taxonID	type	description	language	source
773D2352584D0B618263F9EDFC0FF898.taxon	description	Male (1). As in Fig. 1. Preantennal margin slightly concave, anterior head notch narrow, 0.055 wide. Dorsal anterior plate slightly longer than wide, with a “ U­shaped ” extension posteriad beyond center of plate, tapering on its posterior margin (Fig. 5). Ventral carina with conspicuous guttate process. Pronotum with single lateroposterior seta on each side; each side of metanotum with lateral sides concave, with 6 marginal setae of varying lengths; thoracic sternum with 2 anterior and 3 posterior setae. Each abdominal tergite II (first apparent tergite) – III with 1 median seta, IV – V with 2 setae mediad of spiracle; VI with 3, VII with 4, VIII with only 1, IX with row of 4 – 5 short setae. Total sternal setae on each of II – V, 2; VI, 4. Lateral margin of II – III without setae, IV – V with 1, VI – VII with 2, and VIII with 3. Ventral outline of pleural thickening rounded. Genitalia as in Fig. 2; with widely separated circular parameres, 0.027 long and 0.019 wide. Dimensions: TW, 0.413; HL, 0.419; DAPL, 0.120; DAPW, 0.100; PW, 0.237; MW, 0.332; AWV, 0.466; GW, 0.060; TL, 1.493. Female (1). As in Fig. 3. Most features as in male, except narrower anterior head notch, 0.045 mm wide (all other females of Formicaphagus have the anterior head notch wider than in conspecific males). Each of abdominal tergites VI – VII with 2 setae mediad of spiracle, VIII with 1, and each side of IX with 2. Posterior margin of subgenital plate with almost straight lateral margins, with 19 short submarginal spiniform setae, 8 short fine marginal setae and 17 short fine setae anteriorly; sparse patch of 9 ventral setae on each side lateroposterior to subgenital plate (Fig. 4). Dimensions: TW, 0.435; HL, 0.432; DAPL, 0.118; DAPW, 0.103; PW, 0.248; MW, 0.344; AWV, 0.496; TL, 1.688.	en	Sychra, Old Ì Ich, Literák, Ivan, Apek, Miroslav Č, Martin (2006): Chewing lice (Phthiraptera) from typical antbirds and ground antbirds (Passeriformes: Thamnophilidae, Formicariidae) from Costa Rica, with descriptions of three new species of the genera Formicaphagus and Myrsidea. Zootaxa 1206: 47-61, DOI: 10.5281/zenodo.172392
773D2352584D0B618263F9EDFC0FF898.taxon	materials_examined	Type material. Male holotype, female allotype, ex Cercomacra tyrannina crepera Bangs, COSTA RICA: Hitoy Cerere BR, Provincia Limón (940 ’ N, 8505 ’ W), 26 August 2004, Literák, Č apek & Havlíček coll. Deposited in INBIO — O. Sychra CR 32.	en	Sychra, Old Ì Ich, Literák, Ivan, Apek, Miroslav Č, Martin (2006): Chewing lice (Phthiraptera) from typical antbirds and ground antbirds (Passeriformes: Thamnophilidae, Formicariidae) from Costa Rica, with descriptions of three new species of the genera Formicaphagus and Myrsidea. Zootaxa 1206: 47-61, DOI: 10.5281/zenodo.172392
773D2352584D0B618263F9EDFC0FF898.taxon	discussion	Remarks. The head shape, a dorsal plate longer than wide, and the chaetotaxy of male tergite VIII place Formicaphagus tyrannina n. sp. close to F. arnoldi Price and Clayton 1997 and F. donpetersi Price and Clayton 1997, which were described from gnateaters (Conopophagidae). Formicaphagus tyrannina is distinguished from both species by the shape of the dorsal anterior head plate (Figs 5 – 7), the presence of guttate processes on the ventral carinae, characters of the male genitalia and by the female having a narrower anterior head notch than the male. Both sexes of F. tyrannina may be separated from those of F. arnoldi by their smaller heads and dorsal anterior head plates. Also, females of F. tyrannina are smaller than females of F. donpetersi. The male of F. tyrannina is very similar to that of F. donpetersi, but they can be separated by the shape of the pleural thickening (F. donpetersi have ventral outline of pleural thickening approximately straight), shape of lateral side of metanotum, tergo­central setae on tergite VIII conspisously shorter (0.030 against 0.140), slightly larger parameres (0.027 x 0.019 against 0.022 x 0.016) and that of the anterior head notch. This species is described from single male and female specimens collected from single captured host individual. Further studies of chewing lice from the type host are needed to document the intraspecific variation for this louse. Host: Gymnocichla nudiceps (Cassin) — Bare­crowned Antbird One chewing louse species was found on this host.	en	Sychra, Old Ì Ich, Literák, Ivan, Apek, Miroslav Č, Martin (2006): Chewing lice (Phthiraptera) from typical antbirds and ground antbirds (Passeriformes: Thamnophilidae, Formicariidae) from Costa Rica, with descriptions of three new species of the genera Formicaphagus and Myrsidea. Zootaxa 1206: 47-61, DOI: 10.5281/zenodo.172392
773D2352584A0B678263FECFFCFCFD27.taxon	materials_examined	Type host. Myrmeciza immaculata (Lafresnaye) Material studied. 2 females and 1 nymph ex Gymnocichla nudiceps, COSTA RICA: Hitoy Cerere BR, Provincia Limón (940 ’ N, 8505 ’ W), 24 August 2004, Literák, Č apek & Havlíček. O. Sychra CR 34 – 35. Only one bird was examined. Host: Gymnopithys leucaspis (P. L. Sclater) — White­cheeked Antbird One chewing lice species was found on this host.	en	Sychra, Old Ì Ich, Literák, Ivan, Apek, Miroslav Č, Martin (2006): Chewing lice (Phthiraptera) from typical antbirds and ground antbirds (Passeriformes: Thamnophilidae, Formicariidae) from Costa Rica, with descriptions of three new species of the genera Formicaphagus and Myrsidea. Zootaxa 1206: 47-61, DOI: 10.5281/zenodo.172392
773D2352584A0B678263FD25FCFCFBCD.taxon	materials_examined	Type host. Thamnophilus doliatus (Linnaeus) Material studied. 2 males, 5 females and 1 nymph ex Gymnopithys leucaspis, COSTA RICA: Hitoy Cerere BR, Provincia Limón (940 ’ N, 8505 ’ W), 25 August 2004, Literák, Č apek & Havlíček. O. Sychra CR 18 – 21. Only one of five birds examined were parasitised by M. laticorpus. Host: Phaenostictus mcleannani (Lawrence) — Ocellated Antbird One chewing lice species was found on this host.	en	Sychra, Old Ì Ich, Literák, Ivan, Apek, Miroslav Č, Martin (2006): Chewing lice (Phthiraptera) from typical antbirds and ground antbirds (Passeriformes: Thamnophilidae, Formicariidae) from Costa Rica, with descriptions of three new species of the genera Formicaphagus and Myrsidea. Zootaxa 1206: 47-61, DOI: 10.5281/zenodo.172392
773D2352584A0B658263FB02FD6FFAD5.taxon	description	Male (2). Head without lateral notch or slit; lateral sides of preantennal region conspicuously concave; hypopharyngeal sclerites strongly developed. Mean length of head seta 10, 0.055 (n = 2); seta 11, 0.114 (n = 2); ratio 10 / 11, 0.48. Latero­ventral fringe with 8 – 9 setae; inner occipital setae long, outer very short; each side of temple margin with 1 long and 3 very long setae; gula with 3 – 4 setae on each side, posteriormost much thicker and longer than others. Pronotum with 6 moderately long posterior marginal setae, 2 short spiniform setae, and 1 finer seta at each lateral corner; well­developed elongate prosternal plate rounded anteriorly, with pair of minute anterior setae (Fig. 9). Mesonotum with median division, with pair of minute setae adjacent to postnotum; segment ringed by sclerite, with large mesosternal plate. Metanotum not enlarged, with very long corner setae and 4 setae on posterior margin. Pleural margin of metathorax with 3 (rarely 4) short, robust setae; metasternal plate large, triangular, with 2 – 3 long setae on each side. First tibia with 3 – 4 outer lateral ventral and 4 – 5 dorsal setae; femur III with 13 – 14 setae in ventral setal brush. Tergal setae (including the postspiracular setae and all setae between them): I, 10 – 12; II, 16; III – IV, 20 – 21; V – VI, 18 – 19; VII, 16; VIII, 8. Postspiracular setae extremely long on II, IV and VIII, long on III and VI – VII and shorter on I and V. Abdomen without anterior tergal or pleural setae; margin of pleurites I – IV with only short spiniform setae, pleurites V – VII with 1 – 2 longer finer setae; margin of pleurite VIII with median long seta flanked on each side by shorter seta. Sternal setae on II, 24 – 25 marginal, including cluster of 5 – 6 robust setae on each side. Marginal setae of sternite: III, 12 – 17; IV – VI, 19 – 21; VII, 13 – 14; VIII of subgenital plate, 7; remainder of plate, 10 – 14. Lateral anterior sternal setae: III, 1 – 2; IV – VI, 3 – 6; VII, 2, without medioanterior setae. Sternite VII separate from subgenital plate of fused VIII – IX; with 10 internal anal setae. Length of inner posterior seta of last tergum 0.047 (n = 1); short lateral marginal seta of last segment 0.023 (n = 2). Male genitalia of typical Myrsidea type. Genital sclerite comprises a flattened plate with ventral and dorsal arm each side. Genitalia and character of genital sclerite as in Figs. 12 – 13. Dimensions: TW, 0.46 – 0.47; HL, 0.31 – 0.32; PW, 0.30 – 0.31; PSL, 0.12 – 0.13; MW, 0.41 – 0.42; MSL, 0.15; AWIV, 0.53; TL, 1.44 – 1.45; GL, 0.47 – 0.53; GW, 0.12; GPL, 0.09. Female (2). As in Fig. 8. Most features as in male. Mean length of head seta 10, 0.052 (n = 4); seta 11, 0.113 (n = 4); ratio 10 / 11, 0.45. Setae of femoral brush, 17 – 19. Abdomen without conspicuously enlarged tergites; with tergal setae: I, 10 – 12; II, 17 – 21; III – IV, 20 – 22; V – VI, 19 – 20; VII, 14 – 15; VIII, 8. Sternal setae on II, 5 – 6 medioanterior; 26 – 30 marginal, including cluster of 6 – 7 robust setae on each side. Marginal setae of sternite: III, 15 – 19; IV – VI, 21 – 25; VII of subgenital plate, 14 – 17; remainder of plate with 8 – 10 marginal, 15 – 16 anterior setae. Subgenital plate formed by fusion of sternites VII – IX, with posterior margin finely serrated. Lateral anterior sternal setae: III, 1 – 3; IV – VI, 7 – 10; VII, 2 – 5; without medioanterior setae. Anal fringe of 30 – 38 dorsal and 39 – 40 ventral setae. Mean length of inner posterior seta of last tergum 0.054 (n = 4); short lateral marginal seta of last segment 0.032 (n = 4). Dimensions: TW, 0.51; HL, 0.33 – 0.36; PW, 0.33; PSL, 0.14; MW, 0.50; MSL, 0.16; AWIV, 0.69 – 0.73; TL, 1.70 – 1.78; ANW, 0.31 – 0.32.	en	Sychra, Old Ì Ich, Literák, Ivan, Apek, Miroslav Č, Martin (2006): Chewing lice (Phthiraptera) from typical antbirds and ground antbirds (Passeriformes: Thamnophilidae, Formicariidae) from Costa Rica, with descriptions of three new species of the genera Formicaphagus and Myrsidea. Zootaxa 1206: 47-61, DOI: 10.5281/zenodo.172392
773D2352584A0B658263FB02FD6FFAD5.taxon	materials_examined	Type material. Female holotype, male allotype, ex Phaenostictus mcleannani, COSTA RICA: Hitoy Cerere BR, Limón province (940 ’ N, 8505 ’ W), 25 August 2004, Literák, Č apek, Havlíček. Paratypes: 1 female and 1 male with same data as holotype. Deposited in INBIO — O. Sychra CR 11 – 13.	en	Sychra, Old Ì Ich, Literák, Ivan, Apek, Miroslav Č, Martin (2006): Chewing lice (Phthiraptera) from typical antbirds and ground antbirds (Passeriformes: Thamnophilidae, Formicariidae) from Costa Rica, with descriptions of three new species of the genera Formicaphagus and Myrsidea. Zootaxa 1206: 47-61, DOI: 10.5281/zenodo.172392
773D2352584A0B658263FB02FD6FFAD5.taxon	discussion	Remarks. Both sexes of this species are characterized by a combination of large dimensions, lateral margins of preantennal region conspicuously concave, the relative lengths of their postspiracular setae, and the absence of medioanterior sternal setae. Furthermore, males are distinguished by having 10 internal anal setae and by features of the genital sclerite. The male genital sclerite comprising ventral and dorsal arms, is similar to that of furnarid­ and icterid­parasitizing Myrsidea or M. ishizawai Uchida, 1926 from Zoothera dauma (Latham) (Turdidae), but it can be separated from that species by the shape of the distal part of the dorsal arm, which is not toothed. Two birds were examined and both were parasitised by Myrsidea mcleannani. 9	en	Sychra, Old Ì Ich, Literák, Ivan, Apek, Miroslav Č, Martin (2006): Chewing lice (Phthiraptera) from typical antbirds and ground antbirds (Passeriformes: Thamnophilidae, Formicariidae) from Costa Rica, with descriptions of three new species of the genera Formicaphagus and Myrsidea. Zootaxa 1206: 47-61, DOI: 10.5281/zenodo.172392
773D235258480B6A8263F98BFB64FE32.taxon	description	15	en	Sychra, Old Ì Ich, Literák, Ivan, Apek, Miroslav Č, Martin (2006): Chewing lice (Phthiraptera) from typical antbirds and ground antbirds (Passeriformes: Thamnophilidae, Formicariidae) from Costa Rica, with descriptions of three new species of the genera Formicaphagus and Myrsidea. Zootaxa 1206: 47-61, DOI: 10.5281/zenodo.172392
773D235258470B688263F968FB57FBE8.taxon	description	Male (2). Head without lateral notch or slit; lateral sides of preantennal region convex; hypopharyngeal sclerites reduced. Mean length of head seta 10, only 0.020 (n = 4); seta 11, 0.103 (n = 4); ratio 10 / 11, 0.19. Latero­ventral fringe with 8 – 10 setae; outer occipital setae as long as inner one; short fine seta 23 (according to Clay 1969) present and anterolateral to 22. Each side of temple margin with 1 long and 3 very long setae; gula with 3 – 5 setae on each side, posteriormost much heavier and longer than others. Pronotum with 6 long and 2 short posterior marginal setae, and only 1 short spiniform setae plus 1 finer seta on each lateral corner. Anterior margin of prosternal plate is straight with pair of minute anterior setae (Fig. 11). Mesonotum with median division, with pair of minute setae adjacent to postnotum; segment ringed by sclerite, with large mesosternal plate. Metanotum not enlarged, with very long corner setae and 4 setae on posterior margin. Pleural margin of metathorax with 3 (rarely 4) short, stout setae; metasternal plate large, triangular, with 4 – 5 long setae on each side. First tibia with 3 outer lateral ventral and 4 – 5 dorsal setae; femur III with 12 setae in ventral setal brush. Tergal setae (including postspiracular setae and all setae between them): I, 8; II, 10 – 11; III, 14 – 15; IV – VII, 9 – 13; VIII, 8 – 9. Postspiracular setae extremely long on II – IV and VIII, long on I and VII, and shorter V – VI. Abdomen without anterior tergal or pleural setae; margin of pleurites I – IV with only short spiniform setae, pleurites V – VII also with 1 – 2 longer finer setae; margin of pleurite VIII with median long seta flanked on each side by shorter seta. Sternite I rounded posteriorly and situated inside wide notch of sternite II. Sternal setae on II, 10 medioanterior; 18 – 20 marginal, including cluster of 4 robust setae on each side. Marginal setae of sternite: III, 17, including another 1 – 2 robust setae on each side; IV – V, 19 – 20; VI, 17 – 18; VII, 12 – 14; VIII, 7 – 10; subgenital plate (only sternite IX), 10 – 13. Lateral anterior sternal setae: III, 0; IV, 2 – 4; V – VI, 3 – 6; VII, 1 – 3, with 2 – 5 medioanterior setae on IV – VII. Length of inner posterior seta of last tergum 0.021 (n = 3); short lateral marginal seta of last segment 0.031 (n = 3) Genitalia and distal end of genital sclerite as in Figs. 14 and 15 respectively. Dimensions: TW, 0.39 – 0.40; HL, 0.24 – 0.25; PW, 0.24; PSL, 0.10; MW, 0.29 – 0.31; MSL, 0.15; AWIV, 0.39 – 0.40; TL, 1.13 – 1.18; GL, 0.30 – 0.32; GW, 0.09 – 0.10; GPL, 0.07. Female (4). As in Fig. 10. Most features as in male. Mean length of head seta 10, 0.023 (n = 7); seta 11, 0.110 (n = 7); ratio 10 / 11, 0.21. Setae of femoral brush, 13 – 15. Abdomen without conspicuously enlarged tergites; with tergal setae: I, 8 – 10; II – IV, 13 – 17; V – VII, 11 – 14; VIII, 8. Pleurites III – VI with one short spiniform anterior seta. Sternal setae on II, 8 – 11 medioanterior; 21 – 23 marginal, including cluster of 5 – 6 robust setae on each side. Marginal setae of sternite: III, 22; including another 3 robust setae on each side; IV, 21 – 23; V, 19 – 22; VI, 17 – 19; VII, 14 – 17; subgenital plate is forming by fusion of sternites VIII – IX with 12 – 14 marginal and 24 – 31 anterior setae. Lateral anterior sternal setae: III, 1; IV, 3 – 8; V – VI, 6 – 10; VII, 4 – 7; with 4 – 8 medioanterior setae on IV – VII. Posterior margin of subgenital plate is plain, not serrated. Anal fringe of 21 – 33 dorsal and 28 – 41 ventral setae. Mean length of inner posterior seta of last tergum 0.032 (n = 6); short lateral marginal seta of last segment 0.042 (n = 6). Dimensions: TW, 0.42 – 0.43; HL, 0.26 – 0.27; PW, 0.25 – 0.27; PSL, 0.11; MW, 0.35 – 0.37; MSL, 0.17 – 0.18; AWIV, 0.49 – 0.52; TL, 1.35 – 1.46; ANW, 0.19 – 0.20.	en	Sychra, Old Ì Ich, Literák, Ivan, Apek, Miroslav Č, Martin (2006): Chewing lice (Phthiraptera) from typical antbirds and ground antbirds (Passeriformes: Thamnophilidae, Formicariidae) from Costa Rica, with descriptions of three new species of the genera Formicaphagus and Myrsidea. Zootaxa 1206: 47-61, DOI: 10.5281/zenodo.172392
773D235258470B688263F968FB57FBE8.taxon	materials_examined	Type material. Female holotype, male allotype, ex Formicarius analis, COSTA RICA: Hitoy Cerere BR, Provincia Limón (940 ’ N, 8505 ’ W), 27 and 31 August 2004, Literák, Č apek & Havlíček. Paratypes: 1 female and 1 male with same data as holotype. Deposited in INBIO — O. Sychra CR 14 – 16.	en	Sychra, Old Ì Ich, Literák, Ivan, Apek, Miroslav Č, Martin (2006): Chewing lice (Phthiraptera) from typical antbirds and ground antbirds (Passeriformes: Thamnophilidae, Formicariidae) from Costa Rica, with descriptions of three new species of the genera Formicaphagus and Myrsidea. Zootaxa 1206: 47-61, DOI: 10.5281/zenodo.172392
773D235258470B688263F968FB57FBE8.taxon	discussion	Remarks. Both sexes of M. klimesi are readily identified by a unique combination of characters, as follows: (1) reduced hypopharynx; (2) head seta 10 very small and, consequently, a very small ratio of lengths of head setae 10 / 11; (3) presence of head seta 23, and outer occipital seta as long as the inner one; (4) pronotum with 6 long and 2 short posterior marginal setae, and only one short spiniform setae plus one finer seta on each lateral corner; (5) straight anterior margin of prosternal plate; (6) position of sternite I, which lies inside the wide notch of sternite II; (7) relative lengths of the postspiracular setae, especially the extremely long seta on abdominal tergite III; (8) presence of a cluster of robust setae on each side of sternite III, as well as on II; (9) presence of medioanterior setae on sternites IV – VII; (10) male abdominal sternite VIII and female sternite VII both separated from the subgenital plate, which is formed by a single sternite IX (in the male) or fusion of sternites VIII and IX (in the female); (11) female with one anterior pleural seta on pleurites III – VI; (12) posterior margin of female subgenital plate smooth (not serrated); (13) unique feature in the male genital sclerite (Fig. 12). Two of four birds examined were parasitised by M. klimesi.	en	Sychra, Old Ì Ich, Literák, Ivan, Apek, Miroslav Č, Martin (2006): Chewing lice (Phthiraptera) from typical antbirds and ground antbirds (Passeriformes: Thamnophilidae, Formicariidae) from Costa Rica, with descriptions of three new species of the genera Formicaphagus and Myrsidea. Zootaxa 1206: 47-61, DOI: 10.5281/zenodo.172392
773D235258470B688263F968FB57FBE8.taxon	etymology	Etymology. This species is named in honour of Jiří Klimeš, our colleague and friend, in recognition of his friendship and unmatched enthusiasm in the study of zoology.	en	Sychra, Old Ì Ich, Literák, Ivan, Apek, Miroslav Č, Martin (2006): Chewing lice (Phthiraptera) from typical antbirds and ground antbirds (Passeriformes: Thamnophilidae, Formicariidae) from Costa Rica, with descriptions of three new species of the genera Formicaphagus and Myrsidea. Zootaxa 1206: 47-61, DOI: 10.5281/zenodo.172392
