taxonID	type	description	language	source
705ACA4EFFEDFFEDFE4EC7F5FF0EC0C0.taxon	description	(Figs. 2 – 6)	en	Caterino, Michael S., Leschen, Richard A. B., Johnson, Colin (2008): A New Genus of Caenoscelini (Cryptophagidae: Cryptophaginae) from California, with Two New Species. The Coleopterists Bulletin 62 (4): 509-523, DOI: 10.1649/1107.1, URL: http://dx.doi.org/10.1649/1107.1
705ACA4EFFEDFFEDFE4EC7F5FF0EC0C0.taxon	diagnosis	Diagnosis. Body form elongate oval. Eye reduced to single facet. Antennal club apparently 1 - segmented (with the two apical antennomeres fused). Genal carina present. Pronotal sides converging anteriorly with greatest width at base. Pronotum without sublateral lines, but transverse line at middle present. Prosternum and prosternal process vaulted, with longitudinal lines. Weakly projecting anteromedially. Prosternum without transverse pits. Hypomeron with antennal groove. Hind wing absent.	en	Caterino, Michael S., Leschen, Richard A. B., Johnson, Colin (2008): A New Genus of Caenoscelini (Cryptophagidae: Cryptophaginae) from California, with Two New Species. The Coleopterists Bulletin 62 (4): 509-523, DOI: 10.1649/1107.1, URL: http://dx.doi.org/10.1649/1107.1
705ACA4EFFEDFFEDFE4EC7F5FF0EC0C0.taxon	description	Description. Length 1.21 – 1.47 mm; body form elongate oval, strongly convex, testaceous, setae conspicuous, decumbent, uniformly dense over entire body. Head with frontal carina prominent, running from eye to eye, separating convex, setose frons from depressed, glabrous genae, acutely angulate between antennal bases, this angulation continuous with short genal carina or epistomal ridge; eye reduced to single, protruding ommatidium, only faintly differentiated from surrounding cuticle; antennae with antennomeres 10 – 11 fused into 1 - segmented club, with faint constriction and series of blunt setae marking line of fusion; frontoclypeal suture not visible; labrum strongly reduced, mostly hidden by clypeus; maxillae with three segmented palpi, palpomere two about half the length and slightly narrower than palpomere 1, palpomere 3 much narrower than 2, about 1.5 times as long, narrowed to subacute, setose apex; mentum wide, with rounded sides, bi-emarginate at palpal bases, acute at middle, with transverse apical stria; labium with three segmented palpi, basal-most strongly reduced, palpomere 2 bulbous, palpomere 3 narrower at base than 2 and narrowed to subacute apex; gular sutures fused, indistinguishable. Pronotum with sides weakly convergent from base, subangulate just behind middle, thence strongly narrowed to apex and widest at base, margined and finely depressed along lateral edge; sublateral lines absent; pronotal disk with sinuate, crenulate transverse stria, meeting marginal angle at sides, disk faintly depressed behind; pronotal disk punctate, with punctures slightly less dense at middle behind transverse stria. Hypomeron with well developed antennal groove in anterior one-third, delimited by strong lateral and weaker medial carinae; prosternum and prosternal process broad and vaulted, about equal in width to procoxa, with two parallel longitudinal lines, with series of setae along lateral striae, but glabrous at middle, the lateral striae meeting before anterior prosternal margin; prosternum lacking transverse pits; weak chin piece present. Mesoventrite deeply emarginate for reception of prosternal keel, mesocoxae widely separated; mesanepisternum lacking fovea or pits; abdomen with 5 visible abdominal ventrites, none connate, all with marginal striae along lateral and posterior edges; first abdominal ventrite with broad, rounded intercoxal process; metasubcoxal lines present, acutely angulate behind broadest part of metacoxa; setae of ventrites converging to midline, those of ventrite 5 more dense, with dense group of blunt setae near apex. Legs with trochanters relatively long, approximately one-sixth (anterior) to one-fifth of total trochanter + femur length (measured along shorter, dorsal margin); tibiae widened in apical half, with apical ctenidium around inner margin; tarsus 5 - 5 - 5 in female and 5 - 4 - 4 in male, tarsomeres with irregular longitudinal striations; tarsal claws simple. Aedeagus horizontal in abdominal cavity, symmetrical, tegmen narrowly rounded anteriorly, anterior edge without strut; parameres reduced to pore-bearing bracons that are fused to phallobase. Penis anteriorly with single broad strut that is 4 times as long as median lobe; endophallus complex with asperities and lacking internal sclerites.	en	Caterino, Michael S., Leschen, Richard A. B., Johnson, Colin (2008): A New Genus of Caenoscelini (Cryptophagidae: Cryptophaginae) from California, with Two New Species. The Coleopterists Bulletin 62 (4): 509-523, DOI: 10.1649/1107.1, URL: http://dx.doi.org/10.1649/1107.1
705ACA4EFFEDFFEDFE4EC7F5FF0EC0C0.taxon	type_taxon	Type species. Renodesta stephani, new species, here designated.	en	Caterino, Michael S., Leschen, Richard A. B., Johnson, Colin (2008): A New Genus of Caenoscelini (Cryptophagidae: Cryptophaginae) from California, with Two New Species. The Coleopterists Bulletin 62 (4): 509-523, DOI: 10.1649/1107.1, URL: http://dx.doi.org/10.1649/1107.1
705ACA4EFFEDFFEDFE4EC7F5FF0EC0C0.taxon	discussion	Remarks. Among Cryptophagidae, adult Caenoscelini are characterized by the combination of: lack of conspicuous glandular ducts on any body surface, serrate incisor lobe of mandible (most commonly on the left mandible), terminal labial palpomere narrower than penultimate, presence of a ‘ boss’ on the front of the head, absence of gular sutures, pronotum laterally unmodified and lacking basal pits, presence of thickened setae on ventrite 5, a vertically oriented aedeagus, and club shaped tibiae (Leschen 1996). Renodesta agrees in all these characters apart from aedeagal orientation (though this is variable in many cucujoid taxa), and shares additional characters with both Dernostea and Sternodea. All three exhibit brachyptery or aptery (note that some Caenoscelis are also apterous, Johnson and Bowestead 2003). With Dernostea, Renodesta also shares a genal carina, single segmented antennal club and reduced eyes. With Sternodea it shares a transverse pronotal stria, eye reduction (in some species) and lack of deep, annulate prosternal pits. The genus is supported to be sister taxon to the Asian clade of Dernostea + Himascelis. 516 The species of Renodesta are found only in the central coast ranges of California, between Monterey Bay and Santa Barbara (roughly 34 – 37 ° N). Specimens have been sifted from a variety of litters, including leaves of Quercus, Platanus, Ceanothus, Metasequoia, Lithocarpus, and Umbellularia, most frequently within or in close proximity to very rotten wood (Quercus, Platanus and Ceanothus).	en	Caterino, Michael S., Leschen, Richard A. B., Johnson, Colin (2008): A New Genus of Caenoscelini (Cryptophagidae: Cryptophaginae) from California, with Two New Species. The Coleopterists Bulletin 62 (4): 509-523, DOI: 10.1649/1107.1, URL: http://dx.doi.org/10.1649/1107.1
705ACA4EFFEDFFEDFE4EC7F5FF0EC0C0.taxon	etymology	Etymology. The genus name is an anagram of the related genera Dernostea and Sternodea, and is feminine. 518	en	Caterino, Michael S., Leschen, Richard A. B., Johnson, Colin (2008): A New Genus of Caenoscelini (Cryptophagidae: Cryptophaginae) from California, with Two New Species. The Coleopterists Bulletin 62 (4): 509-523, DOI: 10.1649/1107.1, URL: http://dx.doi.org/10.1649/1107.1
705ACA4EFFE7FFEAFE02C559FF43C30D.taxon	description	(Figs. 2, 3 B, 4, 5 C – D, 6 C – D)	en	Caterino, Michael S., Leschen, Richard A. B., Johnson, Colin (2008): A New Genus of Caenoscelini (Cryptophagidae: Cryptophaginae) from California, with Two New Species. The Coleopterists Bulletin 62 (4): 509-523, DOI: 10.1649/1107.1, URL: http://dx.doi.org/10.1649/1107.1
705ACA4EFFE7FFEAFE02C559FF43C30D.taxon	diagnosis	Diagnosis. Funicle slender with antennomere 3 about 1.3 times the length of antennomere 2 (Fig. 5 C). Sides of the prosternal process parallel (Fig. 5 D). Aedeagus in lateral view with apex of phallobase ventrally oriented and penis narrower than its base as posterior quarter (Fig. 6 C – D).	en	Caterino, Michael S., Leschen, Richard A. B., Johnson, Colin (2008): A New Genus of Caenoscelini (Cryptophagidae: Cryptophaginae) from California, with Two New Species. The Coleopterists Bulletin 62 (4): 509-523, DOI: 10.1649/1107.1, URL: http://dx.doi.org/10.1649/1107.1
705ACA4EFFE7FFEAFE02C559FF43C30D.taxon	description	Description. Length (n 5 7) 4.10 – 4.45 mm. Body about 1.85 times as long as wide (BL / EW 5 1.81 – 1.93), and greatest depth about 0.68 times as great as elytral width (GD / EW 5 0.67 – 0.70). Overall color light to tan brown; antennae, mouthparts, and legs yellowish-brown in darker specimens. Dorsal vestiture of suberect setae on pronotum, biseriate on elytra and composed of suberect and decumbent setae that are more or less the same lengths; longest setae at posterior end of elytra about as long as antennal club; setae on ventral surfaces slightly shorter and appressed. Punctation relatively coarse on all surfaces: punctures of head with diameters half that of those on the pronotum and separated by about K their diameter; punctures of pronotum anterior to the transverse stria separated by 1 diameter or less, punctures posterior to transverse stria separated by up to 3 diameters, shallower and less dense at middle; elytral punctation similar to that present in the posterior portion of the pronotum. Antennomere ratios 2.75: 3: 3: 1: 1: 1: 1: 1.25: 1.5: 5.5; funicle slender with antennomere 3 about 1.3 times the length of antennomere 2. Pronotum about 0.61 times as long as wide (PL / PW 5 0.58 – 0.63). Elytra about 1.33 times as long as wide (EL / EW 5 1.28 – 1.32) and about 2.53 times as long as pronotum (EL / PL 5 2.34 – 2.69). Sides of the prosternal process parallel. Aedeagus in lateral view with apex of phallobase ventrally oriented and penis narrower than its base as posterior quarter.	en	Caterino, Michael S., Leschen, Richard A. B., Johnson, Colin (2008): A New Genus of Caenoscelini (Cryptophagidae: Cryptophaginae) from California, with Two New Species. The Coleopterists Bulletin 62 (4): 509-523, DOI: 10.1649/1107.1, URL: http://dx.doi.org/10.1649/1107.1
705ACA4EFFE7FFEAFE02C559FF43C30D.taxon	discussion	Remarks. As discussed above, R. ramsdalei has a nearly parallel-sided prosternal process (Fig. 5 D), and its antennomere 3 is about 1.3 times the length of antennomere 2, the funicle in total being relatively slender (Fig. 5 C). The apex of the penis in this species is (in lateral view) pointed ventrally and deflected anteriorly (Fig. 6 C – D) The two are otherwise extremely similar, and adequately characterized by the generic description.	en	Caterino, Michael S., Leschen, Richard A. B., Johnson, Colin (2008): A New Genus of Caenoscelini (Cryptophagidae: Cryptophaginae) from California, with Two New Species. The Coleopterists Bulletin 62 (4): 509-523, DOI: 10.1649/1107.1, URL: http://dx.doi.org/10.1649/1107.1
705ACA4EFFE7FFEAFE02C559FF43C30D.taxon	etymology	Etymology. We dedicate this species to the memory of Alistair Ramsdale, formerly a graduate student of Montana State University, who assisted in the collection of the type series.	en	Caterino, Michael S., Leschen, Richard A. B., Johnson, Colin (2008): A New Genus of Caenoscelini (Cryptophagidae: Cryptophaginae) from California, with Two New Species. The Coleopterists Bulletin 62 (4): 509-523, DOI: 10.1649/1107.1, URL: http://dx.doi.org/10.1649/1107.1
705ACA4EFFE7FFEAFE02C559FF43C30D.taxon	materials_examined	Type Material. Holotype: ‘‘ CA: Santa Barbara Co., 34.4855 ° N 120.1423 ° W, Arroyo Hondo Preserve, iv. 8.2006, Platanus litter, Caterino & Ramsdale’ ’ / ‘‘ CA BEETLE PROJ, CBP 0044721 ’’; deposited in SBMN. Paratypes (total 24). 14: same data as type, CBP 0008160 (gold coated for SEM), CBP 0071095 (DNA extraction voucher MSC- 1456), CBP 0044734, CBP 0044729, CBP 0078326, CBP 0078327, CBP 0078328, CBP 0078329, CBP 0078330; 7: ‘‘ California: Santa Barbara Co. Arroyo Hondo Preserve, 34.4855 ° N 120.1423 ° W, 08 April 2006, Ramsdale & Caterino leg., berlese Quercus sp. debris & litter’ ’. 1: ‘‘ CA: Santa Barbara Co., 34.4833 ° N 120.1428 ° W, Arroyo Hondo Preserve, 25 mi. W Sta. Barbara, i. 1.2003, M. Caterino, sifted Quercus litter’ ’, CBP 0003795; 1: ‘‘ CA: Santa Barbara Co., 34.4833 ° N 120.1428 ° W, Arroyo Hondo Preserve, 25 mi. W Sta. Barbara, i. 8.2003, M. Caterino, sifted Quercus litter’ ’, CBP 0004774; 1: ‘‘ CA: Santa Barbara Co., 34.5145 ° N 119.8015 ° W, LPNF: E. Camino Cielo, xi. 22 - xii. 5.2003, M. Caterino, unbaited pitfall’ ’, CBP 0016536; deposited in CASC, CJC, MTEC, NZAC, and SBMN.	en	Caterino, Michael S., Leschen, Richard A. B., Johnson, Colin (2008): A New Genus of Caenoscelini (Cryptophagidae: Cryptophaginae) from California, with Two New Species. The Coleopterists Bulletin 62 (4): 509-523, DOI: 10.1649/1107.1, URL: http://dx.doi.org/10.1649/1107.1
705ACA4EFFE1FFEBFE1BC2B5FE0BC378.taxon	description	(Figs. 2, 3 A, C, 4, 5 A – B, 6 A – B)	en	Caterino, Michael S., Leschen, Richard A. B., Johnson, Colin (2008): A New Genus of Caenoscelini (Cryptophagidae: Cryptophaginae) from California, with Two New Species. The Coleopterists Bulletin 62 (4): 509-523, DOI: 10.1649/1107.1, URL: http://dx.doi.org/10.1649/1107.1
705ACA4EFFE1FFEBFE1BC2B5FE0BC378.taxon	diagnosis	Diagnosis. Funicle relatively compact with antennomeres 2 and 3 subequal (Fig. 5 A). Sides of the prosternal process converging anteriorly (Fig. 5 B). Aedeagus in lateral view with apex of phallobase obliquely oriented and penis about as wide as base of posterior quarter (Fig. 6 A – B).	en	Caterino, Michael S., Leschen, Richard A. B., Johnson, Colin (2008): A New Genus of Caenoscelini (Cryptophagidae: Cryptophaginae) from California, with Two New Species. The Coleopterists Bulletin 62 (4): 509-523, DOI: 10.1649/1107.1, URL: http://dx.doi.org/10.1649/1107.1
705ACA4EFFE1FFEBFE1BC2B5FE0BC378.taxon	description	Description. Length 1.19 – 1.44 mm (n 5 10). Body about 1.85 times as long as wide (BL / EW 5 1.81 – 2.00), and greatest depth about 0.70 times as great as elytral width (GD / EW 5 0.68 – 0.75). Overall color light to tan brown; antennae, mouthparts, and legs yellowish-brown in darker specimens. Dorsal vestiture of decumbent setae on pronotum, biseriate on elytra and composed of suberect and decumbent setae that differ in lengths; longest setae at posterior end of elytra about as long as antennal club (specimens from Big Sur with setae more decumbent and slightly shorter); setae on ventral surfaces slightly shorter and appressed. Punctation relatively coarse on all surfaces: punctures of head with diameters subequal to those on the pronotum and separated by 1 diameter or less; punctures of pronotum anterior to the transverse stria separated by 1 – 2 diameters, punctures posterior to transverse stria separated by 1 – 3 diameters; elytral punctation similar to that present in the posterior portion of the pronotum. Antennomere ratios 2.7: 2: 2: 1: 1: 1: 1: 1: 1.2: 4.4; funicle relatively compact with antennomeres 2 and 3 subequal. Pronotum about 0.68 times as long as wide (PL / PW 5 0.68 – 69). Elytra about 1.31 times as long as wide (EL / EW 5 1.25 – 1.39) and about 2.24 times as long as pronotum (EL / PL 5 2.20 – 2.31). Sides of the prosternal process converging anteriorly. Aedeagus in lateral view with apex of phallobase obliquely oriented and penis about as wide as base of posterior quarter.	en	Caterino, Michael S., Leschen, Richard A. B., Johnson, Colin (2008): A New Genus of Caenoscelini (Cryptophagidae: Cryptophaginae) from California, with Two New Species. The Coleopterists Bulletin 62 (4): 509-523, DOI: 10.1649/1107.1, URL: http://dx.doi.org/10.1649/1107.1
705ACA4EFFE1FFEBFE1BC2B5FE0BC378.taxon	discussion	Remarks. The two species of Renodesta are easily distinguished. Renodesta stephani has the sides of the prosternal process converging anteriorly (Fig. 5 B), whereas the prosternal process in R. ramsdalei is nearly or fully parallel-sided (Fig. 5 D). The antennal funicle of R. stephani is slightly thicker, with the difference most noticeable in a comparison of antennomeres 2 and 3, which are subequal in R. stephani (Fig. 5 A). In R. ramsdalei antennomere 3 is about 1.3 times as long as antennomere 2 (Fig. 5 C). The male genitalia of the two are distinct, with the apex of the penis in lateral view pointed ventrally in R. stephani (Fig. 6 A – B).	en	Caterino, Michael S., Leschen, Richard A. B., Johnson, Colin (2008): A New Genus of Caenoscelini (Cryptophagidae: Cryptophaginae) from California, with Two New Species. The Coleopterists Bulletin 62 (4): 509-523, DOI: 10.1649/1107.1, URL: http://dx.doi.org/10.1649/1107.1
705ACA4EFFE1FFEBFE1BC2B5FE0BC378.taxon	etymology	Etymology. We take pleasure naming this species for Karl Stephan, who collected the first known specimens of this species near Big Sur, California many years ago (1969). 520 Type material. Holotype: ‘‘ CA: Monterey Co., 36.0178 ° N 121.5962 ° W, UC Big Creek Reserve, Eagle Pt. Tr., iii. 27.2004, Ceanothus litter, M. Caterino’ ’ / ‘‘ CA BEETLE PROJ, CBP 0018455 ’’; deposited in SBMN. Paratypes (total 56). 25: same data as type, CBP 0043752 (gold coated for SEM), CBP 0018464, CBP 0018465, CBP 0018454, CBP 0018458, CBP 0018457, CBP 0078311, CBP 0078312, CBP 0078313, CBP 0078314, CBP 0078315, CBP 0078316, CBP 0078317, CBP 0078318, CBP 0078319, CBP 0078320, CBP 0078321, CBP 0078322, CBP 0078323, CBP 0078324, CBP 0078325; 1: ‘‘ CA: Monterey Co., 36.0857 ° N 121.5935 ° W, UC Big Creek Reserve, @ Deer Head Spring, ii. 10.2003, M. Caterino, oak / bay litter’ ’, CBP 0005902; 1: ‘‘ CA: Monterey Co., 36.0772 ° N 121.5923 ° W, UC Big Creek Reserve, Redwood Camp, iii. 28.2004, M. Caterino, redwood litter’ ’, CBP 0018436; 17: ‘‘ CA: Monterey Co., 36.0904 ° N 121.5880 ° W, UC Big Creek Reserve, nr. Boronda Camp, iii. 28.2004, M. Caterino, Quercus litter’ ’, CBP 0071172 (DNA extraction voucher MSC- 1457), CBP 0018427, CBP 0018433, CBP 0018434, CBP 0018429, CBP 0078299, CBP 0078300, CBP 0078301, CBP 0078302, CBP 0078303, CBP 0078304, CBP 0078305, CBP 0078306, CBP 0078307, CBP 0078308, CBP 0078309, CBP 0078310; 5: ‘‘ CA: Monterey Co., 36.0712 ° N 121.5854 ° W, UC Big Creek Reserve, ii. 14.2006, Caterino & Chatzimanolis, tanoak litter’ ’, CBP 0041604, CBP 0041594, CBP 0041571, CBP 0041576, CBP 0041603; 1: ‘‘ CA: Monterey Co., 36.0673 ° N 121.5781 ° W, UC Big Creek Reserve, Gamboa Rd., iii. 30.2004, M. Caterino, Quercus litter’ ’, CBP 00018547; 6: ‘‘ California, Big Sur, June 16 1969 ’’, Renodesta stephani gen. sp. n. [handwritten], det. C. Johnson 1982; deposited in CASC, CJC, FSCA, NZAC, MMUE, MTEC and SBMN.	en	Caterino, Michael S., Leschen, Richard A. B., Johnson, Colin (2008): A New Genus of Caenoscelini (Cryptophagidae: Cryptophaginae) from California, with Two New Species. The Coleopterists Bulletin 62 (4): 509-523, DOI: 10.1649/1107.1, URL: http://dx.doi.org/10.1649/1107.1
