laterally in anterior half, lateral marginal borders thin, visible in dorsal view along anterior half; posterior margin nearly straight, posterior marginal border marked, becoming thinner in middle; anterior angles broadly rounded, posterior angles obtuse; disc convex, finely and sparsely punctate. Scutellum widely triangular, glossy, finely punctate. Elytra fusiform, widest in middle, EL / EW = 1.8; strongly convex, highest before middle; with 9 continuous furrowed striae, 8 th ending at apical sixth of elytra; 1 st and 9 th; 2 th, 3 rd, and 7 th converging at ends, 4 th, 5 th successively joined 6 th at each end; intervals strongly convex, sparsely and finely punctate. Prosternum (Fig. 31 X) shagreened, finely and sparsely punctate; prosternal process declivous, truncate at apex; hypomeron weakly rugulose, shagreened. Metasternum glossy, finely wrinkled on metaventral anterior process. Abdomen weakly depressed, surface smooth in middle and weakly wrinkled on sides and anterior margin of each sternite, sparsely and finely punctate, sternites III and IV sulcate on sides. Legs slender. Protibiae (Fig. 32 V) thick, curved in apical fourth, apical 2 / 5 of inner margins concave, bulged in apical 2 / 5, pubescent; mesotibiae (Figs. 31 Z, 32 W) weakly curved in apical sixth, apical 2 / 5 of inner margins concave, bulged in apical 2 / 5, pubescent; metatibiae (Fig. 32 X) sinuous, apical half of inner margins pubescent, outer margins depressed before apices. Aedeagus (Fig. 33 F) elongate, curved in lateral view; parameres slender, 0.21 as long as total length, with ovoid apex. Sternite VIII (Fig. 33 L) with apical lobes constricted at base, and expanded posteriorly in lateral view, oblong, slightly curved. Female (Fig. 30 B, D). Less shiny and wider than male, length 26.5 mm. OI = 48.7, PW / PL = 1.24; elytra more convex, highest near middle, EL / EW = 1.68; abdomen straight in lateral view. Pro- and mesotibiae without bulges, metatibiae without pubescent sinuosity. Ovipositor elongate, gradually narrowing posteriorly, apex of gonocoxite acute.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A28FF92FF5AF9606CD493CD.taxon	discussion	Comments. This species is originally treated as a subspecies of Promorphostenophanes atavus by Kaszab (1980). Later, it was elevated to species rank by Masumoto & Bečvář (2008). The unique bulges on inner sides of male pro- and mesotibiae clearly separate it from every other congeners and verify its species-level status. The known localities of M. vietnamicus are extremely close to Jinping County, southeastern Yunnan, where this species may be found in the future, and thus be recorded in China.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A28FF92FF5AF9606CD493CD.taxon	distribution	Distribution. (Map 1). North VIETNAM.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A2CFF93FF5AFA1F6DE593BB.taxon	discussion	Comments. Masumoto and Bečvář (2008) mentioned M. elegantulus with ‘ the body of Promorphostenphanes in outline, but the elytra bear rows of rounded or ovate impressions as in Morphostenophanes. ’, therefore, Promophostenophanes was synonymized with Morphostenophanes. The author found species within the atavus - and elegantulus - group sharing not only general habitus but also relatively smooth sternites III and IV without distinct impressions and similar elongate ovipositor. These two species groups only differ in elytral sculpturing. Furthermore, some individuals of M. gaoligongensis, M. furvus, M. furvus weishanus and M. sinicus have disconnected and not encircled short segments of striae. The author found a male M. furvus possessing almost complete elytra striae 3 rd and 4 th, but they are still interrupted by elevated, cross-border intervals (marked by yellow arrows in Fig. 40 F). This seems to show a transtional form between continuous elytral striae and centrally convex impressions. In this sense, atavus - and elegantulus - group are potentially closely related.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A2FFF90FF5AFF196C14967F.taxon	description	(Fig. 34 G, H)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A2FFF90FF5AFF196C14967F.taxon	materials_examined	Type material examined. Holotype labelled: ‘ Fang Chiang Mai Thailand, 25. VII. I 994 leg. Native // Coll. Masumoto 2001 // Holotype Morphostenophanes elegantulus MAS. & BEČ’ (male, NSMT, Fig. 34 G), examined through two photograph provided by Kimio Masumoto. Comparative notes. Morphostenophanes elegantulus can be distinguished from other congeners of elegantulus - group by the major part of its elytra and legs colored dark red brown and strongly shortened female elytra.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A2FFF90FF5AFF196C14967F.taxon	distribution	Distribution. (Map 1) THAILAND.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A2FFF9EFF5AF9756A1A933C.taxon	description	(Figs. 34 A – F; 35 A – D; 39 A, E, I, M; 40 A, H – L; 41 A, E, L)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A2FFF9EFF5AF9756A1A933C.taxon	materials_examined	Type materials. CHINA: Yunnan: ♂ (Holotype, SNUC, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C in CZDY), Mount Gaoligong, Baihualing, Jinchanghe, 25 ° 18 ’ 40.68 ’’ N, 98 ° 47 ’ 26.45 ’’ E, 1700 m, 2016. vi. 20, Zhi-Wei Dong. Paratypes (17 ♂♂, 12 ♀♀): CHINA: Yunnan: 1 ♀ (SNUC), Mount Gaoligong, Baihualing, 2100 m, 2012. v. 5, Wen-Xuan Bi; 1 ♂ (CZDY), 1 ♂ (CQL), Mount Gaoligong, Baihualing, Jin- changhe, 25 ° 18 ’ 40.68 ’’ N, 98 ° 47 ’ 26.45 ’’ E, 1700 m, 2016. vi. 20, Zhi-Wei Dong; 1 ♂ (CZDY, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C), Mount Gaoligong, Baihualing, near Ertaipo, 25 ° 17 ’ 57.56 ’’ N, 98 ° 47 ’ 7.94 ’’ E’, 2057 m, 2017. vi. 28, Zhong-Liang Jiang; 1 ♀ (CZJZ), Mount Gaoligong, Baihualing, 25 ° 18 ’ 28.47 ” N, 98 ° 47 ’ 37.80 ” E, 1588 m, 2019. vii. 25, Jia-Zhi Zhang; 4 ♂♂, 1 ♀ (MYNU), Mount Gaoligong, Baihualing, 1900 m, 2015. ii. 23, Hao Xu & Jian-Yue Qiu; 1 ♀ (MYNU), Mount Gaoligong, Baihualing, 1900 m, 2016. ii. 13, Hao Xu & Jian-Yue Qiu; 1 ♂ (MHBU), Mount Gaoligong, Baihualing, 1900 m, 2018. ii. 18, Hao Xu & Jian-Yue Qiu; 1 ♂, 1 ♀ (CZDY, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C), Tengchong County, Gaoligong N. R., Xiaodifang Village, 24 ° 51 ’ 15.43 ” N, 98 ° 45 ’ 10. 73 ” E, 2172 m, 2019. ix. 19 – 20, Liang Ding; 1 ♂, 1 ♀ (SNUC), 3 ♂♂, 4 ♀♀ (CZDY, a female was preserved in 99.7 % ethanol at – 18 ° C), Baoshan City, Tengchong County, Zha- oyun Village, ca. 25 ° 8 ’ 56.60 ’’ N, 98 ° 30 ’ 21.67 ’’ E, 2350 m, 2018. ii, native collector; 2 ♂♂, 1 ♀ (CZDY), Tengchong County, Zhaoyun Village, ca. 25 ° 8 ’ 56.60 ’’ N, 98 ° 30 ’ 21.67 ’’ E, 2350 m, 2019. i, native collector; 1 ♂, 1 ♀ (CZJZ), 1 ♂ (CZDY), Yingjiang County, Taiping Town, 1200 m, 2019. v. 27, native collector.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A2FFF9EFF5AF9756A1A933C.taxon	diagnosis	Diagnosis. Body medium sized, bronze, sometimes greenish, weaklyshiny. Elytra widest near middle, with rows of encircled segments of striae, some interrupted or straight, each encircled area convex. Aedeagus with apex of parameres not curved ventrally.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A2FFF9EFF5AF9756A1A933C.taxon	description	Description. Male (Fig. 34 A, B, D, E; 35 A, C). Bronze, sometimes greenish, shagreened; antennae dark brown, claws reddish brown. Body stout, length 19.7 – 21.4 mm, width 7.4 – 7.7 mm, strongly convex, noticeably constricted between pronotum and elytra. Head (Fig. 39 A, E) transversely subquadrate, sparsely and finely punctate, with outer margin distinctly notched between genae and clypeus; clypeus transversely heptagonal, gently bent downwards in front, with anterior margin nearly straight, slightly emarginate in middle, clypeal transverse impression short, marked to absent; frontoclypeal suture depressed, becoming weaker laterad, widely U-shaped; genae strongly raised, depressed before eyes, strongly and roundly produced anterolaterally; frons broad, smooth to vaguely wrinkled, anterior part gradually sloping forwards; eyes transversely reniform, strongly convex laterally; inner ocular sulci finely depressed; tempora moderately convex. OI = 51.0 – 55.1. Antennae (Fig. 40 A) slender, slightly surpassing basal third of elytra, with antennomeres weakly thickened to apices; relative lengths of antennomeres: 0.68: 0.37: 1.11: 1.19: 1.24: 1.22: 1.22: 1.17: 1.14: 1.04: 1.14. Mentum (Fig. 39 E) subcircular, lateral margins rounded, seamlessly connected posteriorly to the emarginate posterior margin; medial surface sparsely and coarsely punctate, with several large pores bearing long setae, gradually rising anteriorly, depressed along both sides. Pronotum (Fig. 39 I) quadrate, PW / PL = 1.09 – 1.19, widest near anterior third; anterior margin nearly straight, slightly projecting in middle, anterior marginal border marked; lateral margins weakly curved, anterior half slightly projecting laterally, lateral sulci thin, visible in dorsal view along anterior third to half; posterior margin weakly rounded or nearly straight, emarginate in middle, posterior marginal border marked; anterior angles rounded, posterior angles obtuse; disc strongly convex in middle, depressed before posterior margin, shagreened, finely and sparsely punctate. Scutellum (Fig. 40 H, I) widely triangular, glossy to shagreened, finely punctate. Elytra fusiform, widest near middle, EL / EW = 1.75 – 1.85; strongly convex, highest near middle; with rows of round and oval segments of striae, some interrupted or straight, encircled areas convex; intervals strongly convex, surface smooth, sparsely and finely punctate. Prosternum (Fig. 39 M) shagreened, finely and sparsely punctate; prosternal process produced posterioly or declivous, apex truncate; hypomeron weakly rugulose, shagreened. Metasternum glossy, metaventral anterior process weakly wrinkled. Abdomen depressed, surface somewhat rough, densely and finely punctate, with sternites III and IV sulcate in both sides. Legs slender. Protibiae (Fig. 40 J) curved in apical third, apical 3 / 5 of inner margins pubescent; mesotibiae (Fig. 40 K) weakly curved in apical third, apical half of inner margins pubescent; metatibiae (Fig. 40 L) weakly sinuous, apical 3 / 5 of inner margins emarginate and pubescent, outer margins depressed before apices. Aedeagus (Fig. 41 A) elongate, curved in lateral view; parameres slender, straightly producing in lateral view, 0.20 as long as total length, with apex obtusely lance-shaped. Sternite VIII (Fig. 41 E) with apical lobes rectangular in lateral view, superior margin straight, exterior margins straight, inferior margins rounded, cornered at lowest point. Female (Fig. 34 C, F; 35 B, D). Stouter than male, length 20.0 – 23.7 mm. OI = 55.0 – 56.3, PW / PL = 1.14 – 1.23; elytra more convex, EL / EW = 1.57 – 1.75, highest in middle; abdomen straight in lateral view. Ovipositor elongate, gradually narrowing apically, apex of gonocoxite acute or truncate. Variability. Both males and females of this species vary in habitus and body color. The male holotype (Fig. 34 A, D) and one female paratype (Fig. 34 C, F) have relatively stout bodies, while other male paratypes (as shown in Fig. 34 B, E) and another paratype female shown less bloated bodies. Some individuals have greener body, even dark green (Fig. 35 A – D). The male holotype has a pair of impressions on scutellum (Fig. 40 H), while other specimens lack such impressions (as shown in Fig. 40 I). One male paratype (Fig. 34 B, E) has more coarsely microsculptured pronotal disc and scutellum, and less curved protibiae. One female paratype has trcunted apex of gonocoxite (Fig. 41 L), with a smoothly curved apical edge, which seems not caused by damage or abrasion. Another female has a normal, acute gonocoxite as those in all other congeners within the elegantulus - and atavus - group. Comparative notes. Morphostenophanes gaoligongensis resembles M. sinicus, but can be clearly distinguished from the latter by its straight produced apex of parameres in lateral view, compared to that of M. sinicus, which isventrally bent. A preliminary identification of M. gaoligongensis is based on its more glossy body, antennomere XI less than 3 x as long as wide comprared to that of M. sinicus more than 3 x as long as wide, and stouter male elytra with aspect ratios ranging from 1.75 – 1.85, compared to that of M. sinicus in male greater than 1.9.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A2FFF9EFF5AF9756A1A933C.taxon	distribution	Distribution. (Map 1) CHINA: Yunnan.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A2FFF9EFF5AF9756A1A933C.taxon	etymology	Etymology. The new species is named after its type locality, Gaoligong Mountains.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A20FF9CFF5AFF196A6592D4.taxon	description	(Figs. 36 A – E; 39 B, F, J, N; 40 B, G, M – O; 41 B, F)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A20FF9CFF5AFF196A6592D4.taxon	materials_examined	Type materials. CHINA: Yunnan: ♂ (Holotype, SNUC), Lincang City, Manwan Town, Shuibatou Village, 1900 – 2200 m, 2016. i – iii, Zi-Chun Xiong. Paratypes (37 ♂♂, 22 ♀♀): CHINA: Yunnan: 1 ♀ (SNUC), 3 ♂♂, 3 ♀♀ (MHBU), 10 ♂♂, 6 ♀♀ (CZDY, a sample of eggs was preserved in 99.7 % ethanol at – 18 ° C), Lincang City, Manwan Town, Shuibatou Village, 1900 – 2200 m, 2016. i – iii, Zi-Chun Xiong; 2 ♂♂ (CBNX), Lincang City, Manwan Town, Shuibatou Village, 1900 – 2200 m, 2016. i – iii, Zi-Chun Xiong; 3 ♂♂ (CYM), Lincang City, Manwan Town, Shuiba- tou Village, 1900 – 2200 m, 2017. viii. 10, Zi-Chun Xiong; 3 ♂♂ (CZDY), Lincang City, Manwan Town, Shuibatou Village, 1900 – 2200 m, 2018. iii, Zi-Chun Xiong; 2 ♂♂, 4 ♀♀ (MYNU), Lincang City, Manwan Town, 2016. ii. 22, Zi-Chun Xiong; 1 ♂, 1 ♀ (MYNU), Lincang City, Manwan Town, Manjiu, 2017. i. 15, Zi-Chun Xiong; 2 ♂♂ (CZDY, a male was preserved in 99.7 % ethanol at – 18 ° C), Lincang City, Daxueshan Nature Reserve, 23 ° 59 ’ 28.40 ” N, 100 ° 18 ’ 46.21 ” E, 2018. xi. 7, Zi-Chun Xiong; 3 ♂♂, 2 ♀♀ (CZDY), Lincang City, Maolan Town, Mount Dabin, 24 ° 40 ’ 21.58 ” N, 100 ° 17 ’ 58.94 ” E, 2018. ii, Zi-Chun Xiong; 1 ♂ (CBWX), Lincang City, Bangdong Country, Mount Wulao, 2600 m, Y. - H. Li; 1 ♂, 4 ♀♀ (CBWX, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C in CZDY), Mount Wuliang, De’an Village, Sheyaoqing, 2300 – 2450 m, 2017. vii. 10 – 11, Wen-Xuan Bi; 3 ♂♂, 1 ♀ (CZDY, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C), Mount Ailao, Qianjiazhai, near 24 ° 17 ’ 1.38 ” N, 101 ° 15 ’ 27.57 ” E, 2225 – 2694 m, 2011. viii, native collector; 3 ♂♂ (CZDY, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C), Yuxi City, Yuanjiang County, Yangchajie Country, ca. 2300 m, 2017. v. 1, Min-Fei Hou.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A20FF9CFF5AFF196A6592D4.taxon	diagnosis	Diagnosis. Body medium-sized to large, slightly greenish. Antennae long, reaching apical third of elytra. Pronotal lateral margins rounded, lateral marginal borders visible in dorsal view along anterior half. Elytra with rows of round or oval segments of striae, some interrupted or straightly presented, with encircled areas convex, elytral aspect ratio greater than 1.9 in male, and greater than 1.6 in female. Apex of parameres bent ventrally in lateral view.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A20FF9CFF5AFF196A6592D4.taxon	description	Description. Male (Fig. 36 A, B, D). Color bronze, with greenish sheen, shagreened; antennae dark brown, claws reddish brown. Body elongate, length 20.0 – 24.4 mm, width 7.0 – 9.5 mm, strongly convex, noticeably constricted between pronotum and elytra. Head (Fig. 39 B, F) transversely subquadrate, sparsely and finely punctate, with outer margin moderately notched between genae and clypeus; clypeus transversely hexagonal, gently bent downwards in front, convex before middle of frontoclypeal suture, with anterior margin nearly straight, clypeal transverse impression marked to absent; frontoclypeal suture moderately grooved, becoming weaker laterally, widely U-shaped; genae strongly raised, depressed before eyes, roundly produced anterolaterally; frons broad, smooth, anterior part gradually sloping forwards; eyes transversely reniform, strongly convex laterally; inner ocular sulci grooved along inner margins; tempora moderately convex. OI = 46.7 – 50.7. Antennae (Fig. 40 B) slender, reaching basal third of elytra, with antennomeres weakly thickened to apices; relative lengths of antennomeres: 0.75: 0.36: 1.12: 1.16: 1.24: 1.24: 1.26: 1.17: 1.11: 1.07: 1.22. Mentum (Fig. 39 F) subcircular, lateral margins rounded, seamlessly connected posteriorly to the emarginate posterior margin; medial surface sparsely and coarsely punctate, with several large pores bearing long setae, gradually rising anteriorly, depressed along both sides. Pronotum (Fig. 39 J) barrel-shaped, PW / PL = 1.07 – 1.12, widest near middle, anterior margin nearly straight, slightly projecting in middle, anterior marginal border marked; lateral margins rounded, lateral marginal borders thin, slightly sinuate, weakly projecting laterally at widest points, visible in dorsal view along anterior half; posterior margin weakly rounded, emarginate in middle, posterior marginal border marked; anterior angles rounded, posterior angles obtuse; disc strongly convex in middle, shagreened, finely and sparsely punctate. Scutellum widely triangular, glossy, finely punctate. Elytra elongate oval, widest near middle, EL / EW = 1.92 – 2.01; strongly convex, highest near middle; with rows of round or oval segments of striae, some interrupted or straight, encircled areas strongly or moderately convex; intervals shagreened and smooth, sparsely and finely punctate. Prosternum (Fig. 39 N) shagreened, finely and sparsely punctate; prosternal process declivous, apex truncate; hypomeron rugulose, shagreened. Metasternum finely wrinkled. Abdomen depressed, surface somewhat rough, densely and finely punctate, with sternites III, IV and anterior 3 / 5 of V sulcate along both sides. Legs slender. Protibiae (Fig. 40 M) curved in apical third, apical 3 / 5 of inner margins pubescent, outer margin emarginate near middle; mesotibiae (Fig. 40 N) weakly curved in apical third, apical 3 / 5 of inner margins pubescent; metatibiae (Fig. 40 O) weakly curved, apical 3 / 5 of inner margins emarginate and pubescent, outer margins depressed before apices. Aedeagus (Fig. 41 B) elongate, curved in lateral view; parameres slender, 0.22 as long as total length, with apex obtusely lance-shaped, weakly bent ventrally in lateral view. Sternite VIII (Fig. 41 F) with apical lobes ovate in lateral view, superior margins projecting dorsad, exterior margins and inferior margins rounded. Female (Fig. 36 C, E). Wider than male, length 20.5 – 23.8 mm. Eyes closer, OI = 42.9; PW / PL = 1.15; elytra stouter, EL / EW = 1.61 – 1.76, more convex, highest in middle; abdomen straight in lateral view. Ovipositor elongate, gradually narrowing apically, apex of gonocoxite acute. Variability. One male from Yuanjiang County has darker and extremely slender body (Fig. 36 B), another Yuanjiang male has pitch black body color almost without metallic luster. One female from Wuliang Mountain shows ridged elytral intervals (Fig. 40 G). Comparative notes. Morphostenophanes sinicus is similar to M. gaoligongensis. Detailed comparison and diagnosis is provided in the comprative notes of M. gaoligongensis.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A20FF9CFF5AFF196A6592D4.taxon	distribution	Distribution. (Map 1) CHINA: Yunnan.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A20FF9CFF5AFF196A6592D4.taxon	etymology	Etymology. The new species is named after the country of its type locality.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A23FF9BFF5AF96D6B5F950C.taxon	description	(Figs. 37 A – D; 39 C, G, K, O; 40 C, E, F, P – R; 41 C, G)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A23FF9BFF5AF96D6B5F950C.taxon	materials_examined	Type materials. CHINA: Yunnan: ♂ (Holotype, SNUC), Diqing Prefecture, Weixi County, above Puduoluo, 27 ° 7 ’ 58.27 ’’ N, 99 ° 6 ’ 26.39 ’’ E, ca. 2900 m, 2017. xii – 2018. ii, native collector. Paratypes (21 ♂♂, 19 ♀♀): CHINA: Yunnan: 1 ♀ (SNUC), 2 ♂♂, 2 ♀♀ (MHBU), 7 ♂♂, 2 ♀♀ (CZDY, one female was disassembled for generic char- acters examined, one male was preserved in 99.7 % ethanol at – 18 ° C), Diqing Prefecture, Weixi County, above Puduoluo, 27 ° 7 ’ 58.27 ’’ N, 99 ° 6 ’ 26.39 ’’ E, ca. 2900 m, 2017. xii – 2018. ii, native collector; 1 ♂, 1 ♀ (CQL), Weixi County, Zhonglu Country, Lagaluo Village, 2018. vii. 10, native collector; 2 ♂♂, 8 ♀♀, Nujiang Prefrecture, Gong- shan County, Puladi Country, Xiqieluo, 2512 m, 2019. ii, native collector; 9 ♂♂, 5 ♀♀ (CZDY), Deqin County, Yunling Country, Yongzhi Village, 2019. iv, native collector.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A23FF9BFF5AF96D6B5F950C.taxon	diagnosis	Diagnosis. Body medium-sized to large, greyish-black, weakly shiny. Pronotum barrel-shaped. Elytra elongate oval, widest near middle, with aspect ratio about 1.83, with rows of round or oval segments of striae, some interrupted, each encircled area strongly or moderately convex. Female much wider than male, with aspect ratio of elytra about 1.73 – 1.84. Protibiae markedly thickened apically; inner margins of pro- and mesotibiae of male pubescent along apical half.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A23FF9BFF5AF96D6B5F950C.taxon	description	Description. Male (Fig. 37 A, C). Color black, weakly shiny; antennae and claws reddish brown. Body elongate, length 20.5 – 22.8 mm, width 7.2 – 8.5 mm, strongly convex, markedly constricted between pronotum and elytra. Head (Fig. 39 C, G) transversely subquadrate, cephalic punctures small, sparsely scattered on frons, and becoming slightly denser on clypeus and genae; outer margin distinctly notched between genae and clypeus; clypeus transversely hexagonal, width-length ratio about 1.75, gently bent downwards in front, convex before middle of the frontoclypeal suture, anterior margin weakly emarginate in middle, clypeal transverse impression short, marked to absent; frontoclypeal suture deeply grooved, becoming weaker laterad, widely U-shaped; genae strongly raised, depressed before eyes, roundly produced anterolaterally; frons broad, vaguely wrinkled, anterior part gradually sloping forwards; eyes transversely reniform, strongly convex laterally; inner ocular sulci finely depressed along inner margins; tempora moderately convex. OI = 49.5 – 51.3. Antennae (Fig. 40 C) slender, reaching basal third of elytra, with antennomeres weakly thickened to apices; relative lengths of antennomeres: 0.66: 0.34: 1.12: 1.19: 1.24: 1.21: 1.19: 1.14: 1.09: 1.00: 1.11. Mentum (Fig. 39 G) square, lateral margins straight or weakly rounded; medial surface sparsely and coarsely punctate, with several large pores bearing long setae; gradually rising anteriorly, depressed along both sides. Pronotum (Fig. 39 K) barrel-shaped, PW / PL = 1.12 – 1.21, widest near middle, anterior margin nearly straight, slightly projecting in middle, with thin marginal border, interrupted in middle; lateral margins weakly rounded, lateral marginal borders thin, visible in dorsal view along anterior half; posterior margin emarginate in middle, posterior marginal border markedly presented; anterior angles rounded; posterior angles obtuse; disc strongly convex, shagreened, finely and sparsely punctate. Scutellum widely triangular, glossy, finely punctate. Elytra elongate oval, widest near middle, EL / EW = 1.80 – 1.86; strongly convex, highest near basal 2 / 5; with rows of round or oval segments of striae, some interrupted or straight, encircled areas strongly or moderately convex; intervals shagreened and smooth, sparsely and finely punctate. Prosternum (Fig. 39 O) shagreened, finely and sparsely punctate; prosternal process declivous, apex truncate; hypomeron rugulose, shagreened. Metasternum finely wrinkled. Abdomen depressed, surface smooth, densely and finely punctate, with sternites III, IV and anterior 3 / 5 of V sulcate at both sides. Legs slender. Protibiae (Fig. 40 P) curved in apical third, apical half of inner margins pubescent; mesotibiae (Fig. 40 Q) curved in apical third, apical half of inner margins pubescent, outer margins weakly depressed before apices; metatibiae (Fig. 40 R) weakly sinuous, apical 3 / 5 of inner margins emarginate and pubescent, outer margins depressed before apices. Aedeagus (Fig. 41 C) elongate, curved in lateral view; parameres slender, ridged along dorsal midline, 0.21 as long as total length, with flabellate apex. Sternite VIII (Fig. 41 G) with apical lobes each ovate in lateral view, superior margins projecting dorsad, exterior margins and inferior margins rounded. Female (Fig. 37 B, D). Wider than male, length 20.0 – 23.1 mm. Distance between eyes wider, OI = 56.5; PW / PL = 1.16 – 1.24, elytra slightly wider than those of males, EL / EW = 1.73 – 1.84; much more convex, highest behind middle; abdomen straight in lateral view. Ovipositor elongate, gradually narrowing apically, apex of gonocoxite acute. Comparative notes. Within the elegantulus - group, Morphostenophanes furvus is distinguished from M. gaoligongensis, and M. sinicus by its evenly black body without metallic luster, shorter antennae and legs, and more elongate basal piece. It is distinguished from M. elegantulus by its pronotum and elytra without reddish areas.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A23FF9BFF5AF96D6B5F950C.taxon	distribution	Distribution. (Map 1). CHINA: Yunnan.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A23FF9BFF5AF96D6B5F950C.taxon	etymology	Etymology. The new species is named from a Latin epithet, ‘ furvus’ referring to its dark body color.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A24FF98FF5AF9F96B0E925A.taxon	description	(Figs. 38 A – D; 39 D, H, L, P; 40 D, S – U; 41 D, H, I, J, K)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A24FF98FF5AF9F96B0E925A.taxon	materials_examined	Type materials. CHINA: Yunnan: ♂ (Holotype, SNUC), Dali Prefecture, Weishan County, Mount Weibao, 2450 – 2580 m, 2017. vii. 7, Wen-Xuan Bi. Paratypes (9 ♂♂, 14 ♀♀): CHINA: Yunnan: 1 ♀ (SNUC), 1 ♂, 1 ♀ (CZDY), 3 ♂♂, 9 ♀♀ (CBWX, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C in CZDY), Dali Prefecture, Weishan County, Mount Weibao, 2450 – 2580 m, 2017. vii. 7, Wen-Xuan Bi; 5 ♂♂, 2 ♀♀ (CBWX), Dali Prefecture, Weishan County, Mount Weibao, 2400 – 2500 m, 2015. viii. 16 – 18, Wen-Xuan Bi; 1 ♀ (CZDY, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C), Dali Prefecture, Weishan County, Ma’anshan Country, 25 ° 19 ’ 59.67 ” N, 100 ° 3 ’ 13.91 ” E, 2019. I, Zhi-Wei Dong. Diagnostic description. Male (Fig. 38 A, C). Greyish-black, shagreened; antennae and claws reddish brown. Body elongate, length 20.6 – 23.2 mm, width 7.5 – 8.3 mm, strongly convex. Head (Fig. 39 E, J) with slightly smaller eyes, frontoclypeal suture less deeply U-shaped, width-length ratio of clypeus about 2.00; OI = 51.6 – 51.9. Antennae (Fig. 40 D) slender, reaching basal third of elytra, with antennomeres weakly thickened to apices; relative lengths of antennomeres: 0.68: 0.34: 1.19: 1.10: 1.24: 1.28: 1.28: 1.19: 1.09: 1.02: 1.09. Pronotum (Fig. 39 L) barrel-shaped, usually broader, sometimes nearly rounded, PW / PL = PW / PL = 1.21 – 1.23, lateral marginal borders thiner, visible in dorsal view along anterior third. Elytra elongate oval, EL / EW = 1.83 – 1.89. Protibiae (Fig. 40 S) more markedly curved, pubescent of inner margins more developed and denser. Female (Fig. 38 B, D) distinctly stouter, length 19.4 – 23.3 mm. Distance between eyes wider, OI = 53.8 – 54.3, PW / PL = 1.18 – 1.26; elytra more convex, EL / EW = 1.60 – 1.64.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A24FF98FF5AF9F96B0E925A.taxon	description	Variation. Some males have more roundly produced pronotal lateral margins; male spicula gastrale occasionally curving dorsally (Fig. 41 J). Comparative notes. Morphostenophanes furvus weishanus can be easily distinguished from the nominate subspecies by its much broader and more rounded male pronotum, and stouter female elytra.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A24FF98FF5AF9F96B0E925A.taxon	distribution	Distribution. (Map 1) CHINA: Yunnan.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A24FF98FF5AF9F96B0E925A.taxon	etymology	Etymology. The new subspecies is named after the locality the type series collected from, Weishan County.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A27FFE7FF5AF8C86F6A90CF.taxon	description	(Figs. 42 A – D; 47 A, F, K, P; 48 A, J – L; 49 A, E)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A27FFE7FF5AF8C86F6A90CF.taxon	materials_examined	Type material. CHINA Yunnan: ♂ (Holotype, SNUC), Diqing Prefecture, Shangri-La City, Hutiaoxia, 27 ° 12 ’ 31.82 ” N, 100 ° 06 ’ 35.75 ” E, 2312 m, 2010. vii. 2, Xiao-Bin Song.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A27FFE7FF5AF8C86F6A90CF.taxon	diagnosis	Diagnosis. Small, stubby and convex species. Head and pronotum densely and coarsely punctate. Pronotum extremely wide, posterior angles vertical, markedly projecting laterally. Elytra with irregularly scattered short strial punctures, intervals convex, with sharp punctures. Inner margin of protibiae gently projected in middle, outer margin of protibiae each concave slightly behind apex.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A27FFE7FF5AF8C86F6A90CF.taxon	description	Description. Male (Fig. 42 A – D). Uniformly reddish brown (originally with green metallic luster shown in Fig. 42 D). Body stubby, length 14.4 mm, width 5.5 mm, strongly convex, noticeably constricted between pronotum and elytra. Head (Fig. 47 A, F) transversely subquadrate, densely and coarsely punctate; clypeus transversely hexagonal, gently bent downwards in front, anterior margin slightly emarginate, clypeal transverse impression short, marked; frontoclypeal suture grooved, widely U-shaped; genae gently raised, depressed before eyes, roundly produced anterolaterally; frons rather broad, gradually sloping forwards; eyes transversely reniform, moderately convex laterally; inner ocular sulci deeply grooved along inner margins, becoming shallower and broader posteriorly; tempora moderately convex, coarsely punctate. OI = 59.8. Antennae (Fig. 48 A) slender, reaching basal fourth of elytra, with antennomeres only weakly thickened to apices; relative lengths of antennomeres: 0.49: 0.24: 0.66: 0.60: 0.65: 0.70: 0.70: 0.65: 0.63: 0.58: 0.75. Mentum (Fig. 47 F) square, lateral margins straight, gradually rising anteriorly, depressed along both sides; surface smooth along midline, wrinkled and coarsely punctate on depressed area, with several large pores bearing long setae. Pronotum (Fig. 47 K) barrel-shaped, PW / PL = 1.3, widest slightly before middle; anterior margin straight, anterior marginal border obsecured, interrupted in middle; lateral margins roundly curved, lateral marginal borders thin; posterior margin gently rounded, posterior marginal border faintly presented; anterior angles rounded, slightly produced anteriorly; posterior angles vertical, slightly projecting laterally; disc moderately convex, presenting a thin longitudinal impression along midline, flank depressed on each side slightly behind the middle, coarsely punctate. Scutellum widely triangular, coarsely punctate. Elytra elongate oval, widest near middle, EL / EW = 1.65; strongly convex, highest slightly before middle; with irregularly distributed short strial punctures; intervals moderately convex, weakly wrinkled and evenly scattered with sharp punctures. Prosternum (Fig. 47 P) densely rugulose, finely and sparsely punctate; prosternal process strongly declivous, apex pointed, strongly dilated in ventral view; hypomeron strongly rugulose, sparsely and finely punctate. Metasternum finely punctate, metaventral anterior process short, transversely wrinkled. Abdomen slightly depressed, surface coarsely punctate, sternites III – V wrinkled. Legs slender. Protibiae (Fig. 48 J) nearly straight, inner margins emarginate and densely pubescent along apical half, gently projecting in middle, outer margins of protibiae concave slightly behind apex; mesotibiae (Fig. 48 K) strongly curved in apical third, apical half of inner margins pubescent; metatibiae (Fig. 48 L) nearly straight, inner margins widely emarginate near apical third, pubescent. Aedeagus (Fig. 49 A) elongate, gently curved in lateral view; parameres slender, very weakly curved towards dorsally in basal half, dorsum weakly ridged along midline, 0.27 as long as total length, with broadly widened and flabellate apex with apical marginal carina. Apical lobes of Sternite VIII (Fig. 49 E) rectangular in lateral view, with interior margins rounded, cornered at lowest point. Female: Unknown. Comparative notes. Morphostenophanes crassus is similar to M. jendeki jendeki. Both are small and stout, have wrinkled elytral disc with irregularly scattered short strial punctures, and strongly dilated apex of prosternal process in ventral view, but can be distinguished from M. jendeki jendeki by its wider pronotum and elytra, longer antenna, extremely widened pronotum with projecting posterior corners, male protibiae with inner margin bulged in middle; straight, wider and flatter metatibiae, and very short metaventral process.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A27FFE7FF5AF8C86F6A90CF.taxon	discussion	Comments. M. crassus is described from a weakly sclerotized (teneral) specimen which has faded to reddish brown. Although the only photograph showing its original color is very faint, strong luster of pronotum and elytra can be seen. Therefore, the real color of M. crassus may be similar to that of M. jendeki jendeki, M. jendeki similis and M. tanikadoi.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A27FFE7FF5AF8C86F6A90CF.taxon	distribution	Distribution. (Map 1) CHINA: Yunnan.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A27FFE7FF5AF8C86F6A90CF.taxon	etymology	Etymology. The new species is named from the Latin epithet ‘ crassus’ referring to its stubby and strongly convex habitus.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A58FFE4FF5AFB526CDC94B4.taxon	description	(Figs. 43 A – D; 47 B, G, L, Q; 48 B, F, M – O)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A58FFE4FF5AFB526CDC94B4.taxon	materials_examined	Type material examined. Paratype labelled: ‘ CHINA, YUNNAN prov. 18.6. – 4.7. 1993 heishui = 35 km N Lijiang 27, 13 N; 100, 19 E let. S. Becvar // Coll. Masumoto 2001 // Paratype Morphostenophane jendeki MASUMOTO [handwritten in pink label with underlines] ’ (male, NSMT, Fig. 43 C). Examined through two photographs taken by Kimio Masumoto.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A58FFE4FF5AFB526CDC94B4.taxon	discussion	Comments on type material. The type series of M. jendeki jendeki consisted of 21 specimens. According to the original description, the holotype is deposited in NMP (= NMNHP). Depository of paratypes was not mentioned in the original description, but at least part of them are currently deposited in NSMT. Additional material examined. CHINA: Yunnan: 1 ♂ (CZDY), Lijiang City, Yulong Snow Mountain, 27 ° 8 ’ 20.40 ” N, 100 ° 13 ’ 37.20 ” E, 3258 m, bark spray in conifer forest, 2012. viii, native collector; 1 ♀ (CZDY, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C), Lijiang City, Yulong Snow Mountain, 27 ° 11 ’ 13.20 ” N, 100 ° 13 ’ 1.20 ” E, 2012. viii, native collector. Comparative notes. Morphostenophanes jendeki jendeki resembles M. crassus. Detailed comparison is provided in the comparative notes of M. crassus. Comments. On the additional male specimen, a pair of small, shallow impressions on the pronotal surfaceand a deep impression on the posterior middle of sternite III was observed. These characters were not mentioned in the original description. The pronotal shape depicted in Fig. 34 A seems accordant with the specimen illustrated in the original publication (Masumoto 1998, Fig. 2), and the form depicted in Fig. 34 C (Paratype) resembles the form of M. jendeki similis. Therefore, whether pronotal shape is reliable for distinguishing M. jendeki jendeki and M. jendeki similis requires study of more specimens.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A58FFE4FF5AFB526CDC94B4.taxon	distribution	Distribution. (Map 1, 3) CHINA: Yunnan.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A5BFFE4FF5AFEC26CDC90E8.taxon	materials_examined	Material examined. None.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A5BFFE4FF5AFEC26CDC90E8.taxon	discussion	Comments on type material. The entire type series of Morphostenophanes jendeki similis was from Jizu Mountain. Based on the author’s correspondence with Kimio Masumoto, the type series is still in the collection of Stanislav Bečvář, and according to the original publication should be transferred and deposited in NMP (= NMNHP) in the future. Unfortunately, the type series was unavailable for study because the author failed to contact Stanislav Bečvář. Comments. Morphostenophanes jendeki similis is very similar to the nominate subspecies. According to the original description (Masumoto, 1998), they share similar body color, body size, aspect ratio of pronotum and elytra, antennae, and aedeagus. However, compared with M. jendeki jendeki, M. jendeki similis has ‘ more noticeably punctate’ head, less dilated genae and tempora; ‘ less densely punctate’ pronotum, ‘ a pair of round impressions slightly before the middle’ of pronotum, less produced lateral margins of pronotum; and ‘ more noticeably wrinkled’ elytra (Masumoto 1998). The above-mentioned differences can be noticed from the photos in the original publication, and are still considered here as main diagnostic characters. However, considering the whole genus, the differences between these two subspecies are quite vague, and some character states might be unstable within the population (as in the case of M. yunnanus). At least the shape of male pronotum in M. jendeki jendeki is here attested to be variable (see comments of M. jendeki jendeki). Moreover, as continuous high altitude ranging from about 2000 meters to over 3500 meters (see Map 3) connects habitats of the two subspecies, potential gene flow might exist between these two populations, and specimens with transitional characters may be found in these mountains. If so, M. jendeki similis will be synonymized with M. jendeki in the future.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A5BFFE4FF5AFEC26CDC90E8.taxon	distribution	Distribution. (Map 1, 3) CHINA: Yunnan.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A5AFFE2FF5AFF196C019030.taxon	description	(Figs. 44 A – D; 47 C, H, M, R; 48 C, G, P – R; 49 B, F, I)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A5AFFE2FF5AFF196C019030.taxon	materials_examined	Type materials. CHINA: Yunnan: ♂ (Holotype, SNUC), Chuxiong City, Dayao County, Mount Baicaoling, Zhuanwanhe forest conservation site, 2870 m, 2013. v. 28, Wen-Xuan Bi. Paratype: CHINA: Yunnan: 1 ♀ (SNUC, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C in CZDY), Chuxiong City, Dayao County, Mount Baicaoling, Zhuanwanhe forest conservation site, 2870 m, 2013. v. 28, Wen-Xuan Bi.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A5AFFE2FF5AFF196C019030.taxon	diagnosis	Diagnosis. Small, aeneous, elongate and moderately convex species. Elytra with rows of encircled segments of striae, some interrupted or straight, encircled areas weakly convex. Prosternal process not declivous. Aedeagal parameres with apical marginal carina.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A5AFFE2FF5AFF196C019030.taxon	description	Description. Male (Fig. 44 A, C). Color light aeneous and shiny; antennae and mouthparts dark brown; tarsi greenish; claws reddish brown. Body elongate, length 15.1 mm, width 5.2 mm, moderately convex dorsad, noticeably constricted between pronotum and elytra. Head (Fig. 47 C, H) transversely subquadrate, densely and markedly punctate, with outer margin strongly notched between genae and clypeus; clypeus transversely hexagonal, gently bent downwards in front, anterior margin nearly straight, weakly emarginate in middle, clypeal transverse impression short, marked; frontoclypeal suture finely impressed, widely U-shaped; genae strongly raised, depressed before eyes, strongly and roundly produced anterolaterally; frons broad, weakly convex in middle and feebly depressed near eyes, gradually sloping forwards; eyes transversely reniform, strongly convex laterally; inner ocular sulci finely grooved along inner margins, becoming broader posteriorly; tempora moderately convex, coarsely punctate. OI = 56.0. Antennae (Fig. 48 C) slender, reaching basal fourth of elytra, with antennomeres weakly thickened to apices; relative lengths of antennomeres: 0.46: 0.27: 0.68: 0.65: 0.71: 0.71: 0.70: 0.71: 0.71: 0.63: 0.79. Mentum (Fig. 47 H) inversely trapezoidal, lateral margins straight; medial surface finely punctate, with several large pores bearing long setae, gradually rising from base to apex, both sides of posterior half depressed. Pronotum (Fig. 47 M) barrel-shaped, PW / PL = 1.17, widest in anterior 2 / 5, anterior margin nearly straight, slightly projecting in middle, anterior marginal border marked; lateral margins weakly rounded, lateral marginal borders thin, visible in dorsal view along anterior half; posterior margin weakly emarginate, posterior marginal bor- der finely presented; anterior and posterior angles rounded; disc moderately convex, surface smooth, densely and finely punctate. Scutellum widely triangular, smooth, sparsely scattered with small punctures. Elytra oblong, widest in basal third, EL / EW = 1.89; moderately convex, highest near basal third; with rows of encircled segments of striae punctures, some interrupted or straight, encircled areas weakly convex; intervals more distinctly convex than encircled areas, smooth. Prosternum (Fig. 47 R) sparsely rugulose, finely and sparsely punctate; prosternal process straightly produced posteriorly, obtusely cuneate at apex; hypomeron rugulose, finely microsculptured, very sparsely and finely punctate. Metasternum glossy, finely punctate, metaventral anterior process transversely wrinkled. Abdomen (Fig. 48 G) depressed, surface weakly wrinkled, sparsely and finely punctate. Legs slender. Protibiae (Fig. 48 P) nearly straight, apical half of inner margins pubescent; mesotibiae (Fig. 48 Q) moderately curved in apical third, apical half of inner margins pubescent; metatibiae (Fig. 48 R) nearly straight, outer margins widely emarginate in middle, more than half of inner margins pubescent. Aedeagus (Fig. 49 B) elongate, curved in lateral view; parameres slender, dorsal margin depressed in apical half in lateral view, 0.22 as long as total length, with flabellate apex with apical marginal carina. Apical lobes of sternite VIII obliquely produced posteriorly in dorsal view (Fig. 49 F), rectangular in lateral view, with interior margins rounded, cornered at lowest point. Female (Fig. 44 B, D). Slightly larger and stouter than male, length 16.2 mm; OI = 56.0, PW / PL = 1.26, elytra more widened and convex, EL / EW = 1.69; abdomen straight in lateral view. Ovipositor (Fig. 49 I) shortened, abruptly narrowing terminally from apical third. Comparative notes. Within the jendeki - group, M. minor is the only species with encircled segments of elytra striae, similar to those found in the aenescens - or elegantulus - group species. However, its evenly thickened apex of parameres and shortened ovipositor confirms its place in the jendeki - group.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A5AFFE2FF5AFF196C019030.taxon	distribution	Distribution. (Map 1) CHINA: Yunnan.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A5AFFE2FF5AFF196C019030.taxon	etymology	Etymology. The new species is named from the Latin epithet ‘ minor’, referring its appearance, which seems like a shrunken elegantulus - group species.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A5DFFE0FF5AFB516B409758.taxon	description	(Figs. 45 A – D; 47 D, I, N, S; 48 D, H, S – U; 49 C, G, J)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A5DFFE0FF5AFB516B409758.taxon	materials_examined	Type materials. CHINA: Yunnan: ♂ (Holotype, SNUC), Diqing Prefecture, Weixi County, 27 ° 6 ’ 18.32 ” N, 99 ° 3 ’ 30.64 ” E, 3500 m, 2018. iii, native collector. Paratypes (8 ♂♂, 3 ♀♀): CHINA: Yunnan: 1 ♀ (SNUC), 2 ♂♂, 1 ♀ (MHBU), 4 ♂♂, 1 ♀ (CZDY, a male was preserved in 99.7 % ethanol at – 18 ° C), Diqing Prefecture, Weixi County, 27 ° 6 ’ 18.32 ” N, 99 ° 3 ’ 30.64 ” E, 3500 m, 2018. iii, native collector; 2 ♂♂ (CZDY), Weixi County, near Paidi, 2019. iv, native collector.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A5DFFE0FF5AFB516B409758.taxon	diagnosis	Diagnosis. Small, elongate and flattened species, with bronzy metallic luster. Pronotum coarsely punctate. Elytra with irregularly scattered short strial punctures, intervals moderately convex. Mesotibiae curved in apical third. Sternites III and IV depressed in posterior middle, with central portions of impressions convex.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A5DFFE0FF5AFB516B409758.taxon	description	Description. Male (Fig. 45 A, C). Bronzy, antennae and mouthparts dark brown; tarsi green; major portions of femora and claws reddish brown. Body elongate, length 15.5 – 17.4 mm, width 5.0 – 6.0 mm, moderately convex dorsad, noticeably constricted between pronotum and elytra. Head (Fig. 47 D, I) somewhat semicircular, densely and coarsely punctate, with outer margin faintly notched between genae and clypeus; clypeus transversely hexagonal, slightly convex in middle, gently bent downwards in front, with apex slightly rounded, emarginate in middle, frontoclypeal suture finely depressed, widely U-shaped; genae gently raised, depressed before eyes, gently and roundly produced anterolaterally; frons rather broad, distinctly sloping forwards; eyes transversely reniform, moderately convex laterally; inner ocular sulci faint in front, sharp and deep along inner margins and becoming shallower and broader posteriorly; tempora moderately convex, coarsely punctate. OI = 57.3 – 59.0. Antennae (Fig. 48 D) slender, reaching basal sixth of elytra, with antennomeres weakly thickened to apices; relative lengths of antennomeres: 0.49: 0.26: 0.71: 0.66: 0.65: 0.68: 0.77: 0.68: 0.66: 0.65: 0.80. Mentum (Fig. 47 I) quadrate, lateral margins rounded; medial surface wrinkled and coarsely punctate, with several large pores bearing long setae, gradually rising anteriorly, depressed in posterior half of both sides. Pronotum (Fig. 47 N) barrel-shaped, PW / PL = 1.12, finely microsculptured, widest in middle or slightly after basal third, anterior margin nearly straight, anterior marginal border faintly presented, obscured or completely interrupted in middle; lateral margins weakly rounded, occasionally broadened in posterior third, lateral marginal borders thin; posterior margin emarginate, posterior marginal border marked; anterior angles rounded, slightly produced anteriorly; posterior angles obtuse; disc weakly convex, coarsely punctate. Scutellum widely triangular, smooth, sparsely scattered with small punctures. Elytra ovate, rather elongate, widest near middle, EL / EW = 2 – 2.05; slightly more convex than pronotum, highest near basal third; disc with irregularly scattered short strial punctures; intervals moderately convex, with sharply marked scattered punctures. Prosternum (Fig. 47 S) rugulose, finely and sparsely punctate; prosternal process declivous, truncate at apex; hypomeron strongly rugulose, sparsely and finely punctate. Metasternum weakly wrinkled and finely punctate, metaventral anterior process transversely wrinkled. Abdomen (Fig. 48 H) depressed, surface somewhat wrinkled, coarsely punctate; sternites III and IV depressed in posterior middle, with central portions weakly convex. Legs slender. Protibiae (Fig. 48 S) curved in near apical third, apical half of inner margins pubescent; mesotibiae (Fig. 48 T) strongly curved in apical third, apical half of inner margins pubescent; metatibiae (Fig. 48 U) nearly straight, outer margins widely emarginate in middle, more than half of apical part of inner margins pubescent, outer margins weakly depressed before apices. Aedeagus (Fig. 49 C) elongate, curved in lateral view, with a pair of large pits on apical flank of basal piece; parameres slender, gently curved towards dorsally in basal half, dorsum weakly ridged along midline, 0.23 as long as total length, with broadly widened and flabellate apex, bearing apical marginal carina. Apical lobes of sternite VIII (Fig. 49 G) with rounded interior margins. Female (Fig. 45 B, D) slightly larger than male, length 16.6 – 18.6 mm. Distance between eyes longer, OI = 60 – 61.3; more constricted between pronotum and elytra; lateral margin of pronotum evenly and weakly rounded, PW / PL = 1.12; elytra more convex and widened, EL / EW = 1.86 – 1.98; abdomen straight in lateral view, without depressed area on sternites III and IV. Ovipositor (Fig. 49 J) shortened, apex beveled. Variability. The holotype has pronotum widest behind basal third, lateral margins distinctly widened at the widest point (as shown in Fig. 45 A), such character states are indistinct or missing in other males (as shown in Fig. 47 N). Considering that the holotype bears the largest body size, these features indicate that the holotype was a well-developed individual. Comparative notes. This new species resembles Morphostenophanes crassus, M. jendeki jendeki and M. jendeki similis. They share small body and similar elytral sculpture. M. planus can be distinguished from these congeners by flattened body with bronzy luster, more coarsely punctate pronotum, more elongate elytra, and more rounded produced apical lobe of male sternite VIII.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A5DFFE0FF5AFB516B409758.taxon	distribution	Distribution. (Map 1) CHINA: Yunnan.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A5DFFE0FF5AFB516B409758.taxon	etymology	Etymology. The new species is named from the Latin epithet ‘ planus’ referring to its flattened body.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A5FFFE0FF5AFCE96CE69139.taxon	description	(Figs. 46 A – D; 47 E, J, O, T; 48 E, I, V – X; 49 D, H)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A5FFFE0FF5AFCE96CE69139.taxon	materials_examined	Material examined. CHINA: Sichuan: 1 ♂ (CZDY, a sample of muscle tissue was preserved in 99.7 % ethanol at – 18 ° C), Shimian County, Yele Dam, 26 ° 55 ’ 22 ’’ N, 102 ° 13 ’ 32 ’’ E, in wilted leaves, 2500 m, 2015. vii. 25, De-Yao Zhou; 1 ♀ (MHBU), 1 ♀ (CDYZ), Liziping, Mamadi, 28 ° 59 ’ 18.41 ’’ N, 102 ° 24 ’ 34.64 ’’ E, 2600 m, 2016. vii. 24, De- Yao Zhou. Comparative notes. Morphostenophanes tanikadoi is unique among jendeki - group species by having slender body with strongly shiny green luster, elytral sculpture consisting of sparsely scattered short segments of striae connecting few deep punctures, sometimes varied into sharply marked large pits, intervals only weakly convex. M. tanikadoi and M. minor may be closely related by sharing similar habitus, but can be easily distinguished by their different body color, different elytral sculpture, and shorter legs. Detailed discussion see in the comments of M. minor.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A5FFFE0FF5AFCE96CE69139.taxon	distribution	Distribution. (Map 1) CHINA: Sichuan.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A5FFFE0FF5AFA3A6CF893B2.taxon	description	(Fig. 46 E, F)	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A5FFFE0FF5AFA3A6CF893B2.taxon	materials_examined	Type material examined. Holotype of Morphostenophanes tuberculatus labelled (translated from Chinese): ‘ 2005 - 5 · 19 Lushui Pianma Yakou [handwritten in white label] // 2005 - V- 19 Yunnan Lushui Pianma Yakou Xiao-Hong Ou Museum of Hebei University [printed label] // Holotype [red label] // Morphostenophane s tuberculatus Gao et Ren, 2009: Acta Zool. Hung., 55 (4): 315 - 318, figs. 31 - 38, 40 ’ (male, MHBU, Fig. 46 E). Examined through two photographs taken by Xing-Long Bai.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A5FFFE0FF5AFA3A6CF893B2.taxon	discussion	Comments. Morphostenophanes tuberculatus has the elytra with densely scattered unevenly sized tubercles, which resemble those of M. chongli chongli and M. chongli glaber. However, these two species are very different in many ways (see comparative notes of M. chongli) so that they clearly not belong to the same species group. Small body (body length 15.5 mm as mentioned in original publication), carinate apical margin of parameres (Gao & Ren 2009, Fig. 37) in M. tuberculatus suggest potential relation between M. tuberculatus and jendeki - group species.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A5FFFE0FF5AFA3A6CF893B2.taxon	distribution	Distribution. (Map 1) CHINA: Yunnan.	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
7F3D87954A50FFEFFF5AFF196ADC9364.taxon	synonymic_list	(# indicates the type species) Morphostenophanes aenescens species group Distribution	en	Zhou, De-Yao (2020): A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini). Zootaxa 4769 (1): 1-81, DOI: 10.11646/zootaxa.4769.1.1
