identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
794B8792FFDCFFFD3E88FED6FBF4FD99.text	794B8792FFDCFFFD3E88FED6FBF4FD99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fagus	<div><p>FAGUS L.</p> <p>Fagus mexicana Martínez, Anales Inst. Biol. Univ. Nac. México 11(1): 85. (1940). Type:— MEXICO. Hidalgo: M. Martínez 1500 (Lectotype (designated here): MEXU00009631 (image)!). Martínez (1940) did not cite a type specimen, but his collection number 1500 at MEXU is marked as the holotype. Specimens of Martínez 800 at F and US are marked as isotypes; these are not isotypes although they are original material.</p> <p>Fagus grandifolia Ehrh. subsp. mexicana (Martínez) A.E. Murray, Kalmia 13: 6 (1983).</p> <p>Fagus grandifolia var. mexicana (Martínez) Little, Castanea 30: 268 (1965).</p> <p>Distribution: — Mexico (Hidalgo, Puebla, San Luis Potosí, Tamaulipas, Veracruz).</p></div> 	https://treatment.plazi.org/id/794B8792FFDCFFFD3E88FED6FBF4FD99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FFDCFFFF3E88FD8AFCE8FEDD.text	794B8792FFDCFFFF3E88FD8AFCE8FEDD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus	<div><p>QUERCUS L.</p> <p>Quercus acherdophylla Trel., Mem. Natl. Acad. Sci. 20: 183, pl. 367 (1924). Type:— MEXICO, Puebla: C. G. Pringle 10008 (p.p.) (Syntypes at B destroyed. Lectotype (designated here): K000512971! based on collections made 25 Sep 1904; isolectotypes A!, BM!, CAS!, E!, ENCB (image)!, F!, G!, GH!, HBG (image)!, ILL!, MEXU!, MO!, NY!, P!, PH!, US!, VT!). Pringle used the same collection number for specimens he collected in flower on 25 Apr 1904 and in fruit on 25 Sep 1904. In 1988 K.C. Nixon marked the K specimen, which has collections from both dates, as the lectotype. We are designating just the 25 Sep 1904 collection as the lectotype, with the 25 Apr 1904 collection of the K specimen remaining as an isosyntype. This species is close to Q. salicifolia, and additional study is needed to determine if it is really distinct.</p> <p>Distribution: Mexico (Hidalgo, Oaxaca, Puebla, Veracruz).</p> <p>Quercus acutifolia Neé, Anales Ci. Nat. 3: 267 (1801). Type:— MEXICO, Guerrero: L. Neé s. n. (1791) (Lectotype: MA25953! designated by Nixon &amp; Barrie (2017, p. 449); isolectotypes: M!, UCSB (fragment)!). We also have seen a possible isolectotype at MPU.</p> <p>Quercus acutifolia var. bonplandii A.DC., Prodr. 16(2.1): 66 (1864). Type:— MEXICO, Michoacán: A. Bonpland 3917 (Lectotype (designated here): P06779600!; isolectotypes: B-W!, P!, UCSB (fragment)!). Trelease (1924, p. 229, pl. 391a) lectotypified the name on Bonpland 3917 at B, but it was destroyed. The specimen of Bonpland 3917 at B-W is not the one illustrated by Trelease.</p> <p>Quercus acutifolia var. conspersa (Benth.) A.DC., Prodr. 16(2.1): 66 (1864).</p> <p>Quercus acutifolia Neé var. longifolia A.DC., Prodr. 16(2.1): 67 (1864). Type:— Mexico, Veracruz: A. Ghiesbreght 124 (Lectotype: G! designated by Trelease (1924, p. 229, pl. 387); isolectotypes: K!, P!, UCSB (fragment)!). Trelease (1924, p. 229, pl. 387) lectotypified the name based on one of the syntypes.</p> <p>Quercus acutifolia var. microcarpa A.DC., Prodr. 16(2.1): 67 (1864). Type:— GUATEMALA: J. R. R. Warszewicz 25 (Holotype: G004100!; isotypes: K!, S (image)!, UCSB (fragment from G holotype)!).</p> <p>Quercus candolleana Trel., Mem. Natl. Acad. Sci. 20: 191, pl. 387 (1924). Nom. nov. for Q. acutifolia Neé var. longifolia A.DC. (1864) non Q. longifolia Raf. (1838) nec Liebm. (1854).</p> <p>Quercus conspersa Benth., Pl. Hartw. 91 (1842). Type:— GUATEMALA, Santa Rosa: T. Hartweg 617 (Lectotype (designated here): K000512983!; isolectotypes: BM!, CGE!, G!, K!, LD (image)!, NY!, UCSB (fragment)!). There are two sheets of Hartweg 617 in the Bentham Herbarium at K, one of which is lectotypified here.</p> <p>Quercus conspersa f. caudata Trel., Mem. Natl. Acad. Sci. 20: 192 (1924). Type:— GUATEMALA, Baja Verapaz: F. C. Lehmann 1320 (Holotype at B destroyed. Lectotype (designated here): G!; isolectotype: BM!).</p> <p>Quercus conspersa f. ovatifolia Trel., Mem. Natl. Acad. Sci. 20: 192, pl. 389b (1924). Type:— GUATEMALA: J. R. R. Warszewicz 28 (Holotype: G!; isotype UCSB (fragment from G holotype)!).</p> <p>Quercus correpta Trel., Mem. Natl. Acad. Sci. 20: 153, pl. 300b (1924). Nom. nov. for Q. acutifolia var. microcarpa (1864) non Q. microcarpa Lapeyr (1813).</p> <p>Quercus grahamii Benth. var. coyulana Trel., Mem. Natl. Acad. Sci. 20: 190 (1924). Type:— MEXICO, Oaxaca: C. Conzatti 3553 (Lectotype (designated here): US00089536!; isolectotype: MEXU!). Trelease (1924, p. 190) did not indicate where the type specimen was deposited.</p> <p>Distribution: Guatemala, Mexico (Chiapas, Ciudad de México, Coahuila, Durango, Guanajuato, Guerrero, Hidalgo,</p> <p>Jalisco, México, Michoacán, Morelos, Nayarit, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa,</p> <p>Tamaulipas, Tlaxcala, Veracruz, Zacatecas). The occurrence of Q. acutifolia in Ciudad de México, Coahuila, Colima,</p> <p>Durango, Guanajuato, Hidalgo, Querétaro, Sinaloa, and Tamaulipas reported by Villaseñor (2016) needs to be confirmed.</p> <p>Quercus affinis Scheidw., Hort. Belge. 4: 321, 17 (1837). Type:— MEXICO, Hidalgo: H. Galeotti 115 (Lectotype (designated here): BR0000006914974!; isolectotypes: BR!, G!, K!, P!). The date of 1840 on Galeotti collections is not the actual collection date, but rather the date the numbers were assigned back in Europe (McVaugh 1978). The actual collection date for what was assigned the number 115 in 1840 was probably 1836.</p> <p>Quercus affinis f. commutata Trel., Mem. Natl. Acad. Sci. 20: 167 (1924). Type:— MEXICO, Veracruz: C. J. W. Schiede 18 (p.p.) (Lectotype: G! designated by Trelease (1924, p. 227, pl. 333a); isolectotype: HAL!). Quercus commutata Liebm. (1854) non Unger (1852) is a later homonym, but the specific epithet of Liebmann was used by Trelease (1924, p. 167), based on the same type, as the basis for Q. affinis f. commutata. The specimen of Schiede 18 (p.p.) at G and one of the two sheets at HAL are from Jalacingo, whereas the other sheet at HAL and the ones at HBG, MO, and the fragment at UCSB are from Xalapa, as is the lectotype of Q. xalapensis f. jalapae.</p> <p>Quercus affinis f. subintegra (A.DC.) Trel., Mem. Natl. Acad. Sci. 20: 167 (1924).</p> <p>Quercus commutata Liebm. non Unger (1852), Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 177 (1854). Although this was lectotypified by Trelease (1924, p. 227, pl. 333a) and treated by him as a forma of Q. affinis, at the specific level it is a later homonym of a fossil oak species.</p> <p>Quercus laurina Bonpl. var. major (A.DC.) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 205 (1884).</p> <p>Quercus laurina var. podocarpa (A.DC.) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 205 (1884).</p> <p>Quercus major (A.DC.) Trel. non Nakai (1916), Contr. U.S. Natl. Herb. 23(2): 193 (1922).</p> <p>Quercus nitens M.Martens &amp; Galeotti non Raf. (1838), Bull. Acad. Roy. Sci. Bruxelles 10(1): 217 (1843). This is an illegitimate name because it is both a later homonym and is based on the same type as Q. affinis Scheidw. (1837).</p> <p>Quercus nitens M.Martens &amp; Galeotti non Raf. (1838) var. major A.DC., Prodr. 16(2.1): 69 (1864). Type:— MEXICO, Veracruz: F. M. Liebmann 3518 (Lectotype (designated here): C10012540!; isolectotypes: CAS!, GH!, K (as s.n.)!, NY!, P (as s.n.)!, UCSB (fragment)!). Although A.de Candolle (1864) did not cite the Liebmann number of the specimens at C for this variety, he did cite it by locality and elevation. The Liebmann collection corresponding to “montis Orizabae, loc Alpatlahua dieto, alt. 7500 p.” is number 3518 collected in Sep 1841. A.de Candolle’s determined, in his hand, two of the three specimens at C plus the sheet now at CAS which was formerly at C as Q. nitens var. major.</p> <p>Quercus nitens M.Martens &amp; Galeotti non Raf. (1838) var. podocarpa A.DC., Prodr. 16(2.1): 69 (1864). Type:— MEXICO, Puebla: F. M. Liebmann 3441 (Lectotype (designated here): C 10012509!; isolectotypes: C!, CAS!, P (as s.n.)!). Trelease (1924) in the caption to pl. 335a wrote “The type, as of Q. nitens podocarpa (Mexico, Liebmann, 3441).” The photo in plate 335a is of unmounted fragments photographed at C which do not correspond to any of the mounted sheets of Liebmann 3441. We are lectotypifying one of the C specimens of Liebmann 3441.</p> <p>Quercus nitens M.Martens &amp; Galeotti non Raf. (1838) var. subintegra A.DC., Prodr. 16(2.1): 69 (1864). Type:— MEXICO, Hidalgo: T. Hartweg 422 (Holotype: G00358099!; isotypes: BM!, CGE!, E!, G!, K!, L!, NY!, P!, UCSB (fragment)!).</p> <p>Quercus ocoteifolia Liebm. f. confusa Trel., Mem. Natl.Acad. Sci. 20: 168, pl. 335b (1924). Type:— MEXICO, Oaxaca: F. M. Liebmann 3514 (Holotype: G (as 60–64)!; isotypes: C (as 62)!, CAS (as 63)!, GH (as 60–65)!). Liebmann 3514 also was assigned individual sheet numbers 60–62 through 60–65. Trelease (1924, p. 168) wrote that the type is 60–64, and in the caption of plate 335b on p. 220 he indicated that the type is at C. The photo in pl. 335b, however, is the specimen labeled 60–64 at G.</p> <p>Quercus ocoteifolia f. podocarpa (A.DC.) Trel., Mem. Natl. Acad. Sci. 20: 168, (1924).</p> <p>Quercus treleaseana A.Camus, Bull. Mus. Natl. Hist. Nat., sér. 2, 4: 124 (1932). Nom. nov. for Q. major (A. DC.) Trelease (1924) non Nakai (1916) based on Q. nitens var. major. Even as a nom. nov. Q. treleaseana is an illegitimate name because it is based on the same type as Q. affinis Scheidw.</p> <p>Distribution: Mexico (Chiapas, Ciudad de México, Coahuila, Guanajuato, Guerrero, Hidalgo, Jalisco, México,</p> <p>Michoacán, Morelos, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Tamaulipas, Veracruz). The occurrence of Q. affinis in Guerrero, Jalisco, and Michoacán reported by Villaseñor (2016) needs to be confirmed.</p> <p>Quercus agrifolia Neé, Anales Ci. Nat. 3: 271 (1801). Type:— U.S.A., California: J. Robredo &amp; M. Esquerra s. n. (1791) (Lectotype: MA25959! designated by Bartholomew &amp; Almeda (2022, p. 28); isolectotypes: MA!, UCSB (fragment from MA lectotype)!, UCSB (fragment from MA isolectotype)!). The locality on the MA labels is between Monterey and Nootka. Although the type locality is unknown, it is most likely in the vicinity of Monterey, California (Bartholomew &amp; Almeda 2022).</p> <p>Distribution: Mexico (Baja California), U.S.A. (California).</p></div> 	https://treatment.plazi.org/id/794B8792FFDCFFFF3E88FD8AFCE8FEDD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FF9EFFBF3E88FF66FD21F7F1.text	794B8792FF9EFFBF3E88FF66FD21F7F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus acherdophylla Trel.	<div><p>Quercus acherdophylla Trel.</p> <p>Quercus achoteana Trel. = Quercus segoviensis Liebm.</p> <p>Quercus achoteana var. sublanosa Trel. = Quercus segoviensis Liebm.</p> <p>Quercus aculcingensis Trel. = Quercus greggii (A.DC.) Trel.</p> <p>Quercus acuminata (Michaux) Sarg. (1895) non Roxb. (1832) = Quercus muehlenbergii Engelm.</p> <p>Quercus acuminata M.Martens &amp; Galeotti (1843) non Roxb. (1832) = Quercus calophylla Schltdl. &amp; Cham. Quercus acutangula Trel. = Quercus elliptica Neé</p> <p>Quercus × acutidens Torr.</p> <p>Quercus acutifolia Neé</p> <p>Quercus acutifolia f. abrupta Trel. = Quercus grahamii Benth.</p> <p>Quercus acutifolia var. angustifolia A.DC. = Quercus xalapensis Bonpl.</p> <p>Quercus acutifolia var. bonplandii A.DC. = Quercus acutifolia Neé</p> <p>Quercus acutifolia var. conspersa (Benth.) A.DC. = Quercus acutifolia Neé</p> <p>Quercus acutifolia var. furfuracea (Liebm.) Oerst. = Quercus furfuracea Liebm.</p> <p>Quercus acutifolia var. longifolia A.DC. = Quercus acutifolia Neé</p> <p>Quercus acutifolia var. microcarpa A.DC. = Quercus acutifolia Neé</p> <p>Quercus aequivenulosa Trel. = Quercus coffeicolor Trel.</p> <p>Quercus × aerea Trel.</p> <p>Quercus affinis M.Martens &amp; Galeotti (1843) non Scheidw. (1837) = Quercus peduncularis Neé</p> <p>Quercus affinis Scheidw.</p> <p>Quercus affinis f. commutata Trel. = Quercus affinis Scheidw.</p> <p>Quercus affinis f. subintegra (A.DC.) Trel. = Quercus affinis Scheidw.</p> <p>Quercus agrifolia Neé</p> <p>Quercus aguana Trel. = Quercus purulhana Trel.</p> <p>Quercus ajoensis C.H.Mull.</p> <p>Quercus alamarensis C.H.Mull. = Quercus canbyi Trel.</p> <p>Quercus alamo Benth. = Quercus calophylla Schltdl. &amp; Cham.</p> <p>Quercus alamosensis Trel. = Quercus arizonica Sarg.</p> <p>Quercus albifolia C.H.Mull. = Quercus gambelii Nutt.</p> <p>Quercus albocincta Trel.</p> <p>Quercus almaguerensis Bonpl. = Quercus humboldtii Bonpl.</p> <p>Quercus alpescens Trel. = Quercus greggii (A.DC.) Trel.</p> <p>Quercus alvarezensis Trel. = Quercus spicata Bonpl.</p> <p>Quercus alveolata Trel.</p> <p>Quercus ambigua Bonpl. = Quercus glabrescens Benth.</p> <p>Quercus ambivenulosa Trel. = Quercus laurina Bonpl.</p> <p>Quercus amissiloba Trel. = Quercus sapotifolia Liebm.</p> <p>Quercus amphioxys Trel. = Quercus benthamii A.DC.</p> <p>Quercus anglohondurensis C.H.Mull. = Quercus cortesii Liebm.</p> <p>Quercus apanecana Trel. = Quercus sapotifolia Liebm.</p> <p>Quercus × apiophylla E.F.Warb.</p> <p>Quercus arachnoidea Trel. = Quercus segoviensis Liebm.</p> <p>Quercus ariifolia Trel.</p> <p>Quercus aristata Hook. &amp; Arn.</p> <p>Quercus aristigera Trel. = Quercus skutchii Trel.</p> <p>Quercus arizonica Sarg.</p> <p>Quercus arizonica var. wootoni Trel. = Quercus arizonica Sarg.</p> <p>Quercus atrescentirhachis Trel. = Quercus elliptica Neé</p> <p>Quercus atriglans E.F.Warb. = Quercus obtusata Bonpl.</p> <p>Quercus aurantiaca Trel. = Quercus tuberculata Liebm.</p> <p>Quercus axillaris E.Fourn. ex Trel. = Quercus castanea Neé</p> <p>Quercus baldoquinae Trel. = Quercus glaucoides M.Martens &amp; Galeotti</p> <p>Quercus balsequillana Trel. = Quercus emoryi Torr.</p> <p>Quercus barbanthera Trel. = Quercus peduncularis Neé</p></div> 	https://treatment.plazi.org/id/794B8792FF9EFFBF3E88FF66FD21F7F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FF82FFA33E88FB52FB71F7F9.text	794B8792FF82FFA33E88FB52FB71F7F9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus acutifolia	<div><p>Q. acutifolia × Q. grahamii Mexico (Guerrero). G. McPherson 964 CAS!, K!, NY!, MICH!, MO!</p> <p>Q. acutifolia × Q. tlapuxahuensis Mexico (Michoacán). G. B. Hinton 12153 GH!, NY!, US!, 13692 GH!, ILL!, NY!, PH!, US!, 13730 GH!, ILL!, NY!, PH!</p> <p>Q. × acutidens Mexico (Baja California). D. E. Breedlove &amp;. C. Burns 71552 CAS!, 71553 71554 CAS!, 71555 CAS!; H. F. Copeland s.n. (1 Apr 1947) DS!; J. T. Howell 30927 CAS!; I. L. Wiggins &amp; Gillespie 4032 A!, CAS!, DS!, MO!, NY!, 4063 A, CAS!, DS!, GH!, MO!, NY!</p> <p>Q. × aerea Mexico (Chihuahua). R. Spellenberg 9381 CAS!, MEXU!, 9807 CAS!, MEXU!</p> <p>Mexico (Durango). R. Spellenberg 11015A CAS!, MEXU!</p> <p>Q. alveolata × Q. grisea Mexico (Chihuahua). H. LeSueur 437 MO!</p> <p>Mexico (Durango). M. Avila H. s. n. (14 Oct 1981) MEXU!</p> <p>Q. benthamii × Q. cortesii Guatemala (Alta Verapaz). P. C. Standley 69935 F!, 90235 A!, F!; 91321 F!, US! Guatemala (Baja Verapaz). P. C. Standley 91077 F!</p> <p>Guatemala (San Marcos). J. A. Steyermark 37193 F!</p> <p>Honduras (Comayagua). T. Hawkins 29 CAS!; D. Mejia 217 CAS!</p> <p>Honduras (Lempira). D. Mejia 190 CAS !</p> <p>Honduras (Francisco Morazán). A. Molina R. 130 CAS!, F!; L. O. Williams 17440 F!, GH!, US!</p> <p>Q. benthamii × Q. salicifolia Costa Rica (Puntarenas). V. L. Sork 42 CAS!</p> <p>Costa Rica (San José). W. C. Burger 8380B F!; L. O. Williams 28637 F!, NY!</p> <p>Honduras (El Paraíso). L. O. Williams 15749 BM!, F!, GH!, MO!, US!, WIS!</p> <p>Panama (Chiriquí). M. D. Correa A. &amp; R. Lazot 1395 MO!, PMA!; W. G. D’Arcy 12716 MO!; E. A. Lao 327 MO!, PMA!; L. S. McIntyre 16 F!; R. J. Schmalzel 1329 MO!; W. L. Stern 52 A!, MO!, US!</p> <p>Q. bolanyosensis × Q. emoryi Mexico (Durango). R. Spellenberg 11010E5 CAS!, MEXU!</p> <p>Q. bumelioides × Q. lancifolia Guatemala (Huehuetenango). J. A. Steyermark 48725 F!, 48906 F!</p> <p>Honduras (Lempira). G. Davidse 34698 CAS!</p> <p>Honduras (Francisco Morazán). C. V. Morton 7490 US!; L.O. Williams 17469 F!, GH!, US!</p></div> 	https://treatment.plazi.org/id/794B8792FF82FFA33E88FB52FB71F7F9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FFDEFFFE3E88FED6FEA1F7FB.text	794B8792FFDEFFFE3E88FED6FEA1F7FB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus ajoensis C. H. Mull., Madrono	<div><p>Quercus ajoensis C.H.Mull., Madroño 12(5): 140, fig. 1 (1954). Type:— U.S.A., Arizona: C. H. Muller &amp; J. M. Tucker</p> <p>9519 (Holotype: BH; isotypes: A!, MICH!, MO!, NY!, RSA (image)!, UC!, UCSB (fragment from BH holotype)!,</p> <p>UCSB!, US!). Quercus turbinella Greene subsp. ajoensis (C.H.Mull.) Felger &amp; C.H.Lowe, J. Arizona Acad. Sci. 6(1): 83 (1970). Quercus turbinella var. ajoensis (C.H.Mull.) Little, Phytologia 42(3): 221 (1979). Distribution: Mexico (Baja California, Sonora), U.S.A. (Arizona).</p> <p>Quercus albocincta Trel., Mem. Natl. Acad. Sci. 20: 193, pl. 394a–b (1924). Type:— MEXICO, Sonora: J. N. Rose et al. 12787 (Lectotype (designated here): US 00089413!; isolectotypes: NY!, UCSB (fragment)!). Trelease (1924) indicated that the type was at both NY and US.</p> <p>Distribution: Mexico (Baja California Sur, Chihuahua, Durango, Sinaloa, Sonora).</p> <p>Quercus alveolata Trel., Mem. Natl. Acad. Sci. 20: 80, pl. 114b (1924). Type:— MEXICO, Puebla: C. A. Purpus 4091 (Holotype: NY00253649!; isotypes: BM!, F!, GH!, MO!, UC!, UCSB (fragment)!, US!).</p> <p>Quercus chihuahuensis Trel., Mem. Natl.Acad. Sci. 20: 85, pl. 130a (1924).Type:— MEXICO, Chihuahua: C. G. Pringle 970 (Lectotype (designated here): MO-473961!; isolectotypes: MEXU!, NY!, UC!, VT!). Trelease (1924, p. 85) wrote “(Pringle, 74, 355, the type)” which would appear to make Pringle 355 the type collection number. He also cited Pringle 970, which based on what he wrote on page 85, would make Pringle 970 a paratype. In the caption to pl. 129a and pl. 129b on p. 220 he wrote “The type collections (Mexico, Pringle: a, no 74; b, no 355). Photographed by permission at Dahlem.” However, in the caption to pl. 130a on the same page he also wrote “The type (Mexico, Pringle 970). Photographed at St. Louis.” If taken all together, our interpretation is that Trelease cited two syntypes for Q. chihuahuensis with one being Pringle 355 at B, now destroyed, and the other being Pringle 970 at MO. We are lectotypifying the name based on the existing syntype at MO. The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. alveolata as broadly interpreted.</p> <p>Quercus chihuahuensis f. amplifolia Trel., Mem. Natl. Acad. Sci. 20: 86, pl. 131b (1924). Type:— MEXICO, San Luis Potosí: W. Schumann 1313 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 131b)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Quercus chihuahuensis f. microphylloides Trel., Mem. Natl. Acad. Sci. 20: 86, pl. 130b (1924). Type:— MEXICO, San Luis Potosí: W. Schumann 1310 (p.p.) (Holotype at B destroyed. Lectotype (designated here): US00089452!). Trelease (1924, p. 86) cited Schumann 1310 (p.p.) as the type of Q. chihuahuensis f. tenuis, but on p. 220 in the caption for pl. 131a, he cited Schumann 1300 (p.p.) as the type at B. The second citation is most likely a typographic error and should have been 1310 (p.p.). The specimens at US of 1310 which we are designating as the lectotype match the description in the original protologue.</p> <p>Quercus chihuahuensis f. tenuis Trel., Mem. Natl. Acad. Sci. 20: 86, pl. 131a (1924). Type:— MEXICO, San Luis Potosí: W. Schumann 1310 (p.p.) (Holotype at B destroyed. Lectotype (designated here): P06852494!; isolectotype: M!). The specimens at P and M match the description of Q. chihuahuensis f. tenuis.</p> <p>Quercus deserticola Trel. var. incisa Trel., Mem. Natl. Acad. Sci. 20: 80, pl. 114a (1924). Type:— MEXICO, Hidalgo: Uhde 265 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 114a)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Quercus infralutea Trel., Mem. Natl. Acad. Sci. 20: 87, pl. 136 (1924). Type:— MEXICO, Sonora: J. N. Rose et al. 12788 (Holotype: US00089550!; isotypes: NY!, UCSB (fragment)!).</p> <p>Quercus panduriformis Trel. f. colimensis Trel., Mem. Natl. Acad. Sci. 20: 63, pl. 63b (1924). Type:— MEXICO, Colima: H. Ross 497 (Holotype: M-0244473!; isotype UCSB (fragment from M holotype)!).</p> <p>Quercus tamiapensis C.H.Mull., Amer. Midl. Naturalist 27(2): 480 (1942). Type:— MEXICO, Sinaloa: H. S. Gentry 5863 (Holotype: US00089669!; isotypes: DS!, GH!, MEXU!, MO!, NY!, RSA (image)!, UCSB (fragment)!).</p> <p>Distribution: Mexico (Aguascalientes, Chihuahua, Ciudad de México, Colima, Durango, Guanajuato, Hidalgo,</p> <p>Jalisco, México, Michoacán, Nayarit, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa, Sonora,</p> <p>Tlaxcala, Zacatecas), U.S.A. (Texas).</p> <p>Quercus ariifolia Trel., Mem. Natl. Acad. Sci. 20: 74, pl. 97b (1924). Type:— MEXICO, San Luis Potosí: E. Palmer 82 (Holotype: MO-144181!; isotypes: A!, BM!, CAS!, F!, GH!, MEXU!, NY!, UC!, UCSB (fragment from MO holotype)!, US!). This species is close to Q. gravesii, and additional study is needed to determine if it is really distinct.</p> <p>Distribution: Mexico (San Luis Potosí).</p> <p>Quercus aristata Hook. &amp; Arn., Bot. Beechey Voy. 444 (1841). Type:— MEXICO, Nayarit: A. Sinclair s. n. (1845) (Holotype: K000512917!; isotype: E!).</p> <p>Quercus planipocula Trel. f. subsessilifolia Trel., Mem. Natl. Acad. Sci. 20: 137 (1924). Type:— MEXICO, Sinaloa: M. Narvaez M. &amp; A. E. Salazar 1169 (Lectotype (designated here): ILL00007622!; isolectotype: ILL!). Two specimens at ILL are marked as types in Trelease’s hand.</p> <p>Quercus productipes Trel., Mem. Natl. Acad. Sci. 20: 140, pl. 264b (1924). Type:— MEXICO, Nayarit: J. Gregg 977 (Holotype: MO-3076316!; isotype UCSB (fragment from MO holotype)!).</p> <p>Distribution: Mexico (Aguascalientes, Colima, Durango, Guanajuato, Guerrero, Jalisco, México, Nayarit, Oaxaca,</p> <p>Querétaro, Sinaloa, Sonora, Veracruz, Zacatecas). The occurrence of Q. aristata in Guerrero, Oaxaca, and Veracruz reported by Villaseñor (2016) needs to be confirmed.</p> <p>Quercus arizonica Sarg., Gard. &amp; Forest 8: 92 (1895). Type:— U.S.A., Arizona: C. S. Sargent s. n. (23 Mar 1894) (Lectotype (designated here): A00034033!; isolectotype: A!). The two specimens at A were collected from the same tree on the same date and marked as the type. Breedlove considered that Q. arizonica could be treated in the broad sense as being the same species as Q. laeta, but here we are treating them as distinct species.</p> <p>Quercus alamosensis Trel., Mem. Natl. Acad. Sci. 20: 179, pl. 361b (1924). Type:— MEXICO, Sonora: E. Palmer 370 (Holotype: GH00055620!; isotype: US!).</p> <p>Quercus arizonica var. wootoni Trel., Mem. Natl. Acad. Sci. 20: 89 (1924). Type:— U.S.A., New Mexico: E. O. Wooton s. n. (1 Oct 1905) (Holotype: NY00248612!; isotype: US!).</p> <p>Quercus sacame Trel., Mem. Natl. Acad. Sci. 20: 89, pl. 142a (1924). Type:— MEXICO, Chihuahua: R. Endlich 1282a (Syntypes at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 142b)!). No extant original material exists, so we are lectotypifying the name based on the image of one of the syntypes.</p> <p>Distribution: Mexico (Chihuahua, Coahuila, Durango, Sonora), U.S.A. (Arizona, New Mexico, Texas). The report of</p> <p>Q. arizonica from Baja California Sur, Nayarit, Nuevo León, Sinaloa, and Tamaulipas by Villaseñor (2016) is actually</p> <p>Q. laeta.</p> <p>Quercus benthamii A.DC., Prodr. 16(2.1): 29 (1864). Type:— GUATEMALA, Quezaltenango: T. Hartweg 563 (Holotype: K000512928!; isotypes: BM!, CGE!, E!, G!, LD (image)!, NY!, OXF!, P!, UCSB (fragment)!). Nom. nov. for Q. undulata Benth. (1841) non Torr. (1827).</p> <p>Quercus amphioxys Trel., Mem. Natl. Acad. Sci. 20: 141, pl. 268a (1924). Type:— EL SALVADOR: G. Niederlein s. n. (9 Jan 1898) (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 268a)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Quercus baruensis C.H.Mull., Trop. Woods 108: 75, f. 1, 2 (1958). Type:— PANAMA, Chiriquí: W. L. Stern &amp; K. L. Chambers 55 (Holotype: WIS00000212 MAD!). The holotype was formerly at Y.</p> <p>Quercus crispifolia Trel., Mem. Natl. Acad. Sci. 20: 147, pl. 286 (1924). Type:— MEXICO, Chiapas: E. Reeves 4 (Holotype: ILL00007523! and ILL00010194!). The holotype is mounted on two sheets, one with just leaves and the other with just acorns and cups. The sheet with acorns and cups has the label from the collector. The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. benthamii as broadly interpreted.</p> <p>Quercus esesmilensis Tucker &amp; C.H.Mull., Madroño 8(4): 115, pl. 14 (1945). Type:— EL SALVADOR, Chalatenango: J. M. Tucker 1198 (Holotype: UC694107!; isotypes: CAS!, F!, K!, LL (image)!, MICH!, NY!, PH!, UC!, UCSB (fragment)!, US!). The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. benthamii as broadly interpreted.</p> <p>Quercus gemmata Trel., Mem. Natl. Acad. Sci. 20: 152, pl. 299b (1924). Type:— EL SALVADOR: G. Niederlein s. n. (9 Jan 1898) (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 299b)!). No extant original material exists, so we are lectotypifying that name based on the image of the holotype.</p> <p>Quercus lowilliamsii C.H.Mull., Madroño 10(5): 138, pl. 11 (1950). Type:— HONDURAS, Morazán: L. O. Williams &amp; A. Molina 12760 (Holotype: BH; isotypes: CAS!, EAP (image)!, F!, GH!, MO!, UC!, UCSB (fragment from BH holotype)!, US!).</p> </div>	https://treatment.plazi.org/id/794B8792FFDEFFFE3E88FED6FEA1F7FB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FF9FFFBE3E88FF67FBA8F7F1.text	794B8792FF9FFFBE3E88FF67FBA8F7F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus barbanthera var. calva Trel.	<div><p>Quercus barbanthera var. calva Trel. = Quercus peduncularis Neé</p> <p>Quercus barbeyana Trel. = Quercus purulhana Trel.</p> <p>Quercus barbinervis Benth. = Quercus laurina Bonpl.</p> <p>Quercus barrancana Spellenb. = Quercus toumeyi Sarg.</p> <p>Quercus baruensis C.H.Mull. = Quercus benthamii A.DC.</p> <p>Quercus × basaseachicensis C.H.Mull.</p> <p>Quercus benthamii A.DC.</p> <p>Quercus bipedalis Trel. = Quercus laeta Liebm.</p> <p>Quercus bocoynensis C.H.Mull. = Quercus depressipes Trel.</p> <p>Quercus bolanyosensis Trel.</p> <p>Quercus bonplandiana Sweet = Quercus glabrescens Benth.</p> <p>Quercus boqueronae Trel. = Quercus lancifolia Schltdl. &amp; Cham.</p> <p>Quercus boquetensis Standl. = Quercus salicifolia Neé</p> <p>Quercus borucasana Trel. = Quercus salicifolia Neé</p> <p>Quercus botryocarpa Trel. = Quercus elliptica Neé</p> <p>Quercus bourgaei Oerst. ex Trel. = Quercus laurina Bonpl.</p> <p>Quercus bourgaei var. ilicifolia Trel. = Quercus laurina Bonpl.</p> <p>Quercus boyacensis Cuatrec. = Quercus humboldtii Bonpl.</p> <p>Quercus brachystachys Benth. = Quercus crassifolia Bonpl.</p> <p>Quercus brachystachys f. caerulea Trel. = Quercus skutchii Trel.</p> <p>Quercus brachystachys f. venulosa Trel. = Quercus skutchii Trel.</p> <p>Quercus brandegeei Goldman</p> <p>Quercus brayi Small = Quercus muehlenbergii Engelm.</p> <p>Quercus breedloveana Nixon &amp; Barrie</p> <p>Quercus brenesii Trel. = Quercus cortesii Liebm.</p> <p>Quercus breviloba (Torr.) Sarg. = Quercus sinuata Walter</p> <p>Quercus breviloba f. laceyi (Small) Trel. = Quercus laceyi Small</p> <p>Quercus breviloba subsp. laceyi (Small) A.Camus = Quercus laceyi Small</p> <p>Quercus bumelioides Liebm.</p> <p>Quercus caeruleocarpa Trel. = Quercus laurina Bonpl.</p> <p>Quercus californica (Torrey) J.G.Cooper = Quercus kelloggii Newb.</p> <p>Quercus callosa Benth. = Quercus peduncularis Neé</p> <p>Quercus calophylla Schltdl. &amp; Cham.</p> <p>Quercus calophylla f. acuminata Seemen ex Trel. = Quercus calophylla Schltdl. &amp; Cham.</p> <p>Quercus calophylla f. alamo (Benth.) Trel. = Quercus calophylla Schltdl. &amp; Cham.</p> <p>Quercus calophylla f. flavida (Liebm.) Trel. = Quercus calophylla Schltdl. &amp; Cham.</p> <p>Quercus calophylla f. intermedia Seemen ex Trel. = Quercus calophylla Schltdl. &amp; Cham.</p> <p>Quercus calophylla f. schiedeana Trel. = Quercus calophylla Schltdl. &amp; Cham.</p> <p>Quercus calophylla f. willdenovii Trel. = Quercus calophylla Schltdl. &amp; Cham.</p> <p>Quercus campanariensis Trel. = Quercus scytophylla Liebm.</p> <p>Quercus canbyi Trel.</p> <p>Quercus canbyi f. ascendens Trel. = Quercus canbyi Trel.</p> <p>Quercus canbyi f. attenuata C.H.Mull. = Quercus canbyi Trel.</p> <p>Quercus canbyi f. berlandieri Trel. = Quercus canbyi Trel.</p> <p>Quercus canbyi f. concolor Trel. &amp; C.H.Mull. = Quercus canbyi Trel.</p> <p>Quercus canbyi f. karwinskii (Trel.) C.H.Mull. = Quercus canbyi Trel.</p> <p>Quercus canbyi f. pedunculata C.H.Mull. = Quercus canbyi Trel.</p> <p>Quercus canbyi f. setacea C.H.Mull. = Quercus canbyi Trel.</p> <p>Quercus canbyi f. subovatifolia C.H.Mull. = Quercus canbyi Trel.</p> <p>Quercus cancellata Trel. = Quercus glaucoides M.Martens &amp; Galeotti</p> <p>Quercus candicans Neé = Roldana candicans (Neé) Villaseño, S. Valencia &amp; Coombes (see Appendix 1, Excluded Names)</p> <p>Quercus candicans auct. non Neé f. alligata Trel. = Quercus calophylla Schltdl. &amp; Cham.</p> <p>Quercus candicans auct. non Neé f. incurva Trel. = Quercus calophylla Schltdl. &amp; Cham.</p></div> 	https://treatment.plazi.org/id/794B8792FF9FFFBE3E88FF67FBA8F7F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FF9DFFBC3E88FC6BFD21F7F1.text	794B8792FF9DFFBC3E88FC6BFD21F7F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus barrancana Spellenb., Phytoneuron 2014	<div><p>Quercus barrancana Spellenb. = Q. toumeyi Sarg.</p> <p>Quercus breedloveana Nixon &amp; Barrie</p> <p>Quercus centenaria L.M.González = Q. vicentensis Trel.</p> <p>Quercus coahuilensis Nixon &amp; C.H.Mull.</p> <p>Quercus cualensis L.M.González</p> <p>Quercus delgadoana S.Valencia, Nixon &amp; L.M.Kelly</p> <p>Quercus hintoniorum Nixon &amp; C.H.Mull.</p> <p>Quercus hirtifolia L.M.Vázquez, S.Valencia &amp; Nixon</p> <p>Quercus huicholensis R.A.McCauley</p> <p>Quercus iltisii L.M.González</p> <p>Quercus mcvaughii Spellenb.</p> <p>Quercus meavei Valencia-A., Sabas &amp; O.J.Soto</p> <p>Quercus melissae Nixon &amp; Barrie</p> <p>Quercus mexiae L.M.González = Q. coffeicolor Trel.</p> <p>Quercus miquihuanensis Nixon &amp; C.H.Mull.</p> <p>Quercus nixoniana S.Valencia &amp; Lozada-Pérez = Q. salicifolia Neé</p> <p>Quercus sarahmariae Nixon &amp; Barrie</p> <p>Quercus tarahumara Spellenb., J.D.Bacon &amp; Breedlove 1995</p> <p>Quercus tuitensis L.M.González</p> <p>APPENDIX 6. Synonymized list of Fagaceae taxa that occur in Latin America (recognized names are in bold face).</p> <p>Colombobalanus excelsa (Lozano, Hern.Cam. &amp; Henao) Nixon = Trigonobalanus excelsa Lozano, Hern.Cam. &amp; Henao</p> <p>Erythrobalanus duqueana O.Schwarz = Quercus humboldtii Bonpl.</p> <p>Erythrobalanus tolimensis (Bonpl.) O.Schwarz = Quercus humboldtii Bonpl.</p> <p>Fagus grandifolia Ehrh. subsp. mexicana (Martínez) A.E. Murray = Fagus mexicana Martínez</p> <p>Fagus grandifolia var. mexicana (Martínez) Little = Fagus mexicana Martínez</p> <p>Fagus mexicana Martínez</p> <p>Quercus aaata C.H.Mull. = Quercus lancifolia Schltdl. &amp; Cham.</p> <p>Quercus acapulcensis Trel. = Quercus salicifolia Neé</p> <p>Quercus acatenangensis Trel. = Quercus laurina Bonpl.</p></div> 	https://treatment.plazi.org/id/794B8792FF9DFFBC3E88FC6BFD21F7F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FFD8FFCC3E88FC9CFC5DFD09.text	794B8792FFD8FFCC3E88FC9CFC5DFD09.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus brandegeei Goldman, Contr. U. S. Natl. Herb.	<div><p>Quercus brandegeei Goldman, Contr. U.S. Natl. Herb. 16: 321 (1916). Type:— MEXICO, Baja California Sur:</p> <p>E. W. Nelson &amp; E. A. Goldman 7475 (Holotype: US00089438!; isotypes: A!, NY!, UCSB (fragment)!). Distribution: Mexico (Baja California, Baja California Sur).</p> <p>Quercus breedloveana Nixon &amp; Barrie, Novon 25(4): 444, fig. 1 (2017). Type:— MEXICO, Guerrero: J. L. Contreras 1207 (Holotype: CAS (accn. 752961)!; isotypes: CAS!, FCME, MEXU!). This species is similar to Q. cortesii, and further study is necessary to determine if they are really different.</p> <p>Distribution: Mexico (Chiapas, Guerrero).</p> <p>Quercus bumelioides Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 188 (1854). Type:— PANAMA, Veraguas: B. Seemann s. n. (Feb 1849) (Holotype: K000575038!; isotype: BM!).</p> <p>Quercus copeyensis C.H.Mull., U.S.D.A. Bur. Pl. Industr. Misc. Publ. 477: 30, pl. 31, 32 (1942). Nom. nov. for Q. costaricensis f. kuntzei Trel. (1924) non Q. kunzei Gard. (1890).</p> <p>Quercus costaricensis Liebm. f. kuntzei Trel., Mem. Natl. Acad. Sci. 20: 146, pl. 283b (1924). Type:— COSTA RICA, Cartago: O. Kuntze 2282 (Holotype: NY; isotype: K!). The specimen at US of Kuntze 2282 does not match the specimen at K or the images in Trelease (1924), pl. 283b of the NY specimen so we do not recognize it as an isotype.</p> <p>Quercus pacayana C.H.Mull., U.S.D.A. Bur. Pl. Industr. Misc. Publ. 477: 30, pl. 30 (1942). Type:— GUATEMALA, Guatemala: P. C. Standley 58467 (Holotype: F0060040 F!; isotypes: F!, NA, UCSB (fragment)!).</p> <p>Distribution: Costa Rica, El Salvador, Guatemala, Honduras, Mexico (Chiapas), Nicaragua, Panama.</p> <p>Quercus calophylla Schltdl. &amp; Cham., Linnaea 5(1): 79 (1830). Type:— MEXICO, Veracruz: C. J. W. Schiede 24 (Holotype at B destroyed. Lectotype (designated here): HAL0060349!; isolectotypes: G (as s.n.)!, M!). At the time of publication of Q. calophylla, D.F.L. Schlechtendal was at B and was not transferred to HAL until 1833 (Stafleu &amp; Cowan 1985, p.190). For this reason, the Schiede 24 holotype would have almost certainly been at B and destroyed; the specimen at HAL should be considered an isotype. It is for this reason that we are lectotypifying the specimen as HAL rather than considering it to be the holotype.</p> <p>Quercus acuminata M.Martens &amp; Galeotti non Roxb. (1832), Bull. Acad. Roy. Sci. Bruxelles 10(1): 217 (1843). See Q. calophylla f. acuminata below.</p> <p>Quercus alamo Benth., Pl. Hartw. 55 (1840). Type:— MEXICO, Puebla: T. Hartweg 423 (Holotype: K000512985!; isotypes: A!, BM!, C (as s.n.)!, CAS!, CGE!, E!, FI!, G!, L!, LD (image)!, LE!, NY!, OXF!, P!, UCSB (fragment)!).</p> <p>Quercus calophylla f. acuminata Seemen ex Trel., Mem. Natl. Acad. Sci. 20: 203 (1924). Type:— MEXICO, Veracruz: H. Galeotti 96 (Lectotype: BR0000006912253! designated by Trelease (1924, p. 230, pl. 413); isolectotypes: BR!, G!, K!, P!). Quercus acuminata M.Martens &amp; Galeotti (1843) non Roxb. (1832) is a later homonym as well as having two syntypes, but the specific epithet of M.Martens &amp; Galeotti was used by Trelease, based on the same type which he effectively lectotypified, as the basis for Q. calophylla f. acuminata.</p> <p>Quercus calophylla f. alamo (Benth.) Trel., Mem. Natl. Acad. Sci. 20: 204 (1924).</p> <p>Quercus calophylla f. flavida (Liebm.) Trel., Mem. Natl. Acad. Sci. 20: 203 (1924).</p> <p>Quercus calophylla f. intermedia Seemen ex Trel., Mem. Natl. Acad. Sci. 20: 204, pl. 415 (1924). Type:— MEXICO, Veracruz: H. Galeotti 132 (Holotype: BR0000006913434!; isotypes: BR!, G!, K!, P!, UCSB (fragment)!). This form is based on the later homonym Q. intermedia M. Martens &amp; Galeotti (1843) non Boenn. ex Rchb. (1831), but is a legitimate form based on the same type.</p> <p>Quercus calophylla f. schiedeana Trel., Mem. Natl. Acad. Sci. 20: 203 (1924). Type:— MEXICO, Veracruz: C. A. Purpus s. n. (Dec 1914) (Lectotype: ILL00007467! designated by Valencia-A. et al. (1916, p. 256)).</p> <p>Quercus calophylla f. willdenovii Trel., Mem.Natl.Acad.Sci. 20: 204, pl. 411(1)(1924). Type:— MEXICO, Guanajuato: A. Bonpland 4270 (Holotype: B-W17613010!; isotypes: P!, UCSB (fragment)!).</p> <p>Quercus candicans auct. non Neé f. alligata Trel., Repert. Spec. Nov. Regni Veg. 33: 319 (1934). Type:— MEXICO, Guerrero: C. Toledo &amp; R. Landa 910 (Syntypes at B destroyed. Neotype: FCME designated by Valencia-A. et al. (2018, p. 256); isoneotype: MEXU (image)!).</p> <p>Quercus candicans auct. non Neé f. incurva Trel., Mem. Natl.Acad. Sci. 20: 205, pl. 418(2) (1924). Type:— MEXICO, Jalisco: J. N. Rose 2999 (Lectotype (designated here): US01013695!; isolectotypes: GH!, NY!, US!). Trelease (1924, p. 205, 230) cited just one collection number, but he did not indicate where the specimen was deposited.</p> <p>Quercus candicans auct. non Neé f. michoacana Trel., Mem. Natl. Acad. Sci. 20: 204, pl. 418(1) (1924). Type:— MEXICO, Michoacán: C. G. Pringle 3955 (Lectotype: US01013694! designated by Valencia-A. et al. (1918, p. 256); isolectotypes: A!, BM!, BR!, E!, F!, G!, HBG (image)!, K!, KFTA (image)!, M!, MEL (image)!, MEXU!, MICH!, MO!, MU (image)!, NY!, P!, PH!, S (image)!, UC!, VT!). Although Trelease designated Pringle 3955 as the type collection, we have not found any specimens that match the photograph in plate 418(1).</p> <p>Quercus chimaltenangana Trel. f. gemmata C.H.Mull., Amer. Midl. Naturalist 18(5): 855 (1937). Type:— GUATEMALA, Quiche: A. F. Skutch 1654 (Holotype: ILL00007436!; isotypes: A!, F!, NY!, UCSB (fragment)!, US!).</p> <p>Quercus flavida Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 178 (1854). Type:— MEXICO, Puebla: F. M. Liebmann 3478 (Lectotype (designated here): C 10012466!; isolectotype: CAS!). Liebmann 3478 was represented at C by two specimens with sheet numbers of Cupulif. Libm. 1 and Cupulif. Libm. 2. On p. 203 Trelease (1924) wrote “ Liebmann, 2, 3478, May, 1841 the type of Q. flavida Liebmann ” and on p. 230 in the caption to pl. 412(2) he wrote “The type of Q. flavida (Mexico, Liebmann, 2). Photographed by permission at Copenhagen.” The image in pl. 412(2), however, Liebmann 3478 Cupulif. Lbm. 1 is now at CAS and Liebmann 3478 Cupulif. Lbm. 2 is at C. Because of this ambiguity, we are lectotypifying the C specimen.</p> <p>Quercus intermedia M.Martens &amp; Galeotti non Boenn. ex Rchb. (1831), Bull. Acad. Roy. Sci. Bruxelles 10(1): 223 (1843). See Q. calophylla f. intermedia above.</p> <p>Quercus tonaguiae Trel., Mem. Natl. Acad. Sci. 20: 190, pl. 385 (1924). Type:— MEXICO, Oaxaca: H. Galeotti 105 (Holotype at B destroyed. Lectotype (designated here): BR0000030524842!; isolectotypes: G!, K!, P!). The holotype at B was Galeotti s. n. Trelease (1924, p. 190) wrote that the specimen in Berlin was “referred to be Martens and Galeotti, p. 224, under no. 105, to Q. acutifolia.” For this reason we are lectotypifying the BR number as 105 which were the specimens referred to by M. Martens &amp; Galeotti (1843, p. 224). The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. calophylla as broadly interpreted.</p> <p>Distribution: Guatemala, Mexico (Chiapas, Chihuahua, Ciudad de México, Colima, Durango, Guanajuato, Guerrero,</p> <p>Hidalgo, Jalisco, México, Michoacán, Morelos, Nayarit, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa, Sonora,</p> <p>Tamaulipas, Tlaxcala, Veracruz, Zacatecas). The occurrence of Q. calophylla in Tamaulipas reported by Villaseñor</p> <p>(2016) needs to be confirmed.</p> <p>Quercus canbyi Trel., Mem. Natl. Acad. Sci. 20: 188, pl. 379b (1924). Type:— MEXICO, Nuevo León: C. G. Pringle 2393 (Holotype at B destroyed. Lectotype (designated here): M-0244476!; isolectotypes: AC!, BM!, BR!, CAS!, E!, ENCB (image) !, F!, G!, GH!, HBG (image) !, K!, MEL (image)!, MEXU!, MICH!, MO!, NDG (image)!, NY!, P!, PH!, S (image) !, UC!, UCSB (fragment) !, US!, VT !).</p> <p>Quercus alamarensis C.H.Mull., J. Arnold Arbor. 17(3): 174 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 613 (Holotype: A00055614!; isotypes: MICH!, UCSB (fragment)!).</p> <p>Quercus canbyi f. ascendens Trel., Mem. Natl. Acad. Sci. 20: 188, pl. 380a (1924). Type:— MEXICO, Nuevo León: C. G. Pringle 11705 (Holotype at B destroyed. Lectotype (designated here): US00089467!; isolectotypes: A!, C!, CAS!, GH!, K!, L!, LE!, MICH!, MO!, L!, VT!).</p> <p>Quercus canbyi f. attenuata C.H.Mull., J.Arnold Arbor. 17(3): 175 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 1347 (Holotype: A00055638!; isotypes: TEX (image)!, UCSB (fragment)!).</p> <p>Quercus canbyi f. berlandieri Trel., Mem. Natl. Acad. Sci. 20: 188, pl. 380b (1924). Type:— MEXICO, Tamaulipas: J. Berlandier 2221 (Holotype: G; isotypes: CAS!, G!, GH!, K!, NY!, P!, UCSB (fragment)!). There are several sheets at G marked as isotypes, but none of them match the holotype shown in Trelease (1924) pl. 380b.</p> <p>Quercus canbyi f. concolor Trel. &amp; C.H.Mull., Bull. Torrey Bot. Club 63(3): 152 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 590 (p.p.) (Holotype: ILL00007466!; isotypes:TEX (image)!,UCSB (fragment)!). The specimen of C. H. Muller &amp; M. T. Muller 590 at A is Q. glaucoides, so it is not an isotype.</p> <p>Quercus canbyi f. karwinskii (Trel.) C.H.Mull., J. Arnold Arbor. 17(3): 175 (1936).</p> <p>Quercus canbyi f. pedunculata C.H.Mull., J. Arnold Arbor. 17(3): 175 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 1009 (Holotype: A00055641!; isotype: UCSB (fragment)!).</p> <p>Quercus canbyi f. setacea C.H.Mull., J. Arnold Arbor. 17(3): 175 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 1011 (Holotype: A00055640!; isotype: UCSB (fragment)!).</p> <p>Quercus canbyi f. subovatifolia C.H.Mull., J. Arnold Arbor. 17(3): 175 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 1346 (Lectotype (designated here): A00069937!; isolectotype: UCSB (fragment)!). Muller (1936) gave this form a name and cited three collection numbers, but he did not designate a type.</p> <p>Quercus karwinskii Trel., Mem. Natl. Acad. Sci. 20: 188, pl. 379a (1924). Type:— MEXICO: W. F. Karwinski s. n. (Holotype: M!). Trelease (1924. p. 189) wrote “Without data (Karwinski, the type in the Munich herbarium)” and on p. 228 for the caption to pl. 379b he wrote “The type (Mexico, Karwinski).” Plate 379b shows that the sheet at M has the number 464 written on the linen tape holding down the specimen. The tradition of writing numbers on the linen tape is seen on other M specimens and is not to be considered a collection number but rather a consecutive numbering done at M some time during the 19th century for specimens in each family (H.-J. Esser, pers. comm.).</p> <p>Distribution: Mexico (Chihuahua, Coahuila, Hidalgo, Nuevo León, San Luis Potosí, Tamaulipas).</p> <p>Quercus castanea Neé, Anales Ci. Nat. 3: 276 (1801). Type:— MEXICO, Hidalgo: L. Neé s. n. (1791) (Lectotype: MA25950! designated by Bartholomew &amp; Almeda (2022, p. 28); isolectotypes: MA!, UCSB (as to larger leaf from MA lectotype).</p> <p>Quercus axillaris E.Fourn. ex Trel., Mem. Natl. Acad. Sci. 20: 176, pl. 352a (1924). Type:— MEXICO, Ciudad de México: E. Bourgeau 1138 (Syntypes at B destroyed and at G. Lectotype (designated here): G (00410066) !; isolectotypes: BR!, C!, GH!, K!, MO (fragment)!, MPU!, P!, S (image) !, UCSB (image)!, US!). Trelease (1924, p. 227, pl. 352) cites the type as being both at B and G. There is a sheet at G marked as an isotype which is in fact an isosyntype as it does not match the syntype in Trelease (1924) pl. 352a. We are lectotypifying the name based on the existing isosyntype at G.</p> <p>Quercus castanea var. sublobata A.DC., Prodr. 16(2.1): 72 (1864). Type:— GUATEMALA: J. R. R. Warszewicz 10 (Holotype: G!).</p> <p>Quercus circummontana Trel., Mem. Natl. Acad. Sci. 20: 177, pl. 356 (1924). Type:— MEXICO, Veracruz: F. M. Liebmann 3455 (Lectotype (designated here): C 10012544!; isolectotypes: C!, CAS!, MO!, NY!, UCSB (fragment)!). Of the three syntype sheets of 3455 originally at C, one is now at CAS. Trelease (1924, pl. 356) shows closeup images of two sheets of 3455 at C. The bottom image in plate 356 is selected as the lectotype, and the top image is the isolectotype at CAS.</p> <p>Quercus consociata Trel., Mem. Natl. Acad. Sci. 20: 172, pl. 345a (1924). Type:— GUATEMALA: J. R. R. Warszewicz 9 (Holotype at B destroyed. Lectotype (designated here): JE00006775!).</p> <p>Quercus crassivenosa Trel., Repert. Spec. Nov. Regni Veg. 33: 316 (1934). Type:— MEXICO, Guerrero: L. Schultze 33 (Holotype at B destroyed. Lectotype (designated here): ILL00007524! (fragment)).</p> <p>Quercus impressa Trel., Mem. Natl. Acad. Sci. 20: 181, pl. 364a (1924). Type:— MEXICO, Oaxaca: C. G. Pringle 6029 (Holotype at B destroyed. Lectotype (designated here): P00754104 !; isolectotypes: A!, AC!, BM!, BR!, CAS!, E!, ENCB (image)!, G!, GH!, HBG (image) !, K!, KFTA (image)!, MEL (image)!, MEXU!, MO!, NDG (image)!, NY!, P!, PH!, S (image) !, UCSB (fragment) !, UC!, US!, VT !).</p> <p>Quercus lanigera M.Martens &amp; Galeotti, Bull.Acad. Roy. Sci. Bruxelles 10(1): 215 (1843). Type:— MEXICO, Oaxaca: H. Galeotti 104 (Lectotype: BR0000006912321! designated by Trelease (1924, p. 228, pl. 354); isolectotypes: BR!, G!, K!, P!, UCSB (fragment)!). Although M. Martens &amp; Galeotti (1843) cite only one collection, there are two sheets of this number at BR marked as types, one of which one was effectively lectotypified by Trelease.</p> <p>Quercus lanigera f. sideroxyloides Trel., Mem. Natl. Acad. Sci. 20: 177, pl. 355 (1924). Type:— MEXICO, Puebla: F. G. Nicolas s. n. (1912) (Holotype: F!).</p> <p>Quercus mucronata Willd., Sp. Pl., ed. 4 4(1): 436 (1805). This is a superfluous name as the older name Q. castanea (1801) is erroneously cited as a synonym.</p> <p>Quercus pulchella Bonpl., Pl. Aequinoct. 2: 44, pl. 88 (1809). Type:— MEXICO, Michoacán: A. Bonpland 4230 (Holotype: P00129710!; isotypes: P (as s.n.)!, UCSB (fragment)!).</p> <p>Quercus rossii Trel., Mem. Natl. Acad. Sci. 20: 179, pl. 362b (1924). Type:— MEXICO, Michoacán: H. Ross 374 (Holotype at B destroyed. Lectotype (designated here): M-0244474!; isolectotypes UCSB (fragment from M lectotype)!).</p> <p>Quercus scherzeri Trel., Mem. Natl. Acad. Sci. 20: 172, pl. 343b (1924). Type:— HONDURAS, Comayagua: K. R. von Scherzer s. n. (Apr 1854) (Lectotype (designated here): image in Trelease (1924, pl. 343b)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Quercus seleri Trel., Mem. Natl. Acad. Sci. 20: 181, pl. 364b (1924). Type:— MEXICO, Morelos: C. Seler &amp; E. G. Seler 4302 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 364b)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Quercus serrulata Trel., Mem. Natl. Acad. Sci. 20: 179, pl. 361a (1924). Type:— MEXICO, Jalisco: M. E. Jones 445 (Holotype: US 00089640!).</p> <p>Quercus spathulistipula Trel., Repert. Spec. Nov. Regni Veg. 33: 319 (1934). Type:— MEXICO, Guerrero: L. Schultze 332 (Holotype: at B destroyed). The determination of Q. spathulistipula as Q. castanea is based on the Trelease (1934) description because no images or original specimens exist.</p> <p>Quercus subcrispata Trel., Mem. Natl. Acad. Sci. 20: 176, pl. 353 (1924). Type:— MEXICO, Puebla: G. Arsène 5926 (Holotype: US 00089662!; isotypes: BM!, ILL!, MPU!, PH!).</p> <p>Quercus tepoxuchilensis Trel., Mem. Natl. Acad. Sci. 20: 180, pl. 363a (1924). Type:— MEXICO, Puebla: G. Arsène 201 (Holotype at B destroyed. Lectotype (designated here): US 00089678!; isolectotypes: MEXU!, US!).</p> <p>Quercus tepoxuchilensis f. perplexans Trel., Mem. Natl.Acad. Sci. 20: 180 (1924). Type:— MEXICO, Puebla: G. Arsène 2068 (Holotype: ILL00007569!; isotypes: MO!, MPU!, US!).</p> <p>Quercus tristis Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 174 (1854). Type:— MEXICO, Veracruz: F. M. Liebmann 3455 (Lectotype (designated here): C 10012544!; isolectotypes: C!, CAS!, MO!, NY!). Trelease (1924, p. 227), in the caption to pl. 341, indicated two collections as the types, so this did not constitute lectotypification. The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. castanea as broadly interpreted.</p> <p>Quercus tristis f. mixcoensis Trel., Mem. Natl. Acad. Sci. 20: 172 (1924). Type:— GUATEMALA, Sacatepequez: W. Trelease 31 (Holotype: ILL00007553!).</p> <p>Quercus tristis f. niederleini Trel., Mem.Natl.Acad.Sci. 20: 171, pl.343a (1924). Type:— EL SALVADOR: G. Niederlein s. n. (9 Jan 1898) (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 343a)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Quercus tristis f. sublobata (A.DC.) Trel., Mem. Natl. Acad. Sci. 20: 171 (1924).</p> <p>Quercus tristis f. vulcani Trel., Mem. Natl. Acad. Sci. 20: 171, pl. 342b (1924). Type:— GUATEMALA, Escuintla: K. R. von Scherzer s. n. (Holotype at W destroyed. Lectotype (designated here): image in Trelease (1924, pl. 342b)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Quercus verrucosirama Trel., Repert. Spec. Nov. Regni Veg. 33: 319 (1934). Type:— MEXICO, Guerrero: L. Schultze 382 (Holotype: at B destroyed). The determination of Q. verrucosirama as probably Q. castanea is based on the Trelease (1934) description as no images or original specimens exist.</p> <p>Distribution: El Salvador, Guatemala, Honduras, Mexico (Aguascalientes, Chiapas, Chihuahua, Ciudad de México,</p> <p>Coahuila, Colima, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, México, Michoacán, Morelos, Nayarit, Nuevo</p> <p>León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa, Sonora, Tamaulipas, Tlaxcala, Veracruz, Zacatecas).</p> <p>The occurrence of Q. castanea in Chihuahua, Colima, and Nuevo León reported by Villaseñor (2016) needs to be confirmed.</p> <p>Quercus cedrosensis C.H.Mull., Madroño 16(6): 188, fig. 1–3, 5 (1962). Type:— MEXICO, Baja California Sur: C. H. Muller 10775 (Holotype: UCSB000021!; isotypes: CAS!, G!, UCSB (fragment)!, US!, VT!).</p> <p>Distribution: Mexico (Baja California, Baja California Sur). Quercus chrysolepis Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 173 (1854). Type:— U.S.A., California: T. Hartweg 1962 (Lectotype: K000832349! designated by Trelease (1924, p. 222, pl. 209); isolectotypes: K!, UCSB (fragment from K lectotype)!). Liebmann (1854, p. 173) cited six syntypes.</p> <p>Distribution: Mexico (Baja California, Chihuahua, Sonora), U.S.A. (Arizona, California, Nevada, New Mexico,</p> <p>Oregon).</p> <p>Quercus coahuilensis Nixon &amp; C.H.Mull., Brittonia 45(2): 150, fig. 2 (1993). Type:— MEXICO, Coahuila: I. M. Johnston &amp; C. H. Muller 567 (Holotype: BH (image)!). This species is part of the Q. hypoxantha complex that could also be treated in a broad sense as a single species.</p> <p>Distribution: Mexico (Chihuahua, Coahuila).</p> <p>Quercus coffeicolor Trel., Mem. Natl. Acad. Sci. 20: 137, pl. 261a (1924). Type:— MEXICO, Sinaloa: J. N. Rose 1758 (Holotype: US00089482!; isotypes: GH!, K!, MO!, NY!, UCSB (fragment)!).</p> <p>Quercus aequivenulosa Trel., Mem. Natl. Acad. Sci. 20: 138, pl. 261b (1924). Type:— MEXICO, Nayarit: J. N. Rose 2231 (Holotype: US00089387!; isotypes: GH!, NY!, UCSB (fragment)!).</p> <p>Quercus mexiae L.M.González, Ibugana 9: 60, fig. 5–7 (2018). Type:— MEXICO, Jalisco: L.M.González V. &amp; J.A. Pérez de la Rosa 5017 (Holotype: IBUG0177437). Although we have not seen the holotype, from the description and from those paratypes seen by Breedlove and us, we consider that Q. mexiae is not sufficiently different to justify it as a distinct species from Q. coffeicolor.</p> <p>Quercus prainiana Trel. Non H. Lév. (1913),Mem. Natl.Acad. Sci. 20:138, pl. 262a, pl. 262b (1924). Type:— MEXICO, Jalisco: C. G. Pringle 8854 (Syntypes: C 10012523!, K000512914!; isosyntypes: A!, BM!, CAS!, COLO (image)!, E!, ENCB (image)!, F!, G!, GH!, HBG (image)!, ILL!, L!, LL (image)!, MEXU!, MICH!, MO!, MSC (image)!, NY!, P!, PH!, UC!, UCSB (fragment)!, US!, VT!). Trelease (1924, p. 138) designated a type collection number, but on p. 224 in the caption to pl. 262 he did not specify which herbarium, C or K, was to house the holotype. In Plate 262 he has images of one specimen at C and one at K, both of which he called the type which makes both the C and K specimens syntypes. Because this species is a later homonym we are not lectotypifying the name.</p> <p>Distribution: Mexico (Colima, Durango, Jalisco, Nayarit, Querétaro, San Luis Potosí, Sinaloa, Zacatecas).</p> <p>Quercus confertifolia Bonpl., Pl. Aequinoct. 2: 53, pl. 94 (1809). Type:— MEXICO, Guanajuato: A. Bonpland 4407 (Holotype: P00129738!; isotypes: P!, UCSB (fragment)!). Although lectotypified by Valencia-A. &amp; Coombes (2020, p. 89), this was not necessary because the holotype at P has the specimen label in Bonpland’s hand. The isotype most likely was added to P or identified as such at a later date.</p> <p>Quercus castanea Neé var. glabrata (Liebm. ex Seem.) A.DC., Prodr. 16(2.1): 72 (1864).</p> <p>Quercus gentryi C.H.Mull., Amer. Midl. Naturalist 27(2): 474 (1942). Type:— MEXICO, Sinaloa: H. S. Gentry 5686 (Holotype: US 00089541!; isotypes: DS!, GH!, MEXU!, MICH!, MO!, NY!, RSA (image)!, UCSB (fragment from US holotype)!, UCSB (fragment)!). The holotype at US was formerly at NA.</p> <p>Quercus mexicana Bonpl. f. confertifolia (Bonpl.) Trel., Mem. Natl. Acad. Sci. 20: 174 (1924).</p> <p>Quercus mexicana f. glabrata Trel., Mem. Natl. Acad. Sci. 20: 174 (1924). Type:— MEXICO, Jalisco: H. Galeotti 118 (Lectotype (designated here): BR0000005105380!; isolectotypes: BR!, G!, P!). We are lectotypifying one of the syntypes.</p> <p>Quercus mexicana var. confertifolia (Bonpl.) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 209 (1884).</p> <p>Quercus mexicana var. glabrata Liebm. ex Seem., Bot. Voy. Herald 332 (1856). Type:— MEXICO, Sinaloa: B. C. Seemann 1974 (Holotype: K000512942!; isotypes: BM!, C!, GH!). Valencia-A. &amp; Coombes (2020, p. 90) designated Seemann 1974 at BM (BM000643467!) as the lectotype, but this is an invalid lectotypification, because the holotype as designated by Seemann is in the Hooker Herbarium at K and is the specimen with the herbarium name of Quercus mexicana var. glabrata in Liebmann’s hand.</p> <p>Distribution: Mexico (Aguascalientes, Chihuahua, Colima, Durango, Guanajuato, Hidalgo, Jalisco, México,</p> <p>Michoacán, Nayarit, Querétaro, Sinaloa, Zacatecas).</p> <p>Quercus conzattii Trel., Proc. Amer. Philos. Soc. 60: 33, pl. 4 (1921). Type:— MEXICO, Oaxaca: C. Conzatti 1900 (Holotype: F0060021 F!; isotype: MEXU!).</p> <p>Quercus urbani Trel. f. parvifolia Martínez, Anales Inst. Biol. Univ. Nac. Autón. México 37(1 &amp; 2): 93 (1966). This form is based on Q. conzattii, so the name has the same type.</p> <p>Distribution: Mexico (Durango, Guerrero, Jalisco, Nayarit, Oaxaca, Zacatecas). Quercus cornelius-mulleri Nixon &amp; K.P.Steele, Madroño 28(4): 210, fig. 4, 5 (1981). Type:— U.S.A., California: K. C. Nixon &amp; K. P. Steele 2765 (Holotype: BH; isotypes: DAV, GH!, MO!, NY!, RSA (image)!, SD (image)!, TEX (image)!, UC!).</p> <p>Distribution: Mexico (Baja California), U.S.A. (California).</p> <p>Quercus corrugata Hook., Icon. Pl. 5: sub pl. 403, 404 (1842). Type:— GUATEMALA, San Marcos: G. V. Skinner 5 (Holotype: K000512592!; isotypes: C (as s.n.?)!, K!, MPU!). The occurrence in Mexico by Villaseñor (2016) may actually be collections of Q. lancifolia Schltdl. &amp; Cham., but additional study is needed.</p> <p>Quercus corrugata var. granulifera Trel., Mem. Natl. Acad. Sci. 20: 45, pl. 24 (1924). Type:— GUATEMALA: J. R. R. Warszewicz 11 (Holotype at B destroyed. Lectotype (designated here): G00410063!; isolectotypes: K!, P!, UCSB (fragment)!).</p> <p>Quercus corrugata var. ipalensis Trel., Mem. Natl. Acad. Sci. 20: 45 (1924). Type:— GUATEMALA, Chiquimula: H. Pittier 1869 (Holotype: US00089474!). The specimen at US is marked by Trelease and in his hand as “ Q. corrugata iipalensis Trel. ”</p> <p>Quercus corrugata var. microcarpa Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 192 (1884). Type:— COSTA RICA, San José: Polakovsky s. n. (Holotype: at B destroyed). The determination is based on the Wenzig (1884) description as no images or original specimens exist.</p> <p>Distribution: Costa Rica, Guatemala.</p> <p>Quercus cortesii Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 175 (1854). Type:— MEXICO, Veracruz: F. M. Liebmann 3463 (Lectotype: C10013674! designated by Trelease (1924, p. 228, pl. 373b); isolectotypes: G! UCSB (fragment from C lectotype)!).</p> <p>Quercus anglohondurensis C.H.Mull., U.S.D.A. Bur. Pl. Industr. Misc. Publ. 477: 76, pl. 114, 115 (1942). Type:— BELIZE, El Cayo: C. L. Lundell 6615 (Holotype: MICH1109366!; isotypes: DS!, GH!, MEXU!, MO!, NY!, TEX (image)!, UCSB (fragment)!, US!, WIS!).</p> <p>Quercus brenesii Trel., Mem. Natl. Acad. Sci. 20: 186, pl. 377b (1924). Type:— COSTA RICA, Alajuela: A. Brenes 14520 (Holotype at B destroyed. Lectotype (designated here): US 00089436!; isolectotypes: GH!, K!, UCSB (fragment)!, US!).</p> <p>Quercus gracilior C.H.Mull., U.S.D.A. Bur. Pl. Industr. Misc. Publ. 477: 77, pl. 116 (1942). Type:— HONDURAS, Comayagua: J. B. Edwards 285 (Holotype: A00055575!; isotypes: F!, UCSB (fragment)!).</p> <p>Quercus gulielmi-treleasei C.H.Mull., U.S.D.A. Bur. Pl. Industr. Misc. Publ. 477: 58, pl. 79, 80 (1942). Type:— PANAMA, Chiriquí: R. J. Seibert 226 (Holotype: MO-143075!).</p> <p>Quercus huitamalcana Trel., Contr. U.S. Natl. Herb. 23(2): 196 (1922). Type:— MEXICO, Puebla: F. M. Liebmann 3560 (Lectotype: C10013681! designated by Trelease (1924, p. 186); isolectotypes: C!, CAS!, UCSB (fragment from C lectotype)!). Nom. nov. for Q. serra Liebm. (1854) non Unger (1845). The lectotype is sheet Cupulif. Liebm. 31 of Liebmann 3560.</p> <p>Quercus paxtalensis C.H.Mull., U.S.D.A. Bur. Pl. Industr. Misc. Publ. 477: 75, pl. 113 (1942). Type:— MEXICO, Chiapas: E. Matuda 1724 (Holotype: MICH1109345!; isotypes: A!, K!, LL (image)!, MO!, NY!, US!, VT!).</p> <p>Quercus serra Liebm. non Unger (1845), Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 174 (1854). See Q. huitamalcana above.</p> <p>Quercus tenuiaristata Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 17(4): 358 (1938). Type:— HONDURAS, Comayagua: T. G. Yuncker et al. 6204 (Holotype: ILL00007574!; isotypes: F!, G!, GH!, K!, MICH!, MO!, NY!, UCSB (fragment)!).</p> <p>Distribution: Belize, Costa Rica, El Salvador, Honduras, Mexico (Chiapas, Colima, Guerrero, Jalisco, Oaxaca, Puebla,</p> <p>Tamaulipas, Veracruz), Nicaragua, Panama.</p> <p>Quercus costaricensis Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 184 (1854). Type:— COSTA RICA, Cartago: A. S. Oersted 3465 (Lectotype (designated here): C 10012470!; isolectotypes: BM!, C!, CAS!, G!, UCSB (fragment)!). Trelease (1924, p. 225) in the caption to pl. 281 wrote “The type (Costa Rica, Oersted, 3465).” In pl. 281 he shows photos taken of the two syntypes now at C, so Trelease’s statement cannot be considered effective lectotypification. We are lectotypifying the specimen shown in the bottom photo of pl. 281.</p> <p>Quercus endresii Trel., Mem. Natl. Acad. Sci. 20: 145, pl. 280 (1924). Type:— COSTA RICA: A. R. Endres s. n. (Holotype at W destroyed. Lectotype (designated here): image in Trelease (1924, pl. 280)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Quercus irazuensis Kuntze, Revis. Gen. Pl. 2: 641 (1891). Type:— COSTA RICA, Cartago: O. Kuntze 2344 (Holotype: NY00253700!; isotypes: K!, UCSB (fragment)!).</p> <p>Quercus wesmaelii Trel., Mem. Natl. Acad. Sci. 20: 172, pl. 344a (1924). Type:— COSTA RICA, Alajuela: H. Pittier 773 (Holotype at B destroyed. Lectotype (designated here): BR0000006914523!; isolectotype: BR!).</p> <p>Distribution: Costa Rica, Panama.</p> <p>Quercus crassifolia Bonpl., Pl. Aequinoct. 2: 49, pl. 91 (1809). Type:— MEXICO, Guerrero: A. Bonpland 4142 (Holotype: P00129735!; isotype: P!).</p> <p>Quercus brachystachys Benth., Pl. Hartw. 91 (1842). Type:— GUATEMALA, Guatemala: T. Hartweg 618 (Holotype: K000512503!; isotypes: BM!, CGE!, K!, LD (image)!).</p> <p>Quercus chicamolensis Trel., Contr. U.S. Natl. Herb. 23(2): 190. (1922). Type:— MEXICO, Oaxaca: H. Galeotti 102 (Holotype: BR0000005104734!; isotypes: BR!, G!, P!, UCSB (fragment)!). Nom. nov. for Q. mollis M.Martens &amp; Galeotti (1843) non Raf. (1838). Although Trelease (1924, p. 224) stated that the specimen in pl. 246 at BR was the type, this specimen is an isotype. The holotype is the specimen from M. Martens’ own herbarium now at BR.</p> <p>Quercus crassifolia var. errans (Trel.) E.F.Warb., Bull. Misc. Inform. Kew 1939 (2): 90 (1939).</p> <p>Quercus errans Trel., Mem. Natl. Acad. Sci. 20: 131, pl. 245a (1924). Type:— MEXICO, Hidalgo: T. Hartweg 424 (Holotype at B destroyed. Lectotype (designated here): G00177569!; isolectotypes: A!, CGE!, G!, K!, LE!, NY!, OXF!, P!). There is no extant collection of the holotype, Hahn 667, which was collected at Perote in Veracruz, Mexico. From the number sequence, Hahn 667 would have been collected in 1865. There is a specimen of Hahn s. n. (1 Jul 1866) both at K and P, but this collection cannot be considered an isotype, as it was collected a year later than the holotype, even though it was collected at the same locality and is an excellent match for the photo of the holotype in Trelease (1924), pl. 245a. On page Trelease (1924) cites several paratypes which include Ehrenberg 895, 896, 920, 1021, and 1284 which, although not stated, were probably all seen by him at B and were destroyed. Of these five collections, the only one that is extant is Ehrenberg 896 from Hidalgo, Mexico now at NY and VT. The photo in pl. 245b is of Hartweg 424 which Trelease also states is “a second collection” of Q. errans, so it is also a paratype. The photo of this Hartweg collection was also taken of the specimen at B which is now destroyed. We are choosing Hartweg 424 at G as the lectotype, which is also the lectotype of Q. splendens var. pallidior.</p> <p>Quercus felipensis Trel., Mem. Natl. Acad. Sci. 20: 129, pl. 237 (1924). Type:— MEXICO, Oaxaca: C. L. Smith 776 (Holotype: MO-144156!; isotype: VT!).</p> <p>Quercus miguelitensis Trel., Mem. Natl. Acad. Sci. 20: 132, pl. 249b (1924). Type:— MEXICO, San Luis Potosí: J. G. Schaffner 897 (Holotype: GH00055673!; isotype: K!).</p> <p>Quercus mollis M.Martens &amp; Galeotti non Raf. (1838), Bull. Acad. Roy. Sci. Bruxelles 10(1): 216 (1843). See Q. chicamolensis above.</p> <p>Quercus moreliana Trel., Mem. Natl. Acad. Sci. 20: 130, pl. 241 (1924). Type:— MEXICO, Michoacán: G. Arsène 6681 (Holotype: US 00089579!).</p> <p>Quercus orbiculata Trel., Mem. Natl.Acad. Sci. 20: 132, pl. 249a (1924). Type:— MEXICO, San Luis Potosí: C. C. Parry &amp; E. Palmer 836 (Holotype: GH00062252!; isotypes: E!, F!, K!, MO!, NY!, P!, PH!, UCSB (fragment)!, US!, WIS!).</p> <p>Quercus spinulosa M.Martens &amp; Galeotti, Bull. Acad. Roy. Sci. Bruxelles 10(1): 218 (1843). Type:— MEXICO, Veracruz: H. Galeotti 114 (Holotype: BR0000006916114!; isotypes: G!, P!, UCSB (fragment)!).</p> <p>Quercus splendens Neé var. pallidior A.DC., Prodr. 16(2.1): 57 (1864). Type:— MEXICO, Hidalgo: T. Hartweg 424 (Lectotype: G00177569! designated by Trelease (1924, p. 224, pl. 243a); isolectotypes: A!, CGE!, G!, K!, LE!, NY!, OXF!, P!). A. de Candolle (1864, p. 57) cited two syntypes.</p> <p>Quercus stipularis Bonpl., Pl. Aequinoct. 2: 47, pl. 90 (1809). Type:— MEXICO, Veracruz: A. Bonpland s. n. (Holotype: P00129748!; isotypes: P!, UCSB (fragment)!).</p> <p>Distribution: Guatemala, Honduras, Mexico (Aguascalientes, Chiapas, Chihuahua, Ciudad de México, Coahuila,</p> <p>Colima, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, México, Michoacán, Morelos, Nayarit, Nuevo León, Oaxaca,</p> <p>Puebla, Querétaro, San Luis Potosí, Sinaloa, Sonora, Tamaulipas, Tlaxcala, Veracruz, Zacatecas). The occurrence of Q.</p> <p>crassifolia in Coahuila, Colima, Sinaloa, and Tamaulipas reported by Villaseñor (2016) needs to be confirmed.</p> <p>Quercus crassipes Bonpl., Pl. Aequinoct. 2: 37, pl. 83 (1809). Type:— MEXICO, Guanajuato: A. Bonpland s. n. (Holotype: P00129734!; isotypes: HAL!, P!).</p> <p>Quercus colimae Trel., Mem. Natl. Acad. Sci. 20: 174, pl. 349a (1924). Type:— MEXICO, Jalisco: Burnett s. n. (1913) (Holotype: USFS). We have been unable to find the holotype. The synonomy is based on Trelease (1924) pl. 349a and the description.</p> <p>Quercus colimae var. zauzillo Trel., Mem. Natl. Acad. Sci. 20: 175, pl. 349b (1924). Type:— MEXICO, Jalisco: Burnett s. n. (1913) (Holotype: USFS). We have been unable to find the holotype. The synonomy is based on Trelease (1924) pl. 349b and the description.</p> <p>Quercus crassipes var. angustifolia Bonpl., Pl. Aequinoct. 2: 38, pl. 84 (1809). Type:— MEXICO, Guanajuato: A. Bonpland s. n. (Holotype: P00129729!; isotypes: HAL!, P!, UCSB (fragment)!).</p> <p>Quercus cuajimalpana Trel., Mem. Natl. Acad. Sci. 20: 165, pl 324 (1924). Type:— MEXICO, Ciudad de México: G. Arsène 8866 (Holotype: US00089472!; isotypes: A!, F!, GH!, ILL!, L!, MO!, MPU!, NY!, UCSB (fragment)!).</p> <p>Quercus malifolia Trel., Mem. Natl. Acad. Sci. 20: 175, pl. 348b (1924). Type:— MEXICO, Veracruz: H. Ross 1256 (Holotype at B destroyed. Lectotype (designated here): M-0244472!; isolectotype UCSB (fragment from M lectotype)!). It is likely that the holotype at B was actually Ross 1256 rather than Ross 256 which was also the conclusion of C. H. Muller who removed some fragments of the M specimens that are now at UCSB.</p> <p>Quercus mexicana Bonpl. f. angustifolia (Bonpl.) Trel., Mem. Natl. Acad. Sci. 20: 174, pl. 345 (1924).</p> <p>Quercus mexicana f. lanosa Trel., Mem. Natl. Acad. Sci. 20: 174, pl. 347a (1924). Type:— MEXICO, Michoacán: T. Hartweg 431 (Holotype at B destroyed. Lectotype (designated here): K000512959!; isolectotypes: BM!, CGE!, E!, G!, K!, L!, LE!, NY!, OXF!, P!, UCSB (fragment)!).</p> <p>Quercus mexicana f. perfertilis Trel., Mem. Natl. Acad. Sci. 20: 174 (1924). Type:— MEXICO, Veracruz: J. N. Rose &amp; R. Hay 5675 (Lectotype (designated here): US00089580!; isolectotype: A!). Trelease (1924) did not specify the place of deposit for the type collections.</p> <p>Quercus obovalifolia E.Fourn. ex Trel., Mem. Natl. Acad. Sci. 20: 175, pl. 351b (1924). Type:— MEXICO, Ciudad de México: E. Bourgeau 1014 (Holotype: C 10012505!; isotypes: BM!, G!, K!, P!, UCSB (fragment from C holotype)!, UCSB (fragment)!).</p> <p>Distribution: Mexico (Aguascalientes, Chiapas, Ciudad de México, Coahuila, Colima, Durango, Guanajuato,</p> <p>Guerrero, Hidalgo, Jalisco, México, Michoacán, Morelos, Nayarit, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis</p> <p>Potosí, Tamaulipas, Tlaxcala, Veracruz, Zacatecas).</p> <p>Quercus crispipilis Trel., Mem. Natl. Acad. Sci. 20: 184, pl. 370b (1924). Type:— GUATEMALA, Huehuetenango: C. Seler &amp; E. G. Seler 2667 (Holotype at B destroyed. Lectotype (designated here): GH00069875!; isolectotypes: NY!, UCSB (fragment from GH lectotype)!, UCSB (fragment)!).</p> <p>Quercus cerifera Trel., Mem. Natl. Acad. Sci. 20: 184, pl. 370a (1924). Type:— MEXICO, Chiapas: O. F. Cook 116 (Holotype: US 00089459!).</p> <p>Quercus cinnamomea Trel., Mem. Natl. Acad. Sci. 20: 184, pl. 371 (1924). Type:— GUATEMALA, Totonicapan: O. F. Cook 27 (Holotype: US00089485!). The protologue says Cook 27 which Breedlove recorded as seeing at US, but the US database says Cook 127 and indicates it is an isotype. Unfortunately, there is no image available for the US specimen to compare to the image of the holotype published by Trelease (1924, pl. 371).</p> <p>Distribution: Guatemala, Mexico (Chiapas).</p> <p>Quercus cualensis L.M.González, Brittonia 55(1): 49, figs. 1–3, 5 (2003). Type:— MEXICO, Jalisco: L. M. González V. et al. 3076 (Holotype: IBUG0070352 (image)!; isotypes: CAS!, F (image)!, IEB, MEXU, MICH (image)!, NMC, NY (image)!, TEX (image)!, UC, US (image)!, WIS (image)!, ZEA). This narrowly endemic species is similar to the more widespread species Q. iltisii, but there are a number of distinct character states presented in the protologue which justify its recognition.</p> <p>Distribution: Mexico (Jalisco).</p> <p>Quercus delgadoana S.Valencia, Nixon &amp; L.M.Kelly, Novon 21(2): 275, fig. 1 (2011). Type:— MEXICO, Hidalgo: S. Valencia A. et al. 3748 (Holotype: FCME; isotypes: BH, HUAP, IBUG (image)!, K, MEXU (image)!, MO, NY). The illustration in Valencia et al. (2011, Fig. 1) and the isotype image seen from MEXU appear to be similar to Q. perseifolia, a species not mentioned in the protologue. This possible synonymy needs additional study.</p> <p>Distribution: Mexico (Chiapas, Hidalgo, Oaxaca, Puebla, Querétaro, San Luis Potosí, Tabasco, Veracruz).</p> <p>Quercus depressa Bonpl., Pl.Aequinoct. 2: 50, pl. 92 (1809). Type:— MEXICO, Hidalgo: A. Bonpland 4145 (Holotype: P00129727!; isotypes: B-W (as s.n.)!, CGE (as s.n.)!, HAL!, P!, UCBG (fragment)!).</p> <p>Quercus carnerosana Trel., Mem. Natl. Acad. Sci. 20: 183, pl. 369b (1924). Type:— MEXICO, Coahuila: C. G. Pringle 2802 (Holotype: A00055603!; isotypes: GH!, MEXU!).</p> <p>Quercus charcasana Trel. ex A.Camus, Chênes Texte 3: 216, 1269 (1953). Type:— MEXICO, San Luis Potosí: C. L. Lundell 5382 (Lectotype (designated here): ILL00007535!; isolectotypes: A!, DS!, K!, S (image)!, US!). Camus did not specify where the type collection was deposited. The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. depressa as broadly interpreted.</p> <p>Quercus subavenia Trel., Mem. Natl. Acad. Sci. 20: 169, pl. 338b (1924). Type:— MEXICO, Puebla: C. G. Pringle 8908 (Holotype: G!; isotypes: A!, BM!, C!, CAS!, E!, F!, FI!, GH!, HBG (image)!, ILL!, K!, L!, LL (image)!, M!, MO!, NY!, P!, PH!, UC!, UCSB (fragment)!, US!, VT!).</p> <p>Distribution: Mexico (Ciudad de México, Coahuila, Hidalgo, México, Nuevo León, Oaxaca, Puebla, San Luis Potosí,</p> <p>Tamaulipas, Veracruz, Zacatecas). The occurrence of Q. depressa in Ciudad de México and México reported by</p> <p>Villaseñor (2016) needs to be confirmed.</p> <p>Quercus depressipes Trel., Mem. Natl. Acad. Sci. 20: 90, pl. 144a (1924). Type:— MEXICO. Durango: P. Tenorio L. &amp; C. Romero de T. 1196 Buenos Aires, 36 km al W de Tepehuanes, por la Brecha a topia, Mpio. Tepehuanes, 22 Jul 1982 (Holotype at B destroyed. Neotype (designated here): MEXU00345812!; isoneotype: CAS!)..</p> <p>Quercus bocoynensis C.H.Mull., Amer. Midl. Naturalist 19(3): 584 (1938). Type:— MEXICO, Chihuahua: F. Shreve 8022 (Holotype: BH; isotypes: UCSB (fragment)!, US!). An isotype in the protologue is also cited as being at the Desert Laboratory, Tucson, Arizona.</p> <p>Quercus oblongifolia Torr. var. pallidinervis Trel., Mem. Natl. Acad. Sci. 20: 91, pl. 146b (1924). Type:— MEXICO, Chihuahua: E. W. Nelson 6309 (Holotype: US 00169742!; isotypes: GH!, K!).</p> <p>Distribution: Mexico (Chihuahua, Durango, Guanajuato, Hidalgo, Jalisco, Nuevo León, Querétaro, San Luis Potosí,</p> <p>Sinaloa, Sonora, Tamaulipas, Zacatecas), U.S.A. (Texas).</p> <p>Quercus devia Goldman, Contr. U.S. Natl. Herb. 16: 322 (1916). Type:— MEXICO, Baja California Sur: E. W. Nelson &amp; E. A. Goldman 7454 (Holotype: US 00089501!; isotype: A!).</p> <p>Quercus viminea Trel., Mem. Natl. Acad. Sci. 20: 123, pl. 222a (1924). Type:— MEXICO, Sonora: R. Hartman 342 (Holotype at B destroyed. Lectotype (designated here): US00898088!; isolectotypes: CAS!, GH!, K!, NY!). The holotype was Endlich 1280 from Chihuahua, Mexico, and no isotypes have been found. The lectotype is selected from one of the paratypes from Sonora, Mexico. Although Villaseñor (2016) considered this species to be distinct, we consider it to fall within Q. devia as broadly interpreted.</p> <p>Distribution: Mexico (Aguascalientes, Baja California, Baja California Sur, Chihuahua, Durango, Guanajuato, Jalisco,</p> <p>Nayarit, San Luis Potosí, Sinaloa, Sonora, Zacatecas), U.S.A. (Arizona). The occurrence of Q. devia in Aguascalientes reported by Villaseñor (2016) needs to be confirmed.</p> <p>Quercus diversifolia Neé, Anales Ci. Nat. 3: 270 (1801). Type:— MEXICO, Guerrero: L. Neé s. n. (1791) (Lectotype: MA26468! designated by Bartholomew &amp; Almeda (2022, p. 28); isolectotype: UCSB (fragment from MA lectotype)!).</p> <p>Quercus centralis Trel., Mem.Natl.Acad.Sci. 20: 61, pl. 60a (1924).Type:— MEXICO, Ciudad de México: C. G. Pringle 10394 (Holotype at B destroyed. Lectotype (designated here): CAS0655142!; isolectotypes: GH!, LE!, UC!, US!, VT!, WIS!). The holotype was Endlich 1365c, which was probably collected in Ciudad de México, but no isotypes are extant. We are lectotypifying the name based on a paratype also from Ciudad de México.</p> <p>Quercus sanchez-colinii Martínez, Anales Inst. Biol. Univ. Nac. Autón. México 24(2): 269, fig. 15 (1953[1954]). Type:— MEXICO, México: E. Matuda 28760 (Holotype: MEXU 01231768!; isotypes: CAS!, MEXU!, UCSB (fragment)!).</p> <p>Quercus texcocana Trel., Mem. Natl. Acad. Sci. 20: 81, pl. 117a (1924). Type:— MEXICO, México: R. Endlich 653 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 117a)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype. The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. diversifolia as broadly interpreted.</p> <p>Quercus texcocana var. ampla Trel., Mem. Natl. Acad. Sci. 20: 81, pl. 117b (1924). Type:— MEXICO, México: R. Endlich 655 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 117b)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Quercus tomentosa Willd. nom. illigit. var. diversifolia (Neé) A.DC., Prodr. 16(2.1): 33 (1864).</p> <p>Distribution: Mexico (Ciudad de México, Coahuila, Guanajuato, Guerrero, Hidalgo, México, Michoacán, Morelos, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Tamaulipas, Tlaxcala, Veracruz). The occurrence of Q. diversifolia in Coahuila, Nuevo León, and Tamaulipas reported by Villaseñor (2016) needs to be confirmed.</p> <p>Quercus dumosa Nutt., N. Amer. Sylv. 1(1): 7 (1842). Type:— U.S.A., California: T. Nuttall s. n. (May 1819) (Holotype: BM000092132!; isotype UCSB (fragment from BM holotype)!).</p> <p>Quercus dumosa f. diversifolia Trel., Mem. Natl.Acad.Sci. 20: 117, pl. 205a (1924). Type:— MEXICO, Baja California: T. Brandegee s. n. (21 Apr 1893) (Holotype: A00055679!).</p> <p>Quercus dumosa f. microcarpa Trel., Mem. Natl. Acad. Sci. 20: 117, pl. 203b (1924). Type:— MEXICO, Baja California: E. Palmer 640 (Holotype: MO).</p> <p>Quercus dumosa f. populifolia Trel., Mem. Natl.Acad. Sci. 20: 117, pl. 204b (1924). Type:— MEXICO, Baja California: T. Brandegee s. n. (6 May 1893) (Holotype: A).</p> <p>Distribution: Mexico (Baja California, Chihuahua, Jalisco, Michoacán), U.S.A. (California).</p> <p>Quercus durifolia Seemen, Bot. Jahrb. Syst. 29(1): 95 (1900). Type:— MEXICO, Durango: E. Palmer 771 (Holotype at B destroyed. Lectotype (designated here): US00089491!; isolectotypes: A!, BM!, CAS!, F!, GH!, K!, MO!, NY!, UC!, UCSB (fragment)!).</p> <p>Quercus flocculenta C.H.Mull., J. Arnold Arbor. 17(3): 172 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 1277 (Holotype: A00055663!; isotypes: TEX (image)!, UCSB (fragment)!). The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. durifolia as broadly interpreted.</p> <p>Quercus flocculenta f. incisa C.H.Mull., J.Arnold Arbor. 17(3): 173 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 1276 (Holotype: A00055664!; isotype: UCSB (fragment)!).</p> <p>Quercus flocculenta f. oblongifolia C.H.Mull., J. Arnold Arbor. 17(3): 173 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 1229 (Holotype: A00055665!; isotypes: BM!, G!, MEXU!, NY!, P!, S (image)!, US!).</p> <p>Quercus sipuraca Trel., Mem. Natl. Acad. Sci. 20: 182, pl. 366b (1924). Type:— MEXICO, Chihuahua: R. Endlich 1286 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 366b)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Distribution: Mexico (Chihuahua, Coahuila, Durango, Nuevo León, Oaxaca, Querétaro, San Luis Potosí, Sinaloa,</p> <p>Sonora, Tamaulipas, Zacatecas).</p> <p>Quercus dysophylla Benth., Pl.Hartw. 55(1840).Type:— MEXICO, Veracruz: T. Hartweg421 (Holotype:K000512275!; isotypes: BM!, CGE!, E!, LD (image)!, OXF (as mislabel #1963)!, UCSB (fragment)!). This species has been considered by some to be of hybrid origin between Q. crassifolia and Q. crassipes (Govaerts &amp; Frodin 1998, Villaseñor 2016, López C. et al. 2017), but we consider the type collection to not be a hybrid.</p> <p>Quercus esperanzae Trel., Mem. Natl. Acad. Sci. 20: 132, pl. 248 (1924). Type:— MEXICO, Puebla: C. A. Purpus 5332 (Holotype: F0060028 F!; isotypes: BM!, GM!, BM!, MEXU!, MO!, UC!, UCSB (fragment)!). Although Trelease (1924, p. 224) in the caption for pl. 248 wrote “The type (Mexico, Purpus 5332). Photographed by permission at N. Y.” the actual specimen in plate 248 is at F.</p> <p>Quercus floccosa Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 178 (1854). Type:— MEXICO, Veracruz: F. M. Liebmann 3479 (Lectotype: C 10012481! Designated by Trelease (1924, p. 223, pl. 232a); isolectotypes: CAS! UCSB (fragment from C lectotype)!). The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. dysophylla as broadly interpreted.</p> <p>Quercus fournieri Trel., Mem. Natl. Acad. Sci. 20: 177, pl. 357 (1924). Type:— MEXICO, Veracruz: L. Hahn s. n. (Jun 1866) (Holotype: P00077457!; isotypes: BM (as 255)!, K (as 255)!, UCSB (fragment from P holotype)!). At P there are three sheets with the name Q. ferruginea, two of which have the number 258 and one which is s.n. This name was apparently being proposed by Fournier as a new species as “n sp.” is written on the s.n. sheet. Trelease (1924, p. 178) realized that this would be a later homonym of Q. ferruginea Michaux (1812) and published the new name, Q. fournieri, (Trelease 1924, p. 178, pl. 357). He cites “ Hahn 255, June, 1866” at P as the type although there is no Quercus specimen at P with the number Hahn 255. In the caption to plate 357 Trelease states “The type (Mexico, Hahn, 255)” which is in fact a close-up of the s.n. specimen at P. On the label for the P specimen of Hahn s. n. the date was originally written as “ Juillet 1866 ” with the month crossed out and in the same hand changed to “Juin.” Both BM and K have specimens distributed from P which are labeled Hahn 255 and dated as being collected in July 1866. With the mixup of numbers and dates but clearly Hahn s. n. at P (1924, pl. 357) being indicated as “the type”, the P specimen of Hahn s. n. at P is the holotype and the specimens of Hahn 255 at BM and K are probably isotypes. There is no mixup in the identity of any of these specimens as they are all Q. dysophylla. The type fragment at UCSB taken by C.H. Muller from a specimen at P is labeled “ Hahn 255?” indicating that Muller was aware of this number mixup by Trelease and also that P did not in fact have a collection of Hahn 255. The UCSB fragment which consists of a single leaf is undoubtedly a fragment from the Hahn s. n. holotype.</p> <p>Quercus hahnii Trel., Mem. Natl. Acad. Sci. 20: 131, pl. 247 (1924). Type:— MEXICO, Ciudad de México: L. Hahn 347 (Holotype: C 10012474!; isotypes: P!, UCSB (fragment from C holotype)!).</p> <p>Quercus imbricariifolia Trel., Mem. Natl. Acad. Sci. 20: 174, pl. 348ª (1924). Type:— MEXICO, Ciudad de México: E. Bourgeau 425 (Lectotype (designated here): P06814111!; isolectotypes: BM!, K!, MPU!, UCSB (fragment)!). The holotype, Endlich 1012, was destroyed at B; no isotypes have been found in any herbarium. We are lectotypifying the name based on the paratype. One of the specimens at P of Bourgeau 425 is Q. crassipes so it cannot be considered an isolectotype, but alternative interpretation would place this species as a synonym of Q. crassipes.</p> <p>Quercus sagata E.F.Warb., Bull. Misc. Inform. Kew 1939 (2): 90 (1939). Type:— MEXICO, México: G. B. Hinton 6577 (Holotype: K000575022!; isotypes: A!, F!, GBH (image)!, UCSB (fragment)!, US!). The status of this taxon needs additional study as the type has some similarities to Q. hintonii.</p> <p>Distribution: Mexico (Chiapas, Ciudad de México, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, México,</p> <p>Michoacán, Morelos, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Tamaulipas, Tlaxcala, Veracruz,</p> <p>Zacatecas).</p> <p>Quercus eduardi Trel., Contr.U.S. Natl. Herb. 23(2): 189 (1922).Type:— MEXICO, Durango: E. Palmer 956 (Holotype at B destroyed. Lectotype (designated here): US 00089572!; isolectotypes: A!, CAS!, DS!, GH!, K!, MO!, NY!, UC!, UCSB!, US!). Nom. nov. for Q. oligodonta Seemen ex Loes. (1900) non Saporta (1879). This species is close to Q. laurina, and additional study is needed to determine if it is really distinct.</p> <p>Quercus eduardi f. cespitifera Trel., Mem. Natl. Acad. Sci. 20: 122, pl. 219b (1924). Type:— MEXICO, Guanajuato: W. Schumann 1311 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 219a)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Quercus oligodonta Seemen non Saporta (1879), Bot. Jahrb. Syst. 29(1): 96 (1900). See Q. eduardi above.</p> <p>Distribution: Mexico (Durango, Guanajuato).</p> <p>Quercus elliptica Neé, Anales Ci. Nat. 3: 278 (1801). Type:— MEXICO, Hidalgo: L. Neé s. n. (1791) (Lectotype: MA25956! designated by Bartholomew &amp; Almeda (2022, p. 28); isolectotypes: MA!, UCSB (fragment from MA lectotype)!, UCSB (fragment)!). Although the type is reported to be from Hidalgo, no subsequent collections have been made from there. It is possible that Neé’s record was not correctly reported.</p> <p>Quercus acutangula Trel., Repert. Spec. Nov. Regni Veg. 33: 318 (1934). Type:— MEXICO, Guerrero: L. Schultze 483 (Holotype: at B destroyed). The tentative determination of Q. acutangula as probably Q. elliptica is based on the Trelease (1934) description as no images, isotype, or other original material exist. Trelease, however, does mention the twigs being tomentose which is a characteristic of Q. elliptica, so the synonymy is uncertain.</p> <p>Quercus atrescentirhachis Trel., Repert. Spec. Nov. Regni Veg. 33: 317 (1934). Type:— MEXICO, Guerrero: L. Schultze 347 (Syntypes at B destroyed. Lectotype (designated here): ILL00007471! (fragment)). ILL also has a fragment from Schulze 384 which Trelease (1934, p. 317) cites as “Also? (no. 384), with rather shallow cupules, 10–15 mm in diameter with apressed somewhat keeled scales and round-ovoid acorns.” Both L. Schultze 447 and 384 are Q. elliptica.</p> <p>Quercus botryocarpa Trel., Mem. Natl. Acad. Sci. 20: 150, pl. 295 (1924). Type:— MEXICO, Nayarit: J. N. Rose 1971 (Holotype: US00089443!; isotypes: GH!, MO!, NY!, UCSB (fragment!)).</p> <p>Quercus castanea Neé var. elliptica (Neé) Trel., Mem. Natl. Acad. Sci. 20: 178, pl. 359a (1924).</p> <p>Quercus chiquihuitillonis Trel., Mem. Natl. Acad. Sci. 20: 139, pl. 264a (1924). Type:— MEXICO, Michoacán: E. Langlassé 212 (Holotype at B destroyed. Lectotype (designated here): P00754084!; isolectotypes: G!, GH!, K!, MPU!, P!, UCSB (fragment)!, US!).</p> <p>Quercus coccinata Trel., Repert. Spec. Nov. Regni Veg. 33: 317 (1934). Type:— MEXICO, Guerrero: L. Schultze 305 (Holotype at B destroyed. Lectotype (designated here): ILL00007526! (fragment)).</p> <p>Quercus comayaguana Trel., J. Arnold Arbor. 11(1): 25 (1930). Type:— HONDURAS, Comayagua: P. C. Standley 56229 (Holotype: F (accn. 581423)!; isotypes: A!, G!, ILL!, US!).</p> <p>Quercus exaristata Trel., Mem. Natl. Acad. Sci. 20: 140, pl. 266a (1924). Type:— MEXICO, Nayarit: J. N. Rose s. n. (3 Aug 1897) (Holotype: US 00089516!).</p> <p>Quercus guayabalana Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 8(1): 6 (1930). Type:— EL SALVADOR, Cuscatlán: S. Calderon 1982 (Holotype: US 00089528!; isotypes: GH!, ILL!, MO!).</p> <p>Quercus hondurensis Trel., Mem. Natl. Acad. Sci. 20: 140, pl. 266b (1924). Type:— HONDURAS, Cortés: C. Thieme 5440 (Holotype: US 00089556!).</p> <p>Quercus ignaciensis C.H.Mull., Amer. Midl. Naturalist 27(2): 475 (1942). Type:— MEXICO, Sonora: H. T. Green s. n. (26 Feb 1938) (Holotype: PH00023359!; isotype UCSB (fragment from PH holotype)!). The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. elliptica as broadly interpreted.</p> <p>Quercus incrassata Trel., Mem. Natl. Acad. Sci. 20: 147, pl. 287 (1924). Type:— MEXICO, Chiapas: E. Reeves 7 (Holotype: ILL00009824!). The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. elliptica as broadly interpreted.</p> <p>Quercus langlassei Trel., Mem. Natl. Acad. Sci. 20: 138, pl. 263 (1924). Type:— MEXICO, Guerrero: E. Langlassé 1006 (Holotype at B destroyed. Lectotype (designated here): K000512915!; isolectotypes: G!, GH!, P!, US!, UCSB (fragment)!).</p> <p>Quercus linguifolia Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 180 (1854). Type:— MEXICO, Oaxaca: F. M. Liebmann 3506 (Lectotype: C10014103! designated by Trelease (1924, p. 225, pl. 290); isolectotypes: CAS!, GH!, K (as s.n.)!, UCSB (from C lectotype)!). The K s.n. sheet is almost certainly a syntype as it is the right taxon from the right locality on the right date.</p> <p>Quercus nectandrifolia Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 175 (1854). Type:— MEXICO, Oaxaca: F. M. Liebmann 3509 (Lectotype: C10014106! designated by Trelease (1924, p. 225 pl. 289a); isolectotypes: CAS!, UCSB (fragment from C lectotype)!). The lectotype is sheet Cupulif. Liebm. 100.</p> <p>Quercus oajacana Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 178 (1854). Type:— MEXICO, Oaxaca: H. Galeotti 108 (Holotype: K000512934!; isotypes: BR!, G!, P!, UCSB (fragment)!).</p> <p>Quercus peradifolia E.F.Warb., Bull. Misc. Inform. Kew 1939 (2): 92 (1939). Type:— MEXICO, México: G. B. Hinton 6062 (Holotype: K000575017!; isotypes: A!, BM!, CAS!, F!, G!, GBH (image)!, K!, NY!, UCSB (fragment)!).</p> <p>Quercus porriginosa Trel., Repert. Spec. Nov. Regni Veg. 33: 316 (1934). Types: MEXICO, Guerrero. L. Schultze 330 and 333 (Syntypes: at B destroyed). No original material exists, so the only information is Trelease’s (1934) description. Quercus porriginosa to be a synonym of Q. elliptica, but Breedlove thought that it might be Q. peduncularis.</p> <p>Quercus pubinervis M.Martens &amp; Galeotti, Bull. Acad. Roy. Sci. Bruxelles 10(1): 211 (1843). Type:— MEXICO, Veracruz: H. Galeotti 134 (Holotype: BR!; isotypes: G!, P!, UCSB (fragment from BR holotype)!).</p> <p>Quercus salicifolia Neé var. oajacana (Liebm.) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 207 (1884).</p> <p>Quercus yoroensis Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 9(4): 282 (1940). Type:— HONDURAS, Lempira: T. G. Yuncker et al. 8181 (Holotype: ILL00007541!; isotypes: BM!, F!, G!, GH!, K!, MO!, NY!, S (image)!, UCSB (fragment)!, US!).</p> <p>Quercus yoroensis var. aguanana Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 9(4): 282 (1940). Type:— HONDURAS, Yoro: T. G. Yuncker et al. 8185 (Holotype: ILL00007540!; isotypes: BM!, F!, G!, GH!, K!, MO!, NY!, S (image)!, UCSB (fragment)!, US!).</p> <p>Distribution: Belize, El Salvador, Guatemala, Honduras, Mexico (Chiapas, Chihuahua, Colima, Durango, Guerrero,</p> <p>Hidalgo, Jalisco, México, Michoacán, Nayarit, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa, Veracruz),</p> <p>Nicaragua.Although the type of Q. elliptica is from Hidalgo, no subsequent collections of this species have been found in this state. The occurrence of Q. elliptica in Querétaro reported by Villaseñor (2016) also needs to be confirmed.</p> <p>Quercus emoryi Torr., Notes Milit. Reconn. 152, pl. 9 (1848). Type:— U.S.A., Texas: W. H. Emory s. n. (17 Oct 1846) (Lectotype: NY00248636! designated by Landrum (1992, p. 240)).</p> <p>Quercus balsequillana Trel., Mem. Natl. Acad. Sci. 20: 123, pl. 220a (1924). Type:— MEXICO, Chihuahua: R. Endlich 1278 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 220a)!). Trelease (1924, p. 123) wrote that the type is Endlich 1275 but on p. 223 in the caption to pl. 220a he says the type is Endlich 1278. One or the other is probably a typographic error, but in any case, the type at B was destroyed.</p> <p>Quercus duraznillo Trel., Mem. Natl. Acad. Sci. 20: 122, pl. 220b (1924). Type:— MEXICO, Chihuahua: C. G. Pringle 2112 (Holotype at B destroyed. Lectotype (designated here): A00970050!). The holotype was Endlich 1277b, and there are no extant isotypes. The lectotype is chosen from the only extant paratype which has spiny leaves like Q. duraznillo f. pinetorum, but both are Q. emoryi.</p> <p>Quercus duraznillo f. bullata Trel., Mem. Natl. Acad. Sci. 20: 123, pl. 221a (1924). Type:— MEXICO, Chihuahua: R. Endlich 1277a (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 221a)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Quercus duraznillo f. cochutensis Trel., Mem. Natl.Acad. Sci. 20: 123 (1924). Type:— MEXICO, Sonora: C. V. Hartman 72 (Holotype at B destroyed. Lectotype (designated here): GH00071010!).</p> <p>Quercus duraznillo f. pinetorum Trel., Mem. Natl. Acad. Sci. 20: 123, pl. 221b (1924). Type:— MEXICO, Sonora: C. V. Hartman 131 (Holotype: GH00055597!).</p> <p>Quercus hastata Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 171 (1854). Type:— U.S.A., Texas: C. Wright 666 (Holotype: K000832269!; isotypes: G!, NY!, OXF!, US!). Liebmann (1854, p. 171) wrote “ Coll. Chr. Wright n. 666. (hb. Hook.!)” although the specimen marked as the holotype at K has a “Herbarium Benthamianum” stamp, it does have a determination in Liebmann’s hand.</p> <p>Distribution: Mexico (Baja California, Chihuahua, Coahuila, Durango, Jalisco, Nuevo León, San Luis Potosí, Sonora,</p> <p>Tamaulipas, Zacatecas), U.S.A. (Arizona, New Mexico, Texas).</p> <p>Quercus engelmannii Greene, Ill. W. Amer. Oaks 1: [33], pl. 15 figs. 2 &amp; 3, pl. 17 (1889). Type:— U.S.A., California: E.L. Greene s. n. (16 Feb 1877) (Lectotype (designated here): NDG01270!). It is uncertain whether there was ever an Albert Kellogg holotype of this species, but if there ever was one at CAS, it was not saved from the 1906 fire. The illustrations in Greene’s publication are by Kellogg. Greene collected this species in 1877 which is the one we are designating as the lectotype. Other collections by Greene at NDG were collected after the publication of the species, so the collection available to Greene at the time of the publication is the one appropriate for lectotypification.</p> <p>Distribution: Mexico (Baja California), U.S.A. (California).</p> <p>Quercus epileuca Trel., Mem. Natl. Acad. Sci. 20: 126, pl. 229a (1924). Type:— MEXICO, Sonora: C. V. Hartman 337 (Holotype: NY00253687!; isotypes: A!, GH!, K!, UCSB (fragment from NY holotype)!, US!).</p> <p>Quercus hypoleuca Engelm. non Miq. (1858), Trans. Acad. Sci. St. Louis 3: 384 (1876). See Q. hypoleucoides below.</p> <p>Quercus hypoleucoides A.Camus, Bull. Mus. Natl. Hist. Nat., sér. 2, 4: 124 (1932). Type:— U.S.A., Arizona: A. Schott s. n. (28 Jul 1855) (Lectotype: NY00248698! designated by Trelease (1924, p. 223, pl. 226a) as the type of Q. hypoleuca Engelm. (1876) non Miq. (1855); isolectotypes: GH!, MO!). Nom. nov. for Q. hypoleuca Engelm (1876) non Miq. (1858). The type is similar to Q. scytophylla Liebm., a species described from Oaxaca. Additional study is necessary to confirm our proposed synonymy.</p> <p>Distribution: Mexico (Chihuahua, Coahuila, Durango, México, Sinaloa, Sonora, Zacatecas), U.S.A. (Arizona, New</p> <p>Mexico, Texas).</p> <p>Quercus flagellifera Trel., Mem. Natl. Acad. Sci. 20: 162, pl. 319 (1924). Type:— GUATEMALA, Alta Verapaz: O. F. Cook &amp; R. F. Griggs 607 (Holotype: US 00089512!). This species is similar to Q. cortesii, and additional study may show that they should be treated as the same species.</p> <p>Distribution: Guatemala.</p> <p>Quercus frutex Trel., Mem. Natl.Acad. Sci. 20: 82, pl. 120a (1924). Type:— MEXICO, Ciudad de México: E. Bourgeau 68 (Holotype: C 10012503!; isotype: UCSB (fragment from C holotype)!).</p> <p>Quercus frutex var. uhdeana Trel., Mem. Natl. Acad. Sci. 20: 83 (1924). Type:— MEXICO: Uhde 295 (Holotype: at B destroyed). Because no images or original material of Q. frutex var. uhdeana exist, the determination of this variety as probably Q. frutex is based on the Trelease (1924, p. 83) description and the fact that he also cited Uhde 295 as possibly a paratype of Q. vallicola (Trelease 1924, p. 81) which is a later homonym and treated here as a synonym of Q. frutex. It is also possible that Q. frutex var. uhdeana may be a synonym of Q. microphylla according to an anonymous reviewer (per. commun.).</p> <p>Quercus intricata Trel. f. angusta Trel., Mem. Natl. Acad. Sci. 20: 85, pl. 127b (1924). Type:— MEXICO, Hidalgo: C. G. Pringle 13286 (Holotype: US00089548!; isotypes: CAS!, F!, GH!, LL (image)!, MEXU!, MICH!, TEX (image)!, UC!, VT!).</p> <p>Quercus vallicola Trel. non Gand. (1890), Mem. Natl. Acad. Sci. 20: 81, pl. 116b (1924). Type:— Mexico, Ciudad de México: Uhde 296 (Holotype: B destroyed). There are apparently no isotypes, but because Q. vallicola Trel. is a later homonym, we are not lectotypifying the image of the holotype in Trelease (1924, pl. 116b). The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. frutex as broadly interpreted.</p> <p>Distribution: Mexico (Chihuahua, Ciudad de México, Guanajuato, Hidalgo, Jalisco, México, Michoacán, Oaxaca,</p> <p>Puebla, Querétaro, San Luis Potosí, Sonora, Tlaxcala, Veracruz, Zacatecas).</p> <p>Quercus fulva Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 183 (1854). Type:— MEXICO, Sinaloa: B. Seemann 1973 (Holotype: K000512901!; isotypes: BM!, C (as s.n.)!, GH!, UCSB (fragment from K holotype)!).</p> <p>Quercus rosei Trel., Mem. Natl. Acad. Sci. 20: 137, pl. 260a (1924). Type:— MEXICO, Nayarit: J. N. Rose 3458 (Holotype: US 00089618!).</p> <p>Distribution: Mexico (Aguascalientes, Chihuahua, Coahuila, Durango, Guanajuato, Jalisco, Michoacán, Nayarit,</p> <p>Nuevo León, Oaxaca, Querétaro, San Luis Potosí, Sinaloa, Sonora, Zacatecas).The occurrence of Q. fulva in Michoacán and Querétaro reported by Villaseñor (2016) needs to be confirmed.</p> </div>	https://treatment.plazi.org/id/794B8792FFD8FFCC3E88FC9CFC5DFD09	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FF83FFA23E88FF67FB6CF7F1.text	794B8792FF83FFA23E88FF67FB6CF7F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus calophylla	<div><p>Q. calophylla × Q. hintonii Mexico (México). G. B. Hinton 6072 F!, G!, K!, NY!, P!, US!</p> <p>Q. confertifolia × Q. crassipes Mexico (Jalisco). R. Cuevas 2819 WIS!</p> <p>Q. cortesii × Q. benthamii Honduras (Comayagua). D. Mejia 247 CAS!; L. O. Williams 18417 F!, GH!, US!</p> <p>Honduras (Francisco Morazán). P. H. Allen 6390 F!; S. F. Glassman 1967 F!, GH!, ILL!, NY!, UC!; A. Molina R. 1025 F!, GH!, 1111 BM!, F!, GH!, MO!, US!, 10110 F!, NY!, US!, 10647 F!, GH!, US!, 12699 F!, NY!, 12703 F!, NY!; L. O. Williams 12636 F!, GH!, 13721 F!, GH!, 15795 BM!, F!, GH!, MO!, US!, 18568 DS!, F!, US! Honduras (Lempira). G. Davidse 34718 CAS!; D. Mejia 191 CAS!</p> <p>Honduras (Santa Bárbara). P. H. Allen 6374 F!; L. O. Williams 10392 F!, GH!, MO!, 17985 F!</p> <p>Q. cortesii × Q. salicifolia Costa Rica (Heredia). L. J. Poveda 97 8 F!</p> <p>Costa Rica (Puntarenas). A. Jimenez M. 1276B CR!, F!, NY!, 1276E CR!, DS!, F!, NY!, US!, 1276G CR!, F!, NY!</p> <p>Q. crassipes × Q. confertifolia Mexico (Puebla). P. S. Martin 2751966 CAS!</p> <p>Q. crassipes × Q. mexicana Mexico (Hidalgo). C. H. Muller 9126 A!, US!; C. G. Pringle 10290 BM!, C!, CAS!, F!, GH!, ILL!, K!, MEXU!, NY!, P!, PH!, UC!, US!, VT!</p> <p>Q. crassifolia × Q. acutifolia Mexico (Chiapas). L. Kaplan 171 F!</p> <p>Q. crassifolia × Q. calophylla Mexico (Ciudad de México). E. Roman 717 MEXU!</p> <p>Q. crassifolia × Q. sideroxyla Mexico (Chihuahua). R. Spellenberg 8415C CAS!, MEXU!</p> <p>Q. crispipilis × Q. skutchii Guatemala (Chimaltenango). A. Molina R. 12463 F!, NY!, US!; A. F. Skutch 61 US!, ILL!; L. O. Williams 14225 F!, GH!, MO!</p> <p>Guatemala (Huehuetenango). P. C. Standley 82672 F!, US!</p> <p>Guatemala (Quiché). V. Grant 647 A!, F!; A. F. Skutch 1653 A!!, F!, ILL!, US!; P. C. Standley 62452 A!, F!, G! Mexico (Chiapas). D. E. Breedlove 34756 DS!, MO!; L. Kaplan 31 F!, 324 F!, G!, 346 F!, G!</p> <p>Q. depressa × Q. perseifolia Mexico (Veracruz). M. Nee 32900 CAS!</p> <p>Q. depressa × Q.? Mexico (Nuevo León). C. H. Muller &amp; M. T. Muller 802 A!</p> <p>Q. depressipes × Q. laeta Mexico (Durango). R. Spellenberg 10278 CAS!, 10284 CAS!, 10392 CAS!</p> <p>Q. devia × Q. epileuca Mexico (Chihuahua). R. Spellenberg 9805 CAS!, MEXU!, 10920 CAS!, MEXU!</p> <p>Q. × duratifolia El Salvador (San Salvador). M. C. Carlson 486 A!, F!, UC!</p> <p>Mexico (Chiapas). D. E. Breedlove 24708 CAS!, F!, GH!, MO!, NY!; M. Heath &amp; A. Long 1112 CAS!; E. Matuda 2073 A!, CAS!, MEXU!, K!, NY!, US!, 2102 A!, MEXU!, NY!, 4232 F!, GH!, MEXU!, MO!</p> <p>Q. dysophylla × Q. crassifolia Mexico (San Luis Potosí). C. C. Parry &amp; E. Palmer 837 GH!, K!, MO!, NY!, P!, US!, WIS!</p> <p>Q. dysophylla × Q.? Mexico (Puebla). C. H. Muller 9351 CAS!, GH!, MEXU!</p> <p>Q. elliptica × Q. cortesii Mexico (Puebla). F. M. Liebmann 3482 C!, CAS!, K!</p> <p>Q. elliptica × Q. sapotifolia El Salvador (Chalatenango). P. H. Allen 7045 F!, GH!, NY!, US!</p> <p>Q. emoryi × Q. durifolia Mexico (Chihuahua). R. Spellenberg 10900 CAS!, MEXU!, 10903 CAS!, MEXU!</p> <p>Q. epileuca × Q. bolanyosensis Mexico (Sinaloa). D. E. Breedlove 16515 CAS!, MO!</p> <p>Q. frutex × Q. subtriloba Mexico (Hidalgo). C. G. Pringle 10308 CAS!, F!, GH!, UC!, US!, VT!, WIS!</p> <p>Q. fusiformis × Q. oleoides Mexico (Tamaulipas). H. H. Bartlett 10001 DS!, GH!, US!</p> <p>Q. × galeottii Costa Rica (San José). A. Jimenez M. 296 F!</p> <p>Mexico (Chiapas). D. E. Breedlove 25025 DS!, MO!; E. W. Lathrop 5298 DS!</p> <p>Mexico (Veracruz). F. M. Liebmann 3480 C!, CAS!, GH!, MO!, NY!, UC!; L. I. Nevling 569 F!</p> <p>Panama (Chiriquí). E. A. Lao 380 MO!</p> <p>Q. glabrescens × Q. toxicodendrifolia Mexico (Hidalgo). F. Miranda 889 MEXU!</p> <p>Q. grahamii × Q. xalapensis Mexico (Jalisco). R. McVaugh 26158 MICH!</p> <p>Mexico (Michoacán). R. McVaugh 24886 MICH!</p> <p>Mexico (Nuevo León). L. A. Kenoyer 168 F!</p> <p>Q. greggii × Q. diversifolia Mexico (Hidalgo). F. Gonzalez M. 9614 MEXU!, 9621 MEXU!</p> <p>Q. greggii × Q.? Mexico (Veracruz). L. I. Nevling 1760 F!, GH!, MEXU!</p> <p>Q. grisea × Q. frutex Mexico (Hidalgo). C. G. Pringle 10309 CAS!, F!, GH!, UC!, US!, VT!, WIS!</p> <p>Q. grisea × Q. laeta Mexico (Chihuahua). S. S. White 2511 GH!, MEXU!</p> <p>Q. grisea × Q. laeta ? Mexico (Durango). G. L. Fisher 44258 GH!, MO!; R. Spellenberg 10333 CAS!</p> <p>Q. grisea × Q. microphylla Mexico (Coahuila). L. R. Stanford et al. 342 GH!, MO!, NY!</p> <p>Mexico (Tamaulipas). L. J. Dorr 2349 CAS!, MEXU!, NY!, WIS!</p> <p>Q. grisea × Q. oblongifolia ? Mexico (Sonora). C. H. Muller 3737A GH!, UC!, 3737B GH!, UC!</p> </div>	https://treatment.plazi.org/id/794B8792FF83FFA23E88FF67FB6CF7F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FF98FFB93E88FF67FC7FF7F1.text	794B8792FF98FFB93E88FF67FC7FF7F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus candicans	<div><p>Quercus candicans auct. non Neé f. michoacana Trel. = Quercus calophylla Schltdl. &amp; Cham.</p> <p>Quercus candolleana Trel. = Quercus acutifolia Neé</p> <p>Quercus carmenensis C.H.Mull. = Quercus undulata Torr.</p> <p>Quercus carnerosana Trel. = Quercus depressa Bonpl.</p> <p>Quercus castanea Neé</p> <p>Quercus castanea var. elliptica (Neé) Trel. = Quercus elliptica Neé</p> <p>Quercus castanea var. glabrata (Liebm. ex Seem.) A.DC. = Quercus confertifolia Bonpl.</p> <p>Quercus castanea var. integra Oerst. = Quercus mexicana Bonpl.</p> <p>Quercus castanea var. mexicana (Bonpl.) A.DC. = Quercus mexicana Bonpl.</p> <p>Quercus castanea var. sublobata A.DC. = Quercus castanea Neé</p> <p>Quercus castanea var. tridens (Bonpl.) A.DC. = Quercus laurina Bonpl.</p> <p>Quercus cedrosensis C.H.Mull.</p> <p>Quercus centenaria L.M. González = Quercus vicentensis Trel.</p> <p>Quercus centralis Trel. = Quercus diversifolia Neé</p> <p>Quercus centralis f. heterophylla Trel. = Quercus obtusata Bonpl.</p> <p>Quercus cerifera Trel. = Quercus crispipilis Trel.</p> <p>Quercus ceripes Trel. = Quercus sebifera Trel.</p> <p>Quercus charcasana Trel. ex A.Camus = Quercus depressa Bonpl.</p> <p>Quercus chartacea Trel. = Quercus magnoliifolia Neé</p> <p>Quercus chesosensis (Sarg.) C.H.Mull. = Quercus gravesii Sudw.</p> <p>Quercus chiapasensis Trel. = Quercus skinneri Benth.</p> <p>Quercus chiapasensis f. cuneifolia Trel. = Quercus skinneri Benth.</p> <p>Quercus chiapasensis f. falcilobata Trel. = Quercus skinneri Benth.</p> <p>Quercus chiapasensis f. flagellata Trel. = Quercus skinneri Benth.</p> <p>Quercus chiapasensis f. longipes Trel. = Quercus skinneri Benth.</p> <p>Quercus chiapasensis f. petiolata Trel. = Quercus skinneri Benth.</p> <p>Quercus chiapasensis f. subcuneata Trel. = Quercus skinneri Benth.</p> <p>Quercus chicamolensis Trel. = Quercus crassifolia Bonpl.</p> <p>Quercus chichavacana Trel. = Quercus skutchii Trel.</p> <p>Quercus chichavacana f. oblanceolata C.H.Mull. = Quercus skutchii Trel.</p> <p>Quercus chichavacana f. sublobata C.H.Mull. = Quercus skutchii Trel.</p> <p>Quercus chichavacana f. undulata C.H.Mull. = Quercus skutchii Trel.</p> <p>Quercus chihuahuensis Trel. = Quercus alveolata Trel.</p> <p>Quercus chihuahuensis f. amplifolia Trel. = Quercus alveolata Trel.</p> <p>Quercus chihuahuensis f. microphylloides Trel. = Quercus alveolata Trel.</p> <p>Quercus chihuahuensis f. tenuis Trel. = Quercus alveolata Trel.</p> <p>Quercus chimaltenangana Trel. = Quercus skutchii Trel.</p> <p>Quercus chimaltenangana f. gemmata C.H.Mull. = Quercus calophylla Schltdl. &amp; Cham.</p> <p>Quercus chinantlensis Liebm. = Quercus glaucescens Bonpl.</p> <p>Quercus chiquihuitillonis Trel. = Quercus elliptica Neé</p> <p>Quercus chrysolepis Liebm.</p> <p>Quercus chrysolepis subsp. palmeri Engelm. = Quercus palmeri Engelm.</p> <p>Quercus chrysolepis subsp. tomentella (Engelm.) A.E.Murray = Quercus tomentella Engelm.</p> <p>Quercus chrysolepis var. tomentella (Engelm.) A.E.Murray = Quercus tomentella Engelm.</p> <p>Quercus chrysophylla Bonpl. = Quercus laurina Bonpl.</p> <p>Quercus chuhuichupensis C.H.Mull. = Quercus toumeyi Sarg.</p> <p>Quercus cinnamomea Trel. = Quercus crispipilis Trel.</p> <p>Quercus circinata Neé = Quercus magnoliifolia Neé</p> <p>Quercus circummontana Trel. = Quercus castanea Neé</p> <p>Quercus citrifolia Liebm. = Quercus salicifolia Neé</p> <p>Quercus clivicola Trel. &amp; C.H.Mull. = Quercus laeta Liebm.</p> <p>Quercus clivicola f. consanguinea C.H.Mull. = Quercus laeta Liebm.</p> <p>Quercus clivicola f. crenifolia Trel. &amp; C.H.Mull. = Quercus laeta Liebm.</p> <p>Quercus clivicola f. dentata Trel. &amp; C.H.Mull. = Quercus laeta Liebm.</p></div> 	https://treatment.plazi.org/id/794B8792FF98FFB93E88FF67FC7FF7F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FF99FFB83E88FF67FD3EF7F1.text	794B8792FF99FFB83E88FF67FD3EF7F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus clivicola fm. elongata C. H. Mull.	<div><p>Quercus clivicola f. elongata C.H.Mull. = Quercus laeta Liebm.</p> <p>Quercus coahuilensis Nixon &amp; C.H.Mull.</p> <p>Quercus coccinata Trel. = Quercus elliptica Neé</p> <p>Quercus coccinea Trel. var. microcarpa Torr. = Quercus gravesii Sudw.</p> <p>Quercus coccolobifolia Trel. = Quercus jonesii Trel.</p> <p>Quercus coffeicolor Trel.</p> <p>Quercus colimae Trel. = Quercus crassipes Bonpl.</p> <p>Quercus colimae var. zauzillo Trel. = Quercus crassipes Bonpl.</p> <p>Quercus colombiana Cuatrec. = Quercus humboldtii Bonpl.</p> <p>Quercus comasaguana Trel. = Quercus peduncularis Neé</p> <p>Quercus comayaguana Trel. = Quercus elliptica Neé</p> <p>Quercus commutata Liebm. (1854) non Unger (1852) = Quercus affinis Scheidw.</p> <p>Quercus confertifolia Bonpl.</p> <p>Quercus conglomerata Trel. (1924) non Pers. (1807) = Quercus rugosa Neé</p> <p>Quercus conjugens Trel. = Quercus glaucoides M.Martens &amp; Galeotti</p> <p>Quercus consociata Trel. = Quercus castanea Neé</p> <p>Quercus conspersa Benth. = Quercus acutifolia Neé</p> <p>Quercus conspersa f. caudata Trel. = Quercus acutifolia Neé</p> <p>Quercus conspersa f. ovatifolia Trel. = Quercus acutifolia Neé</p> <p>Quercus convallata Trel. = Quercus spicata Bonpl.</p> <p>Quercus conzattii Trel.</p> <p>Quercus copeyensis C.H.Mull. = Quercus bumelioides Liebm.</p> <p>Quercus cordata M.Martens &amp; Galeotti = Quercus glaucoides M.Martens &amp; Galeotti</p> <p>Quercus cordifolia Trel. = Quercus microphylla Neé</p> <p>Quercus cornelius-mulleri Nixon &amp; K.P.Steele</p> <p>Quercus correpta Trel. = Quercus acutifolia Neé</p> <p>Quercus corrugata Hook.</p> <p>Quercus corrugata var. granulifera Trel. = Quercus corrugata Hook.</p> <p>Quercus corrugata var. ipalensis Trel. = Quercus corrugata Hook.</p> <p>Quercus corrugata var. microcarpa Wenz. = Quercus corrugata Hook.</p> <p>Quercus cortesii Liebm.</p> <p>Quercus costaricensis Liebm.</p> <p>Quercus costaricensis f. kuntzei Trel. = Quercus bumelioides Liebm.</p> <p>Quercus crassifolia Bonpl.</p> <p>Quercus crassifolia var. errans (Trel.) E.F.Warb. = Quercus crassifolia Bonpl.</p> <p>Quercus crassipes Bonpl.</p> <p>Quercus crassipes var. angustifolia Bonpl. = Quercus crassipes Bonpl.</p> <p>Quercus crassivenosa Trel. = Quercus castanea Neé</p> <p>Quercus crenatifolia Trel. = Quercus obtusata Bonpl.</p> <p>Quercus crispifolia Trel. = Quercus benthamii A.DC.</p> <p>Quercus crispipilis Trel.</p> <p>Quercus crispipilis var. pannosifolia (C.H.Mull.) C.H.Mull. = Quercus skutchii Trel.</p> <p>Quercus cuajimalpana Trel. = Quercus crassipes Bonpl.</p> <p>Quercus cualensis L.M.González</p> <p>Quercus cubana A.Rich = Quercus sagrana Nutt.</p> <p>Quercus cuneifolia Liebm. (1854) non Raf. (1838) = Quercus glaucescens Bonpl.</p> <p>Quercus cupreata Trel. &amp; C.H.Mull. = Quercus gravesii Sudw.</p> <p>Quercus cupreata f. brachystachys C.H.Mull. = Quercus gravesii Sudw.</p> <p>Quercus cupreata f. serrata Trel. &amp; C.H.Mull. = Quercus gravesii Sudw.</p> <p>Quercus cyclobalanoides Trel. = Quercus lancifolia Schltdl. &amp; Cham.</p> <p>Quercus davidsoniae Standl. = Quercus insignis M.Martens &amp; Galeotti</p> <p>Quercus dawsonii Trel. = Meliosma sp. (see Appendix 1, Excluded Names)</p> <p>Quercus decipiens M.Martens &amp; Galeotti = Quercus rugosa Neé</p> <p>Quercus delgadoana S.Valencia, Nixon &amp; L.M.Kelly</p></div> 	https://treatment.plazi.org/id/794B8792FF99FFB83E88FF67FD3EF7F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FF9AFFBB3E88FF67FD2CF7F1.text	794B8792FF9AFFBB3E88FF67FD2CF7F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus deliquescens C. H. Mull.	<div><p>Quercus deliquescens C.H.Mull. = Quercus intricata Trel.</p> <p>Quercus depressa Bonpl.</p> <p>Quercus depressipes Trel.</p> <p>Quercus derrumbaderoensis C.H.Mull. = Quercus i ntricata Trel.</p> <p>Quercus deserticola Trel. = Quercus grisea Liebm.</p> <p>Quercus deserticola Trel. var. incisa Trel. = Quercus alveolata Trel.</p> <p>Quercus devia Goldman</p> <p>Quercus diversicolor Trel. = Quercus rugosa Neé</p> <p>Quercus diversicolor var. mearnsii Trel. = Quercus rugosa Neé</p> <p>Quercus diversicolor var. socorronis Trel. = Quercus rugosa Neé</p> <p>Quercus diversifolia Neé</p> <p>Quercus dolichopus E.F.Warb. = Quercus peduncularis Neé</p> <p>Quercus donnell-smithii Trel. = Quercus sapotifolia Liebm.</p> <p>Quercus douglasii Hook. &amp; Arn. var. gambelii (Nutt.) A.DC. = Quercus gambelii Nutt.</p> <p>Quercus douglasii var. novomexicana A.DC = Quercus gambelii Nutt.</p> <p>Quercus dugesii (Trel.) A. Nelson = Quercus spicata Bonpl.</p> <p>Quercus dumosa Nutt.</p> <p>Q. dumosa var. acutidens (Torr.) Wenz. = Quercus × acutidens Torr.</p> <p>Quercus dumosa f. diversifolia Trel. = Quercus dumosa Nutt.</p> <p>Quercus dumosa f. linearis Trel. = Quercus × acutidens Torr.</p> <p>Quercus dumosa f. microcarpa Trel. = Quercus dumosa Nutt.</p> <p>Quercus dumosa f. populifolia Trel. = Quercus dumosa Nutt.</p> <p>Quercus dumosa subsp. turbinella (Greene) A.E.Murray = Quercus turbinella Greene</p> <p>Quercus dumosa var. turbinella (Greene) Jeps. = Quercus turbinella Greene</p> <p>Quercus dumosa var. turbinella f. crispata Trel. = Quercus turbinella Greene</p> <p>Quercus dunnii Kellogg = Quercus palmeri (Engelm.) Engelm.</p> <p>Quercus durandii Buckley = Quercus sinuata Walter</p> <p>Quercus durangensis Trel. = Quercus rugosa Neé</p> <p>Quercus × duratifolia C.H.Mull.</p> <p>Quercus duraznillo Trel. = Quercus emoryi Torr.</p> <p>Quercus duraznillo f. bullata Trel. = Quercus emoryi Torr.</p> <p>Quercus duraznillo f. cochutensis Trel. = Quercus emoryi Torr.</p> <p>Quercus duraznillo f. pinetorum Trel. = Quercus emoryi Torr.</p> <p>Quercus durifolia Seemen</p> <p>Quercus dysophylla Benth.</p> <p>Quercus eduardi Trel.</p> <p>Quercus eduardi f. cespitifera Trel. = Quercus eduardi Trel.</p> <p>Quercus edwardsiae C.H.Mull. = Quercus tuberculata Liebm.</p> <p>Quercus elliptica Neé</p> <p>Quercus elliptica var. microcarpa (Liebm.) A.DC. = Quercus sapotifolia Liebm.</p> <p>Quercus emoryi Torr.</p> <p>Quercus endemica C.H.Mull. = Quercus undulata Torr.</p> <p>Quercus endlichiana Trel. = Quercus jonesii Trel.</p> <p>Quercus endlichiana f. minor C.H.Mull. = Quercus jonesii Trel.</p> <p>Quercus endlichiana f. serrata C.H.Mull. = Quercus jonesii Trel.</p> <p>Quercus endresii Trel. = Quercus costaricensis Liebm.</p> <p>Quercus engelmannii Greene</p> <p>Quercus epileuca Trel.</p> <p>Quercus errans Trel. = Quercus crassifolia Bonpl.</p> <p>Quercus errans f. graciliramis C.H.Mull. = Quercus hypoxantha Trel.</p> <p>Quercus erubescens Trel. = Quercus magnoliifolia Neé</p> <p>Quercus esesmilensis Tucker &amp; C.H.Mull. = Quercus benthamii A.DC.</p> <p>Quercus esperanza e Trel. = Quercus dysophylla Benth.</p> <p>Quercus eugeniifolia Liebm. = Quercus salicifolia Neé</p></div> 	https://treatment.plazi.org/id/794B8792FF9AFFBB3E88FF67FD2CF7F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FFF4FFD63E88FF67FD5FFA31.text	794B8792FFF4FFD63E88FF67FD5FFA31.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus elliptica var. microcarpa (Liebm.) A. DC., Prodr.	<div><p>Quercus elliptica Neé var. microcarpa (Liebm.) A.DC., Prodr. 16(2.1): 71 (1864).</p> <p>Quercus microcarpa Liebm. non Lapeyr. (1813), Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 184 (1854). See Q. parviglans below.</p> <p>Quercus parviglans Trel., Proc.Amer.Philos. Soc. 54: 8 (1915). Type:— GUATEMALA: J. R. R. Warszewicz 8 (Lectotype (designated here): G!; isolectotype: UCSB (fragment from G lectotype)!). Nom. nov. for Q. microcarpa Liebm. (1854) non Lapeyr. (1813). Trelease (1924, p. 225, pl. 298a, pl. 298b) indicated that types are at B and G.</p> <p>Quercus parviglans f. polycarpa Trel., Mem. Natl. Acad. Sci. 20: 152, pl. 299a (1924). Type:— GUATEMALA, Baja Verapaz: O. F. Cook 291 (Holotype: US 00089597!).</p> <p>Quercus parviglans f. tejadana Trel., Mem. Natl. Acad. Sci. 20: 152 (1924). Type:— GUATEMALA, Sacatepequez: R.Tejada 209 (Holotype: US 00089596!).</p> <p>Quercus perseifolia Liebm. var. achoteana Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 17(4): 357 (1938). Type:— HONDURAS, Comayagua: T. G. Yuncker et al. 6203 (Holotype: ILL00007608!; isotypes: F!, G!, GH!, K!, MO!, NY!, UCSB (fragment)!, US!).</p> <p>Quercus siguatepequeana Trel., J. Arnold Arbor. 11(1): 25 (1930). Type:— HONDURAS, Comayagua: P. C. Standley 56393 (Holotype: F (accn. 581663)!; isotypes: A!, G!, ILL!, UCSB (fragment)!, US!).</p> <p>Quercus totutlensis A.DC., Prodr. 16(2.1): 62 (1864). Type:— MEXICO, Veracruz: F. M. Liebmann 3568 (Lectotype: CAS0004231! designated by Trelease (1924, p. 225, pl. 297b); isolectotypes: C!, GH!, K (as s.n.)!, P (as s.n.)!, UCSB (fragment)!).</p> <p>Distribution: Belize, Costa Rica, El Salvador, Guatemala, Honduras, Mexico (Chiapas, Guerrero, Hidalgo, Oaxaca,</p> <p>Puebla, San Luis Potosí, Tabasco, Veracruz), Nicaragua.</p> <p>Quercus sarahmariae Nixon &amp; Barrie, Novon 25(4): 446, fig. 2 (2017). Type:— COSTA RICA, Puntarenas: G. Davidse 24503 (Holotype: MO-138404!; isotypes: CAS!, CR). This species needs to be compared with Q. benthamii of which it may be a synonym.</p> <p>Distribution: Costa Rica.</p> <p>Quercus sartorii Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 177 (1854). Type:— MEXICO, Veracruz: F. M. Liebmann 3554 (Lectotype: C10013683! designated by Trelease (1924, p. 228, pl. 381a); isolectotypes: C!, CAS!, F (as s.n.)!, GH!, K (as s.n.)!, MEL? (as s.n., s.d.) (image)!, MO!, NY!, UCSB (fragment from C lectotype)!, UCSB (fragment)!, US (as s.d.)!). The lectotype is sheet Cupulif. Liebm. 14 of Liebmann 3554.</p> <p>Quercus ghiesbreghtii M.Martens &amp; Galeotti, Bull. Acad. Roy. Sci. Bruxelles 10(1): 212 (1843). Type:— MEXICO, Veracruz: H. Galeotti 128 (Lectotype: BR0000006913557! designated by Trelease (1924, p. 226, pl. 322b); isolectotypes: BR!, G!, L!, LE!, P!, UCSB (fragment)!). M. Martens &amp; Galeotti cite only one collection number, two sheets of which are at BR marked as types; one was designated by Trelease as “The type.” The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. sartorii as broadly interpreted.</p> <p>Quercus sartorii f. magma Trel., Mem. Natl. Acad. Sci. 20: 189 (1924). Type:— MEXICO, Veracruz: F. M. Liebmann 3552 (Holotype: C 10012525!; isotypes: BM!, C!, CAS!, MO!, NY!, UCSB (fragment from C holotype)!, UCSB (fragment)!, US!). Although Trelease (1924, p. 189) did not specify where the holotype was deposited, by citing “ Liebmann, 12, 3552 ” he actually designated a specific sheet. The 12 refers to a Cubulif. Lbm. number which is different for each sheet, and Liebm. Cupulif. 12 is a C.</p> <p>Quercus vexans Trel., Mem. Natl. Acad. Sci. 20: 190, pl. 386 (1924). Type:— MEXICO, Veracruz: C. A. Purpus s. n. (Dec 1914) (Lectotype (designated here): ILL00007550!; isolectotypes: ILL!, UC (as 7468)). The UC specimen numbered 7468 is undoubtedly a type collection as it has the correct date, locality, and identity, although none of the seven sheets at ILL have a collection number. The label on the UC specimen is in Purpus’ hand including the phrase “part of Type.” Purpus apparently sent most of the specimens from this set to Trelease for determination who kept them. Evidently, the only specimen that has the collection number on the label is the sheet at UC.</p> <p>Distribution: Mexico (Chiapas, Coahuila, Guerrero, Hidalgo, México, Nuevo León, Oaxaca, Puebla, Querétaro, San</p> <p>Luis Potosí, Tamaulipas, Veracruz). The occurrence of Q. sartorii in Guerrero and México reported by Villaseñor</p> <p>(2016) needs to be confirmed.</p> <p>Quercus scytophylla Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 180 (1854). Type:— MEXICO, Oaxaca: F. M. Liebmann 3557 (Lectotype (designated here): C 10012528!; isolectotypes: CAS!, G!, UCSB (fragment from C lectotype)!). Trelease (1924) in pl. 227 shows two different specimens. In the caption to pl. 227 on p. 223 he states “The type (Mexico, Liebmann, 144-5 = 3557).” The lower image in pl. 227 is sheet 144 which we are designating as the lectotype, and the upper image in pl. 227 is sheet 145, an isolectotype.</p> <p>Quercus campanariensis Trel., Mem. Natl.Acad. Sci. 20: 126, pl. 228 (1924). Type:— MEXICO, Michoacán: G. Arsène 6004 (Holotype: US 00089469!). The specimens of Arsène 6004 at BR, ILL, MO, and US collected on 13 Jul 1911 are Q. laeta so they are not isotypes of Q. campanariensis.</p> <p>Distribution: Mexico (Chiapas, Chihuahua, Colima, Durango, Guerrero, Hidalgo, Jalisco, México, Michoacán,</p> <p>Morelos, Nayarit, Oaxaca, Puebla, Sinaloa, Sonora, Veracruz, Zacatecas).</p> <p>Quercus sebifera Trel., Mem. Natl. Acad. Sci. 20: 93, pl. 149a (1924). Type:— MEXICO, Chiapas: O. F. Cook 79 (Holotype: US 00089638!; isotypes: MO!, UCSB (fragment)!, US!, VT!).</p> <p>Quercus ceripes Trel., Mem. Natl. Acad. Sci. 20: 93, pl. 150b (1924). Type:— MEXICO, Puebla: H. Schenk 239 (Holotype at B destroyed. Lectotype (designated here): ILL00007460! (fragment)).</p> <p>Quercus schenckiana Trel., Mem. Natl. Acad. Sci. 20: 93, pl. 150a (1924). Type:— MEXICO, Puebla: H. Schenk 238 (Holotype at B destroyed. Lectotype (designated here): ILL00007590!).</p> <p>Quercus sebifera f. comitanensis Trel., Mem. Natl. Acad. Sci. 20: 93, pl. 149b (1924). Type:— MEXICO, Chiapas: O. F. Cook 78 (Holotype: US 00089639!).</p> <p>Quercus trinidadensis C.H.Mull., Amer. Midl. Naturalist 27(2): 481 (1942). Type:— MEXICO, Nuevo León: C. H. Muller 2803 (Holotype: US 00089671!; isotypes: F!, GH!, LL (image)!, MICH!, MO!, NY!, UC!, UCSB (fragment)!, US!).</p> <p>Distribution: Mexico (Chiapas, Chihuahua, Coahuila, Durango, Hidalgo, Nuevo León, Oaxaca, Puebla, Querétaro,</p> <p>San Luis Potosí, Sonora, Tamaulipas, Veracruz).</p> <p>Quercus segoviensis Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 186 (1854). Type:— NICARAGUA, Segovia: A. S. Oersted 3559 (Lectotype: C 10012528! designated by Trelease (1924, 217, pl. 54a); isolectotypes: CAS!, UCSB (fragment)!).</p> <p>Quercus achoteana Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 17(4): 356 (1938). Type:— HONDURAS, Comayagua: T. G. Yuncker et al. 6136 (Holotype: ILL00007478!; isotypes: F!, G!, GH!, K!, MICH!, MO!, NY!, UCSB (fragment)!, US!).</p> <p>Quercus achoteana var. sublanosa Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 17(4): 356 (1938). Type:— HONDURAS, Comayagua: T. G. Yuncker et al. 5961 (Holotype: ILL00007477!; isotypes: F!, G!, GH!, K!, MICH!, MO!, NY!, UCSB (fragment)!, US!, WIS!).</p> <p>Quercus arachnoidea Trel., Mem. Natl. Acad. Sci. 20: 59, pl. 55 (1924). Type:— EL SALVADOR: G. Niederlein s. n. (9 Jan 1898) (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 55)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Quercus matagalpana Trel., Mem. Natl. Acad. Sci. 20: 58, pl. 54b (1924). Type:— NICARAGUA, Matagalpa: E. Rothschuh 611 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 54b)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Quercus peduncularis Neé var. sublanosa (Trel.) C.H.Mull., U.S.D.A. Bur. Pl. Industr. Misc. Publ. 477: 34. (1942).</p> <p>Quercus reticulata Bonpl. var. segoviensis (Liebm.) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 195 (1884).</p> <p>Distribution: Belize, El Salvador, Guatemala, Honduras, Mexico (Chiapas, Oaxaca), Nicaragua. The occurrence of Q.</p> <p>segoviensis in Michoacán reported by Villaseñor (2016) is probably incorrect.</p> <p>Quercus sideroxyla Bonpl., Pl. Aequinoct. 2: 39, pl. 85 (1809). Type:— MEXICO, Guanajuato: A. Bonpland 4219 (p.p.) (Holotype: P00129702!; isotypes: P!, UCSB (fragment)!). Three sheets at P of Bonpland 4219 from Villapando are Q. sideroxyla, the holotype and two isotypes. Collections with the same number but from Santa Rosa are Q. tridens and are not types.</p> <p>Quercus incarnata Trel., Mem. Natl. Acad. Sci. 20: 126, pl. 230a (1924). Type:— MEXICO, Chihuahua: R. Endlich 782b (p.p.) (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 230a)!). No isotypes exist of this species or any of the forms described by Trelease, and the other Endlich paratypes of the species at B were also destroyed. The two Arsène paratypes of Q. incarnata are other species (Arsène 5807 = Q. scytophylla and Arsène 6036 = Q. castanea) so they cannot be used as possible lectotypes.</p> <p>Quercus incarnata f. ampla Trel., Mem. Natl. Acad. Sci. 20: 127, pl. 229b (1924). Type:— MEXICO, Chihuahua: R. Endlich 782b (p.p.) (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 229b)!). One must assume that Trelease’s (1924) pl. 229b is of a branch from a different plant than the branch shown as pl. 230a which is of the holotype of the species.</p> <p>Quercus incarnata f. grosse-serrata Trel., Mem.Natl.Acad.Sci. 20:127,pl. 231a (1924). Type:— MEXICO, Chihuahua: R. Endlich 782d (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 231a)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Quercus incarnata f. longa Trel., Mem. Natl. Acad. Sci. 20: 127, pl. 230b (1924). Type:— MEXICO, Chihuahua: R. Endlich 782a (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 230b)!). Trelease (1924, p. 127) cites Endlich 728a as the type, but on p. 223 in the caption to pl. 230b he cites Endlich 782a as the type. It is likely that the first citation is a typographic error, but in any event the collection at B was destroyed. Because no extant original material exists, we are lectotypifying the name based on the image of the holotype.</p> <p>Quercus omissa A.DC., Prodr. 16(2.1): 28 (1864). Type:— MEXICO, Sinaloa: B. Seemann 1969 (Lectotype: K000512899! designated by Trelease (1924, p. 223, pl. 231b); isolectotypes: BM!, C (as s.n.), G!, GH!, UCSB (fragment from K lectotype)!, UCSB (fragment as s.n. from C)!). A. de Candolle (1864, p. 28) cites the type as being at both C and K.</p> <p>Distribution: Mexico (Aguascalientes, Chihuahua, Coahuila, Durango, Guanajuato, Hidalgo, Jalisco, México,</p> <p>Michoacán, Morelos, Nayarit, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa, Sonora, Tamaulipas,</p> <p>Veracruz, Zacatecas). The occurrence of Q. sideroxyla in Morelos and Oaxaca reported by Villaseñor (2016) needs to be confirmed.</p> <p>Quercus sinuata Walter, Fl. Carol. 235 (1788). Type:— U.S.A., Illinois: D. B. Ward 10586 (Neotype: FLAS218678 (image)! designated by Ward (2007, p. 180); isoneotypes: FLAS (image)!, GA, GH (image)!, NY (image)!, USCH (image)!).</p> <p>Quercus breviloba (Torr.) Sarg., Gard. &amp; Forest 8: 93 (1895).</p> <p>Quercus durandii Buckley, Proc. Acad. Nat. Sci. Philadelphia 12: 445 (1860[1861]). Type:— U.S.A., Alabama: C. H. Muller 36898 (Neotype: NA, designated by Dorr &amp; Nixon (1985, p. 218); isoneotype: BH).</p> <p>Quercus obtusifolia D. Don var. breviloba Torr., Rep. U.S. Mex. Bound., Bot. 2(1): 206 (1859). Type:— U.S.A., Texas: C. C. Parry et al. 1377 (Lectotype (designated here): NY00253713!; isolectotypes: GH!, NY!). There are two syntypes at NY. The sheet with handwritten label information that is consistent with the protologue is chosen as the lectotype.</p> <p>Quercus sinuata subsp. breviloba (Torr.) A.E.Murray, Kalmia 13: 27 (1983).</p> <p>Quercus sinuata var. breviloba (Torr.) C.H.Mull., J. Arnold Arbor. 25(4): 439 (1944).</p> <p>Distribution: Mexico (Coahuila, Nuevo León, Tamaulipas, Veracruz), U.S.A. (Alabama, Arkansas, Florida, Georgia,</p> <p>Louisiana, Mississippi, North Carolina, Oklahoma, South Carolina, Texas).</p> <p>Quercus skinneri Benth., Gard. Chron. 1841: 116 (1841). Type:— GUATEMALA, Quetzaltenango: T. Hartweg 615 (Holotype: K000512974!; isotypes: BM!, CGE!, E!, G!, GH (as s.n.)!, LD (image)!, MO!, NY!, OXF!, P!, UCSB (fragment)!, US (frag.)).</p> <p>Quercus chiapasensis Trel., Proc. Amer. Philos. Soc. 54: 9, pl. 2 (1915). Type:— MEXICO, Chiapas: C. A. Purpus 6999 (Lectotype: ILL00007531! and ILL00010196! designated by Trelease (1924, p. 228, pl. 376); isolectotypes: BM!, F!, GH!, ILL!, MO!, NY!, UC!, UCSB (fragment)!, US!, VT!). The lectotype is mounted on two sheets, one with just leaves and the other with just acorns and cups. The sheet with leaves has the label from the collector.</p> <p>Quercus chiapasensis f. cuneifolia Trel., Mem. Natl. Acad. Sci. 20: 186 (1924). Type:— MEXICO, Chiapas: E. Reeves 3 (Holotype: ILL00007530! and ILL00010195!). The holotype is mounted on two sheets, one with just leaves and the other with just acorns and cups. The sheet with acorns and cups has the label from the collector.</p> <p>Quercus chiapasensis f. falcilobata Trel., Mem. Natl. Acad. Sci. 20: 186, pl. 375 (1924). Type:— MEXICO, Chiapas: E. Reeves 5 (Holotype: ILL00007558! and ILL00010199!). The holotype is mounted on two sheets, one with just leaves and the other with just acorns and cups. The sheet with acorns and cups has the label from the collector.</p> <p>Quercus chiapasensis f. flagellata Trel., Mem. Natl. Acad. Sci. 20: 186 (1924). Type:— MEXICO, Chiapas: E. Reeves 6 (Holotype: ILL00007557! and ILL00010198!). The holotype is mounted on two sheets, one with just leaves and the other with just acorns and cups. The sheet with acorns and cups has the label from the collector.</p> <p>Quercus chiapasensis f. longipes Trel., Mem. Natl. Acad. Sci. 20: 186 (1924). Type:— MEXICO, Chiapas: E. Reeves 1 (Holotype: ILL00007563! and ILL00010203!). The holotype is mounted on two sheets, one with just leaves and the other with just acorns and cups. The sheet with acorns and cups has the label from the collector.</p> <p>Quercus chiapasensis f. petiolata Trel., Mem. Natl. Acad. Sci. 20: 186 (1924). Type:— MEXICO, Chiapas: E. Reeves 8 (Holotype: ILL00007562! and ILL00010202!). The holotype is mounted on two sheets, one with just leaves and the other with just acorns and cups. The sheet with acorns and cups has the label from the collector.</p> <p>Quercus chiapasensis f. subcuneata Trel., Mem. Natl.Acad. Sci. 20: 186 (1924). Type:— MEXICO, Chiapas: E. Reeves 9 (Holotype: ILL00007561! and ILL00010201!). The holotype is mounted on two sheets, one with just leaves and the other with just acorns and cups. The sheet with acorns and cups has the label from the collector.</p> <p>Quercus hemipteroides C.H.Mull., Amer. Midl. Naturalist 18(5): 853 (1937). Type:— GUATEMALA, Quiché: A. F. Skutch 1875 (Holotype: ILL00007456!; isotypes: A!, F!, NY!, UCSB (fragment)!, US).</p> <p>Quercus trichodonta Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 17(4): 358 (1938). Type:— HONDURAS, Comayagua: T. G. Yuncker et al. 6116 (Holotype: ILL00007555!; isotypes: F!, G!, GH!, K!, MO!, NY!, UCSB (fragment)!, WIS!). The placement of this species needs additional study. It might be synonymous with Q. xalapensis.</p> <p>Distribution: El Salvador, Guatemala, Honduras, Mexico (Chiapas, Hidalgo, Oaxaca, Tamaulipas, Veracruz). The occurrence of Q. skinneri in Hidalgo reported by Villaseñor (2016) needs to be confirmed.</p> <p>Quercus skutchii Trel., J. Wash. Acad. Sci. 23(1): 61 (1933). Type:— GUATEMALA, Chimaltenango. A. F. Skutch 44 (Holotype: US00089648!; isotype: ILL!). The synonomy of Q. skutchii presented here needs additional study and may be shown to represent more than one species.</p> <p>Quercus aristigera Trel., J. Wash. Acad. Sci. 23(1): 63 (1933). Type:— GUATEMALA, Chimaltenango: A. F. Skutch 86 (Holotype: US 00089429!; isotype: ILL!).</p> <p>Quercus brachystachys Benth.f. caerulea Trel., Mem.Natl.Acad.Sci. 20: 130,pl.240b (1924). Type:— GUATEMALA, Quezaltenango: W. Trelease 30 (Holotype: ILL00007441!; isotype: ILL!).</p> <p>Quercus brachystachys f. venulosa Trel., Mem. Natl. Acad. Sci. 20: 130, pl. 240a (1924). Type:— GUATEMALA, Quezaltenango: W. Trelease 28 (Holotype: ILL00007446!; isotype: ILL!).</p> <p>Quercus chichavacana Trel., J. Wash.Acad. Sci. 23(1): 62 (1933). Type:— GUATEMALA, Chimaltenango: A. F. Skutch 57 (Holotype: US 00089457!; isotype: ILL!).</p> <p>Quercus chichavacana f. oblanceolata C.H.Mull., Amer. Midl. Naturalist 18(5): 852 (1937). Type:— GUATEMALA, Chimaltenango: A.F. Skutch 681 (Holotype: US 00089456!; isotype: A!).</p> <p>Quercus chichavacana f. sublobata C.H.Mull., Amer. Midl. Naturalist 18(5): 852 (1937). Type:— GUATEMALA, Chimaltenango. A. F. Skutch 712 (Holotype: ILL00007440!; isotypes: A!, DS!, F!, US!).</p> <p>Quercus chichavacana f. undulata C.H.Mull., Amer. Midl. Naturalist 18(5): 852 (1937). Type:— GUATEMALA, Chimaltenango: A. F. Skutch 689 (Holotype: US 00089454!; isotypes: A!, F!, K!).</p> <p>Quercus chimaltenangana Trel., J. Wash. Acad. Sci. 23(1): 62 (1933). Type:— GUATEMALA, Chimaltenango: A. F. Skutch 62 (Holotype: US 00089451!; isotype: ILL!).</p> <p>Quercus crispipilis Trel. var. pannosifolia (C.H.Mull.) C.H.Mull., U.S.D.A. Bur. Pl. Industr. Misc. Publ. 477: 79 (1942).</p> <p>Quercus pannosifolia C.H.Mull., Amer. Midl. Naturalist 18(5): 855 (1937). Type:— GUATEMALA, Chimaltenango: A. F. Skutch 698 (Holotype: ILL00007448!; isotypes: A, F, UC, US!).</p> <p>Distribution: Guatemala, Mexico (Chiapas).</p></div> 	https://treatment.plazi.org/id/794B8792FFF4FFD63E88FF67FD5FFA31	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FF9BFFBA3E88FF67FD11F7F1.text	794B8792FF9BFFBA3E88FF67FD11F7F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus eugeniifolia fm. petiolata Trel.	<div><p>Quercus eugeniifolia f. petiolata Trel. = Quercus salicifolia Neé</p> <p>Quercus exaristata Trel. = Quercus elliptica Neé</p> <p>Quercus excelsa Liebm. = Quercus lancifolia Schltdl. &amp; Cham.</p> <p>Quercus felipensis Trel. = Quercus crassifolia Bonpl.</p> <p>Quercus fendleri Liebm. = Quercus undulata Torr.</p> <p>Quercus filiformis C.H.Mull. = Quercus pringlei Seemen</p> <p>Quercus flagellifera Trel.</p> <p>Quercus flavida Liebm. = Quercus calophylla Schltdl. &amp; Cham.</p> <p>Quercus floccosa Liebm. = Quercus dysophylla Benth.</p> <p>Quercus flocculenta C.H.Mull. = Quercus durifolia Seemen</p> <p>Quercus flocculenta f. incisa C.H.Mull. = Quercus durifolia Seemen</p> <p>Quercus flocculenta f. oblongifolia C.H.Mull. = Quercus durifolia Seemen</p> <p>Quercus fournieri Trel. = Quercus dysophylla Benth.</p> <p>Quercus frutex Trel.</p> <p>Quercus frutex var. uhdeana Trel. = Quercus frutex Trel.</p> <p>Quercus fulva Liebm.</p> <p>Quercus furfuracea Liebm.</p> <p>Quercus fusiformis Small</p> <p>Quercus galeanensis C.H.Mull.</p> <p>Quercus × galeottii M.Martens</p> <p>Quercus gambelii Nutt.</p> <p>Quercus gambelii var. gunnisonii (Torr. &amp; A. Gray) Wenz. = Quercus gambelii Nutt.</p> <p>Quercus gemmata Trel. = Quercus benthamii A.DC.</p> <p>Quercus gentryi C.H.Mull. = Quercus confertifolia Bonpl.</p> <p>Quercus germana Schltdl. &amp; Cham.</p> <p>Quercus germana var. echinata Trel. = Quercus germana Schltdl. &amp; Cham.</p> <p>Quercus germana Schltdl. &amp; Cham. var. lemmonii Trel. = Quercus polymorpha Schltdl. &amp; Cham.</p> <p>Quercus ghiesbreghtii M.Martens &amp; Galeotti = Quercus sartorii Liebm.</p> <p>Quercus glabrescens Benth.</p> <p>Quercus glabrescens var. integrifolia Liebm. ex A.DC. = Quercus glabrescens Benth.</p> <p>Quercus glauca Oerst. (1869) non Thunb. (1784) = Quercus sororia Liebm.</p> <p>Quercus glaucescens Bonpl.</p> <p>Quercus glaucoides M.Martens &amp; Galeotti</p> <p>Quercus glaucophylla Seemen = Quercus glaucoides M.Martens &amp; Galeotti</p> <p>Quercus glaucophylla f. lobata C.H.Mull. = Quercus laceyi Small</p> <p>Quercus glaucophylla f. longifolia C.H.Mull. = Quercus laceyi Small</p> <p>Quercus glaucophylla var. macropetiolata C.H.Mull. = Quercus laceyi Small</p> <p>Quercus glaucophylla var. subrotundifolia C.H.Mull. = Quercus laceyi Small</p> <p>Quercus glaucophylla f. tlacolulana Trel. = Quercus glaucoides M.Martens &amp; Galeotti</p> <p>Quercus gracilior C.H.Mull. = Quercus cortesii Liebm.</p> <p>Quercus × graciliramis C.H.Mull.</p> <p>Quercus grahamii Benth.</p> <p>Quercus grahamii f. brevipes Trel. = Quercus grahamii Benth.</p> <p>Quercus grahamii var. coyulana Trel. = Quercus acutifolia Neé</p> <p>Quercus grahamii var. nelsonii Trel. = Quercus grahamii Benth.</p> <p>Quercus grandis Liebm. = Quercus xalapensis Bonpl.</p> <p>Quercus grandis var. tenuipes Trel. = Quercus xalapensis Bonpl.</p> <p>Quercus granulata Liebm. (1854) non Raf. (1838) = Quercus salicifolia Neé</p> <p>Quercus gravesii Sudw.</p> <p>Quercus greggii (A.DC.) Trel.</p> <p>Quercus greggii Trel. f. subglabrata C.H.Mull. = Quercus undulata Torr.</p> <p>Quercus grisea Liebm.</p> <p>Quercus guatimalensis A.DC = Quercus polymorpha Schltdl. &amp; Cham.</p> <p>Quercus guayabalana Trel. = Quercus elliptica Neé</p></div> 	https://treatment.plazi.org/id/794B8792FF9BFFBA3E88FF67FD11F7F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FFEDFFCC3E88FCFAFBF0FC29.text	794B8792FFEDFFCC3E88FCFAFBF0FC29.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus furfuracea Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854	<div><p>Quercus furfuracea Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 189 (1854).</p> <p>Type:— MEXICO, Puebla: F. M. Liebmann 3438 (Lectotype: C10013679! designated by Trelease (1924, p. 228,</p> <p>pl. 382a); isolectotypes: C!, CAS!, GH!, K (as s.n.)!, L!, LE!, NY!, P (as s.n.)!, UCSB (fragment)!). The K and P</p> <p>s.n. sheets are almost certainly syntypes as they are the right taxon from the right locality on the right date. Quercus acutifolia Neé var. furfuracea (Liebm.) Oerst., Chênes Amér. Trop. 28, pl. C, pl. 12(3, 3) (1869). Distribution: Mexico (Hidalgo, Michoacán, Oaxaca, Puebla, Querétaro, San Luis Potosí, Tamaulipas). The occurrence of Q. furfuracea in Querétaro reported by Villaseñor (2016) needs to be confirmed.</p> </div>	https://treatment.plazi.org/id/794B8792FFEDFFCC3E88FCFAFBF0FC29	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FFEDFFCC3E88FBDAFB62F7F1.text	794B8792FFEDFFCC3E88FBDAFB62F7F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus fusiformis Small, Bull. Torrey Bot.	<div><p>Quercus fusiformis Small, Bull. Torrey Bot. Club 28(6): 357 (1901). Type:— U.S.A., Texas: H. Lacey s. n. (1899–1900)</p> <p>(Holotype: NY00248638!; isotypes: CAS!, UCSB (fragment)!). Quercus sillae Trel., Mem. Natl. Acad. Sci. 20: 102, pl. 171b (1924). Type:— MEXICO, Nuevo León: C. G. Pringle</p> <p>11332 (Holotype: US 00089646!; isotypes: A!, GH!, LL (image)!, MEXU!, S (image)!, UCSB (fragment)!, VT!). Quercus virginiana Mill. subsp. fusiformis (Small) A.E.Murray, Kalmia 13: 28 (1983). Quercus virginiana var. fusiformis (Small) Sarg., Bot. Gaz. 65(5): 448 (1918). Distribution: Mexico (Chihuahua, Coahuila, Nuevo León, San Luis Potosí, Tamaulipas, Veracruz), U.S.A. (Oklahoma, Texas).</p> <p>Quercus galeanensis C.H.Mull., J. Arnold Arbor. 17(3): 173 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 2168 (Holotype: A00069895!; isotypes: F!, ILL!, MO!, UCSB (fragment)!). This species is close to Q. depressa, and additional study is needed to determine if it is really distinct.</p> <p>Distribution: Mexico (Coahuila, Nuevo León, San Luis Potosí, Tamaulipas).</p> <p>Quercus gambelii Nutt., Proc. Acad. Nat. Sci. Philadelphia 4(1): 22 (1848). Type:— U.S.A.: W. Gambel s. n. (Holotype: PH?; isotypes: GH?!, K!). It is uncertain whether the actual collection locality of Gambel s.n. was in New Mexico or Texas. The type locality is just “On the banks of the Rio del Norte” which is an alternate name for the Rio Grande. For additional synonyms based on non Latin American types see Nixon (1997).</p> <p>Quercus albifolia C.H.Mull., Amer. Midl. Naturalist 27(2): 470 (1942). Type:— MEXICO, Chihuahua: C. H. Muller 3535 (Holotype: US 00089414!; isotypes: GH!, UC!).</p> <p>Quercus douglasii Hook. &amp; Arn. var. gambelii (Nutt.) A.DC., Prodr. 16(2.1): 23 (1864).</p> <p>Quercus douglasii var. novomexicana A.DC, Prodr. 16(2.1): (1864). Type:— U.S.A., New Mexico: A. Fendler 809 (Lectotype (designated here): MO-204683(image)!; isolectotypes: CAS!, K(image)!, MU(image)!, NY(image)!, S(image)!, US (image)!). There were two syntypes for this name.</p> <p>Quercus gambelii Nutt. var. gunnisonii (Torr. &amp; A. Gray) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 190 (1884).</p> <p>Quercus lesueuri C.H.Mull., Amer. Midl. Naturalist 27(2): 476 (1942). Type:— MEXICO, Chihuahua: H. LeSueur 1498 (Holotype: BH; isotypes: CAS!, GH!, MO!, TEX (image)!, UCSB (fragment from BH holotype)!).</p> <p>Quercus marshii C.H.Mull., Amer. Midl. Naturalist 18(5): 848 (1937). Type:— MEXICO, Coahuila: F. L. Wynd &amp; C. H. Muller 640 (Holotype: ILL00007506!; isotypes: ILL!, K!, MEXU!, MO!, NY!, WIS!).</p> </div>	https://treatment.plazi.org/id/794B8792FFEDFFCC3E88FBDAFB62F7F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FFE8FFDD3E88F8C2FAD9F935.text	794B8792FFE8FFDD3E88F8C2FAD9F935.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus greggii (A. DC.) Trel., Contr. U. S. Natl. Herb.	<div><p>Quercus greggii (A.DC.) Trel., Contr. U.S. Natl. Herb. 23(2): 185 (1922).</p> <p>Quercus aculcingensis Trel., Contr. U.S. Natl. Herb. 23(2): 185 (1922). Nom. nov. for Q. reticulata Bonpl. var. crassifolia Oerst. (1869) non Q. crassifolia Benth. (1809).</p> <p>Quercus alpescens Trel., Mem. Natl. Acad. Sci. 20: 83, pl. 122b (1924). Type:— MEXICO: C. Ehrenberg 109 2 (Holotype at B destroyed. Lectotype (designated here): P00754039!). Trelease (1924, p. 83) wrote “ Ehrenberg, 1091–3, Jan., 1840, the types” and on p. 219 for the caption of pl. 122b he wrote “The type (Mexico, Ehrenberg, 1093)” for a photograph of a specimen at B. The first statement indicates that there were three syntypes, but the second statement indicates that there is just the holotype. In either case all three specimens would have been at B and were destroyed.</p> <p>Quercus loeseneri Trel., Mem. Natl. Acad. Sci. 20: 79, pl. 110b (1924). Type:— MEXICO, Coahuila: C. G. Pringle 10120 (Holotype at B destroyed. Lectotype (designated here): US 00089566!; isolectotypes: C!, E!, G!, GH!, HBG (image) !, ILL!, K!, L!, MEXU!, MO!, MP!, NY!, P !, PH!, UC!, UCSB (feagment) !, VT !).</p> <p>Quercus reticulata Bonpl. var. crassifolia Oerst., Chênes Amér. Trop. 29, pl. 34 (10–16) (1869). Type:— MEXICO, Puebla: F. M. Liebmann 3548 (p.p. as to Cupulif. Lbm. 151) (Lectotype (designated here): C 10012458!). Trelease (1924, p. 219, pl. 110a) wrote “The types of Q. reticulata crassifolia (Mexico, Liebmann, 151).” The plate shows mostly Liebmann 3548 (Cupulif. Lbm. 151), plus a small part of Liebmann 3548 (Cuplulif. Lbm. 155) mounted on the same sheet, which were given different barcode numbers at C and were collected at different dates although from the same locality. Because Trelease wrote types rather than type, even though he mentioned Cupulif. Lbm. 151, he did not effectively lectotypify specimen Cupulif. Lbm. 151 which we are now lectotypfying. Because Liebmann 3548 (Cupulif. Lbm. 155) was collected on a different date, it cannot be considered an isolectotype.</p> <p>Quercus reticulata var. greggii A.DC., Prodr. 16(2.1): 34 (1864). Type:— MEXICO, Coahuila: J. Gregg 380 (Holotype: G!; isotypes: GH!, MO!, UCSB (fragment)!).</p> <p>Quercus revoluta Trel. f. dysophyllopsis Trel., Mem. Natl.Acad. Sci. 20: 79, pl. 112 (1924). Type:— MEXICO, Hidalgo: J. Berlandier 411 (Holotype: G00087110!; isotypes: BM!, FI!, G!, P!, UCSB (fragment from G holotype)!, UCSB (fragment)!).</p> <p>Distribution: Mexico (Chihuahua, Coahuila, Durango, Hidalgo, Jalisco, México, Michoacán, Morelos, Nayarit,</p> <p>Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Tamaulipas, Tlaxcala, Veracruz, Zacatecas). The occurrence of Q. greggii in Nayarit reported by Villaseñor (2016) needs to be confirmed.</p> <p>Quercus grisea Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 171 (1854). Type:— U.S.A., Texas: C. Wright 665 (Holotype: K000832239!; isotypes: G!, GH!, K!, MO!, NY!, P!, UCSB (fragment)!, US!).</p> <p>Quercus deserticola Trel., Mem. Natl. Acad. Sci. 20: 79, pl. 113 (1924). Type:— MEXICO, Hidalgo: C. A. Purpus 1233 (Holotype at B destroyed. Lectotype (designated here): GH02003320!; isolectotypes: GH!, MO!, NY!, UC!, VT!). Neither an isotype of Uhde 309, nor duplicates of any of the Uhde paratypes from B have been found. The lectotype is designated from among the other paratypes that are the same species (Q. grisea) as the holotype photograph in Trelease (1924) pl. 113.</p> <p>Quercus jaralensis Trel. f. berlandieri Trel., Mem. Natl. Acad. Sci. 20: 86, pl. 134 (1924). Type:— MEXICO, San Luis Potosí: J. Berlandier 1285 (Holotype: GH00062249!; isotype: UCSB (fragment probably from GH holotype)!).</p> <p>Quercus santaclarensis C.H.Mull., Amer. Midl. Naturalist 19(3): 583 (1938). Type:— MEXICO, Chihuahua: F. Shreve 7951 (Holotype: BH; isotypes: MICH!, UCSB (fragment from BH holotype)!, US!). An isotype in the protologue is also cited as being at the Desert Laboratory, Tucson, Arizona.</p> <p>Quercus undulata Torr. var. grisea (Liebm.) Engelm., Trans. Acad. Sci. St. Louis 3: 393 (1877).</p> <p>Distribution: Mexico (Aguascalientes, Chiapas, Chihuahua, Ciudad de México, Coahuila, Colima, Durango,</p> <p>Guanajuato, Guerrero, Hidalgo, Jalisco, México, Michoacán, Morelos, Nuevo León, Oaxaca, Puebla, Querétaro, San</p> <p>Luis Potosí, Sinaloa, Sonora, Tamaulipas, Tlaxcala, Veracruz, Zacatecas), U.S.A. (Arizona, New Mexico, Texas).</p> <p>Quercus hinckleyi C.H.Mull., Cont. Texas Res. Found., Bot. Stud. 1: 40, pl. 11 (1951). Type:— U.S.A., Texas: C. H. Muller &amp; L. C. Hickley 8901 (Holotype: BH; isotypes: A!, BRIT (image)!, F!, LL (image)!, MO!, NY!, TEX (image)!, UC!, UCSB (fragment)!, US!).</p> <p>Distribution: Mexico (Chihuahua), U.S.A. (Texas).</p> <p>Quercus hintonii E.F.Warb., Bull. Misc. Inform. Kew 1939 (2): 91 (1939). Type:— MEXICO, México: G. B. Hinton 6359 (Holotype: K000575008!; isotypes: A!, BM!, F!, G!, K!, NY!, UCSB (fragment)!).</p> <p>Quercus ochroesthes E.F.Warb., Bull. Misc. Inform. Kew 1939 (2): 94 (1939). Type:— MEXICO, México: G. B. Hinton 6576 (Lectotype (designated here): K000575014!; isolectotypes: BM!, G!, GBH (image)!, K!, UCSB (fragment)!). There are two specimens at K, both of which are annotated by E.F.Warburg as “type.”</p> <p>Distribution: Mexico (Guerrero, México, Michoacán).</p> <p>Quercus hintoniorum Nixon &amp; C.H.Mull., Brittonia 45(2): 147, fig. 1 (1993). Type:— MEXICO, Coahuila: R. M. Straw &amp; M. Forman 1361 (Holotype: US; isotype: CAS!).</p> <p>Distribution: Mexico (Coahuila).</p> <p>Quercus hirtifolia L.M.Vázquez, S.Valencia &amp; Nixon, Brittonia 56(2): 137 (2004). Type:— MEXICO, Puebla: M.L.Vázquez 3069 (Holotype: MEXU (image)!; isotypes:BH, FCME).The specimen from Coahuila of C. B. Cowan 3566 at CAS! may possibly be a hybrid between Q. jonesii and Q. hirtifolia although there are no reported collections of Q. hirtifolia from Coahuila.</p> <p>Distribution: Mexico (Hidalgo, Puebla, Veracruz).</p> <p>Quercus huicholensis R.A.McCauley, Phytotaxa 471(3): 249. (2020). Type:— MEXICO, Jalisco: J. D. Bacon &amp; R. D. Spellenberg 1475 (Holotype: NMC (NMC-065317) (image)!; isotypes: CAS (not yet received), CIIDIR, COLO, DAV, IBUG).Although we have not seen any of the type specimens, based on the description, illustrations, and the paratypes that we have seen, Q. huicholensis is very close to Q. urbani. Additional study is needed to determine whether they are different species.</p> <p>Distribution: Mexico (Durango, Jalisco).</p> <p>Quercus humboldtii Bonpl., Pl. Aequinoct. 2: 155, pl. 130 (1809). Type:— COLOMBIA, Cundinamarca: A. Bonpland 2083 (Holotype: P00669716!; isotype: P!).</p> <p>Erythrobalanus duqueana O.Schwarz, Notizbl. Bot. Gart. Berlin-Dahlem 13: 495 (1937). Type:— COLOMBIA, Caldas: J. M. Duque 567 (Holotype: at B destroyed). The only specimen cited by Schwarz (1937) was Duque 567, but no isotypes have been found. There is a collection of Duque 1136 apparently from the same area, Río Nima, and also with the same determination at both F! and US!. Erythrobalanus duqueana is certainly Q. humboldtii but we have chosen not to neotypify the name based on Duque 1136 because it is not a recognized species.</p> <p>Erythrobalanus tolimensis (Bonpl.) O.Schwarz, Notibl. Bot. Gart. Berlin-Dahlem 13: 496. 1937.</p> <p>Quercus almaguerensis Bonpl., Pl. Aequinoct. 2: 157, pl. 131 (1809). Type:— COLOMBIA, Cuaca: A. Bonpland s. n. (Lectotype (designated here): image in Trelease (1924, pl. 269)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype. There is no doubt that Q. almaguerensis is Q. humboldtii, but no extant Bonpland collection has been found from Almaguer which is in Cauca Department, Colombia.</p> <p>Quercus boyacensis Cuatrec., Notas Fl. Colombia 6: 37 (1944). Type:— COLOMBIA, Boyacá: J. Cuatrecasas &amp; H. Garcia G. 9816 (Holotype: COL000000750 (image)!; isotypes: F!, UCSB (fragment)!, US!).</p> <p>Quercus colombiana Cuatrec., Notas Fl. Colombia 6: 37 (1944). Type:— COLOMBIA, Boyacá: E. Pérez A. &amp; J. Cuatrecasas 8100 (Holotype: COL000000751 (image)!; isotypes: F!, P!, UCSB (fragment)!, US!).</p> <p>Quercus humboldtii var. lehmanniana Hieron. ex Trel., Mem. Natl. Acad. Sci. 20: 142, pl. 273b (1924). Type:— COLOMBIA, Antioquia: F. C. Lehmann 7453 (Holotype at B destroyed. Lectotype (designated here): K000648583!).</p> <p>Quercus lindenii A.DC., Prodr. 16(2.1): 32 (1864). Type:— COLOMBIA, Tunja: J. Linden 1325 (Holotype: G00410065!; isotypes: BM!, CGE!, G!, K!, P!, UCSB (fragment from G holotype)!, USCS (fragment)!).</p> <p>Quercus tolimensis Bonpl., Pl. Aequinoct. 2: 153, pl. 129 (1809). Type:— COLOMBIA, Quindío: A. Bonpland 4165 (Holotype: P00669733!; isotypes: P (as s.n.)!, UCSB (as s.n.) (fragment)!).</p> <p>Distribution: Colombia, Panama.</p> <p>Quercus hypoxantha Trel., Mem. Natl. Acad. Sci. 20: 170, pl. 339b (1924). Type:— MEXICO, Coahuila: C. G. Pringle 10227 (Holotype: GH00444446!; isotypes: ASU (image)!, CAS!, ENCB (image)!, LL (image)!, NY!, PH!, UC!, UCSB (fragment)!, UNM (image)!, US!, VT!, WIS!). Quercus hypoxantha forms a species complex in northern Mexico that has been studied by Nixon and Muller (1993). Several segregate species have been recognized. The complex could be treated in a broad sense as mostly a single species, but within this complex, Q. coahuilensis and Q. miquihuanensis are here treated as distinct.</p> <p>Quercus errans Trel. f. graciliramis. C.H.Mull., J. Arnold Arbor. 17(3): 169 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 1273 (Lectotype (designated here): A00069935!; isolectotypes: MEXU!, P!, UCSB (fragment)!). Muller (1936) designated two syntypes.</p> <p>Distribution: Mexico (Chihuahua, Coahuila, Guanajuato, Nuevo León, Querétaro, San Luis Potosí, Tamaulipas,</p> <p>Zacatecas). The occurrence of Q. hypoxantha in Guanajuato reported by Villaseñor (2016) needs to be confirmed.</p> <p>Quercus iltisii L.M.González, Brittonia 55(1): 54, figs. 1, 3, 4 (2003). Type:— MEXICO, Jalisco: L. M. González V. et al. 4657 (Holotype: IBUG0160259 (image)!; isotypes: BH, CHAPA, ENCB, F (image)!, IEB, MEXU, MICH (image)!, NMC, NY (image)!, UC, US (image)!, WIS (image)!, ZEA).</p> <p>Distribution: Mexico (Colima, Jalisco).</p> <p>Quercus insignis M.Martens &amp; Galeotti, Bull.Acad. Roy.Sci.Bruxelles 10(1): 219 (1843). Type:— MEXICO, Veracruz: H. Galeotti 125 (Lectotype: BR0000006912260! designated by Trelease (1924, p. 216, pl. 14); isolectotypes: G!, P!). H. Galeotti 123, 124, and 125 at BR are all syntypes, but Trelease (1924, p. 216) effectively lectotypified the name in the caption to his pl. 14.</p> <p>Quercus davidsoniae Standl., Publ. Field Mus. Nat. Hist., Bot. Ser. 22(1): 14 (1940). Type:— PANAMA, Chiriquí: M. E. Davidson 864 (Holotype: F!; isotypes: A!, MO!, US!).</p> <p>Quercus insignis subsp. oocarpa (Liebm.) A.E.Murray, Kalmia 13: 26 (1983).</p> <p>Quercus insignis var. oocarpa (Liebm.) A.E.Murray, Kalmia 13: 26 (1983).</p> <p>Quercus insignis var. strombocarpoides (Liebm.) Oerst., Chênes Amér. Trop. 29, pl. H, pl. 28 (1869).</p> <p>Quercus oocarpa Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 184 (1854). Type:— GUATEMALA: J. R. R. Warszewicz50 (Holotype at B destroyed. Lectotype (designated here): G00386111!; isolectotype: UCSB (fragment from G lectotype)!). Warszewicz 50a at B was both the holotype of Q. oocarpa and a syntype of Q. warsewiczii.</p> <p>Quercus schippii Standl., Publ. Carnegie Inst. Wash. 461: 53 (1935). Type:— BELIZE, El Cayo: W. A. Schipp 1248 (Holotype: F (accn. 733679)!; isotypes: BM!, CAS!, G!, GH!, K!, LL (image)!, MICH!, MO!, NY!, S (image)!, UCSB (fragment)!).</p> <p>Quercus seibertii C.H.Mull., U.S.D.A. Bur. Pl. Industr. Misc. Publ. 477: 19, pl. 6, 7 (1942). Type:— PANAMA, Chiriquí: R. J. Seibert 225 (Holotype: A00055576!; isotypes: K!, MO!, NY!, UCSB (fragment)!).</p> <p>Quercus strombocarpa Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 176 (1854). Type:— MEXICO, Veracruz: F. M. Liebmann 3563 (Lectotype (designated here): C 10012531!; isolectotypes: C!, CAS!, GH!, M!, UCSB (fragment from C lectotype)!). Trelease (1924, p. 216) in the caption for pl. 17 wrote “Type material (Mexico foliage, Liebmann, 174=3563: fruit, Liebmann, Nov., 1841).” The Liebmann 3563 Cupulif. Lbm. 174 sheet at C is sterile, but presumably the acorn and cup also shown on pl. 17 are stored separately. Because he called this sheet type material rather than the type, we are lectotypifying the name based on this collection.</p> <p>Quercus tomentocaulis C.H.Mull., Madroño 10(5): 130, pl. 9 (1950). Type:— HONDURAS, Morazán: L. O. Williams &amp; A. Molina 12756 (Holotype: BH; isotypes: EAP (image)!, F!, GH!, MEXU!, MO!, UC!, UCSB (fragment from BH holotype)!, US!).</p> <p>Quercus warscewiczii Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 187 (1854). Type:— PANAMA, Veraguas: B. Seemann 1230 (Lectotype (designated here): K000575035!; isolectotype: BM!). Trelease (1924, p.43) had effectively lectotypified the type of Q. warscewiczii as Warscewicz 50a at B (destroyed) but this was also his lectotype of Q. oocarpa in the same publication, so we are relectotypifying the name based on a different syntype.</p> <p>Quercus yunckeri Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 17(4): 358 (1938). Type:— HONDURAS, Comayagua: T. G. Yuncker et al. 6180 (Holotype: ILL00007538!; isotypes: F!, G!, GH!, K!, MO!, NY!, S (image)!, UCSB (fragment)!).</p> <p>Distribution: Belize, Costa Rica, Guatemala, Honduras, Mexico (Chiapas, Colima, Guanajuato, Guerrero, Hidalgo,</p> <p>Jalisco, México, Michoacán, Nayarit, Oaxaca, Puebla, Tabasco, Veracruz), Nicaragua, Panama. The occurrence of Q.</p> <p>insignis in Guanajuato, Hidalgo, México, and Michoacán reported by Villaseñor (2016) needs to be confirmed.</p> <p>Quercus intricata Trel., Contr. U.S. Natl. Herb. 23(2): 185 (1922). Nom. nov. for Q. microphylla Née var. crispata A. DC (1864) non Q. crispata Steven. (1857).</p> <p>Quercus deliquescens C.H.Mull., Phytologia 42: 289 (1979). Type:— Mexico, Chihuahua: M. C. Johnston et al. 11388 (Holotype: TEX00370409 (image)!; isotype: UCSB (fragment)!).</p> <p>Quercus derrumbaderoensis C.H.Mull., Amer. Midl. Naturalist 27(2): 471 (1942). Nom. nov. for Q. reticulata f. pungens C.H.Mull. (1936) non Q. pungens Liebm. (1854).</p> <p>Quercus intricata f. erratica Trel., Mem. Natl. Acad. Sci. 20: 85, pl. 128b (1924). Type:— MEXICO, Coahuila: C. G. Pringle 2862 (Holotype: CAS474678!; isotypes: A!, VT!). Trelease (1924) in the caption to plate 128b states that Pringle 2862 is the type of Q. intricata f. erratica and that the image was photographed at St. Louis. The specimen of Pringle 2862 now at CAS was received as part of an exchange of duplicate type specimens with Harvard, and the GH specimen now at CAS was assumed at the time to be a duplicate isotype. However, the specimen now at CAS is the specimen in Trelease’s plate 128b, so it is the holotype of this form.</p> <p>Quercus intricata f. ovata Trel., Mem. Natl. Acad. Sci. 20: 85, pl. 128a (1924). Type:— MEXICO, Coahuila: E. Palmer 746 (Holotype: MO-144164!; isotypes: GH!, NY!, UC!, UCSB (fragment from MO holotype)!, US!).</p> <p>Quercus microphylla Neé var. crispata A.DC., Prodr. 16(2.1): 36 (1864). Type:— MEXICO, Coahuila: J. Gregg 296 (Holotype: G00358136!; isotypes: CAS!, GH!, UCSB (fragment)!).</p> <p>Quercus reticulata Bonpl. f. pungens C.H.Mull., J. Arnold Arbor. 17(3): 166 (1936). Type:— MEXICO, Nuevo León: C. H. Muller 2417 (Holotype: A00055634!; isotypes: CAS!, MEXU!, MO!, UCSB (fragment)!).</p> <p>Quercus revoluta Trel. f. acuta C.H.Mull., J. Arnold Arbor. 17(3): 166 (1936). Type:— MEXICO, Nuevo León: C. H. Muller 2169 (Holotype: A00055628!; isotypes: CAS!, MEXU!, MO!, UCSB (fragment)!).</p> <p>Quercus supranitida C.H.Mull., Amer. Midl. Naturalist 27(2): 479 (1942). Nom. nov. for Q. revoluta f. acuta C.H.Mull. (1936) non Q. acuta Thunb. (1784).</p> <p>Distribution: Mexico (Chihuahua, Coahuila, Durango, Nuevo León, Oaxaca, San Luis Potosí, Tamaulipas, Zacatecas),</p> <p>U.S.A. (Texas). The occurrence of Q. intricata in Oaxaca reported by Villaseñor (2016) needs to be confirmed.</p> <p>Quercus invaginata Trel., Mem. Natl.Acad. Sci. 20: 87, pl. 137, 138a (1924). Type:— MEXICO, Coahuila: C. A. Purpus 5029 (Holotype at B destroyed. Lectotype (designated here): MO-144167!; isolectotypes: BM!, GH!, NY!, UC!, UCSB (fragment from MO lectotype)!, US!).</p> <p>Quercus invaginata f. purpusiana Trel., Mem. Natl. Acad. Sci. 20: 87, pl. 138b (1924). Type:— MEXICO, Coahuila: C. A. Purpus 5030 (Holotype at B destroyed. Lectotype (designated here): MO-144168!; isolectotypes: BM!, E!, F!, GH!, NY!, UC!, UCSB (fragment from MO lectotype)!, US!).</p> <p>Distribution: Mexico (Coahuila, Nuevo León, San Luis Potosí).</p> <p>Quercus jonesii Trel., Mem. Natl. Acad. Sci. 20: 136, pl. 257b (1924). Type:— MEXICO, Jalisco: M. E. Jones 446 (Holotype: US00089543!; isotypes: RSA (image)!).</p> <p>Quercus coccolobifolia Trel., Mem. Natl.Acad. Sci. 20: 136, pl. 258 (1924).Type:— MEXICO, Guanajuato: A. Bonpland 4269 (Holotype at B destroyed. Lectotype (designated here): P00754118!; isolectotype UCSB (fragment from P lectotype)!). The type collection number is not the same as the syntypes for Q. reticulata which are Bonpland s. n. but from the same locality. Trelease (1924, p. 136) was aware of the distinction and that Bonpland had used the specific epithet “cocolobaefolia” as an ined. name for what became the syntypes Q. reticulata. Martínez (1966) was the first person to synonymize Q. coccolobifolia under Q. jonesii.</p> <p>Quercus endlichiana Trel., Mem. Natl. Acad. Sci. 20: 141, pl. 267b (1924). Type:— MEXICO, Durango: R. Endlich 6 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 267b)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Quercus endlichiana f. minor C.H.Mull., J. Arnold Arbor. 17(3): 169 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 1157 (Holotype: A00069873!; isotype: P!).</p> <p>Quercus endlichiana f. serrata C.H.Mull., J. Arnold Arbor. 17(3): 170 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 1330 A (Holotype: A00069871!).</p> <p>Distribution: Mexico (Aguascalientes, Chihuahua, Coahuila, Durango, Guanajuato, Jalisco, Michoacán, Nayarit,</p> <p>Nuevo León, Oaxaca, San Luis Potosí, Sinaloa, Sonora, Zacatecas). The occurrence of Q. jonesii in Oaxaca reported by Villaseñor (2016) needs to be confirmed.</p> <p>Quercus kelloggii Newb., Pacif. Railr. Rep. 6(3): chapter 2, 28, 89, fig. 6 (1858). Nom. nov. at the specific level for Q. tinctoria Michx. var. californica. Newberry (1859, p. 28–29) cited Q. tinctoria var. californica as a synonym and indicated that he considered it to be neither Q. tinctoria nor Q. coccinea Münchh. although similar to both. He would also have been aware that Torrey (1857, p 138) thought that Q. tinctoria var. californica could be a new species when he wrote “it is probably a distinct species.” For these reasons we consider Q. kelloggii to be a nom. nov. for Q. tinctoria var. californica based on the same type (Bartholomew &amp; Almeda 2022).</p> <p>Quercus californica (Torrey) J.G.Cooper, Smithson. Rep. 1858: 261. (1859). Nom. illeg. because Cooper cites Q. kelloggii Newb. (1857) as a synonym which is an older name.</p> <p>Quercus sonomensis Benth. ex A.DC., Prodr. 60(2.1): 62 (1864). Type:— U.S.A., California: T. Hartweg 1966 (139) (Holotype: G00719751 (image)!; isotypes: GH (image)!, K (image)!, NY (image)!, P (image)!).</p> <p>Quercus tinctoria W. Bartram var. californica Torr., Pacif. Railr. Rep. 4(5): 138. (1857). Type:— U.S.A., California: J. M. Bigelow s. n. (1854) (Lectotype: NY01470684 (image)! designated by Bartholomew &amp; Almeda (2022, p. 28); isolectotype: GH (image)!). Torrey noted that “The greater part of the botanical collections made by D. J. M. Bigelow, in the Pacific Railroad Survey, under the charge of Captain Whipple, were submitted to me for examination in accordance with the instructions of the War Department.” Torrey also had available to him other collections which were in the Torrey Herbarium now at NY, including examples of the same oak made during the Wilkes Expedition of 1838–42, Frémont’s Expedition to California in 1845–47, and the Emory expeditions with a collection by C.C. Parry in 1850. It is clear that he relied on the collection localities of all these specimens when he wrote “It occurs throughout the valley of the Sacramento, and as far south as San Diego.” All of these NY specimens are syntypes including the collection J. M Bigelow made in the Napa Valley of California in 1854 which we are designating as the lectotype.</p> <p>Distribution: Mexico (Baja California), U.S.A. (California, Oregon).</p> <p>Quercus laceyi Small, Bull. Torrey Bot. Club 28(6): 358 (1901). Type:— U.S.A., Texas: H. Lacey s. n. (Holotype: NY). Breedlove considered Q. laceyi to be a synonym of Q. glaucoides, but Nixon &amp; Muller (1992) convincingly show that they are different species.</p> <p>Quercus breviloba (Torr.) Sarg. f. laceyi (Small) Trel., Mem. Natl. Acad. Sci. 20: 102 (1924).</p> <p>Quercus breviloba subsp. laceyi (Small) A.Camus, Chênes Texte 2: 680 (1939).</p> <p>Quercus glaucophylla f. lobata C.H.Mull., J. Arnold Arbor. 17(3): 163 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 1281 (Holotype: A00055590!; isotype: MEXU!).</p> <p>Quercus glaucophylla f. longifolia C.H.Mull., J. Arnold Arbor. 17(3): 163 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 589 (p.p.) (Holotype: A00055591!). The specimen of C. H. Muller &amp; M. T. Muller 589 at ILL is Q. calophylla, so it is not an isotype.</p> <p>Quercus glaucophylla var. macropetiolata C.H.Mull., Bull. Torrey Bot. Club 63(3): 151 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 574 (Holotype: ILL!; isotype: MEXU!).</p> <p>Quercus glaucophylla var. subrotundifolia C.H.Mull., J. Arnold Arbor. 17(3): 163 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 1282 (Holotype: A00283983!).</p> <p>Distribution: Mexico (Coahuila, Nuevo León, Tamaulipas), U.S.A. (Texas).</p> <p>Quercus laeta Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 179 (1854). Type:— MEXICO, Hidalgo: T. Hartweg 419 (Lectotype (designated here): K000512627!; isolectotypes: BM!, CAS!, CGE!, E!, G!, K!, NY!, OXF!, P!, UCSB (fragment)!). Liebmann cited syntypes of T. Hartweg 419 at B and K. Trelease (1924, p. 218, pl. 86b) effectively lectotypified the name based on Hartweg 419 at B. Because the B specimen was destroyed, we are re-lectotypifying the name based on the other syntype which is from the Hooker Herbarium at K and has a determination by Liebmann in his hand. Further study is needed regarding the relationship of Q. laeta and Q. arizonica. Breedlove believed that these two species might best be treated in a broad sense as a single species.</p> <p>Quercus bipedalis Trel., Mem. Natl. Acad. Sci. 20: 89, pl. 143 (1924). Type:— MEXICO, Nayarit: J. N. Rose s. n. (10 Aug 1897) (Holotype: US 00089417!).</p> <p>Quercus clivicola Trel. &amp; C.H.Mull., Bull. Torrey Bot. Club 63(3): 149 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 584 (Holotype: ILL00007560!; isotypes: A!, MEXU!, TEX (image)!, UCSB (fragment)!).</p> <p>Quercus clivicola f. consanguinea C.H.Mull., Bull. Torrey Bot. Club 63(3): 150 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 597 (Holotype: ILL00007529!; isotype: MEXU!).</p> <p>Quercus clivicola f. crenifolia Trel. &amp; C.H.Mull., Bull. Torrey Bot. Club 63(3): 150 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 585 (Holotype: ILL00007528!; isotypes: MEXU!, TEX (image)!, UCSB (fragment)!).</p> <p>Quercus clivicola f. dentata Trel. &amp; C.H.Mull., Bull. Torrey Bot. Club 63(3): 150 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 583 (Holotype: ILL00007527!; isotypes: A!, CAS!, TEX (image)!, UCSB (fragment)!).</p> <p>Quercus clivicola f. elongata C.H.Mull., J. Arnold Arbor. 17(3): 162 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 1323 (Holotype: A00969952!; isotypes: MEXU!, NY!, P!).</p> <p>Quercus laeta f. heterophylla Trel., Mem. Natl. Acad. Sci. 20: 71, pl. 87 (1924). Type:— MEXICO, Ciudad de México: E. Bourgeau 1009 (Holotype: P00754119!; isotypes: P!, UCSB (fragment)!). Trelease (1924, p. 71) stated that the type is “Bourgeau, without number” at P, but the photograph in pl. 87, which Trelease designated in the caption to pl. 87 as the type, is of Bourgeau 1009 and also was annotated by Trelease as Q. laeta in 1913.</p> <p>Quercus obscura Trel. non Seemen (1897), Mem. Natl. Acad. Sci. 20: 71, pl. 88 (1924). Type:— MEXICO: B.Seemann 1971 (Syntypes: C, GH00034124!, K000512630!; isosyntypes: BM!, UCSB (fragment)!). Because Q. obscura Trel. is a later homonym, we are not lectotypifying the name, but see below where we lectotypify the name Q. transmontana var. obscura based on the K specimen of Seemann 1971.</p> <p>Quercus obscura Trel. non Seemen (1897) var. perpusilla Trel., Mem. Natl. Acad. Sci. 20: 71, pl. 89a (1924). Type:— MEXICO: B. Seemann 1976 (Lectotype (designated here): K000512631!; isolectotypes: BM!, E (as s.n.)!, GH!). Trelease (1924, p. 71) cited a single collection but indicated that it was at both GH and K. In his pl. 89b he showed an image of the K specimen, but he did not indicate that it was “the type” as was his usual practice.</p> <p>Quercus pallescens Trel., Mem. Natl. Acad. Sci. 20: 89, pl. 141b (1924). Type:— MEXICO, Jalisco: J. N. Rose 2960 (Holotype: US00089603!; isotypes: NY!, UCSB (fragment)!).</p> <p>Quercus pandurata Bonpl. var. laeta (Liebm.) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 197 (1884).</p> <p>Quercus prinopsis Trel., Mem. Natl. Acad. Sci. 20: 61, pl. 58b (1924). Type:— MEXICO, San Luis Potosí: C. A. Purpus 4941 (Holotype: MO-144170!; isotypes: BM!, E!, F!, GH!, P!, UC!, UCSB (fragment from MO holotype)!, US!).</p> <p>Quercus rossii Trel. f. arsenei Trel., Mem. Natl. Acad. Sci. 20: 180 (1924). Type:— MEXICO, Michoacán: G. Arsène 5913 (Lectotype (designated here): ILL00007610!; isolectotypes: BR!, MEXU!, MO!, P!, US!). Trelease (1924, p. 180) cited three Arsène collections without indicating which is the type.</p> <p>Quercus transmontana Trel., Mem. Natl.Acad. Sci. 20: 71, pl. 89b (1924). Type:— MEXICO, Michoacán: C. G. Pringle 4116 (Holotype at B destroyed. Lectotype (designated here): US 00089673!; isolectotypes: A!, BM!, BR!, CAS!, E!, F!, G!, GH!, K!, KFTA (image) !, MEL (image) !, MEXU!, MO!, NY!, P!, PH!, UC!, UCSB (fragment) !, US!, VT !).</p> <p>Quercus transmontana f. lata (Trel.) A.Camus, Chênes Texte 2: 654 (1939).</p> <p>Quercus transmontana var. lata Trel., Mem. Natl. Acad. Sci. 20: 72 (1924). Type:— MEXICO, Michoacán: G. Arsène 2975 (Syntypes at M lost. Lectotype (designated here): US00089672!; isolectotype: MPU!). Trelease (1924, p. 72) cited two Arsène syntypes at M that were destroyed. We are lectotypifying the name based on an isosyntype.</p> <p>Quercus transmontana var. obscura A.Camus, Chênes Atlas 2: 97 (1935). Type:— Mexico: B. Seemann 1971 (Lectotype (designated here): K000512630!; isolectotypes: BM!, C!, GH (as s.n.)!, UCSB (fragment)!). Quercus obscura Trel. (1924) non Seemen (1897) is a later homonym, but the specific epithet of Trelease was used by A.Camus, based on the same type of which Trelease cited syntypes at C, GH, and K, as the basis for Q. transmontana var. obscura.</p> <p>Distribution: Mexico (Aguascalientes, Baja California Sur, Chihuahua, Ciudad de México, Coahuila, Colima,</p> <p>Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, México, Michoacán, Morelos, Nayarit, Nuevo León, Oaxaca,</p> <p>Puebla, Querétaro, San Luis Potosí, Sinaloa, Sonora, Tamaulipas, Tlaxcala, Veracruz, Zacatecas).</p> <p>Quercus lancifolia Schltdl.&amp;Cham., Linnaea 5(1):78(1830). Type:— MEXICO, Veracruz: C. J. W. Schiede 15 (Holotype at B destroyed. Lectotype: P00754120! designated by Valencia-A. &amp; Coombes (2020, p. 89); isolectotypes: BM (as s.n.)!, P!, UCSB (fragment from P holotype)!, UCSB (fragment from BM as s.n.)!). The holotype at B would have been number 20. The species of oaks are numbered in accordance with the Linnaean treatment, 17 through 24. Other oaks were apparently numbered but not in this consecutive numbering system.</p> <p>Quercus aaata C.H.Mull., U.S.D.A. Bur. Pl. Industr. Misc. Publ. 477: 27, pl. 23–26. (1942). Type:— GUATEMALA, Guatemala: P. C. Standley 58466 (Holotype: F0060002 F!; isotypes: F!, NA, UCSB (fragment)!, US).</p> <p>Quercus boqueronae Trel., Mem. Natl. Acad. Sci. 20: 48, pl. 33b (1924). Type:— MEXICO, Chiapas: C. A. Purpus 6997 (Holotype: ILL00009820!; isotypes: BM!, F!, GH!, HBG (image)!, ILL!, MO!, NY!, UC!, UCSB (fragment)!, US!, VT!).</p> <p>Quercus cyclobalanoides Trel., Proc. Amer. Philos. Soc. 54: 11, pl. 3 (1915). Type:— MEXICO, Chiapas: C. A. Purpus 6998 (Lectotype: ILL00007515! and ILL00010193!designated by Trelease (1924, p. 216, pl. 20–21); isolectotypes: BM!, GH!, F!, ILL!, MO!, NY!, UC!, UCSB (fragment)!, US!). The lectotype is mounted on two sheets, one with just leaves and the other with just acorns and cups. The sheet with leaves has the collector’s label.</p> <p>Quercus excelsa Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 174 (1854). Type:— MEXICO, Veracruz: F. M. Liebamnn 3477 (Lectotype: C 10012479! designated by Trelease (1924, p. 47); isolectotypes: C!, CAS!, GH!, K (as s.n.)!, M!, UCSB (fragment)!). Trelease (1924, p. 47) wrote “ Liebmann, 205, 3477, Jan., 1943, the type)” by which he effectively lectotypified the Cupulif. Lbm. 205 sheet at C of Liebmann 3477. On p. 216 in the caption to pl. 29 he again wrote “The type (Mexico, Liebmann, 3477)” although in pl. 29 he shows photos taken of the two syntypes at C, neither of which are the Cupulif. Lbm. 205 sheet, so the caption to pl. 29 cannot be considered effective lectotypification. The three s.n. collections at K are almost certainly syntypes of Q. excelsa as they have the correct date and locality.</p> <p>Quercus insignis M.Martens &amp; Galeotti var. pilarius (Trel.) A.E.Murray, Kalmia 13: 26 (1983).</p> <p>Quercus insignis subsp. pilarius (Trel.) A.E.Murray, Kalmia 13: 26 (1983).</p> <p>Quercus lancifolia f. pilosiuscula Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 192 (1884). Type:— MEXICO, Veracruz: C. J. W. Schiede s. n. (Sep 1829) (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 32b)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Quercus lancifolia var. monocarpa Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 192 (1884). Type:— MEXICO, Veracruz: F. M. Liebmann 3503 (Holotype: C 10012494!; isotypes: C!, CAS!, GH!, K (as s.n.)!, P!, UCSB (fragment)!). The holotype of Q. lancifolia var. monocarpa is also the lectotype of Q. leiophylla.</p> <p>Quercus leiophylla A.DC., Prodr. 16(2.1): 71 (1864). Type:— MEXICO, Veracruz: F. M. Liebmann 3503 (Lectotype: C10012494! designated by Valencia-A. &amp; Coombes (2020, p. 89); isolectotypes: C!, CAS!, GH!, K (as s.n.)!, P!, UCSB (fragment)!). Collections cited by A. de Candolle (1864) correspond to Liebmann 3505, some of which were collected in Oct 1841 and some in Nov 1841, but all at the same locality of S. Bartholome. A. de Candolle cites specimens as “ Q. lancifolia Liebm. ! in h. Copenh. et DC., non Cham. et Schl.” Of the six specimens originally at C, one is now at CAS. All six specimens were determined in A. de Candolle’s hand as Q. leiophylla. We have chosen one of the C specimens collected in Oct 1841 as the lectotype which is also the holotype of Q. lancifolia var. monocarpa Wenz.</p> <p>Quercus leiophylla f. subintegra Trel., Mem. Natl. Acad. Sci. 20: 47 (1924). Type:— MEXICO, Veracruz: H. Galeotti 129 (Lectotype (designated here): G00358129!; isolectotypes: BR!, G!, L!, P!, UCSB (fragment)!). Although Trelease (1924) cited a single specimen, he did not designate the herbarium.</p> <p>Quercus molinae C.H.Mull., Madroño 10(5): 132, pl. 10 (1950). Type:— HONDURAS, Morazán: L. O. Williams &amp; A. Molina 12753 (Holotype: BH; isotypes: EAP (image)!, F!, GH!, MEXU!).</p> <p>Quercus ovandensis Matuda, Amer. Midl. Naturalist 43(1): 198 (1950). Type:— MEXICO, Chiapas: E. Matuda 17835 (Holotype: MEXU00013001!; isotypes: F!, MEXU!, UCSB (as s.n.) (fragment)!).</p> <p>Quercus pilarius Trel., Mem. Natl.Acad. Sci. 20: 44, pl. 19 (1924). Type:— MEXICO, Chiapas: E. Reeves 11 (Holotype: ILL00007605! and ILL00010205!). The holotype is mounted on two sheets. One sheet has just leaves and the other has the acorns and cups. The original label from the collection is on the sheet with just acorns and cups.</p> <p>Quercus pilgeriana Seemen, Bull. Herb. Boissier, sér. 2, 4(7): 655 (1904). Type:— COSTA RICA, Alajuela: A. Tonduz 2197 (Holotype: BR0000006916176!; isotypes: BR!, UCSB (fragment)!, US!). The label on the BR specimen indicates that it was collected by H. Pittier, but the protologue give the collection is by A. Tonduz 2197.</p> <p>Quercus reevesii Trel., Mem. Natl. Acad. Sci. 20: 45, pl. 22 (1924). Type:— GUATEMALA, Quetzaltenango: E. Reeves s. n. (Feb 1918) (Holotype: ILL00007619! and ILL00010207!; isotype: UCSB (fragment from ILL holotype)!). The holotype is mounted on two sheets. One sheet with just leaves and the other with just acorns and cups. The sheet with leaves has the label from the collector.</p> <p>Quercus yousei Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 17(4): 358 (1938). Type:— HONDURAS, Comayagua: T. G. Yuncker et al. 6297 (Holotype: ILL00007539!; isotypes: F!, G!, GH!, K!, MO!, NY!, S (image)!, UC!, UCSB (fragment)!, US!).</p> <p>Distribution: Belize, Costa Rica, El Salvador, Guatemala, Honduras, Mexico (Chiapas, Colima, Guerrero, Hidalgo,</p> <p>Jalisco, México, Nayarit, Oaxaca, Puebla, San Luis Potosí, Tamaulipas, Veracruz), Nicaragua, Panama. The occurrence of Q. lancifolia in Colima, Nayarit, and Tamaulipas reported by Villaseñor (2016) needs to be confirmed.</p> <p>Quercus laurina Bonpl., Pl. Aequinoct. 2: 32, pl. 80 (1809). Type:— MEXICO, Hidalgo: A. Bonpland 4143 (Holotype: P00129720!; isotypes: CGE (as s.n.)!, G!, P!, UCSB (fragment)!).</p> <p>Quercus acatenangensis Trel., Mem.Natl.Acad.Sci. 20: 163, pl. 320 (1924). Type:— GUATEMALA: J. R. R. Warszewicz 48 (Syntypes at B destroyed. Lectotype (designated here): K000512943!; isolectotypes:G!, P!, UCSB (fragment)!). Nom. nov. for Q. longifolia Liebm. (1854) non Raf. (1838).</p> <p>Quercus ambivenulosa Trel., J. Wash.Acad. Sci. 23(1): 62 (1933). Type:— GUATEMALA, Chimaltenango: A. F. Skutch 14 (Holotype: US 00089407!; isotypes: ILL!, US!).</p> <p>Quercus barbinervis Benth., Pl.Hartw. 56 (1840). Type:— MEXICO, Hidalgo: T. Hartweg 427 (Holotype: K000512950!; isotypes: A!, BM!, CAS!, CGE!, E!, FI!, G!, K!, LD (image)!, LE!, NY!, P!).</p> <p>Quercus bourgaei Oerst. ex Trel., Mem. Natl. Acad. Sci. 20: 168, pl. 336a (1924). Type:— MEXICO, Ciudad de México: E. Bourgeau 1013 (Holotype: G00358130!; isotypes: BM!, BR!, C!, G!, GH!, K!, LE!, MPU!, P!, UCSB (fragment)!, US!).</p> <p>Quercus bourgaei var. ilicifolia Trel., Mem. Natl. Acad. Sci. 20: 168, pl. 336b (1924). Type:— MEXICO, Ciudad de México: C. G. Pringle 10315 (Lectotype (designated here): MO-144175!; isolectotypes: BM!, CAS!, F!, GH!, K!, ILL!, MEXU!, NY!, OKLA (image)!, P!, PH!, UC!, US!, VT!). Trelease (1924, p. 227) in the caption for plate 336 states that the type is at G, but because it could not be found there, we are lectotypifying an isotype specimen at MO which has a determination in Trelease’s hand.</p> <p>Quercus caeruleocarpa Trel., Mem. Natl. Acad. Sci. 20: 163, pl. 321a (1924). Type:— MEXICO, Ciudad de México: R. Endlich 1365a (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 321a)!). No extant original material exists, so we are lectotypifying the name based on an image of the holotype.</p> <p>Quercus castanea Neé var. tridens (Bonpl.) A.DC., Prodr. 16(2.1): 72 (1864).</p> <p>Quercus chrysophylla Bonpl., Pl. Aequinoct. 2: 42, pl. 87 (1809). Type:— MEXICO, Hidalgo: A. Bonpland 4062 (Holotype: P00669720!; isotypes: B-W!, CGE (as s.n.)!, G!, P!, UCSB (fragment)!).</p> <p>Quercus lanceolata Bonpl., Pl. Aequinoct. 2: 34, pl. 81 (1809). Type:— MEXICO, Guanajuato: A. Bonpland s. n. (Holotype: P00129724!; isotypes: B-W!, G!, P!, UCSB (fragment)!).</p> <p>Quercus lanceolata var. undulatodentata A.DC., Prodr. 16(2.1): 70 (1864). Type:— MEXICO, Veracruz: F. M. Liebmann 3496 (Lectotype (designated here): C 10012513!; isolectotypes: C!, CAS!). A.de Candolle (1864) cites two Liebmann collections at C by locality, not collection number. His citation of “In monte Orizaba alt. 8000 p.” corresponds to Liebmann 3496, and his citation of “prov. Oaxaca da Pico de Pelado alt. 9000 p.” corresponds to Liebmann 3495. Collections of both numbers at C are annotated in A.de Candolle’s hand. The placement of this variety needs additional study, but we consider the type to fall within the variation found within Q. laurina as broadly interpreted.</p> <p>Quercus laurina var. barbinervis (Benth.) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 205 (1884).</p> <p>Quercus laurina var. ocoteifolia (Liebm.) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 205 (1884).</p> <p>Quercus longifolia Liebm. non Raf. (1838), Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 185 (1854). See Q. acatenangensis above.</p> <p>Quercus malinaltepecana Trel., Repert. Spec. Nov. Regni Veg. 33: 318 (1934). Type:— MEXICO, Guerrero: L. Schultze 387 (Holotype: at B destroyed). No original material exists, so the only information is Trelease’s (1934) description. Quercus malinaltepecana might be synonymous with Q. laurina, but Breedlove thought that it might be Q. planipocula.</p> <p>Quercus nitens M.Martens &amp; Galeotti (1843) non Raf. (1838) var. ocoteifolia (Liebm.) A.DC., Prodr. 16(2.1): 69 (1864).</p> <p>Quercus nitidissima Trel., Mem. Natl. Acad. Sci. 20: 122, pl. 218b (1924). Type:— MEXICO, Querétaro: J. N. Rose et al. 9720 (Holotype: US 00089577!).</p> <p>Quercus ocoteifolia Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 176 (1854). Type:— MEXICO, Oaxaca: F. M. Liebmann 3520 (Lectotype: C 10012510! designated by Trelease (1924, p. 227, pl. 334a); isolectotypes: C!, CAS!, GH (as s.n.)!, UCSB (fragment)!).</p> <p>Quercus orizabae Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 189 (1954). Type:— MEXICO, Veracruz: F. M. Liebmann 3539 (Lectotype (designated here): C 10012521!; isolectotypes: C!, CAS!, GH (as s.n.)!, K (as s.n.)!, NY!, P (as s.n.)!, UC!, UCSB (fragment from C lectotype)!, UCSB (fragment)!). Trelease (1924) in pl. 233 shows two different specimens. In the caption to pl. 233 on p. 223 he states “The type specimens. (Mexico, Liebmann, 83-84=3539).” The upper image in pl. 233 is sheet 84 which we are designating as the lectotype, and the lower image in pl. 233 is sheet 83, an isolectotype also at C.</p> <p>Quercus roseovenulosa Trel., Mem. Natl. Acad. Sci. 20: 182, pl. 366a (1924). Type:— MEXICO, México: C. A. Purpus 1797 (Holotype: MO-144176!; isotypes: F!, GH!, NY!, UC!, UCSB (fragment from MO holotype)!, US!).</p> <p>Quercus sideroxyla Bonpl. f. ciliifera Trel., Mem. Natl. Acad. Sci. 20: 170 (1924). Type:— MEXICO, Hidalgo: C. A. Ehrenberg 1279 (Lectotype (designated here): NY00023599!; isolectotypes: BM!, P!, UCSB (fragment)!, VT!). Trelease (1924, p. 170) cited two different collections. The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. laurina as broadly interpreted.</p> <p>Quercus simillima Trel., Mem. Natl.Acad. Sci. 20: 180, pl. 363b (1924). Type:— MEXICO, San Luis Potosí: C. C. Parry &amp; E. Palmer s. n. (1878) (Holotype: MO-143908!; isotype: UCSB (fragment from MO holotype)!).</p> <p>Quercus tlapuxahuensis A.DC. var. obconica Trel., Mem. Natl. Acad. Sci. 20: 165, pl. 325b (1924). Type:— MEXICO, Ciudad de México: E. Bourgeau 1013 (Holotype: GH00055708!; isotypes: BM!, BR!, C!, G!, K!, LE!, MO!, MPU!, P!, UCSB (fragment)!, US!). Trelease (1924, p. 165, pl. 325b) considered “ Bourgeau, 1013 in part” at GH to be the type of Q. tlapuxahuensis var. obconica; it is different from Bourgeau 1013 at G which is the holotype of Q. bourgaei.</p> <p>Quercus tridens Bonpl., Pl. Aequinoct. 2: 56, pl. 96 (1809). Type:— MEXICO, Hidalgo: A. Bonpland 4061 (p.p.) (Holotype: P00669717!; isotypes: P!, UCSB (fragment)!). There are two additional sheets of Bonpland 4061 (p. p.) at P that are isotypes of Q. tridens (P00129746! and P0012974!).</p> <p>Quercus xalapensis Bonpl. var. longifolia (Liebm.) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 210 (1884).</p> <p>Quercus zempoaltepecana Trel., Mem. Natl.Acad. Sci. 20: 164, pl. 323a (1924). Type:— MEXICO, Oaxaca: E. W. Nelson 672 (Holotype: US00089682!). The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. laurina as broadly interpreted.</p> <p>Distribution: El Salvador, Guatemala, Honduras, Mexico (Aguascalientes, Chiapas, Ciudad de México, Coahuila,</p> <p>Colima, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, México, Michoacán, Morelos, Nayarit, Nuevo León,</p> <p>Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa, Tamaulipas, Tlaxcala, Veracruz, Zacatecas).</p> <p>Quercus macdougallii Martínez,Anales Inst. Biol. Univ. Nac.Autón. México 34(1 &amp; 2): 147 (1964). Type:— MEXICO, Oaxaca: T. B. MacDougall 503 (Holotype: MEXU00051895!; isotypes: CAS!, GH!, ENCB (image)!, F!, MEXU!, MICH!, US!).</p> <p>Distribution: Mexico (Oaxaca).</p> <p>Quercus magnoliifolia Neé, Anales Ci. Nat. 3: 268 (1801). Type:— MEXICO, Guerrero: L. Neé s. n. (1791) (Lectotype: MA23387! designated by Bartholomew &amp; Almeda (2022, p. 29); isolectotype: MA!).</p> <p>Quercus chartacea Trel., Mem. Natl. Acad. Sci. 20: 61, pl. 59a (1924). Type:— MEXICO, Oaxaca: C. Seler &amp; E. G. Seler 1631 (Holotype at B destroyed. Lectotype (designated here): NY01450026!; isolectotype: GH!). The holotype was E. G. Seler 62, but no isotypes have been found. The lectotype is selected from the paratypes.</p> <p>Quercus circinata Neé, Anales Ci. Nat. 3: 272 (1801). Type:— MEXICO, Guerrero: L. Neé s.n. (1791) (Lectotype: MA26472! designated by Muller &amp; McVaugh (1972, p. 518); isolectotype: MA!).</p> <p>Quercus erubescens Trel., Repert. Spec. Nov. Regni Veg. 33: 315 (1934). Type:— MEXICO, Guerrero: L. Schultze 341 (Holotype: at B destroyed). The determination of Q. erubescens as probably Q. magnoliifolia is based on the Trelease (1934) description as no images or original specimens exist.</p> <p>Quercus haematophlebia Trel., Mem. Natl. Acad. Sci. 20: 66, pl. 74 (1924). Type:— MEXICO, Nayarit: J. N. Rose 3363 (Holotype: US 00089527!).</p> <p>Quercus liebmannii Oerst. ex Trel., Mem. Natl. Acad. Sci. 20: 66, pl. 75a (1924). Type:— MEXICO, Oaxaca: F. M. Liebmann 3505 (Lectotype (designated here): C10012498!; isolectotypes: C!, CAS!). Trelease (1924) on p. 67 wrote “ Liebmann, 89, 3505, 5742, May 1842, the type”, and on p. 67 in the caption to pl. 75a he wrote “The type (Mexico, Liebmann, 89=5742).” The specimen photographed in pl. 75a is, however, Liebmann 3505 not 5742. In fact, no specimen numbered 5742 has been found at C. Because of this ambiguity we are designating Liebmann 3505 at C as the lectotype. The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. magnoliifolia as broadly interpreted.</p> <p>Quercus lutea Neé, Anales Ci. Nat. 3: 269 (1801). Type:— MEXICO, Guerrero: L. Neé s. n. (1791) (Lectotype: MA233384! designated by Bartholomew &amp; Almeda (2022, p. 29); isolectotype: MA!).</p> <p>Quercus macrophylla Neé, Anales Ci. Nat. 3: 274 (1801). Type:— MEXICO, Guerrero: L. Neé s. n. (1791) (Lectotype: MA26466! designated by Muller &amp; McVaugh (1972, p. 518); isolectotypes: MA!).</p> <p>Quercus magnoliifolia var. lutea (Neé) A.DC., Prodr. 16(2.1): 27 (1864).</p> <p>Quercus magnoliifolia var. macrophylla (Neé) A.DC., Prodr. 16(2.1): 27 (1864).</p> <p>Quercus poculifer Trel., Mem. Natl. Acad. Sci. 20: 64, pl. 69 (1924). Type:— MEXICO, Oaxaca: C. G. Pringle 4760 (Holotype: MO-144177!; isotypes: A!, BM!, BR!, CAS!, E!, ENCB (image)!, F!, G!, GH!, HBG (image)!, K!, KFTA (image)!, MEXU!, NDG (image)!, NY!, P!, PH!, UC!, UCSB (fragment)!, US!, VT!). The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. magnoliifolia as broadly interpreted.</p> <p>Quercus rubescens Trel., Repert. Spec. Nov. Regni Veg. 33: 316 (1934). Type:— MEXICO, Guerrero: L. Schultze 328 (Holotype: at B destroyed). The determination of Q. rubescens as Q. magnoliifolia is based on the Trelease (1934) description as no images or original specimens exist.</p> <p>Quercus tepicana Trel., Mem. Natl. Acad. Sci. 20: 135, pl. 355 (1924). Type:— MEXICO, Nayarit: J. N. Rose s. n. (3 Aug 1897) (Holotype: US 00089679!).</p> <p>Distribution: Mexico (Aguascalientes, Baja California Sur, Chiapas, Ciudad de México, Coahuila, Colima, Durango,</p> <p>Guanajuato, Guerrero, Hidalgo, Jalisco, México, Michoacán, Morelos, Nayarit, Oaxaca, Puebla, Querétaro, San Luis</p> <p>Potosí, Sinaloa, Veracruz, Zacatecas).</p> <p>Quercus mcvaughii Spellenb.,Amer. J. Bot. 79(10): 1200, fig. 1 (1992). Type:— MEXICO, Chihuahua: R. D. Spellenberg et al. 9821 (Holotype: NMC (accn. 059633) (image)!; isotypes: ASU (image)!, CAS!, CIIDIR (image)!, IBUG (image)!, MEXU!, NY!, MO!, US!, K!). This species is very close to Q. crassifolia and could be treated at the infraspecific level.</p> <p>Distribution: Mexico (Chihuahua, Durango, Jalisco, San Luis Potosí, Sinaloa, Sonora).</p> <p>Quercus meavei Valencia-A., Sabas &amp; O. J.Soto, Phytotaxa 269(2): 121 (2016). Type:— MEXICO, Hidalgo: Valencia &amp; Flores 5286b (Holotype: FCME; isotypes: HUMO, HGMO, HUAP). The holotype has not been seen. Some of the paratypes that have been seen are Q. xalapensis, but based on the protologue and illustrations, the type collection seems to represent a distinct species.</p> <p>Distribution: Mexico (Hidalgo).</p> <p>Quercus melissae Nixon &amp; Barrie, Novon 25(4): 447, fig. 3 (2017). Type:— MEXICO, Chiapas: L. Kaplan &amp; E. L. Davis 358 (p.p.) (Holotype: F!; isotypes: A!, F!, G!). There are also collections of Kaplan &amp; Davis 358 at F which are Q. acutifolia so they are not isotypes. This species is close to Q. segoviensis, and additional study is needed to determine if it is distinct.</p> <p>Distribution: Guatemala, Mexico (Chiapas).</p> <p>Quercus mexicana Bonpl., Pl.Aequinoct. 2: 35, pl. 82 (1809). Type:— MEXICO, Hidalgo: A. Bonpland 4060 (Holotype: P00129716!; isotypes: CGE (as s.n.)!, P!, UCSB (fragment)!).</p> <p>Quercus castanea Neé var. integra Oerst., Bidra Kundsk. Egefamil. 362 (1871). Nom. nov. for Q. mexicana at the varietal level.</p> <p>Quercus castanea var. mexicana (Bonpl.) A.DC., Prodr. 16(2.1): 72 (1864).</p> <p>Quercus mexicana f. bonplandii Trel., Mem. Natl. Acad. Sci. 20: 174, pl. 347 (1924). Type:— MEXICO: A. Bonpland s. n. (Syntypes at B destroyed. Lectotype (designated here): P00754160!).</p> <p>Quercus pablillensis C.H.Mull.,Amer. Midl. Naturalist 27(2): 477 (1942). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 1100 (Holotype: A00055712!; isotypes: BM!, F!, G!, P!, UCSB (fragment from A holotype)!, US!).</p> <p>Quercus rugulosa M.Martens &amp; Galeotti, Bull. Acad. Roy. Sci. Bruxelles 10(1): 209 (1843). Type:— MEXICO, Hidalgo: H. Galeotti 116 (Holotype: BR0000006913618!; isotypes: K!, P!, UCSB (fragment)!).</p> <p>Quercus rugulosa f. subtruncata Trel., Mem. Natl. Acad. Sci. 20: 182 (1924). Type:— MEXICO, San Luis Potosí: E. Palmer 70 (Lectotype (designated here): A00969951!; isolectotypes: BM!, F!, GH!, MEXU!, MO!, NY!, UC!, UCSB (fragment)!, US!). Trelease (1924, p. 182) cites two specimens without designating a type. The specimen at A has a determination in Trelease’s hand.</p> <p>Distribution: Mexico (Chiapas, Ciudad de México, Coahuila, Guanajuato, Hidalgo, Jalisco, México, Michoacán,</p> <p>Morelos, Nayarit, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa, Tamaulipas, Tlaxcala, Veracruz).</p> <p>The occurrence of Q. mexicana in Jalisco, Michoacán, Nayarit, and Sinaloa reported by Villaseñor (2016) needs to be confirmed. The reports from Nayarit may possibly involve introgression with some other species.</p> <p>Quercus microphylla Neé, Anales Ci. Nat. 3: 264 (1801). Type:— MEXICO, Guanajuato: L. Neé s. n. (1791) (Lectotype: MA26469! designated by Muller &amp; McVaugh (1972, p. 519); isolectotypes: MA!, UCSB (fragment)!). Although there are differences between Q. microphylla, Q. cordifolia, and Q. striatula, they all form a broadly defined species. Further study would be important for this group.</p> <p>Quercus cordifolia Trel., Mem. Natl. Acad. Sci. 20: 84, pl. 125b (1924). Type:— MEXICO, Coahuila: E. Palmer 1278 (Holotype: GH00055707!; isotypes: AC!, K!, UCSB (fragment)!, US!).</p> <p>Quercus striatula Trel., Mem. Natl. Acad. Sci. 20: 93, pl. 151 (1924). Type:— MEXICO, Durango: R. Nelson 5001 (Holotype at B destroyed. Lectotype (designated here): GH00969979!; isolectotypes: K!, US!). No isotypes of the original type, Endlich 3 (p.p.), have been found. The lectotype is selected from the paratypes.</p> <p>Quercus striatula f. otinapensis Trel., Mem. Natl. Acad. Sci. 20: 94, pl. 152a (1924). Type:— MEXICO, Durango: E. Palmer 445 (Holotype: GH00055599!; isotype: US!).</p> <p>Distribution: Mexico (Aguascalientes, Chihuahua, Ciudad de México, Coahuila, Durango, Guanajuato, Hidalgo,</p> <p>Jalisco, México, Nayarit, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa, Tamaulipas, Tlaxcala,</p> <p>Veracruz, Zacatecas).</p> <p>Quercus miquihuanensis Nixon &amp; C.H.Mull., Brittonia 45(2): 152, fig. 3 (1993). Type:— MEXICO, Tamaulipas: K. C. Nixon et al. 4051-I (Holotype: BH (image)!). This species is part of the Q. hypoxantha complex of northeastern Mexico, but Q. miquihuanensis is sufficiently distinct to be treated here as a separate species.</p> <p>Distributon: Mexico (Nuevo León, Tamaulipas).</p> <p>Quercus mohriana Buckley ex Rydb., Bull. New York Bot. Gard. 2(6): 219, pl. 31, figs. 1, 2 (1901). Type:— U.S.A., Texas: S. B. Buckley s. n. (1875) (Lectotype: NY00248752! designated by Trelease (1924 p. 220, pl. 153b) and redundantly relectotypificed by Dorr &amp; Nixon (1985, p. 218); isolectotype: NY!). The distinction between Q. mohriana and Q. tinkhamii needs additional study. If Q. tinkhamii is included as a synonym, the distribution of Q. mohriana becomes much broader in Mexico.</p> <p>Distribution: Mexico (Chihuahua, Coahuila, Nuevo León), U.S.A. (New Mexico, Oklahoma, Texas).</p> <p>Quercus muehlenbergii Engelm., Trans. Acad. Sci. St. Louis 3: 391 (1877). Type:— U.S.A.: F. T. Pursh s. n. (1806) (Holotype: PH 00038773 (image)!).</p> <p>Quercus acuminata (Michaux) Sarg. non Roxb. (1832), Silva 8: 55 (1895).</p> <p>Quercus brayi Small, Bull. Torrey Bot. Club 28(6): 358 (1901). Type:— U.S.A., Texas: H. Lacey s. n. (1899–1900) (Holotype: NY00248621!; isotype: CAS!).</p> <p>Quercus muehlenbergii var. brayi (Small) Sarg., Bot. Gaz. 65(5): 442 (1918).</p> <p>Quercus sentenelensis C.H.Mull., Amer. Midl. Naturalist 18(5): 849 (1937). Type:— MEXICO, Coahuila: F. L. Wynd &amp; C. H. Muller 629 (Lectotype (designated here): ILL00007587!; isolectotypes: A!, ILL!, K!, MO!, NY!, UCSB (fragment)!). The two specimens at ILL were marked as co-types.</p> <p>Distribution: Canada (Ontario), Mexico (Chihuahua, Coahuila, Nuevo León, Sonora, Tamaulipas), U.S.A. (Alabama,</p> <p>Arkansas,Connecticut,Florida,Georgia,Illinois,Indiana,Iowa,Kansas,Kentucky,Louisiana,Maryland,Massachusetts,</p> <p>Michigan, Minnesota, Mississippi, Missouri, Nebraska, New Jersey, New Mexico, New York, North Carolina, Ohio,</p> <p>Oklahoma, Pennsylvania, South Carolina, Tennessee, Texas, Vermont, Virginia, West Virginia, Wisconsin).</p> <p>Quercus nanchititlensis B.M.Barthol. &amp; Almeda, nom. nov. here designated for Q. platyphylla W.F. Warb. (1939) non Morogues (1877). Type:— MEXICO, México: G. B. Hinton 6360 (Holotype: K000575020! and K000575021!; isotypes: A!, BM!, F!, G!, K!, NY!). The holotype is mounted on two sheets with the second sheet consisting of additional leaves that would not fit on the first sheet.</p> <p>Quercus platyphylla E.F.Warb. non Morogues(1877), Bull.Misc. Inform.Kew1939(2):85 (1939).See Q. nanchititlensis above.</p> <p>Distribution: Mexico (Guerrero, Jalisco, México, Michoacán).</p> <p>Quercus nudinervis Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 182 (1854). Type:— MEXICO, Sinaloa: B. Seemann 1457 (Holotype: K000512617!; isotypes: BM!, GH (as s.n.)!, K!, UCSB (fragment)!). The holotype at K has the determination in Liebmann’s hand.</p> <p>Distribution: Mexico (Durango, Jalisco, Nayarit, Sinaloa).</p> <p>Quercus oblongifolia Torr., Rep. Exped. Zuni &amp; Colorado Rivers 173, pl. 19 (1853). Type:— U.S.A., New Mexico: D. Woodhouse s. n. (Holotype: NY00253711!; isotype: MO (fragment)!).</p> <p>Quercus perpallida Trel., Mem. Natl. Acad. Sci. 20: 91, pl. 147 (1924). Type:— MEXICO, Sonora: J. N. Rose et al. 13089 (Holotype: US 00089592!; isotype: NY!).</p> <p>Distribution: Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Durango, Nuevo León, Sinaloa,</p> <p>Sonora, Tamaulipas, Zacatecas), U.S.A. (Arizona, New Mexico, Texas).</p> <p>Quercus obtusata Bonpl., Pl. Aequinoct. 2: 26, pl. 76 (1809). Type:— MEXICO, Michoacán: A. Bonpland 4329 (Holotype: P00129713!; isotypes: P!, UCSB (fragment from P holotype)!, UCSB (fragment)!).</p> <p>Quercus atriglans E.F.Warb., Bull. Misc. Inform. Kew 1939 (2): 88 (1939). Type:— MEXICO, México: G. B. Hinton 6549 (Holotype: K000512995!; isotypes: BM!, F!, G!, GH!, K!, MO!, UCSB (fragment)!, VT!).</p> <p>Quercus centralis Trel. f. heterophylla Trel., Mem. Natl. Acad. Sci. 20: 62 (1924). Type:— MEXICO, Puebla: G. Arsène 7052 (Holotype at B destroyed. Lectotype (designated here): US 00089460!; isolectotypes: MO!, MPU!).</p> <p>Quercus crenatifolia Trel., Mem. Natl. Acad. Sci. 20: 57, pl. 52b (1924). Type:— MEXICO, Jalisco: M. E. Jones 440 (Holotype: MO-143884!; isotypes: DS!, UCSB (fragment)!, US!).</p> <p>Quercus innuncupata Trel., Mem. Natl. Acad. Sci. 20: 77, pl. 108 (1924). Type:— MEXICO, Michoacán: G. Arsène 5826 (Holotype: F (accn. 416692)!; isotypes: ILL!, MPU!, PH!, US!).</p> <p>Quercus obtusata var. pandurata (Bonpl.) A.DC., Prodr. 16(2.1): 27 (1864).</p> <p>Quercus pandurata Bonpl., Pl. Aequinoct. 2: 28, pl. 77 (1809). Type:— MEXICO, Michoacán: A. Bonpland 4330 (Holotype: P00129711!; isotypes: B-W!, P!, USSB (fragment)!).</p> <p>Quercus panduriformis Trel., Mem. Natl. Acad. Sci. 20: 62, pl. 62b (1924). Type:— MEXICO, Colima: H. Ross 519 (Lectotype (designated here): M-0244466!). Trelease (1924, p. 63) cites Endlich 1357 as the type of Q. panduriformis which was undoubtedly at B and destroyed, but in the caption to pl. 62b he also cites Ross 579 at M as the type. No specimen of Ross 579 was found at M, but there is a specimen of Ross 519 which is the specimen photographed in pl. 62b. Trelease misread the “1” as a “7” which is an easy mistake to make. We have determined the M specimen of Ross 519 to be Q. obtusata. Note that Trelease (1924, p. 62) in the specimens examined statement for Q. obtusata does cite “? Ross, 519 ” indicating that he was uncertain, at least at one time, if Ross 519 was actually Q. obtusata, but he decided that it belonged to his new species Q. panduriformis. Because two different collections were cited by Trelease as the type of Q. panduriformis, one of which is now lost, as well as a typographic error in citing the extant specimen, we are designating Ross 519 as the lectotype of Q. panduriformis.</p> <p>Quercus panduriformis f. rubrinervis Trel., Mem. Natl. Acad. Sci. 20: 63 (1924). Type:— MEXICO, Guerrero: E. Langlassé 600 (Lectotype (designated here): US00089600!; isolectotypes: G!, GH!, K!, MPU!, P!). Trelease (1924, p. 63) cites two Langlassé collections without designating one as the type.</p> <p>Quercus reticulata Bonpl. subsp. rhodophlebia (Trel.) A.Camus f. crenata (Trel.) A.Camus, Chênes Texte 2: 581 (1939).</p> <p>Quercus rhodophlebia Trel. f. crenata Trel., Mem. Natl. Acad. Sci. 20: 74 (1924). Type:— MEXICO, Michoacán: G. Arsène 6018 (Lectotype (designated here): ILL00007616!; isolectotypes: A!, CAS!, MO!, MPU!, UCSB (fragment)!, US!). Trelease (1924, p. 74) cited two different Arsène collections without indicating where they were deposited.</p> <p>Distribution: Mexico (Aguascalientes, Chiapas, Ciudad de México, Colima, Durango, Guanajuato, Guerrero, Hidalgo,</p> <p>Jalisco, México, Michoacán, Morelos, Nayarit, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa,</p> <p>Tamaulipas, Tlaxcala, Veracruz, Zacatecas).</p> <p>Quercus oleoides Schltdl. &amp; Cham., Linnaea 5(1): 79 (1830). Type:— MEXICO, Veracruz: C. J. W. Schiede 23 (Holotype at B destroyed. Lectotype (designated here): HAL0060354!; isolectotypes: HAL!, LE!).</p> <p>Quercus lutescens M.Martens&amp;Galeotti,Bull.Acad.Roy.Sci.Bruxelles 10(1): 219(1843). Type:— MEXICO, Veracruz: H. Galeotti 91 (Lectotype: BR0000006915636! designated by Trelease (1924, p. 222, pl. 194); isolectotypes: BR!, G!, K!, P!, UCSB (fragment)!). Although M. Martens &amp; Galeotti (1843) cited only one collection number, there are two sheets of this number at BR marked as types, one of which was designated by Trelease as “The type.”</p> <p>Quercus oleoides var. australis Trel., Mem. Natl. Acad. Sci. 20: 114, pl. 192b, 193 (1924). Type:— COSTA RICA, Heredia: H. Pittier 2607 (Holotype: BR0000006911669!; isotypes: US!, UCSB (fragment from BR holotype)!, UCSB (fragment)!).</p> <p>Quercus oleoides f. lutescens (M.Martens &amp; Galeotti) Trel., Mem. Natl. Acad. Sci. 20: 114, pl. 194 (1924).</p> <p>Quercus oleoides var. lutescens (M. Martens &amp; Galeotti) A.Camus, Chênes Atlas 2: 64 (1935).</p> <p>Quercus retusa Liebm. non Raf. (1838), Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 187 (1854). Type:— COSTA RICA, Heredia: A. Oersted 4 (Holotype: C 10012516!; isotype UCSB (fragment from C holotype)!).</p> <p>Distribution: Belize, Costa Rica, Guatemala, Honduras, Mexico (Chiapas, Ciudad de México, Guerrero, Hidalgo,</p> <p>Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Tabasco, Tamaulipas, Veracruz), Nicaragua.</p></div> 	https://treatment.plazi.org/id/794B8792FFE8FFDD3E88F8C2FAD9F935	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FF9CFFBD3E88FF67FD59F7F1.text	794B8792FF9CFFBD3E88FF67FD59F7F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus grisea Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854	<div><p>Q. grisea × Q. undulata Mexico (Coahuila). R. M. Stewart 167 GH!, 182 GH!; F. L. Wynd &amp; C. H. Muller 540 A!, ILL!,</p> <p>MEXU!, NY!, P!, VT!, 565 A! Q. grisea × Q. xylina ? Mexico (Durango). R. Spellenberg 11114D CAS!, MEXU!, 11115 CAS!, MEXU! Q. grisea × Q.? Mexico (Coahuila). R. M. Stewart 2273 GH!, 2273A GH!, 2274 GH!</p> <p>Mexico (San Luis Potosí). J. Rzedowski 5719 MEXU! Q. hintonii × Q. acutifolia Mexico (México). G. B. Hinton 6073 BH!, GH!, K!, US!, 6376 BM!, F!, G!, GH!, K!, MO!,</p> <p>NY!, US!; E. Matuda 30866 CAS!, MEXU! Q. hintonii × Q. crassifolia Mexico (México). G. B. Hinton 3451 GH!, NY!, US! Q. hypoxantha × Q. hirtifolia ? Mexico (Tamaulipas). L. I. Nevling 1759 F!, GH!, MEXU! Q. hypoxantha × Q.? Mexico (Coahuila). J. Hinton 18428 GH!</p> <p>Mexico (Nuevo León). R. Torres C. 1042 MEXU! Q. intricata × Q. microphylla Mexico (Zacatecas). F. E. Lloyd 130 GH! Q. jonesii × Q. albocincta Mexico (Sinaloa). D. E. Breedlove 19264 CAS!, F!, MO! Q. jonesii × Q. devia Mexico (Chihuahua). R. Spellenberg 8602 CAS! Q. jonesii × Q. durifolia Mexico (Durango). C. H. Muller 9022 US! Q. jonesii × Q.? Mexico (Nuevo León). C. H. Muller &amp; M. T. Muller 587 A!</p> <p>Mexico (San Luis Potosí). A. Gomez 59 MEXU! Q. jonesii × Q. hirtifolia Mexico (Coahuila). C. P. Cowan 3566 CAS! Q. laceyi × Q. oblongifolia Mexico (Nuevo León). C. H. Muller &amp; M. T. Muller 1353 DS! Q. laeta × Q. grisea Mexico (Chihuahua). R. Spellenberg 10898 CAS!, MEXU! Q. laeta × Q. spicata Mexico (Durango). R. Spellenberg 10292 CAS!, 10371 CAS!, 10394 CAS!, 10395 CAS! Q. laeta × Q. undulata Mexico (Nuevo León). Mexico (Michoacán). C. H. Muller 427 A! Q. laeta × Q.? Mexico (Hidalgo). H. E. Moore 3771 A!, BM!, MEXU!, UC!, US! Q. lancifolia × Q. bumelioides Costa Rica (Cartago). P. C. Standley 34395 US! Q. laurina × Q. affinis Mexico (México). E. Ventura V. 388 MEXU! Q. laurina × Q. crassifolia Mexico (Oaxaca). S. R. Hill 1743 NY!, VT! Q. laurina × Q. mexicana Mexico (Hidalgo). C. H. Muller 9131 A!, NY! Q. laurina × Q.? Mexico (Veracruz). H. Narave F. 950 NY!; Q. magnoliifolia × Q. nanchititlensis Mexico (Guerrero). G. B. Hinton 10613 GH!, ILL!, NY!, PH!, US! Q. magnoliifolia × Q. spicata Mexico (Hidalgo). C. H. Muller 9325 MEXU!, US!</p> <p>Mexico (Oaxaca). C. Solano S. 193 WIS! Q. × manzanillana Mexico (Querétaro). H. Puig 2716 MEXU! Q. mexicana × Q.? Mexico (Nayarit). D. E. Breedlove 61442 CAS!, 61448 CAS!, 61449 CAS!, 61467 CAS! Q. mohriana × Q. sinuata Mexico (Coahuila). E. G. Marsh 33 6 GH!; R. M. Stewart 1369 GH! Q. nudinervis × Q. magnoliifolia Mexico (Sinaloa). J. N. Rose 1644 NY!, US! Q. oleoides × Q. fusiformis Mexico (Tamaulipas). H. H. Bartlett 11028 DS!, GH! Q. perseifolia × Q. laurina Mexico (Hidalgo). R. Hernandez M. 3422 CAS!, MEXU! Q. pungens × Q.? Mexico (Coahuila). M. C. Johnston et al. 12030 CAS!, NY!</p> <p>Mexico (Tamaulipas). R. L. Dressler 1929 GH! Q. purulhana × Q. segoviensis Mexico (Chiapas). D. E. Breedlove 20435 DS!, MO!, 34723 DS!, MO!, NY!; L. Kaplan</p> <p>177 F! Q. rekonis × Q. magnoliifolia Mexico (Guerrero). G. B. Hinton 10024 GH!, K!, NY!, PH!, US! Q. rugosa × Q. obtusata Mexico (Michoacán). W. A. Eggler 159 WIS! Q. rugosa × Q. spicata Mexico (Puebla). L. S. McIntyre 219 F! Q. rugosa × Q.? Mexico (Nuevo León). C. H. Muller &amp; M. T. Muller 747 A! Q. salicifolia × Q. benthamii Honduras (Francisco Morazán). P. C. Standley 746 F!</p> <p>Panama (Chiriquí). P.H.Allen 1595 GH!, ILL!, MO!, US!; F. Almeda 6275 CAS!; R. J. Schmalzel 1977 CAS!, 1982</p> <p>MO! Q. salicifolia × Q. laurina Guatemala (Huehuetenango). J. A. Steyermark 51955 A!, F!, US! Q. sapotifolia × Q. elliptica Mexico (Veracruz). F. M. Liebmann 3510 BM!, C!, CAS!, GH!, P! Q. sartorii × Q. laurina Mexico (Veracruz). J. Dorantes L. 352 CAS!, F! Q. sebifera × Q.? Mexico (Veracruz). F. Muller 1853A GH! Q. segoviensis × Q. purulhana Honduras (Francisco Morazán). L. O. Williams 18607 F!, GH!, US!</p> <p>Mexico (Chiapas). E. L. Little 9818 CAS!</p></div> 	https://treatment.plazi.org/id/794B8792FF9CFFBD3E88FF67FD59F7F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FF94FFB53E88FF67FD7AF7F1.text	794B8792FF94FFB53E88FF67FD7AF7F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus gulielmi-treleasei C. H. Mull.	<div><p>Quercus gulielmi-treleasei C.H.Mull. = Quercus cortesii Liebm.</p> <p>Quercus haematophlebia Trel. = Quercus magnoliifolia Neé</p> <p>Quercus hahnii Trel. = Quercus dysophylla Benth.</p> <p>Quercus harmsiana Trel. = Quercus glaucoides M.Martens &amp; Galeotti</p> <p>Quercus hartmanii Trel. = Quercus toumeyi Sarg.</p> <p>Quercus hartwegii Benth. = Quercus spicata Bonpl.</p> <p>Quercus hartwegii f. glabrata Trel. = Quercus × warburgii A. Camus</p> <p>Quercus hastata Liebm. = Quercus emoryi Torr.</p> <p>Quercus hemipteroides C.H.Mull. = Quercus skinneri Benth.</p> <p>Quercus hinckleyi C.H.Mull.</p> <p>Quercus hintonii E.F.Warb.</p> <p>Quercus hintoniorum Nixon &amp; C.H.Mull.</p> <p>Quercus hirtifolia L.M.Vázquez, S.Valencia &amp; Nixon</p> <p>Quercus hondurensis Trel. = Quercus elliptica Neé</p> <p>Quercus huicholensis R.A.McCauley</p> <p>Quercus huitamalcana Trel. = Quercus cortesii Liebm.</p> <p>Quercus humboldtii Bonpl.</p> <p>Quercus humboldtii var. lehmanniana Hieron. ex Trel. = Quercus humboldtii Bonpl.</p> <p>Quercus hypoleuca Engelm. (1876) non Miq. (1858) = Quercus epileuca Trel.</p> <p>Quercus hypoleucoides A.Camus = Quercus epileuca Trel.</p> <p>Quercus hypoxantha Trel.</p> <p>Quercus idonea Goldman = Quercus tuberculata Liebm.</p> <p>Quercus ignaciensis C.H.Mull. = Quercus elliptica Neé</p> <p>Quercus iltisii L.M.González</p> <p>Quercus imbricariifolia Trel. = Quercus dysophylla Benth.</p> <p>Quercus impressa Trel. = Quercus castanea Neé</p> <p>Quercus incarnata Trel. = Quercus sideroxyla Bonpl.</p> <p>Quercus incarnata f. ampla Trel. = Quercus sideroxyla Bonpl.</p> <p>Quercus incarnata f. grosse-serrata Trel. = Quercus sideroxyla Bonpl.</p> <p>Quercus incarnata f. longa Trel. = Quercus sideroxyla Bonpl.</p> <p>Quercus incrassata Trel. = Quercus elliptica Neé</p> <p>Quercus infralutea Trel. = Quercus alveolata Trel.</p> <p>Quercus innuncupata Trel. = Quercus obtusata Bonpl.</p> <p>Quercus insignis M.Martens &amp; Galeotti</p> <p>Quercus insignis subsp. oocarpa (Liebm.) A.E.Murray = Quercus insignis M.Martens &amp; Galeotti</p> <p>Quercus insignis var. oocarpa (Liebm.) A.E.Murray = Quercus insignis M.Martens &amp; Galeotti</p> <p>Quercus insignis subsp. pilarius (Trel.) A.E.Murray = Quercus lancifolia Schltdl. &amp; Cham.</p> <p>Quercus insignis var. pilarius (Trel.) A.E.Murray = Quercus lancifolia Schltdl. &amp; Cham.</p> <p>Quercus insignis var. strombocarpoides (Liebm.) Oerst. = Quercus insignis M.Martens &amp; Galeotti</p> <p>Quercus intermedia M.Martens &amp; Galeotti (1843) non Boenn. ex Rchb. (1831) = Quercus calophylla Schltdl. &amp; Cham.</p> <p>Quercus intricata Trel.</p> <p>Quercus intricata f. angusta Trel. = Quercus frutex Trel.</p> <p>Quercus intricata f. erratica Trel. = Quercus intricata Trel.</p> <p>Quercus intricata f. ovata Trel. = Quercus intricata Trel.</p> <p>Quercus invaginata Trel.</p> <p>Quercus invaginata f. purpusiana Trel. = Quercus invaginata Trel.</p> <p>Quercus irazuensis Kuntze = Quercus costaricensis Liebm.</p> <p>Quercus jaliscensis Trel. = Quercus xylina Scheidw.</p> <p>Quercus jaralensis Trel. = Quercus potosina Trel.</p> <p>Quercus jaralensis f. berlandieri Trel. = Quercus grisea Liebm.</p> <p>Quercus jonesii Trel.</p> <p>Quercus juergensenii Liebm. = Quercus polymorpha Schltdl. &amp; Cham.</p> <p>Quercus karwinskii Trel. = Quercus canbyi Trel.</p></div> 	https://treatment.plazi.org/id/794B8792FF94FFB53E88FF67FD7AF7F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FF95FFB43E88FF66FC06F7F1.text	794B8792FF95FFB43E88FF66FC06F7F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus kelloggii Newb.	<div><p>Quercus kelloggii Newb.</p> <p>Quercus × knoblochii C.H.Mull. = Quercus ×aerea Trel.</p> <p>Quercus laceyi Small</p> <p>Quercus laeta Liebm.</p> <p>Quercus laeta f. heterophylla Trel. = Quercus laeta Liebm.</p> <p>Quercus lanceolata Bonpl. = Quercus laurina Bonpl.</p> <p>Quercus lanceolata var. undulatodentata A.DC. = Quercus laurina Bonpl.</p> <p>Quercus lancifolia Schltdl. &amp; Cham.</p> <p>Quercus lancifolia f. pilosiuscula Wenz. = Quercus lancifolia Schltdl. &amp; Cham.</p> <p>Quercus lancifolia var. monocarpa Wenz. = Quercus lancifolia Schltdl. &amp; Cham.</p> <p>Quercus langlassei Trel. = Quercus elliptica Neé</p> <p>Quercus lanigera M.Martens &amp; Galeotti = Quercus castanea Neé</p> <p>Quercus lanigera f. sideroxyloides Trel. = Quercus castanea Neé</p> <p>Quercus laurina Bonpl.</p> <p>Quercus laurina var. barbinervis (Benth.) Wenz. = Quercus laurina Bonpl.</p> <p>Quercus laurina var. major (A.DC.) Wenz. = Quercus affinis Scheidw.</p> <p>Quercus laurina var. ocoteifolia (Liebm.) Wenz. = Quercus laurina Bonpl.</p> <p>Quercus laurina var. podocarpa (A.DC.) Wenz. = Quercus affinis Scheidw.</p> <p>Quercus laxa Liebm. = Quercus xylina Scheidw.</p> <p>Quercus lecomteana Trel. = Quercus subtriloba Trel.</p> <p>Quercus leiophylla A.DC. = Quercus lancifolia Schltdl. &amp; Cham.</p> <p>Quercus leiophylla f. subintegra Trel. = Quercus lancifolia Schltdl. &amp; Cham.</p> <p>Quercus lesueuri C.H.Mull. = Quercus gambelii Nutt.</p> <p>Quercus liebmannii Oerst. ex Trel. = Quercus magnoliifolia Neé</p> <p>Quercus liebmannii f. brevipes Trel. = Quercus peduncularis Neé</p> <p>Quercus lindenii A.DC. = Quercus humboldtii Bonpl.</p> <p>Quercus linguifolia Liebm. = Quercus elliptica Neé</p> <p>Quercus loeseneri Trel. = Quercus greggii (A.DC.) Trel.</p> <p>Quercus longifolia Liebm. (1854) non Raf. (1838) = Quercus laurina Bonpl.</p> <p>Quercus lowilliamsii C.H.Mull. = Quercus benthamii A.DC.</p> <p>Quercus lutea Neé = Quercus magnoliifolia Neé</p> <p>Quercus lutescens M.Martens &amp; Galeotti = Quercus oleoides Schltdl. &amp; Cham.</p> <p>Quercus macdougallii Martínez</p> <p>Quercus macrophylla Neé = Quercus magnoliifolia Neé</p> <p>Quercus macrophylla var. rugosa (Neé) Wenz. = Quercus rugosa Neé</p> <p>Quercus magnoliifolia Neé</p> <p>Quercus magnoliifolia var. lutea (Neé) A.DC. = Quercus magnoliifolia Neé</p> <p>Quercus magnoliifolia var. macrophylla (Neé) A.DC. = Quercus magnoliifolia Neé</p> <p>Quercus major (A.DC.) Trel. (1922) non Nakai (1916) = Quercus affinis Scheidw.</p> <p>Quercus malifolia Trel. = Quercus crassipes Bonpl.</p> <p>Quercus malinaltepecana Trel. = Quercus planipocula Trel.</p> <p>Quercus × manzanillana Trel.</p> <p>Quercus marshii C.H.Mull. = Quercus gambelii Nutt.</p> <p>Quercus martensiana Trel. = Quercus peduncularis Neé</p> <p>Quercus martensiana f. berlandieri Trel. = Quercus spicata Bonpl.</p> <p>Quercus martensiana f. perplexans Trel. = Quercus peduncularis Neé</p> <p>Quercus martinezii C.H.Mull. = Quercus vicentensis Trel.</p> <p>Quercus matagalpana Trel. = Quercus segoviensis Liebm.</p> <p>Quercus mcvaughii Spellenb.</p> <p>Quercus meavei Valencia-A., Sabas &amp; O.J.Soto</p> <p>Quercus melissae Nixon &amp; Barrie</p> <p>Quercus mexiae L.M.González = Q. coffeicolor Trel.</p> <p>Quercus mexicana Bonpl.</p> <p>Quercus mexicana f. angustifolia (Bonpl.) Trel. = Quercus crassipes Bonpl.</p></div> 	https://treatment.plazi.org/id/794B8792FF95FFB43E88FF66FC06F7F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FF96FFB73E88FF67FC0CF7F1.text	794B8792FF96FFB73E88FF67FC0CF7F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus mexicana fm. bonplandii Trel.	<div><p>Quercus mexicana f. bonplandii Trel. = Quercus mexicana Bonpl.</p> <p>Quercus mexicana f. confertifolia (Bonpl.) Trel. = Quercus confertifolia Bonpl.</p> <p>Quercus mexicana var. confertifolia (Bonpl.) Wenz. = Quercus confertifolia Bonpl.</p> <p>Quercus mexicana f. glabrata Trel. = Quercus confertifolia Bonpl.</p> <p>Quercus mexicana var. glabrata Liebm. ex Seem. = Quercus confertifolia Bonpl.</p> <p>Quercus mexicana f. lanosa Trel. = Quercus crassipes Bonpl.</p> <p>Quercus mexicana f. perfertilis Trel. = Quercus crassipes Bonpl.</p> <p>Quercus microcarpa Liebm. (1854) non Lapeyr. (1813) = Quercus sapotifolia Liebm.</p> <p>Quercus microlepis Trel. &amp; C.H.Mull. (1936) non Vuk. (1883) = Quercus porphyrogenita Trel.</p> <p>Quercus microphylla Neé</p> <p>Quercus microphylla var. crispata A.DC. = Quercus intricata Trel.</p> <p>Quercus miguelitensis Trel. = Quercus crassifolia Bonpl.</p> <p>Quercus miquihuanensis Nixon &amp; C.H.Mull.</p> <p>Quercus mixtecana Trel. = Quercus glaucoides M.Martens &amp; Galeotti</p> <p>Quercus mohriana Buckley ex Rydb.</p> <p>Quercus molinae C.H.Mull. = Quercus lancifolia Schltdl. &amp; Cham.</p> <p>Quercus mollis M.Martens &amp; Galeotti (1843) non Raf. (1838) = Quercus crassifolia Bonpl.</p> <p>Quercus monserratensis C.H.Mull. = Quercus xalapensis Bonpl.</p> <p>Quercus monterreyensis Trel. &amp; C.H.Mull. = Quercus tuberculata Liebm.</p> <p>Quercus moreliana Trel. = Quercus crassifolia Bonpl.</p> <p>Quercus mucronata Willd. = Quercus castanea Neé</p> <p>Quercus muehlenbergii Engelm.</p> <p>Quercus muehlenbergii var. brayi (Small) Sarg. = Quercus muehlenbergii Engelm.</p> <p>Quercus mulleri Martínez = Quercus salicifolia Neé</p> <p>Quercus nanchititlensis B.M.Barthol. &amp; Almeda</p> <p>Quercus nectandrifolia Liebm. = Quercus elliptica Neé</p> <p>Quercus nigrirhachis Trel. = Quercus glaucescens Bonpl.</p> <p>Quercus nitens M.Martens &amp; Galeotti (1843) non Raf. (1838) = Quercus affinis Scheidw.</p> <p>Quercus nitens M.Martens &amp; Galeotti (1843) non Raf. (1838) var. major A.DC. = Quercus affinis Scheidw.</p> <p>Quercus nitens M.Martens &amp; Galeotti (1843) non Raf. (1838) var. ocoteifolia (Liebm.) A.DC. = Quercus laurina Bonpl.</p> <p>Quercus nitens M.Martens &amp; Galeotti (1843) non Raf. (1838) var. podocarpa A.DC. = Quercus affinis Scheidw.</p> <p>Quercus nitens M.Martens &amp; Galeotti (1843) non Raf. (1838) var. subintegra A.DC. = Quercus affinis Scheidw.</p> <p>Quercus nitida M.Martens &amp; Galeotti (1843) non Raf. (1838) = Quercus tlapuxahuensis A.DC.</p> <p>Quercus nitidissima Trel. = Quercus laurina Bonpl.</p> <p>Quercus nixoniana S.Valencia &amp; Lozada-Pérez = Quercus salicifolia Neé</p> <p>Quercus novomexicana (A.DC.) Rydb. = Quercus gambelii Nutt.</p> <p>Quercus nudinervis Liebm.</p> <p>Quercus oajacana Liebm. = Quercus elliptica Neé</p> <p>Quercus oblongifolia Torr.</p> <p>Quercus oblongifolia var. pallidinervis Trel. = Quercus depressipes Trel.</p> <p>Quercus obovalifolia E.Fourn. ex Trel. = Quercus crassipes Bonpl.</p> <p>Quercus obscura Trel. (1924) non Seemen (1897) = Quercus laeta Liebm.</p> <p>Quercus obscura Trel. (1924) non Seemen (1897) var. perpusilla Trel. = Quercus laeta Liebm.</p> <p>Quercus obscurirhachis Trel. = Quercus vicentensis Trel.</p> <p>Quercus obtusanthera Trel. = Quercus salicifolia Neé</p> <p>Quercus obtusata Bonpl.</p> <p>Quercus obtusata var. hartwegii (Benth.) A.DC. = Quercus spicata Bonpl.</p> <p>Quercus obtusata var. pandurata (Bonpl.) A.DC. = Quercus obtusata Bonpl.</p> <p>Quercus obtusifolia D. Don var. breviloba Torr. = Quercus sinuata Walter</p> <p>Quercus ochroesthes E.F.Warb. = Quercus hintonii E.F.Warb.</p> <p>Quercus ocoteifolia Liebm. = Quercus laurina Bonpl.</p> <p>Quercus ocoteifolia f. confusa Trel. = Quercus affinis Scheidw.</p> <p>Quercus ocoteifolia f. podocarpa (A.DC.) Trel. = Quercus affinis Scheidw.</p></div> 	https://treatment.plazi.org/id/794B8792FF96FFB73E88FF67FC0CF7F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FFEEFFC93E88FF67FBA8F8D1.text	794B8792FFEEFFC93E88FF67FBA8F8D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus novomexicana (A. DC.) Rydb., Bull.	<div><p>Quercus novomexicana (A.DC.) Rydb., Bull. New York Bot. Gard. 2(6): 208 (1901).</p> <p>Quercus stellata Wagenh. var. utahensis A.DC, Prodr. 16(2.1): 22 (1864). Type:— U.S.A. (Holotype: MO?). According to A. de Candolle (1864, p. 22) the type was collected between Salt Lake and the Sierra Nevada. The specimen was from the “(h. Engelm.!)” so it should be at MO.</p> <p>Quercus undulata Torr. var. gambelii (Nutt.) Engelm., Trans. Acad. Sci. St. Louis 3: 382 (1876).</p> <p>Quercus utahensis (A.DC.) Rydberg, Bull. New York Bot. Gard. 2(6): 202 (1901).</p> <p>Distribution: Mexico (Chihuahua, Coahuila, Nuevo León, Sonora, Veracruz), U.S.A. (Arizona, Colorado, Nevada,</p> <p>New Mexico, Oklahoma, Texas, Utah, Wyoming).</p> <p>Quercus germana Schltdl. &amp; Cham., Linnaea 5(1): 78 (1830). Type:— MEXICO, Veracruz: C. J. W. Schiede 21 (Holotype at B destroyed. Lectotype (designated here): HAL0060343!; isolectotypes: GOET (image)!, HBG (image)!, LE!, M!).</p> <p>Quercus germana var. echinata Trel., Mem. Natl. Acad. Sci. 20: 52 (1924). Type:— MEXICO, Veracruz: L. Hahn 344 (Lectotype (designated here): P00754016!). Trelease (1924, p. 52) did not indicate place of deposit for Hahn 344, but most of the Hahn specimens he cites were at B and now destroyed. The only specimen of Hahn that we found was at P; it was annotated by Trelease in 1913 as an isotype.</p> <p>Quercus substenocarpa Trel., Mem. Natl. Acad. Sci. 20: 52, pl. 41 (1924). Type:— MEXICO, San Luis Potosí: C. G. Pringle 3969 (Holotype: BR0000006915407!; isotypes: A!, BM!, E!, F!, G!, GH!, K!, MEL (image)!, MO!, NY!, P!, PH!, UC!, UCSB (fragment)!, US!, VT!).</p> <p>Distribution: Mexico (Coahuila, Guanajuato, Hidalgo, Jalisco, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis</p> <p>Potosí, Tamaulipas, Veracruz).</p> <p>Quercus glabrescens Benth., Pl. Hartw. 56 (1840). Type:— MEXICO, Hidalgo: T. Hartweg 428 (Lectotype (designated here): K000512603!; isolectotypes: A!, BM!, CGE!, E!, FI!, G!, K!, L!, LD (image)!, LE!, NY!, OXF!, P!, UCSB (fragment)!). There are two sheets of Hartweg 428 at K in the Bentham Herbarium, one of which is here lectotypified.</p> <p>Quercus ambigua Bonpl., Pl.Aequinoct. 2: 51, pl. 93 (1809). Type:— MEXICO, Hidalgo: A. Bonpland 4162 (Holotype: P00129745!; isotypes: B-W (as s.n.)!, UCSB (fragment from P holotype)!).</p> <p>Quercus bonplandiana Sweet, Hort. Brit. 370 (1826). This is an invalid name as the holotype is the same as Q. ambigua Bonpl.</p> <p>Quercus glabrescens var. integrifolia Liebm. ex A.DC., Prodr. 16(2.1): 35 (1864). Type:— MEXICO, Veracruz: F. M. Liebmann 3488 (Lectotype: C 10012484! designated by Trelease (1924, p. 217, pl. 34b); isolectotypes: C!, CAS!, UCSB (fragment)!). A. de Candolle cited “Liebm.! h. Copenh. f. 165–167.” These numbers are three Cupulif. Lbm. sheet numbers for Liebmann 3488. Trelease effectively lectotypified sheet 166 at C. Sheet 167 is at C as well as a fragment at UCSB, and sheet 165 is now at CAS.</p> <p>Quercus radlkoferiana Trel., Mem. Natl. Acad. Sci. 20: 49, pl. 35a (1924). Type:— MEXICO, Puebla: F. M. Liebmann 3483 (Holotype at B destroyed. Lectotype (designated here): C 10012485!; isolectotypes: C!, CAS!, GH!, M!, UCSB (fragment from C lectotype)!, UCSB (fragment)!). Trelease (1924, p. 49) indicated that “ Liebmann, 156, 3483 ” is the type, so the specimen of Cupulif. Lbm. 156 at C would apparently be the holotype. However, in the caption of pl. 35a on p. 217 Trelease wrote “The type Mexico, Liebmann, 156 ” and indicates that this image was taken at B. It would seem more likely that the specimen at B would have had just the number 3483 on it, which is an assigned number for a particular collection, rather than just Liebmann 156 which is an assigned number for individual sheets. Because the specimen at B was destroyed, and the label is not shown in Trelease’s pl. 35a, it is impossible to know for certain what was on the B label. For this reason, we are lectotypifying the C specimen Cupulif. Lbm. 156.</p> <p>Distribution: Mexico (Chiapas, Ciudad de México, Guanajuato, Guerrero, Hidalgo, Jalisco, México, Michoacán,</p> <p>Morelos, Oaxaca, Puebla, San Luis Potosí, Sinaloa, Tlaxcala, Veracruz, Zacatecas). The occurrence of Q. glabrescens in Querétaro and Tamaulipas reported by Villaseñor (2016) needs to be confirmed.</p> <p>Quercus glaucescens Bonpl., Pl. Aequinoct. 2: 29, pl. 78 (1809). Type:— MEXICO, Guerrero: A. Bonpland 3921 (Holotype: P00129726!; isotypes: B-W!, CGE (as s.n.)!, UCSB (fragment from P holotype)!). Although the specimen at P is marked as an isotype, it is actually the holotype and has a description in Bonpland’s hand.</p> <p>Quercus chinantlensis Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 179 (1854). Type:— MEXICO, Oaxaca: F. M. Liebmann 3471 (Lectotype: C 10012469! designated by Trelease (1924, p. 41, pl. 28b); isolectotypes: C!, CAS!).</p> <p>Quercus cuneifolia Liebm. non Raf. (1838), Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 182 (1854). See Q. pinalensis below.</p> <p>Quercus nigrirhachis Trel., Repert. Spec. Nov. Regni Veg. 33: 314 (1934). Type:— MEXICO, Guerrero: L. Schultze 493 (Holotype: at B destroyed). The determination of Q. nigrirhachis as probably Q. glaucescens is based on the Trelease (1934) description as no images or original specimens exist.</p> <p>Quercus pinalensis Trel., Mem. Natl. Acad. Sci. 20: 46, pl. 28a (1924). Type:— MEXICO, Sinaloa: B. Seemann 1456 (Holotype: K000512594!; isotypes: BM!, K!). Nom. nov. for Q. cuneifolia Liebm. (1854) non Raf. (1838). The holotype is the K specimen with the determination by Liebmann in his hand.</p> <p>Quercus synthetica Trel., Mem. Natl. Acad. Sci. 20: 151, pl. 297a (1924). Type:— MEXICO, Oaxaca: A. Ghiesbreght 121 (Holotype: C 10012534!; isotypes: P!, UCSB (fragment from C holotype)!, UCSB (fragment)!).</p> <p>Quercus synthetica var. crenifolia Trel., Mem. Natl. Acad. Sci. 20: 151, pl. 296 (1924). Type:— MEXICO, Veracruz: A. Ghiesbreght 117 (Holotype: P00754020!; isotypes: P!, UCSB (fragment)!).</p> <p>Quercus texquitzinae Trel., Repert. Spec. Nov. Regni Veg. 33: 315 (1934). Type:— MEXICO, Guerrero: L. Schultze 32 (Holotype: at B destroyed). We suspect that Q. texquitzinae is probably Q. glaucescens based on the Trelease (1934) description as no images or original specimens exist.</p> <p>Distribution: Mexico (Colima, Guerrero, Jalisco, México, Michoacán, Nayarit, Oaxaca, Puebla, Querétaro, San Luis</p> <p>Potosí, Sinaloa, Tamaulipas, Veracruz).</p> <p>Quercus glaucoides M.Martens &amp; Galeotti, Bull. Acad. Roy. Sci. Bruxelles 10(1): 209 (1843). Type:— MEXICO, Oaxaca: H. Galeotti 103 (Lectotype: BR0000009816220!designated by Trelease (1924 p. 217.pl. 42);isolectotypes: BR!, G!, K!, P!, UCSB (fragment from BR lectotype)!, UCSB (fragment)!). M. Martens &amp; Galeotti cite only one collection number, two sheets of which are at BR; either one could be considered the type. Breedlove annotated the better of these two specimens as the holotype in 1987, but Trelease had already effectively lectotypified the other specimen. The specimen which is the lectotype would be a much better specimen had C. H. Muller not removed a small branch, apparently in 1958, which is now at UCSB.</p> <p>Quercus baldoquinae Trel., Mem. Natl. Acad. Sci. 20: 53, pl. 43b (1924). Type:— MEXICO, Michoacán: R. Endlich 1354 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 43b)!). No extant original material exists, so we are lectotypifying the name on the image of the holotype.</p> <p>Quercus cancellata Trel., Mem. Natl. Acad. Sci. 20: 55, pl. 49a (1924). Type:— MEXICO, Puebla: H. Schenk 236 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 49a)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Quercus conjugens Trel., Mem. Natl.Acad. Sci. 20: 55, pl. 48 (1924). Type:— MEXICO, Guanajuato: C. G. Pringle 8841 (Lectotype (designated here): G00410068!; isolectotypes: A!, BM!, CAS!, E!, F!, FI!, G!, GH!, HBG (image)!, ILL!, K!, L!, M!, MEXU!, MO!, NY!, P!, PH!, UC!, UCSB (fragment)!, US!, VT!). Trelease (1924) indicated that the type was at B, now destroyed, and at G.</p> <p>Quercus cordata M.Martens &amp; Galeotti, Bull. Acad. Roy. Sci. Bruxelles 10(1): 211 (1843). Type:— MEXICO, Oaxaca: H. Galeotti 111 (Holotype: BR0000009816237!; isotype: UCSB (fragment from BR holotype)!).</p> <p>Quercus glaucophylla Seemen, Bot. Jahrb. Syst. 29(1): 95 (1900). Type:— MEXICO, Oaxaca: C. G. Pringle 4843 (Holotype: BR0000009816374!; isotypes: A!, BM!, CAS!, E!, ENCB (image)!, G!, GH!, K!, KFTA (image)!, MEL (image)!, MEXU!, MO!, NDG (image)!, NY!, P!, PH!, UC!, UCSB (fragment)!, US!, VT!).</p> <p>Quercus glaucophylla f. tlacolulana Trel., Mem. Natl. Acad. Sci. 20: 53 (1924). Type:— MEXICO, Oaxaca: C. Seler &amp; E. G. Seler 1752 (Holotype at B destroyed. Lectotype (designated here): GH00062248!).</p> <p>Quercus harmsiana Trel., Mem. Natl. Acad. Sci. 20: 54, pl. 45b (1924). Type:— MEXICO, Michoacán: R. Endlich 1356 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 45b)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Quercus mixtecana Trel., Mem. Natl. Acad. Sci. 20: 54 (1924). Type:— MEXICO, Oaxaca: C. Conzatti 1416 (Holotype at B destroyed. Lectotype (designated here): GH00969954!; isolectotype: A!). No extant specimens of the type number, Schenck 235, from Puebla have been found, so the lectotype is based on the paratype, Conzatti 1416, from Oaxaca.</p> <p>Distribution: Mexico (Chiapas, Chihuahua, Ciudad de México, Coahuila, Durango, Guanajuato, Guerrero, Hidalgo,</p> <p>Jalisco, México, Michoacán, Morelos, Oaxaca, Puebla, Querétaro, San Luis Potosí, Tamaulipas, Tlaxcala, Veracruz).</p> <p>Quercus grahamii Benth., Pl. Hartw. 57 (1840). Type:— MEXICO: G. J. Graham 326 (Holotype: K000512981!; isotypes: BM!, CGE!, GH!).</p> <p>Quercus acutifolia Neé f. abrupta Trel., Mem. Natl. Acad. Sci. 20: 192 (1924). Type:— MEXICO, Colima: H. Ross 534 (Lectotype (designated here): M!). Although Trelease (1924, p. 192) cited Ross 534, he did not specify the herbarium.</p> <p>Quercus grahamii f. brevipes Trel., Mem. Natl. Acad. Sci. 20: 190, pl. 384a (1924). Type:— MEXICO, Veracruz: F. M. Liebmann 3437 (Holotype: C 10012490!; isotypes: CAS!, K (as s.n.)!, P (as s.n.)!). The caption of Trelease’s plate (1924, p. 229, pl. 384a) states “The type (Mexico, Liebmann, 33=3437). Photographed by permission at Kew.” The photograph in the plate of Liebmann 3437 (Cupulif. Lbm. 33) is of the specimen at C.</p> <p>Quercus grahamii var. nelsonii Trel., Mem. Natl. Acad. Sci. 20: 190, pl. 384b (1924). Type:— MEXICO, Oaxaca: E. W. Nelson 533 (Holotype: US 00089534!; isotype: US!).</p> <p>Quercus xalapensis Bonpl. f. surculina Trel., Mem. Natl. Acad. Sci. 20: 193, pl. 393b (1924). Type:— MEXICO: W. F. Karwinski s. n. (Holotype: M!; isotype: UCSB (fragment from M holotype)!).</p> <p>Distribution: Mexico (Colima, Guerrero, Hidalgo, Jalisco, México, Michoacán, Nuevo León, Oaxaca, Puebla,</p> <p>Tamaulipas, Tlaxcala, Veracruz).</p> <p>Quercus gravesii Sudw., Check List For. Trees U.S. 86 (1927). Type:— U.S.A., Texas: G. Sudworth s. n. (5 Nov 1913) (Holotype: A00034105!). Sudworth (1927) published Q. gravesii as a nom. nov. at the specific level for Q. texana var. chesoensis.</p> <p>Quercus chesosensis (Sarg.) C.H.Mull., Amer. Midl. Naturalist 18(5): 850 (1937). This is an illegitimate name because Q. gravesii was already a nom. nov. at the specific level for Q. texana var. chesosensis.</p> <p>Quercus coccinea Trel. var. microcarpa Torr., Rep. U.S. Mex. Bound., Bot. 2(1): 206 (1859). Type:— U.S.A., Texas: C. C. Parry et al. 1375 (Lectotype (designated here): NY00248625!; isolectotypes: CAS!, UCSB (fragment)!, US!). There were two syntypes for this name.</p> <p>Quercus cupreata Trel. &amp; C.H.Mull., Bull. Torrey Bot. Club 63(3): 152 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 586 (Holotype: ILL00007521!; isotypes: MEXU!, TEX (image)!, UCSB (fragment)!).</p> <p>Quercus cupreata f. brachystachys C.H.Mull., J. Arnold Arbor. 17(3): 176 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 1327 (Holotype: A00055577!; isotypes: MEXU!, NY!, P!, UCSB (fragment)!).</p> <p>Quercus cupreata f. serrata Trel. &amp; C.H.Mull., Bull. Torrey Bot. Club 63(3): 153 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 595 (Holotype: ILL00007519!; isotypes: MEXU!, TEX (image)!, UCSB (fragment)!).</p> <p>Quercus runcinatifolia Trel. &amp; C.H.Mull., Bull. Torrey Bot. Club 63(3): 153 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 591 (Holotype: ILL00007571!; isotype: UCSB (fragment)!).</p> <p>Quercus runcinatifolia f. lata Trel. &amp; C.H.Mull., Bull. Torrey Bot. Club 63(3): 153 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 593 (Holotype: ILL00007572!; isotypes: TEX (image)!, UCSB (fragment)!).</p> <p>Quercus shumardii Buckley var. microcarpa (Torr.) Shinners, Field &amp; Lab. 24(1): 37 (1956).</p> <p>Quercus stellapila (Sarg.) H.B. Parks ex Cory, Rhodora 38(455): 405 (1936).</p> <p>Quercus tardifolia C.H.Mull., Bull. Torrey Bot. Club 63(3): 154 (1936). Type:— U.S.A., Texas: C. H. Muller s. n. (3 Jul 1932) (Holotype: ILL00007575!; isotypes: CAS!, UCSB (fragment)!).</p> <p>Quercus texana Buckley var. c hesosensis Sarg., Bot. Gaz. 65(5): 423 (1918). Type:— U.S.A., Texas: G. Sudworth s. n. (5 Nov 1913) (Holotype: A00034105!). The protologue says 15 November but the date on the specimens is 5 November. This is also the type of Q. gravesii.</p> <p>Quercus texana var. stellapila Sarg., Bot. Gaz. 65(5): 424 (1918). Type:— U.S.A., Texas: D. M. Andrews 74 (Lectotype (designated here): A00034106!; isolectotype: A!). There are two specimens of Andrew 74 at A marked as the type by Sargent.</p> <p>Distribution: Mexico (Chihuahua, Coahuila, Colima, Jalisco, Nuevo León, Tamaulipas), U.S.A. (Texas). The occurrence of Q. gravesii in Colima reported by Villaseñor (2016) needs to be confirmed.</p> </div>	https://treatment.plazi.org/id/794B8792FFEEFFC93E88FF67FBA8F8D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FF97FFB63E88FF66FC92F7F1.text	794B8792FF97FFB63E88FF66FC92F7F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus oleoides Schltdl. & Cham.	<div><p>Quercus oleoides Schltdl. &amp; Cham.</p> <p>Quercus oleoides var. australis Trel. = Quercus oleoides Schltdl. &amp; Cham.</p> <p>Quercus oleoides f. lutescens (M.Martens &amp; Galeotti) Trel. = Quercus oleoides Schltdl. &amp; Cham.</p> <p>Quercus oleoides var. lutescens (M. Martens &amp; Galeotti) A.Camus = Quercus oleoides Schltdl. &amp; Cham.</p> <p>Quercus oleoides subsp. sagrana (Nutt.) Borhdi = Quercus sagrana Nutt.</p> <p>Quercus oleoides var. sagrana (Nutt.) C.H.Mull. = Quercus sagrana Nutt.</p> <p>Quercus oligodonta Seemen (1900) non Saporta (1879) = Quercus eduardi Trel.</p> <p>Quercus omissa A.DC. = Quercus sideroxyla Bonpl.</p> <p>Quercus oocarpa Liebm. = Quercus insignis M.Martens &amp; Galeotti</p> <p>Quercus opaca Trel. = Quercus pringlei Seemen</p> <p>Quercus orbiculata Trel. = Quercus crassifolia Bonpl.</p> <p>Quercus orizabae Liebm. = Quercus laurina Bonpl.</p> <p>Quercus ovandensis Matuda = Quercus lancifolia Schltdl. &amp; Cham.</p> <p>Quercus pablillensis C.H.Mull. = Quercus mexicana Bonpl.</p> <p>Quercus pacayana C.H.Mull. = Quercus bumelioides Liebm.</p> <p>Quercus pallescens Trel. = Quercus laeta Liebm.</p> <p>Quercus pallidifolia C.H.Mull. = Quercus subspathulata Trel.</p> <p>Quercus palmeri (Engelm.) Engelm.</p> <p>Quercus panamandinaea C.H.Mull. = Quercus benthamii A.DC.</p> <p>Quercus pandurata Bonpl. = Quercus obtusata Bonpl.</p> <p>Quercus pandurata var. hartwegii (Benth.) Wenz. = Quercus spicata Bonpl.</p> <p>Quercus pandurata var. laeta (Liebm.) Wenz. = Quercus laeta Liebm.</p> <p>Quercus panduriformis Trel. = Quercus obtusata Bonpl.</p> <p>Quercus panduriformis f. colimensis Trel. = Quercus alveolata Trel.</p> <p>Quercus panduriformis f. rubrinervis Trel. = Quercus obtusata Bonpl.</p> <p>Quercus pannosifolia C.H.Mull. = Quercus skutchii Trel.</p> <p>Quercus parviglans Trel. = Quercus sapotifolia Liebm.</p> <p>Quercus parviglans f. polycarpa Trel. = Quercus sapotifolia Liebm.</p> <p>Quercus parviglans f. tejadana Trel. = Quercus sapotifolia Liebm.</p> <p>Quercus × pastorensis C.H.Mull.</p> <p>Quercus paxtalensis C.H.Mull. = Quercus cortesii Liebm.</p> <p>Quercus peduncularis Neé</p> <p>Quercus peduncularis subsp. callosa (Benth.) A.Camus = Quercus peduncularis Neé</p> <p>Quercus peduncularis subsp. hurteri (Trel.) A.Camus = Quercus rugosa Neé</p> <p>Quercus peduncularis subsp. pedicularis var. callosa (Benth.) A.Camus = Quercus peduncularis Neé</p> <p>Quercus peduncularis subsp. pilicaulis (Trel.) A.Camus = Quercus peduncularis Neé</p> <p>Quercus peduncularis subsp. pilicaulis var. macrodonta (Trel.) A.Camus = Quercus peduncularis Neé</p> <p>Quercus peduncularis subsp. pilicaulis var. obovalis (Trel.) A.Camus = Quercus peduncularis Neé</p> <p>Quercus peduncularis var. sublanosa (Trel.) C.H.Mull. = Quercus segoviensis Liebm.</p> <p>Quercus peninsularis Trel.</p> <p>Quercus pennivenia Trel. = Q. urbani Trel.</p> <p>Quercus peradifolia E.F.Warb. = Quercus elliptica Neé</p> <p>Quercus perpallida Trel. = Quercus oblongifolia Torr.</p> <p>Quercus perseifolia Liebm.</p> <p>Quercus perseifolia var. achoteana Trel. = Quercus sapotifolia Liebm.</p> <p>Quercus perseifolia f. microcarpa Trel. = Quercus perseifolia Liebm.</p> <p>Quercus pertessellata Trel. = Billia hippocastanum Peyr. (see Appendix 1, Excluded Names)</p> <p>Quercus petiolaris Benth. = Quercus polymorpha Schltdl. &amp; Cham.</p> <p>Quercus petiolata (Trel.) A.E.Murray (1983) non Schur (1857) = Quercus salicifolia Neé</p> <p>Quercus pilarius Trel. = Quercus lancifolia Schltdl. &amp; Cham.</p> <p>Quercus pilgeriana Seemen = Quercus lancifolia Schltdl. &amp; Cham.</p> <p>Quercus pilicaulis Trel. = Quercus peduncularis Neé</p> <p>Quercus pilicaulis f. armata Trel. = Quercus rugosa Neé</p> <p>Quercus pilicaulis f. concava C.H.Mull. = Quercus rugosa Neé</p></div> 	https://treatment.plazi.org/id/794B8792FF97FFB63E88FF66FC92F7F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FFFCFFDA3E88F8EEFEE6F7F1.text	794B8792FFFCFFDA3E88F8EEFEE6F7F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus palmeri (Engelm.) Engelm., Bot.	<div><p>Quercus palmeri (Engelm.) Engelm., Bot. California 2: 97 (1880).</p> <p>Quercus chrysolepis Liebm. subsp. palmeri Engelm., Trans. Acad. Sci. St. Louis 3: 393. (1877). Type:— U.S.A., California: E. Palmer 30 (Holotype: MO-204133!; isotypes: A!, GH!).</p> <p>Quercus dunnii Kellogg, Pacific Rural Press 17(23): 371 (1879). This is an invalid name because Kellogg cited it as a provisional name.</p> <p>Distribution: Mexico (Baja California, Baja California Sur, Chihuahua, Sonora), U.S.A. (Arizona, California).</p> <p>Quercus peduncularis Neé, Anales Ci. Nat. 3: 270 (1801). Type:— MEXICO, Guerrero: L. Neé s. n. (1791) (Lectotype: MA26474! designated by Valencia-A. &amp; Coombes (2020, p. 87); isolectotypes: MA!, UCSB (fragment p.p. as to smaller leaf from MA lectotype)!, UCSB (fragment)!).</p> <p>Quercus affinis M.Martens &amp; Galeotti non Scheidw. (1837), Bull. Acad. Roy. Sci. Bruxelles 10(1): 222 (1843). See Q. martensiana below.</p> <p>Quercus barbanthera Trel., Mem. Natl. Acad. Sci. 20: 68, pl. 81 (1924). Type:— GUATEMALA: G. V. Skinner s. n. (1845) (Holotype at B destroyed. Lectotype (designated here): K000648587!). Valencia-A. &amp; Coombes (2020, p. 87) invalidly lectotypified the name based on Trelease’s image of the holotype at B which was destroyed. A paratype exists at K and according to Article 9.12 of the Code “If no isotype, syntype or isosyntype is extant, the lectotype must be chosen from among the paratypes if such exist” (Turland et al. 2018). The epitype, Miranda 6429 (MEXU10381 as image!) which they selected is also invalid, because of the existence of the paratype which is now the lectotype.</p> <p>Quercus barbanthera var. calva Trel., Mem. Natl. Acad. Sci. 20: 68 (1924). Type:— GUATEMALA, Chimaltenango: E. T. Heyde &amp; E. Lux 3152 (Lectotype (designated here): K000512621!; isolectotypes: NY!, US!). Trelease (1924. p. 68) cited two different collections in the protologue.</p> <p>Quercus callosa Benth., Pl. Hartw. 91 (1842). Type:— GUATEMALA, Guatemala: T. Hartweg 616 (Lectotype: K000512622! designated by Trelease (1824, p. 218, pl. 82a); isolectotypes: CGE!, K!, LD (image)!, NY!). There are two sheets of Hartweg 616 in the Bentham Herbarium at K, one of which was effectively lectotypified by Trelease.</p> <p>Quercus comasaguana Trel., Mem. Natl. Acad. Sci. 20: 43 (1924). Type:— EL SALVADOR, La Libertad: S. Calderon 1430 (Lectotype (designated here): ILL00007525!; isolectotypes: GH!, US!). There is no indication in the protologue as to where the holotype was deposited, and both the ILL and US specimens are labeled as a type.</p> <p>Quercus dolichopus E.F.Warb., Bull. Misc. Inform. Kew 1939 (2): 87 (1939). Type:— MEXICO, México: G. B. Hinton 6378 (Lectotype: K000575005! designated by Valencia-A. &amp; Coombes (2020, p. 87); isolectotypes: BM!, F!, G!, GH!, K!, MEXU!, MO!, NY!, UCSB (fragment)!, US!).</p> <p>Quercus liebmannii Oerst. ex Trel. f. brevipes Trel., Mem. Natl. Acad. Sci. 20: 67, pl. 75b (1924). Type:— MEXICO, Oaxaca: F. M. Liebmann 3526 (Holotype: C not located; isotypes: C!, CAS!). The holotype of Q. liebmannii f. brevipes at C is missing although isotypes exist at C and CAS. A photo of part of the holotype is in Trelease (1924) pl. 75b which is Liebmann 3526 (Cupulif. Lbm. 136); hopefully the specimen with the correct Cupulif. Lbm. number and matching the photo will eventually be found. The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. peduncularis as broadly interpreted.</p> <p>Quercus martensiana Trel., Contr. U.S. Natl. Herb. 23(2): 182 (1922). Type:— MEXICO, Veracruz: H. Galeotti 90 (Holotype: BR0000009816213!; isotypes: G!, K!, NY!, P!, UCSB (fragment)!). Nom. nov. for Q. affinis M. Martens &amp; Galeotti (1843) non Scheidw. (1837).</p> <p>Quercus martensiana f. perplexans Trel., Mem. Natl. Acad. Sci. 20: 60 (1924). Type:— MEXICO, Veracruz: F. M. Liebmann 3567 (Holotype: C 10012502!).</p> <p>Quercus peduncularis subsp. callosa (Benth.) A.Camus, Chênes Texte 2: 599 (1939).</p> <p>Quercus peduncularis subsp. pedicularis var. callosa (Benth.) A.Camus, Chênes Atlas 2: 91 (1935).</p> <p>Quercus peduncularis subsp. pilicaulis (Trel.) A.Camus, Chênes Atlas 2: 91 (1935).</p> <p>Quercus peduncularis subsp. pilicaulis var. macrodonta (Trel.) A.Camus, Chênes Atlas 2: 91 (1935).</p> <p>Quercus peduncularis subsp. pilicaulis var. obovalis (Trel.) A.Camus, Chênes Atlas 2: 91 (1935).</p> <p>Quercus pilicaulis Trel., Mem. Natl. Acad. Sci. 20: 67, 77b (1924). Nom. nov. for Q. tomentosa var. abbreviata A. DC (1864) non Q. abreviata Vuk. (1889).</p> <p>Quercus pilicaulis f. macrodonta Trel., Mem. Natl. Acad. Sci. 20: 68, pl. 78 (1924). Type:— GUATEMALA: J. R. R. Warszewicz 15 (Holotype at B destroyed. Lectotype (designated here): G00390238!; isolectotype; UCSB (fragment from G lectotype)!). Trelease (1924, p. 218) in the caption to pl. 78a designated Warscewicz 15 at B as the holotype of Q. pilicaulis f. macrodonta and treated Q. tomentosa var. bullata A.DC. as a synonym for which Warsczwicz 15 at G was a syntype and which he effectively lectotypified in the caption to pl. 78b. We are lectotypifying Q. plicaulis f. macrodonta based on the specimen of Warsczwicz 15 at G.</p> <p>Quercus pilicaulis f. obovalis Trel.,Mem.Natl.Acad.Sci. 20:68,pl.79b(1924). Type:— GUATEMALA, Chimaltenango: F. C. Lehmann 1725 (Holotype at B destroyed. Lectotype: UCSB000351 (fragment)! designated by Valencia-A. &amp; Coombes (2020, p. 87); isolectotypes: BM!, G). Lectotypification of this name was based on a fragment at UCSB rather than the isotype at G from which the fragment was reportedly taken or the other isotype at BM.</p> <p>Quercus schultzei Trel., Repert. Spec. Nov. Regni Veg. 33: 316 (1934). Type:— MEXICO, Guerrero: L. Schultze 115 (Holotype at B destroyed. Lectotype (designated here): ILL00007589! (fragment)).</p> <p>Quercus splendens Neé, Anales Ci. Nat. 3: 275 (1801). Type:— MEXICO, Guerrero: L. Neé s. n. (1791) (Lectotype: MA25972! designated by Valencia-A. &amp; Coombes (2020, p. 87); isolectotypes: MA!, UCSB (fragment)!).</p> <p>Quercus tomentosa Willd. nom. illeg., Sp. Pl., ed. 4(1): 437 (1805). This is a superfluous name because Willdenow cites Q. peduncularis Neé (1801), which is an older name, in synonomy.</p> <p>Quercus tomentosa Willd. nom. illeg. var. abbreviata A.DC., Prodr. 16(2.1): 33 (1864). Type:— GUATEMALA: J. R. R. Warszewicz 43 (Holotype: G00421035!; isotype: UCSB (fragment from G holotype)!). Although Valencia- A. &amp; Coombes (2020, p. 87) lectotypified this name, it was not necessary as A. de Candolle clearly specified the holotype.</p> <p>Quercus tomentosa Willd. nom. illeg. var. bullata A.DC., Prodr. 16(2.1): 33 (1864). Type:— GUATEMALA: J. R. R. Warszewicz 15 (Lectotype: G00390238! designated by Trelease (1924, p. 218, pl.78b); isolectotype: UCSB (fragment from the G lectotype)!). A. de Candolle (1864, p. 33) cited two syntypes.</p> <p>Quercus tomentosa Willd. nom. illeg. var. communis A.DC., Prodr. 16(2.1): 33 (1864). Type:— GUATEMALA: J. Warszewicz 17 (Lectotype: G00390235! designated by Valencia-A. &amp; Combes (2020, p. 87)). A. de Candolle (1864, p. 33) cites several syntypes.</p> <p>Distribution: El Salvador, Guatemala, Honduras, Mexico (Aguascalientes, Chiapas, Ciudad de México, Colima,</p> <p>Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, México, Michoacán, Morelos, Nayarit, Oaxaca, Puebla, Querétaro,</p> <p>San Luis Potosí, Sinaloa, Tamaulipas, Veracruz, Zacatecas).</p> <p>Quercus peninsularis Trel., Mem. Natl. Acad. Sci. 20: 124, pl. 225a (1924). Type:— MEXICO, Baja California: T. Brandegee s. n. (23 May 1893) (Holotype: A00062253!; isotypes: UCSB (fragment)!, US).</p> <p>Distribution: Mexico (Baja California).</p> <p>Quercus perseifolia Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 188 (1854). Type:— MEXICO, Veracruz: F. M. Liebmann 3476 (Lectotype: CAS0006172! designated by Trelease (1924, p. 225, pl. 292a); isolectotypes: C!, UCSB (fragment from the CAS lectotype)!). The lectotype is sheet Cupulif. Liebm. 98. Trelease (1924, p. 149) described the fruit as annual although most collections we have seen are biennial. It is known that some species of Quercus have both annual and biennial fruit, not only within the same species, but even within the same population (Pons &amp; Pausas 2012).</p> <p>Quercus perseifolia f. microcarpa Trel., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 188 (1854). Type:— MEXICO, Veracruz: H. Galeotti 112 (Lectotype (designated here): BR!). Trelease cited two specimens without designating which one was the type.</p> <p>Distribution: Mexico (Hidalgo, México, Michoacán, Nuevo León, Oaxaca, Puebla, San Luis Potosí, Tamaulipas,</p> <p>Veracruz).</p> <p>Quercus pinnativenulosa C.H.Mull., J. Arnold Arbor. 17(3): 171 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 1342 (Holotype: A00055690!; isotypes: CAS!, F!, MEXU!, TEX (image)!). This species may be the same as Q. perseifolia in the broad sense. Additional study is needed.</p> <p>Distribution: Mexico (Coahuila, Hidalgo, Nuevo León, Oaxaca, Puebla, Qurerétaro, San Luis Potosí, Tamaulipas,</p> <p>Veracruz).</p> <p>Quercus planipocula Trel., Mem. Natl. Acad. Sci. 20: 136, pl. 259a (1924). Type:— MEXICO, Nayarit: J. N. Rose 1970 (Holotype: US00089613!; isotypes: GH!, MO!, NY!, UCSB (fragment)!).</p> <p>Distribution: Mexico (Colima, Guerrero, Jalisco, México, Michoacán, Nayarit, Oaxaca, San Luis Potosí, Sinaloa,</p> <p>Tamaulipas). The occurrence of Q. planipocula in Oaxaca, San Luis Potosí, and Tamaulipas reported by Villaseñor</p> <p>(2016) needs to be confirmed.</p> <p>Quercus polymorpha Schltdl. &amp; Cham., Linnaea 5(1): 78 (1830). Type:— MEXICO, Veracruz: C. J. W. Schiede 20 (Holotype at B destroyed. Lectotype (designated here): HAL0060344!; isolectotypes: BR!, G!).</p> <p>Quercus germana Schltdl. &amp; Cham. var. lemmonii Trel., Mem. Natl. Acad. Sci. 20: 52 (1924). Type:— MEXICO, Veracruz: J. G. Lemmon &amp; J. G. Lemmon 103 (Holotype: US 00089539!; isotypes: E!, F!, GH!, MO!, NY!, UC!).</p> <p>Quercus guatimalensis A.DC, Prodr. 16(2.1): 78 (1864). Type:— GUATEMALA: J. R. R. Warszewicz 37 (Holotype at B destroyed. Lectotype (designated here): P00754147!). Nom. nov. for Q. turbinata Liebm. (1854) non Blume (1825).</p> <p>Quercus juergensenii Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 188 (1854). Type:— MEXICO, Puebla: F. M. Liebmann s. n. (Holotype: C (missing); isotype: UCSB (fragment from C holotype)!). Although the foliage is described by Liebmann, the only remaining specimens at C when Trelease visited there were acorns. These acorns are now missing at C and the only extant ones are a few now at UCSB. Trelease also considered this be a problematic species but perhaps synonymous with Q. polymorpha, which is a name that has priority.</p> <p>Quercus petiolaris Benth., Pl. Hartw. 55 (1840). Type:— MEXICO, Jalisco: T. Hartweg 420 (Holotype: K000648582!; isotypes: A!, BM!, CAS!, CGE!, E!, FI!, G!, LD (image)!, LE!, NY!, P!, UCSB (fragment)!).</p> <p>Quercus polymorpha f. angustifolia C.H.Mull., J. Arnold Arbor. 17(3): 161 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 1144 (Holotype: A00283982!).</p> <p>Quercus turbinata Liebm. non Blume (1825), Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 186 (1854). See Q. guatimalensis above.</p> <p>Quercus varians M.Martens &amp; Galeotti, Bull.Acad. Roy. Sci. Bruxelles 10(1): 214 (1843). Type:— MEXICO, Veracruz: H. Galeotti 92 (Lectotype: BR0000006916251! designated by Trelease (1924, p. 217, pl. 37); isolectotypes: BR!, G!, K!, L!, LE!, P!, UCSB (fragment)!). M. Martens &amp; Galeotti (1843, p. 214) cited only one collection number, four sheets of which are at BR; one of these was designated by Trelease as “The type.”</p> <p>Distribution: Guatemala, Honduras, Mexico (Chiapas, Coahuila, Guanajuato, Hidalgo, Jalisco, México, Michoacán,</p> <p>Morelos, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa, Tamaulipas, Veracruz).</p> <p>Quercus porphyrogenita Trel., Mem. Natl. Acad. Sci. 20: 51, pl. 39a (1924). Type:— MEXICO, Nuevo León: W. Trelease 117 (Holotype: MO-144179!; isotype: ILL!). The distinction between Q. porphyrogenita and Q. laceyi needs additional study.</p> <p>Quercus microlepis Trel. &amp; C.H.Mull. non Vuk. (1883), Bull. Torrey Bot. Club 63(3): 150 (1936). Type:— MEXICO, Nuevo León: C.H. Muller 578 (Holotype: ILL00007500!; isotypes: MEXU (a image!), UCSB (fragment)!).</p> <p>Distribution: Mexico (Nuevo León).</p> <p>Quercus potosina Trel., Mem. Natl. Acad. Sci. 20: 84, pl. 123 (1924). Type:— MEXICO, San Luis Potosí: C. C. Parry &amp; E. Palmer 838 (Holotype: MO-143894!; isotypes: BM!, GH!, K!, MO!, NY!, P!, PH!, UCSB (fragment)!, US!, WIS!).</p> <p>Quercus jaralensis Trel., Mem. Natl. Acad. Sci. 20: 86, pl. 133 (1924). Type:— MEXICO, Guanajuato: W. Schumann 1316 (Holotype at B destroyed. Lectotype (designated here): P00754117!; isolectotype: P!).</p> <p>Quercus potosina f. aperta Trel., Mem. Natl. Acad. Sci. 20: 84, pl. 125a (1924). Type:— MEXICO, San Luis Potosí: C. C. Parry &amp; E. Palmer 838b (Holotype: MO-143893!).</p> <p>Quercus potosina f. exilis Trel., Mem. Natl. Acad. Sci. 20: 84, pl. 124a (1924). Type:— MEXICO, San Luis Potosí: C. C. Parry &amp; E. Palmer 838 (Holotype: MO-143892!; isotypes: BM!, GH!, K!, MO!, NY!, P!, PH!, UCSB (fragment)!, US!, WIS!).</p> <p>Distribution: Mexico (Aguascalientes, Chihuahua, Coahuila, Colima, Durango, Guanajuato, Hidalgo, Jalisco, México,</p> <p>Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Tamaulipas, Veracruz, Zacatecas). The occurrence of Q.</p> <p>potosina in México and Veracruz reported by Villaseñor (2016) needs to be confirmed.</p> <p>Quercus praeco Trel., Mem. Natl. Acad. Sci. 20: 88, pl. 139 (1924). Type:— MEXICO, Jalisco: J. N. Rose 2590 (Holotype: US00089611!; isotypes: GH!, NY!, UCSB (fragment)!). This species is close to Q. xylina, and additional study is needed to determine if it is distinct.</p> <p>Distribution: Mexico (Jalisco, Nayarit, Zacatecas).</p> <p>Quercus pringlei Seemen, Bot. Jahrb. Syst. 29(1): 96 (1900). Type:— MEXICO, Coahuila: C. G. Pringle 2382 (Holotype at B destroyed. Lectotype (designated here): US00089609!; isolectotypes: A!, AC!, BM!, BR!, CAS!, COLO (image)!, E!, F!, G!, GH!, K!, KFTA (image)!, LL (image)!, MEL (image)!, MEXU!, MO!, NDG (image)!, NY!, P!, PH!, PUL (image)!, UC!, UCSB (fragment)!, US!, VT!, YU (image)!). Trelease (1924, p. 124) in the caption to pl. 148a designated Pringle 2382 at B as “The type” which was destroyed.</p> <p>Quercus filiformis C.H.Mull., Amer. Midl. Naturalist 27(2): 473 (1942). Type:— MEXICO, Coahuila: C. H. Muller 3150 (Holotype: US00089513!; isotypes: GH!, UC!, UCSB (fragment)!). This name was also used in 1818 as a nom. nud. and as nom. inval. as a synonym of Q. bicolor var. platanoides.</p> <p>Quercus opaca Trel., Mem. Natl. Acad. Sci. 20: 92, pl. 148b (1924). Type:— MEXICO, Hidalgo: J. N. Rose 9030 (Holotype: US00089571!; isotypes: GH!, NY!, UCSB (fragment)!).</p> <p>Quercus pringlei f. dentata C.H.Mull., J. Arnold Arbor. 17(3): 168. (1936). Type:— MEXICO, Nuevo León: C. H. Muller 881 (Holotype: A00969953!).</p> <p>Distribution: Mexico (Chihuahua, Coahuila, Durango, Hidalgo, Nuevo León, Querétaro, San Luis Potosí, Tamaulipas,</p> <p>Veracruz, Zacatecas).</p> <p>Quercus pungens Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 171 (1854). Type:— U.S.A., Texas: C. Wright 664 (Lectotype: NY00248762 (image)! designated by Trelease (1924, p. 221, pl. 157a); isolectotypes: BM!, GH!, K (image)!, NY!, P!, UCSB (fragment)!, US!, W!). The Hooker Herbarium collection of Wright 664 at K, which was a syntype of this species, was sent to NY, and this is the specimen Trelease effectively lectotypified. The only specimen of Wright 664 currently at K is the Bentham Herbarium specimen which becomes a lectotype. The other syntype is Lobb 106 from the Hooker Herbarium at K (K000648585!).</p> <p>Quercus pungens var. vaseyana (Buckley) C.H.Mull., Cont. Texas Res. Found., Bot. Stud. 1:70 (1951).</p> <p>Quercus undulata Torr. var. pungens (Liebm.) Engelm., Trans. Acad. Sci. St. Louis 3: 392 (1877).</p> <p>Quercus undulata var. vaseyana (Buckley) Rydb., Bull. New York Bot. Gard. 2(6): 218 (1901).</p> <p>Quercus vaseyana Buckley, Bull. Torrey Bot. Club 10(8): 91 (1883). Type:— U.S.A., Texas: S. B. Buckley s. n. (Lectotype: NY00253631! designated by Trelease (1924, p. 221, pl. 156a) and redundantly relectotypified by Dorr &amp; Nixon (1985, p. 223); isolectotype: NY!).</p> <p>Distribution: Mexico (Chihuahua, Coahuila, Durango, Guanajuato, Hidalgo, Nuevo León, Querétaro, San Luis Potosí,</p> <p>Sonora, Tamaulipas, Veracruz), U.S.A. (Arizona, New Mexico, Texas).</p> <p>Quercus purulhana Trel., Mem. Natl.Acad. Sci. 20: 63, pl. 64 (1924). Type:— GUATEMALA, Baja Verapaz: H. Pittier 163 (Holotype: US 00089604!; isotypes: NY!, UCSB (fragment)!, US!).</p> <p>Quercus aguana Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 9(4): 281 (1940). Type:— HONDURAS, Lempira: T. G. Yuncker et al. 8180 (Holotype: ILL00007476!; isotypes: BM!, F!, G!, GH!, K!, MICH!, MO!, NY!, S (image)!, US!).</p> <p>Quercus barbeyana Trel., Mem. Natl. Acad. Sci. 20: 67, pl. 76 (1924). Type:— GUATEMALA, Chiquimula: F. C. Lehmann 1711 (Holotype: G00358171!; isotype: BM!).</p> <p>Distribution: Belize, Guatemala, Honduras, Mexico (Chiapas), Nicaragua.</p> <p>Quercus radiata Trel., Proc. Amer. Philos. Soc. 60: 33, pl. 3 (1921). Type:— MEXICO, Nayarit: J. N. Rose 2230 (Holotype: US 00089601!; isotypes: GH!, NY!).</p> <p>Distribution: Mexico (Durango, Jalisco, Nayarit, Zacatecas).</p> <p>Quercus rekonis Trel., Mem. Natl. Acad. Sci. 20: 60, pl. 57 (1924). Type:— MEXICO, Oaxaca: C. Conzatti et al. 3121 (Holotype: US00089631!; isotypes: MEXU!, MO!). The collectors of the type were C. Conzatti, B.P. Reko &amp; Makrinius, but only the the name B.P. Reko is on the US holotype.</p> <p>Distribution: Mexico (Chihuahua, Durango, Guerrero, Jalisco, México, Michoacán, Nayarit, Oaxaca, Sinaloa).</p> <p>Quercus resinosa Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 182 (1854). Type:— MEXICO: B. Seemann 1972 (Holotype: K000512615!; isotypes: GH!, K!, UCSB (fragment)!). There are two specimens of Seemann 1972 in the Hooker Herbarium at K, but only one of them has a determination in Liebmann’s hand which we consider to be the holotype.</p> <p>Distribution: Mexico (Aguascalientes, Colima, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, México, Michoacán,</p> <p>Nayarit, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa, Sonora, Tamaulipas, Zacatecas). The occurrence of Q.</p> <p>resinosa in Guerrero and Hidalgo reported by Villaseñor (2016) needs to be confirmed.</p> <p>Quercus rugosa Neé, Anales Ci. Nat. 3: 275 (1801). Type:— MEXICO, Guerrero: L. Neé s. n. (1791) (Lectotype: MA26471! designated by Bartholomew &amp; Almeda (2022, p. 29); isolectotypes: MA!, UCSB (fragment from MA lectotype)!, UCSB (fragment)!).</p> <p>Quercus conglomerata Trel. non Pers. (1807), Mem. Natl. Acad. Sci. 20: 77, pl. 106a (1924). Type:— MEXICO, Michoacán: T. Hartweg 429 (Holotype: B destroyed; isotypes: BM!, E!, G!, K!, L!, NY!, P!, UCSB (fragment)!). Because Q. conglomerata Trel. is a later homonym, we are not lectotypifying the name.</p> <p>Quercus decipiens M.Martens &amp; Galeotti, Bull. Acad. Roy. Sci. Bruxelles 10(1): 214 (1843). Type:— MEXICO, Veracruz: H. Galeotti 131 (p.p.) (Lectotype: BR0000006914295! designated by Trelease (1924, p. 219, pl. 107); isolectotypes: BR!, G!, P!, UCSB (fragment)!). M.Martens &amp; Galeotti cite only one collection, Galeotti 131, four sheets of which are at BR. Two of them are types of Q. decipiens, one (BR000006915612) is a mixed sheet in which only the upper left specimen on the sheet is a type of Q. decipiens, and the other one (BR0000006913953) is not a type of Q. decipiens. The lectotype is the sheet designated by Trelease as “The type.”</p> <p>Quercus diversicolor Trel., Mem. Natl. Acad. Sci. 20: 73, pl. 92a (1924). Type:— MEXICO, Chihuahua: C. G. Pringle 2021 (Holotype at B destroyed. Lectotype (designated here): US 01013683!; isolectotypes: BM!, BR!, CAS!, G!, GH!, K!, M!, MEL (image) !, MO!, NDG (image) !, NY!, P!, PH!, UC!, UCSB (fragment) !, US!, VT !).</p> <p>Quercus diversicolor var. mearnsii Trel., Mem. Natl. Acad. Sci. 20: 74, pl. 94 (1924). Type:— U.S.A., Arizona: E. A. Mearns 2230 (Holotype: NY01465003!; isotype: US!).</p> <p>Quercus diversicolor var. socorronis Trel., Mem. Natl. Acad. Sci. 20: 74 (1924). Type:— U.S.A., New Mexico: O. B. Metcalfe 669 (Lectotype (designated here): US 00089497!; isolectotypes: NDG (image)!, UCSB (fragment)!). Trelease (1924, p. 74) did not indicate where the type collections were deposited.</p> <p>Quercus durangensis Trel., Mem. Natl. Acad. Sci. 20: 73, pl. 91a (1924). Type:— MEXICO, Durango: Mathews s. n. (Nov 1904) (Holotype: MO-143914!; isotype: UCSB (fragment from MO holotype)!).</p> <p>Quercus macrophylla Neé var. rugosa (Neé) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 196 (1884).</p> <p>Quercus peduncularis Neé subsp. hurteri (Trel.) A.Camus, Chênes Atlas 2: 92 (1935).</p> <p>Quercus pilicaulis Trel. f. armata Trel., Mem. Natl. Acad. Sci. 20: 68, pl. 79b (1924). Type:— GUATEMALA, Quezaltenango: E. W. Nelson 3721 (Holotype: US 00089588!).</p> <p>Quercus pilicaulis f. concava C.H.Mull., Amer. Midl. Naturalist 18(5): 851 (1937). Type:— GUATEMALA, Chimaltenango: A. F. Skutch 685 (Holotype: US 00089587!; isotypes: A!, F!, ILL!, UC!).</p> <p>Quercus pilicaulis f. elongata C.H.Mull., Amer. Midl. Naturalist 18(5): 851 (1937). Type:— GUATEMALA, Chimaltenango: A. F. Skutch 687 (Holotype: US00089586!; isotypes: A!, BR!, DS!, F!, ILL!, K!, UC!).</p> <p>Quercus pilicaulis f. exserta C.H.Mull., Amer. Midl. Naturalist 18(5): 852 (1937). Type:— GUATEMALA, Chimaltenango: A. F. Skutch 661 (Holotype: US 00089585!; isotypes: A!, UC!).</p> <p>Quercus pilicaulis f. hurteri Trel., Mem. Natl.Acad. Sci. 20: 68, pl. 80 (1924). Type:— GUATEMALA, Quezaltenango: W. Trelease 37 (Holotype: ILL00007598!).</p> <p>Quercus purpusii Trel., Mem. Natl. Acad. Sci. 20: 76, pl. 105 (1924). Type:— MEXICO, México: C. A. Purpus 1794 (Holotype: MO-143916!; isotypes: F!, GH!, NY!, UC!, UCSB (fragment)!, US!).</p> <p>Quercus reticulata Bonpl., Pl. Aequinoct. 2: 40, pl. 86 (1809). Type:— MEXICO, Guanajuato: A. Bonpland s. n. (Lectotype (designated here): P00129698!; isolectotypes: P!, UCSB (fragment)!). There are four Bonpland s. n. collections from Santa Rosa, Guanajuato, Mexico which are syntypes of Q. reticulata. The lectotype has the name “ Quercus coccolobaefolia sp. n. ” in Bonpland’s hand. Bonpland, however, used the name Q. reticulata in his publication. The syntypes of Q. reticulata are not the same collection as Bonpland 4269 which is the collection number for the type of Q. coccolobifolia Trel.</p> <p>Quercus reticulata f. longa Trel., Mem. Natl.Acad. Sci. 20: 75, pl. 99a (1924). Type:— MEXICO, Hidalgo: C. Ehrenberg 897 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 99a)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Quercus reticulata subsp. reticulata var. conglomerata (Trel.) A.Camus, Chênes Atlas 2: 93 (1935).</p> <p>Quercus reticulata subsp. reticulata var. squarrosa (Trel.) A.Camus, Chênes Atlas 2: 94 (1935).</p> <p>Quercus reticulata subsp. rhodophlebia (Trel.) A.Camus, Chênes Atlas 2: 94 (1935).</p> <p>Quercus reticulata subsp. rhodophlebia f. concava (Trel.) A.Camus, Chênes Texte 2: 581 (1939).</p> <p>Quercus reticulata f. squarrosa Trel., Mem. Natl. Acad. Sci. 20: 75, pl. 99b (1924). Type:— MEXICO, Hidalgo: G. J. Graham 328 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 99b)!). Trelease (1924. p. 219) wrote in the caption to pl. 99b that the type was an Ehrenberg collection at B. He cited five of Ehrenberg’s collections earlier on p. 75. Because they were all destroyed, it is impossible to know which collection number (903, 904, 905, 906, or 908) was the holotype. The image in pl. 99b is probably of a collection made in Hidalgo, Mexico, because all the Ehrenberg collections around these numbers are from Hidalgo. Trelease also cited two Graham collections, of which we have found only Graham 328 which is a different species, so we are not lectotypifying the name based on either of the paratypes but rather on the image of the holotype in plate 99b.</p> <p>Quercus rhodophlebia Trel., Mem. Natl. Acad. Sci. 20: 74, pl. 95a (1924). Type:— MEXICO, Zacatecas: J. N. Rose 2806 (Holotype: US 00089626!).</p> <p>Quercus rhodophlebia f. concava Trel., Mem. Natl. Acad. Sci. 20: 74, pl. 95b (1924). Type:— MEXICO, Jalisco: J. N. Rose 2998 (Holotype: US00089623!; isotypes: GH!, MO!, NY!, UCSB (fragment)!).</p> <p>Quercus sideroxyla Bonpl. f. aquifolia Trel., Mem. Natl. Acad. Sci. 20: 170 (1924). Type:— MEXICO, Guanajuato: A. Dugès 4 (Holotype: GH00969956!). The specimen of Dugès 4 at US is Q. tridens so it cannot be considered an isotype.</p> <p>Quercus suchiensis Warb., J. Roy. Hort. Soc. 58(1): 187, fig. 55 (1933). Type:— MEXICO, México / Michoacán: R. Heatley s. n. (Holotype: K000575026! and K000575027!). The holotype was originally mounted on one sheet but now it is mounted on two sheets. The type collection is “on the boundaries of the states of Mexico and Michoacan.”</p> <p>Quercus uhdeana Trel., Mem. Natl. Acad. Sci. 20: 75, pl. 100 (1924). Type:— MEXICO: Uhde 276 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 100)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Distribution: Guatemala, Honduras, Mexico (Aguascalientes, Baja California, Baja California Sur, Chiapas, Chihuahua,</p> <p>Ciudad de México, Coahuila, Colima, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, México, Michoacán, Morelos,</p> <p>Nayarit, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa, Sonora, Tamaulipas, Tlaxcala, Veracruz,</p> <p>Zacatecas), U.S.A. (Arizona, New Mexico, Texas).</p> <p>Quercus rysophylla Weath., Proc. Amer. Acad. Arts 45: 423 (1910). Type:— MEXICO, Nuevo León: C. G. Pringle 10225 (Lectotype: GH00069922! designated by Trelease (1924, p. 225, pl. 279a); isolectotypes: ASU (image)!, CAS!, DAO (image)!, DUKE (image)!, ENCB (image)!, F!, IBUG (image)!, LE!, LL (image)!, MICH!, MSC (image)!, RSA (image)!, UC!, UCR (image)!, UCSB (fragment)!, US!, VT!, WIS!, YU (image)!).</p> <p>Quercus tenuiloba f. hirsuta C.H.Mull., J.Arnold Arbor. 17(3): 178 (1936). Type:— MEXICO, Nuevo León: C. H. Muller 2122 (Holotype: A00055647!; isotypes: MEXU!, MO!, P!, TEX (image)!, UCSB (fragment)!, US!). Although the species epithet is based on a probable hybrid collection (see Appendix 2), the type of this forma is Q. rysophylla.</p> <p>Distribution: Mexico (Coahuila, Hidalgo, Nuevo León, Oaxaca, Querétaro, San Luis Potosí, Tamaulipas, Veracruz).</p> <p>Quercus sagrana Nutt., N. Amer. Sylv. 1(1): 17 (1842). Type:— CUBA, Pinal del Rio: R. de la Sagra 418 (Holotype: PH00023400!; isotypes: G!, P (as s.n.)!, UCSB (as s.n.) (fragment)!).</p> <p>Quercus cubana A.Rich., Hist. Fis. Cuba 11: 230 (1850). This is an illegitimate name because it is based on an isotype of Q. sagrana Nutt., R. de la Sagra [J. M. Valenzuela] s. n., which is at P as P00754080!. J.M. Valenzuela was a collector for R.de la Sagra and all of the type specimens of both Q. sagrana and Q. cubana were almost certainly collected by him. Richard (1850) cited J.M.Valenzuela as the collector of Q. cubana, but Valenzuela’s name is not on the any of the specimens distributed by de la Sagra.</p> <p>Quercus oleoides Schltdl. &amp; Cham. var. sagrana (Nutt.) C.H.Mull., Amer. Midl. Naturalist 65(1): 37. (1961).</p> <p>Quercus oleoides subsp. sagrana (Nutt.) Borhidi, Acta Bot. Acad. Sci. Hung. 17: 6 (1971[1972]).</p> <p>Quercus virginiana Mill. var. sagrana (Nutt.) Trel., Mem. Natl. Acad. Sci. 20: 113, pl. 189b (1924).</p> <p>Distribution: Cuba.</p> <p>Quercus salicifolia Neé, Anales Ci. Nat. 3: 265 (1801). Type:— MEXICO, Guerrero: L. Neé s. n. (1791) (Lectotype: MA25957! designated by Muller &amp; McVaugh (1972, p. 520); isolectotypes: MA!, UCSB (fragment)!). We are treating Q. salicifolia in a broad sense to include what could be recognized as many separate species if a narrower species concept were adopted. Further study of this species complex is needed.</p> <p>Quercus acapulcensis Trel., Mem. Natl. Acad. Sci. 20: 153, pl. 302a (1924). Type:— MEXICO, Guerrero: E. Palmer 334 (Holotype: GH00069901!; isotypes: CAS!, DS!, F!, K!, NY!, UCSB (fragment)!, US!).</p> <p>Quercus boquetensis Standl., Publ. Field Mus. Nat. Hist., Bot. Ser. 22(1): 13 (1940). Type:— PANAMA, Chiriquí: M. E. Davidson 780 (Holotype: F!; isotypes: GH!, MO!, US!). Additional study is needed to determine if Q. boquetensis might actually be a good species.</p> <p>Quercus borucasana Trel., Mem. Natl.Acad. Sci. 20: 161, pl. 315a (1924). Type:— COSTA RICA, Cartago: A. S. Oersted 3491 (Holotype: C10014102!). Nom. nov. for Q. granulata Liebm. (1854) non Raf. (1838).</p> <p>Quercus citrifolia Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 187 (1854). Type:— COSTA RICA, Cartago: A. S. Oersted 3461 (Lectotype (designated here): C 10012469!; isolectotypes: C!, CAS!, G!, UCSB (fragment)!). Trelease (1924) in pl. 288 shows two different specimens, but in his caption for pl. 288 states “The type (Costa Rica, Oersted, 6=3461).” Specimen 6=3461 is the bottom image of pl. 288 and is being lectotypified here, and 8=3461, the top image, which is now at CAS, becomes an isolectotype. A third specimen, 7=3461, is an isolectotype at C.</p> <p>Quercus eugeniifolia Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 185 (1854). Type:— COSTA RICA: J. R. R. Warszewicz s. n. (C) (Lectotype (designated here): image in Trelease (1924, pl. 316a)!). Liebmann (1854, p. 185) wrote “Guatimala [sic] et Costa rica [sic] leg. Warsewicz [sic] (in hb. reg. Berol.!) Species affinis Q. salicifoliae Née. ” Trelease (1924, p. 161, pl. 316a) effectively lectotypified Warszewicz C at B. The photograph in pl. 316a is Q. salicifolia. The lectotype at B was destroyed, and because there are no extant specimens of Warszewicz at C, we are lectotypifying the name based on pl. 316a. Trelease cites Warszewicz 29 as also being Q. eugeniifolia, although without stating where he saw the specimens. If this specimen was at B it was destroyed, and the specimens of Warszewicz 29 at G and P are Q. acutifolia.</p> <p>Quercus eugeniifolia f. petiolata Trel., Mem. Natl. Acad. Sci. 20: 161, pl. 316b (1924). Type:— COSTA RICA: C. Hoffmann 863 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 316b)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Quercus granulata Liebm. non Raf. (1838), Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 186 (1854). See Q. borucasana above.</p> <p>Quercus mulleri Martínez, Anales Inst. Biol. Univ. Nac. Autón. México 24(2): 51, fig. 1, 2 (1953[1954]). Type:— MEXICO, Oaxaca: T. MacDougall 2435 (Holotype: MEXU00011238!; isotypes: CAS!, ENCB (image)!, UCSB (fragment)!).</p> <p>Quercus nixoniana S.Valencia &amp; Lozada-Pérez, Novon 13(2): 261, f. 1 (2003). Type:— Mexico, Guerrero: R. de Santiago 747 (Holotype: FCME; isotypes: BH, IBUG, MEXU, MO, XAL). The relegation of this epithet to synonymy is based on the protologue and illustration as well as examination of numerous paratypes.</p> <p>Quercus obtusanthera Trel., Repert. Spec. Nov. Regni Veg. 33: 318 (1934). Type:— MEXICO, Guerrero: L. Schultze 343 (Holotype: at B destroyed). The determination of Q. obtusanthera as probably Q. salicifolia is based on Trelease’s (1933) description as no images or original specimens exist.</p> <p>Quercus petiolata (Trel.) A.E.Murray non Schur (1857), Kalmia 13: 27 (1983).</p> <p>Quercus rubramenta Trel., Repert. Spec. Nov. Regni Veg. 33: 318 (1934). Types: MEXICO, Guerrero. L. Schultze 371 and 372 (Syntypes: at B destroyed). The determination of Q. rubramenta as Q. salicifolia is based on Trelease’s (1934) description as no images or original specimens exist.</p> <p>Quercus salicifolia var. seemannii (Liebm.) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 207 (1884).</p> <p>Quercus seemannii Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 188 (1854). Type:— PANAMA, Veraguas: B. Seemann 1228 (Lectotype: K000575040! designated by Trelease (1924, p. 226, pl. 317a); isolectotypes: BM!, G!, GH? (as s.n.)!, K!, UCSB (fragment)!). Liebmann (1854, p. 188) wrote “leg. Seemann (Coll. n. 1228 in hb. Hook.!)” but there are two collections of Seemann 1228 in the Hooker Herbarium, both of which are named by Liebmann in his hand as Q. seemanni Liebm.</p> <p>Quercus tahuasalana Trel., Mem. Natl.Acad. Sci. 20: 154, pl. 302b (1924). Type:— MEXICO, Michoacán: E. Langlassé 211 (Syntypes at B destroyed. Lectotype (designated here): G00358146!; isolectotypes: GH!, K!, P!, UCSB (fragment from G lectotype)!, UCSB (fragment)!, US!). Trelease (1924, p. 154) wrote “ Langlassé, 217, June 18, 1898, the type” but on p. 226 in the caption to pl. 302b he wrote “The type (Mexico, Langlassé, 211).” The image detail in pl. 302b is from B now lost. Both Langlassé 211 and 217 are Q. salicifolia and are considered to be syntypes.</p> <p>Distribution: Costa Rica, El Salvador, Guatemala, Honduras, Mexico (Chiapas, Colima, Durango, Guerrero, Hidalgo,</p> <p>Jalisco, México, Michoacán, Morelos, Nayarit, Oaxaca, Puebla, San Luis Potosí, Sinaloa, Veracruz, Zacatecas),</p> <p>Nicaragua, Panama. The occurrence of Q. salicifolia in México, San Luis Potosí, Sinaloa, and Zacatecas reported by</p> <p>Villaseñor (2016) needs to be confirmed.</p> <p>Quercus sapotifolia Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 185 (1854). Type:— GUATEMALA: G. V. Skinner 6 (Holotype: K000512931!).</p> <p>Quercus amissiloba Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 17(4): 357 (1938). Type:— HONDURAS, Comayagua: T. G. Yuncker et al. 6379 (Holotype: ILL00007474!; isotypes: F!, G!, GH!, K!, MO!, NY!, US!).</p> <p>Quercus apanecana Trel., Mem. Natl. Acad. Sci. 20: 152 (1924). Type:— EL SALVADOR, Ahuachapan: P. C. Standley 20187 (Holotype: US 00089403!).</p> <p>Quercus donnell-smithii Trel., Mem. Natl. Acad. Sci. 20: 162, pl. 318 (1924). Type:— GUATEMALA, Guatemala: J. D. Smith 1967 (Lectotype (designated here): US00089494!; isolectotypes: GH!, NY!). Although Trelease cited Smith 1967 as the type, there is no indication of a collection locality. In pl. 318 two different specimens are shown which Trelease says “are two variants of the type.” Neither of these is a good match for the NY or US specimens.</p> </div>	https://treatment.plazi.org/id/794B8792FFFCFFDA3E88F8EEFEE6F7F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FFD8FFF93E88FF67FC42FCEB.text	794B8792FFD8FFF93E88FF67FC42FCEB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus panamandinaea C. H. Mull., U. S. D. A. Bur. Pl. Industr. Misc. Publ.	<div><p>Quercus panamandinaea C.H.Mull., U.S.D.A. Bur. Pl. Industr. Misc. Publ. 477: 26, pl. 21, 22 (1942). Type:—</p> <p>PANAMA, Chiriquí: R. E. Woodson Jr. &amp; R. W. Schery 360 (Holotype: MO-144140!; isotypes: GH!, NA, UCSB</p> <p>(fragment)!). Quercus rapurahuensis Pittier ex Seemen, Bull. Herb. Boissier, sér. 2, 4(7): 654. (1904). Type:— COSTA RICA, San</p> <p>José: A. Tonduz 11795 (Lectotype: BR0000006914257!designated by Trelease (1924, p. 225, pl.275);isolectotypes:</p> <p>BM!, BR!, CAS!, G!, GH!, K!, NY!, P!, UCSB (fragment)!, US!). This species was also lectotypified by Burger</p> <p>(1977, p. 77), but it was already effectively lectotypified based on the same type by Trelease (1924). Quercus seemannii Liebm. subsp. gulielmi-treleasei (C.H.Mull.) A.E.Murray, Kalmia 13: 27 (1983). Quercus seemannii var. gulielmi-treleasei (C.H.Mull.) A.E.Murray, Kalmia 13: 27 (1983). Quercus seemannii subsp. rapurahuensis (Trel.) A.E.Murray, Kalmia 13: 27 (1983). Quercus seemannii var. rapurahuensis (Pittier ex Trel.) A.E.Murray, Kalmia 13: 27 (1983). Quercus undulata Benth. non Torr. (1827), Pl. Hartw. 81 (1841). Distribution: Costa Rica, El Salvador, Guatemala, Honduras, Mexico (Chiapas, Guerrero, Jalisco, Oaxaca), Nicaragua, Panama. Villaseñor (2016) also reported Q. benthamii from the Mexican states of Chihuahua and Coahuila, but these reports seem unlikely.</p> <p>Quercus bolanyosensis Trel., Mem. Natl. Acad. Sci. 20: 123, pl. 233 (1924). Type:— MEXICO, Jalisco: J. N. Rose 2958 (Holotype: US00089447!; isotypes: GH!, NY!, UCSB (fragment)!).</p> <p>Distribution: Mexico (Durango, Jalisco, Nayarit, Sinaloa, Zacatecas).</p></div> 	https://treatment.plazi.org/id/794B8792FFD8FFF93E88FF67FC42FCEB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FF90FFB13E88FF67FB76F7F1.text	794B8792FF90FFB13E88FF67FB76F7F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus pilicaulis fm. elongata C. H. Mull.	<div><p>Quercus pilicaulis f. elongata C.H.Mull. = Quercus rugosa Neé</p> <p>Quercus pilicaulis f. exserta C.H.Mull. = Quercus rugosa Neé</p> <p>Quercus pilicaulis f. hurteri Trel. = Quercus rugosa Neé</p> <p>Quercus pilicaulis f. macrodonta Trel. = Quercus peduncularis Neé</p> <p>Quercus pilicaulis f. obovalis Trel. = Quercus peduncularis Neé</p> <p>Quercus pinalensis Trel. = Quercus glaucescens Bonpl.</p> <p>Quercus pinnativenulosa C.H.Mull.</p> <p>Quercus planipocula Trel.</p> <p>Quercus planipocula Trel. f. subsessilifolia Trel. = Quercus aristata Hook. &amp; Arn.</p> <p>Quercus platyphylla E.F.Warb. (1939) non Morogues (1877) = Quercus nanchititlensis B.M.Barthol. &amp; Almeda Quercus poculifer Trel. = Quercus magnoliifolia Neé</p> <p>Quercus polymorpha Schltdl. &amp; Cham.</p> <p>Quercus polymorpha f. angustifolia C.H.Mull. = Quercus polymorpha Schltdl. &amp; Cham.</p> <p>Quercus porphyrogenita Trel.</p> <p>Quercus porriginosa Trel. = Quercus elliptica Neé</p> <p>Quercus potosina Trel.</p> <p>Quercus potosina f. aperta Trel. = Quercus potosina Trel.</p> <p>Quercus potosina f. exilis Trel. = Quercus potosina Trel.</p> <p>Quercus praeco Trel.</p> <p>Quercus prainiana Trel. = Quercus coffeicolor Trel.</p> <p>Quercus pringlei Seemen</p> <p>Quercus pringlei f. dentata C.H.Mull. = Quercus pringlei Seemen</p> <p>Quercus prinopsis Trel. = Quercus laeta Liebm.</p> <p>Quercus productipes Trel. = Quercus aristata Hook. &amp; Arn.</p> <p>Quercus pubinervis M.Martens &amp; Galeotti = Quercus elliptica Neé</p> <p>Quercus pulchella Bonpl. = Quercus castanea Neé</p> <p>Quercus pungens Liebm.</p> <p>Quercus pungens subsp. vaseyana (Buckley) A.E.Murray = Quercus pungens Liebm.</p> <p>Quercus pungens var. vaseyana (Buckley) C.H.Mull. = Quercus pungens Liebm.</p> <p>Quercus purpusii Trel. = Quercus rugosa Neé</p> <p>Quercus purulhana Trel.</p> <p>Quercus radiata Trel.</p> <p>Quercus radlkoferiana Trel. = Quercus glabrescens Benth.</p> <p>Quercus rapurahuensis Pittier ex Seemen = Quercus benthamii A.DC.</p> <p>Quercus ravenscroftiana A. Murray ≠ Fagaceae (see Appendix 1, Excluded Names)</p> <p>Quercus reevesii Trel. = Quercus lancifolia Schltdl. &amp; Cham.</p> <p>Quercus rekonis Trel.</p> <p>Quercus repanda Bonpl. (1809) non Michx. (1801) = Quercus subtriloba Trel.</p> <p>Quercus resinosa Liebm.</p> <p>Quercus reticulata Bonpl. = Quercus rugosa Neé</p> <p>Quercus reticulata var. crassifolia Oerst. = Quercus greggii (A.DC.) Trel.</p> <p>Quercus reticulata f. dugesii Trel. = Quercus spicata Bonpl.</p> <p>Quercus reticulata var. greggii A.DC. = Quercus greggii (A.DC.) Trel.</p> <p>Quercus reticulata var. laxa (Liebm.) Wenz. = Quercus xylina Scheidw.</p> <p>Quercus reticulata f. longa Trel. = Quercus rugosa Neé</p> <p>Quercus reticulata f. pungens C.H.Mull. = Quercus intricata Trel.</p> <p>Quercus reticulata subsp. reticulata var. conglomerata (Trel.) A.Camus = Quercus rugosa Neé</p> <p>Quercus reticulata subsp. reticulata var. dugsei (Trel.) A.Camus = Quercus spicata Bonpl.</p> <p>Quercus reticulata subsp. reticulata var. squarrosa (Trel.) A.Camus = Quercus rugosa Neé</p> <p>Quercus reticulata subsp. rhodophlebia (Trel.) A.Camus = Quercus rugosa Neé</p> <p>Quercus reticulata subsp. rhodophlebia f. applanata (Trel.) A.Camus = Quercus spicata Bonpl.</p> <p>Quercus reticulata subsp. rhodophlebia f. apus (Trel.) A.Camus = Quercus spicata Bonpl.</p> <p>Quercus reticulata subsp. rhodophlebia f. concava (Trel.) A.Camus = Quercus rugosa Neé</p> <p>Quercus reticulata subsp. rhodophlebia f. crenata (Trel.) A.Camus = Quercus obtusata Bonpl.</p></div> 	https://treatment.plazi.org/id/794B8792FF90FFB13E88FF67FB76F7F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FF91FFB03E88FF67FB85F7F1.text	794B8792FF91FFB03E88FF67FB85F7F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus reticulata subsp. rhodophlebia fm. inclusa (Trel.) A. Camus	<div><p>Quercus reticulata subsp. rhodophlebia f. inclusa (Trel.) A.Camus = Quercus spicata Bonpl.</p> <p>Quercus reticulata var. segoviensis (Liebm.) Wenz. = Quercus segoviensis Liebm.</p> <p>Quercus reticulata f. squarrosa Trel. = Quercus rugosa Neé</p> <p>Quercus retusa Liebm. (1854) non Raf. (1838) = Quercus oleoides Schltdl. &amp; Cham.</p> <p>Quercus revoluta Trel. = Quercus subtriloba Trel.</p> <p>Quercus revoluta f. acuta C.H.Mull. = Quercus intricata Trel.</p> <p>Quercus revoluta f. dysophyllopsis Trel. = Quercus greggii (A.DC.) Trel.</p> <p>Quercus rhodophlebia Trel. = Quercus rugosa Neé</p> <p>Quercus rhodophlebia f. applanata Trel. = Quercus spicata Bonpl.</p> <p>Quercus rhodophlebia f. apus Trel. = Quercus spicata Bonpl.</p> <p>Quercus rhodophlebia f. concava Trel. = Quercus rugosa Neé</p> <p>Quercus rhodophlebia f. crenata Trel. = Quercus obtusata Bonpl.</p> <p>Quercus rhodophlebia f. inclusa Trel. = Quercus spicata Bonpl.</p> <p>Quercus rosei Trel. = Quercus fulva Liebm.</p> <p>Quercus roseovenulosa Trel. = Quercus laurina Bonpl.</p> <p>Quercus rossii Trel. = Quercus castanea Neé</p> <p>Quercus rossii f. arsenei Trel. = Quercus laeta Liebm.</p> <p>Quercus rubescens Trel. = Quercus magnoliifolia Neé</p> <p>Quercus rubramenta Trel. = Quercus salicifolia Neé</p> <p>Quercus rugosa Neé</p> <p>Quercus rugulosa M.Martens &amp; Galeotti = Quercus mexicana Bonpl.</p> <p>Quercus rugulosa f. subtruncata Trel. = Quercus mexicana Bonpl.</p> <p>Quercus runcinatifolia Trel. &amp; C.H.Mull. = Quercus gravesii Sudw.</p> <p>Quercus runcinatifolia f. lata Trel. &amp; C.H.Mull. = Quercus gravesii Sudw.</p> <p>Quercus rydbergiana Cockerell = Quercus undulata Torr.</p> <p>Quercus rysophylla Weath.</p> <p>Quercus sacame Trel. = Quercus arizonica Sarg.</p> <p>Quercus sagata E.F.Warb. = Quercus dysophylla E.F.Warb.</p> <p>Quercus sagrana Nutt.</p> <p>Quercus salicifolia Neé</p> <p>Quercus salicifolia var. oajacana (Liebm.) Wenz. = Quercus elliptica Neé</p> <p>Quercus salicifolia var. seemannii (Liebm.) Wenz. = Quercus salicifolia Neé</p> <p>Quercus salicifolia var. tlapuxahuensis (A.DC.) Wenz. = Quercus tlapuxahuensis A.DC.</p> <p>Quercus × saltillensis Trel.</p> <p>Quercus salvadorensis Trel. = Quercus xalapensis Bonpl.</p> <p>Quercus sanchez-colinii Martínez = Quercus diversifolia Neé</p> <p>Quercus santaclarensis C.H.Mull. = Quercus grisea Liebm.</p> <p>Quercus sapotifolia Liebm.</p> <p>Quercus sarahmariae Nixon &amp; Barrie</p> <p>Quercus sartorii Liebm.</p> <p>Quercus sartorii f. magma Trel. = Quercus sartorii Liebm.</p> <p>Quercus schenckiana Trel. = Quercus sebifera Trel.</p> <p>Quercus scherzeri Trel. = Quercus castanea Neé</p> <p>Quercus schippii Standl. = Quercus insignis M.Martens &amp; Galeotti</p> <p>Quercus schultzei Trel. = Quercus peduncularis Neé</p> <p>Quercus scytophylla Liebm.</p> <p>Quercus sebifera Trel.</p> <p>Quercus sebifera f. comitanensis Trel. = Quercus sebifera Trel.</p> <p>Quercus seemannii Liebm. = Quercus salicifolia Neé</p> <p>Quercus seemannii subsp. gulielmi-treleasei (C.H.Mull.) A.E.Murray = Quercus benthamii A.DC.</p> <p>Quercus seemannii var. gulielmi-treleasei (C.H.Mull.) A.E.Murray = Quercus benthamii A.DC.</p> <p>Quercus seemannii subsp. rapurahuensis (Trel.) A.E.Murray = Quercus benthamii A.DC.</p> <p>Quercus seemannii var. rapurahuensis (Pittier ex Trel.) A.E.Murray = Quercus benthamii A.DC.</p> <p>Quercus seemannii subsp. tonduzii (Seemen) A.E.Murray = Quercus × tonduzii Seemen</p></div> 	https://treatment.plazi.org/id/794B8792FF91FFB03E88FF67FB85F7F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FF92FFB33E88FF67FC50F7F1.text	794B8792FF92FFB33E88FF67FC50F7F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus seemannii var. tonduzii (Seemen) A. E. Murray	<div><p>Quercus seemannii var. tonduzii (Seemen) A.E.Murray = Quercus × tonduzii Seemen</p> <p>Quercus segoviensis Liebm.</p> <p>Quercus seibertii C.H.Mull. = Quercus insignis M.Martens &amp; Galeotti</p> <p>Quercus seleri Trel. = Quercus castanea Neé</p> <p>Quercus sentenelensis C.H.Mull. = Quercus muehlenbergii Engelm.</p> <p>Quercus serra Liebm. (1854) non Unger (1845) = Quercus cortesii Liebm.</p> <p>Quercus serrulata Trel. = Quercus castanea Neé</p> <p>Quercus shumardii Buckley var. microcarpa (Torr.) Shinners = Quercus gravesii Sudw.</p> <p>Quercus sideroxyla Bonpl.</p> <p>Quercus sideroxyla f. aquifolia Trel. = Quercus rugosa Neé</p> <p>Quercus sideroxyla f. ciliifera Trel. = Quercus laurina Bonpl.</p> <p>Quercus × sierramadrensis C.H.Mull.</p> <p>Quercus siguatepequeana Trel. = Quercus sapotifolia Liebm.</p> <p>Quercus sillae Trel. = Quercus fusiformis Small</p> <p>Quercus siltepecana Matuda = Quercus vicentensis Trel.</p> <p>Quercus simillima Trel. = Quercus tridens Bonpl.</p> <p>Quercus sinuata Walter</p> <p>Quercus sinuata subsp. breviloba (Torr.) A.E.Murray = Quercus sinuata Walter</p> <p>Quercus sinuata var. breviloba (Torr.) C.H.Mull. = Quercus sinuata Walter</p> <p>Quercus sipuraca Trel. = Quercus durifolia Seemen</p> <p>Quercus skinneri Benth.</p> <p>Quercus skutchii Trel.</p> <p>Quercus sonomensis Benth. ex A.DC. = Quercus kelloggii Newb.</p> <p>Quercus sororia Liebm.</p> <p>Quercus spathulistipula Trel. = Quercus castanea Neé</p> <p>Quercus spicata Bonpl.</p> <p>Quercus spinulosa M.Martens &amp; Galeotti = Quercus crassifolia Bonpl.</p> <p>Quercus splendens Neé = Quercus peduncularis Neé</p> <p>Quercus splendens var. pallidior A.DC. = Quercus crassifolia Bonpl.</p> <p>Quercus standleyi Trel. = Quercus tuberculata Liebm.</p> <p>Quercus stellapila (Sarg.) H.B. Parks ex Cory = Quercus gravesii Sudw.</p> <p>Quercus stellata Wagenh. var. utahensis A.DC = Quercus gambelii Nutt.</p> <p>Quercus stipularis Bonpl. = Quercus crassifolia Bonpl.</p> <p>Quercus striatula Trel. = Quercus microphylla Neé</p> <p>Quercus striatula f. otinapensis Trel. = Quercus microphylla Neé</p> <p>Quercus strombocarpa Liebm. = Quercus insignis M.Martens &amp; Galeotti</p> <p>Quercus subavenia Trel. = Quercus depressa Bonpl.</p> <p>Quercus subcrispata Trel. = Quercus castanea Neé</p> <p>Quercus subspathulata Trel.</p> <p>Quercus substenocarpa Trel. = Quercus germana Schltdl. &amp; Cham.</p> <p>Quercus subtriloba Trel.</p> <p>Quercus subturbinella Trel. = Quercus turbinella Greene</p> <p>Quercus suchiensis Warb. = Quercus rugosa Neé</p> <p>Quercus supranitida C.H.Mull. = Quercus intricata Trel.</p> <p>Quercus synthetica Trel. = Quercus glaucescens Bonpl.</p> <p>Quercus synthetica var. crenifolia Trel. = Quercus glaucescens Bonpl.</p> <p>Quercus tahuasalana Trel. = Quercus salicifolia Neé</p> <p>Quercus tamiapensis C.H.Mull. = Quercus alveolata Trel.</p> <p>Quercus tarahumara Spellenb., J.D.Bacon &amp; Breedlove</p> <p>Quercus tardifolia C.H.Mull. = Quercus gravesii Sudw.</p> <p>Quercus tenuiaristata Trel. = Quercus cortesii Liebm.</p> <p>Quercus × tenuiloba C.H.Mull.</p> <p>Quercus × tenuiloba nothof. gracilis C.H.Mull. = Quercus × sierramadrensis C.H.Mull.</p> <p>Quercus tenuiloba f. hirsuta C.H.Mull. = Quercus rysophylla Weath.</p></div> 	https://treatment.plazi.org/id/794B8792FF92FFB33E88FF67FC50F7F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FF9DFFBC3E88FF67FB3EFCB9.text	794B8792FF9DFFBC3E88FF67FB3EFCB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus sinuata	<div><p>Q. sinuata × Q. pungens Mexico (Nuevo León). J. L. Berlandier 1400 BM!, GH!</p> <p>Q. skutchii × Q. crispipilis Guatemala (Huehuetenango). J. A. Steyermark 50635 A!, CAS!, F!, G!</p> <p>Mexico (Chiapas). D. E. Breedlove 6956 DS!, F!</p> <p>Q. spicata × Q. diversifolia Mexico (Jalisco). L. M. Gonzalez V. 572 MEXU!</p> <p>Q. spicata × Q. rugosa Mexico (Veracruz). R. Hernandez M. 778 F!, MEXU!</p> <p>Q. subtriloba × Q.? Mexico (Michoacán). J. Gregg 795 MO!</p> <p>Q. tridens × Q. depressa Mexico (Hidalgo). H. E. Moore 2816 GH!</p> <p>Q. tridens × Q.? Mexico (Nuevo León). C. H. Muller &amp; M. T. Muller 581 A!</p> <p>Q. tuberculata × Q. grisea ? Mexico (Chihuahua). H. LeSueur 1490 GH!</p> <p>Q. tuberculata × Q. laeta ? Mexico (Baja California Sur). J. H. Thomas 7835 CAS!, GH!, GH!, US!, VT!</p> <p>Q. undulata × Q.? Mexico (Chihuahua). R. M. Stewart &amp; I. M. Johnston 2078 GH!</p> <p>Mexico (Nuevo León). C.H.Muller &amp; M. T. Muller 748 A!</p> <p>Q. xalapensis × Q. cortesii Mexico (Chiapas). D. E. Breedlove 11002 DS!, F!, WIS!</p> <p>Q. xalapensis × Q. grahamii Mexico (Jalisco). R. McVaugh 10258 CAS!, G!, MEXU!, MICH!, US!</p> <p>Mexico (Nuevo León). C. H. Muller 1124 A!, MEXU!</p> <p>Q. xalapensis × Q. skinneri El Salvador (San Vicente). P. C. Standley 21599 GH!, ILL!, MO!, NY!, US!</p> <p>Mexico (Chiapas). R. L. Dressler 1592 GH!</p> <p>APPENDIX 5. Latin American Quercus published since 1990 (recognized names are in bold face).</p> </div>	https://treatment.plazi.org/id/794B8792FF9DFFBC3E88FF67FB3EFCB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FFF7FFD03E88F9E2FD00FD2E.text	794B8792FFF7FFD03E88F9E2FD00FD2E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus sororia Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854	<div><p>Quercus sororia Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 175 (1854).</p> <p>Type:— MEXICO, Oaxaca: F. M. Liebmann 3562 (Holotype: C 10012530!; isotype UCSB (fragment from C</p> <p>holotype)!). Quercus glauca Oerst. non Thunb. (1784), Chênes Amér. Trop. 16, 29, pl. 36 (1869). Distribution: Mexico (Durango, Guerrero, Hidalgo, Jalisco, México, Michoacán, Nayarit, Oaxaca, Puebla, San Luis Potosí, Sinaloa).</p> <p>Quercus spicata Bonpl., Pl. Aequinoct. 2: 46, pl. 89 (1809). Type:— MEXICO, Hidalgo: A. Bonpland 4061 (p.p.) (Holotype: P00129694!; isotypes: B-W!, P!, UCSB (fragment)!). There are four sheets of Bonpland 4061 (p.p.) at P that are isotypes of Q. spicata (P00129695!, P00129696!, P00129697!, and P00129731!).</p> <p>Quercus alvarezensis Trel., Mem. Natl.Acad. Sci. 20: 76, pl. 104 (1924). Type:— MEXICO, San Luis Potosí: E. Palmer 69 (Holotype: MO-143903!; isotypes: A!, BM!, CAS!, F!, GH!, NY!, UCSB (fragment from MO holotype)!, US!).</p> <p>Quercus convallata Trel., Mem. Natl. Acad. Sci. 20: 88, pl. 140 (1924). Type:— MEXICO, Nayarit: M. L. Diguet s. n. (Holotype: P00754073!; isotypes: NY!, P!, UCSB (fragment)!). Although the specimen labels say that this collection is from Jalisco, Sierra de Nayarit (Territoire Huichol), the type locality is actually in Nayarit.</p> <p>Quercus dugesii (Trel.) A. Nelson, Amer. J. Bot. 23(4): 268 (1936). The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. spicata as broadly interpreted.</p> <p>Quercus hartwegii Benth., Pl. Hartw. 56 (1840). Type:— MEXICO, Michoacán: T. Hartweg 432 (Holotype: K000512626!; isotypes: BM!, CAS!, CGE!, E!, FI!, G!, LD (image)!, NY!, OXF!, P!, UCSB (fragment)!).</p> <p>Quercus martensiana Trel. f. berlandieri Trel., Mem. Natl. Acad. Sci. 20: 60 (1924). Type:— MEXICO, Tamaulipas: J. Berlandier 2131 (Lectotype (designated here): GH00062251!; isolectotypes: BM!, G!, K!, P!). Trelease (1924, p. 60) did not designate where the holotype was deposited.</p> <p>Quercus obtusata Bonpl. var. hartwegii (Benth.) A.DC., Prodr. 16(2.1): 27 (1864).</p> <p>Quercus pandurata Bonpl. var. hartwegii (Benth.) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 197 (1884).</p> <p>Quercus reticulata Bonpl. f. dugesii Trel., Mem. Natl.Acad. Sci. 20: 75, pl. 98b (1924). Type:— MEXICO, Guanajuato: A. Dugès 3 (Holotype: US 00089630!; isotype: GH!).</p> <p>Quercus reticulata subsp. reticulata var. dugesii (Trel.) A.Camus, Chênes Atlas 2: 94 (1935).</p> <p>Quercus reticulata subsp. rhodophlebia (Trel.) A.Camus f. applanata (Trel.) A.Camus, Chênes Texte 2: 581 (1939).</p> <p>Quercus reticulata subsp. rhodophlebia f. apus (Trel.) A.Camus, Chênes Atlas 2: 95 (1935).</p> <p>Quercus reticulata subsp. rhodophlebia f. inclusa (Trel.) A.Camus, Chênes Atlas 2: 95 (1935).</p> <p>Quercus rhodophlebia Trel. f. applanata Trel., Mem. Natl. Acad. Sci. 20: 74, pl. 96a (1924). Type:— MEXICO, Zacatecas: J. N. Rose 2735 (Holotype: US 00089625!; isotypes: GH!, NY!).</p> <p>Quercus rhodophlebia f. apus Trel., Mem. Natl. Acad. Sci. 20: 74, pl. 96b (1924). Type:— MEXICO, Zacatecas: J.N. Rose 2721 (Holotype: US 00089624!; isotypes: GH!, NY!).</p> <p>Quercus rhodophlebia f. inclusa Trel., Mem. Natl. Acad. Sci. 20: 74, pl. 97a (1924). Type:— MEXICO, Jalisco: J. N. Rose 3726 (Holotype: US 00089621!).</p> <p>Quercus vellifera Trel., Mem. Natl. Acad. Sci. 20: 73, pl. 90 (1924). Type:— MEXICO: E. Palmer s. n. (9–15 Oct 1891) (Holotype: US 00089690!).</p> <p>Quercus verde C.H.Mull., J. Arnold Arbor. 17(3): 163 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 373 (Holotype: A00055683!; isotypes: TEX (image)!, UCSB (fragment)!).</p> <p>Distribution: Mexico (Chihuahua, Ciudad de México, Coahuila, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco,</p> <p>México, Michoacán, Morelos, Nayarit, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa, Tamaulipas,</p> <p>Veracruz, Zacatecas).</p> <p>Quercus subspathulata Trel., Mem. Natl. Acad. Sci. 20: 56, pl. 52a (1924). Type:— MEXICO, Durango: J. N. Rose 2239 (Holotype: US 00089665!).</p> <p>Quercus pallidifolia C.H.Mull., Contr. Univ. Michigan Herb. 9(7): 522 (1972). Type:— MEXICO, Nayarit: R. Pérez C. 2135 (Holotype: BH; isotypes: MEXU!, MICH!, UCSB (fragment from BH holotype)!).</p> <p>Distribution: Mexico (Chihuahua, Colima, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, México, Michoacán,</p> <p>Nayarit, Oaxaca, Puebla, Querétaro, Sinaloa, Sonora, Zacatecas).</p> <p>Quercus subtriloba Trel., Mem. Natl. Acad. Sci. 20: 81, pl. 118 (1924). Type:— MEXICO, Hidalgo: C. G. Pringle 10303 (Holotype: US00089667!; isotypes: A!, ASU (image)!, CAS!, COLO (image)!, DS!, DUKE (image)!, ENCB (image)!, F!, GH!, LL (image)!, MICH!, OKLA (image)!, TEX (image)!, UC!, UCSB (fragment)!, VT!, WIS!).</p> <p>Quercus lecomteana Trel., Mem. Natl. Acad. Sci. 20: 81, pl. 115 (1924). Type:— MEXICO, Veracruz: L. Hahn 255 (Holotype: P00754126!; isotypes: P!, UCSB (fragment)!).</p> <p>Quercus repanda Bonpl. non Michx. (1801), Pl.Aequinoct. 2: 31, pl. 79 (1809).Type:— MEXICO, Hidalgo: A.Bonpland 4081 (Holotype: P00129705!; isotypes B-W!, CGE (as s.n.)!, P!, UCSB (fragment)!).</p> <p>Quercus revoluta Trel., Mem. Natl. Acad. Sci. 20: 78, pl. 111 (1924). Type:— MEXICO, Puebla: C. G. Pringle 8966 (Holotype at B destroyed. Lectotype (designated here): US 00089629!; isolectotypes: A!, BM!, C!, CAS!, E!, FI!, G!, GH!, HBG (image) !, ILL!, K!, L!, M!, MO!, NY!, P !, PH!, UC!, UCSB (fragment) !, US!, VT !).</p> <p>Distribution: Mexico (Ciudad de México, Guanajuato, Hidalgo, Jalisco, México, Michoacán, Nuevo León, Oaxaca,</p> <p>Puebla, Querétaro, San Luis Potosí, Tamaulipas, Tlaxcala, Veracruz, Zacatecas).</p> <p>Quercus tarahumara Spellenb., J.D.Bacon &amp; Breedlove, Madroño 42(1): 28, figs. 1, 2 (1995). Type:— MEXICO, Chihuahua: R. D. Spellenberg 10830 (Holotype: NMC (image)!; isotypes: CAS!, MEXU!).</p> <p>Distribution: Mexico (Chihuahua, Durango, Sinaloa, Sonora).</p> <p>Quercus tinkhamii C.H.Mull., Amer. Midl. Naturalist 27(2): 481 (1942). Type:— MEXICO, Nuevo León: F. Shreve &amp; E. R. Tinkham 9686 (Holotype: GH00069874!; isotype: UCSB (fragment from GH holotype)!). The distinction between Q. tinkhamii and Q. mohriana needs additional study. If Q. tinkhamii is included as a synonym of Q. mohriana, the distribution of Q. mohriana is much broader in Mexico.</p> <p>Distribution: Mexico (Coahuila, Durango, Guanajuato, Hidalgo, Jalisco, Nuevo León, Querétaro, San Luis Potosí,</p> <p>Tamaulipas, Veracruz, Zacatecas).</p> <p>Quercus tlapuxahuensis A.DC., Prodr. 16(2.1): 29 (1864). Type:— MEXICO, Michoacán: T. Hartweg 430 (Holotype: G00358122!; isotypes: BM!, E!, K!, L!, LE!, P!, UCSB (fragment from G holotype), UCSB (fragment)!).</p> <p>Quercus nitida M.Martens &amp; Galeotti non Raf. (1838), Bull. Acad. Roy. Sci. Bruxelles 10(1): 210 (1843). See Q. uruapanensis below.</p> <p>Quercus salicifolia Neé var. tlapuxahuensis (A.DC.) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 207 (1884).</p> <p>Quercus uruapanensis Trel., Contr. U.S. Natl. Herb. 23(2): 191 (1922). Type:— MEXICO, Michoacán: H. Galeotti 121 (Holotype: BR0000006912444!; isotypes: BR!, G!, K!, P!, UCSB (fragment)!). Nom. nov. for Q. nitida M. Martens &amp; Galeotti (1843) non Raf. (1838). The placement of this species needs additional study, but we consider the type to fall within the variation found in Q. tlapuxahuensis as broadly interpreted.</p> <p>Distribution: Mexico (Jalisco, México, Michoacán).</p></div> 	https://treatment.plazi.org/id/794B8792FFF7FFD03E88F9E2FD00FD2E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FF93FFB23E88FF67FC34F7F1.text	794B8792FF93FFB23E88FF67FC34F7F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus tepicana Trel.	<div><p>Quercus tepicana Trel. = Quercus magnoliifolia Neé</p> <p>Quercus tepoxuchilensis Trel. = Quercus castanea Neé</p> <p>Quercus tepoxuchilensis f. perplexans Trel. = Quercus castanea Neé</p> <p>Quercus texana Buckley var. chesosensis Sarg. = Quercus gravesii Sudw.</p> <p>Quercus texana var. stellapila Sarg. = Quercus gravesii Sudw.</p> <p>Quercus texcocana Trel. = Quercus diversifolia Neé</p> <p>Quercus texcocana var. ampla Trel. = Quercus diversifolia Neé</p> <p>Quercus texquitzinae Trel. = Quercus glaucescens Bonpl.</p> <p>Quercus tinctoria W. Bartram var. californica Torr. = Quercus kelloggii Newb.</p> <p>Quercus tinkhamii C.H.Mull.</p> <p>Quercus tlapuxahuensis A.DC.</p> <p>Quercus tlapuxahuensis var. obconica Trel. = Quercus laurina Bonpl.</p> <p>Quercus tolimensis Bonpl. = Quercus humboldtii Bonpl.</p> <p>Quercus tomentella Engelm.</p> <p>Quercus tomentocaulis C.H.Mull. = Quercus insignis M.Martens &amp; Galeotti</p> <p>Quercus tomentosa Willd. nom. illigit. = Quercus peduncularis Neé</p> <p>Quercus tomentosa Willd. nom. illigit. var. abbreviata A.DC. = Quercus peduncularis Neé</p> <p>Quercus tomentosa Willd. nom. illigit. var. bullata A.DC. = Quercus peduncularis Neé</p> <p>Quercus tomentosa Willd. nom. illigit. var. communis A.DC. = Quercus peduncularis Neé</p> <p>Quercus tomentosa Willd. nom. illigit. var. diversifolia (Neé) A.DC. = Quercus diversifolia Neé</p> <p>Quercus tonaguiae Trel. = Quercus calophylla Schltdl. &amp; Cham.</p> <p>Quercus × tonduzii Seemen</p> <p>Quercus totutlensis A.DC. = Quercus sapotifolia Liebm.</p> <p>Quercus toumeyi Sarg.</p> <p>Quercus toxicodendrifolia Trel.</p> <p>Quercus transmontana Trel. = Quercus laeta Liebm.</p> <p>Quercus transmontana f. lata (Trel.) A.Camus = Quercus laeta Liebm.</p> <p>Quercus transmontana var. lata Trel. = Quercus laeta Liebm.</p> <p>Quercus transmontana var. obscura A.Camus = Quercus laeta Liebm.</p> <p>Quercus treleaseana A.Camus = Quercus affinis Scheidw.</p> <p>Quercus trichodonta Trel. = Quercus skinneri Benth.</p> <p>Quercus tridens Bonpl. = Quercus laurina Bonpl.</p> <p>Quercus trinidadensis C.H.Mull. = Quercus sebifera Trel.</p> <p>Quercus trinitatis Trel.</p> <p>Quercus tristis Liebm. = Quercus castanea Neé</p> <p>Quercus tristis f. mixcoensis Trel. = Quercus castanea Neé</p> <p>Quercus tristis f. niederleini Trel. = Quercus castanea Neé</p> <p>Quercus tristis f. sublobata (A.DC.) Trel. = Quercus castanea Neé</p> <p>Quercus tristis f. vulcani Trel. = Quercus castanea Neé</p> <p>Quercus tuberculata Liebm.</p> <p>Quercus tuberculata f. latifolia Martínez = Quercus tuberculata Liebm.</p> <p>Quercus tuitensis L.M.González</p> <p>Quercus turbinata Liebm. (1854) non Blume (1825) = Quercus polymorpha Schltdl. &amp; Cham.</p> <p>Quercus turbinella Greene</p> <p>Quercus turbinella subsp. ajoensis (C.H.Mull.) Felger &amp; C.H.Lowe = Quercus ajoensis C.H.Mull.</p> <p>Quercus turbinella var. ajoensis (C.H.Mull.) Little = Quercus ajoensis C.H.Mull.</p> <p>Quercus uhdeana Trel. = Quercus rugosa Neé</p> <p>Quercus × undata Trel.</p> <p>Quercus undulata Benth. (1841) non Torr. (1827) = Quercus benthamii A.DC.</p> <p>Quercus undulata Torr.</p> <p>Quercus undulata var. gambelii (Nutt.) Engelm. = Quercus gambelii Nutt.</p> <p>Quercus undulata var. grisea (Liebm.) Engelm. = Quercus grisea Liebm.</p> <p>Quercus undulata var. pungens (Liebm.) Engelm. = Quercus pungens Liebm.</p> <p>Quercus undulata var. vaseyana (Buckley) Rydb. = Quercus pungens Liebm.</p></div> 	https://treatment.plazi.org/id/794B8792FF93FFB23E88FF67FC34F7F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FFF1FFAD3E88FCD8FCDBF9A1.text	794B8792FFF1FFAD3E88FCD8FCDBF9A1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus tomentella Engelm., Trans. Acad. Sci. St. Louis	<div><p>Quercus tomentella Engelm., Trans. Acad. Sci. St. Louis 3: 393 (1877). Type:— MEXICO, Baja California: E. Palmer</p> <p>89 (Lectotype: MO-143905! designated by Trelease (1924, p. 223, pl. 212a); isolectotypes: GH!, K!, MO!, PH!,</p> <p>UCSB (fragment from MO lectotype)!, UCSB (fragment)!). Quercus chrysolepis Liebm. subsp. tomentella (Engelm.) A.E.Murray, Kalmia 13: 25 (1983). Quercus chrysolepis var. tomentella (Engelm.) A.E.Murray, Kalmia 13: 25 (1983). Distribution: Mexico (Baja California), U.S.A. (California).</p> <p>Quercus toumeyi Sarg., Gard. &amp; Forest 8: 92, fig. 13, 14 (1895). Type:— U.S.A., Arizona: J. W. Toumey s. n. (23 Jul 1894) (Holotype: A00034108!; isotypes: K!, UCSB (fragment from A holotype)!, UCSB (fragment)!, US!).</p> <p>Quercus barrancana Spellenb., Phytoneuron 2014 -105:fig.1–8 (2014). Type:— MEXICO, Chihuahua: R. D. Spellenberg &amp; W. Anderson 13487 (Holotype: NY (image)!; isotype: TEX (image)!). There is an image of the holotype in Spellenberg (2014), but as of this writing, NY has not received the holotype.</p> <p>Quercus chuhuichupensis C.H.Mull., Amer. Midl. Naturalist 19(3): 584 (1938). Type:— MEXICO, Chihuahua: H. LeSueur 525 (Holotype: BH; isotype: TEX!, UCSB (fragment from BH holotype)!).</p> <p>Quercus hartmanii Trel., Mem. Natl. Acad. Sci. 20: 95, pl. 154a (1924). Type:— MEXICO, Chihuahua: C. V. Hartman 633 (Holotype at B destroyed. Lectotype (designated here): GH00069942!; isolectotypes: A!, K!).</p> <p>Distribution: Mexico (Chihuahua, Durango, Nuevo León, Sinaloa, Sonora), U.S.A. (Arizona, New Mexico, Texas).</p> <p>Quercus toxicodendrifolia Trel., Mem. Natl.Acad. Sci. 20: 48, pl. 33a (1924). Type:— MEXICO, Hidalgo: J. Berlandier 533 (p.p.) (Holotype at W destroyed. Lectotype (designated here): G00358149!; isolectotypes: BM?!, P?!, UCSB? (fragment)!). The holotype of Q. toxicodendrifolia was collected on 21 May 1827 and is cited by Trelease (1924, p. 48). Collections of Berlandier 533 made on 19 May 1827 cannot be considered type specimens although G has a specimen from each of these dates. The G specimen of Berlandier 533 dated 21 May 1827 is the one here designated the lectotype. In the case of Berlandier 533 at BM which is undated, it can only be considered to be a possible isotype and now becomes a possible isolectotype. In the case of the P specimen there is a date of 21 May 1827 on the label but no number. Because Berlandier made collections of the same oak species using number 429 on this date, the P specimen can also only be considered a possible isolectotype. In the case of the fragments at UCSB, one of the Berlandier 533 fragments is dated 19 May 1827, so it is not a type collection. Another Berlandier 533 fragment is undated so it could be a possible lectotype fragment, but the leaf size is much larger than those of the lectotype and is probably from the G collection of 19 May, so it is not a lectotype fragment.</p> <p>Distribution: Mexico (Hidalgo, Puebla, Veracruz).</p> <p>Quercus trinitatis Trel., Mem. Natl. Acad. Sci. 20: 144, pl. 277b (1924). Type:— MEXICO, Hidalgo: C. G. Pringle 8888 (Holotype: BP; isotypes: A!, BM!, C!, CAS!, E!, ENCB (image)!, G!, GH!, HBG (image)!, ILL!, K!, MEXU!, MICH!, MO!, NY!, P!, PH!, S (image)!, UC!, UCSB (fragment)!, US!, VT!). This species is close to Q. perseifolia, and additional study is needed to determine if it is distinct.</p> <p>Distribution: Mexico (Hidalgo).</p> <p>Quercus tuberculata Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 181 (1854). Type:— MEXICO: B. Seemann 1970 (Holotype: K000512609!; isotypes: BM!, GH (as s.n.), K!). There are two specimens of Seemann 1970 in the Hooker Herbarium at K, but only one of them has a determination in Liebmann’s hand which is the one we consider to be the holotype.</p> <p>Quercus aurantiaca Trel., Mem. Natl. Acad. Sci. 20: 57, pl. 53a (1924). Type:— MEXICO, Chihuahua: R. Endlich 770 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 53a)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.</p> <p>Quercus edwardsiae C.H.Mull.,Amer.Midl.Naturalist 27(2):472(1942). Type:— MEXICO, Nuevo León: M. T. Edwards 435 (Holotype: BH; isotypes: DS!, UCSB (fragment from BH holotype)!).</p> <p>Quercus idonea Goldman, Contr. U.S. Natl. Herb. 16: 321 (1916). Type:— MEXICO, Baja California Sur: E. W. Nelson &amp; E. A. Goldman 7423 (Holotype: US 00089553!; isotype: A!).</p> <p>Quercus monterreyensis Trel. &amp; C.H.Mull., Bull. Torrey Bot. Club 63(3): 151 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 596 (p.p.) (Holotype: ILL00007499!; isotypes: MEXU!, TEX (image)!, UCSB (fragment)!). The specimen of C. H. Muller &amp; M. T. Muller 596 at A is Q. oblongata, so it is not an isotype.</p> <p>Quercus standleyi Trel., Contr. U.S. Natl. Herb. 23(2): 181 (1922). Type:— MEXICO, Sonora: J. N. Rose et al. 12789 (Holotype: US 00089652!; isotypes: NY!, UCSB (fragment)!, US!).</p> <p>Quercus tuberculata f. latifolia Martínez, Anales Inst. Biol. Univ. Nac. Autón. México 25(1 &amp; 2): 45, fig. 6 (1954). Type:— MEXICO, Durango: H. S. Gentry 5226 (Holotype: MEXU00010646!; isotypes: DS!, GH!, MEXU!, MO!, NY!, UC!, VT!).</p> <p>Distribution: Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Durango, Jalisco, México,</p> <p>Michoacán, Nayarit, Nuevo León, Sinaloa, Sonora, Zacatecas).</p> <p>Quercus tuitensis L.M.González, Brittonia 55(1): 42, figs. 1–4 (2003). Type:— MEXICO, Jalisco: L. M. González V. 4660 (Holotype: IBUG0160260 (image)!; isotypes: BH, CAS!, CHAPA, ENCB, F (image)!, IBUG (image)!, MICH (image)!, NY (image)!, TEX (image)!, US (image)!, WIS!).</p> <p>Distribution: Mexico (Jalisco).</p> <p>Quercus turbinella Greene, Ill. W. Amer. Oaks 1: 37 (1889). Type:— MEXICO, Baja California: G. W. Dunn s. n. (25 Sep 1888) (Holotype: NDG13622!; isotypes: NY!, UCSB (fragment)!). There is no date on the Dunn s. n. sheet at NDG, but it was the specimen from which the illustration was prepared for the protologue. The isotypes at NY and US give the collection date as 25 Sep 1888. All the other specimens at NDG were collected too late to be considered syntypes as the species was published on 1 May 1889 and the Greene s. n. collections of Q. turbinella at NDG were made later in 1889.</p> <p>Quercus dumosa Nutt. subsp. turbinella (Greene) A.E.Murray, Kalmia 13: 25 (1983).</p> <p>Quercus dumosa var. turbinella (Greene) Jeps., Fl. Calif. 1(2): 356 (1909).</p> <p>Quercus dumosa var. turbinella f. crispata Trel., Mem. Natl. Acad. Sci. 20: 117 (1924). Type:— MEXICO, Baja California: E. Palmer 29 (Holotype: MO-204123!; isotype: US!).</p> <p>Quercus subturbinella Trel., Mem. Natl. Acad. Sci. 20: 95, pl. 153b (1924). Type:— U.S.A., Arizona: E. L. Greene s. n. (2 Jul 1889) (Holotype: NDG13626!).</p> <p>Distribution: Mexico (Baja California, Baja California Sur, Chihuahua, Sonora), U.S.A. (Arizona, California,</p> <p>Colorado, Nevada, New Mexico, Texas, Utah).</p> <p>Quercus undulata Torr., Ann. Lyceum Nat. Hist. New York 2: 248 (1827). Type:— U.S.A., Colorado: James s. n. (Holotype: NY00073979!).</p> <p>Quercus carmenensis C.H.Mull., Amer. Midl. Naturalist 18(5): 847 (1937). Type:— MEXICO, Coahuila: F. L. Wynd &amp; C. H. Muller 639 (Lectotype (designated here): ILL00007461!; isolectotypes: A!, ILL!, K!, MO!, NY!, P!, UCSB (fragment)!, WIS!). Neither of the two specimens at ILL were marked as types.</p> <p>Quercus endemica C.H.Mull., Amer. Midl. Naturalist 18(5): 846 (1937). Type:— MEXICO, Coahuila: F. L. Wynd &amp; C. H. Muller 638 (Lectotype (designated here): ILL00007498!; isolectotypes: A!, ILL!, K!, MEXU!, MO!, NY!, UCSB (fragment)!). The two specimens at ILL were both marked as types.</p> <p>Quercus fendleri Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 170 (1854). Type:— U.S.A., New Mexio: A. Fendler 805 (Lectotype: K000832235! designated by Trelease (1924, p. 221, pl. 159); isolectotypes: BM!, GH!, MO!, NY!, PH!, UCSB (fragment from K holotype)!, WIS!).</p> <p>Quercus greggii (A.DC.) Trel. f. subglabrata C.H.Mull., J. Arnold Arbor. 25(4): 440 (1944). Type:— MEXICO, Coahuila: C. H. Muller 3238 (Holotype: GH00055662!; isotype: UC!).</p> <p>Quercus rydbergiana Cockerell, Torreya 3(1): 7 (1903). Type:— U.S.A., New Mexico: T. D. A. Cockerell s. n. (1902) (Holotype: NY00248767!).</p> <p>Distribution: Mexico (Chihuahua, Coahuila, Nuevo León, San Luis Potosí, Tamaulipas), U.S.A. (Colorado, New</p> <p>Mexico, Texas).</p> <p>Quercus urbani Trel., Proc. Amer. Philos. Soc. 60: 32, pl. 2 (1921). Type:— MEXICO, Guerrero: E. Langlassé 1066 (Holotype at B destroyed. Lectotype (designated here): GH00055682!; isolectotypes: G!, K!, MPU!, P!, US!).</p> <p>Quercus pennivenia Trel., Mem. Natl. Acad. Sci. 20: 135, pl. 256a (1924). Type:— MEXICO: B. Seemann 1998 (Holotype: K000512911!). There is apparently a typographic error in Trelease (1924, p. 224) in the caption to pl. 256. The holotype of Q. pennivenia is Seemann 1998 not 1978. Both Seemann 1968 and 1998 cited on p. 135 are Q. urbani whereas Seemann 1978 is Q. durifolia. The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. urbani as broadly interpreted.</p> <p>Distribution: Mexico (Chihuahua, Coahuila, Durango, Guanajuato, Guerrero, Jalisco, México, Michoacán, Nayarit,</p> <p>Oaxaca, Querétaro, Sinaloa, Sonora, Zacatecas).</p> <p>Quercus uxoris McVaugh, Contr. Univ. Michigan Herb. 9(7): 513 (1972). Type:— MEXICO, Jalisco: R. McVaugh 20292 (Holotype: MICH1109370!; isotypes: CAS!, G!, K!, LL (image)!, MO!, NY!, TEX (image)!, UCSB (fragment)!, US!).</p> <p>Distribution: Mexico (Aguascalientes, Colima, Guerrero, Jalisco, Michoacán, Oaxaca). The occurrence of Q. uxoris in Augscalientes reported by Villaseñor (2016) needs to be confirmed.</p> <p>Quercus vicentensis Trel., Mem. Natl. Acad. Sci. 20: 59 (1924). Type:— EL SALVADOR, San Vicente: P.C. Standley 21593 (Holotype: US00089688!; isotype: GH!). Breedlove considered some of the Mexican collections of this species from the states of Guerrero Jalisco, México, Michoacán, and Oaxaca to be a new species, but we have determined that the purported distinctions are variable and do not justify the recognition of a segregate species.</p> <p>Quercus centenaria L.M.González, Ibugana 9: 49, fig. 1–4 (2018). Type:— MEXICO, Jalisco: L.M.González V. &amp; L.M.Villarreal de Puga 4817 (Holotype: IBUG0174119; isotype: IBUG). Breedlove considered collections of this to be a new species and was planning to publish it. Although we have not seen the type of Q. centenaria, we have seen many of the paratypes as well as additional specimens collected by Breedlove, and disagree with both Breedlove and González V. We consider Q. centenaria to not be sufficiently distinct from Q. vicentensis.</p> <p>Quercus martinezii C.H.Mull., Anales Inst. Biol. Univ. Nac. Autón. México 24(2): 274, fig. 1 (1953[1954]). Type:— MEXICO, Jalisco: R. McVaugh 13912 (Holotype: BH; isotypes: K!, MEXU!, MICH!, UCSB (fragment from BH holotype)!, US!). The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. vicentenis as broadly interpreted.</p> <p>Quercus obscurirhachis Trel., Repert. Spec. Nov. Regni Veg. 33: 315 (1934). Type:— MEXICO, Guerrero: L. Schultze 345 (Holotype: at B destroyed). The determination of Q. obscurirhachis as probably Q. vicentensis is based on the Trelease (1934) description as no images or original specimens exist.</p> <p>Quercus siltepecana Matuda, Bol. Soc. Bot. México 18: 3 (1955). Type:— MEXICO, Chiapas: E. Matuda 19652 (Holotype: MEXU00085229!).</p> <p>Distribution: El Salvador, Mexico (Chiapas, Colima, Guerrero, Jalisco, México, Michoacán, Nayarit, Oaxaca).</p> <p>Quercus wislizeni A.DC., Prodr. 16(2.1): 67 (1864). Type:— U.S.A., California: F. A. Wislizenus s. n. (Nov 1851) (Holotype: MO-204635 (image)!). The holotype of Q. wislizeni was collected in November 1851 at the American Fork of the Sacramento River near Auburn. It was collected by Friedrich Adolph Wislizenus (1810–1889) and sent to A. de Candolle by G. Engelmann. W. Trelease, C. H. Muller, and K. C. Nixon all considered the MO specimen to be the holotype, and the specimen has label information in A. de Candolle’s hand. There is an inconsistency with the A. de Candolle publication which gives the collection locality as being from Chihuahua in the mountains of western Mexico in the vicinity of “Cosiquiriachi” (which is likely Cusihuiriachi) at an elevation of 7000 ft. Trelease (1924, p. 205) considered that A. de Candolle had erroneously ascribed that type to the Mexican state of Chihuahua. Certainly, the MO specimen is the species that occurs in California and only ranges into the Mexican state of Baja California.</p> <p>Distribution: Mexico (Baja California), U.S.A. (California).</p> <p>Quercus xalapensis Bonpl., Pl. Aequinoct. 2: 24, pl. 75 (1809). Type:— MEXICO, Veracruz: A. Bonpland 4467 (Holotype: P00129754! (as s.n.); isotypes: B-W!, P!, P!, UCSB (fragment)!). The holotype at P (s.n.) has a description on it in Bonpland’s hand. The other two specimens at P (s.n) as well as those with the number 4467 at P and elsewhere are all considered to be isotypes.</p> <p>Quercus acutifolia Neé var. angustifolia A.DC., Prodr. 16(2.1): 66 (1864). Type:— GUATEMALA: J. R. R. Warszewicz 6 (Lectotype (designated here): G00410062!). A. de Candolle designated two syntypes.</p> <p>Quercus grandis Liebm., Overs.Kongel.Danske Vidensk.Selsk.Forh.Medlemmers Arbeider1854:183 (1854). Type:— GUATEMALA: J. R. R. Warszewicz 20 (Holotype at B destroyed. Lectotype (designated here): K000512973!; isolectotypes: G!, P!).</p> <p>Quercus grandis var. tenuipes Trel., Mem. Natl. Acad. Sci. 20: 185, pl. 373a (1924). Type:— MEXICO, Puebla: H. Galeotti 135 (Holotype: G!; isotypes: P!, UCSB (fragment from G holotype)!).</p> <p>Quercus monserratensis C.H.Mull., U.S.D.A. Bur. Pl. Industr. Misc. Publ. 477: 71, pl. 104 (1942). Type:— MEXICO, Chiapas: C. A. Purpus 10091 (p.p.) (Holotype: A00055674!; isotype: UCSB (fragment from A holotype)!). Only the specimens of Purpus 10091 made in Apr 1927 are type collections of Q. monserratensis which is a synonym of Q. xalapensis. Purpus had used this same collection number two years earlier in May 1925 and at approximately the same locality. The 1925 collections are Q. segoviensis rather than Q. xalapensis and are not types.</p> <p>Quercus salvadorensis Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 8(1): 5 (1930). Type:— EL SALVADOR, Cuscatlán: S. Calderon 1981 (Holotype: US00089634!; isotypes: GH!, ILL!, NY!, UCSB (fragment)!).</p> <p>Quercus xalapensis f. jalapae Trel., Mem. Natl. Acad. Sci. 20: 193, pl. 393a (1924). Type:— MEXICO, Veracruz: C. J. W. Schiede 18 (p.p.) (Holotype at B destroyed. Lectotype (designated here): HAL0060347!; isolectotypes: HBG (image)!, MO!, UCSB (fragment)!). Only the collections of Schiede 18 (p.p.) from Jalapa are Q. xalapensis f. jalapa e. The sheets of Schiede 18 (p.p.) at G, and one of the two sheets at HAL are from Jalcingo and are the lectotype and an isolectotype of Q. affinis f. commutata.</p> <p>Distribution: El Salvador, Guatemala, Honduras, Mexico (Chiapas, Chihuahua, Colima, Durango, Guerrero, Hidalgo,</p> <p>Jalisco, México, Michoacán, Nayarit, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa, Sonora,</p> <p>Tamaulipas, Veracruz), Nicaragua. The occurrence of Q. xalapensis in Chihuahua, Colima, Durango, Nayarit, Sinaloa,</p> <p>and Sonora reported by Villaseñor (2016) needs to be confirmed and may involve differences in the interpretation of what should be included in this species.</p> <p>Quercus xylina Scheidw., Hort. Belge 4: 321, pl. 18 (1837). Type:— MEXICO, Jalisco: H. Galeotti 119 (Lectotype (designated here): BR0000006915674!; isolectotype: UCSB (fragment from BR holotype)!). The date of 1840 on Galeotti collections is not the actual collection date but rather the date the numbers were assigned back in Europe (McVaugh 1978). The actual collection date for what was assigned the number 119 in 1840 was probably 1836. Scheidweiler (1837) did not cite the exact specimen on which he was basing Q. xylina.</p> <p>Quercus jaliscensis Trel., Mem. Natl. Acad. Sci. 20: 86, pl. 132 (1924). Type:— MEXICO, Zacatecas: J. N. Rose 2685 (Holotype: US00089544!; isotypes: NY!, UCSB (fragment)!).</p> <p>Quercus laxa Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 181 (1854). Type:— MEXICO, Sinaloa: B. Seemann 1967 (Holotype: K000512629!; isotypes: BM!, C (as s.n.)!, GH (as s.n.)!, UCSB (as s.n.) (fragment)!).</p> <p>Quercus reticulata Bonpl. var. laxa (Liebm.) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 195 (1884).</p> <p>Distribution: Mexico (Ciudad de México, Colima, Durango, Guanajuato, Hidalgo, Jalisco, México, Michoacán,</p> <p>Nayarit, Querétaro, San Luis Potosí, Sinaloa, Zacatecas).</p></div> 	https://treatment.plazi.org/id/794B8792FFF1FFAD3E88FCD8FCDBF9A1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FF8EFFA33E88F90AFE4FFBA1.text	794B8792FF8EFFA33E88F90AFE4FFBA1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus tonduzii Seemen, Bull. Herb. Boissier	<div><p>Quercus × tonduzii Seemen, Bull. Herb. Boissier, sér. 2, 4(7): 656 (1904). Type:— COSTA RICA, Alajuela: A. Tonduz</p> <p>10788 (Holotype: BR!; isotypes: A!, G!, M!, P (as s.n.)!, UCSB (fragment)!, US!). The type collection apparently involves hybridization between Q. benthamii and Q. costaricensis. Quercus seemannii Liebm. subsp. tonduzii (Seemen) A.E.Murray, Kalmia 13: 27 (1983). Quercus seemannii var. tonduzii (Seemen) A.E.Murray, Kalmia 13: 27 (1983). Distribution: Costa Rica.</p> <p>Quercus × undata Trel., Mem. Natl. Acad. Sci. 20: 86, pl 135 (1924). Type:— MEXICO, Durango: R. Endlich 1 (Holotype at B destroyed. Lectotype: image in Trelease (1924, pl. 135)! designated by Bacon et al. (2011)). No extant original material exists, so lectotypification of the name was based on the image of the holotype. This putative hybrid species is believed to involve introgression between Q. chihuahuensis, Q. grisea and Q. arizonica (Bacon et al. 2011).</p> <p>Distribution: Mexico (Durango).</p> <p>Quercus × warburgii A.Camus, Chênes Texte 2: 621, 793 (1939). Nom. nov. for Q. hartwegii f. glabra Trel. (1924) non Q. glabra Thunb. (1784). This is a cultivated hybrid between the Mexican species Q. rugosa and possibly the European species Q. rubur L.</p> <p>Quercus hartwegii Benth. f. glabrata Trel., Mem. Natl. Acad. Sci. 20: 70 (1924). Type:—(Cult.): A. Henry 1 (Lectotype (designated here): ILL00007542!; isolectotype: ILL!). According to Trelease (1924, p. 70) “A quickly glabrate form, f. glabrata, from which the catkins are described, is cultivated at Cambridge Botanical Garden in England (Henry) from acorns said to have been collected by Hartweg. ” There is no indication on either of the ILL specimens indicating which one Trelease considered the holotype.</p> <p>Distribution: Cultivated plant in the United Kingdom.</p> <p>APPENDIX 3. Distribution of recognized species of Fagaceae in Latin America by country and state for Mexico. APPENDIX4. Quercus interspecific hybrids and putative hybrids based on more than 19,000 Latin American collections examined by Breedlove. For hybrid collections that are listed by formula rather than as a named nothospecies, the first putative parent is the one to which the hybrid is morphologically closest. For hybrids that are the types of named hybrid species see Appendix 2.</p> </div>	https://treatment.plazi.org/id/794B8792FF8EFFA33E88F90AFE4FFBA1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FFACFF8D3E88FF66FD30FAC8.text	794B8792FFACFF8D3E88FF66FD30FAC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quercus urbani Trel.	<div><p>Quercus urbani Trel.</p> <p>Quercus urbani f. parvifolia Martínez = Quercus conzattii Trel.</p> <p>Quercus uruapanensis Trel. = Quercus tlapuxahuensis A.DC.</p> <p>Quercus utahensis (A.DC.) Rydberg = Quercus gambelii Nutt.</p> <p>Quercus uxoris McVaugh</p> <p>Quercus vallicola Trel. (1924) non Gand. (1890) = Quercus frutex Trel.</p> <p>Quercus varians M.Martens &amp; Galeotti = Quercus polymorpha Schltdl. &amp; Cham.</p> <p>Quercus vaseyana Buckley = Quercus pungens Liebm.</p> <p>Quercus vellifera Trel. = Quercus spicata Bonpl.</p> <p>Quercus verde C.H.Mull. = Quercus spicata Bonpl.</p> <p>Quercus verrucosirama Trel. = Quercus castanea Neé</p> <p>Quercus vexans Trel. = Quercus sartorii Liebm.</p> <p>Quercus vicentensis Trel.</p> <p>Quercus viminea Trel. = Quercus devia Goldman</p> <p>Quercus virginiana Mill. subsp. fusiformis (Small) A.E.Murray = Quercus fusiformis Small</p> <p>Quercus virginiana var. fusiformis (Small) Sarg. = Quercus fusiformis Small</p> <p>Quercus virginiana var. sagrana (Nutt.) Trel. = Quercus sagrana Nutt.</p> <p>Quercus × warburgii A.Camus</p> <p>Quercus warscewiczii Liebm. = Quercus insignis M.Martens &amp; Galeotti</p> <p>Quercus wesmaelii Trel. = Quercus costaricensis Liebm.</p> <p>Quercus wislizeni A.DC.</p> <p>Quercus xalapensis Bonpl.</p> <p>Quercus xalapensis f. jalapae Trel. = Quercus xalapensis Bonpl.</p> <p>Quercus xalapensis f. surculina Trel. = Quercus grahamii Benth.</p> <p>Quercus xalapensis var. longifolia (Liebm.) Wenz. = Quercus laurina Bonpl.</p> <p>Quercus xylina Scheidw.</p> <p>Quercus yoroensis Trel. = Quercus elliptica Neé</p> <p>Quercus yoroensis var. aguanana Trel. = Quercus elliptica Neé</p> <p>Quercus yousei Trel. = Quercus lancifolia Schltdl. &amp; Cham.</p> <p>Quercus yunckeri Trel. = Quercus insignis M.Martens &amp; Galeotti</p> <p>Quercus zempoaltepecana Trel. = Quercus laurina Bonpl.</p> <p>Trigonobalanus excelsa Lozano, Hern.Cam. &amp; Henao</p></div> 	https://treatment.plazi.org/id/794B8792FFACFF8D3E88FF66FD30FAC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
794B8792FF8CFFAF3E88F952FDF1F919.text	794B8792FF8CFFAF3E88F952FDF1F919.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trigonobalanus Forman	<div><p>TRIGONOBALANUS Forman</p> <p>Trigonobalanus excelsa Lozano, Hern.Cam. &amp; Henao, Caldasia 12(60): 519 (1979). Type:— COLOMBIA, Huila: J. E. Henao S. et al. s. n. (Jan 1979) (Holotype: COL000000754 (image)!; isotype: COL (image)!). We here follow Kubitzki (1993), Govaerts &amp; Frodin (1998), Christenhusz et al. (2017), and The World Flora Online (2022) in accepting a broadly circumscribed Trigonobalanus of three species that includes Colombobalanus.</p> <p>Colombobalanus excelsa (Lozano, Hern.Cam. &amp; Henao) Nixon &amp; Crepet, Amer. J. Bot. 76(6): 841 (1989).</p> <p>Distribution: Colombia.</p> <p>EXCLUDED NAMES</p> <p>Quercus candicans Neé, Anales Ci. Nat. 3: 277 (1801). Type:— MEXICO, Guerrero: L. Neé s. n. (1791) (Lectotype: MA25948! designated by Valencia-A. et al. (2018, p. 256); isolectotypes: MA!, UCSB (fragment)!). Valencia- A. et al. (2018) determined that the lectotype of Q. candicans is actually a Roldana La Llave (Asteraceae). It is possible that the specimens at BM and G as well as the fragments at G and UCSB, all of which are Quercus, may be original material used in part by Neé for his description of Q. candicans rather than the specimen now at MA (Bartholomew &amp; Almeda 2022). Because the leaf description by Neé seems to agree with that for a Roldana, the mixup in the collection may have been done by Neé himself.</p> <p>Quercus dawsonii Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 17(4): 357 (1938). Type:— HONDURAS, Comayagua: T. G. Yuncker et al. 6378 (Holotype: ILL00007514!; isotypes: GH!, K!, MO!, UCSB (fragment)!). This type collection is probably a Meliosma (Sabiaceae).</p> <p>Quercus pertessellata Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 17(4): 357 (1938). Type:— HONDURAS, Comayagua: T. G. Yuncker et al. 6376 (Holotype: ILL00007607!; isotypes: GH!, MO!, NY!, UCSB (fragment)!). The type is not a Quercus but rather Billia hippocastanum Peyr. (Sapindaceae) determined by P.S. Green in Feb 1999.</p> <p>Quercus ravenscroftiana A. Murray, Gard. Chron. 1866: 1018 (1866). The only Quercus in South America is Q. humboldtii which occurs only in Colombia, and the illustration in Gard. Chron. is not that species. The description is of a sterile specimen, and there are no known extant specimens. It is quite likely that this is not even a Quercus.</p> <p>APPENDIX 2. Latin American named hybrid taxa. See also Appendix 4 for non-type collections that probably involve hybridization.</p> <p>Quercus × acutidens Torr., Rep. U.S. Mexico Bound., Bot. 2(1): 207, pl. 51 (1859). U.S.A. (California). C. C. Parry s. n. (1850) (Holotype NY002486802 (image)!; isotype NY (as image)!). The type collection is believed to involve hybridization between Q. cornelius-mulleri and Q. engelmannii (Nixon 1997, Govaerts &amp; Frodin 1998).</p> <p>Quercus dumosa Nutt. var. acutidens (Torr.) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 204 (1884).</p> <p>Quercus dumosa f. linearis Trel., Mem. Natl. Acad. Sci. 20: 117, pl. 205b (1924). Type:— MEXICO, Baja California: T. Brandegee s. n. (21 Apr 1893) (Holotype: A00055678!). The type collection of this forma is apparently a hybrid with the same parentage as Q. × acutidens.</p> <p>Distribution: Mexico (Baja California), U.S.A. (California).</p> <p>Quercus × aerea Trel., Mem. Natl. Acad. Sci. 20: 135, pl. 257 (1924). Type:— MEXICO, Chihuahua: C. G. Pringle 1570 (Holotype: GH00055630!; isotype: A!). The type collection apparently involves hybridization between Q. jonesii and Q. emoryi.</p> <p>Quercus × knoblochii C.H.Mull., Amer. Midl. Naturalist 27(2): 475 (1942). Type:— MEXICO, Chihuahua: C. H. Muller 3600 (Holotype: US00089542!; isotypes: GH!, LL (image)!, UC!, UCSB (fragment)!). This hybrid epithet apparently involves the same parentage as Q. × aerea.</p> <p>Distribution: Mexico (Chihuahua, Durango, Sonora).</p> <p>Quercus × apiophylla E.F.Warb., Bull. Misc. Inform. Kew 1939 (2): 95 (1939). Type:— MEXICO, México: G. B. Hinton 6568 (Lectotype (designated here): K000512997!; isolectotypes: A!, BM!, CAS!, F!, G!, K!, NY!, UCSB (fragment)!). There are two specimens at K, both of which are annotated by E.F.Warburg as “Type.” This may be a hybrid between Q. calophylla and Q. acutifolia or Q. grahamii.</p> <p>Distribution: Mexico (México)</p> <p>Quercus × basaseachicensis C.H.Mull., Amer. Midl. Naturalist 19(3): 582 (1938). Type:— MEXICO, Chihuahua: H. LeSueur 549 (Holotype: BH; isotypes: GH!, MO!, TEX (image)!, UC!). This species has been considered to be of hybrid origin between Q. depressipes and Q. rugosa (Govaerts &amp; Frodin 1998, Villaseñor 2016). The type collection is similar to Q. emoryi, and additional study is necessary to clarify its hybrid status.</p> <p>Distribution: Mexico (Chihuahua, Sonora).</p> <p>Quercus × duratifolia C.H.Mull., U.S.D.A. Bur. Pl. Industr. Misc. Publ. 477: 50, pl. 66, 67 (1942). Type:— MEXICO, Chiapas: E. Matuda 1827 (Holotype: A00055572!; isotypes: K!, MEXU!, MICH!, MO!, NY!, UCSB (fragment)!, VT!). The type collection apparently involves hybridization between Q. sapotifolia and Q. salicifolia.</p> <p>Distribution: El Salvador, Mexico (Chiapas).</p> <p>Quercus × galeottii M.Martens, Bull. Acad. Roy. Sci. Bruxelles 10(1): 220 (1843). Type:— MEXICO, Veracruz: H. Galeotti 126 (Lectotype: BR0000006912567! designated by Trelease (1924, p. 216, pl. 25); isolectotypes: BR!, G!, L!, LE!, P!, UCSB (fragment)!). Although M.Martens &amp; Galeotti cite only one collection number, two sheets of which are at BR and marked as types; one of which was designated by Trelease as “The type.” The type collections apparently involve hybridization between Q. lancifolia and Q. insignis.</p> <p>Distribution: Costa Rica, Mexico (Chiapas, Veracruz), Panama.</p> <p>Quercus × graciliramis C.H.Mull., J. Arnold Arbor. 17(3): 177 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 1106 (Holotype:A00055659!; isotypes: BM!, G!, MEXU!, NY!, P!, TEX (image)!, UCSB (fragment)!, US!). The type collection apparently involves hybridization between Q. gravesii and Q. xalapensis.</p> <p>Distribution: Mexico (Coahuila, Nuevo León).</p> <p>Quercus × manzanillana Trel., Mem. Natl. Acad. Sci. 20: 80, pl. 116a (1924). Type:— MEXICO, Puebla: G. Arsène 5925 (Holotype: US 00089560!; isotypes: ILL!, MPU!). The type collection apparently involves hybridization between Q. diversifolia and Q. laeta.</p> <p>Distribution: Mexico (Puebla, Querétaro).</p> <p>Quercus × pastorensis C.H.Mull., J. Arnold Arbor. 17(3): 164 (1936). Type:— MEXICO, Nuevo León: C. H. Muller 1296 (Holotype: A00055618!; isotype: ILL!). This has been considered to be a hybrid between Q. glaucophylla and Q. laeta (Muller 1936) or between Q. laeta and Q. glaucoides (Gavaerts &amp; Frodin 1998), but according to Breedlove the parentage apprarently involves hybridization between Q. laceyi and Q. laeta.</p> <p>Distribution: Mexico (Nuevo León).</p> <p>Quercus × saltillensis Trel., Mem. Natl. Acad. Sci. 20: 183, pl. 368 (1924). Type:— MEXICO, Coahuila: E. Palmer 1277 (Holotype: GH00069924!; isotypes: A!, K!, MO!, NY!, US!). The type collection apparently involves hybridization between Q. depressa and Q. mexicana.</p> <p>Distribution: Mexico (Coahuila, Nuevo León).</p> <p>Quercus × sierramadrensis C.H.Mull., Amer. Midl. Naturalist 27(2): 478 (1942). Nom. nov. for Q. tenuiloba C.H. Mull. f. gracilis C.H. Mull. (1936) non Q. gracilis Korth. (1844).</p> <p>Quercus × tenuiloba nothof. gracilis C.H.Mull., J. Arnold Arbor. 17(3): 178 (1936). Type:— MEXICO, Nuevo León: C. H. Muller 2048 (Holotype: A00055646!; isotypes: CAS!, MO!, NY!, UCSB (fragment)!). The type collection apparently involves hybridization between Q. rysophylla and Q. canbyi.</p> <p>Distribution: Mexico (Nuevo León).</p> <p>Quercus × tenuiloba C.H.Mull., J. Arnold Arbor. 17(3): 178 (1936). Type:— MEXICO, Nuevo León: C. H. Muller &amp; M. T. Muller 1125 (Holotype: A00055645!; isotypes: BM!, G!, ILL!, MO!, NY!, P!, PH!, UCSB (fragment)!, US!). The type collection apparently involves hybridization between Q. rysophylla and Q. xalapensis.</p> <p>Distribution: Mexico (Nuevo León).</p></div> 	https://treatment.plazi.org/id/794B8792FF8CFFAF3E88F952FDF1F919	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bartholomew, Bruce;Almeda, Frank	Bartholomew, Bruce, Almeda, Frank (2023): Nomenclator botanicus of Fagaceae in Latin America. Phytotaxa 580 (1): 1921-1935, DOI: 10.11646/phytotaxa.580.1.1, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
