taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
7409957DB317B81FD2AB7603FDAFFEFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15263184/files/figure.png	https://doi.org/10.5281/zenodo.15263184	FIGURE 2. Maximum Likelihood (ML) phylogenetic tree of Trimeresurus derived from the analysis of 2,402 bp of cyt b, ND4, and 16S rRNA mitochondrial gene fragments. For voucher specimen information and GenBank accession numbers, see Appendix II. Numbers at the nodes correspond to ML UFBS/BI PP support values, respectively. Colors of clades and locality numbers correspond to those in Figure 1. Photographs by P. Pawangkhanant and T. Woranuch.	FIGURE 2. Maximum Likelihood (ML) phylogenetic tree of Trimeresurus derived from the analysis of 2,402 bp of cyt b, ND4, and 16S rRNA mitochondrial gene fragments. For voucher specimen information and GenBank accession numbers, see Appendix II. Numbers at the nodes correspond to ML UFBS/BI PP support values, respectively. Colors of clades and locality numbers correspond to those in Figure 1. Photographs by P. Pawangkhanant and T. Woranuch.	2025-04-22	Pawangkhanant, Parinya;Idiiatullina, Sabira S.;Nguyen, Tan Van;Ruangsuwan, Thiti;Matsukoji, Tomoya;David, Patrick;Suwannapoom, Chatmongkon;Poyarkov, Nikolay A.		Zenodo	biologists	Pawangkhanant, Parinya;Idiiatullina, Sabira S.;Nguyen, Tan Van;Ruangsuwan, Thiti;Matsukoji, Tomoya;David, Patrick;Suwannapoom, Chatmongkon;Poyarkov, Nikolay A.			
7409957DB317B81FD2AB7603FDAFFEFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15263188/files/figure.png	https://doi.org/10.5281/zenodo.15263188	FIGURE 3. The holotype of Trimeresurus cryptographicus sp. nov. in preservative (ZMMU Re-17934, adult female). A dorsolateral and B ventral view of body; C dorsal, D ventral, E lateral right, and F lateral left aspects of head. Scale bar = 1 cm. Photographs by S. S. Idiiatullina.	FIGURE 3. The holotype of Trimeresurus cryptographicus sp. nov. in preservative (ZMMU Re-17934, adult female). A dorsolateral and B ventral view of body; C dorsal, D ventral, E lateral right, and F lateral left aspects of head. Scale bar = 1 cm. Photographs by S. S. Idiiatullina.	2025-04-22	Pawangkhanant, Parinya;Idiiatullina, Sabira S.;Nguyen, Tan Van;Ruangsuwan, Thiti;Matsukoji, Tomoya;David, Patrick;Suwannapoom, Chatmongkon;Poyarkov, Nikolay A.		Zenodo	biologists	Pawangkhanant, Parinya;Idiiatullina, Sabira S.;Nguyen, Tan Van;Ruangsuwan, Thiti;Matsukoji, Tomoya;David, Patrick;Suwannapoom, Chatmongkon;Poyarkov, Nikolay A.			
7409957DB317B81FD2AB7603FDAFFEFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15263190/files/figure.png	https://doi.org/10.5281/zenodo.15263190	FIGURE 4. Habitat of Trimeresurus cryptographicus sp. nov. A microhabitat of the new species in Tham Kang Kao Cave, Phitsanulok, Thailand; B adult female, holotype (ZMMU Re-17934), in life in situ. Photographs by P. Pawangkhanant.	FIGURE 4. Habitat of Trimeresurus cryptographicus sp. nov. A microhabitat of the new species in Tham Kang Kao Cave, Phitsanulok, Thailand; B adult female, holotype (ZMMU Re-17934), in life in situ. Photographs by P. Pawangkhanant.	2025-04-22	Pawangkhanant, Parinya;Idiiatullina, Sabira S.;Nguyen, Tan Van;Ruangsuwan, Thiti;Matsukoji, Tomoya;David, Patrick;Suwannapoom, Chatmongkon;Poyarkov, Nikolay A.		Zenodo	biologists	Pawangkhanant, Parinya;Idiiatullina, Sabira S.;Nguyen, Tan Van;Ruangsuwan, Thiti;Matsukoji, Tomoya;David, Patrick;Suwannapoom, Chatmongkon;Poyarkov, Nikolay A.			
7409957DB317B81FD2AB7603FDAFFEFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15263192/files/figure.png	https://doi.org/10.5281/zenodo.15263192	FIGURE 5. A An everted right hemipenis of Trimeresurus cryptographicus sp. nov. (RIM-00106) B Everted hemipenes of T. kuiburi (RIM-00107). Photographs by P. Pawangkhanant (A) and T. Ruangsuwan (B).	FIGURE 5. A An everted right hemipenis of Trimeresurus cryptographicus sp. nov. (RIM-00106) B Everted hemipenes of T. kuiburi (RIM-00107). Photographs by P. Pawangkhanant (A) and T. Ruangsuwan (B).	2025-04-22	Pawangkhanant, Parinya;Idiiatullina, Sabira S.;Nguyen, Tan Van;Ruangsuwan, Thiti;Matsukoji, Tomoya;David, Patrick;Suwannapoom, Chatmongkon;Poyarkov, Nikolay A.		Zenodo	biologists	Pawangkhanant, Parinya;Idiiatullina, Sabira S.;Nguyen, Tan Van;Ruangsuwan, Thiti;Matsukoji, Tomoya;David, Patrick;Suwannapoom, Chatmongkon;Poyarkov, Nikolay A.			
7409957DB317B81FD2AB7603FDAFFEFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15263194/files/figure.png	https://doi.org/10.5281/zenodo.15263194	FIGURE 6. Trimeresurus cryptographicus sp. nov. in life from the type locality—Tham Kang Kao Cave, Phitsanulok, Thailand. A–B neonates, sex unkown, released; C–D juvenile male, released; E adult male, paratype (RIM-00056); F adult female, paratype (RIM-00055). Photographs by R. Bilgin (A–D), P. Pawangkhanant (E), and G. Vogel (F).	FIGURE 6. Trimeresurus cryptographicus sp. nov. in life from the type locality—Tham Kang Kao Cave, Phitsanulok, Thailand. A–B neonates, sex unkown, released; C–D juvenile male, released; E adult male, paratype (RIM-00056); F adult female, paratype (RIM-00055). Photographs by R. Bilgin (A–D), P. Pawangkhanant (E), and G. Vogel (F).	2025-04-22	Pawangkhanant, Parinya;Idiiatullina, Sabira S.;Nguyen, Tan Van;Ruangsuwan, Thiti;Matsukoji, Tomoya;David, Patrick;Suwannapoom, Chatmongkon;Poyarkov, Nikolay A.		Zenodo	biologists	Pawangkhanant, Parinya;Idiiatullina, Sabira S.;Nguyen, Tan Van;Ruangsuwan, Thiti;Matsukoji, Tomoya;David, Patrick;Suwannapoom, Chatmongkon;Poyarkov, Nikolay A.			
7409957DB317B81FD2AB7603FDAFFEFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15263196/files/figure.png	https://doi.org/10.5281/zenodo.15263196	FIGURE 7. All known species of the Trimeresurus kanburiensis species group, in life: A T. cryptographicus sp. nov. from Tham Kang Kao Cave, Phitsanulok, Thailand; B T. cardamomensis from Khao Soidao WS., Chanthaburi, Thailand; C T. cyanolabris from Phuoc Binh NP., Ninh Thuan, Vietnam; D T. erythrochloris from Khlong Hat, Sakeo, Thailand; E T. macrops from Tha Luang, Lop Buri, Thailand; F T. rubeus from Cat Tien NP., Dong Nai, Vietnam; G T. ciliaris from Wang Prachan, Satun, Thailand; H T. honsonensis from Hon Son Is., Kien Giang, Vietnam; I T. kanburiensis from Sai Yok NP., Kanchanaburi, Thailand; J T. kraensis from Tham Sanook Cave, Chumphon, Thailand; K T. kuiburi from Sam Roi Yot, Prachuap Khiri Khan, Thailand; L T. venustus from Khao Luang NP., Nakhon Si Thammarat, Thailand. Photographs by P. Pawangkhanant (A), A. Kaosung (B, E, I, J), N. A. Poyarkov (C, K), T. Smith (D), G. Vogel (F, G), L. P. Tran (H), R. Jaihan (L).	FIGURE 7. All known species of the Trimeresurus kanburiensis species group, in life: A T. cryptographicus sp. nov. from Tham Kang Kao Cave, Phitsanulok, Thailand; B T. cardamomensis from Khao Soidao WS., Chanthaburi, Thailand; C T. cyanolabris from Phuoc Binh NP., Ninh Thuan, Vietnam; D T. erythrochloris from Khlong Hat, Sakeo, Thailand; E T. macrops from Tha Luang, Lop Buri, Thailand; F T. rubeus from Cat Tien NP., Dong Nai, Vietnam; G T. ciliaris from Wang Prachan, Satun, Thailand; H T. honsonensis from Hon Son Is., Kien Giang, Vietnam; I T. kanburiensis from Sai Yok NP., Kanchanaburi, Thailand; J T. kraensis from Tham Sanook Cave, Chumphon, Thailand; K T. kuiburi from Sam Roi Yot, Prachuap Khiri Khan, Thailand; L T. venustus from Khao Luang NP., Nakhon Si Thammarat, Thailand. Photographs by P. Pawangkhanant (A), A. Kaosung (B, E, I, J), N. A. Poyarkov (C, K), T. Smith (D), G. Vogel (F, G), L. P. Tran (H), R. Jaihan (L).	2025-04-22	Pawangkhanant, Parinya;Idiiatullina, Sabira S.;Nguyen, Tan Van;Ruangsuwan, Thiti;Matsukoji, Tomoya;David, Patrick;Suwannapoom, Chatmongkon;Poyarkov, Nikolay A.		Zenodo	biologists	Pawangkhanant, Parinya;Idiiatullina, Sabira S.;Nguyen, Tan Van;Ruangsuwan, Thiti;Matsukoji, Tomoya;David, Patrick;Suwannapoom, Chatmongkon;Poyarkov, Nikolay A.			
7409957DB317B81FD2AB7603FDAFFEFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15263182/files/figure.png	https://doi.org/10.5281/zenodo.15263182	FIGURE 1. Distribution of the members of the Trimeresurus kanburiensis species group (including members of the former T. macrops species complex) in Indochina region. Stars denote type localities of each species following by numbers of localities. T. cryptographicus sp. nov. (magenta, 1): Thailand, Phitsanulok Prov., Noen Maprang Dist., Tham Kang Kao Cave; T. kanburiensis (orange, 2): Thailand, Kanchanaburi Prov., Sai Yok Dist., Wat Tham, Phom Lo Khao Yai; T. macrops (neon green, 3): Thailand, Bangkok; T. erythrochloris (red, 4): Thailand, Sa Kaeo Prov., Klong Hat Dist., Tham Khao Siwa Cave; T. cardamomensis (yellow, 5): Cambodia, Koh Kong Prov., Sre Ambel Dist., Cardamom Mountains; T. rubeus (brown, 6): Cambodia, Mondolkiri Prov., O’Rang Dist., Seima Biodiversity Conservation Area; T. cyanolabris (light blue, 7): Vietnam, Ninh Thuan Prov., Ninh Hai Dist., Nui Chua N. P.; T. honsonensis (black, 8): Vietnam, Kien Giang Prov., Hon Son Island; T. kuiburi (dark blue, 9): Thailand, Prachuap Khiri Khan Prov., Kuiburi Dist., Wat Khao Daeng; T. kraensis (purple, 10): Thailand, Chumphon Prov., Wat Tham Sanook; T. venustus (light pink, 11): Thailand, Krabi Prov., Mueang Krabi Dist., Tiger Cave viewpoint; T. ciliaris (dark green, 12): Thailand, Trang Prov., Palian Dist., Thum Khao Ting. Question marks in neon green indicate localities where the distribution of T. macrops sensu stricto is anticipated.	FIGURE 1. Distribution of the members of the Trimeresurus kanburiensis species group (including members of the former T. macrops species complex) in Indochina region. Stars denote type localities of each species following by numbers of localities. T. cryptographicus sp. nov. (magenta, 1): Thailand, Phitsanulok Prov., Noen Maprang Dist., Tham Kang Kao Cave; T. kanburiensis (orange, 2): Thailand, Kanchanaburi Prov., Sai Yok Dist., Wat Tham, Phom Lo Khao Yai; T. macrops (neon green, 3): Thailand, Bangkok; T. erythrochloris (red, 4): Thailand, Sa Kaeo Prov., Klong Hat Dist., Tham Khao Siwa Cave; T. cardamomensis (yellow, 5): Cambodia, Koh Kong Prov., Sre Ambel Dist., Cardamom Mountains; T. rubeus (brown, 6): Cambodia, Mondolkiri Prov., O’Rang Dist., Seima Biodiversity Conservation Area; T. cyanolabris (light blue, 7): Vietnam, Ninh Thuan Prov., Ninh Hai Dist., Nui Chua N. P.; T. honsonensis (black, 8): Vietnam, Kien Giang Prov., Hon Son Island; T. kuiburi (dark blue, 9): Thailand, Prachuap Khiri Khan Prov., Kuiburi Dist., Wat Khao Daeng; T. kraensis (purple, 10): Thailand, Chumphon Prov., Wat Tham Sanook; T. venustus (light pink, 11): Thailand, Krabi Prov., Mueang Krabi Dist., Tiger Cave viewpoint; T. ciliaris (dark green, 12): Thailand, Trang Prov., Palian Dist., Thum Khao Ting. Question marks in neon green indicate localities where the distribution of T. macrops sensu stricto is anticipated.	2025-04-22	Pawangkhanant, Parinya;Idiiatullina, Sabira S.;Nguyen, Tan Van;Ruangsuwan, Thiti;Matsukoji, Tomoya;David, Patrick;Suwannapoom, Chatmongkon;Poyarkov, Nikolay A.		Zenodo	biologists	Pawangkhanant, Parinya;Idiiatullina, Sabira S.;Nguyen, Tan Van;Ruangsuwan, Thiti;Matsukoji, Tomoya;David, Patrick;Suwannapoom, Chatmongkon;Poyarkov, Nikolay A.			
