taxonID	type	description	language	source
7617FA4FFFB6926DFF25F9077A7BFACB.taxon	materials_examined	Type material. Cingula tumidula: holotype (Fig. 1 D), ZMO D 659, single dried specimen damaged by acidic glass; Cingula griegi — lectotype (Fig. 1 G) designated by Warén (1973) and two paralectotypes, all with number ZMB 20857; Punctullum minutum — holotype (Fig. 1 H), ZIN 35666 / 1 and 81 specimens labeled as “ paratypes ” (1 ♀ dissected), ZIN 35667 / 2, however the paratypes are not mentioned in the original description and therefore these specimens cannot be included in the type series. Type localities. Cingula tumidula: littoral zone at Vardø, Northern Norway; Rissoa griegi: Vöringen, st. 353 (77 ° 58 ' N, 05 ° 10 ' E, 2438 m); Punctullum minutum: White Sea, Kandalaksha Bay, 50 m. Other examined material: Southwestern Barents Sea: 1 ♂ (dissected), Malaya Sharkovka Bay, 27 m, 69 ° 12.57 ' N, 34 ° 54.31 ' E, 20 Sept. 2012, MS VIKING- 2, MMBI, no number; 1 ♂ (dissected), Bolshaya Sharkovka Bay, 25 m, 69 ° 12.59 ' N, 34 ° 56.45 ' E, 20 Sept. 2012, MS VIKING- 2, MMBI, no number; 1 spc, Zavalishina Bay, 12 m, 69 ° 11.38 ' N, 35 ° 14.78 ' E, 10 Oct. 2010, MMBI, no number; 2 spc, Orlovka Bay, 23 m, 69 ° 12.35 ' N, 35 ° 16.00 ' E, 11 Oct. 2010, MMBI, no number; 1 spc, Korabelnaya Bay, 11 m, 69 ° 10.22 ' N, 35 ° 08.79 ' E, 10 Oct. 2010, MMBI, no number; 6 spcs (3 ♀ dissected), Yarnishnaya Inlet, 67 m, 69 ° 08.71 ' N, 36 ° 00.44 ' E, 2 June 2009, RV DALNIE ZELENTSY, MMBI, no number; 1 spc, 73 m, 69 ° 07.64 ' N, 36 ° 02.01 ' E, 19 Sept. 2012, MS VIKING- 2; 9 spc (1 ♀ dissected), Dalne-Zelenetskaya Bay, 59 m, 69 ° 07.92 ' N, 36 ° 05.45 ' E, 4 June 2009, RV DALNIE ZELENTSY, MMBI, no number; Eastern Barents Sea (Pechora Sea): 2 spc, 17.5 m, 69 ° 15.61 ' N, 57 ° 22.76 ' E, 15 Nov. 2010, RV BUJNITSKIY, MMBI, no number; 1 spc, 20 m, 69 ° 23.15 ' N, 57 ° 44.62 ' E, 15 Nov. 2010, RV BUJNITSKIY, MMBI, no number; Franz Joseph Land: 13 spcs (2 ♀ and 1 ♂ dissected), 44 m, 79 º 53.4 ' N, 51 º 26.06 ' E, 24 Aug. 2006, RV DALNIE ZELENTSY, MMBI, no number; White Sea: 23 spcs. (1 ♂ and 1 ♀ dissected), 85 m, 66 ° 20 ' N, 30 ° E, 8 June 1926 RV PERSEY, ZIN 35668 / 3; 2 spcs, NE Kara Sea, 1937 RV SEDOV, ZIN 33631 / 2. Shell. (Fig 1 A – H) Ovate-conical, small, white, usually semitransparent, covered by thin transparent periostracum. Up to 3.5 total whorls, well rounded with deep sutures. Sculpture of adult shell consists of slightly prosocline growth lines and thin, numerous (30 or more on body whorl) spiral riblets. Some specimens form wide costae usually limited to the upper parts of whorls. Protoconch (Fig. 2 A – D) of 1 – 1.2 whorls, 420 – 570 µm in diameter, with nucleus of about 130 – 200 µm; larval whorls with more frequent and delicate spiral striae than those on teleoconch. Aperture oval, slightly angulated in upper part. Umbilicus relatively wide, partly covered by edge of lip. Operculum concave, transparent, without peg. Mean values of morphometric characters of 22 adult measured specimens: shell height = 2 ± 0.04 mm, shell width = 1.43 ± 0.03 mm, aperture height = 1.09 ± 0.05 mm, aperture width = 0.74 ± 0.02 mm, last whorl height = 1.37 ± 0.05 mm, shell whorls number = 3.3. The largest measured specimen had shell height = 2.35 mm, shell width = 1.58 mm, aperture height = 0.98 mm, aperture width = 0.80 mm, last whorl height = 1.73 mm, whorls number = 3.5. Radula. (Fig 2 E – F) Rachidian tooth relatively wide with prominent central cusp and 5 lateral denticles; single pairs of both basal and lateral processes are present; U-shaped ventral extension well developed. Lateral teeth with one prominent cusp and 4 to 5 smaller cusps on each side. Both inner and outer marginal teeth elongated with relatively similar-sized denticles.	en	Nekhaev, Ivan O. (2016): Morphology and systematic position of Cingula tumidula G. O. Sars, 1878 (Gastropoda: Rissoidae). Zootaxa 4184 (2): 395-400, DOI: 10.11646/zootaxa.4184.2.13
7617FA4FFFB6926DFF25F9077A7BFACB.taxon	description	Animal. Mantle not pigmented. Cephalic tentacles slightly longer than bilobed snout in preserved state (Fig. 3 E). Small anterior pallial tentacle on the right side and single metapodial tentacle present. Ctenidium elongated, filaments absent osphradium shorter than ctenidium. Male reproductive system. (Fig. 3 E) Penis (PE) flattened, almost parallel-sided, with acute distal end; duct (SD) submarginal. Prostate gland large, approximately twice as long as wide. Female reproductive system. (Fig. 3 A – D) Ventral channel marked, vestibule not developed. Lower oviduct gland (LOG) longer than high. Bursa copulatrix (BC) relatively large, about half size of lower oviduct gland; seminal receptacle (SR) small, elongated, not found in all studied specimens. Upper oviduct gland (UOG) coiled, usually short lobes (L) visible (these probably involved in sperm storage). Digestive tract: contained intact calcareous foraminiferans, detritus and sand.	en	Nekhaev, Ivan O. (2016): Morphology and systematic position of Cingula tumidula G. O. Sars, 1878 (Gastropoda: Rissoidae). Zootaxa 4184 (2): 395-400, DOI: 10.11646/zootaxa.4184.2.13
7617FA4FFFB6926DFF25F9077A7BFACB.taxon	distribution	Distribution. Pusillina tumidula has been reported from Iceland, and Faeroe, Norwegian, Barents, White and Kara Seas including the waters around Svalbard and Franz Joseph Land (Golikov 1987; Warén 1996; Golikov et al. 2001; Sneli et al. 2005; Nekhaev 2014). Study of ZIN collections shows that records of that species from the Bering Sea (Kantor & Sysoev 2006; Sirenko et al. 2013) are based on mis-identification of the another rissoid species. Habitat. Found on substrates with sand, shell rock and stones, occasionally with silt and clay. Several samples of Pusillina tumidula were taken from water as shallow 11 – 12 m, however the holotype of Cingula tumidula is the only known specimen of the species collected in the intertidal zone. The lower reported distributional limit is 500 m (Warén 1996).	en	Nekhaev, Ivan O. (2016): Morphology and systematic position of Cingula tumidula G. O. Sars, 1878 (Gastropoda: Rissoidae). Zootaxa 4184 (2): 395-400, DOI: 10.11646/zootaxa.4184.2.13
7617FA4FFFB6926DFF25F9077A7BFACB.taxon	discussion	Remarks. Both the reproductive system and the radula of Cingula tumidula differ significantly from that of Obtusella intersecta (Wood, 1857), which is the type species of Obtusella: the females of O. intersecta have a welldeveloped vestibule modified for sperm storage, and the seminal receptacle is placed perpendicular to the bursa copulatrix (Ponder 1984; Nekhaev 2015). Males have a long penis which is oval in section and with a long, prominent filament. The central tooth of Obtusella intersecta has two pairs of basal denticles. The adult shell of Obtusella intersecta is similar to that of Cingula tumidula in shape and sculpture, but the protoconch of the former species is multiwhorled with very weak spiral striae (Ponder 1984; Warén 1989), not like that of Cingula tumidula.	en	Nekhaev, Ivan O. (2016): Morphology and systematic position of Cingula tumidula G. O. Sars, 1878 (Gastropoda: Rissoidae). Zootaxa 4184 (2): 395-400, DOI: 10.11646/zootaxa.4184.2.13
