taxonID	type	description	language	source
715487A7FFD9EC65D8D9EFBEFDAE1590.taxon	description	Figs 1 A – E, 107, 151 F	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFD9EC65D8D9EFBEFDAE1590.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of the flagellum, long and tapering, with bristle-like tips (Fig. 151 F); epandrium with two bulging lobes; widened surstyli basally, moderately narrowed to apex in dorsal view (Fig. 1 A); gonopods triangle-shaped with an edge in lateral margin in ventral view (Fig. 1 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFD9EC65D8D9EFBEFDAE1590.taxon	description	Description: MALE: Body length: 3.0 mm. Head. Scape, pedicel dark, flagellum long tapering, brown pollinose, tips narrowed and like bristle (Fig. 151 F). Face silvery brown pollinose with a bright line close to antennae. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 3 – 4 dark different-sized bristles. Mesonotum (viewed obliquely from front) brown pollinose. Dorsocentral and supra-alar bristles short, dark, longer in frontal half. Scutellum silvery brown pollinose and with 4 – 6 bristles along distal edge. Halter, knob light brown, tip brownish, base and stem dark brown. Legs. Mid coxa with 3 – 4 long dark bristles. Trochanters brown, femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae light brown (tibiae otherwise brown), tarsal segments light brown. Hind trochanter with 5 – 8 short dark bristles ventrobasally, brown pollinose. Hind femur without ventroapical spines. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres not flattened, hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 3.2 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 4 – 5 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites subshiny brown-black, tergite 1 brown pollinose Dispersed short dark bristles on all tergites present. Lateral bristles on first tergite present, 6 – 8 dark bristles up to as long as hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.1) with two bulging lobes in each side. Surstyli widened at basal, moderately narrowed to apex, constricted before apex, curved to each other at apex (Fig. 1 A). Genital capsule in ventral view: gonopods extended towards surstyli, pointed at apex with an edge on outer margin; phallic guide without dorso- dorsolateral spines; subepandrial sclerite distinct (Fig. 1 B); genital capsule in lateral view: both surstyli curved towards sternite, (Fig. 1 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 1 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFD9EC65D8D9EFBEFDAE1590.taxon	etymology	Etymology: From the Latin acuminatus, pointed or sharpened, in reference to the long, tapering flagellum. Examined material: HOLOTYPE: AUSTRALIA: Queensland: The Boulders, 5 km W of Babinda, 17 ° 21 ’ S, 145 ° 55 ’ E, 14 – 15. XI. 1981, D. H. Colless, at light, JSS 9254 (1 ♂, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFD9EC65D8D9EFBEFDAE1590.taxon	distribution	Distribution: Australia (Queensland) (Fig. 107). Notes: Unlike most Tomosvaryella species, this appears to be a rainforest species and is conceivably nocturnal given that it was collected at a light.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFD9EC62D8D9E956FCB616C8.taxon	description	Figs 2 A – E, 107, 149 B	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFD9EC62D8D9E956FCB616C8.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter with a keel covered by a row of short bristles in a row along the edge (Fig. 149 B); elongated surstyli in dorsal view (Fig. 2 A); elongated gonopods in ventral view (Fig. 2 B); subepandrial sclerite with 3 – 4 fin-shaped protrusions in middle in ventral view; phallic guide with 3 – 4 dorso- dorsolateral spines; one of ejaculatory ducts with one distinct spine in lateral view (Fig. 2 C).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFD9EC62D8D9E956FCB616C8.taxon	description	Description: MALE: Body length: 2.8 – 3 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 – 2.5 times the length of ocellar triangle. Occiput silvery pollinose, slightly less so on upper 1 / 3. Thorax. Postpronotum pale, with 5 – 6 pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum brown pollinose and with 8 – 10 bristles along the distal edge. Dorsocentral bristles distinct, dark, somewhat longer along frontal edge, numerous long bristles around postpronotum present. Halter, knob pale, stem brown. Legs. Mid coxa with 4 strong, dark bristles, one is shorter. Trochanters and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees yellow-brown (tibiae otherwise dark brown), tarsal segments yellow-brown, ventrally paler, last segment darker. Hind trochanter with a small keel, in a row of short bristles along the edge (Fig. 149 B); ventrobasal spines absent on fore and mid femora. Ventroapical row of spines on fore femur absent; 4 – 5 short dark spines on mid femur; hind femur without ventroapical spines, except 10 – 12 bristles posteroventrally, equally spaced, longer on distal half, longest up to 1 / 3 of the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present. Hind tibia slightly bent in posterior view. Metatarsi not flattened, hind metatarsus almost as long as 2 – 4 combined, all tarsomeres shorter than usual. Pulvilli shorter than last tarsal segment. Wing. Length: 3.0 – 3.1 mm. Upper side of basal costal cell with one long dark bristle. Fourth costal section 2 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2 – 3 distinct brown setulae on tegula. Abdomen. Viewed obliquely from front tergites subshiny black, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (larger on tergite 5). Dispersed short dark bristles on all tergites present, longest on tergite 5 up to 1 / 3 of the width of hind tibia at distal end. Lateral bristles on first tergite present, 5 – 6 dark bristles up to as long as hind femur’s width at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1.1 – 1.3 times as long as ST 8. Genitalia without dissection: ST 8 medium sized, broad, round in dorsal view, dark brown and with some distinct bristles, otherwise velvet-like coverage; membranous area slit-like, widening dorsally; epandrium dark brown, surstyli yellow, long. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.1). Surstyli elongated, broadened at base, straightened in middle, pointed at apex on inner margin, left surstylus slightly longer than right one (Fig. 2 A). Genital capsule in ventral view: gonopods elongated towards surstyli, tapering at apex, equal in height; subepandrial sclerite with 3 – 4 small fin-shaped protrusions in middle (Fig. 2 B); phallic guide with 3 – 4 dorso-dorsolateral spines. Genital capsule in lateral view: both surstyli curved towards sternite, right surstylus narrowed in apical quarter (Fig. 2 C); phallus with three phallic guides, one with one small spine (Fig. 2 E). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFD9EC62D8D9E956FCB616C8.taxon	etymology	Etymology: This species is named in honour of Alexander Skevington, JHS’s son, a keen young entomologist and one of the collectors of several specimens of this species. Alexander collected or was involved in the collection of 7 % of the specimens and 15 % of the species used in this revision (Supplementary file 2). Examined material: HOLOTYPE: AUSTRALIA: Queensland: Carnarvon N [ational] P [ark], summit of Fly Hill near West Branch Camp, 24 ° 58 ’ S, 147 ° 60 ’ E, 900 m, 12. X. 2002, J. Skevington, hand collected, JSS 12799 (1 ♂, QM); PARATYPES: AUSTRALIA: Queensland: Carnarvon National Park, Carnarvon Gorge, Consuelo land, 24 ° 56 ’ S, 148 ° 6 ’ E, land, S. Boucher, sweep net, CNCD 253914 (1 ♂, LEM); Carnarvon National Park, Mount Moffatt Section, southernmost of 3 Sisters, summit, 25 ° 7 ’ S, 148 ° 6 ’ E, hilltop, 23. I. 1998, J. & A. Skevington & S. Winterton, hand collected, JSS 2732; JSS 2735; JSS 2737; JSS 2742; JSS 2749 (5 ♂, QM); Mount Moffatt summit, 25 ° 4 ’ S, 148 ° 3 ’ E, 1097 m, 18. XI. 2014, J. H., A. M. and A. W. Skevington, CNC 374960; CNC 375009 - 10 (3 ♂, CNC); Carnarvon National Park; Summit of Fly Hill Near West Branch Camp, 24 ° 58 ’ S, 147 ° 60 ’ E, 900 m, 12. X. 2002, J. Skevington, JSM 467, CNC 484089 (1 ♂, CNC); Western Australia: Summit of Mount Barker, 34 ° 39 ’ S, 117 ° 39 ’ E, hilltop, 26. I. 2015, J. H., A. M. and A. W. Skevington, CNC 391301 (1 ♂, CNC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFD9EC62D8D9E956FCB616C8.taxon	distribution	Distribution: Australia (Queensland, Western Australia) (Fig. 107). Notes: Two highly disjunct populations have been documented based on specimens from Carnarvon National Park in Queensland and Mount Barker in Western Australia. Most specimens were captured hilltopping. Intraspecific genetic distance ranges from 0.0 – 3.9 % (Supplementary file 3). This may be a cryptic species complex with the same terminalia characters, so future genetic work is encouraged.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFDCEC60D8D9EFF6FF72167C.taxon	description	Figs 3 A – E, 107	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFDCEC60D8D9EFF6FF72167C.taxon	diagnosis	Diagnosis: This species can be recognized by long rectangular-shaped surstyli in dorsal view (Fig. 3 A); gonopods quadratic-shaped, the right one with a small finger-like projection on the right corner at apex in ventral view (Fig. 3 B); long subepandrial sclerite, one of the ejaculatory ducts with three long spines in ventral view (Fig. 3 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFDCEC60D8D9EFF6FF72167C.taxon	description	Description: MALE: Body length: 2.4 mm. Head. Scape, pedicel and arista dark brown. Scape with four short upper bristles and two short lower bristles. Flagellum tapering; brown, grey-brown pollinose. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput silvery pollinose. Thorax. Postpronotum pale, with 3 – 4 long pale bristles. Mesonotum (viewed obliquely from front) silvery pollinose, anterior part more grey, with some medium supra-alar bristles behind postpronotum lobes. Scutellum silvery brown pollinose and with 6 – 12 short dark bristles along distal edge. Halter, knob pale, stem and base dark. Legs. Mid coxa with 3 – 4 long dark bristles. Trochanters brown, femora black except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments dark dorsally and yellow ventrally. Hind trochanter without specific feature, grey pollinose with some scattered tiny bristles; two ventrobasal bristles present on fore and absent on mid femur. Hind femur without ventroapical spines, except 8 – 10 equally spaced sort bristles posteroventrally. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres not flattened, hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 2.5 mm. Upper side of basal costal cell with one short dark brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 1 – 2 long pale setulae on tegula. Abdomen. Viewed obliquely from front tergites brown, tergite 1 silvery grey. Lateral bristles on first tergite present, 4 – 5 light brown bristles up to as long as ¾ of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.2). Surstyli straight and elongated, left is longer than left one and a little widened before the apex (Fig. 3 A). Genital capsule in ventral view: both gonopods equal in height, quadratic-shaped, right one with a small finger-like projection on right corner at apex; phallic guide strong without dorso- dorsolateral spines; subepandrial sclerite long, phallus trifid, one of ejaculatory ducts with three long spins (Fig. 3 B); genital capsule in lateral view: both surstyli curved towards sternite, (Fig. 3 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 3 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFDCEC60D8D9EFF6FF72167C.taxon	etymology	Etymology: This species is named after Alice Daniels, one of the collectors of the holotype. Alice and Greg Daniels are prolific collectors and have contributed immensely to our understanding of Australian insects. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Elanda Point, Lake Cootharaba, 26 ° 14 ’ S, 153 ° 0 ’ E, 2. II. 1986, G. & A. Daniels, JSS 8917 (1 ♂, QM); PARATYPE: AUSTRALIA: New South Wales: Murray River 50 miles west of Wentworth, 34 ° 3 ’ S, 141 ° 3 ’ E, 22. XI. 1967, A. Neboiss, JSS 8798 (1 ♂, MVMA).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFDCEC60D8D9EFF6FF72167C.taxon	distribution	Distribution: Australia (New South Wales, Queensland) (Fig. 107). Notes: Two specimens are known from nearly 2000 km apart. We were unable to sequence the DNA from these two specimens and need more samples to test if this is a widespread species as hypothesized or two cryptic species.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC2EC7FD8D9EFF6FE7B1168.taxon	description	Figs 4 A – E, 108, 145 E	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC2EC7FD8D9EFF6FE7B1168.taxon	diagnosis	Diagnosis: This species can be recognized by two spines along posterior margin of sternites 3 and 4; hind femur with small posterior triangular keel (‘ shark fin’); left surstylus with a depression dorsomedially in lateral view (Fig. 4 A); hypandrium with long hypandrial apodeme in ventral view (Fig. 4 B); phallic guide with four spines dorsolaterally (Fig. 4 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC2EC7FD8D9EFF6FE7B1168.taxon	description	Description: MALE: Body length: 3.6 – 4.0 mm. Head. Flagellum acuminate, brown. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose; eyes touching for distance equal to 2 – 2.5 times the length of ocellar triangle. Occiput silvery pollinose, including upper half. Thorax. Postpronotum pale, with 6 – 8 distinct pale bristles. Mesonotum (viewed obliquely from front) subshining black, anterior part silver pollinose, slightly grayish also from the side. Scutellum silvery pollinose and with 12 – 14 indistinct pale bristles along the distal edge. Dorsocentral bristles firmly developed, dark, longer on frontal part and numerous dark bristles around postpronotum. Halter, knob pale with a brownish tinge distally, stem dark brown. Legs. Mid coxa with 2 – 3 long brown bristles at inner apical corner. Trochanters and base of femora brown, femora brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees, basal 1 / 5 of tibiae yellow (otherwise brown), tarsal segments yellow, last segment dorsally brown. Ventroapical row of spines on fore femur missing, 4 – 6 short, black spines on mid femur; no spines on hind femur, only 10 – 12 pale bristles posteriorly equally distributed, and longer on distal half (half as long as width of hind tibia at the distal end). Subapical (distal) short spines on first four tibiae present. Hind trochanter with small keel and comb-like structure with 8 – 10 spines (Fig. 145 E), one big round tooth posteriorly instead of a last spine; 1 – 3 long ventrobasal spines present on fore femur and mid femur. Hind femur with small triangular keel (‘ shark fin’) on the posterior side close to base. Hind tibia with a distinct row of semi-erect bristles along lateral edge. Hind metatarsus flattened (1, 2 and 3 tarsomeres especially), first tarsomere as long as 2 – 3 combined; ventral bristles scrub-like, yellow, dorsal bristles on tarsomeres 1 and 2 in dark row along posterior edge. Pulvilli as long as 2 / 3 of last tarsal segment. Wing. Length: 3.4 – 3.6 mm. Upper side of basal costal cell with one long distinct bristle. Fourth costal section 4 – 5 times as long as third costal section (3 rd costal section very short). Cross-vein r-m distinctly distal to middle of discal cell. 4 – 5 dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brownish pollinose, tergite 1 silvery grey, sides silvery pollinose on tergites 4 and 5 (the latter with side spots almost touching in the middle). Dispersed short dark bristles on tergites present, the longest on tergite 5 and up to twice as long as width of hind tibia at base. Lateral bristles on first tergite present, 8 – 10 black bristles up to twice as long as width of hind femur at distal tip. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 as long as ST 8. Sternites 3 and 4 with a pair of spines along posterior margin. Genitalia without dissection: ST 8 large, shiny black; membranous area long, slit-like, dividing ST 8 completely; epandrium paler, narrow; yellow surstyli appear long and flat, often visible in lateral view showing can-opener-shaped margin of left surstylus. Genitalia. Genital capsule in dorsal view: epandrium dark brown, surstyli light brown. Epandrium wider than long (MLE: MWE = 0.6). Surstyli rather symmetrical, rectangular-shaped with broad base, left surstylus longer than right one (Fig. 4 A). Genital capsule in ventral view: subepandrial sclerite wide, extended towards epandrium with a small protrusion towards phallic guide; gonopods flat, elongated towards surstyli (Fig. 4 B), phallic guide with four spines dorsolaterally (three in left and one in right side); hypandrium with long hypandrial apodeme (Fig. 4 B). Genital capsule in lateral view: both surstyli curved towards sternite, left surstylus with a depression dorsomedially (Fig. 4 D); phallus with three ejaculatory ducts; ejaculatory apodeme flat, sperm pump elongated, linear (Fig. 4 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC2EC7FD8D9EFF6FE7B1168.taxon	etymology	Etymology: The name is a combination of the Greek amphi, around, on both sides, double and the Latin spina, thorn, in reference to the paired spines along the posterior margin of sternites 3 and 4. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: M [oun] t Magnet, M [oun] t Waramboo, 28 ° 2 ’ S, 117 ° 49 ’ E, hilltop, 14. xii. 1999, J. Skevington, JSS 7434 (WAM); PARATYPES: AUSTRALIA: Western Australia: same data as holotype, JSS 7429 - 30, JSS 7432, JSS 7451 – 2, JSS 7454 – 62, JSS 7465, JSS 7467 – 8, JSS 7472 – 80, JSS 7482 (2 ♂ ANIC, 2 ♂ CNC, 20 ♂ QM, 2 ♂ USNM; 2 ♂ WAM); South Australia: Great Victoria Desert, vehicle net between 183 – 217 km South Vokes Hill Corner, [27 ° 52 ’ S, 130 ° 22 ’ E], 29. viii. 1980, J. Forrest, JSS 8704 (1 ♂, SAM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC2EC7FD8D9EFF6FE7B1168.taxon	distribution	Distribution: Australia (South Australia, Western Australia) (Fig 108). Notes: This desert species is presumably widespread but has only been collected in two widely separated localities. Most of the specimens were captured hilltopping. Based on the DNA barcode, this species is genetically most similar to T. spinifex sp. nov. (1.9 – 3.5 % pairwise divergence). Intraspecific genetic distance ranges from 0.0 % to 0.2 % (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC0EC7DD8D9EFF6FC1C16EC.taxon	description	Figs 5 A – F, 108, 148 B, 154 C	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC0EC7DD8D9EFF6FC1C16EC.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter with a few dark, short spines in a group ventrobasally and the flat ventral side with shorter and fewer spines (Fig. 148 B); elongated surstyli in dorsal view (Fig. 5 A); phallic guide with some dorso-dorsolateral spines (Fig. 5 F); elongated gonopods with two inwards projections (Fig. 5 B); hypandrium with two hairy membranous sheaths (Fig. 5 D – F); subepandrial sclerite with a small fin-shaped protuberance (Fig. 5 B); one of ejaculatory ducts with some small spines at base (Fig. 5 D – F).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC0EC7DD8D9EFF6FC1C16EC.taxon	description	Description: MALE: Body length: 3.1 – 3.3 mm. Head. Flagellum short acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 – 2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 5 – 7 pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum brownish pollinose and with 6 – 8 bristles along the distal edge. Dorsocentral bristles distinct, dark, somewhat longer along frontal edge, dark bristles around postpronotum present. Halter, knob pale, stem brown. Legs. Mid coxa with 3 – 4 long dark bristles. Trochanters and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise black), tarsal segments dark brown, ventrally yellowish, last segment darker. Hind trochanter with few dark short spines (3 – 8) in a group at the base ventrally and with a flat ventral side (a few spines are possible on the flat part as well) (Fig. 148 B); ventrobasal spines (1) present on fore femur. Hind tibia slightly bent in posterior view. Ventroapical row of spines on fore femur 4 – 5; 4 – 5 small, black spines on mid femur; hind femur without ventroapical spines, except 6 – 8 equally spaced bristles posteroventrally on distal half, longest up to 2 / 3 the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres extensively flattened, hind metatarsus almost as long as 2 – 5 combined and with scrub-like bristles ventrally (tarsi 1 – 3). Pulvilli shorter than last tarsal segment. Wing. Length: 3.0 – 3.1 mm. Upper side of basal costal cell with one long dark bristle and 2 – 3 short brown bristles. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2 – 3 distinct brown setulae on tegula. Abdomen. Viewed obliquely from front tergites shining brown, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (larger on t 5). Dispersed strong dark bristles on all tergites present, longest on tergite 5 up to the width of hind tibia at distal end. Lateral bristles on first tergite present, 5 – 6 dark bristles up to 1.2 – 1.4 times as long as hind femur’s width at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1.2 – 1.4 times as long as ST 8. Genitalia without dissection: ST 8 medium sized rounded in dorsal view, brown and with distinct bristles, otherwise velvet-like coverage; membranous area with narrow part narrowing epandrium and rounded part mostly directed dorsally; epandrium brown, surstyli yellow, long and slim, often visible completely in lateral view (nothing is blocking the view). Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.3). Surstyli rather symmetrical, widened at base, elongated with rounded tips, right surstylus smaller than left one, pointed at inner margin before apex (Fig. 5 A). Genital capsule in ventral view: phallic guide straight with 3 – 4 dorso-dorsolateral spines (one is longer), gonopods equal in height, elongated towards surstyli with two small inward projections before apex, hypandrium with two hairy membranous sheaths extended towards gonopods; subepandrial sclerite rather wide with a small fin-shaped protuberance in its center; phallus with three ejaculatory ductus, one with four small spines (Fig. 5 B). Genital capsule in lateral view: both surstyli straight, abruptly bent in 90 ° in apical fourth, left is longer than right one (Fig. 5 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 5 C). FEMALE: Body length: 3.2 mm. As male except for the following characters. Frons with eyes separated, as wide as 1.6 – 1.8 times the width at antennae; completely silver-grey pollinose until up to 1 / 3 of the lower frons, shiny black in front of ocellar triangle, no transition zone; frons depressed along upper 2 / 3 with a groove in the median line. Enlarged ommatidia silvery, shining. Pulvilli and claws about 1.5 – 2 times as long as last tarsal segment, smaller on metatarsi. Female abdomen with silver spots on sides of tergites 3 – 6. Ovipositor. Base of ovipositor brownish pollinose with dispersed small bristles, piercer straight, thin and reaching distal end of 1 st segment. LP: LB = 2.3. LDP: LPP = 2.5. (Fig. 154 C).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC0EC7DD8D9EFF6FC1C16EC.taxon	etymology	Etymology: This species is named in honour of Angelika (Angela) Skevington who collected or helped collect 35 % of the species and 17 % of the specimens used in this revision and several of the specimens of this species (Supplementary file 2). Examined material: HOLOTYPE: AUSTRALIA: Western Australia: Karijini National Park, Juna Downs Road, 22 ° 44 ’ S, 118 ° 25 ’ E, 798 m, dry Turee Creek, grassy open Eucalyptus scrub, 14 – 19. v. 2003, C. Lambkin & T. Weir, Malaise trap, ANIC [Bulk Sample] 2145, JSS 16110 (1 ♂, ANIC); PARATYPES: AUSTRALIA: New South Wales: Warrumbungle National Park, 31 ° 16 ’ S, 149 ° 0 ’ E, in creek bed [in Eucalyptus forest], 17 – 22. xii. 1998, J. & A. Skevington & S. Winterton, Malaise trap, JSS 5064 (1 ♀, QM); Northern Territory: Litchfield, 4 WD camping area; about 300 m in on 4 WD map, 13 ° 2 ’ S, 130 ° 55 ’ E, xii. 1997 – i. 1998, A. Zwick, JSS 8846 (1 ♂, QM); Queensland: 2 km S of Horseshoe lookout, Blackdown land, 23 ° 50 ’ S, 149 ° 3 ’ E, 23 – 24. IV. 1981, D. H. Colless, JSS 8613 (1 ♂, ANIC); 3 km NE of Mount Webb, 15 ° 3 ’ S, 145 ° 9 ’ E, 30. IV. 1981, D. H. Colless, Malaise trap, JSS 8610 (1 ♂, ANIC); Culgoa Floodplain National Park, 11.8 km North of Headquarters (CG 2 AM), 28 ° 54 ’ S, 147 ° 8 ’ E, 151 m, Brigalow Black Box, 17. XII. 2009 – 20. I. 2010, C. Kelly, A. Coward, Malaise trap, 19263, CNC 574521, CNC 576695 – 7, CNC 576679 (4 ♂, CNC, 1 ♂ USNM); 91.2 km west Windorah, 25 ° 22 ’ S, 141 ° 47 ’ E, Malaise in dry creek bed, 31. viii – 5. ix. 1997, S. Winterton, J. & A. Skevington, JSS 8396 (1 ♂, QM); Brisbane, Mount Coot-tha, 27 ° 29 ’ S, 152 ° 57 ’ E, 170 m, hilltop, 15. v. 1999, J. & A. Skevington, hand collected, JSS 5773 (1 ♂, QM) ;; Kumbia-Bunya Mountains Road, 26 ° 48 ’ S, 151 ° 34 ’ E, hilltop, 17. I. 1998, C. Lambkin, JSS 8848 (1 ♂, QM); South Australia: Flinder’s Ranges National Park, Brachina Gorge, Heysen Hilltop, 31 ° 20 ’ S, 138 ° 33 ’ E, hilltop, 9. x. 1997, J. & A. Skevington & C. Lambkin, JSS 8383, JSS 8386 (2 ♂, QM); Gammon Ranges National Park, south end of Weetootla Gorge, 30 ° 29 ’ S, 139 ° 16 ’ E; ~ 537 m, hilltop, 11. x. 1997, J. Skevington & C. Lambkin, JSS 8351, JSS 8359, JSS 8365 (3 ♂, QM); Western Australia: 82 km S junction Karijini Drive on Great Northern Highway, 23 ° 7 ’ S, 119 ° 6 ’ E, 694 m, wash with pools, 23. iv – 6. v. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15673 (1 ♂, CNC); Karijini National Park, Juna Downs Road, 22 ° 44 ’ S, 118 ° 25 ’ E, 798 m, dry Turee Creek, grassy open Eucalyptus scrub, 14 – 19. v. 2003, C. Lambkin & T. Weir, Malaise trap, ANIC [Bulk Sample] 2145, JSS 16105 (1 ♂, ANIC); Karijini National Park, Juna Downs Road, 22 ° 42 ’ S, 118 ° 25 ’ E; 789 m; between drying pools in rocky Turee Creek; near rocky outcrop, Eucalyptus Eremophila scrub, 19 – 25. iv. 2003, C. Lambkin & T. Weir, Malaise trap, ANIC [Bulk Sample] 2059; JSS 16254, (1 ♂, ANIC); Karijini National Park, Juna Downs Road, 22 ° 43 ’ S, 118 ° 25 ’ E, 799 m, ANIC [Bulk Sample] 2039, JSS 16179 (1 ♀, ANIC); Karijini National Park, Juna Downs Road, 22 ° 44 ’ S, 118 ° 25 ’ E, 798 m, dry Turee Creek, grassy open Eucalyptus scrub, 25. iv – 14. v. 2003, C. Lambkin & T. Weir, Malaise trap, ANIC [Bulk Sample] 2044, JSS 16085, JSS 16098, JSS 16100 (1 ♂, 2 ♀, ANIC); Mount Augustus National Park; south slope on Saddle Trail, 24 ° 18 ’ S, 116 ° 48 ’ E, 400 m, small dry wooded wash, 25. iv – 7. v. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15885 (1 ♀, CNC); Martin’s Well, West Kimberley, 16 ° 34 ’ S, 122 ° 51 ’ E, 28. IV. 1977, D. H. Colless, Malaise trap, JSS 9258 (1 ♂, ANIC); Northern Territory: Litchfield National Park, 13 ° 30 ’ S, 130 ° 36 ’ E, 20. I. 1998, A. Zwick, debu 265568 (1 ♀, DEBU); Tasmania: Mount William National Park, 40 ° 53 ’ S, 148 ° 13 ’ E, across dry creek bed, 23. XII. 0 – 10. I. 1999, J. & A. Skevington, Malaise trap, JSS 5073 (1 ♂, QM); Western Australia: Cape Borda to Pender, West Kimberley, 16 ° 0 ’ S, 126 ° 0 ’ E, 24. IV. 1977, D. H. Colless, JSS 9172 (1 ♂, ANIC); Karijini National Park, Weano Gorge Road, 22 ° 22 ’ S, 118 ° 15 ’ E, 775 m, on hilltop, open Eucalyptus, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16139 (1 ♂, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC0EC7DD8D9EFF6FC1C16EC.taxon	distribution	Distribution: Australia (New South Wales, Northern Territory, Tasmania, Queensland, South Australia, Western Australia) (Fig. 108). Notes: This species is found throughout most of Australia in a variety of habitat types, from Eucalyptus - dominated to extremely arid, Acacia - dominated areas. Some specimens have been found hilltopping. This species is genetically closest to T. pterygia sp. nov., differing by 2.5 – 2.7 % (pairwise divergence). Intraspecific genetic distance ranges from 0.0 % to 2.0 % (Supplementary file 3). This may be a species complex with two taxa including JSS 8365, JSS 8386 (species group 1) collected from South Australia and the rest of the specimens along with JSS 8351 from SA (species group 2) (Supplementary Fig. 5). As morphological features, particularly genitalia features are similar, more molecular work and specimens are needed to shed light on this.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC7EC78D8D9EFF6FB25108C.taxon	description	Figs 6 A – D, 109, 149 F, 156 A	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC7EC78D8D9EFF6FB25108C.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter with a keel covering 2 – 3 short dark spines at the proximal end (Fig. 149 F); elongated surstyli in dorsal view (Fig. 6 A); extended gonopods towards surstyli, left longer than right one; distinct subepandrial sclerite with fin-shaped projection; phallic guide with 5 – 6 dorso-dorsolateral spines in ventral view (Fig. 6 B); left surstylus narrowed in apical half in lateral view (Fig. 6 C).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC7EC78D8D9EFF6FB25108C.taxon	description	Description: MALE: Body length: 2.6 – 2.7 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 3 – 4 distinct pale bristles. Mesonotum (viewed obliquely from front) silvery pollinose, anterior part more grey, grayish also from the side. Scutellum silvery brown pollinose and with 5 – 7 bristles along distal edge. Dorsocentral bristles short, dark, longer in frontal half, a few pale bristles around postpronotum present. Halter, knob pale, stem brown. Legs. Mid coxa with 2 – 3 long dark bristles. Trochanters brown, femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 5 of tibiae yellow (tibiae otherwise brown), tarsal segments brown dorsally, yellow ventrally, last segment darker. Hind trochanter with 45 degree keel, 2 – 3 short dark spines at the proximal end (Fig. 149 F); ventrobasal spines absent on fore and mid femur. Ventroapical row of spines on fore femur missing; 4 – 7 distinct, black spines on mid femur; hind femur without ventroapical spines, except 10 – 12 equally spaced bristles posteroventrally, longer on distal half, longest up to 1 / 3 the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres flattened (especially hind metatarsus on distal half), hind metatarsus as long as 2 – 3 combined. Pulvilli shorter than last tarsal segment. Wing. Length: 2.7 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2 – 3 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites subshiny, brown-black, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (spot on tergite 5 larger). Dispersed strong dark bristles on all tergites present, longest on tergite 5 in length up to 1 / 4 the width of hind tibia at distal end. Lateral bristles on first tergite present, 4 – 6 dark bristles up to as long as ¾ of hind femur’s width at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1 – 1.2 times as long as ST 8. Genitalia without dissection: ST 8 small to medium sized, round, globular, brown and with distinct bristles, otherwise with velvet-like coverage; membranous area is round, mostly directed posteriorly; epandrium brown, surstyli brown. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.2). Surstyli elongated, symmetrical, left surstylus longer than right one and wider at apex, right surstylus with a distinct triangle-shaped protrusion at inner margin before apex (Fig. 6 A). Genital capsule in ventral view: both gonopods extended towards surstyli, left is longer than right one, right one with a projection on inner margin; phallic guide with 5 – 6 dorso- dorsolateral spines, one spine longer than others; subepandrial sclerite distinct with a small fin-shaped protrusion extended from basal to its center (Fig. 6 B); genital capsule in lateral view: both surstyli curved towards sternite, left surstylus narrowed in apical half (Fig. 6 C). FEMALE: Body length: 2.5 mm. As male except for the following characters. Frons, eyes separated, as wide as 1.2 – 1.4 times the width at antennae; completely silver-grey pollinose on lower 1 / 3, shiny black only around directly ocellar triangle, smooth change in transition zone (just before ocellar triangle) to shiny black. Enlarged ommatidia silvery shining. Pulvilli and claws about 1.5 – 2.0 times as long as last tarsal segment, smaller on metatarsi. Female abdomen with silver spots on sides of tergites 4 – 6; Ovipositor. Straight in ventral view, straight, dagger-like in lateral view (yellow piercer, brown base), reaching distal end of 2 nd segment; base silvery brown pollinose with dispersed short dark bristles. LP: LB = 1.3. LDP: LPP = 4.30. (Fig. 156 A).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC7EC78D8D9EFF6FB25108C.taxon	etymology	Etymology: From the Latin angustus, narrow, in reference to the narrowed left surstylus. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: Millstream-Chichester Nat [ional] P [ark], Roebourne Road, 21 ° 26 ’ S, 117 ° 9 ’ E, 337 m, sandy creek, Eucalyptus Spinifex grassland, 27. IV – 3. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16196 (1 ♂, WAM); PARATYPES: AUSTRALIA: Western Australia: 158 km S Newman, 9 km N Kumarina Road House, 24 ° 38 ’ S, 117 ° 37 ’ E, 638 m, wide sandy wash, 7 – 18. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15726 (1 ♂, CNC); 171 km East of Marble Bar, 21 ° 17 ’ S, 121 ° 14 ’ E, 300 m, dry wash with flowers, 2 – 15. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15870, JSS 15872 (2 ♀, CNC); Karijini National Park, Karijini Drive, 22 ° 34 ’ S, 118 ° 18 ’ E, 814 m, open Eucalyptus grassland, 19 – 25. IV. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16222 – 3 (1 ♂, 1 ♀, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC7EC78D8D9EFF6FB25108C.taxon	distribution	Distribution: Australia (Western Australia) (Fig. 109). Notes: This species appears to be a Western Australian endemic and is found in Eucalyptus dominated habitats to desert washes. Based on the DNA sequences, this species is genetically close to T. patula sp. nov. (6.7 % pairwise divergence). Intraspecific genetic distance ranges from 0.2 % to 1.4 % (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC5EC79D8D9EFF6FA161738.taxon	description	Figs 7 A – D, 108, 147 C, 151 C – D	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC5EC79D8D9EFF6FA161738.taxon	diagnosis	Diagnosis: This species can be recognized by the peg-like projection and dense dark bristles on the hind trochanter (Fig. 147 C); the unusual shape of the distal tarsomere of the fore tarsus, with a handle-shaped base that gradually widens towards the tip (Fig. 151 C); abdominal sternites 2 – 5 with distinct dark spines along posterior margin; surstyli elongated in dorsal view (Fig. 7 A); left gonopod longer than right one (Fig. 7 B); phallic guide with five spines dorsal-dorsolaterally, one longer than others.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC5EC79D8D9EFF6FA161738.taxon	description	Description: MALE: Body length: 4.1 mm. Head. Flagellum acuminate, yellow. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose with a golden tinge; eyes touching for distance equal to 3.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper half. Thorax. Postpronotum pale, slightly brownish, with 15 – 17 distinct long pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part slightly silver pollinose; grayish also from the side. Scutellum brownish pollinose and with 20 – 22 long bristles along the distal edge. Dorsocentral bristles undeveloped, indistinct, numerous pale bristles around postpronotum. Halter, knob pale, stem brown. Legs. Mid coxa with 4 – 6 extra long, dark bristles, hind coxa 4 – 5 shorter dark bristles. Trochanters and femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees, and distal extremity of tibiae yellow-brown (tibiae otherwise black), tarsal segments dark brown, yellowish ventrally. Hind trochanter without a keel, but with a distinct, blunt peg on the distal end with a few dark bristles (8 – 10) basally to the peg (Fig. 147 C). Ventroapical row of spines on fore femur missing, a patch of 6 – 7 short dark spines are present at 1 / 3 from distal end; 3 – 4 short, black spines on mid femur; no spines on hind femur, only 12 – 14 dark bristles posteriorly on distal half positioned on a slight keel (up to as long as half the width of hind tibia at the distal end). Fore and mid femora with 6 – 8 ventrobasal bristles. Subapical (distal) spines on first four tibiae absent. Hind tibia covered with dense long dark bristles ventroapically; last tarsomere of fore tarsi longer than other tarsomeres, with a long handle-type base and gradually widening towards tip, provided with strong dark bristles dorsally; the segment itself is bent (Fig. 151 C). Hind tarsomeres all flattened, hind metatarsus almost as long as 2 – 4 combined, last tarsomere long and broad, almost as broad and longer than tarsomere 2 (Fig. 151 D); bristles are missing dorsally on hind metatarsus in the median line but are present anterior and extensively on posterior edge. Pulvilli 1 / 3 as long as last tarsomere of fore tarsi, as long as last tarsomere of mid tarsi and half as long as last tarsomere of hind tarsi. Wing. Length: 4.2 mm. Upper side of basal costal cell with two long light bristles. Fourth costal section 3.5 – 4 times as long as third costal section (3 rd section appears to be unusually short). Cross-vein r-m at 3 / 5 of discal cell, distal to middle. 7 – 8 very strong, dark setulae on tegula. Abdomen. Dissected. Tergite 1 silvery grey. Lateral bristles on first tergite present, 16 – 18 long pale bristles in a group (as seen in species of Claraeola Aczél) up to 1.6 – 1.8 times as long as the width of hind femur at distal tip in lateral view. Sternites 2 – 5 with distinct dark spine along posterior margin in each segment. Postabdomen in dorsal view: Dissected. Genitalia. Genital capsule in dorsal view: epandrium and surstyli dark brown. Epandrium wider than long (MLE: MWE = 0.6). Surstyli rather symmetrical, broadened at base, elongated, pointing towards each other at apex, left is wider than right one (Fig. 7 A). Genital capsule in ventral view: gonopods unequal, left is longer than right one, hypandrium broad, subepandrial sclerite rectangular-shaped (Fig. 7 B), phallic guide with five spines dorso- dorsolaterally (the longest one twice as length as others); genital capsule in lateral view: both surstyli curved towards sternite (Figs 7 C – D). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC5EC79D8D9EFF6FA161738.taxon	etymology	Etymology: The Latin ansa for handle or haft, refers to the handle-shaped distal fore tarsomere. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Cudmore National Park (CM 1 M), 22 ° 54 ’ S, 146 ° 21 ’ E, 365 m, Eucalyptus woodland with Spinifex, 27. x. 2010 – 2. viii. 2011, Lambkin, Starick & Bailey, Malaise trap, CNC 591870 (1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC5EC79D8D9EFF6FA161738.taxon	distribution	Distribution: Australia (Queensland) (Fig. 108). Notes: The only known specimen was collected in central Queensland in Eucalyptus woodland with Spinifex.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFCBEC77D8D9EFF6FF5D1624.taxon	description	Figs 8 A – E, 109, 145 A	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFCBEC77D8D9EFF6FF5D1624.taxon	diagnosis	Diagnosis: This species can be recognized by hind trochanter with 6 – 10 short dark bristles in a row from ventrobasally to ventroapically (Fig. 145 A); tiny surstyli, upright in dorsal view (Fig. 8 A); gonopods extended towards surstyli, rounded at apex with a distinct projection on inner margin; phallic guide with 2 – 4 short dorso- dorsolateral spines, one of ejaculatory ducts with some short teeth in basal half in ventral view (Fig. 8 B, D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFCBEC77D8D9EFF6FF5D1624.taxon	description	Description: MALE: Body length: 2.7 mm. Head. Scape, pedicel dark brown, flagellum acuminate, yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum yellow, with 3 – 4 pale bristles. Mesonotum (viewed obliquely from front) silvery brown pollinose, anterior part without pollinosity. Dorsocentral bristles short, pale, numerous pale bristles around postpronotum present. Scutellum silvery brown pollinose and with 4 – 8 short dark bristles. Halter, knob pale, tip brownish, stem brown. Legs. Mid coxa with 3 – 4 long dark bristles. Fore and mid trochanter with two short dark bristles ventroapically; trochanters brown, femora black, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments bright brown. Hind trochanter with 6 – 10 short dark bristles in a row from ventrobasally to ventroapically (Fig. 145 A); ventrobasal spines present (2) on fore and mid femur. Hind femur without ventroapical spines, except 8 – 10 equally spaced bristles posteroventrally. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres gently flattened (t 1 – 3), hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 2.7 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. Abdomen. Viewed obliquely from front tergites brown-black, tergite 1 silvery grey. Dispersed short dark brown bristles on all tergites present. Lateral bristles on first tergite present, 3 – 5 dark bristles up to as long as ¾ of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.7). Surstyli tiny (rather hard to see in dorsal view), upright, widened at the base, angled straight upwards in apical half (Fig. 8 A). Genital capsule in ventral view: both gonopods equal in height, extended towards surstyli, rounded at apex, gonopods with a distinct projection on inner margin; phallic guide with 2 – 4 short dorso- dorsolateral spines; phallus trifid, one of ejaculatory ducts with some teeth in basal half; subepandrial sclerite distinct (Fig. 8 B); Genital capsule in lateral view: both surstyli straight in basal two thirds, bent in apical third (Fig. 8 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 8 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFCBEC77D8D9EFF6FF5D1624.taxon	etymology	Etymology: This species name refers to Arnhem Land, a historical region of the Northern Territory of Australia where the only known specimen was collected. Examined material: HOLOTYPE: AUSTRALIA: Northern Territory: Arnhem Land, Maningrida, 12 ° 3 ’ S, 134 ° 13 ’ E, 5 m, 20 – 21. III. 1961, L. & M. Gressitt, CNCD 5563 (1 ♂, BPBM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFCBEC77D8D9EFF6FF5D1624.taxon	distribution	Distribution: Australia (Northern Territory) (Fig. 109). Notes: This species is known from a single specimen from a coastal area near a river mouth in the Northern Territory. Based on this limited information, it should be looked for in either mangroves or coastal monsoon forest.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC9EC73D8D9EFF6FE9512F0.taxon	description	Figs 9 A – D, 109, 146 A	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC9EC73D8D9EFF6FE9512F0.taxon	diagnosis	Diagnosis: This species can be recognized by a patch of dark spines on hind trochanter (Fig. 146 A); a pair of spines along the posterior margin of sternite 5; elongated surstyli bent in apical third, both surstyli with a small projection at base along inner margin in dorsal view (Fig. 9 A); left surstylus bent towards sternite in apical third in lateral view (Fig. 9 C); three spines on phallic guide (Fig. 9 A); elongated gonopods in ventral view (Fig. 9 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC9EC73D8D9EFF6FE9512F0.taxon	description	Description: MALE: Body length: 2.6 – 3.0 mm. Head. Flagellum acuminate, brown. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose with golden tinge; eyes touching for distance equal to 2 – 2.5 times the length of ocellar triangle. Occiput silvery pollinose, more black on upper half. Thorax. Postpronotum pale, with 3 – 5 distinct brown bristles. Mesonotum (viewed obliquely from front) subshining black, anterior part silver pollinose, slightly grayish also from the side. Scutellum subshining black and with 6 – 7 distinct dark bristles along the distal edge. Dorsocentral bristles firmly developed, dark, longer on frontal part and numerous dark bristles around postpronotum. Halter, knob pale with a brownish tinge distally, stem dark brown. Legs. Mid coxa with 2 – 3 bristles. Trochanters and base of femora brown, femora brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees, basal 1 / 6 of tibiae yellow (otherwise brown), tarsal segments yellow ventrally, brown dorsally, last segment dorsally darker. Fore femur with two distinct ventrobasal bristles, without ventroapical row of spines, 8 – 10 short, black spines on mid femur; no spines on hind femur, only 7 – 9 pale bristles posteriorly equally distributed on distal 1 / 3 (twice as long as width of hind tibia at the distal end). Subapical (distal) spines on first four tibiae present (short). Hind trochanter without a keel, but with a group of short dark spines basally “ echidna ” style (Fig. 146 A). Hind metatarsus not flattened, first tarsomere as long as 2 – 4 combined; ventral bristles less pronounced, but scrub-like, yellow, dorsal bristles on all metatarsi semi-erect and dark. Pulvilli as long as 2 / 3 of last tarsal segment. Wing. Length: 2.5 – 3.0 mm. Upper side of basal costal cell with one long distinct bristle. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2 – 3 dark setulae on tegula. Abdomen. Viewed obliquely from front tergites dark brownish pollinose, tergite 1 silvery grey, sides silvery pollinose on tergites 4 and 5 (the latter with somewhat larger spots). Dispersed short dark bristles on tergites present, the longest on tergite 5 and up to as long as width of hind tibia at base. Lateral bristles on first tergite present, 5 – 6 brown bristles up to 1.5 as long as width of hind femur at distal tip. Sternite 5 with a pair of spines along posterior margin. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 as long as 1.5 times the length of ST 8. Genitalia without dissection: ST 8 medium sized, subshiny black; membranous area broad slit-like, in the shape of a sunflower seed; epandrium paler brown, narrow; yellow surstyli with curly shape discernible. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.8). Surstyli light brown, rather symmetrical, elongated, left surstylus longer and broader than right one, its tip bent toward right one in apical third (Fig. 9 A), both surstyli with a small projection ventrobasally. Genital capsule in ventral view: subepandrial sclerite wide, extended beneath gonopods, covered with transverse streaks; gonopods elongated close to apex of subepandrial sclerite, with a small inward projection (Fig. 9 B), phallic guide with three different-sized spines dorsally (longer one closer to apex); genital capsule in lateral view: left surstylus straight in basal two thirds, bent towards sternite in apical third, right surstylus smoothly curved towards sternite (Fig. 9 D); phallus with three ejaculatory ducts. FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC9EC73D8D9EFF6FE9512F0.taxon	etymology	Etymology: The name is derived from the Latin words atra, black, and spina, spine, in reference to the patch of dark spines on the trochanter. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Barakula State Forest No. 302, Summit Round Mountain, 26 ° 22 ’ S, 150 ° 58 ’ E, 8. i. 2000, J. & A. Skevington & M. Mathieson, JSS 7555 (1 ♂, QM); PARATYPES: AUSTRALIA: Australian Capital Territory: Black Mountain, Canberra, 35 ° 16 ’ S, 149 ° 6 ’ E, 18 – 23. ii. 1985, W. W. Middlekauff, flight interception trap, JSS 8448 (1 ♂, CAS); Queensland: Barakula State Forest No. 302, Summit Round Mountain, 26 ° 22 ’ S, 150 ° 58 ’ E, 8. i. 2000, J. & A. Skevington & M. Mathieson, JSS 7554 (1 ♂, QM); Bribie Island, QDPI Fisheries Site, 27 ° 3 ’ S, 153 ° 11 ’ E, Heathland- Acacia regrowth, 17 – 24. x. 1997, S. Winterton, N. Power & D. White, Malaise trap, JSS 8434 (1 ♀, QM); Brisbane, Mount Coot-tha, 27 ° 29 ’ S, 152 ° 57 ’ E, 170 m, [hilltop], 1. xi. 1997, C. Lambkin, hand collected, JSS 3804 (1 ♂, QM); 170 m, hilltop, 15. v. 1999, J. & A. Skevington, hand collected, JSS 5781 (1 ♂, QM); 16. ix. 1997, J. Skevington, hand collected, JSS 3799 (1 ♂, QM); ~ 170 m, hilltop, 27. i. 1998, J. & A. Skevington, hand collected, JSS 3050 – 1, JSS 3088 (3 ♂, CNC, USNM); Brisbane, Mt. Coot-tha, 27 ° 29 ’ S, 152 ° 57 ’ E, ~ 170 m, hilltop, 7. xii. 1997, J. & A. Skevington, hand collected, JSS 2328, JSS 2331, JSS 2334, JSS 2337 – 8, JSS 2342, JSS 2344 – 9, JSS 2353 – 7 (17 ♂, QM); Carnarvon National Park, Hilltop north of Tombs, 25 ° 5 ’ S, 147 ° 52 ’ E, hilltop, 26. xi. 1997, J. Skevington & C. Lambkin, hand collected, JSS 2203, JSS 2211 (2 ♂, QM); Tasmania: Freycinet National Park, 42 ° 8 ’ S, 148 ° 19 ’ E, in heath with scattered eucalypts, 26. xii. 1998 – 10. i. 1999, J. & A. Skevington, Malaise trap, JSS 4671 (1 ♂, QM); Western Australia: Lake Muir Nature Reserve, Lake Muir Road, 34 ° 26 ’ S, 116 ° 38 ’ E, 202 m, Jarrah woodland with Macrozamia & grasstrees, 3 – 14. xi. 2003, C. Lambkin, Malaise trap, JSS 15954 (1 ♂, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFC9EC73D8D9EFF6FE9512F0.taxon	distribution	Distribution: Australia (Queensland, Tasmania, Western Australia) (Fig. 109). Notes: This species is found in eucalypt dominated habitats and is regularly encountered hilltopping. It is genetically most similar to T. trispina sp. nov. (5.1 – 6.1 % pairwise divergence) and intraspecific genetic distance ranges from 0.0 % to 3.3 % (Supplementary file 3). The specimens JSS 15954 and JSS 7555 collected from Western Australia and Queensland, show a greater pairwise distance (2.9 % and 3.3 %, respectively) compared to the others, although their morphological features are consistent with those of the other specimens. This suggests that they might be part of a cryptic species complex. Additional fresh specimens and molecular work are needed to support this hypothesis.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFCFEC71D8D9EDB7FB3112DC.taxon	description	Figs 10 A – E, 110, 148 A	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFCFEC71D8D9EDB7FB3112DC.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having two spur-like projections (Fig. 148 A); surstyli elongated in dorsal view (Fig. 10 A); phallic guide with six spines; phallus with three short ejaculatory ducts and two long membranous sheaths in ventral view (Fig. 10 E); hypandrial apodeme L-shaped in lateral view (Fig. 10 C – D)	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFCFEC71D8D9EDB7FB3112DC.taxon	description	Description: MALE: Body length: 3.1 – 3.3 mm. Head. Flagellum acuminate; brown. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 – 2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 2 – 3 pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum brownish pollinose and with 4 – 6 weak bristles along the distal edge. Dorsocentral bristles indistinct, somewhat more visible along frontal edge, numerous pale bristles around postpronotum. Halter, knob pale, stem brown. Legs. Mid coxa with 1 – 2 dark spine-like bristles. Trochanters and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 5 of tibiae yellow (tibiae otherwise dark brown), tarsal segments yellow-brown, ventrally somewhat paler, last segment darker. Hind trochanter with double spurs, anterior is straighter and needle-like (Fig. 148 A); ventrobasal bristle (one) on fore and mid femora. Ventroapical row of spines on fore femur absent; 5 – 6 small, black spines on mid femur; hind femur without ventroapical spines, except 12 – 13 equally spaced bristles posteroventrally, longer on the distal half, in length up to 1.2 – 1.4 times the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae absent. Metatarsi: hind metatarsus flattened, and almost as long as 2 nd and both with scrub-like bristles ventrally. Pulvilli shorter than last tarsal segment. Wing. Length: 2.9 – 3.0 mm. Upper side of basal costal cell with one short dark bristle. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m at middle of discal cell. 1 – 2 small brown setulae on tegula. Abdomen. Viewed obliquely from front tergites subshining black, tergite 1 silvery grey, sides silvery pollinose on tergites 3, 4 and 5 (largest on tergite 5). Dispersed strong dark bristles on all tergites present, longest on tergite 5 up to 0.5 times the width of hind tibia at distal end. Lateral bristles on first tergite present, 5 – 6 pale bristles up to 1.5 times as long as hind femur’s width at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1.2 times as long as ST 8. Genitalia without dissection: ST 8 medium sized square shaped in dorsal view, brown and with the occasional abdominal bristles, otherwise velvet-like coverage; membranous area elongated, widening dorsally; epandrium brown, surstyli pale brown, long and broad. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.8); surstyli symmetrical, rectangular shaped with pointing tips at its apical inner corner, left surstylus slightly longer and wider than right one (Fig. 10 A). Genital capsule in ventral view: phallic guide straight and slender, phallus with three ejaculatory ducts and with two membranous sheaths covered by some small condensed hairs at apex; gonopods elongated towards surstyli, equal in height, subepandrial sclerite long at apex and widened at base; hypandrium with a small hairy membranous sheath extended towards gonopod (Fig. 10 B). Genital capsule in lateral view: both surstyli curved towards sternite, more broadened at middle; hypandrial apodeme L-shaped, phallic guide with six dorso- dorsolateral spines (three short and three longer ones, the longest one twice as length as others) (Fig. 10 C – E). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFCFEC71D8D9EDB7FB3112DC.taxon	etymology	Etymology: The name is from the Latin bi for two and calcar for spur, in reference to the hind trochanter having two spur-like projections. Examined material: HOLOTYPE: AUSTRALIA: New South Wales: Culgoa National Park, 4.5 km West-Northwest Cawwell Homestead, Claypan (CGN 3 M), 29 ° 3 ’ S, 147 ° 1 ’ E, Gidgee, 21. xii. 2009 – 30. i. 2010, B. Schiebaan, Malaise trap, CNC 574785 (1 ♂, QM); PARATYPES: AUSTRALIA: Queensland: Culgoa Floodplain National Park, 10 km NE Toulby Gate (CG 3), 28 ° 56 ’ S, 147 ° 0 ’ E, 140 m, Brigalow, 16. ix – 1. x. 2008, Simpson, Coward, Malaise trap, CNC 576530 (1 ♂, CNC); South Australia: Flinder’s Ranges National Park, Brachina Gorge, Heysen Hilltop, 31 ° 20 ’ S, 138 ° 33 ’ E, 9. x. 1997, J. & A. Skevington & C. Lambkin, JSS 8376, JSS 8384, JSS 8387 – 8, JSS 8391 (5 ♂, QM); New South Wales: Culgoa National Park, 4.5 km WNW Cawwell Homestead, Claypan (CGN 3 M), 29 ° 3 ’ S, 147 ° 1 ’ E, 192 m, Gidgee, 22. XI – 21. XII. 2009, C. Lambkin, N. Starick & B. Schiebaan, Malaise trap, CNC 591856 (1 ♂, CNC); Culgoa National Park, 1 km North-Northwest of Cawwell Homestead, Claypan (CGN 4 M), 29 ° 4 ’ S, 147 ° 3 ’ E, 136 m, Gidgee, 30. I – 18. V. 2010, Malaise trap, B. Schiebaan, CNC 574679 (1 ♂, USNM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFCFEC71D8D9EDB7FB3112DC.taxon	distribution	Distribution: Australia (New South Wales, Queensland, South Australia) (Fig. 110). Notes: This species is found in dry habitats dominated by Acacia and is known to hilltop.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFCDEC4FD8D9ED9BFB77150C.taxon	description	Figs. 11 A – E, 111, 146 D, 158 B	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFCDEC4FD8D9ED9BFB77150C.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter with two tufts of bristles ventrobasally and ventroapically (Fig. 146 D); shape of surstyli in dorsal view, both long, left surstylus longer than right one (Fig. 11 A); subepandrial sclerite with a vertical protrusion in middle; gonopods with four projections in ventral view (Fig. 11 B); phallic guide with some spines dorsolaterally covered by two short membranous sheath, each membranous sheath covered with small teeth-like projections in lateral view (Fig. 11 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFCDEC4FD8D9ED9BFB77150C.taxon	description	Description: MALE: Body length: 3 – 3.4 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose; eyes touching for distance equal to 2 – 2.5 times the length of ocellar triangle. Occiput silvery pollinose, more brownish on upper half. Thorax. Postpronotum pale, with 4 – 5 distinct pale brown bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part silver pollinose; slightly grayish also from the side. Scutellum brownish pollinose and without bristles along the distal edge. Dorsocentral bristles well developed, dark, longer on frontal part and numerous dark bristles around postpronotum. Halter, knob pale with a brownish tinge distally, stem dark brown. Legs. Mid coxa with 2 – 3 bristles. Trochanters and base of femora black, femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees yellow, tibia otherwise dark brown, tarsal segments yellow ventrally, brown dorsally, last segment dorsally darker. Hind trochanter with tuft of dark longer bristles (4 – 6) ventrobasally in one group: “ echidna-like ” and 2 – 3 shorter bristles ventroapically (Fig. 146 D). Ventroapical row of spines on fore femur, black spines on mid femur; no spines on hind femur, only 4 – 6 pale bristles posteriorly equally distributed on distal 1 / 3 (as long as 1 / 3 the width of hind tibia at the distal end); hind tibia bent at middle in posterior view. apical spurs on first four tibiae present (short). Hind metatarsus flattened (1 and 2 especially), first tarsomere slightly longer than 2 – 5 combined; ventral bristles scrub-like, yellow. Pulvilli as long as last tarsal segment. Wing. Length: 3.0 – 3.1 mm. Upper side of basal costal cell with 1 – 3 long distinct bristles. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m proximal to middle of discal cell. 4 – 5 dark setulae on tegula. Abdomen. Viewed obliquely from front tergites subshining black, tergite 1 silvery grey, sides silvery pollinose on tergites 4 and 5 (the latter with larger spots), tergites 2 and 3 with faint gray side spots. Dispersed short dark bristles on tergites present, the longest on tergite 5 and ST 8 and up to 1.5 times as long as width of hind tibia at base. Lateral spines on first tergite present, 5 – 6 brown bristles up to as long as width of hind femur at distal tip. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1 – 1.2 times as long as ST 8. Genitalia without dissection: ST 8 medium sized, brownish, very hairy; membranous area slit-like, posteriorly directed; epandrium brown; yellow surstyli appear long and slim. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.7). Surstyli brown, rather symmetrical with broad base and finger-like apex, left slightly longer than right one; both tips bent inward apically (Fig. 11 A). Genital capsule in ventral view: subepandrial sclerite covered with transverse streaks with a vertical protrusion in middle; gonopods with four projections, two longer projections elongated towards surstyli, reaching the basal third of subepandrial sclerite, two shorter projections bent inwards (Fig. 11 B), phallic guide with 5 – 6 different-sized spines dorsolaterally (longer ones extended towards left surstylus) and surrounded by two membranous sheaths. Genital capsule in lateral view: surstyli straight, in apical third bent towards sternite, gradually narrowed to apex (Fig. 11 D – E); phallus with three ejaculatory ducts, some small teeth-like projections arranged on its membranous sheath (Fig. 11 D – E); ejaculatory apodeme flat, sperm pump elongated, linear (Fig. 11 C). FEMALE: Body length (excluding antennae and head): 3.4 mm. As male except for the following characters. Frons, eyes separated, as wide as 1.2 – 1.5 times the width at antennae; completely silver-grey pollinose until up to 3 / 4 of the frons, shiny black in front of ocellar triangle. Pulvilli and claws about 1 – 1.2 times as long as last tarsal segment. Fore and mid femora with 1 – 2 distinct dark spine-like bristles ventrobasally. Female abdomen with silver spots on sides of tergites 3 – 6. Ovipositor. Base dark brown with pale apical margin, piercer straight, slightly up curved in lateral view LP: LB = 2.1. LDP: LPP = 4.5. (Fig. 158 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFCDEC4FD8D9ED9BFB77150C.taxon	etymology	Etymology: From the Latin bi, two, and floccus, tuft of wool or hair, in reference to the two tufts of bristles on the hind trochanter. Examined material: HOLOTYPE: AUSTRALIA: Queensland: 3 km NE of M [oun] t Webb, 15 ° 3 ’ S, 145 ° 9 ’ E, 2. v. 1981, D. H. Colless, Malaise trap, JSS 8594 (1 ♂, ANIC); PARATYPES: AUSTRALIA: New South Wales: Culgoa National Park, 8 km West-Northwest Cawwell Homestead, Diemunga Lagoon (CGN 2 M), 29 ° 3 ’ S, 146 ° 60 ’ E, Coolibah, 30. i – 18. v. 2010, C. Lambkin, R. Olsen & B. Shieban, Malaise trap, CNC 575155 (1 ♂, CNC); Queensland: 1 km W of Cooktown, 15 ° 28 ’ S, 145 ° 15 ’ E, ~ 13. v. 1981, D. H. Colless, Malaise trap, JSS 8587 – 8 (2 ♂, ANIC); 3 km NE of Mount Webb, 15 ° 3 ’ S, 145 ° 9 ’ E, ~ 1. v. 1981, D. H. Colless, Malaise trap, JSS 8599, JSS 8603 – 4 (3 ♂, ANIC); 2. v. 1981, D. H. Colless, Malaise trap, JSS 8593, JSS 8595 – 7, JSS 8605, JSS 8607 – 8 (7 ♂, ANIC); 30. iv. 1981, D. H. Colless, Malaise trap, JSS 8600 (1 ♂, ANIC); 3.5 km SW by S Mount Baird, 15 ° 10 ’ S, 145 ° 7 ’ E, 3. v. 1981, D. H. Colless, Malaise trap, JSS 8579, JSS 8582 (2 ♂, ANIC); 4. v. 1981, D. H. Colless, Malaise trap, JSS 8584 (1 ♂, ANIC); 5. v. 1981, D. H. Colless, Malaise trap, JSS 8572 (1 ♂, ANIC); Brisbane, Mount Coot-tha, 27 ° 29 ’ S, 152 ° 57 ’ E, 170 m, hilltop, 15. v. 1999, J. & A. Skevington, hand collected, JSS 5772, JSS 5775, JSS 5778 (3 ♂, QM); ~ 170 m, hilltop, 19. iv. 1998, J. Skevington, hand collected, 11: 45 – 12: 15 pm, 11: 15 – 11: 45 am, 12: 15 – 12: 45 pm, JSS 3857, JSS 3861, JSS 3864 – 5, JSS 3874, JSS 3879, JSS 3882 (7 ♂, QM); 4. ix. 1999, J. & A. Skevington, hand collected, 10: 00 am to noon, JSS 7192 (1 ♂, QM); ~ 170 m, hilltop, 7. iii. 1998, J. & A. Skevington, hand collected, JSS 3632, JSS 3634, JSS 3638, JSS 3643 – 4, JSS 3648 – 9 (7 ♂, QM); 170 m, hilltop, 7. xii. 1997, J. & A. Skevington, hand collected, JSS 2343 (1 ♂, QM); 8. xi. 1997, J. & A. Skevington, hand collected, 10: 15 am – 10: 45 am, 9: 15 am – 9: 45 am, JSS 982, JSS 994 (2 ♂, QM); Coorado homestead near Expedition Range National Park, 24 ° 30 ’ N, 149 ° 5 ’ E, 3 – 9. iii. 2001, M. Mathieson, Malaise trap, JSS 13983 – 5 (3 ♂, CNC); Proserpine, Thompson Creek, 20 ° 31 ’ S, 148 ° 34 ’ E, 30 m, 3 – 10. v. 2007, J. Stanisic, CNCD 159596 (1 ♀, CNC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFCDEC4FD8D9ED9BFB77150C.taxon	distribution	Distribution: Australia (New South Wales, Queensland) (Fig. 111). Notes: This species is mostly restricted to eastern Queensland, with one outlier from Culgoa in NSW. Some specimens have been captured hilltopping. This species is genetically most similar to T. angelikae sp. nov. (3.4 – 4.8 % pairwise divergence) and intraspecific genetic distances range from 0.0 % to 3.2 % (Supplementary file 3). The specimens JSS 8593, JSS 8600, JSS 8604, JSS 8607 and JSS 8846 have 2 – 3.2 % pairwise divergence from other specimens in this species. It is likely these specimens belong to a new species, but we could not find any morphological characters to support this hypothesis. For now, they are treated as a single species until more specimens can be collected and additional molecular work can be conducted to improve discrimination of the species.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFF3EC4AD8D9EAABFC9C13BC.taxon	description	Figs 12 A – E, 110, 149 E, 155 F	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFF3EC4AD8D9EAABFC9C13BC.taxon	diagnosis	Diagnosis: This species can be recognized by hind trochanter with a distinct keel and some dark bristles along the whole length (Fig. 149 E); symmetrical, tiny, elongated surstyli in dorsal view (Fig. 12 A); gonopods extended towards surstyli with two lobes on inner margin in ventral view (Fig. 12 B); phallic guide with 3 – 4 dorso- dorsolateral spines; subepandrial sclerite with some small teeth dorsally in lateral view (Fig. 12 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFF3EC4AD8D9EAABFC9C13BC.taxon	description	Description: MALE: Body length: 3.3 – 3.6 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 – 2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 3 – 4 pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum silvery pollinose and without bristles along distal edge. Dorsocentral bristles indistinct, dark, more visible in front part, a few pale bristles around postpronotum present. Halter, knob pale, stem brown. Legs. Mid coxa with four long dark bristles. Trochanters brown, femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 5 of tibiae yellow (tibiae otherwise brown), tarsal segments dark brown dorsally, yellow ventrally, last segment darker. Hind trochanter with distinct keel and some dark bristles along the whole (14 – 16), evenly distributed (Fig. 149 E); ventrobasal spines absent on fore and mid femur. Ventroapical row of spines on fore femur absent; 4 – 5 small, black spines on mid femur; hind femur without ventroapical spines, except 12 – 14 equally spaced bristles posteroventrally, longer on distal half, longest up to ½ times the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present (short). Hind metatarsus slightly flattened, as long as 2 – 4 combined and with strong dark bristles dorsally and dense scrub-like bristles ventrally (tarsomeres 1 – 2 only). Pulvilli shorter than last tarsal segment. Wing. Length: 3.2 – 3.3 mm. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m at 3 / 5 of discal cell (distal to middle). 4 – 5 distinct dark setulae on tegula. Abdomen. Viewed obliquely from front, tergites subshiny brown-black, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (spot on tergite 5 larger). Dispersed strong dark bristles on all tergites present, longest on tergite 5 in length up to 1 / 4 the width of hind tibia at distal end. Lateral bristles on first tergite present, 4 – 6 short dark bristles up to as long as hind femur’s width at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1.2 – 1.4 times as long as ST 8. Genitalia without dissection: ST 8 small to medium sized, round in dorsal view, brown and with a few distinct bristles, otherwise with velvet-like coverage; membranous rounded slit, mostly directed posteriorly; epandrium brown, surstyli yellow, short. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.7). Surstyli short, elongated, symmetrical (Fig. 12 A). Genital capsule in ventral view: both gonopods extended towards sternite, rounded at tips with two distinct lobes on inner margin; phallic guide with 3 – 4 dorso- dorsolateral spines, one spine longer than others, subepandrial sclerite distinct, rectangular-shaped (Fig. 12 B); Genital capsule in lateral view: left surstylus straight in basal two thirds, bent towards sternite apically, right surstylus straight, moderately broadened to apex (not bent) subepandrial sclerite with some small teeth dorsally in basal half (Fig. 12 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 12 C). FEMALE: Body length: 3.1 – 3.3 mm. As male except for the following characters. Frons, eyes separated, as wide as 1.1 – 1.2 times the width at antennae; completely silver-grey pollinose on lower 1 / 3, shiny black in front of ocellar triangle, smooth change in transition zone (at 1 / 4 from ocellar triangle) to shiny black, tiny ridge in the median line on upper 1 / 3 of frons. Enlarged ommatidia silvery shining. Pulvilli and claws about 1.5 – 2.5 times as long as last tarsal segment, smaller on metatarsi. Female abdomen with silver spots on sides of tergites 4 – 6. Ovipositor. Straight in ventral view, strongly curved towards sternite, hook-like in lateral view (yellow piercer, dark brown base), reaching distal end of 5 th segment; base silvery pollinose with dispersed short dark bristles. LP: LB = 2.5. LDP: LPP = 3.3. (Fig. 155 F).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFF3EC4AD8D9EAABFC9C13BC.taxon	etymology	Etymology: From the Latin bi, two and lobus, a rounded projection or protuberance, in reference to the two lobes on the gonopods. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: 8 km E [ast of] Pebble Mouse Cr [eek] on Great Northern H [igh] w [a] y, 70 km E [ast of] J [un] ct [ion], Karijini Dr [ive], 23 ° 06.3.4 ’ S [sic], 118 ° 59 ’ E, 710 m, dry wash, 23. IV – 6. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15891 (1 ♂, WAM); PARATYPES: AUSTRALIA: Northern Territory: Kakadu National Park, Mirrai Lookout, 12 ° 52 ’ S, 132 ° 42 ’ E, hilltop, 27. IX. 1998, J. & A. Skevington, hand collected, JSS 4115 (1 ♂, QM); Mary River National Park, 1.8 km Northwest of Bird Billabong Trail Carpark, west of Mary River bridge on Arnhem Highway, 12 ° 52 ’ S, 131 ° 37 ’ E, hilltop, 25. IX. 1998, J. & A. Skevington, hand collected, JSS 4081 (1 ♂, QM); Darwin, 12 ° 27 ’ S, 130 ° 50 ’ E, 8 – 9. XII. 1963, J. Sedlacek, CNCD 5573 – 4 (2 ♂, BPBM); Queensland: Ridgepole Waterhole, 24 km ESE of Musselbrook Resource Centre, Lawn Hill National Park, 18 ° 40 ’ S, 138 ° 22 ’ E, 180 m, 12. V. 1995, G. Daniels & M. A. Schneider, JSS 8282 (1 ♂, QM); Carnarvon National Park, Mount Rugged Summit, 24 ° 54 ’ S, 147 ° 60 ’ E, 1130 m, hilltop, 30. XI. 1997, J. Skevington & C. Lambkin, hand collected, JSS 2224 (1 ♂, QM); Western Australia: 12 miles S by E of Onslow, 21 ° 41 ’ S, 115 ° 8 ’ E, 18. X. 1970, D. H. Colless, JSS 8669 – 70; JSS 8672 (3 ♂, ANIC); Walyunga National Park, 31 ° 44 ’ S, 116 ° 4 ’ E, 19 – 29. XII. 1999, J. & A. Skevington, C. Lambkin & P. Bouchard, Malaise trap, JSS 7673 (1 ♂, QM); 158 km S Newman, 9 km N Kumarina Road House, 24 ° 38 ’ S, 117 ° 37 ’ E, 638 m, in wide sandy wash, 18 – 21. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 16407 (1 ♂, CNC); 47 km S Pardoo Road House on Shay Gap Road, 20 ° 23 ’ S, 120 ° 1 ’ E, 170 m, dry wash near Spinifex, 1 – 14. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15699 – 700 (1 ♂, CNC; 1 ♂, USNM); 67 km SW Pardoo Road House on Shay Gap Road, 20 ° 28 ’ S, 120 ° 10 ’ E, 177 m, dry sandy ravine with flowers, 1 – 14. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15662, JSS 15664 – 6, JSS 15668 – 71 (5 ♂, 2 ♀, CNC, 1 ♂ USNM); 8 km East of Pebble Mouse Creek on Great Northern Highway, 70 km East of Junction, Karijini Drive, 23 ° 06.3.4 ’ S [sic], 118 ° 59 ’ E, 710 m, dry wash, 23. IV – 6. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15888 – 90 (1 ♂, 2 ♀, CNC); Cape Range National Park, Milyering Ranger House, 22 ° 2 ’ S, 113 ° 56 ’ E, 22 m, below cliff near Ficus, 28. IV – 11. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15653 (1 ♂, CNC); Karijini National Park, Hamersley-Mount Bruce Road, 22 ° 37 ’ S, 118 ° 19 ’ E, 762 m, open mallee with Spinifex, 14 – 19. V. 2003, C. Lambkin, J. Recsei, Malaise trap, JSS 15972 (1 ♂, ANIC); 22 ° 34 ’ S, 118 ° 18 ’ E, 757 m, dry rocky creek bed, Eucalyptus grassland, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16116 (1 ♂, ANIC); Karijini National Park, Juna Downs Road, 22 ° 42 ’ S, 118 ° 25 ’ E; 789 m, between drying pools in rocky Turee Creek, near rock outcrop, Eucalyptus Eremophila scrub, dry Turee Creek, grassy open Eucalyptus scrub, 14 – 19. V. 2003, C. Lambkin & T. Weir, Malaise trap, Malaise trap, JSS 16028, 19 – 25. IV. 2003, JSS 16167 – 8, JSS 16170, JSS 16174, 22 ° 44 ’ S, 118 ° 25 ’ E, 14 – 19. V. 2003, JSS 16108, JSS 16113, 798 m, 25. IV – 14. V. 2003, JSS 16086, JSS 16091 – 2, 22 ° 43 ’ S, 118 ° 25 ’ E, 799 m, 25. IV – 14. V. 2003, JSS 16048 (9 ♂, 2 ♀, ANIC); Karijini National Park, Karijini Drive, 22 ° 34 ’ S, 118 ° 18 ’ E, 815 m, across dry gully, Acacia scrub, 19 – 25. IV. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16156 (1 ♂, ANIC); Karijini National Park, Juna Downs Road (between drying pools in rocky Turee Creek), 22 ° 42 ’ S, 118 ° 25 ’ E, 789 m, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, CNCD 3824, CNCD 3840, CNCD 3850, CNCD 3866 (2 ♂, 2 ♀, CNC); Millstream-Chichester National Park, Snappy Gum Drive, 21 ° 34 ’ S, 117 ° 4 ’ E, 329 m, over deep rocky gully, 2 – 11. V. 2003, C. Lambkin, D. Yeates & J. Recsei, Malaise trap, JSS 16180 (1 ♂, ANIC); Western Australia: Karijini National Park, Karijini Dr. 8 km WNW, Great Northern Hwy, 22 ° 37 ’ S, 118 ° 22 ’ E, 750 m, Dry wash, 22. IV. 2003, F. D. Parker & M. E. Irwin, pan trap, CNCD 6878 (1 ♂, CNC); Cape Range National Park; Milyering Ranger House, 22 ° 2 ’ S, 113 ° 55 ’ E, 22 m, below cliff near Ficus, 28. IV – 11. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15654.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFF3EC4AD8D9EAABFC9C13BC.taxon	distribution	Distribution: Australia (Northern Territory, Queensland, Western Australia). Notes: This species is found in the arid north central and northwestern parts of Australia. Many specimens were collected in dry washes and plant associates include Acacia, Eremophila, Eucalyptus and Spinifex. Some specimens have been captured hilltopping. Intraspecific genetic distance ranges from 0.0 % to 2.4 % (Supplementary file 3). The specimens JSS 15888 and JSS 15891 display 1.4 – 2.4 % pairwise divergence from other specimens suggesting that this is a species complex that requires more investigation.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFF6EC4BD8D9EB7BFB5511D8.taxon	description	Figs 13 A – E, 110	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFF6EC4BD8D9EB7BFB5511D8.taxon	diagnosis	Diagnosis: This species can be recognized by slender and long surstyli in dorsal view (Fig. 13 A); long gonopods with a slender projection towards surstyli, phallic guide with 3 – 6 short dorso- dorsolateral spines; ejaculatory ducts biramous at apex; subepandrial sclerite elongated in ventral view (Fig. 13 B); hind femur without long bristles posteroventrally.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFF6EC4BD8D9EB7BFB5511D8.taxon	description	Description: MALE: Body length: 2.5 mm. Head. Flagellum long tapering; brown pollinose. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput silvery pollinose, with scattered short dark bristles. Thorax. Postpronotum pale, with 3 – 4 short pale bristles. Mesonotum (viewed obliquely from front) silvery pollinose, anterior part more grey. Scutellum silvery pollinose, without bristles along distal edge. Halter, knob pale, tip brownish, stem and base dark. Legs. Mid coxa with 3 – 4 long brown bristles. Trochanters brown, femora dark brown, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments bright yellow ventrally. Hind trochanter with a keel with some short teeth along its edge; ventrobasal spines absent on fore and mid femora. Hind femur without ventroapical spines and without bristles posteroventrally. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres not flattened, hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with shirt light brown erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 2.5 mm. Upper side of basal costal cell with one medium and two short brown bristles. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 1 – 3 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brown-black, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (spot on tergite 5 larger). Lateral bristles on first tergite present, 4 – 5 brown bristles up to as long as ¾ of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.5). Surstyli widened at base, then strongly slender and elongated, curved to each other at apex, left surstylus is longer, right one with a small bumper before its apex (Fig. 13 A). Genital capsule in ventral view: both gonopods equal in height, extended towards surstyli with slender pointed projection at apex towards surstyli; phallic guide with 3 – 6 short dorso- dorsolateral spines, one is longer; phallus trifid; ejaculatory ducts biramous at apex; subepandrial sclerite elongated (Fig. 13 B); Genital capsule in lateral view: both surstyli straight in basal two thirds, bent towards sternite in apical third (Fig. 13 D – E). Ejaculatory apodeme tube-like, bent (Fig. 13 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFF6EC4BD8D9EB7BFB5511D8.taxon	etymology	Etymology: From the Latin bi, two and ramus, branch, in reference to the double branched ejaculatory ducts. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Kurumba, 17 ° 28 ’ S, 140 ° 53 ’], 18. IV. 1983, J. F. Donaldson, D-Vac, JSS 8967 (1 ♂, QDPC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFF6EC4BD8D9EB7BFB5511D8.taxon	distribution	Distribution: Australia (Queensland) (Fig. 110). Notes: This species is known from a single specimen from coastal north central Queensland.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFF4EC46D8D9EFF6FBE1167C.taxon	description	Figs 14 A – D, 112, 158 D	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFF4EC46D8D9EFF6FBE1167C.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter with a tuft of short spines ventrobasally in one group and a few bristles at the distal end; some long bristles on mid femur; surstyli constricted in basal fourth, widened in apical fourth in dorsal view (Fig. 14 A); phallic guide with 5 – 6 dorso-dorsolateral spines, gonopods elongated towards surstyli (Fig. 14 B); both surstyli boomerang-shaped, broadened in middle in lateral view (Fig. 14 C – D).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFF4EC46D8D9EFF6FBE1167C.taxon	description	Description: MALE: Body length: 2.9 – 3.0 mm. Head. Flagellum acuminate; yellow. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose; eyes touching for distance equal to 1 – 1.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 3 – 4 pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum brownish pollinose and with 18 – 20 weak bristles along the distal edge. Dorsocentral bristles distinct, dark, somewhat longer along frontal edge, numerous long pale bristles around postpronotum. Halter, knob pale, stem brown. Legs. Mid coxa with 3 – 4 long dark bristles. Trochanters and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal ¼ of tibiae yellow (tibiae otherwise dark brown), tarsal segments yellow-brown, ventrally somewhat paler, last segment darker. Hind trochanter with a tuft of short, dark, longer spines (7 – 10) ventrobasally in one group: “ echidna-like ”; basal spines longer and a few bristles at the distal end; ventrobasal spines (2) present on fore and mid femora. Ventroapical row of spines on fore femur 5 – 6; 8 – 10 small, 3 – 8 long bristles on mid femur; hind femur without ventroapical spines, except 8 – 10 equally spaced bristles posteroventrally on the distal half, longest at 2 / 3 from the base, in length up to 1.5 the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present (short). Metatarsi moderately flattened, first tarsomere almost as long as 2 – 3 combined. Pulvilli as long as last tarsal segment. Hind metatarsus and second tarsomere widened apically. Wing. Length: 3.1 – 3.2 mm. Upper side of basal costal cell with one long dark bristle. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m distal to the middle of discal cell. 3 – 4 distinct brown setulae on tegula. Abdomen. Viewed obliquely from front tergites subshining black, tergite 1 and anterior half of tergite 2 silvery grey, sides silvery pollinose on tergites 4 and 5 (larger on tergite 5). Dispersed strong dark bristles on all tergites present, longest on tergite 5 up to 1.1 – 1.3 times the width of hind tibia at distal end. Lateral bristles on first tergite present, 5 – 6 pale bristles up to 1.5 times as long as hind femur’s width at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1 – 1.2 times as long as ST 8. Genitalia without dissection: ST 8 medium sized square-shaped in dorsal view, brown and with the occasional distinct abdominal bristles, otherwise velvet-like coverage; membranous area elongated, widening dorsally; epandrium brown, surstyli yellow and bent towards abdomen, appear to be a yellow triangle in lateral view (the corner is square). Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.1). Surstyli symmetrical, broadened in basal fourth and rather straight in apical, its medial margin widened, covered by dense light bristles (Fig. 14 A). Genital capsule in ventral view: gonopods elongated towards surstyli, rather unequal in height, left is slightly longer than right one, subepandrial sclerite long, covered with transverse streaks (Fig. 14 B). Genital capsule in lateral view: both surstyli boomerang-shaped, constricted at base, broadened at middle and bent by 90 ° towards sternite; phallic guide pointed at apex, with 5 – 6 lateral spines (the longest one twice as length as others) (Figs 14 C – D). FEMALE: Body length: 2.7 mm. As male except for the following characters. Frons, eyes separated, as wide as 1.5 times the width at antennae; completely silver-grey pollinose until up to 1 / 3 of the lower frons, shiny black in front of ocellar triangle, gradually changing from pollinose to shiny black in the transition zone; frons depressed at the point where it is widest (a groove is visible in the middle line). Enlarged ommatidia silvery shining. Pulvilli and claws about 1.6 – 1.8 times as long as last tarsal segment, smaller on metatarsi. Female abdomen with silver spots on sides of all tergite. Ovipositor. Straight, reaching middle of 3 rd segment; base silvery pollinose with dispersed bristles. LP: LB = 2.07. LDP: LPP = 3.3. (Fig. 158 D).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFF4EC46D8D9EFF6FBE1167C.taxon	etymology	Etymology: The name refers to the boomerang-shaped surstyli. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: Heinsmen Rock, Mt Ragged Track, 30 ° 7 ’ S, 123 ° 28 ’ E, 145 m, woodland near drying pools, 30. x – 18. xi. 2003, C. Lambkin & J. Recsei, Malaise trap, JSS 16242 (1 ♂, WAM); PARATYPES: AUSTRALIA: Australian Capital Territory: Canberra, Black Mountain, 35 ° 16 ’ S, 149 ° 6 ’ E, 11 – 19. i. 1999, G. Gibson, sweeping, CNCD 4464, CNCD 4470 (2 ♂, CNC); New South Wales: Mount Boppy near Cobar, 31 ° 33 ’ S, 146 ° 17 ’ E, hilltop, 16. i. 1999, C. Lambkin, N. Power & P. Bouchard, 10: 00 AM, JSS 8902 (1 ♂, QM); Yathong Nature Reserve near 8 Mile Tank, 32 ° 38 ’ S, 145 ° 41 ’ E, ~ 205 m, field of flowers: open woodland, 3 – 25. x. 2003, C. Lambkin & N. Starick, Malaise trap, CNCD 3623, CNCD 3627, CNCD 3631, JSS 50566 (3 ♂, QM, 1 ♂, ANIC); Tinderry Natl Res; Round Flat Fire Trail 11.2 Km E Michelago, 35 ° 43 ’ S, 149 ° 17 ’ E, 1186 m, Revegetation Site, 26. II – 19. III. 2005, C. Lambkin & N. Starick, Malaise trap, Bulk Sample 2711, JSS 50565 (1 ♂, ANIC); Queensland: Culgoa Floodplain National Park, 11.8 km North of Headquarters (CG 2 AM), 28 ° 54 ’ S, 147 ° 8 ’ E, 151 m, Brigalow, Black box, 17. xii. 2009 – 20. i. 2010, C. Kelly & A. Coward, Malaise trap, CNC 574518 (1 ♂, CNC); 36 m, 15 – 22. viii. 2007, J. Stanisic, Malaise trap, 15036, CNCD 159453 (1 ♀, CNC); Culgoa Floodplain National Park Headquarters, 11.8 km N, 28 ° 55 ’ S, 147 ° 8 ’ E, 150 m, Brigalow, 9. XII. 2007 – 16. I. 2008, N. Starick, C. Lambkin & R. Raven, Malaise trap, CNC 591838 (1 ♂, QM); Culgoa Floodplain National Park, 7 km NNW Toulby Gate (CG 4 AM), 28 ° 56 ’ S, 146 ° 55 ’ E, 160 m, Gidgee, 17. XII. 2009 – 20. I. 2010, Kelly & Coward, Malaise trap, CNC 591825 (1 ♂, QM); Culgoa Floodplain National Park, 7 km WNW Toulby Gate (TOLB 6 M), 28 ° 57 ’ S, 146 ° 52 ’ E, 138 m, Callitris, 19. III – 20. V. 2010, C. Kelly, A. Coward, Malaise trap, CNC 575020, CNC 575022 – 3 (3 ♂, QM); Proserpine, Airport Drive, 20 ° 29 ’ S, 148 ° 34 ’ E, 514 m, 10 – 16. ii. 2007, C. Lambkin & N. Starick, 15002, CNCD 159303 (1 ♂, CNC); Fanning River Homestead, 3.2 km Southeast, 19 ° 45 ’ S, 146 ° 27 ’ E, 330 m, vinescrub, 9. XII. 2006 – 10. II. 2007, S. Wright, Malaise trap, CNC 575198 (1 ♂, QM); South Australia: Flinder’s Ranges National Park, Love Range, Mine Road, 31 ° 24 ’ S, 138 ° 47 ’ E, in dry creek bed, 8 – 9. x. 1997, S. Winterton, J. & A. Skevington & C. Lambkin, Malaise trap, JSS 8407 (1 ♂, QM); Gammon Ranges National Park, south end of Weetootla Gorge, 30 ° 29 ’ S, 139 ° 16 ’ E, 537 m, hilltop, 11. x. 1997, J. Skevington & C. Lambkin, JSS 8346, JSS 8349 - 50, JSS 8358, JSS 8360, JSS 8362 (6 ♂, QM); Mount Lofty, 35 ° 0 ’ S, 138 ° 44 ’ E, 2. I. 1974, 10 th Australian Boy Scout Jamboree, Malaise trap, JSS 8719 (1 ♂, SAM); Wyperfeld NP, East Lookout, 35 ° 36 ’ S, 142 ° 7 ’ E, 22. X. 2014, J. H., A. W., & A. M. Skevington, A. D. Young, A. Wyatt, & S. Namek, CNC 373402, CNC 373424 (2 ♂, CNC); Victoria: Hattah-Kulkyne NP, Warepil Lookout, 34 ° 43 ’ S, 142 ° 18 ’ E, 18. x. 2014, J. H., A. W., & A. M. Skevington, A. D. Young, A. Wyatt, & S. Namek, CNC 373150 – 1, CNC 373172 (3 ♂, CNC); 12 km SE Merrijig, Howqua River, 37 ° 12 ’ S, 146 ° 22 ’ E, 30. XI. 1971, Neboiss, JSS 8809 (1 ♂, MVMA); Northern Territory: Entire Creek, 155 km NE by E of Alice Springs, 22 ° 50 ’ S, 135 ° 16 ’ E, 7. X. 1978, D. H. Colless, JSS 9253 (1 ♂, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFF4EC46D8D9EFF6FBE1167C.taxon	distribution	Distribution: Australia (Australian Capital Territory, New South Wales, Northern Territory, Queensland, South Australia, Western Australia, Victoria) (Fig. 112). Notes: This is a widespread species throughout the outback of Australia. The two specimens from NE Queensland (CNCD 159303 and CNC 575198) are the exception and found in mesic habitats. The DNA sequence of CNCD 159303 specimen is identical with all other specimens and genitalia features of both specimens are similar to others. The specimen CNC 575198 doesn’t have all legs and the color is lighter due to preservation conditions. Despite the apparent ecological differences, support is strong that these outliers are the same species. A few specimens have been collected hilltopping. Intraspecific genetic distance ranges from 0.0 % to 1.4 %. This species is genetically most similar to T. dissidens sp. nov. (1.3 – 1.5 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFFAEC44D8D9E93BFEA1150C.taxon	description	Figs 15 A – E, 112, 144 F	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFFAEC44D8D9E93BFEA1150C.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a patch of short bristles ventrobasally (Fig. 144 F); elongated surstyli with two lobes at outer margin before apex and a small, pointed edge in ventrobasal margin in dorsal view (Fig. 15 A); both gonopods extended towards surstyli, rounded at apex; phallic guide with 3 – 4 short dorso- dorsolateral spines in ventral view (Fig. 15 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFFAEC44D8D9E93BFEA1150C.taxon	description	Description: MALE: Body length: 2.8 – 2.9 mm. Head. Scape and pedicel dark, arista dark brown, flagellum light brown. Flagellum tapering. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2.2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 3 – 4 dark bristles. Mesonotum (viewed obliquely from front) brown pollinose, anterior part of scutum more grey. Scutellum silvery brown pollinose with scattered dark bristles. Dorsocentral and supra-alar bristles short, dark, longer in frontal half, numerous pale bristles around postpronotum present. Halter, knob pale, tip brownish, stem light brown with 2 short bristles in upper side, base dark. Legs. Mid coxa with 3 – 4 long dark bristles. Fore and mid trochanters with 2 – 3 short bristles ventroapically. Trochanters brown, femora black, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments brown dorsally, light brown ventrally. Hind trochanter with a batch of short bristles ventrobasally, brown pollinosity (Fig. 144 F); one long ventrobasal bristle present on fore and 1 – 3 shorter bristles on mid femur. Hind femur without ventroapical spines. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres not flattened, hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 3.0 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 3 – 5 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brown-black, tergite 1 silvery grey. Lateral bristles on first tergite present, 3 – 5 long dark bristles up to as long as ¾ of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium as long as wide (MLE: MWE = 1.20). Surstyli elongated, with two lobes at outer margin before apex, curved to each other at apex, left surstylus is longer, both surstyli with a small pointed edge in ventrobasal margin (Fig. 15 A). Genital capsule in ventral view: both gonopods extended towards surstyli, rounded at apex; phallic guide with 3 – 4 short dorso- dorsolateral spines; subepandrial sclerite large with some transverse sutures (Fig. 15 B); Genital capsule in lateral view: both surstyli straight in basal half, strongly bent towards sternite in basal half; both surstyli with a pointed edge dorsobasally (Fig. 15 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 15 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFFAEC44D8D9E93BFEA1150C.taxon	etymology	Etymology: From the Greek bounites, dweller in the hills, in reference to the fact that all known specimens have been collected on hilltops. Many pipunculids go to hilltops to mate (Skevington, 2008). Examined material: HOLOTYPE: AUSTRALIA: Queensland: Summit of Mount Greville, 28 ° 4 ’ S, 152 ° 30 ’ E, 393 m, 3. I. 2015, J. H., A. M. & A. W. Skevington, CNC 374390 (1 ♂, QM); PARATYPES: AUSTRALIA: Queensland: Carnarvon National Park, Hilltop north of Tombs, 25 ° 5 ’ S, 147 ° 52 ’ E, hilltop, 26. XI. 1997, J. Skevington & C. Lambkin, hand collected, JSS 2217 (1 ♂, QM); Summit of Mount Greville, 28 ° 4 ’ S, 152 ° 30 ’ E, 393 m, 3. I. 2015, J. H., A. M. & A. W. Skevington, CNC 374415 (1 ♂, CNC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFFAEC44D8D9E93BFEA1150C.taxon	distribution	Distribution: Australia (Queensland) (Fig. 112). Notes: The only known specimens have been collected hilltopping in dry sclerophyll forest in SE to SC Queensland.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFF8EC45D8D9EAABFB9017A8.taxon	description	Figs 16 A – E, 113, 155 A	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFF8EC45D8D9EAABFB9017A8.taxon	diagnosis	Diagnosis: This species can be recognized by shortened surstyli, the left surstylus constricted in middle at outer margin in dorsal view (Fig. 16 A); extended hypandrium, small gonopods, shortened subepandrial sclerite in ventral view (Fig. 16 B); phallic guide widened; phallus with three strong ejaculatory ducts, one covered by small spines in basal half in lateral view (Fig. 16 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFF8EC45D8D9EAABFB9017A8.taxon	description	Description: MALE: Body length: 3.1 – 3.2 mm. Head. Flagellum short acuminate; brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 – 2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 2 – 3 short pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum brownish pollinose and with 8 – 10 weak bristles along the distal edge. Dorsocentral bristles weakly developed, indistinct, dark, somewhat more visible along frontal edge, a few pale bristles around postpronotum present. Halter, knob pale with brownish tinge, stem brown. Legs. Mid coxa with three short dark bristles in a row along the edge. Trochanters brown and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 4 of tibiae yellow (tibiae otherwise brown), tarsal segments yellow ventrally brown dorsally, last segment darker. Hind trochanter with a half circle, no distinct bristles; ventrobasal spines absent on fore and mid femora. Ventroapical row of spines on fore femur absent; 4 – 6 small, black spines on mid femur; hind femur without ventroapical spines, except 8 – 10 equally spaced bristles posteroventrally, on distal half more like strong bristles, longest up to 1 / 2 the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae absent. Hind tibia slightly bent in posterior view. Metatarsi moderately flattened, hind metatarsus almost as long as 2 – 5 combined. Pulvilli shorter than last tarsal segment. Wing. Length: 3.6 – 3.7 mm. Upper side of basal costal cell with two short dark bristles. Fourth costal section 2 times as long as third costal section. Cross-vein r-m slightly distal to middle of discal cell. 1 – 2 distinct brown setulae on tegula. Abdomen. Viewed obliquely from front tergites subshiny black, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (larger on tergite 5). Dispersed short dark bristles on all tergites present, very small, indistinct. Lateral bristles on first tergite absent. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1.5 – 2.2 times as long as ST 8. Genitalia without dissection: ST 8 medium sized, broad, rounded triangle in dorsal view, dark brown and with some distinct bristles, otherwise velvet-like coverage; membranous area drop-like, directed posteriorly; epandrium dark brown, surstyli brown, inconspicuous. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.7). Surstyli rather asymmetrical, shorter than epandrium, both surstyli broader basally and distally, left one constricted in middle at outer margin (Fig. 16 A). Genital capsule in ventral view: gonopods small, semicircular in lateral margin; subepandrial sclerite small; hypandrium more extended; phallus with three strong ejaculatory ducts, one covered with small spines in basal half (Fig. 16 B); Genital capsule in lateral view: both surstyli curved towards sternite; phallic guide widened and short, pointed at apex (Fig. 16 D – E). Ejaculatory apodeme rather long, tube-like, with a bulb in its middle (Fig. 16 C). FEMALE: Body length: 3.1 – 3.2 mm. As male except for the following characters. Frons, eyes separated, as wide as 1.2 – 1.3 times the width at antennae; completely silver-grey pollinose until up to 1 / 2 of the lower frons, shiny black in front of ocellar triangle, sudden change in transition zone. Enlarged ommatidia silvery shining. Pulvilli and claws about 1 – 2 times as long as last tarsal segment, smaller on metatarsi. Female abdomen with silver spots on sides of tergites 4 – 6 (sometimes T 3). Ovipositor. Straight in ventral view, curved towards sternite and short in lateral view, reaching distal edge of 4 th segment; base brownish pollinose with occasional bristles; very flat, in level with rest of abdomen. LP: LB = 2.6. LDP: LPP = 1.7. (Fig. 155 A).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFF8EC45D8D9EAABFB9017A8.taxon	etymology	Etymology: Brachy is Greek for short and refers to the short surstyli of the males. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Carnarvon National Park, Mount Moffatt Section, Top Shelter Shed, 24 ° 55 ’ S, 148 ° 3 ’ E, 1100 m, 22. XI. 1995, D. K. Yeates, JSS 8311 (1 ♂, QM); PARATYPES: AUSTRALIA: Queensland: Bogantungan, 13.5 km North, 23 ° 32 ’ S, 147 ° 18 ’ E, 880 m, open forest, 18. XII. 2000 – 27. III. 2001, Cook & Monteith, flight interception trap, 10007, CNC 575165 (1 ♂, CNC); Cudmore National Park (CM 3 M), 22 ° 58 ’ S, 146 ° 23 ’ E, 351 m, Melaleuca heath near drying creek, 28. X. 2010 – 2. VIII. 2011, Lambkin, Starick & Bailey, Malaise trap, CNC 575000, CNC 575003 (2 ♂, QM); Cudmore National Park (CM 3 M), 22 ° 58 ’ S, 146 ° 23 ’ E, 351 m, Melaleuca heath near drying creek, 28. X. 2010 – 2. VIII. 2011, Lambkin, Starick & Bailey, Malaise trap, CNC 574704 (1 ♂, CNC); 3.5 km SW by S Mount Baird, 15 ° 10 ’ S, 145 ° 7 ’ E, 4. V. 1981, D. H. Colless, Malaise trap, JSS 8578 (1 ♂, ANIC); Brisbane Forest Park, 27 ° 25 ’ S, 152 ° 51 ’ E, sclerophyll Eucalyptus forest, 8. XI. 1995, Irwin & Gaimari, JSS 8835 (1 ♀, INHS); Carnarvon National Park, Mount Moffatt Section Headquarters, 25 ° 1 ’ S, 147 ° 57 ’ E, 740 m, 21. XI. 1995, M. E. Irwin & S. D. Gaimari, JSS 8899 (1 ♂, INHS); Carnarvon National Park, Mount Moffatt Section, Top Shelter Shed, 24 ° 55 ’ S, 148 ° 3 ’ E, 1100 m, 22. XI. 1995, D. K. Yeates, JSS 8308, JSS 8313, JSS 8317 (3 ♀, QM); Carnarvon National Park, Mount Moffatt section, 25 ° 3 ’ S, 147 ° 53 ’ E, [dry sclerophyll, savanna opening], 27. XI. 1997, J. Skevington & C. Lambkin, Malaise trap, JSS 2278 (1 ♂, QM); Scrub Road, Brisbane Forest Park, 27 ° 25 ’ S, 152 ° 50 ’ E, 3 – 10. X. 1997, S. Winterton, N. Power & D. White, Malaise trap, JSS 8857 (1 ♂, QM); Northern Territory: Darwin, Fannie Bay, 12 ° 30 ’ S, 130 ° 50 ’ E, Park near beach, 4. IX. 2004, J. Forrest & T. Wheeler, sweeping, LEM _ 0016202 (1 ♂, LEM); New South Wales: 20 mi. Glen Innes to Grafton Highway, 29 ° 43 ’ S, 151 ° 45 ’ E, 20. IV. 1970, D. H. Colless, Coordinates for Glen Innes, JSS 8504 (1 ♂, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFF8EC45D8D9EAABFB9017A8.taxon	distribution	Distribution: Australia (New South Wales, Northern Territory, Queensland) (Fig. 113). Notes: This species has been found in dry sclerophyll forest and heathlands in eastern NSW, Queensland and coastal Northern Territory. Intraspecific genetic distance ranges from 0.0 % to 0.3 %. This species is genetically similar to T. biloba sp. nov. (2.6 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFFFEC41D8D9EFF6FF3712F0.taxon	description	Figs 17 A – E, 113, 153 B	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFFFEC41D8D9EFF6FF3712F0.taxon	diagnosis	Diagnosis: This species can be recognized by the keelless hind trochanter, with only a bump ventrally with some spine-like bristles; elongated surstyli in dorsal view (Fig. 17 A); extended gonopods; phallic guide with 4 – 6 dorso-dorsolateral spines; subepandrial sclerite distinct with a small fin-shaped protrusion in its center (Fig. 17 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFFFEC41D8D9EFF6FF3712F0.taxon	description	Description: MALE: Body length: 3.0 – 3.2 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 – 2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 3 – 4 pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum silvery brown pollinose and with 5 – 7 bristles along distal edge. Dorsocentral bristles indistinct, dark, more visible in frontal half, a few pale bristles around postpronotum present. Halter, knob pale, stem brown. Legs. Mid coxa with 3 – 4 long dark bristles, one is longer. Trochanters brown, femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments dark brown dorsally, yellow ventrally, last segment darker. Hind trochanter without a keel, a bump ventrally with a tuft of dark spine-like bristles, 8 – 10; ventrobasal spines (2) present on fore and absent on mid femur. Ventroapical row of spines on fore femur 4 – 6; 8 – 10 distinct, black spines on mid femur; hind femur without ventroapical spines, except 6 – 8 equally spaced bristles posteroventrally distal half, longest up to 1 / 3 the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres flattened (especially hind metatarsus in the middle, as long as 2 – 4 combined), t 1 with shiny patch ventrally (no bristles). Pulvilli shorter than last tarsal segment. Wing. Length: 3.2 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m at middle of discal cell. 3 – 4 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites subshiny brown-black, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (spot on t 5 larger). Dispersed strong dark bristles on all tergites present, longest on tergite 5 in length up to 1 / 4 the width of hind tibia at distal end. Lateral bristles on first tergite present, 4 – 6 dark bristles up to as long as ¾ of hind femur’s width at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 0.9 – 1.1 times as long as ST 8. Genitalia without dissection: ST 8 small to medium sized, round in dorsal view, brown and with a few distinct bristles, otherwise with velvet-like coverage; membranous rounded slit, mostly directed posteriorly; epandrium brown, surstyli yellow, elongated, medium sized. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.2). Surstyli elongated, symmetrical, left surstylus longer than right one (Fig. 17 A). Genital capsule in ventral view: both gonopods extended towards sternite, rounded at tips; phallic guide with 4 – 6 dorso- dorsolateral spines, one spine longer than others; subepandrial sclerite distinct with a small fin-shaped protrusion in its center (Fig. 17 B); Genital capsule in lateral view: left surstylus straight in basal two thirds, bent towards sternite apically, pointed at apex, right surstylus straight, slightly curved towards sternite (Fig. 17 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 17 C). FEMALE: Body length: 2.8 – 3.0 mm. As male except for the following characters. Frons, eyes separated, as wide as 1.1 – 1.3 times the width at antennae; completely silver-grey pollinose on lower 1 / 3, shiny black in front of ocellar triangle, smooth change in transition zone (at 1 / 3 from ocellar triangle) to shiny black, small groove in the median line on upper 1 / 3 of frons. Enlarged ommatidia silvery shining. Pulvilli and claws about 1.5 – 2.5 times as long as last tarsal segment, smaller on metatarsi. Female abdomen with silver spots on sides of tergites 4 – 6. Ovipositor. Straight in ventral view, straight, dagger-like in lateral view (whitish yellow piercer, dark brown base), reaching distal end of 2 nd segment; base silvery brown pollinose with dispersed short dark bristles. LP: LB = 2.0. LDP: LPP = 3.3. (Fig. 153 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFFFEC41D8D9EFF6FF3712F0.taxon	etymology	Etymology: The name is derived from the Greek branchos for fin, in reference to the fin-shaped protrusion on the subepandrial sclerite. Examined material: HOLOTYPE: AUSTRALIA: Northern Territory: Kakadu National Park, Mirrai Lookout, 12 ° 52 ’ S, 132 ° 42 ’ E ,, hilltop, 27. IX. 1998, J. & A. Skevington, hand collected, JSS 4100 (1 ♂, QM); PARATYPES: AUSTRALIA: Queensland: Black Mountain, 6.7 km west of Croyden, 18 ° 9 ’ S, 142 ° 12 ’ E, 195 m, hilltop, dry, open savanna, 12. VI. 1997, J. & A. Skevington, JSS 8318 (1 ♂, QM); Western Australia: Karijini National Park, Banjima Drive, 10 km W visitors center, 22 ° 29 ’ S, 118 ° 22 ’ E, 743 m, 4 – 16. V. 2003, M. E. Irwin & F. D. Parker, JSS 15822 (1 ♂, CNC); Karijini National Park, Hamersley-Mount Bruce Road, 22 ° 34 ’ S, 118 ° 18 ’ E, 757 m, dry rocky creek bed, Eucalyptus grassland, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16118 (1 ♂, ANIC); Karijini National Park, Juna Downs Road, 22 ° 42 ’ S, 118 ° 25 ’ E, 789 m, between drying pools in rocky Turee Creek, near rock outcrop, Eucalyptus Eremophila scrub, dry Turee Creek, grassy open Eucalyptus scrub, 19 – 25. IV. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16175 (1 ♀, ANIC); 22 ° 44 ’ S, 118 ° 25 ’ E, 798 m, 25. IV – 14. V. 2003, JSS 16087 (1 ♂, ANIC); 22 ° 43 ’ S, 118 ° 25 ’ E, 799 m, 25. IV – 14. V. 2003, JSS 16060 (1 ♀, ANIC); Karijini National Park, Karijini Drive, 22 ° 34 ’ S, 118 ° 18 ’ E, 814 m, open Eucalyptus grassland, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 15582 (1 ♂, ANIC); Karijini National Park, Weano Gorge Road, 22 ° 22 ’ S, 118 ° 15 ’ E, 775 m, hilltop, open Eucalyptus, 20 – 25. IV. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16227 (1 ♂, ANIC); Karijini National Park, above Fortescue Falls, 22 ° 29 ’ S, 118 ° 33 ’ E, over dry creek in low Acacia scrub, 21 – 24. IV. 2003, C. Lambkin, Malaise trap, JSS 16135 (1 ♀, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFFFEC41D8D9EFF6FF3712F0.taxon	distribution	Distribution: Australia (Northern Territory, Queensland, Western Australia) (Fig. 113). Notes: This species is found in open Eucalyptus forest and Acacia scrub across northern Australia. Some specimens have been captured hilltopping. Intraspecific genetic distance ranges from 0.0 % to 0.7 %. This species is genetically closest to T. biloba sp. nov. and T. uncinata sp. nov. (5.0 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFFDEC41D8D9EDB7FCE6178F.taxon	description	Figs 18 A – E, 114	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFFDEC41D8D9EDB7FCE6178F.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a long keel covered by strong teeth along its edge; elongated surstyli, curved to each other before apex in dorsal view (Fig. 18 A); both gonopods extended towards surstyli; phallic guide with 3 – 4 short dorso- dorsolateral spines in ventral view (Fig. 18 B); left surstylus straight in basal two thirds, bent towards sternite in apical fourth in lateral view (Fig. 18 D).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFFDEC41D8D9EDB7FCE6178F.taxon	description	Description: MALE: Body length: 2.8 mm. Head. Scape dark, pedicel brown, flagellum light brown, long tapering, brown pollinose. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching as long as the length of ocellar triangle. Occiput black, somewhat silvery pollinose. Thorax. Postpronotum pale. Mesonotum (viewed obliquely from front) brown pollinose, supra-alar area greyer. Dorsocentral bristles short, dark, longer in frontal half, numerous pale bristles around postpronotum present. Scutellum silvery brown pollinose with 2 – 4 short scattered bristles. Legs. Mid coxa with 1 – 2 long dark bristles. Trochanters brown, femora black, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae light brown (tibiae otherwise brown), tarsal segments light brown. Hind trochanter with long keel covered by strong teeth along its edge; two ventrobasal spines present on fore and mid femora. Hind femur without ventroapical spines. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres distinctly flattened (t 1 – 3), hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 3.0 mm. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2 – 3 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites subshiny brown-black, tergite 1 silvery grey. Dispersed short dark bristles on all tergites present. Lateral bristles on first tergite present, 4 – 6 brown bristles up to as long as ¾ of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.1). Surstyli elongated, straight in basal two thirds, slightly broadened and curved to each other before apex, left surstylus is longer and its apex is wider than right one (Fig. 18 A). Genital capsule in ventral view: both gonopods extended towards surstyli, pointed at apex, both gonopods with a distinct projection on inner margin, right one is wider and left one is smaller; phallic guide with 3 – 4 short dorso- dorsolateral spines; subepandrial sclerite distinct with a fin-shaped projection in its center (Fig. 18 B); Genital capsule in lateral view: left surstylus straight in basal two thirds, bent towards sternite in apical fourth, right surstylus gently curved towards sternite (Fig. 18 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 18 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFFDEC41D8D9EDB7FCE6178F.taxon	etymology	Etymology: From the Latin brochus, with projecting teeth, in reference to the strong teeth on the hind trochanter. Examined material: HOLOTYPE: AUSTRALIA: Tasmania: Launceston, [41 ° 26 ’ S, 147 ° 8 ’ E], JSS 8726 (1 ♂, SAM); PARATYPE: AUSTRALIA: Queensland: Barakula State Forest # 302, 26 ° 22 ’ S, 150 ° 58 ’ E, 8. I. 2000, J. & A. Skevington, M. Mathieson, JSM 71, CNC 483840 (1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFFDEC41D8D9EDB7FCE6178F.taxon	distribution	Distribution: Australia (Queensland, Tasmania) (Fig. 114).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE3EC5FD8D9EFF6FDEB16C8.taxon	description	Figs 19 A – E, 115	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE3EC5FD8D9EFF6FDEB16C8.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli, both are triangle-shaped and the right surstylus is wider than left one in the middle in dorsal view (Fig. 19 A); gonopods asymmetrical, right is longer than left one, both with a short semicircular projection at the apex; each of ejaculatory ducts with 2 – 3 small spines in ventral view (Fig. 19 B); both surstyli straight and widened, pointed at apex in lateral view (Fig. 19 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE3EC5FD8D9EFF6FDEB16C8.taxon	description	Description: MALE: Body length: 2.5 mm. Head. Scape, pedicel and arista dark brown, Flagellum tapering, yellow-brown, light brown pollinose. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.2 times the length of ocellar triangle. Occiput silvery pollinose. Thorax. Postpronotum pale, with 3 – 4 long pale bristles. Prescutum with two triangle-shaped grey marks on anterior part. Mesonotum (viewed obliquely from front) silvery-brown pollinose. Dorsocentral bristles short, dark, longer in frontal half, numerous pale bristles around postpronotum present. Scutellum silvery brown pollinose and with 6 – 8 scattered bristles. Halter, base dark, stem pale, knob pale, tip brownish. Legs. Mid coxa with 3 – 4 dark bristles. Trochanters brown, femora black, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments bright yellow completely, except last tarsomeres. Hind trochanter triangle-shaped grey pollinose without specific feature; ventrobasal spines absent on fore and mid femora. Hind femur without ventroapical spines, except 6 – 10 equally spaced short bristles posteroventrally and anteroventrally, longest up to 1 / 2 the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres not flattened, hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect pale bristles except last tarsomeres with longer dark bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 2.4 mm. Upper side of basal costal cell with one long light brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2 – 3 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brown-black, all tergite silvery-brown pollinose, Dispersed short brown bristles on all tergites present. Lateral bristles on first tergite present, 3 – 6 dark bristles up to as long as half of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: membranous area round and wide, epandrium longer than wide (MLE: MWE = 1.3). Surstyli triangle-shaped, wide at the base, pointed to each other at apex, right surstylus is wider than left one in the middle, left one is longer (Fig. 19 A). Genital capsule in ventral view: gonopods asymmetrical, right is longer than left one, both with a short semicircular projection at the apex, small hypandrium, long subepandrial sclerite; phallic guide long; phallus trifid, each one with 2 – 3 small spines (Fig. 19 B); Genital capsule in lateral view: both surstyli straight and widened, pointed at apex (Fig. 19 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 19 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE3EC5FD8D9EFF6FDEB16C8.taxon	etymology	Etymology: This species is named in honour of Scott E. Brooks. Scott collected one of the two known specimens and is a well-known dipterist and colleague of the authors. Scott works at the CNC in Ottawa, Ontario, Canada. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Carnarvon Nat [ional] Park, Consuelo land, Race Course Spring, 24 ° 56 ’ S, 148 ° 5 ’ E, Vegetation, 12. X. 2002, S. Boucher, sweep net, LEM _ 0016155 (1 ♂, QM); PARATYPE: AUSTRALIA: Queensland: Carnarvon National Park, The Tombs, 25 ° 5 ’ S, 147 ° 52 ’ E, 12. X. 2002, S. E. Brooks, sweep net, LEM _ 0016200 (1 ♂, LEM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE3EC5FD8D9EFF6FDEB16C8.taxon	distribution	Distribution: Australia (Queensland) (Fig. 115). Notes: This species is only known from two specimens collected in Carnarvon National Park in Queensland. The species presumably does not hilltop as there has been a lot of effort collecting on hilltops in Carnarvon and it has not appeared in any of those samples.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE1EC5DD8D9EFF6FD5417CC.taxon	description	Figs 20 A – D, 114, 148 F	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE1EC5DD8D9EFF6FD5417CC.taxon	diagnosis	Diagnosis: This species can be recognized by the bulging and arched hind trochanter with short spines along a keel ventrally (Fig. 148 F); elongated surstyli in dorsal view (Fig. 20 A); elongated gonopods towards surstyli in ventral view (Fig. 20 B); long subepandrial sclerite; long and distinct hypandrial apodeme in ventral view (Fig. 20 B); phallic guide with 5 – 6 dorso-dorsolateral spines (Fig. 20 B – D).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE1EC5DD8D9EFF6FD5417CC.taxon	description	Description: MALE: Body length: 2.6 – 2.7 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 2 – 3 pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum silvery pollinose and with 6 – 8 bristles along the distal edge. Dorsocentral bristles distinct, pale, somewhat longer along frontal edge, numerous dark bristles around postpronotum present. Halter, knob pale, stem brown. Legs. Mid coxa with four same size dark bristles. Trochanters and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 5 of tibiae yellow (tibiae otherwise dark brown), tarsal segments yellow-brown, ventrally paler, last segment darker. Hind trochanter bulging arched ventrally with short spines along a keel ventrally (20 – 22) (Fig. 148 F); ventrobasal spines absent on fore and mid femora. Ventroapical row of spines on fore femur absent; 3 – 4 small, black spines on mid femur; hind femur without ventroapical spines, except 18 – 20 equally spaced bristles posteroventrally, longer and stronger on distal half, longest up to 1.5 times the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present (short). Hind tibia bent in posterior view. Hind metatarsus not flattened and slightly longer than 2 – 4 combined. Pulvilli shorter than last tarsal segment. Wing. Length: 3.0 mm. Upper side of basal costal cell with one short dark bristle. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m at 3 / 5 of discal cell (distal to middle). 2 – 3 distinct brown setulae on tegula. Abdomen. Viewed obliquely from front tergites slivery pollinose, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (similar size). Dispersed strong dark bristles on all tergites present, longest on tergite 5 up to ½ the width of hind tibia at distal end. Lateral bristles on first tergite present, 3 – 4 dark bristles up to half as long as hind femur’s width at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1.3 – 1.6 times as long as ST 8. Genitalia without dissection: ST 8 medium sized, broad, round in dorsal view, brown and with distinct bristles, otherwise velvet-like coverage; membranous area rounded elongate, mostly directed posterodorsally; epandrium and surstyli not visible (view blocked). Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.7). Surstyli elongated, right surstylus smaller than left one, pointed at inner margin before apex; left surstylus curved towards right one (Fig. 20 A). Genital capsule in ventral view: gonopods elongated towards surstyli, tapering at apex with a small edge at inner margin before apex, equal in height; subepandrial sclerite long; hypandrial apodeme distinct, more extended beneath hypandrium (Fig. 20 B). Genital capsule in lateral view: both surstyli narrow, curved towards sternite; phallic guide with 5 – 6 dorso-dorsolateral spines (one is longer) (Fig. 20 C – D). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE1EC5DD8D9EFF6FD5417CC.taxon	etymology	Etymology: The name is based on the Latin bulbus for swelling, in reference to the bulging hind trochanter. Examined material: HOLOTYPE: AUSTRALIA: Queensland: 3 km west Windorah, [- 25.406587, 142.635480], 31. VIII – 5. IX. 1997, S. Winterton, J. & A. Skevington & C. Lambkin, Malaise trap, JSS 8852 (1 ♂, QM); PARATYPES: AUSTRALIA: South Australia: Flinder’s Ranges National Park, Love Range, Mine Road, 31 ° 24 ’ S, 138 ° 47 ’ E, in dry creek bed, 8 – 9. X. 1997, S. Winterton, J. & A. Skevington & C. Lambkin, Malaise trap, JSS 8403 (1 ♂, QM); Flinders Ranges National Park, 31 ° 24 ’ S, 138 ° 47 ’ E, 10 – 11. X. 1997, J. & A. Skevington, S. Winterton & C. Lambkin, Malaise trap, JSS 8409 (1 ♂, QM); Queensland: 20 km E Mareeba, - 17.092489, 145.320180, 15. IV. 1980, G. F. Hevel & J. A. Fortin, JSS 10529 (1 ♂, USNM); Carnarvon Station, Mailman Spring, 24 ° 51 ’ S, 147 ° 42 ’ E, 760 m, Eucalyptus woodland, 16. X. 2014, Wright, Lambkin, Starick, hand net, CNC 597045 (1 ♂, CNC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE1EC5DD8D9EFF6FD5417CC.taxon	distribution	Distribution: Australia (Queensland, South Australia) (Fig. 114). Notes: This is another species that occurs in the arid, inland areas ranging from desert to Eucalyptus woodland. Intraspecific genetic distance ranges from 0.0 % to 2.0 %. This species is genetically similar to T. wintertoni sp. nov. (2.3 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE7EC5BD8D9EFF6FC4E17E0.taxon	description	Figs 21 A – E, 112, 145 C, 152 B	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE7EC5BD8D9EFF6FC4E17E0.taxon	diagnosis	Diagnosis: This species can be recognized by dense long bristles covering the abdominal tergites (Fig. 152 B); hind trochanter with two patches of bristles ventroapically and ventrobasally (Fig. 145 C); elongated surstyli in dorsal view (Fig. 21 A); gonopods with two thumb-like projections extended towards surstyli, one of the ejaculatory ducts with some teeth in basal half in ventral view (Fig. 21 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE7EC5BD8D9EFF6FC4E17E0.taxon	description	Description: MALE: Body length: 4.0 – 4.2 mm. Head. Scape and pedicel brown, Flagellum tapering, yellow-brown. Face silvery-yellow pollinose. Frons, upper part shining black; lower part distinctly silver-yellow pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput silvery pollinose, covered by scattered short light bristles. Thorax. Postpronotum pale, light brown pollinose, with 4 – 5 long dark bristles along upper margin. Mesonotum (viewed obliquely from front) light brown pollinose, with long bristles at anterior supra-alar area in a row, extended to posterior and uniseriate rows of long intra-alar bristles. Scutellum silvery brown pollinose and with 24 – 26 long bristles along distal edge. Halter, knob, stem pale, tip brownish, stem with three short bristles dorsally. Legs. Fore coxa gray pollinose with 10 short brown bristles in a row. Mid coxa with 3 – 4 long dark bristles and 2 – 3 short light brown bristles. Coxae, trochanters, femora, tibiae dark brown, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise dark brown), tarsal segments brown. Hind trochanter with two patches of condensed spine-like dark bristle ventrobasally and ventroapically (Fig. 145 C); ventrobasal spines present on fore and mid femora. Fore and mid femora with some short bristles posteroventrally in basal half. Hind femur without ventroapical spines, except 8 – 12 equally spaced bristles posteroventrally in distal half. Subapical (distal) spines on first four tibiae present (short). Hind metatarsus distinctly flattened, extended toward one side, covered by erect bristles especially along lateral edge. Pulvilli shorter than last tarsal segment. Wing. Length: 3.9 – 4.1 mm. light brown color. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 1 – 3 long dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brown-black, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (spot on tergite 5 larger). All tergites covered by condensed long strong brown bristles as long as width of hind femur at distal end (Fig. 152 B). Lateral bristles on first tergite present, 32 – 38 long dark bristles up to as long as hind femur’s width at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 0.4 – 0.5 times as long as ST 8. Genitalia without dissection: ST 8 grey pollinose, covered by long bristles, round, globular in dorsal view, brown; membranous area is slit-like, mostly directed posteriorly, extended close to epandrium; Genitalia. Genital capsule in dorsal view: epandrium brown without pollinosity, surstyli brown; epandrium wider at base. Surstyli widened at base, elongated, slightly curved to each other at apex, left surstylus is longer (Fig. 21 A). Genital capsule in ventral view: both gonopods equal in height with a thumb-like projection extended towards surstyli, rounded at apex, with a distinct projection in their inner margins before apex; subepandrial sclerite distinct, with a fin-shaped projection in its center, phallus trifid, one of ejaculatory ducts with some teeth at the basal third; phallic guide with 3 – 5 spines dorso-dorsolaterally, one is longer (Fig. 21 B); Genital capsule in lateral view: left surstylus straight in basal two thirds, bent toward sternite in apical thirds, right surstylus short and rather straight (Fig. 21 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 21 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE7EC5BD8D9EFF6FC4E17E0.taxon	etymology	Etymology: From the Latin caesariatus, covered with hair or long-haired, in reference to the densely haired abdomen of this species. Examined material: HOLOTYPE: AUSTRALIA: Queensland: 3.5 km SW by S Mount Baird, 15 ° 10 ’ S, 145 ° 7 ’ E, 3. V. 1981, D. H. Colless, Malaise trap, JSS 8580 (1 ♂, ANIC); PARATYPES: AUSTRALIA: Queensland: 3.5 km SW by S Mount Baird, 15 ° 10 ’ S, 145 ° 7 ’ E, 4. V. 1981, D. H. Colless, Malaise trap, JSS 199 (1 ♂, ANIC); Brisbane, [27 ° 28 ’ S, 153 ° 1 ’ E], 23. VII. 1964, I. C. Yeo, JSS 8266 (1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE7EC5BD8D9EFF6FC4E17E0.taxon	distribution	Distribution: Australia (Queensland) (Fig. 112). Notes: The distribution of this species is suspicious, and the old Brisbane specimen could be mislabeled. It is strange that no other specimens have been found around Brisbane with the intensity of collecting there. More work should be conducted in the type locality to determine habitat preferences.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE5EC57D8D9EFF6FE191520.taxon	description	Figs 22 A – E, 116, 149 A, 150 E, 152 D	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE5EC57D8D9EFF6FE191520.taxon	diagnosis	Diagnosis: This species can be recognized by the presence of a Y-shaped spine along the posterior margin of abdominal sternite 4; hind trochanter with a row of short dark spines (Fig. 152 D); mid tibia with some extremely long apical spurs (Fig. 150 E); first and second mid tarsomeres with condensed bristles ventrally; surstyli elongated, restricted in basal quarter in dorsal view (Fig. 22 A); elongated gonopods, distinct hypandrial apodeme; small subepandrial sclerite in ventral view (Fig. 22 B); phallic guide with 4 – 5 dorso-dorsolateral spines (Fig. 22 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE5EC57D8D9EFF6FE191520.taxon	description	Description: MALE: Body length: 2.3 – 2.5 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1 – 1.5 times the length of ocellar triangle. Occiput silvery pollinose, slightly less so on upper 1 / 3. Thorax. Postpronotum pale, with 5 – 6 pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum brown pollinose and with 8 – 10 bristles along the distal edge. Dorsocentral bristles distinct, dark, somewhat longer along frontal edge, numerous long bristles around postpronotum present. Halter, knob pale, stem brown. Legs. Mid coxa with three long, dark bristles. Trochanters and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 5 of tibia yellow (tibia otherwise dark brown), tarsal segments yellow, ventrally paler, last segment darker. Hind trochanter with a row of short dark spines, longest spines distally (Fig. 149 A); ventrobasal spines absent on fore and mid femora. Ventroapical row of spines on fore femur absent; absent on mid femur; hind femur without ventroapical spines, except 10 – 12 bristles posteroventrally, equally spaced, longer on distal half, longest up to 1 / 3 of the width of hind tibiae at distal end. Subapical (distal) spines on first four tibiae present, three-six extremely long apical spurs only on mid tibia (as long as half the length of mid tibia) (Fig. 150 E). Hind tibia slightly bent in posterior view. First and second mid tarsomeres with condensed bristles ventrally. Metatarsi flattened, hind metatarsus almost as long as 2 – 5 combined. Pulvilli shorter than last tarsal segment. Wing. Length: 2.2 – 2.3 mm. Upper side of basal costal cell with one long dark bristle. Fourth costal section 2.5 – 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2 – 3 distinct brown setulae on tegula. Abdomen. Viewed obliquely from front tergites shiny black, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (larger on tergite 5). Dispersed short dark bristles on all tergites present, longest on tergite 5 up to the width of hind tibia at distal end. Lateral bristles on first tergite present, 5 – 6 dark bristles up to as long as hind femur’s width at base, sternite 4 with a specific Y shaped feature along posterior margin (Fig. 152 D). Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1.2 – 1.4 times as long as ST 8. Genitalia without dissection: ST 8 medium sized, round in dorsal view, dark brown and with some distinct bristles, otherwise velvet-like coverage; membranous area rounded elongate, mostly directed posterodorsally; epandrium dark brown, surstyli yellow, long and broad. Genitalia. Genital capsule in dorsal view: epandrium as long as wide (MLE: MWE = 1.0). Surstyli restricted at basal quarter, broadened in middle, gently curved to each other, right surstylus smaller than left one (Fig. 22 A). Genital capsule in ventral view: gonopods elongated to near surstyli, towards surstyli, tapering at apex, left is longer than right one; subepandrial sclerite small, hypandrium with distinct and rather long hypandrial apodeme (Fig. 22 B); phallic guide with 4 – 5 dorso-dorsolateral spines (one is longer). Genital capsule in lateral view: both surstyli curved towards sternite (Fig. 22 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 22 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE5EC57D8D9EFF6FE191520.taxon	etymology	Etymology: The name is derived from the Latin calcar for spur and longa for long in reference to the extremely long apical spurs on the mid tibia. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: N [ea] r Walyunga National Park, 31 ° 44 ’ S, 116 ° 2 ’ E, 18 – 29. XII. 1999, J. & A. Skevington, C. Lambkin & P. Bouchard, Malaise trap, JSS 7626 (1 ♂, WAM); PARATYPES: AUSTRALIA: Australian Capital Territory: Black Mountain Reserve, 35 ° 16 ’ S, 149 ° 6 ’ E, 4. I. 1980, Z. Liepa, JSS 9184, JSS 9186 (2 ♂, ANIC); Black Mountain, 35 ° 16 ’ S, 149 ° 6 ’ E, 15 – 17. II. 1980, 25. II. 1980, 26. II. 1980, 4 – 5. III. 1980, D. H. Colless, Malaise trap, Malaise site 2, JSS 8506, JSS 8510 – 1, JSS 8516 – 7 (5 ♂, ANIC); I. 1988, M. E. Irwin, Malaise trap, JSS 8533 (1 ♂, ANIC); 25. X. 1981, Z. Liepa, JSS 9189, JSS 9191 (2 ♂, ANIC); Black Mountain, Canberra, 35 ° 16 ’ S, 149 ° 6 ’ E, 18 – 23. II. 1985, W. W. Middlekauff, flight interception trap, JSS 8446, JSS 8449, JSS 8451, JSS 8453, JSS 8456 (5 ♂, CAS); near Condor Creek, 35 ° 19 ’ S, 148 ° 50 ’ E, 6. III. 1970, R. Lindsay, JSS 8496 – 7 (2 ♂, ANIC); New South Wales: Fitzroy Falls, 34 ° 36 ’ S, 150 ° 29 ’ E, 22. III. 1961, D. H. Colless, JSS 8638 (1 ♂, ANIC); Warrumbungle National Park, 31 ° 14 ’ S, 149 ° 1 ’ E, 18. X. 1997, J. Skevington & S. Winterton, Malaise trap, JSS 8398 (1 ♂, QM); 19. X – 3. XI. 1997, J. Skevington & S. Winterton, Malaise trap, JSS 8435 (1 ♂, QM); Warrumbungle National Park, Woolshed at Wombelong Creek, 31 ° 16 ’ S, 148 ° 57 ’ E, 420 m, rocky valley, cedar forest, 12 – 16. XII. 1995, M. E. Irwin, Malaise trap, JSS 14027, JSS 14029 - 30 (3 ♂, INHS); Picadilly Circus, 35 ° 22 ’ S, 148 ° 48 ’ E, 14 – 28. II. 1999, G. Gibson, Malaise trap, CNC 335235 (1 ♂, CNC); Northern Territory: Fogg Dam area, 12 ° 38 ’ S, 131 ° 15 ’ E, 9. IV. 1980, G. F. Hevel, J. A. Fortin, JSS 10534 (1 ♂, USNM); Baroalba Creek Springs, 19 km NE by E of Mount Cahill, 12 ° 48 ’ S, 132 ° 53 ’ E, 13. VI. 1973, D. H. Colless, JSS 8545 (1 ♂, ANIC); Queensland: 3 km NE of Mount Webb, 15 ° 3 ’ S, 145 ° 9 ’ E, 1. V. 1981, D. H. Colless, Malaise trap, JSS 8601 (1 ♂, ANIC); 2. X. 1980, D. H. Colless, Malaise trap, JSS 8571 (1 ♂, ANIC); Carnarvon National Park, Mount Moffatt Section, Mount Moffatt area, 25 ° 4 ’ S, 148 ° 3 ’ E, 19 – 23. I. 1998, J. & A. Skevington & S. Winterton, Malaise trap, JSS 2485 (1 ♂, QM); 24 ° 4 ’ S, 148 ° 3 ’ E, 10. X. 2002, S. Boucher, sweep net, LEM _ 0016181 (1 ♂, LEM); 25 ° 4 ’ S, 148 ° 3 ’ E, 10. X. 2002, S. E. Brooks, sweep net, LEM _ 0016176 (1 ♂, LEM); Eclectus ‚ Iron Range, 12 ° 46 ’ S, 143 ° 17 ’ E, 20 m, 16. VII. 1997, G. & A. Daniels, JSS 8926 (1 ♂, QM); Lonesome National Park, near Lookout (LNP 4 M), 25 ° 30 ’ S, 148 ° 49 ’ E, closed Eucalyptus woodland on rocky ridge, 26. XI. 2010 – 11. I. 2011, D. Beard & B. Sigley, Malaise trap, 19382, CNC 575011 – 12 (1 ♂, CNC); Mount Moffatt summit, 25 ° 4 ’ S, 148 ° 3 ’ E, 18. XI. 2014, J. H., A. M. & A. W. Skevington, CNC 374966 (1 ♂, CNC); Prince of Wales Island, Torres Strait, 10 ° 41 ’ S, 142 ° 11 ’ E, 27 – 30. V. 1969, Neboiss, JSS 8794 (1 ♂, MVMA); Upper Crossing, Teviot Brook, Boona – Killarney Road, SE Queensland, 21 ° 58 ’ S, 148 ° 3 ’ E, 23. III. 1975, B. K. Cantrell, JSS 8934 (1 ♂, QDPC); Victoria: 12 km southeast Merrijig, Howqua River, 37 ° 12 ’ S, 146 ° 22 ’ E, 30. XI. 1971, Neboiss, JSS 8810 – 1, JSS 8813, JSS 8816 (4 ♂, MVMA); Dartmouth Survey, Dart-Mitta River Junction, Locality L, 36 ° 31 ’ S, 147 ° 27 ’ E, 4. III. 1973, JSS 8761 – 3 (3 ♂, MVMA); Dart – Mitta River Junction, Locality L, 36 ° 31 ’ S, 147 ° 27 ’ E, 4. III. 1973, Dartmouth Survey, JSS 8759; JSS 8770; JSS 8775 (3 ♂, MVMA); Eight Mile Creek, Locality JA, 37 ° 13 ’ S, 146 ° 26 ’ E, 12. IV. 1973, Dartmouth Survey, JSS 8752 – 3 (2 ♂, MVMA); Mitta River, 546493 Loc. CB, 36 ° 31 ’ S, 147 ° 27 ’ E, 28. XI – 3. XII. 1973, Dartmouth Survey, JSS 8758 (1 ♂, MVMA); Western Australia: Near Walyunga National Park, 31 ° 44 ’ S, 116 ° 2 ’ E, 18 – 29. XII. 1999, J. & A. Skevington, C. Lambkin & P. Bouchard, Malaise trap, JSS 7625 (1 ♂, QM); Neerabup Lake National Park, 31 ° 38 ’ S, 115 ° 43 ’ E, 18 – 29. XII. 1999, J. & A. Skevington, C. Lambkin & P. Bouchard, Malaise trap, JSS 7634 (1 ♂, QM); Walyunga National Park, 31 ° 44 ’ S, 116 ° 4 ’ E, 19 – 29. XII. 1999, J. & A. Skevington, C. Lambkin & P. Bouchard, Malaise trap, JSS 7674 (1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE5EC57D8D9EFF6FE191520.taxon	distribution	Distribution: Australia (Australian Capital Territory, New South Wales, Northern Territory, Queensland, Western Australia, Victoria) (Fig. 116). Notes: The Y-shaped spine along the posterior margin of abdominal sternite 4 is similar to Tomosvaryella neboissi sp. nov. but terminalia characters are different. Tomosvaryella calcarilonga sp. nov. is restricted to forested parts of Australia and occurs in both wet and dry forests. Intraspecific genetic distance ranges from 0.0 % to 1.6 %. This species is genetically most similar to T. cona sp. nov. (0.2 – 2.7 % pairwise divergence). Despite the overlap, these species are morphologically divergent and in the absence of additional data are treated as separate species (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFEBEC55D8D9EAC7FCAC12F0.taxon	description	Figs 23 A – E, 116	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFEBEC55D8D9EAC7FCAC12F0.taxon	diagnosis	Diagnosis: This species can be recognized by the distinct ventroapical spine on the hind trochanter; elongated surstyli, left one longer and club-shaped in dorsal view (Fig. 23 A); both gonopods with a thumb-like projection towards surstyli; phallic guide with 3 – 4 different-sized dorso- dorsolateral spines, one is longer and sinuous; subepandrial sclerite with condensed transverse wrinkles in ventral view (Fig. 23 B); phallus bearing some teeth apically before ejaculatory ducts in lateral view (Fig. 23 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFEBEC55D8D9EAC7FCAC12F0.taxon	description	Description: MALE: Body length: 2.6 mm. Head. Scape, pedicel, flagellum light brown. Flagellum tapering. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput silvery pollinose with scattered short dark bristles. Thorax. Postpronotum pale, with 2 – 3 short pale bristles. Mesonotum (viewed obliquely from front) silvery pollinose, anterior part close to postpronotum, greyer. Scutellum silvery pollinose and with 6 – 8 very short bristles along distal edge. Dorsocentral bristles short, dark, longer in frontal half, numerous pale bristles around postpronotum present. Halter, knob and stem pale, tip brownish. Legs. Mid coxa with 3 – 6 different-sized dark bristles. Trochanters brown, femora darker, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments light brown. Hind trochanter grey pollinose with a distinct ventroapical spine; ventrobasal spines present (1) on fore and mid femur. Hind femur without ventroapical spines. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres not flattened, hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect dark bristles. Pulvilli shorter than last tarsal segment. Wing. (damaged). Upper side of basal costal cell with 2 – 3 short to medium dark brown bristles. Abdomen. Viewed obliquely from front tergites brown-black, tergite 1 silvery grey. Lateral bristles on first tergite present, 4 – 6 dark bristles up to as long as ¾ of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium rather quadratic-shaped (MLE: MWE = 1.02). Surstyli elongated, left longer and wider than right one, both surstyli widened at the basal fourth, narrowed in the middle, broadened at the apex, left one club-shaped, both curved to each other at apex (Fig. 23 A). Genital capsule in ventral view: both gonopods widened at base with a thumb-like projection towards surstyli; phallic guide with 3 – 4 different-sized dorso- dorsolateral spines, one is longer and sinuous; subepandrial sclerite widened in the middle with condensed transverse wrinkles (Fig. 23 B); Genital capsule in lateral view: both surstyli straight in basal two thirds, left one bent towards sternite apically, right one slightly curved towards sternite apically. Phallus trifid with some teeth apically before ejaculatory ducts (Fig. 23 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 23 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFEBEC55D8D9EAC7FCAC12F0.taxon	etymology	Etymology: The name is from the Latin clava, club or cudgel and refers to the club-shaped left surstylus. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: Millstream-Chichester Nat [ional] P [ark], Roebourne Road, 21 ° 26 ’ S, 117 ° 9 ’ E, 337 m, sandy creek, Eucalyptus Spinifex grassland, 27. IV – 3. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16075 (1 ♂, WAM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFEBEC55D8D9EAC7FCAC12F0.taxon	distribution	Distribution: Australia (Western Australia) (Fig. 116).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE9EC53D8D9EDB7FD0E11FC.taxon	description	Figs 24 A – E, 117, 148 D, 154 E	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE9EC53D8D9EDB7FD0E11FC.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter with few dark tiny spines in one group ventrobasally (Fig. 148 D); elongated surstyli in dorsal view (Fig. 24 A); elongated gonopods fused with subepandrial sclerite, with two inward projections (Fig. 24 B); hypandrium with two hairy membranous sheaths (Fig. 24 B); subepandrial sclerite with a small fin-shaped protrusion (Fig. 24 B); one ejaculatory duct with some spines (Fig. 24 D – E); phallic guide with some dorso- dorsolateral spines in lateral view (Fig. 24 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE9EC53D8D9EDB7FD0E11FC.taxon	description	Description: MALE: Body length: 2.6 – 2.9 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 2 – 3 pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum silvery pollinose and with 6 – 8 bristles along the distal edge. Dorsocentral bristles distinct, dark, somewhat longer along frontal edge, numerous dark bristles around postpronotum present. Halter, knob pale with brownish tinge, stem brown. Legs. Mid coxa with 2 – 3 long brown bristles. Trochanters and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments brown, ventrally yellow-brown, last segment darker. Hind trochanter with few dark tiny spines in one group ventrobasally (“ echidna ” - like), elongated troch with flat ventral part (Fig. 148 D); ventrobasal spines (1 or 2) present on fore and present on mid femur. Ventroapical row of spines on fore femur 5 – 7; 6 – 8 small, black spines on mid femur; hind femur without ventroapical spines, except 14 – 16 equally spaced bristles posteroventrally, longer on distal half, longest up to 1 / 3 the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present. Hind tarsomeres flattened, hind metatarsus almost as long as 2 – 5 combined. Pulvilli shorter than last tarsal segment. Wing. Length: 3.0 mm. Upper side of basal costal cell with one long dark bristle and three short light brown bristles. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m slightly distal to middle of discal cell. 2 – 3 distinct brown setulae on tegula. Abdomen. Viewed obliquely from front tergites brownish pollinose, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (similar size). Dispersed strong dark bristles on all tergites present, longest on tergite 5 up to ½ the width of hind tibia at distal end. Lateral bristles on first tergite present, 4 – 5 dark bristles up to 1.1 – 1.3 times as long as hind femur’s width at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 0.8 – 1.0 times as long as ST 8. Genitalia without dissection: ST 8 large, broad, round in dorsal view, brown and with distinct bristles, otherwise velvet-like coverage; membranous area with narrow part approaching epandrium and rounded part mostly directed dorsally; epandrium brown, surstyli yellow, long. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.7). Surstyli symmetrical, elongated, right surstylus smaller than left one, right one with a small protrusion at inner margin before apex (Fig. 24 A). Genital capsule in ventral view: gonopods equal in height, elongated towards surstyli and fused with subepandrial sclerite, with two small inward projection before apex, subepandrial sclerite rather wide with a small fin-shaped protrusion in its center; hypandrium with two hairy membranous sheaths extended towards phallus (Fig. 24 B). Genital capsule in lateral view: both surstyli curved towards sternite, phallus with three short ejaculatory ducts, one with three small spines, phallic guide with 6 – 7 dorso-dorsolateral spines (one is longer) (Fig. 24 D – E); Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 24 C). FEMALE: Body length: 2.5 – 2.6 mm. As male except for the following characters. Frons, eyes separated, as wide as 1.7 – 1.9 times the width at antennae; completely silver-grey pollinose until up to 1 / 3 of the lower frons, shiny black in front of ocellar triangle, no transition zone; frons depressed along upper 2 / 3 with a groove in the median line. Enlarged ommatidia silvery shining. Pulvilli and claws about 1.5 – 2 times as long as last tarsal segment, smaller on metatarsi. Female abdomen with silver spots on sides of tergites 4 – 6. Ovipositor. Piercer straight in ventral view, straight and robust in lateral view, reaching distal end of 2 nd segment; base brownish pollinose with occasional bristles. LP: LB = 2.0. LDP: LPP = 2.0. (Fig. 154 E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE9EC53D8D9EDB7FD0E11FC.taxon	etymology	Etymology: This species is named after Don Colless, who collected the holotype and several other specimens. Don collected 8 % of the specimens used in this revision and 28 % of the species (Supplementary file 2). Examined material: HOLOTYPE: AUSTRALIA: Queensland: 14 km W by N Hope Vale Mission, 15 ° 16 ’ S, 144 ° 59 ’ E, 8. V. 1981, D. H. Colless, Malaise trap, JSS 8561 (1 ♂, ANIC); PARATYPES: AUSTRALIA: Queensland: 1 km North of Eclectus ‚ Iron Range, 12 ° 46 ’ S, 143 ° 17 ’ E, 5. VII. 1997 20 m, G. & A. Daniels, JSS 8925 (1 ♂, QM); 14 km W by N Hope Vale Mission, 15 ° 16 ’ S, 144 ° 59 ’ E, 9. V. 1981, D. H. Colless, Malaise trap, JSS 8559 - 60, JSS 8562 (3 ♂, ANIC); Cooloola National Park, East Mullen hilltop, 26 ° 0 ’ S, 152 ° 59 ’ E, dry sclerophyll forest, 15. XI. 2014, J. H., A. M. & A. W. Skevington, CNC 374117 – 8, CNC 384971 (3 ♂, CNC); Cape York, Stewart Range, 14 ° 4 ’ S, 143 ° 41 ’ E, 500 m, 29. IX – 3. V. 1961, L. & M. Gressitt, CNCD 5551 (1 ♂, BPBM); Mount Cook National Park, 15 ° 29 ’ S, 145 ° 16 ’ E, 10. V. 1981, D. H. Colless, Malaise trap, JSS 183 (1 ♂, ANIC); Proserpine, near Kelsey Substation, XY 19, 20 ° 23 ’ S, 148 ° 32 ’ E, 21 m, closed forest, 3 – 10. V. 2007, E. J. Stanisic, Malaise trap, CNCD 160423 – 34 (12 ♂, QM); Buderim, SE Queensland, 26 ° 41 ’ S, 153 ° 3 ’ E, 14 – 22. III. 1985, G. K. Waite, Malaise trap, JSS 8948 (1 ♂, QDPC); Carnarvon National Park, Consuelo land, Race Course Spring, 24 ° 56 ’ S, 148 ° 5 ’ E, Vegetation, 12. X. 2002, S. Boucher, sweeping, LEM _ 0016156 – 7 (2 ♂, LEM); Cooloola National Park, East Mullen hilltop, 26 ° 0 ’ S, 152 ° 59 ’ E, dry sclerophyll forest, 15. XI. 2014, J. H., A. M. & A. W. Skevington, CNC 374119 (1 ♂, CNC); Northern Territory: Barramundi Creek, 13 ° 19 ’ S, 132 ° 26 ’ E, 14. V. 1998, R. Lynch, Malaise trap, JSS 7905 (1 ♀, QM); Fogg Dam area, 12 ° 38 ’ S, 131 ° 15 ’ E, 9. IV. 1980, G. F. Hevel, J. A. Fortin, JSS 10533 (1 ♂, USNM); Queensland: Carnarvon National Park, Mount Moffatt section, 25 ° 4 ’ S, 148 ° 2 ’ E, 27 – 28. XI. 1997, [dry sclerophyll, along permanent creek], J. Skevington & C. Lambkin, Malaise trap, JSS 2293 (1 ♂, QM); Gunshot Creek, 13 km NW Heathlands Homestead, 11 ° 42 ’ S, 142 ° 26 ’ E, 23. III. 1992, G. Daniels, M. A. Schneider, JSS 8270 (1 ♂, QM); Mount Cook National Park, 15 ° 29 ’ S, 145 ° 16 ’ E, 10. V. 1981, D. H. Colless, Malaise trap, JSS 8557 (1 ♂, ANIC); Mica Creek, Mount Isa, 20 ° 49 ’ S, 139 ° 27 ’ E, 17 – 22. III. 2001, 360 m, D. C. Darling, Malaise trap, JSS 19188 (1 ♂, QM); 20 km E Mareeba, 17 ° 0 ’ S, 145 ° 26 ’ E, 15. IV. 1980, G. F. Hevel & J. A. Fortin, JSS 10523, JSS 10532 (2 ♂, USNM); Gordonvale, 17 ° 5 ’ S, 145 ° 47 ’ E, VII. 1918, JSS 10512 (1 ♂, USNM); New South Wales: Warrumbungle National Park, 31 ° 16 ’ S, 149 ° 0 ’ E, 17 – 22. XII. 1998, in creek bed [in Eucalyptus forest], J. & A. Skevington & S. Winterton, Malaise trap, JSS 5063 (1 ♂, QM); Arnhem Land, Maningrida, 12 ° 3 ’ S, 134 ° 13 ’ E, 5 m, 20 – 21. III. 1961, L. & M. Gressitt, CNCD 5562 (1 ♂, BPBM); Darwin, 12 ° 27 ’ S, 130 ° 50 ’ E, 8 – 9. XII. 1963, J. Sedlacek, CNCD 5572 (1 ♂, ANIC); Victoria: Dartmouth Survey, Dart-Mitta River Junction, Locality L, 36 ° 31 ’ S, 147 ° 27 ’ E, 4. III. 1973, JSS 8772 (1 ♂, MVMA); Dartmouth Survey, Mitta Mitta River, Loc. EA, 36 ° 31 ’ S, 147 ° 27 ’ E, 9. II. 1973, JSS 8768 – 9 (2 ♂, MVMA); 12 km SE Merrijig, Howqua River, 37 ° 12 ’ S, 146 ° 22 ’ E, 30. XI. 1971, Neboiss, JSS 8781 (1 ♂, MVMA); Dartmouth Survey, Dart-Mitta River Junction, Locality L, 36 ° 31 ’ S, 147 ° 27 ’ E, 4. III. 1973, JSS 8754 – 5, JSS 8766, JSS 8773, JSS 8777 (5 ♂, MVMA); South Australia: Mount Lofty, 35 ° 0 ’ S, 138 ° 44 ’ E, 28. XII. 0 – 6. I. 1974, 10 th Australian Boy Scout Jamboree, JSS 8718 (1 ♂, SAM). Non-type material examined: Australia: Queensland: Sybella Creek, 20 ° 53 ’ S, 139 ° 27 ’ E, 17 – 22. III. 2001, Malaise trap, JSS 50563 (1 ♂, QM); Western Australia: 158 km S Newman, 9 km N Kumarina Road House, 24 ° 38 ’ S, 117 ° 37 ’ E, 638 m, in wide sandy wash, 18 – 21. V. 2003, M. E. Irwin, F. D. Parker, Malaise trap, JSS 16415 (1 ♂, CNC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFE9EC53D8D9EDB7FD0E11FC.taxon	distribution	Distribution: Australia (New South Wales, Northern Territory, Queensland, South Australia; Victoria, Western Australia) (Fig. 117). Notes: This is a coastal forest species found from Darwin to Adelaide in the eastern half of the country. The two specimens from Western Australia and western Queensland that are found in arid regions break this pattern and were not included in the type series. These two specimens, JSS 50563 and JSS 16415, are similar to the holotype but there is a little difference in the shape of the gonopods and subepandrial sclerites in ventral view. The left gonopod is a little longer and the subepandrial sclerite is missing the two small lobes in the middle of outer margin. This is suggestive that these specimens should be treated as a distinct new species but as these characters vary in some other Tomosvaryella species we have chosen to treat them as part of T. collessi sp. nov. for now. We need fresh samples to allow us to compare DNA sequence data and to see if these characters hold up across more specimens. Intraspecific genetic distance ranges from 0.0 % to 3.7 %. Specimen JSS 8562 shows the same morphological features as genetically divergent specimens (pairwise distance 1.9 – 3.7 %). This could be another species complex so more specimens and molecular work are needed. This species is genetically most similar to T. biflocca sp. nov. (3.2 – 5.8 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFECEC2ED8D9EFF6FAA11624.taxon	description	Figs 25 A – E, 118, 157 C	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFECEC2ED8D9EFF6FAA11624.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a keel covered with erect bristles along its edge; surstyli gradually broadened towards apex in dorsal view (Fig. 25 A); gonopods cone-shaped, extended towards surstyli in ventral view (Fig. 25 B); phallic guide with some spines in ventral view (Fig. 25 B); both surstyli straight at basal two thirds and bent towards sternite in lateral view (Fig. 25 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFECEC2ED8D9EFF6FAA11624.taxon	description	Description: MALE: Body length: 2.0 – 2.2 mm. Head. Flagellum tapering; yellow-brown, as long as wide. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver-brown pollinose; eyes touching for distance equal to 1.2 times the length of ocellar triangle. Occiput silvery pollinose. Thorax. Postpronotum light yellow, with 2 – 3 short pale bristles. Mesonotum (viewed obliquely from front) brown, upper side darker; brown pollinose. Scutellum light brown, silvery brown pollinose and without bristles along distal edge. Dorsocentral bristles distinct, dark, mostly in frontal half, as long as ¾ width of hind femur at base, pale bristles around postpronotum absent; with 3 – 5 postalar bristles. Halter, knob and stem light yellow, base brown, stem with 3 – 4 short pale bristles. Legs. Mid coxa with 3 – 5 long dark bristles. Trochanters brown, femora black, silvery pollinose posteriorly, tarsal segments yellow-brown dorsally, yellow ventrally, last segment darker. Hind trochanter with small keel, and some short, erect and condensed dark bristles along its edge, ventrobasal spines absent on fore and mid femur. Hind femur without ventroapical spines, with some bristles posteroventrally. Hind tarsomeres obviously flattened. Pulvilli shorter than last tarsal segment. Wing. Length: 2.0 – 2.1 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m at middle of discal cell. 1 – 2 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brown, tergite 1 silvery grey except a narrow band at its lower margin. Dispersed short dark bristles on all tergites present. Lateral bristles on first tergite present, 2 – 6 dark bristles up to as long as hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium rather longer than wide (MLE: MWE = 1.08). Surstyli symmetrical, elongated, constricted before the base, gradually broadened towards apex, curved to each other, left surstylus a little longer than right one (Fig. 25 A). Genital capsule in ventral view: gonopods equal in height, extended towards surstyli, cone-shaped, both gonopods with a distinct projection on inner margin, right one is clearly distinct; phallic guide with 4 – 6 dorso- dorsolateral spines, one is longer than others; subepandrial sclerite short (Fig. 25 B); Genital capsule in lateral view: both surstyli straight in basal three fourths and bent towards sternite apically (Fig. 25 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 25 C). FEMALE: Body length: 2.6 – 2.7 mm. As male except for the following characters. Frons, eyes separated, as wide as 1.9 – 2.3 times the width at antennae, wider in the middle; completely silver-grey pollinose, shiny black in front of ocellar triangle, no transition zone. Enlarged ommatidia silvery shining. For femur with two distinct bristles ventrobasally. Hind tibia with condensed short bristles ventrobasally. Tarsi covered with erected bristles, longer on metatarsi. Pulvilli and claws about 1.5 times as long as last tarsal segment. Ovipositor. Piercer rather straight in ventral view, very long, rather straight and upwards at tips in lateral view, reaching distal end of 2 nd segment; base brownish pollinose with occasional bristles. LP: LB = 2.0. LDP: LPP = 3.0. (Fig. 157 C).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFECEC2ED8D9EFF6FAA11624.taxon	etymology	Etymology: From the Latin conus, cone-like, in reference to the cone-shaped gonopods. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Nairana National Park (NR 2 M), 21 ° 34 ’ S, 146 ° 55 ’ E, 196 m, closed Acacia woodland, 10. XI – 7. XII. 2010, R. Raven, H. & D. Hanrahan, Malaise trap, 18497, CNC 576506 (1 ♂, QM); PARATYPES: AUSTRALIA: Queensland: same details as holotype, CNC 576500, CNC 576503 (2 ♂, CNC, QM); Near Rosedale, 24 ° 37 ’ S, 151 ° 54 ’ E, 21. VII. 1975, H. Frauca, JSS 8524 (1 ♂, ANIC); Carnarvon National Park, Mount Moffatt section, 25 ° 3 ’ S, 147 ° 53 ’ E, dry sclerophyll, savanna opening, 1 – 2. XII. 1997, J. Skevington & C. Lambkin, Malaise trap, JSS 2276 (1 ♂, QM); Ridgepole Waterhole, 24 km eastsoutheast of Musselbrook Resource Centre, Lawn Hill National Park, 18 ° 40 ’ S, 138 ° 22 ’ E, 180 m, 1. V. 1995; 12. V. 1995; 4. V. 1995; 6. V. 1995; 8. V. 1995, G. Daniels & M. A. Schneider, JSS 8281; JSS 8286; JSS 8288 – 89; JSS 8297 (4 ♂, 1 ♀, QM); Scrub Road, Brisbane Forest Park, 27 ° 25 ’ S, 152 ° 50 ’ E, 10 – 17. X. 1997; 2 – 9. I. 1998, S. Winterton, N. Power & D. White, Malaise trap, JSS 8412, JSS 8440 (2 ♂, QM); New South Wales: Colo Heights, Putty Road, 33 ° 23 ’ S, 150 ° 45 ’ E, 6. II. 1968, D. H. Colless, JSS 8666 (1 ♂, ANIC); Coolabah, 31 ° 0 ’ S 146 ° 42 ’ E, 12. X. 1963, D. H. Colless, JSS 8663 (1 ♂, ANIC); Nelligen (tidal flat), 35 ° 38 ’ S, 150 ° 8 ’ E ,, 1. II. 1973, D. H. Colless, JSS 8500 – 1 (2 ♂, ANIC); Rotary Lookout, Pigeon House Range NW of Milton, 19 ° 30 ’ S, 140 ° 19 ’ E; 22. II. 1979, Z. Liepa, JSS 9167 – 8 (2 ♂, ANIC); Culgoa National Park, 8 km West-Northwest Cawwell Homestead, Diemunga Lagoon (CGN 2 M), 29 ° 3 ’ S, 146 ° 60 ’ E, Coolibah, 30. I – 18. V. 2010, C. Lambkin, R. Olsen, B. Shieban, Malaise trap, CNC 575133 (1 ♂, QM); Ledknapper National Reserve, 5 km N Beulah Homestead (LDN 1 M), 28 ° 19 ’ S, 147 ° 10 ’ E, Allocasuarina, 18. III – 15. V. 2010, S. O’Sullivan, C. Lambkin, Malaise trap, CNC 576717, CNC 576719 (2 ♂, CNC, QM); Mount Kaputar National Park, Dawson ‚ Spring, 30 ° 17 ’ S, 150 ° 10 ’ E, 1 – 5. XII. 1987, G. R. Brown, yellow pan trap, Moericke trap, JSS 8745 (1 ♀, ASCU); Warrumbungle National Park, 31 ° 14 ’ S, 149 ° 1 ’ E, 18. X. 1997, J. Skevington & S. Winterton, Malaise trap, JSS 8399 (1 ♂, QM); Northern Territory: Namatijira Drive, 20 km W Hugh River Crossing, 23 ° 49 ’ S, 133 ° 11 ’ E, 760 m, 14 – 15. III. 2002, C. J. Burwell, JSS 13933 (1 ♀, QM); Standley Chasm, 23 ° 44 ’ S, 133 ° 28 ’ E, 750 m, 14. III. 2002, C. J. Burwell, sweep net, JSS 13931 (1 ♂, QM); South Australia: Koonchera Dune near Koonchera Waterhole, 26 ° 41 ’ S, 139 ° 30 ’ E, 3. IX. 1997, J. & A. Skevington, JSS 8341 (1 ♀, QM); Tasmania: Chauncy Vale Wildlife Sanctuary Near Bagdad, 42 ° 37 ’ S, 147 ° 15 ’ E, 17 – 27. XII. 1998, J. & A. Skevington, Malaise trap, JSS 4836; JSS 4839; JSS 4843 (3 ♂, QM); S. Winterton, J. & A. Skevington, D. Yeates, Malaise trap, JSS 4833 (1 ♂, QM); Cradle Mountain National Park, 41 ° 38 ’ S, 145 ° 57 ’ E, in flowering heath, 22. XII. 1998 – 8. I. 1999, J. & A. Skevington, Malaise trap, JSS 4685; JSS 4725; JSS 4735 (2 ♂, 1 ♀, QM); Western Australia: Karijini National Park, Juna Downs Road, 22 ° 42 ’ S, 118 ° 25 ’ E; 22 ° 43 ’ S, 118 ° 25 ’ E, 789 m; 799 m, between drying pools in rocky Turee Creek; near rock outcrop, Eucalyptus Eremophila scrub, 19 – 25. IV. 2003; 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, ANIC [Bulk Sample] 2059; ANIC [Bulk Sample] 2040, JSS 16058; JSS 16253 (2 ♂, ANIC); Karijini National Park, Weano Gorge Road, 22 ° 22 ’ S, 118 ° 15 ’ E, 775 m, on hilltop, open Eucalyptus, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, ANIC Bulk Sample 2052, JSS 15210; JSS 16138; JSS 16143; JSS 16147 (4 ♂, ANIC); Millstream – Chichester National Park, Black Hill Pool, 21 ° 20 ’ S, 117 ° 15 ’ E, 176 m, over wide dry rocky creek bed, Eucalyptus, 7 – 12. V. 2003, C. Lambkin & D. Yeates, Malaise trap, ANIC [Bulk Sample] 2106, JSS 16315 (1 ♂, ANIC); 82 km S junction Karijini Drive on Great Northern Highway, 23 ° 7 ’ S, 119 ° 6 ’ E, 694 m, wash with pools, 23. IV – 6. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15674 (1 ♂, CNC); Karijini National Park, Banjima Road, 22 ° 24 ’ S, 118 ° 16 ’ E, 690 m, in creek with pools, unburnt Eucalyptus grassland, 15 – 18. V. 2003, C. Lambkin, J. Recsei & N. Starick, Malaise trap, JSS 15975 (1 ♂, ANIC); Karijini National Park, Juna Downs Road, 22 ° 42 ’ S, 118 ° 25 ’ E; 22 ° 44 ’ S, 118 ° 25 ’ E, 789 m; between drying pools in rocky Turee Creek; dry Turee Creek, grassy open Eucalyptus scrub, 19 – 25. IV. 2003; 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16166; JSS 16090; JSS 16093; JSS 16103 (3 ♂, 1 ♀, ANIC); Karijini National Park: Juna Downs Road (between drying pools in rocky Turee Creek), 22 ° 42 ’ S, 118 ° 25 ’ E, 789 m, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, CNCD 3825; CNCD 3832 – 3; CNCD 3839; CNCD 3862 (3 ♂, 2 ♀, CNC); Near Walyunga National Park, 31 ° 44 ’ S, 116 ° 2 ’ E, 18 – 29. XII. 1999, J. & A. Skevington, C. Lambkin & P. Bouchard, Malaise trap, JSS 7627 (1 ♂, QM); Neerabup Lake National Park, 31 ° 38 ’ S, 115 ° 43 ’ E, 18 – 29. XII. 1999, J. & A. Skevington, C. Lambkin & P. Bouchard, Malaise trap, JSS 7637 (1 ♂, QM); Pebble Mouse Creek on Great Northern Highway, 62 km E Junction, Karijini Drive, 23 ° 5 ’ S, 118 ° 55 ’ E, 713 m, damp river bed, 23. IV – 6. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15880 (1 ♂, CNC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFECEC2ED8D9EFF6FAA11624.taxon	distribution	Distribution: Australia (New South Wales, Queensland, South Australia, Tasmania, Western Australia) (Fig. 118). Notes: This species occurs from the coast to Acacia - dominated habitats in inland Queensland and New South Wales. Tomosvaryella cona is morphologically and genetically similar to T. calcarilonga sp. nov. (1.1 – 2.1 % pairwise distance). In T. calcarilonga sp. nov., the surstyli are wider in the middle in dorsal view, there is a Y-shaped spine along the posterior margin of abdominal sternite 4 and extremely long apical spurs are present on the mid tibia (Fig. 150 E). These character differences are consistent so despite other similarities we recognize two species.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF92EC2CD8D9E9C3FB6514B0.taxon	description	Figs 26 A – D, 119	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF92EC2CD8D9E9C3FB6514B0.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of its surstyli in dorsal view, straight in the basal half, falciform in the apical half (Fig. 26 A); gonopods extended towards surstyli; phallic guide with three equal-sized dorsolateral downward-directed spines; subepandrial sclerite and hypandrial apodeme distinct in ventral view (Fig. 26 B); phallic guide widened at the middle and sinuous in lateral view (Fig. 26 C – D).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF92EC2CD8D9E9C3FB6514B0.taxon	description	Description: MALE: Body length: 2.5 mm (from head to first segment of abdomen). Head. Scape and pedicel dark brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1 – 1.2 times the length of ocellar triangle. Occiput grey pollinose, covered by short dark bristles. Thorax. Postpronotum light yellow, with 2 – 6 distinct light brown bristles. Mesonotum (viewed obliquely from front) silvery pollinose, anterior part greyer; scutum with scattered bristles at anterior supra-alar area. Postalar callus light brown with 3 – 4 brown bristles in upper margin. Scutellum black, silvery pollinose with 8 – 10 bristles along distal edge. Halter, knob pale, stem brown. Legs. Mid coxa with two dark bristles. Trochanters brown, femora dark brown, shining ventrally, knees and basal 1 / 4 of tibiae yellow (tibiae otherwise brown), tarsal segments yellow ventrally and brown dorsally, last segment darker. Hind trochanter silvery pollinose, without specific feature; ventrobasal spines absent on fore and mid femur, two rows ventroapical present. Hind femur without ventroapical spines. Tibiae with two rows of short black bristle on anterior and three rows on posterior side. Hind tarsomeres distinctly flattened (especially tarsomeres 1 – 3), hind metatarsus as long as 2 – 4 combined; posterior corners with distinct bristles on all tarsomeres. Pulvilli shorter than last tarsal segment. Wing. Length: 3.5 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 1.5 – 2 times as long as third costal section. Cross-vein r-m at middle of discal cell. 1 – 2 short dark setulae on tegula. Abdomen. Dissected except tergite 1. Tergites 1 dark, silvery pollinose. Lateral bristles on first tergite present, 6 – 10 dark bristles up to as long as 3 / 4 hind femur’s width at base, distributed from lateral to its middle. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.2). Surstyli symmetrical, basal half straight, small depression ventromedially, apical half falciform (Fig. 26 A). Genital capsule in ventral view: gonopods equal in height, both gonopods extended towards surstyli; phallic guide with three same sized dorsolateral downward spines; subepandrial sclerite wide and distinct with a crescent-shaped wrinkle in its center; hypandrial apodeme long and distinct (Fig. 26 B); Genital capsule in lateral view: both surstyli widened and rather rounded at basal two thirds, narrowed and curved towards sternite at apical third; phallic guide widened at the middle and sinuous (Fig. 26 C – D). FEMALE: unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF92EC2CD8D9E9C3FB6514B0.taxon	etymology	Etymology: From the Greek kreagra, meathook, in reference to the hooked surstyli. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: Pine Hill, Cape Arid N [ational] P [ark], Balladonia Road, 33 ° 18 ’ S, 123 ° 22 ’ E, 130 m, mallee woodland, 31. X – 18. XI. 2003, C. Lambkin & J. Recsei, Malaise trap, JSS 15595 (1 ♂, WAM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF92EC2CD8D9E9C3FB6514B0.taxon	distribution	Distribution: Australia (Western Australia) (Fig. 119). Notes: The only known specimen was collected in mallee woodland in Cape Arid National Park. This species is genetically most similar to T. irwini sp. nov. (8.2 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF90EC2AD8D9EA77FC42142C.taxon	description	Figs 27 A – D, 116	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF90EC2AD8D9EA77FC42142C.taxon	diagnosis	Diagnosis: This species can be recognized by a few short dark bristles on the hind trochanter ventromedially; elongated and slender surstyli in dorsal view (Fig. 27 A); gonopods more elongated towards surstyli in ventral view (Fig. 27 B); phallic guide with four spines dorsomedially, one is longer than others (Fig. 27 B – D).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF90EC2AD8D9EA77FC42142C.taxon	description	Description: MALE: Body length: 2.2 – 2.4 mm. Head. Flagellum acuminate; yellow. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose; eyes touching for distance equal to 1.2 – 1.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper half. Thorax. Postpronotum pale, with 3 – 4 pale bristles. Mesonotum (viewed obliquely from front) silvery pollinose, anterior part more grey, grayish also from the side. Scutellum brownish pollinose and without bristles along the distal edge. Dorsocentral bristles undeveloped, indistinct, somewhat visible along frontal edge, a few pale bristles around postpronotum. Halter, knob pale, stem brown. Legs. Mid coxa with two long and two short, pale bristles. Trochanters and femora brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees, basal 1 / 5 and distal extremity of tibiae yellow (tibiae otherwise brown), tarsal segments yellow, base of last segment slightly brown. Hind trochanter without a keel, but with a few short dark bristles (5 – 7) in a groups ventromedially. Ventroapical row of spines on fore femur 3 – 4 very weak; 5 – 7 short, black spines on mid femur; no spines on hind femur, only 4 – 5 stronger dark bristles at 2 / 3 from base, where femur is wider (up to as long as 1.2 – 1.4 times the width of hind tibia at the distal end). Ventrobasal spines (2) present on fore and mid femora and absent on hind femur (both legs: one strong, one weaker). Subapical (distal) spines on first four tibiae present (short). Hind metatarsus slightly flattened. Pulvilli as long as the last tarsal segment, shorter on metatarsi. Wing. Length: 2.4 – 2.5 mm. Upper side of basal costal cell with one dark and short bristles. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m at middle of discal cell. 4 – 5 distinct brown setulae on tegula, mostly along the edges. Abdomen. Viewed obliquely from front tergites subshining black, tergite 1 silvery grey, sides silvery pollinose on tergites 4 and 5 (spot on 5 larger). Dispersed strong dark bristles on all tergites absent, only occasional indistinct bristles present. Lateral bristles on first tergite present, 4 – 5 short pale bristles in a row up to 0.5 – 0.7 times as long as the width of hind femur at distal tip in lateral view. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 as long as ST 8. Genitalia without dissection: ST 8 medium sized, brownish; membranous area elongated, broadening on dorsal part; epandrium brown, surstyli yellow, slim. Genitalia. Genital capsule in dorsal view: epandrium and surstyli brown, gray pollinose. Epandrium wider than long (MLE: MWE = 0.8). Surstyli rather symmetrical, broadened at base, elongated and slender, pointing towards each other at apex, left is wider than right one (Fig. 27 A). Genital capsule in ventral view: gonopods more elongated towards surstyli (reach cerci), equal in height, hypandrium broad, subepandrial sclerite rectangular-shaped covered with transverse streaks (Fig. 27 B), phallic guide with four spines dorsolaterally (one left spines longer than others; the longest one twice as length as others). Genital capsule in lateral view: both surstyli rather straight, left apically bent towards sternite (Figs 27 C – D). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF90EC2AD8D9EA77FC42142C.taxon	etymology	Etymology: This species is named after the only place that specimens have been collected, Culgoa National Park. Examined material: HOLOTYPE: AUSTRALIA: New South Wales: Culgoa National Park, 8 km West-Northwest Cawwell Homestead, Diemunga Lagoon (CGN 2 M), 29 ° 3 ’ S, 146 ° 60 ’ E, Coolibah, 30. i – 18. v. 2010, C. Lambkin, R. Olsen & B. Shieban, Malaise trap, CNC 575143 (1 ♂, QM); PARATYPES: AUSTRALIA: New South Wales: Culgoa National Park, 8 km West-Northwest Cawwell Homestead, Diemunga Lagoon (CGN 2 M), 29 ° 3 ’ S, 146 ° 60 ’ E, Coolibah, 30. i – 18. v. 2010, C. Lambkin, R. Olsen & B. Shieban, Malaise trap, CNC 575148, CNC 575156 (1 ♂, QM, 1 ♂ CNC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF90EC2AD8D9EA77FC42142C.taxon	distribution	Distribution: Australia (New South Wales) (Fig. 116). Notes: Culgoa National Park has a rich diversity of Tomosvaryella with 20 species known from only 68 specimens. Tomosvaryella culgoaensis is currently endemic to the park and named to raise awareness of this remote borderland park that includes part of Queensland and New South Wales.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF96EC28D8D9EBCBFB36134C.taxon	description	Figs 28 A – E, 117	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF96EC28D8D9EBCBFB36134C.taxon	diagnosis	Diagnosis: This species can be recognized by the two patches of spines on the hind trochanter; surstyli elongated in dorsal view (Fig. 28 A); phallic guide with some different-sized spines; both gonopods with a finger-like projection in ventral view (Fig. 28 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF96EC28D8D9EBCBFB36134C.taxon	description	Description: MALE: Body length: 3.5 mm. Head. Flagellum acuminate, brown. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose; eyes touching for distance equal to 2 – 2.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum dark brown, with 3 – 4 distinct pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part silver pollinose, slightly grayish also from the side. Scutellum brown pollinose and with 6 – 8 weak bristles along the distal edge. Dorsocentral bristles developed, dark, longer on frontal part and numerous dark bristles around postpronotum. Halter, knob brown, stem black. Legs. Mid coxa with two long dark bristles. Trochanters and femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees yellow-brown, tibiae black, tarsal segments dark brown, only yellowish ventrally. Hind trochanter with a few short dark spines (5 – 7) in a group at the base ventrally (“ echidna ” - like), at distal end with 1 strong and 2 – 3 weaker bristles in a small spot (appears to be one needle). Ventroapical row of spines on fore femur missing, 4 – 6 very short, black spines on mid femur; no spines on hind femur, only 6 – 8 pale bristles posteriorly on distal half (as long as the half the width of hind tibia at the distal end); fore and mid femora with two dark bristles ventrobasally; hind femur bent in a slight arch when observed from above; hind tibia slightly bent at middle in posterior view. Hind metatarsus flattened, slightly longer than 2 – 4 combined covered by semi-erect bristles dorsally. Pulvilli as long as last tarsal segment. Wing. Length: 4.0 mm. Upper side of basal costal cell with one dark and long bristles. Fourth costal section 4 – 5 times as long as third costal section, third section appears to be unusually short. Cross-vein r-m distal to middle of discal cell. 5 – 7 dark setulae on tegula. Abdomen. Dissected: tergite 1 silvery grey, Lateral bristles on first tergite present, 6 – 8 brown bristles in a row up to as long as 1.5 – 1.8 times the width of hind femur at distal tip in lateral view. Postabdomen in dorsal view: Dissected. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.5). Surstyli symmetrical, elongated with a broad base, left slightly longer than right one (Fig. 28 A). Genital capsule in ventral view: subepandrial sclerite covered with some transverse streaks except apical third close to surstyli; both gonopods with a finger-like projection extended towards surstyli, a small protrusion presented on these finger-like projections extended towards phallus, left gonopod longer than right one (Fig. 28 B), phallic guide straight, with 5 – 6 different-sized spines dorsolaterally before apex (longer ones as wide as left surstylus); phallus with three short ejaculatory ducts. Genital capsule in lateral view: both surstyli curved towards sternite, with a small projection in basal towards hypandrium (Figs 28 D – E); phallic guide with a dorsally hooked apex (turned in the photo) (Fig. 28 D – E); ejaculatory apodeme flat, sperm pump elongated, linear (Fig. 28 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF96EC28D8D9EBCBFB36134C.taxon	etymology	Etymology: This species is named after the finger-like projections on the gonopods (from the Greek daktylos for finger). Examined material: HOLOTYPE: AUSTRALIA: South Australia: Great Vict [oria] Desert, vehicle net bet [ween] 145 – 165 km N [orth] Cook, 27 ° 52 ’ S, 130 ° 22 ’ E, 30. viii. 1980, J. Forrest, JSS 8703 (1 ♂, SAM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF96EC28D8D9EBCBFB36134C.taxon	distribution	Distribution: Australia (South Australia), Fig (117). Notes: Based on the genitalia characters, this species is similar to Tomosvaryella mathiesoni sp. nov., where the surstyli in dorsal view lack small triangular projections at base along outer margin (Fig. 61 A).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF94EC26D8D9ECEBFBBE1360.taxon	description	Figs 29 A – E, 119, 144 A	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF94EC26D8D9ECEBFBBE1360.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having some ventrobasal bristles and some shorter bristles in a row towards the apical margin; hind femur with 8 – 10 long bristles posteroventrally; surstyli elongated, with a patch of distinct bristles in its middle (Fig. 29 A); both gonopods extended towards surstyli, pointed at apex in ventral view (Fig. 29 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF94EC26D8D9ECEBFBBE1360.taxon	description	Description: MALE: Body length: 2.9 – 3.1 mm. Head. Pedicel with two short dark upper bristles and one lower bristle. Flagellum yellow-brown, long tapering. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 times the length of ocellar triangle. Vertex dark, lacking pollinosity. Occiput silvery pollinose with scattered sort dark bristles. Thorax. Postpronotum pale, with two long and one short dark bristles. Mesonotum (viewed obliquely from front) silvery brown pollinose, posterior part more brown. Scutellum silvery brown pollinose and with 5 – 8 bristles along distal edge. Halter, knob pale, tip brownish, stem dark. Legs. Mid coxa with three long and 3 – 5 short dark bristles. Trochanters and femora black; knees and basal 1 / 6 of tibiae and tarsomeres yellow. Hind trochanter with some longer black ventrobasal bristles and some shorter bristles in a row towards apical margin; ventrobasal short spines present on all femora. Fore and mid femora with two distinct bristles ventrobasally. Hind femur without ventroapical spines, except 8 – 10 equally spaced sort bristles posteroventrally. Subapical (distal) spines on first four tibiae present (short). Hind metatarsus flattened, almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 3.1 mm. Upper side of basal costal cell with one long dark bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 3 – 4 long dark setulae on tegula. Abdomen. Viewed obliquely from front tergites black, all tergites silvery grey pollinose covered with short brown bristles. Lateral bristles on first tergite present, 5 – 7 dark bristles up to as long as ¾ of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.5). Surstyli elongated, both surstyli with a patch of distinct bristles in the middle (Fig. 29 A), left surstylus broadened in the middle. Genital capsule in ventral view: both gonopods extended towards surstyli, pointed at apex; phallic guide with 3 – 4 short dorso- dorsolateral spines, one is longer than others; subepandrial sclerite short, with a fin-shaped projection in its center (Fig. 29 B); Genital capsule in lateral view: both surstyli curved towards sternite (Fig. 29 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 29 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF94EC26D8D9ECEBFBBE1360.taxon	etymology	Etymology: This species in named in honour of Greg Daniels. Greg is one of the best collectors in Australia and has contributed massively to our understanding of the continental fauna. He contributed 56 specimens and 22 species to this revision (Supplementary file 2). Examined material: HOLOTYPE: AUSTRALIA: Queensland: Lake Broadwater n [ea] r Dalby, site B, 27 ° 21 ’ S, 151 ° 6 ’ E, feeding on blossom, Acacia spectabilis, 30. VIII. 1986, G. & A. Daniels, JSS 8924 (1 ♂, QM); PARATYPES: AUSTRALIA: Queensland: Brisbane, 27 ° 28 ’ S, 153 ° 1 ’ E, 28. III. 1952, J. Davis, JSS 8278 (1 ♂, QM); Kholo, 27 ° 29 ’ S, 152 ° 48 ’ E, 9. III. 1973, Colbo, JSS 9708 (1 ♂, QM); Charleville, 26 ° 24 ’ S, 146 ° 14 ’ E, 15. II. 1970, A. L. Dyce, car net, ex. spirit via ethyl acetate, JSS 8618 (1 ♂, ANIC); Western Australia: Karijini National Park, Juna Downs Road, 22 ° 44 ’ S, 118 ° 25 ’ E, 798 m, dry Turee Creek, grassy open Eucalyptus scrub, 14 – 19. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16114 (1 ♂, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF94EC26D8D9ECEBFBBE1360.taxon	distribution	Distribution: AUSTRALIA (Queensland, Western Australia) (Fig. 119). Notes: Although the holotype specimen notes that the specimen was feeding on an Acacia blossom, it was more likely simply sitting on it. There is no confirmed evidence of nectar feeding in Pipunculidae despite a paper to the contrary (Kehlmaier 2014). The paper documents two cases of pipunculids on flowers but does not conclusively document nectar feeding. JHS has observed Pipunculidae on flowers in a few instances and close observation confirmed that they were not feeding. Although we failed to obtain a sequence of JSS 16114 from Western Australia, its external and genitalia features are the same as the holotype and support treatment of these widely disjunct specimens as a single species. More samples are needed to test the species concept.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF9AEC24D8D9EC07FB4F12DC.taxon	description	Figs 30 A – E, 120	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF9AEC24D8D9EC07FB4F12DC.taxon	diagnosis	Diagnosis: This species can be recognized by hind trochanter with a keel covered by short erect bristles along its edge; extremely large and hairy cerci, gradually widened towards epandrium in lateral view (Fig. 30 D – E); asymmetrical surstyli in dorsal view (Fig. 30 A); phallus with some small spines emerging before the ejaculatory ducts; both surstyli curved toward sternite, covered by long bristles in lateral view (Fig. 30 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF9AEC24D8D9EC07FB4F12DC.taxon	description	Description: MALE: Body length: 4.2 – 4.5 mm. Head. Flagellum long, acuminate; brown, light brown at frontal edge, grey pollinose. Scape, pedicel dark. Upper side of pedicel with 3 – 4 short dark bristles. Face silvery pollinose. Frons, upper part shining black; eyes touching for distance equal to 1.2 – 1.4 times the length of ocellar triangle. Occiput silvery pollinose with scattered short dark bristles. Thorax. Postpronotum pale, with 3 – 4 short, pale bristles. Mesonotum (viewed obliquely from front) silvery-brown pollinose, anterior part greyer. Scutellum brownish pollinose and with 3 – 6 weak bristles along the distal edge. Dorsocentral bristles short and dark, somewhat longer along frontal edge, postalar callus with two short and dark bristles. Halter, knob and stem brown at apex, black at base. Legs. Mid coxa with 3 – 4 long, dark bristles. Trochanters, coxae and femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 5 of tibiae yellow, tarsal segments dark brown, ventrally somewhat paler, last segment same dark brown. Hind trochanter with a keel, extended from basal to apical covered with short erect bristles along its edge. Ventroapical row of 4 – 5 short spines on all femora present, all femora with 10 – 12 long bristles dorsally, 2 – 3 dark short spines present on fore and mid femora ventrobasally. Subapical (distal) spines on first four tibiae absent. Hind tarsi slightly elongated, metatarsus almost as long as 2 – 5 combined; stronger, peg-like dark spines on metatarsus laterally along anterior edge. Pulvilli as long as last tarsal segment. Wing. Length: 4.3 – 4.6 mm. Upper side of basal costal cell with two dark bristle, one is longer. Fourth costal section 3 – 3.5 times as long as third costal section (section 3 appears short). Cross-vein r-m distinctly distal to middle of discal cell. 6 – 8 distinct dark bristles on tegula. Abdomen. Viewed obliquely from front tergites black, tergite 1 silvery grey. Dispersed strong dark bristles on all tergites present. Lateral bristles on first tergite present, 18 – 20 different-sized and strong bristles. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.6). Surstyli asymmetrical, widened at base, right surstylus wider, pointed towards left one before apex at inner margin; left surstylus curved towards right side away from right one, both surstyli covered by dense short bristles in basal half (Fig. 30 A). Genital capsule in ventral view: gonopods pointed at apex, equal in height; hypandrium sclerotized, subepandrial sclerite long (Fig. 30 B); phallus with three short ejaculatory ducts. Genital capsule in lateral view: extremely large cerci, as large as surstyli covered with long dense bristles at the end; both surstyli curved towards sternite, broadened at the middle, covered with long bristles; phallus with some short spines before emerging ejaculatory ducts; phallic guide with some spines, one is longer. (Figs 30 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 30 C). FEMALE: unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF9AEC24D8D9EC07FB4F12DC.taxon	etymology	Etymology: The name is derived from the Greek dasys, hairy, shaggy, tufted or dense and proktos for anus, rectum or tail, in reference to the extremely hairy cerci. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Carnarvon National Park, Hilltop north of Tombs, 25 ° 5 ’ S, 147 ° 52 ’ E, 26. XI. 1997, J. Skevington & C. Lambkin, hand collected, JSS 2200 (1 ♂, QM); PARATYPES: AUSTRALIA: Queensland: same data as holotype, JSS 2199; JSS 2201 – 2; JSS 2204 – 5; JSS 2219 – 20; JSS 2223 (1 ♂, CNC; 6 ♂, QM; 1 ♂, USNM); Isla Gorge National Park, 25 ° 11 ’ S, 149 ° 58 ’ E, 320 m, 3. XI. 1991, G. Daniels, JSS 8904 (1 ♂, QM); South Australia: Flinder’s Ranges National Park, Brachina Gorge, Heysen Hilltop, 31 ° 20 ’ S, 138 ° 33 ’ E, 9. X. 1997, J. & A. Skevington & C. Lambkin, JSS 8378 (1 ♂, QM); Western Australia: Karijini National Park, Juna Downs Road, 22 ° 43 ’ S, 118 ° 25 ’ E, 799 m, near rock outcrop, Eucalyptus Eremophila scrub, 25. IV – 14. V. 2003, C. Lambkin, T. Weir, Malaise trap, ANIC [Bulk Sample] 2040, JSS 16046 (1 ♂, ANIC); 158 km S Newman, 9 km N Kumarina Road House, 24 ° 38 ’ S, 117 ° 37 ’ E, 638 m, wide sandy wash, 7 – 18. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15720 (1 ♂, CNC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF9AEC24D8D9EC07FB4F12DC.taxon	distribution	Distribution: Australia (Queensland, South Australia, Western Australia) (Fig. 120). Notes: This species appears to favour dry sclerophyll forest and has been found hilltopping.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF98EC22D8D9ED9BFE19150C.taxon	description	Figs 31 A – D, 121, 155 E	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF98EC22D8D9ED9BFE19150C.taxon	diagnosis	Diagnosis: This species can be recognized by triangle-shaped surstyli in dorsal view (Fig. 31 A); epandrium with a depression in its middle divided into two parts by epandrial groove in dorsal view (Fig. 31 A); membranous area approaching epandrium; broadened gonopods, phallic guide strong with one longer spine before apex and some condensed small spines in middle in lateral view (Fig. 31 C – D); phallus with a small membranous sheath close to ejaculatory ducts, one of ejaculatory ducts with a small denticulated sheath at its base (Fig. 31 C – D); subepandrial sclerite separated into two parts from its middle towards surstyli (Fig. 31 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF98EC22D8D9ED9BFE19150C.taxon	description	Description: MALE: Body length: 2.2 – 2.3 mm. Head. Flagellum long acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 – 2.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 5 – 6 black erect bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum brownish silvery pollinose and with 8 – 10 erect black bristles along the distal edge, in length up to the width of hind femur at base. Dorsocentral bristles massive, black, uniform in length, almost as long as bristles on scutellum, numerous black bristles around postpronotum present. Halter, knob pale, stem brown. Legs. Mid coxa with 2 – 3 long dark bristles. Trochanters brown, femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 5 of tibiae yellow (tibiae otherwise brown), tarsal segments brown dorsally, yellow ventrally, last segment darker. Hind trochanter gray pollinose, with 2 – 4 bristles close to hind coxa, ventrally slightly flattened half circle (completely flat dorsally); ventrobasal spines (2) present on fore and missing on mid femur. Ventroapical row of spines on fore femur absent; 4 – 5 small, black spines on mid femur; hind femur without ventroapical spines, except 14 – 16 equally spaced, equally long bristles, posteroventrally, longest up to 1 / 4 times the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present (short). Hind metatarsus not flattened, slightly longer than 2 – 4 combined and with strong dark bristles dorsally, scrub-like bristles ventrally (t 1 only). Pulvilli shorter than last tarsal segment. Wing. Length: 2.2 mm. Upper side of basal costal cell with one long dark bristle and 1 – 3 shorter bristles. Fourth costal section 2 – 2.2 times as long as third costal section. Cross-vein r-m at middle of discal cell. 3 – 4 distinct black setulae on tegula. Abdomen. Viewed obliquely from front tergites subshiny brown-black, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (spot on t 5 larger). Dispersed strong dark bristles on all tergites present, longest on tergite 5, in length up to 1 / 3 width of hind tibia at distal end. Lateral bristles on first tergite present, 4 – 6 short dark bristles up to as long as 1 / 3 width of hind femur at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1.6 – 1.8 times as long as ST 8. Genitalia without dissection: ST 8 large, rounded triangle in dorsal view, brown and with a few distinct bristles, otherwise with velvet-like coverage; membranous area with round part extended dorsally and narrow part approaching epandrium, mostly directed posteriorly; epandrium brown, surstyli yellow, broad at base. Genitalia. Genital capsule in dorsal view: epandrium slightly as long as wide (MLE: MWE = 1.07), with a depression in its middle divided into two parts by epandrial groove. Surstyli triangle-shaped, broadened in basal half, strongly narrowed in apical half (Fig. 31 A). Genital capsule in ventral view: gonopods widened, equal in height; phallic guide strong and wide, with one upward spine dorsally before apex, subepandrial sclerite separated into two parts from its middle towards surstyli, phallus with a small membranous sheath close to ejaculatory ducts, one of ejaculatory ducts with a small denticulated sheath at its base (Fig. 31 B); Genital capsule in lateral view: both surstyli broadened at middle and narrowed apically, curved towards sternite; phallic guide strong with some condensed small spines dorsally in its middle, pointed at apex (Fig. 31 C – D). FEMALE: As male except for the following characters. Frons, eyes separated, as wide as 1.5 – 1.8 times the width at antennae; completely silver-grey pollinose on lower 1 / 3, shiny black in front of ocellar triangle, smooth change in transition zone (at 1 / 3 from ocellar triangle) to shiny black. Enlarged ommatidia silvery shining. Pulvilli and claws about 1 – 3 times as long as last tarsal segment, smaller on metatarsi. Female abdomen with silver spots on sides of tergites 4 – 6. Ovipositor. straight in ventral view, down curved at distal end and robust in lateral view (yellow piercer, brown base), reaching distal end of 3 rd segment; base silvery pollinose with dispersed short dark bristles. LP: LB = 2.5. LDP: LPP = 1.8. (Fig. 155 E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF98EC22D8D9ED9BFE19150C.taxon	etymology	Etymology: After the Greek capital letter delta, shaped like a triangle, in reference to the triangle shaped surstyli. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: Karijini National Park, Juna Downs Road, 22 ° 44 ’ S, 118 ° 25 ’ E, 798 m, dry Turee Creek, grassy open Eucalyptus scrub, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16089 (1 ♂, WAM); PARATYPES: AUSTRALIA: Western Australia: Karijini National Park, Juna Downs Road, 22 ° 44 ’ S, 118 ° 25 ’ E, 798 m, dry Turee Creek, grassy open Eucalyptus scrub, 14 – 19. V. 2003; 25. IV – 14. V. 2003, C. Lambkin, T. Weir, Malaise trap, JSS 16101, JSS 16106 (1 ♂, 1 ♀, ANIC); Queensland: Cudmore National Park (CM 3 M), 22 ° 58 ’ S, 146 ° 23 ’ E, 351 m, Melaleuca heath near drying creek, 28. X. 2010 – 2. VIII. 2011, Lambkin, Starick & Bailey, Malaise trap, CNC 574703 (1 ♂, QM); Cudmore National Park (CM 1 M), 22 ° 54 ’ S, 146 ° 21 ’ E, 365 m, Eucalyptus woodland with Spinifex, 27. X. 2010 – 2. VIII. 2011, Lambkin, Starick & Bailey, Malaise trap, CNC 576550, CNC 591868 (2 ♂, CNC, QM); New South Wales: Ledknapper National Reserve, 5 km N Beulah Homestead (LDN 1 M), 28 ° 19 ’ S, 147 ° 10 ’ E, Allocasuarina, 18. III – 15. V. 2010, S. O’Sullivan & C. Lambkin, Malaise trap, CNC 576713 (1 ♂, QM); Ledknapper National Reserve, 7.3 km ENE Beulah Headquarters (LDN 2 M), 29 ° 21 ’ S, 146 ° 13 ’ E, E. melanopholia, Spinifex, 10. XII. 2009 – 18. III. 2010, S. O’Sullivan, K. Taylor, Malaise trap, CNC 576515 (1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF98EC22D8D9ED9BFE19150C.taxon	distribution	Distribution: Australia (New South Wales, Queensland, Western Australia) (Fig. 121). Notes: This is another species with a wide distribution in the Australian outback. It will be nice to fill in some of the gaps in the range over time to help understand its host, ecology and distribution in more detail. Intraspecific genetic distance is 0.0 %. This species is genetically closest to T. ngarrbekiota sp. nov. (8.3 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF9EEC20D8D9EAABFA9C1284.taxon	description	Figs 32 A – E, 117	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF9EEC20D8D9EAABFA9C1284.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a distinct edge and some short bristles; elongated surstyli in dorsal view (Fig. 32 A); phallic guide with two long spines dorsolaterally (Fig. 32 B); gonopods extended towards surstyli (Fig. 32 B); both surstyli broadened with a small hump in dorsomedial margin in lateral view (Fig. 32 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF9EEC20D8D9EAABFA9C1284.taxon	description	Description: MALE: Body length: 2.6 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 2 – 3 indistinct bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum brownish silvery pollinose and with 10 – 12 short bristles along the distal edge. Halter, knob pale, stem brown. Legs. Mid coxa with 3 – 4 long dark bristles. Trochanters brown, femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 5 of tibiae yellow (tibiae otherwise brown), tarsal segments brown dorsally, yellow ventrally, last segment darker. Hind trochanter with edge, 45 degrees to the long axis of trochanter, distal part shiny, 3 – 4 short bristles on proximal part, otherwise velvet-like coverage; ventrobasal spines absent on fore and mid femur. Ventroapical row of spines on fore femur absent; 4 – 5 small, black spines on mid femur; hind femur without ventroapical spines, except 10 – 12 equally spaced, equally long bristle, posteroventrally, longest up to 1 / 3 times the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present (short). Hind metatarsi not flattened, as long as 2 – 4 combined, scrub-like bristles ventrally (t 1 only). Pulvilli shorter than last tarsal segment. Wing. Length: 2.7 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 2 times as long as third costal section. Cross-vein r-m slightly distal to middle of discal cell. 2 – 3 short dark setulae on tegula. Abdomen. Dissected. tergite 1 silvery grey, Lateral bristles on first tergite present, 4 – 6 short dark bristles up to as long as hind femur’s width at base. Postabdomen in dorsal view: Dissected. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.2). Surstyli elongated (Fig. 32 A). Genital capsule in ventral view: both gonopods extended towards sternite, equal in height; phallic guide strong with two small lateral lobes before apex, with 3 – 4 dorso- dorsolateral spines, two spines longer than others, subepandrial sclerite distinct (Fig. 32 B); Genital capsule in lateral view: both surstyli broadened with a small hump at dorsomedial margin, smoothly curved towards sternite (Fig. 32 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 32 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF9EEC20D8D9EAABFA9C1284.taxon	etymology	Etymology: Named after the small town closest to where the only known specimen was collected. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: ~ 3 km N [orth] of Denham, 25.909 ° S, 113.531 ° E, hilltop, 12. XII. 1999, J. Skevington, JSS 7375 (1 ♂, WAM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF9EEC20D8D9EAABFA9C1284.taxon	distribution	Distribution: Australia (Western Australia) (Fig. 117). Notes: One hundred specimens of 10 species of Tomosvaryella were collected on the amazing little hilltop where the only specimen of this species was found. The site is extremely arid and only one other pipunculid genus was caught here (four specimens of Clistoabdominalis trochanteratus (Becker 1900 )). Tomosvaryella T. denhamensis sp. nov. is genetically closest to T. bulbosa sp. nov. (8.1 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF9CEC3ED8D9EC23FB451248.taxon	description	Figs. 33 A – F, 121	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF9CEC3ED8D9EC23FB451248.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter bearing a few short dark bristles in a row ventrally; surstyli elongated in dorsal view (Fig. 33 A); gonopods extended towards surstyli in ventral view (Fig. 33 – B); phallic guide with four spines dorsomedially (one is longer); both surstyli straight, curved towards sternite in apical third in lateral view (Fig. 33 D – F).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF9CEC3ED8D9EC23FB451248.taxon	description	Description: MALE: Body length: 2.6 – 3 mm. Head. Flagellum acuminate; yellow. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 3 – 4 extra-long, pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum brownish pollinose and with 10 – 12 bristles along the distal edge. Dorsocentral bristles well developed, somewhat longer along frontal edge, numerous long pale bristles around postpronotum. Halter, knob pale, stem brown. Legs. Mid coxa with 2 – 3 long bristles. Trochanters and femora brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees, basal 1 / 4 and distal extremity of tibiae yellow (tibiae otherwise brown), tarsomeres yellow ventrally and brown dorsally, last segment same, brown. Hind trochanter without a keel, but with a few short dark bristles (7 – 8) in a row ventrally, equally spaced. Ventrobasal spines (2) present on fore and absent on mid femur. Ventroapical row of spines on fore femur absent; 5 – 6 short, black spines on mid femur; no spines on hind femur, only 22 – 24 long dark bristles equally spaced, longest at 2 / 3 from base (up to as long as 1.5 – 1.8 times the width of hind tibia at the distal end). Subapical (distal) spines on first four tibiae present (very short). Hind tarsi flattened (1 – 3), metatarsus almost as long as 2 – 3 combined. Pulvilli slightly shorter than the last tarsal segment. Wing. Length: 3.0 – 3.1 mm. Upper side of basal costal cell with one short dark bristle. Fourth costal section 2 times as long as third costal section. Cross-vein r-m distal to middle of discal cell. 3 – 4 distinct brown setulae on tegula, mostly along the edges. Abdomen. Viewed obliquely from front tergites subshining black, tergite 1 silvery grey, sides silvery pollinose on tergites 4 and 5 (both very small). Dispersed strong dark bristles on all tergites, longest on tergite 5, up to 1.2 – 1.4 times as long as the width of hind tibia at base. Lateral bristles on first tergite present, 4 – 5 short pale bristles in a row up to 1 – 1.2 times as long as the width of hind femur at distal tip in lateral view. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1 – 1.2 times as long as ST 8. Genitalia without dissection: ST 8 large and wide, brown and with the occasional abdominal bristles; membranous area elongated, slim, directed posteroventrally; epandrium brown, surstyli yellow, elongated. Genitalia. Genital capsule in dorsal view: epandrium and surstyli brown. Epandrium longer than wide (MLE: MWE = 1.1). Surstyli rather symmetrical, broadened at base, elongated, right surstylus pointed towards left one at apex, left is wider and longer than right one (Fig. 33 A). Genital capsule in ventral view: gonopods equal in height, more extended towards surstyli and surrounding outer margin of subepandrial sclerite; hypandrium broad, subepandrial sclerite widened and covered by scattered short bristles in its upper half close to surstyli (Fig. 33 B); phallus with three short ejaculatory ducts; phallic guide with four spines dorso- dorsolaterally (the longest one twice as length as others). Genital capsule in lateral view: both surstyli widened at base, straight, curved towards sternite in apical third (Figs 33 D – F). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF9CEC3ED8D9EC23FB451248.taxon	etymology	Etymology: From the Latin dirigo, set straight or arrange in a straight line, in reference to the straight surstyli of the males. Examined material: HOLOTYPE: AUSTRALIA: Australian Capital Territory: Black M [oun] t [ai] n, 35 ° 16 ’ S, 149 ° 6 ’ E, xii. 1987, M. Irwin, Malaise trap, JSS 8531 (1 ♂, ANIC); PARATYPES: AUSTRALIA: South Australia: Wilpena Pound, Flinders Range, 31 ° 25 ’ S, 138 ° 45 ’ E, 1. x. 1975, Z. Liepa, JSS 9163 – 4 (1 ♂, ANIC; 1 ♂ CNC); Gammon Ranges National Park, south end of Weetootla Gorge, 30 ° 29 ’ S, 139 ° 16 ’ E, 537 m, hilltop, 11. X. 1997, J. Skevington & C. Lambkin, JSS 8355 (1 ♂, QM); New South Wales: 20 km E of Forbes, 33 ° 23 ’ S, 148 ° 1 ’ E, in Callitris forest, 9. IV. 1977, Z. Liepa, JSS 9179 (1 ♂, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF9CEC3ED8D9EC23FB451248.taxon	distribution	Distribution: Australia (Australian Capital Territory, New South Wales, South Australia) (Fig. 121). Notes: This is a southeastern Australian species. One specimen was captured hilltopping. This species is genetically closest to T. macrostyla sp. nov. (4.4 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF82EC3CD8D9EDEFFC561520.taxon	description	Figs 34 A – E, 122, 147 F	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF82EC3CD8D9EDEFFC561520.taxon	diagnosis	Diagnosis: This species can be recognized by having two differently shaped spur-like projections on the hind trochanter (Fig. 147 F); surstyli quadratic-shaped at base with rather straight apical margin in dorsal view (Fig. 34 A); gonopods unequal in ventral view (Fig. 34 B); phallic guide with 5 – 6 different-sized lateral spines (Fig. 34 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF82EC3CD8D9EDEFFC561520.taxon	description	Description: MALE: Body length: 2.7 – 2.9 mm. Head. Flagellum acuminate; yellow. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose; eyes touching for distance equal to 2 – 2.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 3 – 4 short, pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum brownish pollinose and with 8 – 10 weak bristles along the distal edge. Dorsocentral bristles distinct, short and dark, somewhat longer along frontal edge, numerous pale bristles around postpronotum. Halter, knob pale, stem brown. Legs. Mid coxa with 2 – 3 long dark bristles. Trochanters and femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees yellow (tibiae otherwise black), tarsal segments yellow-brown, ventrally somewhat paler, last segment same dark brown. Hind trochanter with double spurs (pegs), anterior is thick, bent medially, posterior one is thin, very pointy (Fig. 147 F). Ventroapical row of spines on fore femur absent; 4 – 5 small, black spines on mid femur; hind femur without ventroapical spines, except 22 – 24 equally space bristles posteroventrally, longest up to 2 / 3 the width of hind tibia at distal end. Subapical (distal) spines (2 – 3) on first four tibiae present (on mid femur longer). Hind metatarsus and second tarsomere flattened, metatarsus as long as 2 – 3 combined. Pulvilli as long as last tarsal segment. Wing. Length: 3.0 – 3.1 mm. Upper side of basal costal cell with one dark bristle. Fourth costal section 2 – 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2 – 3 distinct brown setulae on tegula. Abdomen. Viewed obliquely from front tergites subshining black, tergite 1 silvery grey, sides silvery pollinose on tergites 4 and 5 (larger on tergite 5). Dispersed strong dark bristles on all tergites present, longest on tergite 5 up to 1.5 times the width of hind tibia at distal end. Lateral spines on first tergite present, 5 – 6 dark bristles up to 1.5 times as long as hind femur’s width at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1 – 1.2 times as long as ST 8. Genitalia without dissection: ST 8 medium sized triangle shaped in dorsal view, brown and with the occasional indistinct bristles, otherwise velvet-like coverage; membranous area elongated, widening dorsally; epandrium brownish at tip, surstyli yellow and bent towards abdomen, appear to be a yellow triangle in lateral view (the corner is rounded). Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.1). Surstyli rather symmetrical, quadratic-shaped in basal third and rather straight in apical two thirds, its medial margin covered by dense bristles, pointing towards each other at apex; left surstylus wider than left one (Fig. 34 A). Genital capsule in ventral view: gonopods unequal in height, left is slightly longer than right one, subepandrial sclerite long (Fig. 34 B). Genital capsule in lateral view: both surstyli broadened in middle and curved towards sternite, right surstylus constricted in dorsomedial margin; phallic guide pointed at apex, with 5 – 6 lateral spines (the longest one twice as length as others), with three distinct short ejaculatory ducts (Figs 34 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 34 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF82EC3CD8D9EDEFFC561520.taxon	etymology	Etymology: From the Latin dissidens, differing or disagreeing, in reference to the two differently shaped spur-like projections on the hind trochanter. Examined material: HOLOTYPE: AUSTRALIA: South Australia: Kangaroo Is [land], Flinders Chase N [ational] P [ark], 36 ° 1 ’ S, 136 ° 44 ’ E, 28. x. 1993, D. K. Yeates, Malaise trap, JSS 8661 (1 ♂, ANIC); PARATYPES: AUSTRALIA: New South Wales: Mount Boppy near Cobar, 31 ° 33 ’ S, 146 ° 17 ’ E, hilltop, 16. i. 1999, C. Lambkin, N. Power & P. Bouchard, 10: 00 AM, JSS 8903 (1 ♂, QM); Queensland: Brisbane, Mount Coot-tha, 27 ° 29 ’ S, 152 ° 57 ’ E, 170 m, hilltop, 15. v. 1999, J. & A. Skevington, hand collected, JSS 5779; JSS 15387 (2 ♂, QM, CNC); 17. vii. 1999, J. & A. Skevington, hand collected, 12: 45 to 2: 00 pm, JSS 6469 – 70 (2 ♂, QM); 19. iv. 1998, J. Skevington, hand collected, 11: 45 – 12: 15 pm, JSS 3871, JSS 3876 (2 ♂, QM); 19. vi. 1999, J. & A. Skevington, hand collected, 11: 15 am to 1: 00 pm, JSS 6474 (1 ♂, QM); 7. iv. 1997, C. Lambkin, JSS 8305 (1 ♂, QM); Carnarvon National Park, hilltop north of Tombs, 25 ° 5 ’ S, 147 ° 52 ’ E, 26. xi. 1997, J. Skevington & C. Lambkin, hand collected, JSS 2197 – 8, JSS 2206 – 10, JSS 2213 – 4, JSS 2216, JSS 2218, JSS 2221 – 2 (13 ♂, QM); Cooloola National Park, East Mullen hilltop, 26 ° 0 ’ S, 152 ° 59 ’ E, dry sclerophyll forest, 15. xi. 2014, J. H., A. M. and A. W. Skevington, CNC 374120 (1 ♂, CNC); Culgoa Floodplain National Park, 7 km Northwest of Toulby Gate (TOLB 7 M), 28 ° 57 ’ S, 146 ° 53 ’ E, 138 m, reedy swamp, 20. i – 19. iii. 2010, C. Kelly, A. Coward, Malaise trap, 19269, CNC 575361, CNC 575367, CNC 575377 (3 ♂, QM); Kroombit Tops National Park, hilltop, 24 ° 27 ’ N, 150 ° 56 ’ E, 21. xii. 2014, J. H., A. M. & A. W. Skevington, CNC 384417 (1 ♂, CNC); Mount Moffatt National Park, Mount Moffatt Section, Summit of Fly Hill, 24 ° 58 ’ S, 147 ° 60 ’ E, 17. xi. 2014, J. H., A. M. & A. W. Skevington, CNC 374169 – 72, CNC 374780 – 2 (7 ♂, CNC); South Australia: Flinder’s Ranges National Park, Brachina Gorge, Heysen Hilltop, 31 ° 20 ’ S, 138 ° 33 ’ E, 9. x. 1997, J. & A. Skevington & C. Lambkin, JSS 8375, JSS 8377, JSS 8382 (3 ♂, QM); Victoria: Wyperfeld NP, East Lookout, 35 ° 36 ’ S, 142 ° 7 ’ E, 22. x. 2014, J. H., A. W., & A. M. Skevington, A. D. Young, A. Wyatt, & S. Namek, CNC 373404, CNC 373408, CNC 373439, CNC 373457 (4 ♂, CNC); Hattah-Kulkyne NP, Warepil Lookout, 34 ° 43 ’ S, 142 ° 18 ’ E, 18. x. 2014, J. H., A. W., & A. M. Skevington, A. D. Young, A. Wyatt, & S. Namek, CNC 373060, CNC 373171, CNC 373178 (3 ♂, CNC); Western Australia: Karijini National Park, Weano Gorge Road, 22 ° 22 ’ S, 118 ° 15 ’ E; 22 ° 24 ’ S, 118 ° 15 ’ E, 775 m, 718 m, on hilltop, in open Eucalyptus, 25. iv – 14. v. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16141, JSS 16146, JSS 16230 – 1 (4 ♂, ANIC); Yanchep National Park, 31 ° 35 ’ S, 115 ° 42 ’ E, 18 – 29. xii. 1999, J. & A. Skevington, C. Lambkin & P. Bouchard, Malaise trap, JSS 7700 (1 ♂, QM); ~ 3 km north of Denham, 25 ° 55 ’ S, 113 ° 32 ’ E, hilltop, 12. xii. 1999, J. Skevington, JSS 7337, JSS 7345, JSS 7353, JSS 7371, JSS 7381 (5 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF82EC3CD8D9EDEFFC561520.taxon	distribution	Distribution: Australia (New South Wales, Queensland, South Australia, Victoria) (Fig. 122). Notes: This species is widespread throughout the southern half of Australia and is easily surveyed by searching on hilltops. Intraspecific genetic distance ranges from 0.2 % to 1.5 %. This species is genetically closest to T. boomerang sp. nov. (0 – 1.3 % pairwise divergence) (Supplementary file 3). As these two species differ only by the shape of surstyli in lateral view and having two differently shaped spur-like projections on the hind trochanter in T. dissidens sp. nov., it is likely that these two species are recently-diverged.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF80EC3AD8D9EAC7FCB4126C.taxon	description	Figs 35 A – E, 118	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF80EC3AD8D9EAC7FCB4126C.taxon	diagnosis	Diagnosis: This species can be recognized by the elongated surstyli, strongly narrowed at the apical half, their tips extended outwards in dorsal view (Fig. 35 A); gonopods triangle-shaped; one ejaculatory duct covered by some teeth in basal half (Fig. 35 D – E); ejaculatory apodeme funnel-shaped (Fig. 35 C).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF80EC3AD8D9EAC7FCB4126C.taxon	description	Description: MALE: Body length: 3.5 mm. Head. Scape, pedicel, arista dark, Flagellum acuminate; yellow-brown, grey pollinose, arista with four short dark upper bristles and one lower bristle. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.2 times the length of ocellar triangle. Occiput silvery pollinose with scattered short dark bristles. Thorax. Postpronotum pale, with 3 – 4 short pale bristles. Mesonotum (viewed obliquely from front) silvery brown (golden) pollinose. Dorsocentral and supra-alar bristles short, dark, Scutellum silvery brown pollinose with 4 – 6 bristles along distal edge. Halter, knob pale, stem light brown with 2 – 3 short dark bristles dorsally. Legs. Mid coxa with four long and 2 short dark bristles. Trochanters brown, femora black, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments light brown, last tarsomeres brown. Hind trochanter without distinct feature, gray pollinose; ventrobasal bristles present (5 – 8) on fore and mid femur. Mid femur with two rows of short bristles posteroventrally and posterodorsally. Hind femur with 2 – 3 ventroapical spines. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres not flattened, hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 3.5 mm. Upper side of basal costal cell with 2 – 3 long dark brown bristles. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2 – 3 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brown-black (golden), tergite 1 silvery grey. Lateral bristles on first tergite present, 4 – 10 short dark bristles up to as long as one quarter of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.1). Surstyli elongated, rather widened in basal half, strongly narrowed at the apical half, their tips extended outwards, left surstylus is longer than right one (Fig. 35 A). Genital capsule in ventral view: gonopods equal in height, triangle-shaped; phallic guide without spines; subepandrial sclerite short, one of ejaculatory ducts with some teeth in basal half (Fig. 35 B); Genital capsule in lateral view: both surstyli curved towards sternite (Fig. 35 D – E). Ejaculatory apodeme funnel shaped (Fig. 35 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF80EC3AD8D9EAC7FCB4126C.taxon	etymology	Etymology: From the Latin divaricatus, spread apart, in reference to the divergent tips of the surstyli. Examined material: HOLOTYPE: AUSTRALIA: Northern Territory: near Darwin, 1 km E Manton Dam, 12 ° 53 ’ S, 131 ° 8 ’ E, grass along stream near road, 3. IX. 2004, J. Forrest & T. Wheeler, sweep net, LEM _ 0016204 (1 ♂, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF80EC3AD8D9EAC7FCB4126C.taxon	distribution	Distribution: Australia (Northern Territory) (Fig. 118).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF86EC38D8D9ED0BFAE4150C.taxon	description	Figs 36 A – E, 123	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF86EC38D8D9ED0BFAE4150C.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a distinct keel covered with spines on the edge; distal half of hind femur with long bristles; elongated surstyli with an extended lobe in middle in dorsal view (Fig. 36 A); elongated gonopods with two small triangular lobes on inner margin (Fig. 36 B); phallic guide with 4 – 6 dorso- dorsolateral spines (Fig. 36 B, D).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF86EC38D8D9ED0BFAE4150C.taxon	description	Description: MALE: Body length: 3.3 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 – 2.2 times the length of ocellar triangle. Occiput silvery pollinose, slightly less so on upper 1 / 3. Thorax. Postpronotum pale, with 5 – 6 pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum brown pollinose and with 8 – 10 bristles along the distal edge. Dorsocentral bristles distinct, dark, somewhat longer along frontal edge, numerous long bristles around postpronotum present. Halter, knob pale, stem brown. Legs. Mid coxa with 2 – 3 strong, dark bristles. Trochanters and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 5 of tibiae yellow-brown (tibiae otherwise dark brown), tarsal segments yellow-brown, ventrally paler, last segment darker. Hind trochanter with a distinct keel, short spines on the edge; ventrobasal spines present on fore and absent on mid femur. Ventroapical row of spines on fore femur 3 – 4; 5 – 6 short dark spines on mid femur; hind femur without ventroapical spines, except 10 – 12 long bristles posteroventrally, bent towards distal tip of femur (sometimes adpressed), equally spaced on distal half, longest up to 2 times the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present (short). Hind tibia bent in posterior view. Hind metatarsus flattened (especially distal half), metatarsi almost as long as 2 – 5 combined, with scrub-like bristles along posterior edge. Pulvilli shorter than last tarsal segment. Wing. Length: 3.2 mm. Upper side of basal costal cell with one long dark bristle. Fourth costal section 2 times as long as third costal section. Cross-vein r-m at middle of discal cell. 5 – 6 distinct brown setulae on tegula. Abdomen. Viewed obliquely from front tergites brown pollinose, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (larger on tergite 5). Dispersed short dark bristles on all tergites present, longest on tergite 5 up to 1 / 3 of the width of hind tibia at distal end. Lateral bristles on first tergite present, 4 – 5 dark bristles up to as long as hind femur’s width at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 0.9 – 1.1 times as long as ST 8. Genitalia without dissection: ST 8 medium sized, round in dorsal view, pale brown and with some distinct bristles, otherwise velvet-like coverage; membranous area round dorsally, narrow and approaching epandrium ventrally, posteriorly directed; epandrium brown, surstyli yellow, long. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.5). Both surstyli elongated, broadened in basal third, narrowed in middle, bent apically to each other; both surstyli with extended lobe in middle (Fig. 36 A). Genital capsule in ventral view: gonopods elongated towards surstyli with two small triangular lobes on inner margin, equal in height, subepandrial sclerite distinct and long; phallic guide with 4 – 6 dorso-dorsolateral spines, one is longer than others (Fig. 36 B); Genital capsule in lateral view: both surstyli curved towards sternite, right one is wider than left one (Fig. 36 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 36 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF86EC38D8D9ED0BFAE4150C.taxon	etymology	Etymology: The name is derived from the Greek word dolichos, long and lobos, lobe, in reference to the extended lobe in the middle of the surstylus. Examined material: HOLOTYPE: AUSTRALIA: Tasmania: Freycinet National Park, 42 ° 8 ’ S, 148 ° 19 ’ E, in heath with scattered eucalypts, 26. XII. 1998 – 10. I. 1999, J. & A. Skevington, Malaise trap, JSS 4665 (1 ♂, QM); PARATYPES: AUSTRALIA: Queensland: Carnarvon National Park, Mount Moffatt Summit, 25 ° 4 ’ S, 148 ° 3 ’ E, 1097 m, hilltop, 2. XII. 1997, J. Skevington & C. Lambkin, hand collected, JSS 1824 (1 ♂, QM); 18. XI. 2014, J. H., A. M. & A. W. Skevington, CNC 374959, CNC 374980, CNC 374995, CNC 375003 (4 ♂, CNC); Culgoa Floodplain National Park, 11.8 km N Headquarters (CG 2 AM), 28 ° 54 ’ S, 147 ° 8 ’ E, 151 m, Brigalow, 20. I – 18. III. 2010, C. Kelly & A. Coward, Malaise trap, CNC 576680, CNC 576683 (2 ♂, QM); Hilltop ~ 21 km South of Coen, 14 ° 8 ’ S, 143 ° 14 ’ E, 945 m, Eucalyptus forest, 2. XII. 2014, J. H., A. M. & A. W. Skevington, CNC 373639 (1 ♂, CNC); Western Australia: Lochada, 5.5 km Southeast of Bolada Camp, 29 ° 14 ’ S, 116 ° 33 ’ E, 293 m, Callitris Acacia woodland, 18 – 24. IX. 2009, Lambkin & Monteith, Malaise trap, CNC 575166 (1 ♂, QM); Heinsmen Rock, Mt Ragged Track, 30 ° 7 ’ S, 123 ° 28 ’ E, 145 m, woodland near drying pools, 30. X – 18. XI. 2003, C. Lambkin & J. Recsei, Malaise trap, JSS 16244 (1 ♂, ANIC); New South Wales: Culgoa National Park, 13.5 km SW Cawwell Homestead (CGN 1 M), 29 ° 7 ’ S, 146 ° 56 ’ E, Callitrus, 21. XII. 2009 – 30. I. 2010, B. Schiebaan, Malaise trap, 19314, CNC 576559 – 60, CNC 576562, CNC 576566, CNC 576569, CNC 576573 – 4, CNC 576583 – 4, CNC 576586 – 7, CNC 576589 (12 ♂, QM); Culgoa National Park, 13.5 km SW Cawwell Homestead (CGN 1 M), 29 ° 7 ’ S, 146 ° 56 ’ E, Blackbox, 22. XI – 21. XII. 2009, C. Lambkin, B. Shiebaan, N. Starick, Malaise trap, 11 DB, 19300, CNC 576659 (1 ♂, QM); Blackbox, 22. XI – 21. XII. 2009, C. Lambkin, B. Shiebaan & N. Starick, Malaise trap, 11 DB, 19300, CNC 576654, CNC 576656, CNC 576660 – 3 (6 ♂, QM); Callitris, 30. I – 18. V. 2010, QM Team, B. Shiebaan, R. Ohlsen, Malaise trap, 19325, CNC 576599, CNC 576603, CNC 576607, CNC 576611, CNC 576613, CNC 576616, CNC 576622, CNC 576627, CNC 576632, CNC 576642 – 3 (11 ♂, QM); Culgoa National Park, 8 km West-Northwest Cawwell Homestead, Diemunga Lagoon (CGN 2 M), 29 ° 3 ’ S, 146 ° 60 ’ E, Coolibah, 30. I – 18. V. 2010, C. Lambkin, R. Olsen & B. Shieban, Malaise trap, 19322, CNC 575121, CNC 575124, CNC 575131 (3 ♂, QM); 29 ° 3 ’ S, 146 ° 1 ’ E, 136 m, Coolibah, 21. XII. 2009 – 30. I. 2010, B. Schiebaan, Malaise trap, 19312, CNC 574752 (1 ♂, CNC); Queensland: Culgoa Floodplain National Park Headquarters, 11.8 km N, 28 ° 55 ’ S, 147 ° 8 ’ E, 150 m, Brigalow, 9. XII. 2007 – 16. I. 2008, N. Starick, C. Lambkin & R. Raven, Malaise trap, 15683, CNC 591839 (1 ♂, QM); Culgoa Floodplain National Park, 10 km NE Toulby Gate (CG 3), 28 ° 56 ’ S, 147 ° 0 ’ E, 140 m, Brigalow, 16. IX – 1. X. 2008, Simpson, Coward, Malaise trap, 4 DB, 17220, CNC 576529 (1 ♂, QM); Culgoa Floodplain National Park, 11.8 km N Headquarters (CG 2 AM), 28 ° 54 ’ S, 147 ° 8 ’ E, 151 m, Brigalow, 20. I – 18. III. 2010, C. Kelly & A. Coward, Malaise trap, 19271, CNC 576674, CNC 576693 (2 ♂, QM); Lonesome National Park, near Lookout (LNP 4 M), 25 ° 30 ’ S, 148 ° 49 ’ E, 585 m, closed Eucalyptus woodland on rocky ridge, 26. XI. 2010 – 11. I. 2011, D. Beard & B. Sigley, Malaise trap, 19382, CNC 575014 (1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF86EC38D8D9ED0BFAE4150C.taxon	distribution	Distribution: Australia (New South Wales; Queensland, Tasmania, Western Australia) (Fig. 123). Notes: This widespread species has been collected in a variety of habitats (Acacia, Callitris, Eucalyptus) and also collected hilltopping. The distribution is suggestive that this is a species complex of as many as four species. We were unable to obtain DNA barcodes from populations in the different locations. Although the surstyli and the inner lobe of New South Wales specimens are longer, we have elected to treat this as a single species for now. Based on the terminalia characters and features of the hind trochanter, this species is similar to T. patula sp. nov. Both species have identical surstyli in dorsal and ventral view and similar keel on hind trochanter. Tomosvaryella dolicholoba sp. nov. differs by having long bristles on the hind femur. This species is genetically close to T. unda sp. nov. and T. patula sp. nov. differing by 2.6 – 5.0 % and 3.1 – 4.0 % pairwise divergence, respectively (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF84EC36D8D9EAABFE9210A0.taxon	description	Figs 37 A – E, 124, 156 C	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF84EC36D8D9EAABFE9210A0.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of the surstyli, in dorsal view the right surstylus is wider and the left one is longer (Fig. 37 A); widened gonopods, pointed hypandrium in lower margin in ventral view (Fig. 37 B); phallic guide short and broad (Fig. 37 D – E); one of ejaculatory ducts with tooth-like projection in basal third in lateral view (Fig. 37 D – E); ejaculatory apodeme funnel-shaped (Fig. 37 C).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF84EC36D8D9EAABFE9210A0.taxon	description	Redescription: MALE: Body length: 3.2 – 3.4 mm. Head. Scape, pedicel and arista brown. Pedicel with 2 – 3 upper bristles and 1 – 2 lower bristles; flagellum brown, gray pollinose and tapering. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance 1.3 times the length of ocellar triangle. Occiput grey pollinose covered by scattered short brown bristles. Thorax. Postpronotum light yellow, with 1 – 3 short pale bristles in upper margin. Mesonotum (viewed obliquely from front) brown pollinose; scutum with scattered dark bristles at anterior supra-alar area, extended to posterior and uniseriate rows of intra-alar short bristles. Postalar callus dark brown with 1 – 2 short light brown bristles in the middle. Scutellum silvery pollinose and with 8 – 10 short dark bristles along the distal edge and a few scattered bristles in its middle. Halter, knob pale, stem light brown, base dark. Legs. Mid coxa with 1 – 2 different-sized dark brown bristles. Coxae, trochanters, femora dark, hind femur shining ventrally, knees and basal 1 / 3 of tibiae yellow (tibiae otherwise dark brown), tarsomeres brown ventrally, dark dorsally. Hind trochanter silvery pollinose covered by some scattered short pale bristles ventrally, its apical third shining; two ventrobasal spines absent on fore mid femur. All femora with two rows tiny spines ventroapically. Hind tibia with two rows of short brown bristle on anterior and posterior side, all tibiae gray pollinose in apical half of both anterior and posterior sides. Hind tarsomeres slightly flattened to one side, covered by short dark brown bristles, hind metatarsus as long as 2 – 4 combined; pulvilli shorter than last tarsal segment, last tarsomeres with longer dark bristles. Wing. Length: 3.2 – 3.3 mm. Upper side of basal costal cell with 1 – 2 long dark brown bristles. Fourth costal section 1.5 – 2 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2 – 3 short dark setulae on tegula. Abdomen. Tergites 1 silvery pollinose with 5 – 7 different-sized lateral bristles; tergite 2 – 5 brown pollinose, covered by scattered short dark bristles. Syntergosternite 8 enlarged, dark brown and grey-brown pollinose. Membranous area ovate. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.8). Surstyli different in height, left is longer, right is wider, left one broadened in basal half and narrowed in apical half, right one broadened in basal three fourths and narrowed in apical fourth (Fig. 37 A). Genital capsule in ventral view: both gonopods widened, equal in height, hypandrium pointed in lower side, upper side with two feather like projections; phallic guide too broad and short; phallus with three ejaculatory ducts, one with tooth-like projection in basal third (Fig. 37 B); Genital capsule in lateral view: both surstyli curved to sternite, pointed at apex with a small projection towards gonopods at the base, phallic guide too broad with two depressions in the middle of ventral margin (Fig. 37 D – E); ejaculatory apodeme funnel-shaped with a bubble in its middle (Fig. 37 C). FEMALE: Body length: 3.1 mm. As male except for the following characters. Frons, eyes separated; completely silver-grey pollinose on lower 1 / 3, wider in middle, shiny black only around ocellar triangle, deep depression in just before ocellar triangle. Enlarged ommatidia shining; pulvilli and claws longer than last tarsal segment. Lateral bristles on the tergite 1 shorter than male. Ovipositor. Straight in ventral view, straight and short in lateral view (piercer brown), reaching sternite 3; basal two thirds silvery brown pollinose. LP: LB = 2.1. LDP: LPP = 3.3. (Fig. 156 C). Examined material: LECTOTYPE [HERE DESIGNATED]: AUSTRALIA: Queensland: Cairns, [16 ° 55 ’ S, 145 ° 46 ’ E], VII. 1904, JSS 4154 (1 ♂, ZMAN); PARALECTOTYPES: AUSTRALIA: Queensland: Cairns, 16 ° 55 ’ S, 145 ° 46 ’ E, VII. 1904, CNCD 5492 – 3 (2 ♀, BPBM); JSS 4155 (1 ♀, ZMAN); Other Material Examined: AUSTRALIA: Brisbane, Department of Primary Industries, Indooroopilly site, 27 ° 30 ’ S, 152 ° 59 ’ E, I. 1977; II. 1977, Malaise trap, JSS 8935, JSS 8937 (2 ♂, QDPC); Brisbane, 27 ° 28 ’ S, 153 ° 1 ’ E, 2. X. 1962, J. W. Turner, date recorded as 2 / 10 / 1962, JSS 8251 (1 ♂, QM); 5. IV. 1962, T. Brooks, JSS 454 (1 ♂, QM); Iron Range, North Queensland, 12 ° 44 ’ S, 143 ° 17 ’ E, 12. VI. 1971, J. Feehan, JSS 8700 (1 ♂, ANIC); James Cook University, 27 ° 28 ’ S, 153 ° 2 ’ E, dry savannah, 21 – 26. V. 1988, A. D. Austin, JSS 8645 (1 ♂, ANIC); Mareeba Shire, Kuranda, Russet Park, North Queensland, 16 ° 49 ’ S, 145 ° 38 ’ E, 460 m, 13. X. 1987, Thomas W. Davies, flight interception trap, Thomas W. Davies Collection donated to California Academy of Sciences 1987, JSS 8470 (1 ♂, CAS); CAMBODIA: Siem Reap, 13.365545, 103.836101, XI. 1958, N. L. H. Krauss, USNM _ ENT 01261318 (1 ♂, USNM); PHILIPPINES: Mountain Province: Abatan, Buguias, 60 km S of Bontoc, 16 ° 43 ’ N, 120 ° 49 ’ E, 2000 m, 25. V. 1964, H. M. Torrevillas, CNCD 5488 – 9 (1 ♂, 1 ♀, BPBM); Palawan: 13 km N of Puerto Princesa, ex. grass, 18. IV. 1968, D. E. Hardy, CNCD 5483 – 7 (3 ♂, 2 ♀, BPBM); 9.916624 ° N, 118.713946 ° E, 18. IV. 1968, D. E. Hardy, JSS 8488 – 9 (1 ♂, 1 ♀, BPBM); 15 – 17. IV. 1968, M. D. Delfinado, CNCD 5467 – 82 (13 ♂, 3 ♀, BPBM); Luzon: La Trinidad, 1388 m, 4 – 5. IV. 1968, D. E. Hardy, CNCD 5490 (1 ♂, BPBM); Mindanao: Maninit Str., Manolo Fortich Bukidnon, 24. IV. 1968, M. D. Delfinado, CNCD 5491 (1 ♂, BPBM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF84EC36D8D9EAABFE9210A0.taxon	distribution	Distribution: AUSTRALIA (Queensland), CAMBODIA, PHILIPPINES (Luzon, Mindanao, Mountain Province, Palawan) (Fig. 124). Notes: This is one of the few Australian species of Tomosvaryella that occurs in other countries. Unlike most Australian Tomosvaryella which favour dry habitats, T. epichalca is coastal. It should be expected in Indonesia and New Guinea as well. Perkins (1905) described this species and never indicated the holotype. There are four specimens in the syntype series including one male (JSS 4154) and three females (CNCD 5492 – 3 and JSS 4155). As the male specimens present more diagnostic characteristics than female, specimen JSS 4154 is hereby nominated as the lectotype to fix and ensure consistent interpretation of the name. Unlike most of Perkins species, this one was not reared.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF8BEC37D8D9EFF6FD77178E.taxon	description	Figs 38 A – E, 118	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF8BEC37D8D9EFF6FD77178E.taxon	diagnosis	Diagnosis: This species can be recognized by the line of bristles at the base of the hind trochanter; elongated surstyli in dorsal view (Fig. 38 A); extended gonopods towards surstyli, right one with a triangular lobe on inner margin in ventral view; phallus with a small membranous sheath at base of ejaculatory ducts, one of ejaculatory ducts divided into branches (Fig. 38 D); gonopods with a small hump-like projection dorsally in lateral view (Fig. 38 E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF8BEC37D8D9EFF6FD77178E.taxon	description	Description: MALE: Body length: 2.9 – 3.1 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 2 – 3 indistinct bristles. Mesonotum (viewed obliquely from front) greyish pollinose, anterior part more grey, grayish also from the side. Scutellum brownish silvery pollinose and with 4 – 6 weak bristles along the distal edge. Dorsocentral bristles indistinct, pale, longer along frontal edge, numerous short pale bristles around postpronotum present. Halter, knob pale with brown tinge, stem brown. Legs. Mid coxa with 2 – 3 long brown bristles. Trochanters brown, femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 4 of tibiae yellow (tibiae otherwise brown), tarsal segments yellow-brown dorsally, yellow ventrally, last segment darker. Hind trochanter with a shiny band ventrally (like a helix) and a line of bristles gradually turning into small bristles at the base; weak ventrobasal spines (2) present on fore and absent on mid femur. Ventroapical row of spines on fore femur absent; 4 – 5 small, black spines on mid femur; hind femur without ventroapical spines, except 10 – 12 equally spaced bristles posteroventrally, longer on distal half, longest up to 1 / 2 the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present (distinct). Hind tibia bent in posterior view. Hind metatarsus not flattened, almost as long as 2 – 4 combined, all segments with semi-erect bristles dorsally. Pulvilli shorter than last tarsal segment. Wing. Length: 3.0 – 3.1 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m at middle of discal cell (can be slightly distal to middle). 3 – 4 distinct, dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brown pollinose, tergite 1 silvery grey, large silvery pollinose spots on tergites 4 and 5 (spot on t 5 larger). Dispersed strong dark bristles on all tergites present, short and indistinct. Lateral spines on first tergite present, 4 – 6 dark bristles up to as long as hind femur’s width at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 0.9 – 1.1 times as long as ST 8. Genitalia without dissection: ST 8 large, rounded in dorsal view, brown and with a few distinct bristles, otherwise with velvet-like coverage; membranous area with round part extended dorsally and narrow part approaching epandrium, mostly directed posteriorly; epandrium dark brown, surstyli brown, broad. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.5). Surstyli elongated, curved to each other at apex (Fig. 38 A). Genital capsule in ventral view: both gonopods widened at base, extended towards sternite, right one with a small triangular lobes on inner margin; phallic guide with 3 – 4 dorso- dorsolateral spines, one spine longer than others, subepandrial sclerite distinct, phallus with a small membranous sheath at the base of ejaculatory ducts, one of ejaculatory ducts divided into branches (Fig. 38 B); Genital capsule in lateral view: both surstyli broadened at basal two thirds and moderately narrowed to apex, gonopods with a small hump-like projection dorsally, curved towards sternite (Fig. 38 DE). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 38 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF8BEC37D8D9EFF6FD77178E.taxon	etymology	Etymology: Eremia is Greek for desert and refers to the desert habitat that this species occurs in. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: ~ 3 km N [orth] of Denham, 25.909 ° S, 113.531 ° E, hilltop, 12. XII. 1999, J. Skevington, JSS 7340 (1 ♂, WAM); PARATYPES: AUSTRALIA: Western Australia: Mount Magnet, Mount Waramboo, 28 ° 2 ’ S, 117 ° 49 ’ E, hilltop, 14. XII. 1999, J. Skevington, JSS 7481 (1 ♂, QM); ~ 3 km North of Denham, 25.909 ° S, 113.531 ° E, hilltop, 12. XII. 1999, J. Skevington, JSS 7322, JSS 7326, JSS 7328, JSS 7330, JSS 7332 – 5, JSS 7341 – 2, JSS 7344, JSS 7346, JSS 7349, JSS 7351, JSS 7354, JSS 7355, JSS 7358 - 9, JSS 7361, JSS 7364, JSS 7382 – 5, JSS 7391 – 2, JSS 7395, JSS 7399, JSS 7403, JSS 7410 – 1 (2 ♂ CNC, 26 ♂, QM, 1 ♂ USNM; 2 ♂ WAM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF8BEC37D8D9EFF6FD77178E.taxon	distribution	Distribution: Australia (Western Australia) (Fig. 118). Notes: This is another possible Western Australian endemic. All known specimens have been encountered on hilltops. It is genetically most similar to T. brancha sp. nov., T. bulbosa sp. nov. and T. cona sp. nov. differing by 6.2 – 7.6 % pairwise divergence (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF89EC35D8D9EFF6FB681624.taxon	description	Figs 39 A – E, 121	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF89EC35D8D9EFF6FB681624.taxon	diagnosis	Diagnosis: This species can be recognized by having 2 – 4 dark bristles ventrobasally on the hind trochanter; elongated surstyli, moderately narrowed before apex, bent to each other at the apex, both surstyli with a small lobe at base along inner margin in dorsal view (Fig. 39 A); both gonopods extended towards surstyli with a semicircular projection on inner margin; phallic guide long and pointed at the apex with 3 – 4 dorso- dorsolateral spines in ventral margin (Fig. 39 B); both surstyli widened at basal half, strongly narrowed at the apical half in lateral view: (Fig. 39 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF89EC35D8D9EFF6FB681624.taxon	description	Description: MALE: Body length: 2.8 mm. Head. Flagellum long tapering, yellow-brown, brown pollinose. Scape and pedicel dark. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2.2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 3 – 4 medium dark bristles. Mesonotum (viewed obliquely from front) silvery-brown pollinose, anterior part greyer. Dorsocentral bristles short, dark, longer in frontal half, numerous pale bristles around postpronotum present. Scutellum silvery brown pollinose and with 6 – 8 short dark bristles along distal edge. Halter, knob pale, tip brownish, stem brown. Legs. Mid coxa with 2 – 3 long dark bristles. Trochanters brown, femora black, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments light brown. Hind trochanter with 2 – 4 dark bristles ventrobasally; 2 – 4 ventrobasal spines present on fore and shorter on mid femur. Hind femur without ventroapical spines. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres moderately flattened, hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 2.7 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 3 – 4 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brown-black, tergite 1 silvery grey with 8 – 10 long dark lateral bristles; other tergites brown pollinose; membranous area rounded. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.2). Surstyli equal in height, elongated, widened at the base, moderately narrowed before apex, bent to each other at the apex, both surstyli with a small lobe ventrobasally (Fig. 39 A). Genital capsule in ventral view: both gonopods extended towards surstyli, pointed at apex, gonopods with a semicircular projection on inner margin; phallic guide long and pointed at the apex with 3 – 4 dorso- dorsolateral spines, one is longer; subepandrial sclerite distinct (Fig. 39 B); Genital capsule in lateral view: both surstyli widened at basal half, curved to the sternite, strongly narrowed at the apical half (Fig. 39 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 39 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF89EC35D8D9EFF6FB681624.taxon	etymology	Etymology: This species is named after the Eucalyptus forest habitat that the only known specimen was captured in. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Lonesome National Park, near Lookout (LNP 4 M), 25 ° 30 ’ S, 148 ° 49 ’ E, 535 m, closed Eucalyptus woodland on rocky ridge, 3 – 26. XI. 2010, Lambkin et al., Malaise trap, CNC 591805 (1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF89EC35D8D9EFF6FB681624.taxon	distribution	Distribution: Australia (Queensland) (Fig. 121). Notes: Based on the terminalia characters, this species is close to Tomosvaryella ledknapperensis sp. nov. and Tomosvaryella prolata sp. nov. The shape of surstyli in lateral view differs from these two species.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF8FEC30D8D9EFF6FD201520.taxon	description	Figs 40 A – E, 119	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF8FEC30D8D9EFF6FD201520.taxon	diagnosis	Diagnosis: This species can be recognized by a distinct triangular protrusion on the hind femur ventrobasally, tiny surstyli covered by long and distinct bristles on the dorsal and central margin before the apex in dorsal view (Fig. 40 A), straight in lateral view (Fig. 40 D – E); gonopods small, wide hypandrium, phallic guide with some dorso-dorsolateral spines in ventral view (Fig. 40 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF8FEC30D8D9EFF6FD201520.taxon	description	Description: MALE: Body length: 2.4 mm. Head. Scape, pedicel and Flagellum dark brown, flagellum tips rather light brown, grey pollinose. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with three long light brown bristles and one short one. Mesonotum (viewed obliquely from front) silvery pollinose, posterior part more brownish. Scutellum silvery brown pollinose without bristles along distal edge. Dorsocentral bristles short, dark, longer in frontal half, some dark bristles around postpronotum present. Legs. Mid coxa with three long and one short dark bristle. Trochanters brown, femora black, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments light brown. Hind trochanter grey pollinose without distinct feature; ventrobasal spines present (1) on fore and absent on mid femur. Hind femur without ventroapical spines, with a distinct triangular protuberance ventrobasally. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres not flattened, hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 2.4 mm. Upper side of basal costal cell with one long brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2 – 3 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brown-black, tergite 1 brown in its middle, grey-brown pollinose. Lateral bristles on first tergite present, 4 – 5 dark bristles up to as long as ¾ of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium wider at base, longer than wide (MLE: MWE = 1.2). Surstyli tiny, broadened at base, then strongly constricted and broadened at apex with condensed long and distinct bristles on central and outer margin before apex, some bristles as wide as surstyli; both surstyli pointed to each other at apex (Fig. 40 A). Genital capsule in ventral view: gonopods small, equal in height; wide hypandrium; phallic guide with 3 – 6 short dorso- dorsolateral spines; subepandrial sclerite distinct (Fig. 40 B); Genital capsule in lateral view: both surstyli short and straight covered by some bristles apically, gonopods with two long membranous sheaths towards surstyli (Fig. 40 D – E). Ejaculatory apodeme tube-like, bent (Fig. 40 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF8FEC30D8D9EFF6FD201520.taxon	etymology	Etymology: From the Latin exiguus, little, short, scanty, poor or meagre, in reference to the tiny surstyli of the male. Examined material: HOLOTYPE: AUSTRALIA: Queensland: n [ea] r Teddington Weir SE of Maryborough, 25 ° 39 ’ S, 152 ° 42 ’ E, rainforest margin, 9. XI. 1990, G. & A. Daniels, JSS 8909 (1 ♂, QM); PARATYPE: AUSTRALIA: Western Australia: 8 km E Salmon Gums, 32 ° 58 ’ S, 121 ° 42 ’ E, 6. I. 1987, G. & A. Daniels, JSS 8915 (1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF8FEC30D8D9EFF6FD201520.taxon	distribution	Distribution: Australia (Queensland, Western Australia) (Fig. 119). Notes: This is another widespread species collected from two different habitats far from each other, from rainforest in the east to Eucalyptus forest in the west. Because of the distinctive triangular protuberance on the hind femur and the similarity of their genitalia, we treat these specimens as the same species. Fresh specimens are needed to allow for DNA sequencing and comparison.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF8CEC0ED8D9EAC7FE191464.taxon	description	Figs 41 A – E, 120	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF8CEC0ED8D9EAC7FE191464.taxon	diagnosis	Diagnosis: This species can be recognized by hind femur with 6 – 10 long bristles posterventrally; hind tibia covered with dense short light brown bristles ventroapically; surstyli symmetrical, straight in basal third, then falcate in dorsal view (Fig. 41 A); gonopods with two small triangle-shaped projections in ventral view (Fig. 41 B); one of ejaculatory ducts with two teeth at base (Fig. 41 D); phallic guide long with one long spine before apex (Fig. 41 D).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF8CEC0ED8D9EAC7FE191464.taxon	description	Description: MALE: Body length: 2.5 mm. Head (detached, stuck on the point). Scape and pedicel black, flagellum brown. Face silvery-brown pollinose; eyes touching for distance equal to 1.2 – 1.4 times the length of ocellar triangle. Occiput dark, silvery pollinose with scattered short dark bristles. Thorax. Postpronotum pale, with 3 – 4 long pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose. Scutellum brownish pollinose and with 7 – 8 dark bristles along the distal edge. Scutum with two uniseriate rows of conspicuous intra-alar bristles and two uniseriate rows of dorsocentral bristles, some long bristles on upper side of supra-alar area; postalar callus with three short dark bristles. Halter, knob yellow, stem brownish. Legs. Mid coxa with 3 – 5 long, brown bristles. Trochanters and femora dark brown, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 4 of tibiae yellow (tibia otherwise black), tarsal segments dark brown, ventrally somewhat paler. Hind trochanter light brown, broadened distally, without a keel, only a few short bristles ventrally. Ventroapical row of 4 – 8 very weak spines on fore femur present; 5 – 6 dark spines on distal third on hind femur; hind femur flattened apically, with 6 – 10 long, brown bristles posterventrally. Subapical (distal) spines on mid tibiae present, hind tibia covered with dense short light brown bristles ventroapically. Hind tarsi flattened, metatarsus almost as long as 2 – 5 combined; stronger, peg-like brown spines on metatarsus ventrally. Pulvilli as long as last tarsal segment. Wing. Length: 2.5 mm. Upper side of basal costal cell with 1 – 2 short dark bristle. Fourth costal section 3 – 3.5 times as long as third costal section (section 3 appears short). Cross-vein r-m distinctly distal to middle of discal cell. Two dark bristles on tegula. Abdomen. Tergites black, tergite 1 silvery grey, 6 – 10 lateral bristles on first tergite. Genitalia. Genital capsule in dorsal view: epandrium and surstyli brown. Epandrium longer than wide (MLE: MWE = 1.6). Surstyli symmetrical, straight in basal third, then falcate, left surstylus a little longer and wider than right one; (Fig. 41 A). Genital capsule in ventral view: gonopods with two small triangular projections extended towards surstyli, hypandrium broad and sclerotized, subepandrial sclerite long (Fig. 41 B); phallus with three ejaculatory ducts, one of ejaculatory ducts with two teeth at base. Genital capsule in lateral view: left surstylus broadened toward middle, bent in the middle; right surstylus straight in basal two third, then bent towards sternite; phallic guide long with one long spine before apex, (Fig. 41 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 41 C). FEMALE: unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF8CEC0ED8D9EAC7FE191464.taxon	etymology	Etymology: Falcatus is Latin for sickle-shaped or hooked and refers to the shape of the male surstyli. Examined material: HOLOTYPE: Australia: Queensland: near Brisbane, 27 ° 30 ’ S, 153 ° 1 ’ E, X – XI. 2000, JSS 13720 (1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF8CEC0ED8D9EAC7FE191464.taxon	distribution	Distribution: Australia (Queensland) (Fig. 120). Notes: The surstyli are shaped like those of Tomosvaryella leipoa sp. nov. (Fig. 56 A – D). Tomosvaryella falcata sp. nov. differs by having only one spine on the phallic guide, two spines on one of ejaculatory ducts, two triangle-shaped projections on the gonopods in ventral view (Fig. 41 D – E), and lack of the large triangular protuberance on the hind trochanter. This species is genetically closest to T. luma sp. nov. (3.5 – 4.7 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB2EC0CD8D9EB03FBD712B8.taxon	description	Figs 42 A – E, 123, 144 B	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB2EC0CD8D9EB03FBD712B8.taxon	diagnosis	Diagnosis: This species can be recognized by the two distinct spines on the hind trochanter (Fig. 144 B); abdominal sternites 3 – 4 with some spine-like bristles along posterior margin; surstyli slender and long in dorsal view (Fig. 42 A); gonopods widened with a distinct lobe towards each other in ventral view (Fig. 42 B); two ejaculatory ducts with spines (Fig. 42 B, D); surstyli with a distinct triangle-shaped lobe dorsobasally in lateral view (Fig. 42 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB2EC0CD8D9EB03FBD712B8.taxon	description	Description: MALE: Body length: 2.6 mm. Head. Scape, pedicel and Flagellum dark brown; scape with two short upper bristles. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput silvery-brown pollinose with scattered pale short bristles. Thorax. Postpronotum pale, with 4 – 5 long pale bristles. Mesonotum (viewed obliquely from front) silvery-brown pollinose, with some longer supra-alar bristles behind postpronotum. Scutellum brown pollinose and with 4 – 6 bristles along distal edge. Halter, knob pale, tip yellow, stem brown. Legs. Mid coxa with 3 – 4 long brown bristles. Trochanters brown, femora dark brown, shining ventrally; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments bright brown. Fore coxa with three short pale bristles ventromedially. Hind trochanter with two distinct spines (Fig. 144 B), ventromedial one darker, posteroventral one lighter and longer, hook like shape; ventrobasal spines present (1) on fore and absent on mid femur. Hind femur without ventroapical spines, two rows short light bristles posteroventrally. Fore femur with two distinct and long bristles ventrobasally. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres distinctly flattened except last one, hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 2.5 mm. Upper side of basal costal cell with one long light brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 3 – 5 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brown-black, covered with short light brown bristles, tergite 1 brown-silvery grey. Lateral bristles on first tergite present, 4 – 5 dark bristles up to as long as ¾ of hind femur’s width at base. Sternite 3 – 4 with 4 – 6 short spine-like bristles along posterior margin. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.5). Surstyli slender and elongated, curved to each other at apex, left surstylus is little longer (Fig. 42 A), both surstyli with a distinct lobe at the base. Genital capsule in ventral view: both gonopods extended towards surstyli and widened, pointed at apex with a distinct lobe on inner margin towards each other; phallic guide long, pointed at apex; phallus with three ejaculatory ducts, two ejaculatory ducts with spines, one spine is longer; subepandrial sclerite long (Fig. 42 B); Genital capsule in lateral view: both surstyli gently curved towards sternite, both with a distinct triangle-shaped lobe dorsobasally (Fig. 42 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 42 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB2EC0CD8D9EB03FBD712B8.taxon	etymology	Etymology: Named in honour of Stephen D. Gaimari, one of the collectors of the only known specimen and a major contributor to Diptera research. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Brisbane Forest Park, top of Scrub Road, 27 ° 25 ’ S, 152 ° 51 ’ E, 370 m, sclerophyll Eucalyptus forest, 1 – 7. XII. 1995, Irwin, Gaimari, JSS 16633 (1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB2EC0CD8D9EB03FBD712B8.taxon	distribution	Distribution: Australia (Queensland) (Fig. 123). Notes: It is surprising that only a single specimen of this species has been found as it is in a well-collected area. Different people have run long term Malaise trapping programs here and there has been a lot of hand collecting as well. The area contains a mixture of dry sclerophyll and wet sclerophyll (vine) forest.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB0EC0AD8D9EC7FFC1D11A4.taxon	description	Figs 43 A – E, 120	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB0EC0AD8D9EC7FFC1D11A4.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a semicircular keel, with 2 – 3 short spines along its edge; surstyli elongated, left one longer, broadened at apex in dorsal view (Fig. 43 A); hypandrium distinctly widened, subepandrial sclerite with three protrusions in its center; one of ejaculatory ducts bifurcate; phallic guide with a dorsal spine before apex in ventral view (Fig. 43 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB0EC0AD8D9EC7FFC1D11A4.taxon	description	Description: MALE: Body length: 2.9 mm. Head. Scape, pedicel, base of flagellum ark brown, otherwise light brown; flagellum tapering; grey pollinose; pedicel with two short upper bristles and one short lower bristle. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.2 times the length of ocellar triangle. Occiput silvery pollinose. Thorax. Postpronotum pale, with 3 – 4 long pale bristles. Mesonotum (viewed obliquely from front) silvery pollinose, with some long supra-alar bristles behind postpronotum lobes. Scutellum light grey, silvery pollinose and with 2 – 4 tiny dark bristles along distal edge. Halter, knob pale, tip brownish, stem light brown, base dark brown. Legs. Mid coxa with 3 – 4 long light brown bristles. Trochanters brown, femora black, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments dark brown dorsally, light brown ventrally. Hind trochanter with a semicircular keel, provided with 2 – 3 short spines along its edge; two ventrobasal spines on fore and mid femora, two row ventroapical spines present on fore and mid femora. Hind femur without ventroapical spines, except 4 – 6 equally spaced short bristles posteroventrally, longer on distal half, longest up to 1 / 2 the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres slightly flattened, hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 3.0 mm. Upper side of basal costal cell with two different-sized dark bristles. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 3 – 4 dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brown-black, tergite 1 brown, silvery brown pollinose. Lateral bristles on first tergite present, 7 – 8 brown bristles up to as long as half of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium wider at its base (MLE: MWE = 1.0). Surstyli slender and elongated, widened at the base, narrowed in the middle; left surstylus slightly broadened at apex; right surstylus shorter, bent towards left one at apex. (Fig. 43 A). Genital capsule in ventral view: both gonopods widened at base, extended towards surstyli, pointed at apex; hypandrium distinctly widened, subepandrial sclerite with three small protrusions in its center; phallus with three ejaculatory ducts, one divided two branches at the base; phallic guide with a small spine laterally before apex (Fig. 43 B); Genital capsule in lateral view: both surstyli straight in basal two thirds, bent towards sternite apically, phallic guide long, pointed at apex (Fig. 43 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 43 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB0EC0AD8D9EC7FFC1D11A4.taxon	etymology	Etymology: Named after the gibber (desert pavement) habitat the only known specimen was collected in. Gibber is a rocky desolate habitat with a specialized flora and fauna and intrinsic beauty. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Plevna Downs, 2.3 km SSW Arima (PD 2), 26 ° 32 ’ S, 142 ° 31 ’ E, 134 m, gibber, 16. IX – 20. X. 2008, Starick, Lambkin & Mackenzie, Malaise trap, CNC 575037 (1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB0EC0AD8D9EC7FFC1D11A4.taxon	distribution	Distribution: Australia (Queensland) (Fig. 120). Notes: Few pipunculids have been collected in gibber desert. More work is needed to find out if this species is a gibber specialist or is more widespread and simply under-collected.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB6EC0AD8D9ED43FD6F1738.taxon	description	Figs 44 A – E, 122	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB6EC0AD8D9ED43FD6F1738.taxon	diagnosis	Diagnosis: This species can be recognized by the triangle-shaped protrusion on the hind femur ventrobasally; surstyli tiny covered by long bristles basally in dorsal view (Fig. 44 A); both gonopods extended towards surstyli, phallic guide with 2 – 3 short dorso-dorsolateral spines in ventral view (Fig. 44 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB6EC0AD8D9ED43FD6F1738.taxon	description	Description: MALE: Body length: 2.6 – 2.7 mm. Head. Scape, pedicel, flagellum brown, arista dark, Flagellum tapering; yellow-brown at tips, brown pollinose. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching as long as the length of ocellar triangle. Occiput silvery pollinose. Thorax. Postpronotum pale, with 2 – 3 long pale bristles. Mesonotum (viewed obliquely from front) silvery pollinose with a row of short supra-alar bristles and some longer bristles after postpronotum. Scutellum silvery brown pollinose with 6 – 10 short dark bristles along distal edge. Halter, knob pale, tip brownish, stem brown. Legs. Mid coxa with 2 long and 1 – 2 short dark brown bristles. Trochanters brown, femora black, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments bright yellow completely. Hind trochanter with 4 – 5 dark bristles ventrobasally; ventrobasal spines absent on femora. Hind femur with a triangle-shaped protrusion ventrobasally. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres not flattened, hind metatarsus almost as long as 2 – 4 combined, dorsally. Pulvilli shorter than last tarsal segment. Wing. Length: 2.6 mm. Upper side of basal costal cell with one brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 1 – 2 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brown, tergite 1 silvery grey, silvery pollinose lateral spots on tergites 4 and 5. Lateral bristles on first tergite present, 2 – 4 long dark bristles up to as long as ¾ of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium widened at base, longer than wide (MLE: MWE = 1.7). Surstyli tiny, widened at the base with some condensed long bristles, moderately narrowed toward apex, pointed to each other at apex. (Fig. 44 A). Genital capsule in ventral view: both gonopods equal in height, extended towards surstyli, gonopods with a distinct projection on inner margin; phallic guide with 2 – 3 short dorso- dorsolateral spines; subepandrial sclerite distinct with transverse suture in the middle, (Fig. 44 B); Genital capsule in lateral view: both surstyli tiny, triangle-shaped (Fig. 44 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 44 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB6EC0AD8D9ED43FD6F1738.taxon	etymology	Etymology: Gidgee may refer to any of a number of species of Acacia native to arid or semi-arid regions of Australia, or to the vegetation communities in which these species dominate. This is the most typical habitat of Tomosvaryella species in Australia and is noted on the labels of all known specimens of this species. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Plevna Downs, Tompilly Hill base (PD 6 M), 26 ° 44 ’ S, 142 ° 39 ’ E, 187 m, gidgee, 25. XI – 18. XII. 2008, R. Mackenzie, Malaise trap, CNC 576416 (1 ♂, QM); PARATYPES: AUSTRALIA: Queensland: Plevna Downs, Tompilly Hill base (PD 6 M), 26 ° 44 ’ S, 142 ° 39 ’ E, 187 m, gidgee, 25. XI – 18. XII. 2008, R. Mackenzie, Malaise trap, CNC 576410, CNC 576414 (2 ♂, CNC, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB6EC0AD8D9ED43FD6F1738.taxon	distribution	Distribution: Australia (Queensland) (Fig. 122).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB4EC09D8D9EFF6FDE31544.taxon	description	Figs 45 A – D, 122	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB4EC09D8D9EFF6FDE31544.taxon	diagnosis	Diagnosis: This species can be recognized by elongated surstyli with two distinct triangle-shaped projections at the inner margin before the apex in dorsal view (Fig. 45 A); gonopods equal in height, extended towards surstyli, right gonopod wider with three dentations at apex in ventral view (Fig. 45 B); phallus with a small triangular projection before emerging ejaculatory ducts in lateral view (Fig. 45 C – D).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB4EC09D8D9EFF6FDE31544.taxon	description	Description: MALE: Body length: 1.1 mm (without head and abdomen). Thorax. Postpronotum light yellow, with 2 – 3 short pale bristles. Mesonotum (viewed obliquely from front) silvery-brown pollinose; scutum with scattered dark bristles at anterior supra-alar area, extended to posterior. Postalar callus dark brown with 1 – 2 short light brown bristles in the middle. Scutellum silvery pollinose and with a few (3 – 5) short dark bristles along the distal edge. Halter, knob pale, stem light brown, base dark. Legs. Mid coxa with 2 – 3 dark bristles. Coxae, trochanters, femora dark, hind femur shining ventrally, knees and basal 1 / 3 of tibiae light brown (tibiae otherwise dark brown), tarsomeres brown. Fore and mid trochanter with two short dark bristles; hind trochanter silvery pollinose covered by scattered short light brown bristles; two ventrobasal spines absent on fore mid femur; two ventroapical spines present on fore mid femur. Hind femur with two rows short spines ventrally. Hind tibia with two rows of short brown bristle on anterior and posterior side. Hind tarsomeres not flattened, covered by short light brown bristles ventrally and short brown bristles dorsally, hind metatarsus as long as 2 – 4 combined; pulvilli shorter than last tarsal segment. Wing. (one wing missing). Length: 3.1 mm. Upper side of basal costal cell with two short dark brown bristles and two same-sized light brown bristles. Fourth costal section 1.5 – 2 times as long as third costal section. Cross-vein r-m at middle of discal cell. Abdomen. Tergites 1 silvery pollinose with 5 – 6 different-sized lateral bristles; Genitalia. Genital capsule in dorsal view: epandrium slightly longer than wide (MLE: MWE = 1.1). Surstyli symmetrical, elongated, widened at base, moderately narrowed before apex, with two distinct triangle-shaped projections at inner margin before apex (Fig. 45 A). Genital capsule in ventral view: gonopods equal in height, extended towards surstyli, right gonopod wider with three dentations at apex (Fig. 45 B); phallic guide strong; subepandrial sclerite small and wide (Fig. 45 B); Genital capsule in lateral view: both surstyli widened at the base, narrowed and straight at the middle, then bent towards sternite, phallus with a small triangular projection before emerging ejaculatory ducts (Fig. 45 C – D). FEMALE: unknown. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Cairns, 16 ° 55 ’ S, 145 ° 46 ’ E, VIII. 1904, CNCD 5550 (1 ♂, BPBM)	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB4EC09D8D9EFF6FDE31544.taxon	etymology	Etymology: This new species is named in honor of D. Elmo Hardy, who discovered it as a new species but never published a description.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB4EC09D8D9EFF6FDE31544.taxon	distribution	Distribution: AUSTRALIA: (Queensland) (Fig. 122). Notes: The sole specimen of this species is a syntype of T. synadelpha but Hardy recognized it was different from the others and likely a new species.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB5EC07D8D9EAE3FC2814D4.taxon	description	Figs 46 A – E, 123, 153 C	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB5EC07D8D9EAE3FC2814D4.taxon	diagnosis	Diagnosis: This species can be recognized by scattered long dark bristles on scutellum; flattened hind tibia with condensed bristles apically; hind trochanter with a long keel and 3 – 10 short bristles; sternites 2 – 3 with different-sized spines, sternite 4 with a long keel in posterior margin; right gonopod with a small projection at inner margin before apex; phallic guide with 3 – 6 different-sized spines dorso- dorsolaterally in ventral view (Fig. 46 B); phallus with two downward small spines in lateral view (Fig. 46 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB5EC07D8D9EAE3FC2814D4.taxon	description	Description: MALE: Body length: 3.8 – 3.9 mm (without head). Thorax. Postpronotum light yellow, with 10 – 12 brown bristles in upper margin. Mesonotum (viewed obliquely from front) silvery-brown pollinose, anterior part greyer; scutum with scattered bristles at anterior supra-alar area, extended to posterior and uniseriate rows of intra-alar bristles. Postalar callus dark brown with three dark bristles in upper margin. Scutellum black, silvery pollinose covered with scattered long dark bristles entirely (not posterior margin). Halter, knob pale, stem light brown, base dark. Legs. Mid coxa with 3 – 5 distinct, long dark bristles and 4 – 5 long light brown bristles; hind coxa with 2 – 4 short brown bristles. Coxae, trochanters, femora dark brown, hind femur shining ventrally, knees and basal 1 / 4 of tibiae yellow (tibiae otherwise brown), tarsomeres light brown ventrally and dark dorsally. Mid trochanter with 2 – 6 different-sized bristles ventroapically. Hind trochanter silvery pollinose, with a distinct and long keel and 3 – 10 short bristles ventromedially and ventrobasally; ventrobasal spines absent on fore and mid femur. Hind femur with two rows ventroapical spines. Hind tibia moderately flattened from basal to apical, with a wrinkled indentation in the middle, two rows of short black bristle on anterior and posterior side covered by dense and long dark bristles (longer bristles condensed apically). Hind tarsomeres distinctly flattened, covered by dense dark bristles dorsally and brown bristles ventrally, last hind tarsomere rather rounded, hind metatarsus as long as 2 – 4 combined; pulvilli shorter than last tarsal segment. Wing. Length: 5 mm. Upper side of basal costal cell with one or two long dark brown bristle and two short light brown bristles. Fourth costal section 1.5 – 2 times as long as third costal section. Cross-vein r-m at middle of discal cell. 8 – 10 short dark setulae on tegula. Abdomen. Tergites 1 silvery pollinose with 20 – 30 long dark bristles, extending from lateral onto dorsal surface, moderately shortened from lateral to dorsal; tergite 4 – 5 with two gray spots laterally, all tergites with scattered distinct dark bristles. sternite 2 with 3 – 6 small dark spines in the middle of posterior margin. Sternite 3 with two long spines and sternite 4 with two distinct rectangular-shaped keels along posterior margin. Syntergosternite 8 enlarged, dark brown and grey pollinose. Membranous area long and broad in middle. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.9). Surstyli small, left surstylus longer and wider than right one, right surstylus with a triangular edge at inner margin before apex (Fig. 46 A). Genital capsule in ventral view: both gonopods widened, unequal in height, left gonopod longer than left one, right gonopod with a small projection at inner margin before apex; phallic guide with 3 – 6 different-sized spines dorso- dorsolaterally, one is longer than others; subepandrial sclerite small (Fig. 46 B); Genital capsule in lateral view: both surstyli rather straight in basal two thirds, curved to sternite in basal third (Fig. 46 D – E); phallus with two downward small spines; ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 46 C). FEMALE: Body length: 3.9 mm. As male except for the following characters. Frons, eyes separated, as wide as the width at antennae; completely silver-grey pollinose on lower 1 / 3, shiny black only around ocellar triangle, smooth depression in just before ocellar triangle. Enlarged ommatidia silvery shining. Length of bristles on hind tibia shorter than male; last hind tarsomere not rounded (elongated); pulvilli and claws about 1.2 – 1.5 times as long as last tarsal segment. Abdominal sternites without spines. Ovipositor. Straight in ventral view, straight, in lateral view (yellow piercer, dark brown base), reaching middle of sternite 3; base silvery brown pollinose. LP: LB = 4.1. LDP: LPP = 2.7. (Fig. 153 C).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB5EC07D8D9EAE3FC2814D4.taxon	etymology	Etymology: The specific name is derived from the Latin word ‘ hirta’, meaning hairy, referring to bristles on the scutellum and hind tibia. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Isla Gorge Nat [ional] Park, 25 ° 11 ’ S, 149 ° 58 ’ E, 320 m, 4. X. 1992, G. Daniels, JSS 8906 (1 ♂, QM); PARATYPES: AUSTRALIA: Queensland: Isla Gorge National Park, 25 ° 11 ’ S, 149 ° 58 ’ E, 320 m, 3. III. 1991; 4. X. 1992, G. Daniels, JSS 8905, JSS 8907 (1 ♂, 1 ♀ QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB5EC07D8D9EAE3FC2814D4.taxon	distribution	Distribution: Australia (Queensland) (Fig. 123). Notes: The only known specimens are all from Isla Gorge National Park, a spectacular area of sandstone cliffs and dry sclerophyll forest. This species is genetically most similar to T. millstreamensis sp. nov. (2.2 – 3.2 % pairwise divergence) and T. tribula sp. nov. (2.7 – 3.2 %) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFBBEC05D8D9EB93FE191284.taxon	description	Figs 47 A – E, 125	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFBBEC05D8D9EB93FE191284.taxon	diagnosis	Diagnosis: This species can be recognized by shape of surstyli in dorsal view, broadened at base, gradually narrowed toward apex in dorsal view (Fig. 47 A); long hypandrium, round gonopods at apex, indistinct subepandrial sclerite and sinuous ejaculatory ducts in ventral view (Fig. 47 B); broadened surstyli in lateral view (Fig. 47 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFBBEC05D8D9EB93FE191284.taxon	description	Description: MALE: Body length: 3.8 mm. Head. Flagellum tapering; brown, gray pollinose. Scape, pedicel black. Face silvery pollinose. Frons shining black; eyes touching for distance equal to 1.1 times the length of ocellar triangle. Occiput silvery pollinose. Thorax. Postpronotum pale, with 4 – 6 extra short, pale bristles. Mesonotum (viewed obliquely from front) greyish pollinose, anterior part greyer. Scutellum silvery pollinose and with 7 – 8 weak bristles along the distal edge. Dorsocentral bristles with very short and pale in a row, somewhat longer along frontal edge. Halter, knob pale, stem brown. Legs. Mid coxa with three long, dark bristles. Coxae, trochanters and femora black, shining ventrally, silvery pollinose posteriorly; knees and basal 1 / 5 of tibiae and femora yellow, tarsal segments brown, ventrally somewhat paler, last segment same dark brown. Mid trochanter with 6 – 8 short bristles. Hind trochanter without a keel, only some short bristles ventrally. Ventroapical row of 4 – 5 very weak spines on fore femur present; 5 – 6 small, black spines on mid femur. Ventrobasal spines absent on fore and mid femora; hind tibia with a wrinkle in the middle. Hind tarsi long, metatarsus almost as long as 2 – 5 combined; stronger, peg-like dark spines on metatarsus laterally along anterior edge. Pulvilli as long as last tarsal segment. Wing. Length: 3.8 mm. Upper side of basal costal cell with three long pale and three shorter bristles short dark bristle. Fourth costal section 3 – 3.5 times as long as third costal section. Cross-vein r-m distinctly distal to middle of discal cell. Abdomen. detached. Genitalia. capsule in dorsal view: epandrium and surstyli brown. Epandrium as long as wide (MLE: MWE = 0.0). Surstyli rather symmetrical, elongated; both surstyli broadened at base, moderately narrowed towards apex; right surstylus wider than left one basally, left surstylus damaged before apex (Fig. 47 A). Genital capsule in ventral view: gonopods rounded at apex, equal in height; hypandrium long and sclerotized, subepandrial sclerite short and indistinct, only short distance between gonopod and base of surstyli (Fig. 47 B); phallus with three distinct long sinuous ejaculatory ducts. Genital capsule in lateral view: both surstyli broadened and gently curved towards sternite before apex; phallic guide broad, pointed at apex (Figs 47 D – E). Ejaculatory apodeme spade-shaped (Fig. 47 C). FEMALE: unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFBBEC05D8D9EB93FE191284.taxon	etymology	Etymology: Indistinctus is Latin for obscure or dim and refers to the indistinct subepandrial sclerite found in the male genitalia. Examined material: HOLOTYPE: Australia: New South Wales: Kinchega National Park, near Ranger h [ead] q [uarters], 32 ° 24 ’ S, 142 ° 23 ’ E, on paper daisy flower, 15. X. 1997, J. Skevington, S. Winterton & C. Lambkin, JSS 8368 (1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFBBEC05D8D9EB93FE191284.taxon	distribution	Distribution: Australia (New South Wales) (Fig. 125). Note: This species is genetically most similar to T. biloba sp. nov. (9.8 – 18.1 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB9EC00D8D9EC23FE6312F0.taxon	description	Figs 48 A – E, 125	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB9EC00D8D9EC23FE6312F0.taxon	diagnosis	Diagnosis: This species can be recognized by elongated surstyli; broadened in basal half, narrowed before apex, apex slightly broadened in dorsal view (Fig. 48 A); equal gonopods in height, rounded at apex; phallic guide short, broadened at base with final hook-like projection in lateral view; one of ejaculatory ducts covered by teeth in basal half (Fig. 48 D); ejaculatory apodeme funnel-shape (Fig. 48 C).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB9EC00D8D9EC23FE6312F0.taxon	description	Description: MALE. Body length: 3.0 – 3.5 mm. Head. Pedicel brown with 2 – 3 upper bristles and 1 – 2 lower bristles; flagellum acuminate; brown. Face silvery pollinose. Frons, upper part close to antennae black, lower part grey pollinose; eyes touching for distance equal to 2.5 – 3 times ocellar triangle. Occiput silvery pollinose, slightly less so on upper half. Thorax. Postpronotum pale brown, with 2 – 3 short whitish bristles. Mesonotum (viewed obliquely from front) subshining black, anterior part indistinctly brown pollinose, slightly brownish also from the side. Scutellum with dorsocentral bristles weakly developed, dark, somewhat longer on frontal part. Halter, knob pale, stem dark brown. Legs. Mid coxa with 2 – 3 dark bristles. Trochanters and base of femora brown, femora dark brown, shining ventrally, femora silvery pollinose posteriorly except shiny hind femur; knees, basal 1 / 3 of tibiae yellow, tarsal segments pale brown, last segment dorsally darker. Ventroapical row of spines on fore femur tiny, 4 – 6, 8 – 10 short, black spines on mid femur; 6 – 8 spines on hind femur, no bristles posteriorly. Subapical (distal) spines on first four tibiae absent. Hind trochanter grey pollinose, without bristles on smoothly curved ventral side, ventrobasal spines absent on fore and mid femora. Hind tarsi not distinctly flattened but wider than fore and mid tarsi, first tarsomere slightly shorter than 2 – 5 combined; ventral bristles scrub-like. Pulvilli as long as last tarsal segment. Wing. Length: 2.8 – 3.1 mm. Upper margin of basal costal cell with two long distinct bristles. Fourth costal section 2 – 3 times as long as third costal section. Cross-vein r-m distinctly distal to middle of discal cell. Setulae on tegula absent. Abdomen. Viewed obliquely from front tergites golden-brownish pollinose, tergite 1 silvery grey pollinose with 6 – 8 short lateral bristles (up to 0.15 mm), sides faintly silvery on tergites 4 and 5 (the latter with larger spot, but weakly colored). Dispersed short dark bristles on tergites absent. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 twice as long as ST 8. Genitalia without dissection: ST 8 small, mainly shiny brown; membranous area lanceolate, shaped like a sunflower seed, vertical; surstyli relatively small, brown; hairy cerci visible without dissection in lateral view. Genitalia. Surstyli hairy, wide at base in ventral view, narrowing at 2 / 3 of the length and broadening at tip, both uniformly wide in lateral view and pointed at tip; epandrium short and wide; cerci symmetrically placed, small; ST 8 with a narrowing membranous area towards epandrium. Genital capsule in dorsal view: epandrium and surstyli dark brown. Epandrium wider than long (MLE: MWE = 0.6). Both surstyli broadened in basal half, then strongly narrowed before apex, apex rather broadened less than basal (Fig. 48 A). Genital capsule in ventral view: subepandrial sclerite short and narrow, hidden behind phallic guide, without spines; gonopods symmetrical broad, apex rounded; phallus trifid, one of ejaculatory ducts covered by teeth in basal half (Fig. 48 B). Genital capsule in lateral view: both surstyli straight in basal half, strongly bent (right angle) towards sternite (Figs 48 D – E). phallic guide broadened with final thin hook-like projection, visible in lateral view (Fig. 48 D); ejaculatory apodeme funnel shape (Fig. 48 C). FEMALE. Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB9EC00D8D9EC23FE6312F0.taxon	etymology	Etymology: From the Latin infundibulum, funnel, in reference to the funnel-shaped ejaculatory apodeme. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: Millstream-Chichester Nat [ional] P [ark], Roebourne Road, 21 ° 26 ’ S, 117 ° 9 ’ E, 337 m, sandy creek, Eucalyptus Spinifex grassland, 7 – 12. V. 2003, C. Lambkin & D. Yeates, Malaise trap, ANIC [Bulk Sample] 2090, JSS 15570 (1 ♂, WAM); Paratypes: AUSTRALIA: Western Australia: Millstream-Chichester National Park near Crossing Pool, 21 ° 35 ’ S, 117 ° 4 ’ E, 276 m, over creek, Melaleuca riverine forest, 28. iv – 5. v. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 15965; JSS 16190 (2 ♂, ANIC); 304 m, Melaleuca riverine forest, 5 – 11. v. 2003, C. Lambkin, D. Yeates & J. Recsei, Malaise trap, JSS 15598 (1 ♂, ANIC); New South Wales: Crowdy Bay National Park, 40 km northeast of Taree, 31 ° 39 ’ S, 152 ° 46 ’ E, 16. iii. 1981, B. J. Loudon, JSS 8740 (1 ♂, ASCU); Queensland: JSS 4188 (1 ♂, ZMAN) [previous syntype of T. pseudophanes]; Carnarvon National Park, north of Ranger Station near Mount Rugged, 24 ° 55 ’ S, 148 ° 0 ’ E, near road, 12. X. 2002, S. Boucher, sweeping, LEM _ 0016213 (1 ♂, LEM); Carnarvon Station, near Homestead, 24 ° 48 ’ S, 147 ° 45 ’ E, 753 m, flowering herbs, 12 – 15. X. 2014, Lambkin, Wright, J. & K. Wilson, Malaise trap, burnt 2014, 37270, CNC 597136 – 7 (2 ♂, CNC); Halifax, 18 ° 35 ’ S, 146 ° 17 ’ E, IV. 1920, F. Muir, CNCD 5557 – 8 (2 ♂, BPBM); 1 km W Cooktown, 15 ° 28 ’ S 145 ° 15 ’ E, 12. v. 1981, D. H. Colless, JSS 177 (1 ♂, ANIC); 29 km S of Lakeland, 16 ° 5 ’ S, 144 ° 47 ’ E, 16. iv. 1980, G. F. Hevel and J. A. Fortin, JSS 10538 (1 ♂, USNM); 5 miles west Ravenshoe, 17 ° 38 ’ S, 145 ° 29 ’ E, 780 m, 6. xi. 1962, E. S. Ross & D. Q. Cavagnaro, JSS 8475 (1 ♂, USNM); 6 km North of Mt. Molloy, 16 ° 41 ’ S, 145 ° 20 ’ E, 16. iv. 1980, G. F. Hevel & J. A. Fortin, JSS 10519 (1 ♂, USNM); Big Mitchell Creek Mareeba-Molloy Road, North Queensland, 16 ° 48 ’ S, 145 ° 21 ’ E, 04. v. 1967, D. H. Colless, JSS 135 (1 ♂, ANIC), Rocky Creek, 7 m [11.2 km] North Atherton, North Queensland, 17 ° 7 ’ S, 145 ° 29 ’ E, 3. V. 1967, D. H. Colless, JSS 8530 (1 ♂, ANIC); Brisbane Forest Park, top of Scrub Road, 27 ° 25 ’ S, 152 ° 51 ’ E, sclerophyll Eucalyptus forest, 04. xi. 1995, Irwin & Gaimari, JSS 8833, JSS 8900 (2 ♂, INHS); Brisbane Forest Park, 27 ° 25 ’ S, 152 ° 51 ’ E, sclerophyll Eucalyptus forest, 8. xi. 1995, Irwin & Gaimari, JSS 8838 (1 ♂, INHS); Brisbane, 27 ° 28 ’ S, 153 ° 01 ’ E, 12. iii. 1960, E. M. Exley, JSS 8263 (1 ♂, QM); Brown Lake, Stradbroke Island, 27 ° 29 ’ S 153 ° 25 ’ E, 01. iii. 1980, G. Daniels, JSS 8918 (1 ♂, QM); Bunya Mountains, 26 ° 51 ’ S, 151 ° 33 ’ E, 20. v. 1962, G. Monteith, JSS 8260 (1 ♂, QM); Burnett River, Bundaberg, 24 ° 52 ’ S, 152 ° 21 ’ E, 15 – 17. iii. 1972, H. Frauca, JSS 8538 – 9 (2 ♂, ANIC); Cape York, Stewart Range, 14 ° 04 ’ S, 143 ° 41 ’ E, 500 m, 3. v – 29. ix. 1961, L. & M. Gressitt, CNCD 5553 (1 ♂, BPBM); Carnarvon National Park, Consuelo land, 24 ° 57 ’ S 148 ° 6 ’ E, mahogany forest, 12. x. 2002, M. von Tschirnhaus, JSS 13951 (1 ♂, CNC); Carnarvon National Park, Mount Moffatt Section, 3 km SE Headquarters, 25 ° 5 ’ S, 148 ° 01 ’ E, 740 m, 20. xi. 1995, M. E. Irwin & S. D. Gaimari, JSS 8901 (1 ♂, INHS); Carnarvon National Park, Mount Moffatt Section, 24 ° 52 ’ S, 148 ° 01 ’ E, 840 m, next to water pool, 24. xi. 1995, M. E. Irwin, Malaise trap, JSS 8831, JSS 8843 (2 ♂, INHS); Carnarvon National Park, Mount Moffatt Section, Malaise traps near Mount Moffatt, 25 ° 04 ’ S, 148 ° 03 ’ E, 19 – 23. i. 1998, J. & A. Skevington & S. Winterton, Malaise trap, JSS 2717 (1 ♂, QM); Carnarvon National Park, Mount Moffatt Section, Top Shelter Shed, 24 ° 55 ’ S, 148 ° 03 ’ E, 1100 m, hilltop, 22. xi. 1995, D. K. Yeates, JSS 8310, JSS 8314 – 6 (4 ♂, QM); Carnarvon Station, near Piebald Spring (CN 1 M 1), 24 ° 50 ’ S, 147 ° 45 ’ E, 821 m, Eucalyptus / Callistemon, in rocky gully, 13. xii. 2010 – 15. vi. 2011, C. Zwick & C. Wilson, Malaise trap, CNC 576485, CNC 576488 (2 ♂, CNC); Durham downs, Nokandra, 26 ° 07 ’ S, 149 ° 04 ’ E, 13. xi. 1949, S. J. Paramonov, JSS 8677 (1 ♂, ANIC); 29. iii – 4. iv. 1998, N. Power, Malaise trap, JSS 7900 (1 ♂, QM); Noosa, 26 ° 23 ’ S, 153 ° 06 ’ E, 27. viii. 1959, F. A. Perkins, JSS 8254 (1 ♂, QM); Peawaddy Gorge, Mount Moffatt National Park, 24 ° 56 ’ S, 148 ° 04 ’ E, 1100 m, 26. ii. 1996, C. J. Burwell, S. Evans, JSS 9745 (1 ♂, QM); SW Brisbane College, 27 ° 28 ’ S, 153 ° 02 ’ E, Eucalyptus, 1. ii. 1961, J. L. Gressitt, CNCD 5556 (1 ♂, BPBM); 16 – 23. i. 1998, S. Winterton, N. Power & D. White, Malaise trap, JSS 8431 – 3 (3 ♂, QM); 26. ix – 3. x. 1997, S. Winterton, N. Power & D. White, Malaise trap, JSS 8849 (1 ♂, QM); Flinder’s Peak, 27 ° 49 ’ S, 152 ° 48 ” E 10. iii. 1963, G. Monteith, CNCD 5569 (1 ♂, BPBM); Western Australia: 11 km E Marble Bar at Brockman Creek, 21 ° 09 ’ S, 119 ° 52 ’ E, 187 m, damp, sandy creek bed, 2 – 14. v. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15684 (1 ♂, CNC); 22 km E Cobra Station, 24 ° 13 ’ S, 116 ° 33 ’ E, 381 m, small dry wash, 26. iv – 10. v. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15681 (1 ♂, CNC); 4 miles SE by S of Minilya, 23 ° 49 ’ S, 114 ° 36 ’ E, 17. x. 1970, D. H. Colless, JSS 8701 (1 ♂, ANIC); Karijini National Park, Hamersley-Mount Bruce Road, 22 ° 34 ’ S, 118 ° 18 ’ E, 757 m, dry rocky creek bed, Eucalyptus grassland, 25. iv – 14. v. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 15968, JSS 16117 (2 ♂, ANIC); 25. iv – 14. v. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 15986, JSS 15989 – 90, JSS 15992, JSS 15994 – 6, JSS 15998, JSS 16000 (9 ♂, ANIC); Karijini National Park, Weano Gorge Road, 22 ° 21 ’ S, 118 ° 15 ’ E, 695 m, grassy dry creek, Eucalyptus Acacia scrub, 20 – 25. iv. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16032 – 4 (3 ♂, ANIC); 25. iv – 15. v. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16079 (1 ♂, ANIC); Karijini National Park: Juna Downs Road (between drying pools in rocky Turee Creek), 22 ° 42 ’ S, 118 ° 25 ’ E, 789 m, 25. iv – 14. v. 2003, C. Lambkin & T. Weir, Malaise trap, CNCD 3830, CNCD 3847 (2 ♂, QM); Millstream-Chichester National Park, Roebourne Road, 21 ° 26 ’ S, 117 ° 09 ’ E, 337 m, sandy creek, Eucalyptus Spinifex grassland, 27. iv – 3. v. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16074, JSS 16195, JSS 16197, JSS 16199 (4 ♂, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFB9EC00D8D9EC23FE6312F0.taxon	distribution	Distribution: Australia (New South Wales, Northern Territory, Queensland, Western Australia) (Fig. 125). Notes: Specimen JSS 4188 was previously recognized as a syntype of T. pseudophanes. Based on the different shape of terminalia and external characters, we include it in T. infundibula sp. nov. The hind femur of T. infundibula sp. nov. has no bristles posteriorly while T. pseudophanes has posterior bristles. T. infundibula sp. nov. also has two long distinct bristles on the upper margin of basal costal cell. The shape of surstyli is also different in dorsal and lateral view (Fig. 48 A, D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFBCEC1ED8D9EDB7FF631248.taxon	description	Figs 49 A – F, 126	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFBCEC1ED8D9EDB7FF631248.taxon	diagnosis	Diagnosis: This species can be recognized by the small patch of bristles on the hind trochanter; elongated surstyli, left surstylus broader than right one in middle in dorsal view (Fig. 49 A); elongated gonopods; subepandrial sclerite with some horizontal wrinkles (Fig. 49 B); phallic guide with 3 – 4 dorso- dorsolateral spines (Fig. 49 F); both surstyli curved to sternite, straight in middle in lateral view (Fig. 49 D – F).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFBCEC1ED8D9EDB7FF631248.taxon	description	Description: MALE: Body length: 2.2 mm. Head. Flagellum acuminate, brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 times the length of ocellar triangle. Occiput silvery pollinose, slightly less so on upper half. Thorax. Postpronotum pale, with 4 – 5 pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, greyish also from the side. Scutellum not visible (glue blocks the view). Dorsocentral bristles distinct, dark, somewhat longer on frontal edge, numerous long bristles around postpronotum present. Halter, knob brown, stem brown. Legs. Mid coxa with 3 long, dark bristles. Trochanters and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow-brown (tibiae otherwise dark brown), tarsal segments yellow-brown, ventrally paler, last segment darker. Hind trochanter with a small patch of extra short bristles; ventrobasal spines absent on fore and mid femora. Ventroapical row of spines on fore femur absent; absent on mid femur; hind femur without ventroapical spines, bristles not visible on specimen. Subapical (distal) spines on first four tibiae absent. Hind trochanter with a small patch of extra short bristles; ventrobasal spines absent on fore and mid femora. Hind tibia not bent in posterior view. Metatarsi slightly flattened, almost as long as 2 – 4 combined. Pulvilli shorter than last tarsal segment. Wing. Length: 3.1 mm. Upper side of basal costal cell with one light bristle. Fourth costal section 2 times as long as third costal section. Cross-vein r-m at 2 / 5 of discal cell (proximal to middle). 2 – 3 distinct brown setulae on tegula. Abdomen. Dissected. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.8). Surstyli elongated towards surstyli, broadened at base, straightened in middle, left surstylus wider than right one, distally curved to each other (Fig. 49 A). Genital capsule in ventral view: gonopods elongated, tapering at apex, unequal in height, left is longer than right one; subepandrial sclerite with 3 – 4 horizontal wrinkles (Fig. 49 B); phallic guide with 3 – 4 dorso- dorsolateral spines (Fig. 49 F). Genital capsule in lateral view: both surstyli curved towards sternite, straight in middle (Fig. 49 D – F). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 49 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFBCEC1ED8D9EDB7FF631248.taxon	etymology	Etymology: This species is named in honour of Mike Irwin, one of the collectors of the only known specimen. Mike collected 9 % of the specimens used in this revision and 23 % of the species (Supplementary file 2). Examined material: HOLOTYPE: AUSTRALIA: Queensland: Brisbane Forest Park, 27 ° 25 ’ S, 152 ° 51 ’ E, sclerophyl [l] Eucal [y] ptus forest, 4. XI. 1995, Irwin & Gaimari, JSS 8842 (1 ♂, QM); PARATYPES: New South Wales: 8 km S Mendooran, 31 ° 50 ’ S, 149 ° 6 ’ E, 28. I. 1987, G. & A. Daniels, JSS 8913 (1 ♂, QM); Queensland: Brisbane Forest Park, Enoggera Creek at Scrub Road Crossing, 27 ° 26 ’ S, 152 ° 50 ’ E; 27 ° 26 ’ S, 152 ° 51 ’ E; 27 ° 29 ’ S, 152 ° 50 ’ E, 200 m, tropical rainforest with Eucalyptus, 30. X – 2. XI. 1995; 2 – 6. XI. 1995; 6 – 9. XI. 1995, 7 – 27. XII. 1995, M. E. Irwin, Malaise trap; CNCD 3774, CNCD 134860, CNCD 134863 – 4, CNCD 134867, CNCD 134873, CNCD 134889, CNCD 134898, CNCD 134900, CNCD 137146 – 7, CNCD 137154 – 5, CNCD 137157, CNCD 137167; CNCD 137176, CNCD 137191, CNCD 137198, CNCD 137200, CNCD 137202, CNCD 137404, CNCD 137409, CNCD 147446, CNCD 147461, CNCD 147469 – 70, CNCD 147486, CNCD 147495 – 96 (18 ♂, 1 ♀, CNC; 8 ♂, QM; 2 ♂, USNM); Brisbane Forest Park, 27 ° 26 ’ S, 152 ° 51 ’ E, 200 m, 9 – 14. XI. 1995, M. E. Irwin, CNCD 136727, CNCD 136740, CNCD 136746, CNCD 136760, CNCD 136764 (5 ♂, CNC); Mount Glorious, Property of Tony Hiller, 27 ° 20 ’ S, 152 ° 46 ’ E, 30. X – 15. XI. 1995, M. E. Irwin, CNCD 136079, CNCD 136087 (1 ♂, 1 ♀, CNC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFBCEC1ED8D9EDB7FF631248.taxon	distribution	Distribution: Australia (New South Wales, Queensland) (Fig. 126). Notes: This species has a very narrow range in NE New South Wales and SE Queensland. Intraspecific genetic distance is 0.9 %. It is genetically most similar to T. creagra sp. nov. (8.2 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA2EC1FD8D9EDEFFA0B10A0.taxon	description	Figs 50 A – E, 125	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA2EC1FD8D9EDEFFA0B10A0.taxon	diagnosis	Diagnosis: This species can be recognized by the hind femur being broadened at the base; surstyli with a hump-like projection in the dorsomedial margin, covered by long and distinct bristles along outer margin in dorsal view; gonopods extended towards surstyli, pointed at apex; phallic guide with at least two distinct spines in ventral view (Fig. 50 B); both surstyli straight in lateral view (Fig. 50 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA2EC1FD8D9EDEFFA0B10A0.taxon	description	Description: MALE: Body length: 2.4 mm. Head. (damaged). Thorax. Postpronotum pale, with 3 – 4 short pale bristles. Mesonotum (viewed obliquely from front) silvery pollinose, anterior part more grey, grayish also from the side. Scutellum silvery pollinose. Halter, knob and stem pale, tip brownish. Legs. Mid coxa with 3 – 4 dark bristles. Trochanters brown, femora black, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments brown. Hind trochanter gray pollinose without any features. Ventrobasal spines present (1) on fore and mid femur. Hind femur without ventroapical spines. Hind femur shinny posteroventrally, broadened at base and moderately narrow towards knee. Hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 2.5 mm. Upper side of basal costal cell with one long brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 4 – 5 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites subshiny brown-black, tergite 1 silvery grey. Lateral bristles on first tergite present, 4 – 6 dark bristles up to as long as ¾ of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.1). Surstyli symmetrical, broadened at the base, then narrowed strongly, widened in the middle, curved to each other towards apex, both surstyli with long condensed bristles at outer margin, longer at the middle (Fig. 50 A). Genital capsule in ventral view: both gonopods unequal in height, left is slightly longer than right one, extended towards surstyli, pointed at apex; phallic guide with at least two long dorso- dorsolateral spines; subepandrial sclerite rectangular-shaped (Fig. 50 B); Genital capsule in lateral view: both surstyli straight, broadened at base, moderately narrow to apex (Fig. 50 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 50 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA2EC1FD8D9EDEFFA0B10A0.taxon	etymology	Etymology: This species is named after the type locality, Koonchera Waterhole. This location is in the northeastern corner of South Australia, a remote location south of Birdsville, Queensland near the Simpson Desert. This was Jeff Skevington’s first major field trip as a graduate student at the University of Queensland and the team of Christine Lambkin, Shaun Winterton, Angela Skevington, Andreas Zwick and Hugh and Jeri Dingle all collected and studied the natural history of this region together. The trip proved to be a bit too early for most pipunculids, but several species of arid land specialists were collected, including this singleton specimen of a new species. Examined material: HOLOTYPE: AUSTRALIA: South Australia: Koonchera Dune near Koonchera Waterhole, 26 ° 41 ’ S, 139 ° 30 ’ E, on dune top under large tree, 3. IX. 1997, J. & A. Skevington, JSS 8320 (1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA2EC1FD8D9EDEFFA0B10A0.taxon	distribution	Distribution: Australia (South Australia) (Fig. 125). Notes: Pipunculid specimens of a number of species collected in this location appeared to be hilltopping along the dune ridge. More specimens are needed to confirm this behaviour in this species. Based on the DNA sequencing, this species is genetically most similar to T. pterygia sp. nov. (1.1 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA0EC1CD8D9EFF6FB8D1704.taxon	description	Figs 51 A – E, 126,147 A	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA0EC1CD8D9EFF6FB8D1704.taxon	diagnosis	Diagnosis: This species can be recognized by having the hind trochanter without a keel, but with some dark spines in a row ventrobasally (Fig. 147 A); hind femur with some long bristles posteriorly (Fig. 147 A); elongated surstyli in dorsal view (Fig. 51 A); gonopods elongated towards surstyli with two small projection towards phallus (Fig. 51 B); phallic guide with six spines dorsolaterally; subepandrial sclerite elongated in ventral view (Fig. 51 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA0EC1CD8D9EFF6FB8D1704.taxon	description	Description: MALE: Body length: 3.3 mm. Head. Flagellum short acuminate; yellow. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose; eyes touching for distance equal to 2.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 3 – 4 distinct pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part silver pollinose, slightly grayish also from the side. Scutellum brownish pollinose and with 6 – 8 weak bristles along the distal edge. Dorsocentral bristles indistinctly developed, dark, more visible on frontal part and numerous dark bristles around postpronotum. Halter, knob pale with brownish spot at tip, stem brown. Legs. Mid coxa with two long and one short bristles. Trochanters and femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees, basal 1 / 5 and distal 1 / 6 of tibia yellow (tibia otherwise brown), tarsal segments yellow dorsally, whitish yellow ventrally, last segment brown dorsally. Fore and mid trochanters with 1 – 2 dark anterior bristles. Hind trochanter without a keel, but with dark spines (22 – 24) in a row ventrobasally (Fig. 147 A). Ventroapical row of spines on fore femur 4 – 5 indistinct; 8 – 10 very short, black spines on mid femur; no spines on hind femur, hind femur with only 22 – 24 pale bristles equally spaced posteriorly, longer on distal half (longest up to as long as 1 – 1.2 times the width of hind tibia at the distal end) (Fig. 147 A). Subapical (distal) spines on first four tibiae present (medium sized); ventrobasal spines (1) present on fore and absent on mid femur. Hind metatarsus flattened almost as long as 2 – 5 combined, with dorsal bristles missing in the middle, arranged in rows along the edges; with scrub-like bristles ventrally. Pulvilli as long as or slightly longer than last tarsal segment. Wing. Length: 3.1 mm. Upper side of basal costal cell with one dark and long bristles. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 10 – 11 dark setulae on tegula, mostly along the distal edge. Abdomen. Dissected. tergite 1 silvery grey. Lateral spines on first tergite present, 6 – 8 brown bristles in a double row up to as long as the width of hind femur at distal tip in lateral view. Postabdomen in dorsal view: Dissected. Genitalia without dissection: Dissected. Genitalia. Genital capsule in dorsal view: surstyli rather symmetrical, broadened at base, slenderized after basal third. Left surstylus longer than right one (Fig. 51 A). Genital capsule in ventral view: gonopods unequal, left slightly longer than right one, elongated towards sternite, with two small projections extended towards phallus; lobes of hypandrium sclerotized, subepandrial sclerite elongated and covered with transverse streaks; phallic guide with six spines dorsolaterally (three left spines longer than others; the longest one twice as length as others) (Fig. 51 B); Genital capsule in lateral view: both surstyli gently curved toward sternite (Fig. 51 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 51 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA0EC1CD8D9EFF6FB8D1704.taxon	etymology	Etymology: This species is named in honour of Christine Lambkin, who collected the only known specimen and is a major contributor to Australian Diptera research. Chris collected or was involved in collecting 26 % of the known Australian Tomosvaryella specimens and 58 percent of the Tomosvaryella species treated in this paper, making her the top contributor to our knowledge of this genus in Australia (Supplementary file 2). This revision clearly could not have been completed without her input. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: Pine Hill, Cape Arid N [ational] P [ark], Balladonia Road, 33 ° 18 ’ S, 123 ° 22 ’ E, 130 m, mallee woodland, 31. x – 18. xi. 2003, C. Lambkin & J. Recsei, Malaise trap, JSS 15593 (1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA0EC1CD8D9EFF6FB8D1704.taxon	distribution	Distribution: Australia (Western Australia) (Fig. 126). Notes: The only known specimen was collected in mallee habitat (a dry, savanna-like habitat dominated by shrubby species of Eucalyptus). Based on the DNA sequencing, this species is genetically most similar to T. ngarrbekiota sp. nov. (7.7 – 13.6 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA6EC19D8D9EFF6FDA411FC.taxon	description	Figs 52 A – E, 126, 153 D	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA6EC19D8D9EFF6FDA411FC.taxon	diagnosis	Diagnosis: This species can be recognized by the shiny thorax and abdominal tergites; wide surstyli at basal two thirds in dorsal view (Fig. 52 A); both gonopods equal in height, with a finger-like projection towards surstyli in ventral view (Fig. 52 B); phallic guide broadened in the middle, with two protrusions ventromedially, pointed at apex in lateral view (Fig. 52 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA6EC19D8D9EFF6FDA411FC.taxon	description	Description: MALE: Body length: 2.9 – 3.2 mm. Head. Scape, pedicel and arista dark brown. Pedicel with two upper bristles; flagellum brown, gray pollinose and tapering. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance twice as the length of ocellar triangle. Occiput grey pollinose. Thorax. Postpronotum light yellow, with 4 – 5 long brown bristles in upper margin. Mesonotum (viewed obliquely from front) gray pollinose, rather shiny; scutum with scattered dark bristles at anterior supra-alar area, extended to posterior and uniseriate rows of intra-alar dark bristles. Postalar callus dark brown with 1 – 2 light brown bristles in the middle. Scutellum rather shiny, with 15 – 20 short pale bristles, scattered throughout it. Halter, knob pale, stem light brown, base dark. Legs. Fore coxa with 2 – 3 light brown bristles, mid coxa with four long dark bristles. Coxae, trochanters, femora dark, all femora shining ventrally, knees and basal 1 / 3 of tibiae yellow (tibiae otherwise dark brown), tarsomeres brown ventrally, dark dorsally. Hind trochanter silvery pollinose covered by some (8 – 12) scattered short brown bristles ventrally; two ventrobasal spines present on fore mid femur. Hind tibia with two rows of short brown bristle on anterior and posterior side. Hind metatarsus not flattened, covered by short dark bristles, as long as 2 – 4 combined; pulvilli shorter than last tarsal segment. Wing. Length: 3.0 mm. Upper side of basal costal cell with 2 – 3 long brown bristles. Fourth costal section 1.5 – 2 times as long as third costal section. Cross-vein r-m at middle of discal cell. 3 – 4 short dark setulae on tegula. Abdomen. Tergites 1 silvery pollinose with 5 – 9 different-sized lateral bristles; tergite 2 – 3 brown spots on dorsal margin, otherwise shiny; all tergites covered by scattered short pale bristles. Genitalia. Genital capsule in dorsal view: epandrium as long as wide (MLE: MWE = 1.0). Surstyli symmetrical, widened at base, moderately narrowed to apex (Fig. 52 A). Genital capsule in ventral view: both gonopods equal in height, with a finger-like projection towards surstyli, right one wider than left one; subepandrial sclerite long (Fig. 52 B); Genital capsule in lateral view: both surstyli wide, curved to sternite; phallic guide broadened in the middle, with two protrusions ventromedially, pointed at apex (Fig. 52 D – E); ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 52 C). FEMALE: Body length: 3.0 mm. As male except for the following characters. Frons, eyes separated; completely silver-grey pollinose on lower 1 / 3, wider in middle, shiny black only around ocellar triangle, depression in just before ocellar triangle. Enlarged ommatidia shining; pulvilli and claws longer than last tarsal segment. Lateral bristles on the tergite 1 shorter than male, tergite 2 – 3 without brown spots (4 – 8). Ovipositor. Straight in ventral view, straight and short in lateral view (piercer brown), reaching sternite 3; basal two thirds silvery brown pollinose. LP: LB = 3.5. LDP: LPP = 3.0. (Fig. 153 D). Examined material: HOLOTYPE: AUSTRALIA: Western Australia: Neerabup Lake National Park, 31 ° 38 ’ S, 115 ° 43 ’ E, 18 – 29. XII. 1999, J. & A. Skevington, C. Lambkin & P. Bouchard, Malaise trap, JSS 7642 (1 ♂, WAM); PARATYPES: AUSTRALIA: New South Wales: Culgoa National Park, 13.5 km SW Cawwell Homestead (CGN 1 M), 29 ° 7 ’ S, 146 ° 56 ’ E, Callitrus, 21. XII. 2009 – 30. I. 2010, B. Schiebaan, Malaise trap, CNC 576567, CNC 576577 (2 ♂, QM); Callitris, 30. I – 18. V. 2010, QM Team, B. Shiebaan & R. Ohlsen, Malaise trap, CNC 576602, CNC 576605, CNC 576609, CNC 576614 – 15 (5 ♂, QM), C. Lambkin, R. Olsen & B. Shieban, CNC 575141, CNC 575149 (2 ♂, USNM); Culgoa National Park, 8 km WNW Cawwell Homestead, Diemunga Lagoon (CGN 2 M), 29 ° 3 ’ S, 146 ° 60 ’ E, Coolibah, 22. XI – 21. XII. 2009, C. Lambkin, B. Shiebaan & N. Starick, Malaise trap, DB 9 (2 nd bottle), CNC 593680 – 81 (2 ♂, QM); Culgoa National Park, 8 km West – Northwest Cawwell Homestead, Diemunga Lagoon (CGN 2 M), 29 ° 3 ’ S, 146 ° 60 ’ E, Coolibah, 30. I – 18. V. 2010, C. Lambkin, R. Olsen & B. Shieban, Malaise trap, CNC 575123; CNC 575129; CNC 575135 (3 ♂, QM); Urila Rd, Tilembeya 14.2 km E Williamsdale, 35 ° 34 ’ S, 149 ° 17 ’ E, 804 m, in dry ravine, 20. XI – 19. XII. 2004, C. Lambkin, N. Starick, Malaise trap, ANIC Bulk sample 2559, JSS 50569 (1 ♀, ANIC); Queensland: Carnarvon, National Park Marlong Creek, 24 ° 56 ’ S, 147 ° 57 ’ E, 13. X. 2002, C. Chenard, sweep net, LEM _ 0016193 (1 ♂, LEM); Culgoa Floodplain National Park, 7 km Northwest of Toulby Gate (TOLB 7 M), 28 ° 57 ’ S, 146 ° 53 ’ E, 138 m, reedy swamp, 20. I – 19. III. 2010, C. Kelly & A. Coward, Malaise trap, 19269, CNC 575378 (1 ♂, QM); Culgoa Floodplain National Park, 7 km WNW Toulby Gate (TOLB 6 M), 28 ° 57 ’ S, 146 ° 52 ’ E, 138 m, Callitris, 19. III – 20. V. 2010, C. Kelly, A. Coward, Malaise trap, 19280, CNC 575029 (1 ♂, QM); Plevan Downs, 2.3 km South – Southwest of Arima (PD 2 M), 26 ° 32 ’ S, 142 ° 31 ’ E, 134 m, Gidgee, 13. I – 3. IV. 2008, R. Mackenzie, Malaise trap, 17284, CNC 595390 (1 ♂, QM); Plevna Downs, 2.3 km SSW Arima (PD 2 M), 26 ° 32 ’ S, 142 ° 31 ’ E; 28 ° 32 ’ S, 142 ° 31 ’ E, 134 m, Gidgee, 25. XI – 17. XII. 2008; 13. I – 3. IV. 2008, R. Mackenzie, Malaise trap, 17292; 17284, CNC 595265, CNC 595267, CNC 575360 (3 ♂, QM); Plevna Downs, 2.3 km South – Southwest Arima (PD 2 M), 28 ° 32 ’ S, 142 ° 31 ’ E, 134 m, Gidgee, 13. I – 3. IV. 2008, R. Mackenzie, Malaise trap, 17284, CNC 575359 (1 ♂, QM); Plevna Downs, 2.5 km WNW Homestead (PD 3), 26 ° 40 ’ S, 142 ° 35 ’ E, 133 m, Gidgee, 16. IX – 2. X. 2008, Starick, Lambkin & Mackenzie, Malaise trap, 16272, CNC 591860 (1 ♂, QM); Plevna Downs, Tompilly Hill base (PD 6 M), 26 ° 44 ’ S, 142 ° 39 ’ E, 187 m, Gidgee, 25. XI – 18. XII. 2008, R. Mackenzie, Malaise trap, 17329, CNC 576411, CNC 576413 (2 ♂, QM); 23 km SE Tamworth, 31 ° 13 ’ S, 151 ° 7 ’ E, 25. XI. 1982, D. S. Horning, Jr., JSS 9155 (1 ♂, ANIC); Murray River 50 miles west of Wentworth, 34 ° 3 ’ S, 141 ° 3 ’ E, 22. XI. 1967, A. Neboiss, JSS 8799 (1 ♂, MVMA); Culgoa National Park, 8 km WNW Cawwell Homestead, Diemunga Lagoon (CGN 2 M), 29 ° 3 ’ S, 146 ° 60 ’ E, Coolibah, 22. XI – 21. XII. 2009, C. Lambkin, B. Shiebaan, N. Starick, Malaise trap, DB 9 (2 nd bottle), 19302, CNC 593678 – 9 (2 ♂, CNC); Queensland: Lolworth National Park, site 2, 19 ° 50 ’ S, 146 ° 5 ’ E, 280 m, vinescrub on basalt, 13 – 14. XII. 2006, S. Wright, yellow pan trap, CNC 593654 (1 ♂, CNC); 23 km SE Tamworth, 31 ° 13 ’ S, 151 ° 7 ’ E, 26. XI. 1982, D. S. Horning, Jr., JSS 9148, JSS 9152 (2 ♂, ANIC); Culgoa National Park, 8 km West-Northwest of Cawwell Homestead, Diemunga Lagoon (CGN 2 M), 29 ° 3 ’ S, 146 ° 1 ’ E, 136 m, Coolibah, 21. XII. 2009 – 30. I. 2010, B. Schiebaan, Malaise trap, CNC 574750 – 1 (2 ♂, CNC); Carnarvon Station, near Piebald Spring (CN 1 M 1), 24 ° 50 ’ S, 147 ° 45 ’ E, 821 m, Eucalyptus / Callistemon, in rocky gully, 13. XII. 2010 – 15. VI. 2011, C. Zwick & C. Wilson, Malaise trap, CNC 595241, CNC 595257 (2 ♂, QM); Cudmore National Park (CM 1 M), 22 ° 54 ’ S, 146 ° 21 ’ E, 365 m, Eucalyptus woodland with Spinifex, 27. X. 2010 – 2. VIII. 2011, Lambkin, Starick & Bailey, Malaise trap, CNC 591851 (1 ♂, QM); Culgoa Floodplain National Park, 7 km Northwest of Toulby Gate (TOLB 7 M), 28 ° 57 ’ S, 146 ° 53 ’ E, 138 m, reedy swamp, 20. I – 19. III. 2010, C. Kelly & A. Coward, Malaise trap, CNC 575365 (1 ♂, QM); 12.7 km South-Southeast of Plevna Downs Homestead (PD 8 M), 26 ° 47 ’ S, 142 ° 39 ’ E, 145 m, sandy creek bank, 25. XI – 18. XII. 2008, R. Mackenzie, Malaise trap, CNC 593801 (1 ♂, QM); 204 km west Windorah, 25 ° 42 ’ S, 140 ° 53 ’ E, Malaise in dry creek bed, 4. IX. 1997, S. Winterton, J. & A. Skevington, JSS 8337 (1 ♂, QM); 91.2 km west Windorah, 25 ° 22 ’ S, 141 ° 47 ’ E, Malaise in dry creek bed, 31. VIII – 5. IX. 1997, S. Winterton, J. & A. Skevington, JSS 8343 (1 ♂, QM); South Australia: Sulphur Peninsular, Madigan Gulf, Lake Eyre N., 28 ° 22 ’ S, 137 ° 22 ’ E, 30. X. 1966, G. F. Gross, JSS 8707 (1 ♂, SAM); Koonchera Dune near Koonchera Waterhole, 26 ° 41 ’ S, 139 ° 30 ’ E, 2. IX. 1997, J. & A. Skevington, JSS 8429 (1 ♂, QM); SE corner Yumbarra CP, 31 ° 39 ’ S, 133 ° 22 ’ E, 27. IX. 1988, J. A. Forrest, JSS 8824 (1 ♂, SAM); Scorpion Springs CP along main North-South track, 35 ° 27 ’ S, 140 ° 53 ’ E, 15. XII. 1983, Museum party, JSS 8825 – 6 (2 ♂, SAM); Simpson Desert between Purni Bore & 54 km ENE on French Track, 26 ° 7 ’ S, 138 ° 37 ’ E, 8. X. 1987, J. A. Forrest, car net, JSS 8820 – 1 (2 ♂, SAM); Simpson Desert, 25 km west Poeppel Corner, 26 ° 7 ’ S, 138 ° 37 ’ E, 8. X. 1987, J. A. Forrest, at light, JSS 8823 (1 ♂, SAM); Western Australia: Cadda Road, 30 ° 24 ’ S, 115 ° 25 ’ E, 244 m, Eucalyptus Woodland, 20. IX – 10. X. 2003, C. Lambkin, N. Starick & J. Recsei, Malaise trap, ANIC BULK Sample 2177, JSS 50567 – 68; JSS 50570 (1 ♂, 2 ♀, ANIC); 23 km E. Marble Bar, 21 ° 11 ’ S 119 ° 57 ’ E, 222 m, Sandy wash, 15. V. 2003, F. D. Parker, M. E. Irwin, pan trap, CSIRO – SCHLINGER PILBARA EXPEDITION 19. iv. – 23. v. 2003, CNCD 6950, CNCD 6952 (2 ♂, ANIC); 158 km S Newman, 9 km N Kumarina Road House, 24 ° 38 ’ S, 117 ° 37 ’ E, 638 m, in wide sandy wash, 18 – 21. V. 2003; 24. IV – 7. V. 2003; 7 – 18. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15706, JSS 15709, JSS 15715 – 6, JSS 15722 – 3, JSS 15732, JSS 16416 (6 ♂, 2 ♀, CNC); 70 km S Newman on Great Northern Highway, 23 ° 54 ’ S, 119 ° 45 ’ E, 626 m, small wash with Eucalyptus, 24. IV – 6. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15881, JSS 15883 (1 ♂, 1 ♀, CNC); Cape Range National Park; Milyering Ranger House, 22 ° 2 ’ S, 113 ° 56 ’ E, 22 m, below cliff near Ficus, 28. IV – 11. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15655 – 6 (2 ♂, CNC); Karijini National Park, Hamersley-Mount Bruce Road, 22 ° 37 ’ S, 118 ° 21 ’ E, 755 m, dry rocky creek bed, open Eucalyptus scrub, 20 – 25. IV. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 15969 (1 ♂, ANIC); Karijini National Park, Juna Downs Road, 22 ° 40 ’ S, 118 ° 26 ’ E; 22 ° 44 ’ S, 118 ° 25 ’ E, 797 m; 798 m, in open mulga with flowering Ptilotus; dry Turee Creek, grassy open Eucalyptus scrub, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16002, JSS 16097 (1 ♂, 1 ♀, ANIC); Millstream-Chichester National Park near Crossing Pool, 21 ° 35 ’ S, 117 ° 4 ’ E, 276 m, over creek, Melaleuca riverine forest, 28. IV – 5. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16189 (1 ♂, ANIC); Neerabup Lake National Park, 31 ° 38 ’ S, 115 ° 43 ’ E, 18 – 29. XII. 1999; 8 – 18. XII. 1999, J. & A. Skevington, C. Lambkin & P. Bouchard, Malaise trap, JSS 7635 – 6, JSS 7639 – 40, JSS 7647 – 9, JSS 7652 (8 ♂, QM); base of “ The Governor ” on Great Northern Highway, 23 ° 3 ’ S, 118 ° 50 ’ E, 722 m, dry wash in wooded area, 23. IV – 6. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15688, JSS 15690 – 2, JSS 15694 – 7 (1 ♂, 7 ♀, CNC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA6EC19D8D9EFF6FDA411FC.taxon	etymology	Etymology: The specific name is derived from the Latin word ‘ lanterna’, meaning shine, referring to the shiny thorax and abdominal tergites.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA6EC19D8D9EFF6FDA411FC.taxon	distribution	Distribution: Australia (New South Wales, Queensland, South Australia, Western Australia) (Fig. 126). Notes: This is a widespread common species in the Australian outback. It apparently does not use hilltops as a mating site as it appears to be common enough not to need to employ a landmark mating system. Intraspecific genetic distance ranges from 0.0 % to 2.5 %. It is genetically most similar to T. quadrata sp. nov. (8.2 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA5EC17D8D9EEBBFBB713BC.taxon	description	Figs 53 A – E, 127, 151 A	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA5EC17D8D9EEBBFBB713BC.taxon	diagnosis	Diagnosis: This species can be recognized by a distinct ventrobasal protuberance on the hind femur (Fig. 151 A); broadened surstyli and tapering in basal third with patch of bristles in its medial in dorsal view (Fig. 53 A); subepandrial sclerite covered with transverse streaks in ventral view; gonopods flat, right one with a lobe extended towards surstyli (Fig. 53 B); phallic guide with three to five spines dorsally before apex.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA5EC17D8D9EEBBFBB713BC.taxon	description	Description: MALE: Body length: 2.3 – 2.6 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black, lower part silvery pollinose; eyes touching for distance equal to 1 – 1.5 times ocellar triangle. Occiput silvery pollinose, slightly less so on upper half. Thorax. Postpronotum pale brown, with 2 – 3 whitish bristles. Mesonotum (viewed obliquely from front) subshining black, anterior part silvery pollinose; also silvery from the side. Scutellum gray pollinose, without bristles. Dorsocentral bristles weakly developed, pale, somewhat longer on frontal part. Halter, knob pale, stem dark black. Legs. Trochanters and base of femora brown, femora dark brown, shining ventrally (hind femur also posteriorly), knees, basal 1 / 6 of tibiae, tarsal segments yellow (otherwise brown), last segment dorsally brown. Fore femur with two distinct ventrobasal spines, mid femur with 4 – 6 very short, black ventroapical spines; no spines on hind femur, only 3 – 4 flattened thick bristles on keel. Hind femur with a distinct ventrobasal protuberance bearing three to six spines (Fig. 151 A). Hind trochanter smooth with 4 – 5 indistinct dark bristles on ventral side. Subapical (distal) spines on first four tibiae absent. Hind metatarsus slightly flattened, first tarsomere longer than 2 – 4 combined. Pulvilli slightly shorter than last tarsal segment. Wing. Length: 2.6 – 2.7 mm. Upper side of basal costal cell with one long distinct bristle. Fourth costal section 2.5 – 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. Setulae on tegula absent. Abdomen. Viewed obliquely from front tergites brownish pollinose, hind margin silvery gray on tergite 1, sides completely silvery on tergites 4 and 5. Dispersed short dark bristles absent. Lateral bristles on first tergite: 2 – 3, pale brown, as long as width of hind femur at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 as long as ST 8. Genitalia without dissection: ST 8 appears large, mainly dark brown and forming a right-angle corner (90 degrees) in lateral view at the lower end of membranous area; membranous area slit-like, vertical, distal part of epandrium paler and surstylus tips whitish, surstyli relatively small. Genitalia. Genital capsule in dorsal view: epandrium brown, surstyli light brown. Epandrium wider than long (MLE: MWE = 0.9), ca. 0.5 times as long as ST 8, cerci symmetrically placed, small; ST 8 with slit-like membranous area. Genital capsule in dorsal view: surstyli, both surstyli broadened at base, pointing tips at its apical inner corner, medially covered with patch of long bristles (Fig. 53 A). Genital capsule in ventral view: Subepandrial sclerite covered with transverse streaks; gonopods asymmetrical and flat with right gonopod bearing a lobe pointing towards surstyli, phallic guide with a membranous projection to the side (Fig. 53 B); phallus trifid, branches short (Fig. 53 B). Genital capsule in lateral view: both surstyli broadened in basal two thirds, tapering in apical third (Fig. 53 D – E). Phallic guide with three to five spines dorsally before apex, phallus with three ejaculatory ducts; ejaculatory apodeme flat, sperm pump elongated, linear (Fig. 53 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA5EC17D8D9EEBBFBB713BC.taxon	etymology	Etymology: From the Latin latus, broad or wide, in reference to the flattened gonopods of the male genitalia. Examined material: HOLOTYPE: AUSTRALIA: Victoria: Wyperfeld NP, East Lookout, 35 ° 36 ’ S, 142 ° 7 ’ E, 21. x. 2014, J. H., A. W., & A. M. Skevington, A. D. Young, A. Wyatt, & S. Namek, CNC 373275 (1 ♂, QM); PARATYPES: AUSTRALIA: Victoria: Wyperfeld NP, East Lookout, 35 ° 36 ’ S, 142 ° 7 ’ E, 22. x. 2014, J. H., A. W., & A. M. Skevington, A. D. Young, A. Wyatt, & S. Namek, CNC 373438 (1 ♂, CNC); South Australia: vehicle net between Wynbring Rocks RS & Malbooma RS, 30 ° 33 ’ S, 133 ° 32 ’ E, 2. x. 1988, J. A. Forrest, JSS 8819 (1 ♂, SAM); New South Wales: Culgoa National Park, 8 km West-Northwest Cawwell Homestead, Diemunga Lagoon, 29 ° 3 ’ S, 146 ° 60 ’ E, Coolibah, 30. i – 18. v. 2010, C. Lambkin, R. Olsen & B. Shieban, Malaise trap, CNC 575145 – 6 (2 ♂, QM); Culgoa National Park, 8 km West-Northwest of Cawwell Homestead, Diemunga Lagoon, 29 ° 3 ’ S, 146 ° 1 ’ E, 136 m, Coolibah, 21. xii. 2009 – 30. i. 2010, B. Schiebaan, Malaise trap, CNC 574747 – 8, CNC 574753 – 4 (4 ♂, QM); Northern Territory: 56 km S by E of Alice Springs, 23 ° 42 ’ S, 133 ° 53 ’ E, 3. x. 1978, D. H. Colless, Malaise trap, JSS 9250 (1 ♂, ANIC); 4. x. 1978, D. H. Colless, Malaise trap, JSS 9251 (1 ♂, ANIC); Queensland: 15.1 km Southwest of Currawinya National Park Headquarters, 28 ° 55 ’ S, 144 ° 26 ’ E, 131 m, Eremophila scrub, 18. ii – 4. iii. 2009, A. Townsend & J. Burke, Malaise trap, CNC 574711 (1 ♂, QM); 204 km west Windorah, 25 ° 42 ’ S, 140 ° 53 ’ E, Malaise in dry creek bed, 4. ix. 1997, S. Winterton, J. & A. Skevington, JSS 8339 (1 ♂, QM); Carbrook, Logan River Delta, 27 ° 40 ’ S, 153 ° 15 ’ E, mangrove, 25. x. 2002, M. Földvari, JSS 13226 (1 ♂, HNHM); Culgoa Floodplain National Park, 11.8 km N Headquarters, 28 ° 54 ’ S, 147 ° 8 ’ E, 151 m, Brigalow Black box, 17. xii. 2009 – 20. i. 2010, C. Kelly & A. Coward, Malaise trap, CNC 574517 (1 ♂, QM); Currawinya National Park, 15.1 km Southwest of Headquarters, 28 ° 55 ’ S, 144 ° 26 ’ E, 131 m, Eremophila, 12 – 26. ix. 2008, Lambkin, Starick & Townsend, Malaise trap, CNC 574664 (1 ♂, QM); Port Douglas, North Queensland, 16 ° 29 ’ S, 145 ° 28 ’ E, 9. vii. 1971, Z. Liepa, JSS 9207 (1 ♂, ANIC); Western Australia: ~ 3 km N. of Denham, 25 ° 55 ’ S, 113 ° 32 ’ E, hilltop, 12. XII. 1999, J. Skevington, CNC 944146 (1 ♂, CNC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA5EC17D8D9EEBBFBB713BC.taxon	distribution	Distribution: Australia (New South Wales, Northern Territory, Queensland, South Australia, Victoria, Western Australia) (Fig. 127). Notes: This is a widespread species that has been captured hilltopping. Most specimens have been collected in mallee and other scrubby desert areas but two have been collected in very unexpected habitats. The specimen from Carbrook was in mangroves and the Port Douglas specimen would have most likely been in mangroves or coastal rainforest. These unusual outliers are morphologically consistent with the other specimens and we believe that they are the same species. To test this, we need fresh specimens from these areas that we can obtain DNA from. This species is closest to Tomosvaryella pterygia sp. nov., see the notes under the latter.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFABEC15D8D9EB7BFC311464.taxon	description	Figs 54 A – E, 128	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFABEC15D8D9EB7BFC311464.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a keel and 2 – 3 short dark bristles proximally; elongated surstyli, left one flattened at apex in dorsal view (Fig. 54 A); wide gonopods at base, pointed at apex, with a distinct projection on inner margin, their projections with small finger-like process in the middle; phallic guide with some dorso- dorsolateral spines, four same size spines longer than others (Fig. 54 B, D – E); one of ejaculatory ducts with a short spine (Fig. 54 D).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFABEC15D8D9EB7BFC311464.taxon	description	Description: MALE: Body length: 3.1 – 3.3 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 3 – 4 distinct pale bristles. Mesonotum (viewed obliquely from front) silvery pollinose, anterior part more grey, grayish also from the side. Scutellum silvery brown pollinose and with 10 – 12 bristles along distal edge. Dorsocentral bristles short, dark, longer in frontal half, numerous pale bristles around postpronotum present. Halter, knob pale, slightly brownish, stem brown. Legs. Mid coxa with 2 – 3 long dark bristles. Trochanters brown, femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 4 of tibiae yellow (tibiae otherwise brown), tarsal segments yellow-brown dorsally, yellow ventrally, last segment darker. Hind trochanter with 45 degree keel, and 2 – 3 short dark bristles proximally; ventrobasal spines absent on fore and mid femur. Hind femur without ventroapical spines, except 8 – 10 equally spaced bristles posteroventrally, longer on distal half, longest up to 1 / 2 the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres moderately flattened, hind metatarsus almost as long as 2 – 4 combined, with semi-erect dark bristles dorsally. Pulvilli shorter than last tarsal segment. Wing. Length: 3.4 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m at middle of discal cell (can be more distal). 2 – 3 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites subshiny brown-black, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (spot on t 5 larger). Dispersed strong dark bristles on all tergites present, longest on tergite 5 in length up to 1 / 4 the width of hind tibia at distal end. Lateral bristles on first tergite present, 4 – 5 dark bristles up to as long as ¾ of hind femur’s width at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1.2 – 1.3 times as long as ST 8. Genitalia without dissection: ST 8 small to medium sized, round, globular, brown and with distinct bristles, otherwise with velvet-like coverage; membranous area is slit-like, mostly directed posteriorly; epandrium brown, surstyli brown. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.3). Surstyli elongated, broaden at base, left surstylus constricted after basal fourth and flattened at its tip, right surstylus curved towards left surstylus (Fig. 54 A). Genital capsule in ventral view: both gonopods widened at base, extended towards surstyli, pointed at apex, equal in height, gonopods with a distinct projection on inner margin, both projections with an upwards small finger-like process in the middle; phallic guide with 5 – 6 dorso- dorsolateral spines, four same size spines longer than others; subepandrial sclerite widened in the middle (Fig. 54 B); Genital capsule in lateral view: both surstyli straight, moderately broadened to apex, curved towards sternite apically; phallus with three ejaculatory ducts, one of ejaculatory ducts with a short spine (Fig. 54 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 54 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFABEC15D8D9EB7BFC311464.taxon	etymology	Etymology: The name is derived from the Latin latus, broad or wide, in reference to the flattened tip of the left surstylus. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Barakula State Forest No. 302, Summit Round Mountain, 26 ° 22 ’ S, 150 ° 58 ’ E, hilltop, 8. I. 2000, J. & A. Skevington & M. Mathieson, JSS 7546 (1 ♂, QM); PARATYPES: AUSTRALIA: Queensland: same data as holotype, JSS 7547 – 51, JSS 7557, (6 ♂, QM); hilltop, 21 – 25. II. 2000, M. Mathieson, JSS 9326 (1 ♂, CNC); Western Australia: Mount Magnet, Mount Waramboo, 28 ° 2 ’ S, 117 ° 49 ’ E, hilltop, 14. XII. 1999, J. Skevington, JSS 7469 (1 ♂, CNC); ~ 3 km North of Denham, 25 ° 55 ’ S, 113 ° 32 ’ E, hilltop, 12. XII. 1999, J. Skevington, JSS 7377, JSS 7393 (2 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFABEC15D8D9EB7BFC311464.taxon	distribution	Distribution: Australia (Queensland, Western Australia) (Fig. 128). Notes: All specimens of this species have been collected at hilltops. Given the distribution it is presumably widespread but rare. Intraspecific genetic distance ranges from 0.0 % to 2.4 %. It is genetically most similar to T. cona sp. nov. (4.4 – 10.7 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA9EC13D8D9EB03FAB812DC.taxon	description	Figs 55 A – E, 128	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA9EC13D8D9EB03FAB812DC.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a keel covered with small teeth along its edge; elongated surstyli, widened at the base and bent toward each other at the apex in dorsal view (Fig. 55 A); both surstyli bent towards sternite in apical third in lateral view (Fig. D – E); subepandrial sclerite with fin-shaped projection in its center, both gonopods elongated and pointed at the apex, hypandrial apodeme distinct in ventral view (Fig. 55 C).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA9EC13D8D9EB03FAB812DC.taxon	description	Description: MALE: Body length: 2.5 mm. Head. Scape, pedicel dark brown, flagellum light yellow, pollinose. Pedicel with three short upper bristles. Flagellum tapering. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3 with scattered short dark bristles. Thorax. Postpronotum pale, with 2 – 3 long pale bristles. Mesonotum (viewed obliquely from front) silvery pollinose, anterior part more grey, grayish also from the side. Scutellum silvery brown pollinose with 2 – 4 short bristles along distal edge. Halter, knob and tip light brown. Legs. Mid coxa with 3 – 4 different-size dark bristles. Trochanters light brown, femora brown, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae light yellow (tibiae otherwise light brown), tarsal segments bright yellow completely. Hind trochanter with a keel covering small teeth along its edge in a row; ventrobasal spines present on fore and mid femora. Hind femur without ventroapical spines. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres not flattened, hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 2.4 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 1 – 2 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brown, tergite 1 silvery grey. Lateral bristles on first tergite present, 4 – 5 long dark bristles up to as long as hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.3). widened at the base, elongated, moderately narrowed to apex, pointed, bent to each other at the apex (Fig. 55 A). Genital capsule in ventral view: both gonopods extended towards surstyli, pointed at apex with a distinct projection on inner margin; phallic guide with 3 – 4 short dorso- dorsolateral spines, one is longer; subepandrial sclerite widened, with a fin-shaped projection in its center, hypandrial apodeme distinct (Fig. 55 B); Genital capsule in lateral view: both surstyli straight in basal two thirds, then bent towards sternite in apical third (Fig. 55 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 55 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA9EC13D8D9EB03FAB812DC.taxon	etymology	Etymology: This species is named after the type locality. Eight species of Tomosvaryella have been collected in this reserve and the only known specimen of this species is from there. Examined material: HOLOTYPE: AUSTRALIA: New South Wales: Ledknapper National Reserve, 7.3 km ENE Beulah Headquarters (LDN 2 M), 29 ° 21 ’ S 146 ° 13 ’ E, E. melanopholia, Spinifex, 10. XII. 2009 – 18. III. 2010, S. O’Sullivan, K. Taylor, Malaise trap, CNC 576517 (1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFA9EC13D8D9EB03FAB812DC.taxon	distribution	Distribution: Australia (New South Wales) (Fig. 128). Notes: Based on the shape of surstyli in dorsal view, this species is close to T. prolata sp. nov. It differs by the shape of surstyli, which are bent towards the sternites in the apical third in lateral view, with a fin-shaped projection in the middle of the subepandrial sclerite and a distinct hypandrial apodeme in ventral view (Fig. 55 A – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFAFEC11D8D9ED9BFA031110.taxon	description	Figs 56 A – E, 129, 150 F	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFAFEC11D8D9ED9BFA031110.taxon	diagnosis	Diagnosis: This species can be recognized by the mid tibia with one distinct and long apical spur, as long as the mid metatarsus (Fig. 150 F); surstyli elongated, apical margin flattened in dorsal view (Fig. 56 A); both gonopods extended towards surstyli with a small triangular projection in its middle; phallic guide with 3 – 6 dorso-dorsolateral spines before apex in ventral view (Fig. 56 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFAFEC11D8D9ED9BFA031110.taxon	description	Redescription: MALE: Body length: 2.6 mm. Head. Scape, pedicel and arista drak brown. Pedicel with 2 – 3 upper bristles; flagellum brown, gray pollinose and tapering. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance as long as the length of ocellar triangle. Occiput grey pollinose covered by scattered short dark bristles. Thorax. Postpronotum light yellow, with 4 – 5 long brown bristles in upper margin. Mesonotum (viewed obliquely from front) gray pollinose; scutum with scattered dark bristles at anterior supra-alar area, extended to posterior and uniseriate rows of intra-alar same sized dark bristles. Postalar callus dark brown with 2 – 3 long, as long as bristles in scutum, light brown bristles in the middle. Scutellum silvery pollinose and with 25 – 30 long, as long as scutum bristles, dark bristles, scattered throughout it. Halter, knob pale, stem light brown, base dark. Legs. Mid coxa with 3 – 4 long dark brown bristles. Coxae, trochanters, femora dark, all femora shining ventrally, knees and basal 1 / 3 of tibiae yellow (tibiae otherwise dark brown), tarsomeres brown ventrally, dark dorsally. Mid trochanter with 4 – 8 short dark bristles ventrally. Hind trochanter silvery pollinose covered by some (10 – 15) scattered short brown bristles ventrally; two ventrobasal spines absent on fore mid femur. Hind tibia with two rows of short brown bristle on anterior and posterior side. Mid tibia with one distinct and long apical spur, as long as mid metatarsus (Fig. 150 F). Hind tarsomeres slightly flattened to one side, covered by short dark brown bristles, hind metatarsus as long as 2 – 4 combined; pulvilli shorter than last tarsal segment. Wing. Length: 2.7 mm. Upper side of basal costal cell with one long and two short dark brown bristles. Fourth costal section 1.5 – 2 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2 – 3 short dark setulae on tegula. Abdomen. Tergites 1 silvery pollinose with 15 – 17 different-sized lateral bristles; tergite 2 – 5 brown pollinose, covered by scattered short dark bristles. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.6). Surstyli symmetrical, left surstylus longer than right one, broadened at base, then strongly constricted, elongated to apex, apical margin flattened (Fig. 56 A). Genital capsule in ventral view: both gonopods equal in height, extended towards surstyli with a small triangular projection in its middle; phallic guide with 3 – 6 dorso-dorsolateral spines before apex, one is longer than others; subepandrial sclerite small (Fig. 56 B); Genital capsule in lateral view: both surstyli curved to sternite, widened in the middle (Fig. 56 D – E); ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 56 C). FEMALE: unknown. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Brisbane, 27 ° 28 ’ S, 153 ° 1 ’ E, VI. 1904, JSS 4150 (1 ♂, ZMAN); Non-type material examined: New South Wales: Nelligen, 35 ° 38 ’ S, 150 ° 8 ’ E, (tidal flat), 1. II. 1973, D. H. Colless, JSS 8499 (1 ♂, ANIC). Distribution: Australia (New South Wales, Queensland) (Fig. 129). Notes: It appears that Perkins only based this species on one specimen, so this is the holotype. The New South Wales specimen was taken on a tidal flat so efforts to find this species should likely focus on mangrove habitats.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFADECEFD8D9EED7FF6311A4.taxon	description	Figs 57 A – D, 127	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFADECEFD8D9EED7FF6311A4.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a large triangular protuberance covered with numerous dark bristles; both surstyli strongly curved, hook- shaped in dorsal view (Fig. 57 A); gonopods equal, elongated towards surstyli in ventral view; subepandrial sclerite wide (Fig. 57 B); phallic guide with 5 – 6 dorso-dorsolateral spines (one is longer and sinuous-shaped) (Fig. 57 B – D).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFADECEFD8D9EED7FF6311A4.taxon	description	Description: MALE: Body length: 3.2 – 3.5 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 – 2.5 times the length of ocellar triangle. Occiput silvery pollinose, slightly less so on upper 1 / 3. Thorax. Postpronotum pale, with 10 + pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum silvery pollinose and with 7 – 9 bristles along the distal edge. Dorsocentral bristles distinct, dark, somewhat longer along frontal edge, numerous dark bristles around postpronotum present. Halter, knob pale, stem brown. Legs. Mid coxa with 3 long, broad, dark bristles. Trochanters and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise dark brown), tarsal segments dark brown, ventrally yellow, last segment darker. Hind trochanter with a large triangle in posterior view, covered with numerous dark bristles; ventrobasal spines absent on fore and mid femora. Ventroapical row of spines on fore femur absent; 12 – 14 small, black spines on mid femur; hind femur without ventroapical spines, except 12 – 14 bristles posteroventrally, longer and closer to each other on distal half, longest up to 1 / 3 of the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present (very short). Hind tibia bent in posterior view. Hind tarsomeres slightly flattened, metatarsus as long as 2 – 4 combined, ventrally with scrub-like bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 3.5 mm. Upper side of basal costal cell with one long dark bristle. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m distinctly distal to middle of discal cell. 2 – 3 distinct brown setulae on tegula. Abdomen. Viewed obliquely from front tergites slivery pollinose, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (larger on tergite 5). Dispersed short dark bristles on all tergites, longest on tergite 5 up to 1 / 3 the width of hind tibia at distal end. Lateral bristles on first tergite present, 5 – 6 dark bristles up to as long as hind femur’s width at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 2.3 – 2.5 times as long as ST 8. Genitalia without dissection: ST 8 medium sized, round in dorsal view, dark brown and with some distinct bristles, otherwise velvet-like coverage; membranous area rounded elongate, forming a ridge ventrally on ST 8, mostly directed posteriorly; epandrium dark brown, surstyli yellow. Genitalia. Genital capsule in dorsal view: epandrium slightly longer than wide (MLE: MWE = 1.07). Surstyli broadened in basal fourth and rectangular-shaped then strongly curved to each other, hookliked shaped, right surstylus smaller than left one (Fig. 57 A). Genital capsule in ventral view: gonopods elongated towards surstyli, tapering at apex, equal in height; subepandrial sclerite wide, egg-like shaped (Fig. 57 B); phallic guide with 5 – 6 dorso-dorsolateral spines (one is longer and sinuous-shaped). Genital capsule in lateral view: both surstyli curved towards sternite (Fig. 57 C – D). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFADECEFD8D9EED7FF6311A4.taxon	etymology	Etymology: This species carries the genus name given to the monotypic mound-building bird, Malleefowl (Leipoa ocellata). Leipoa is a Finnish word that refers to the alternating pressing motion with the two front paws of a cat, a behaviour like that of the Malleefowl as they tend to their mounds. All the specimens of this new pipunculid species were collected in in old growth mallee habitat where Malleefowl were plentiful. Mallee is a dry, semi-open habitat dominated by multi-stemmed Eucalyptus under 10 m in height. Examined material: HOLOTYPE: AUSTRALIA: New South Wales: Yathong Nature Res [erve], Malleefowl conservation area, 32 ° 35 ’ S, 145 ° 24 ’ E, 155 m, across sandy track in closed low mallee, 4 – 25. X. 2003, C. Lambkin & N. Starick, Malaise trap, JSS 16053 (1 ♂, ANIC); PARATYPES: AUSTRALIA: New South Wales: same data as holotype, JSS 16051 – 2, JSS 16054 (3 ♂, ANIC); South Australia: Gluepot Reserve, 33 ° 47 ’ S 140 ° 13 ’ E, 1. IX. 2004, J. Skevington, CNCD 1319 – 21 (2 ♂, CNC, 1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FFADECEFD8D9EED7FF6311A4.taxon	distribution	Distribution: Australia (New South Wales, South Australia) (Fig. 127). Notes: All known specimens have been collected in mallee reserves that contain significant old growth mallee areas. This species is genetically most similar to T. cona sp. nov. (3.1 – 7.7 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF53ECEFD8D9ED43FCAD17A8.taxon	description	Figs 58 A – E, 127, 149 C	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF53ECEFD8D9ED43FCAD17A8.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a pronounced hump ventrobasally covered by crest-like bristles and a few short bristles ventroapically (Fig. 149 C); surstyli slender in dorsal view (Fig. 58 A); phallic guide with 5 – 6 dorso-dorsolateral spines; two long membranous sheaths associated with the ejaculatory ducts; hypandrium with a small projection and a small hairy membranous sheath (Fig. 58 B, D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF53ECEFD8D9ED43FCAD17A8.taxon	description	Description: MALE: Body length: 2.8 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 4 – 5 pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum brownish pollinose and with 4 – 6 weak bristles along the distal edge. Dorsocentral bristles distinct, dark, somewhat longer along frontal edge, numerous long dark bristles around postpronotum. Halter, knob pale, stem brown. Legs. Mid coxa with 2 – 3 dark bristles. Trochanters and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 5 of tibiae yellow (tibiae otherwise dark brown), tarsal segments yellow-brown, ventrally somewhat paler, last segment darker. Hind trochanter with a pronounced little hump and crest-like bristles, distal part with 1 – 2 short spines (Fig. 149 C); ventrobasal (one) long bristle on fore and mid femora. Ventroapical row of indistinct spines on fore femur 3 – 4; 8 – 10 small, black spines on mid femur; hind femur without ventroapical spines, except 12 – 14 equally spaced bristles posteroventrally, longer on the distal half, in length up to 0.8 times the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present (distinct, longer on mid tibia). Hind metatarsus flattened and almost as long as 2 – 4 combined and with strong, thick bristles ventrally. Pulvilli as long as last tarsal segment. Wing. Length: 3.0 mm. Upper side of basal costal cell with one long dark bristle. Fourth costal section 2.5 times as long as third costal section. Cross-vein r-m at middle of discal cell. 3 – 4 small brown setulae on tegula. Abdomen. Dissected. tergite 1 and anterior 2 / 3 of tergite 2 silvery grey. Lateral bristles on first tergite present, 4 – 5 pale bristles up to 1.5 times as long as hind femur’s width at base. Postabdomen in dorsal view: Dissected. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.3). Surstyli symmetrical, broadened at base, slender and elongated, both curved to each other in middle, left surstylus longer than right one (Fig. 58 A). Genital capsule in ventral view: phallus with three short ejaculatory ducts and two long membranous sheath covered by small hair; gonopods elongated towards surstyli, equal in height, right is slightly wider than left one, subepandrial sclerite long, hypandrium with a small projection towards gonopod and a small hairy membranous sheath (Fig. 58 B). Genital capsule in lateral view: both surstyli curved towards sternite, more broadened at middle; phallic guide with 5 – 6 lateral spines (the longest one twice as length as others) (Fig. 58 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 58 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF53ECEFD8D9ED43FCAD17A8.taxon	etymology	Etymology: The name lophia is Greek for a mane, crest, comb, tuft or ridge and refers to the crest-like bristles on the hind trochanter. Specimen examined: HOLOTYPE: AUSTRALIA: Western Australia: Heinsmen Rock, M [oun] t Ragged Track, 30 ° 7 ’ S, 123 ° 28 ’ E, 145 m, woodland near drying pools, 30. x – 18. xi. 2003, C. Lambkin & J. Recsei, Malaise trap, JSS 16241 (1 ♂, WAM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF53ECEFD8D9ED43FCAD17A8.taxon	distribution	Distribution: Australia (Western Australia) (Fig. 127).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF51ECEAD8D9EFF6FDA41694.taxon	description	Figs 59 A – E, 130, 157 A	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF51ECEAD8D9EFF6FDA41694.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having 2 – 3 short bristles ventrobasally; elongated surstyli in dorsal view (Fig. 59 A); widened gonopods with inwards extended edge on inner margin in ventral view (Fig. 59 B); phallic guide long without spines; one of the ejaculatory ducts with a distinct and long downwards spine in lateral view (Fig. 59 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF51ECEAD8D9EFF6FDA41694.taxon	description	Description: MALE: Body length: 2.2 – 2.5 mm. Head. Scape, pedicel and arista dark brown. Pedicel with 2 – 3 upper bristles; flagellum brown, gray pollinose and tapering. Face silvery pollinose. Frons distinctly silver pollinose; eyes touching for distance equal the length of ocellar triangle. Occiput grey pollinose, covered by short dark bristles. Thorax. Postpronotum light yellow, with 2 – 4 brown bristles in upper margin. Mesonotum (viewed obliquely from front) silvery-brown pollinose, anterior part greyer; scutum with scattered dark bristles at anterior supra-alar area, extended to posterior and uniseriate rows of intra-alar bristles. Postalar callus dark brown with 2 – 3 short dark bristles in the middle. Scutellum silvery pollinose and with a few (6 – 8) short dark bristles along the distal edge. Halter, knob pale, stem light brown, base dark. Legs. Mid coxa with 3 – 4 dark bristles, 1 – 2 bristles longer than others. Coxae, trochanters, femora dark, hind femur shining ventrally, knees and basal 1 / 4 of tibiae yellow (tibiae otherwise dark brown), tarsomeres light brown ventrally and dark dorsally. Hind trochanter silvery pollinose, with 2 – 3 short bristles ventrobasally; two ventrobasal spines present on fore and absent on mid femur. Hind femur with two rows ventroapical spines. Hind tibia with two rows of short black bristle on anterior and posterior side. Hind tarsomeres not flattened, covered by short dark bristles dorsally and brown bristles ventrally, last hind tarsomere not rounded as long as 3 – 4 combined; pulvilli shorter than last tarsal segment. Wing. Length: 2.2 mm. Upper side of basal costal cell with one dark brown bristle. Fourth costal section 1.5 – 2 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2 – 3 short light brown setulae on tegula. Abdomen. Tergites 1 silvery pollinose with 4 – 8 long brown bristles; tergite 2 – 5 covered by scattered dark bristles. Syntergosternite 8 enlarged, dark brown and grey-brown pollinose. Membranous area long and broad in middle. Genitalia. Genital capsule in dorsal view: epandrium as long as wide (MLE: MWE = 1.0). Surstyli elongated (somewhat rectangular-shaped), curved to each other at apex; left surstylus slightly longer (Fig. 59 A). Genital capsule in ventral view: both gonopods widened, right gonopod longer; both with an extended inwards edge on inner margin; subepandrial sclerite long and distinct with patch of short bristles basally (Fig. 59 B); Genital capsule in lateral view: both surstyli curved towards sternite, right surstylus widened at basal half, then narrowed, left surstylus widened in basal third, strongly narrowed, one of the ejaculatory ducts with a distinct downwards spine; phallic guide long, bent towards surstyli in apical third and pointed, without spines (Fig. 59 D – E); ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 59 C). FEMALE: Body length: 2.2 – 2.3 mm. As male except for the following characters. Frons, eyes separated, wider in middle; completely silver-grey pollinose on lower 1 / 3, shiny black only around ocellar triangle, deep depression in just before ocellar triangle. Enlarged ommatidia silvery shining; Upper side of basal costal cell with one long dark and 2 – 3 short bristles; pulvilli and claws as long as last tarsal segment. Tergites 4 – 6 with two grey lateral spots of gray pollinosity. Ovipositor. Straight in ventral view, rather curved towards sternite in lateral view (piercer dark at its base and brown in apical two thirds), reaching sternite 1; base silvery brown pollinose. LP: LB = 2.5. LDP: LPP = 4.2. (Fig. 157 A).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF51ECEAD8D9EFF6FDA41694.taxon	etymology	Etymology: The specific name is derived from the Latin word ‘ luma’, meaning thorn, referring to the distinct thorn on one of the ejaculatory ducts. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: Karijini National Park, Juna Downs Road, 22 ° 40 ’ S, 118 ° 26 ’ E, 797 m, in open mulga with flowering Ptilotus, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16003 (1 ♂, WAM); PARATYPES: AUSTRALIA: Australian Capital Territory: Black Mountain Reserve, 35 ° 16 ’ S, 149 ° 6 ’ E, 4. I. 1980, Z. Liepa, JSS 9185 (1 ♂, ANIC); Black Mountain, 35 ° 16 ’ S, 149 ° 6 ’ E, 21. II. 1980; 29. XII. 1979; 30. I. 1980; 5. II. 1980, D. H. Colless, Malaise trap, JSS 8508, JSS 8518 – 20 (4 ♂, ANIC); 29. II. 1968, I. F. B. Common, light trap, JSS 8503 (1 ♂, ANIC); 2. II. 1967, Z. Liepa, JSS 9182 (1 ♂, ANIC); Canberra, Black Mountain, 35 ° 16 ’ S, 149 ° 6 ’ E, 11 – 19. I. 1999, G. Gibson, sweep net, CNCD 4468 (1 ♂, CNC); Black Mountain, 35 ° 16 ’ S, 149 ° 6 ’ E, 11 – 13. I. 1980; 15. I. 1980; 18. I. 1980; 21 – 22. IV. 1980; 21. II. 1980; 23. I. 1980; 28 – 30. III. 1980; 29. XII. 1979; 30. XII. 1979; 5 – 6. I. 1980, D. H. Colless, Malaise site 1, JSS 8678, JSS 8681 – 3, JSS 8685, JSS 8689, JSS 8691, JSS 8693 – 5 (10 ♂, ANIC); Black Mountain, Canberra, 35 ° 16 ’ S, 149 ° 6 ’ E, 18 – 23. II. 1985, W. W. Middlekauff, flight interception trap, JSS 8452 (1 ♂, CAS); Blundells Creek, 35 ° 22 ’ S, 148 ° 50 ’ E, ex undergrowth; II. 1987, D. H. Colless, sweeping; Malaise trap, JSS 8627 – 9, JSS 8632 – 3 (5 ♂, ANIC); near Condor Creek, 35 ° 19 ’ S, 148 ° 50 ’ E, 6. III. 1970, R. Lindsay, JSS 8675 (1 ♂, ANIC); New South Wales: 23 km SE Tamworth, 31 ° 13 ’ S, 151 ° 7 ’ E, 26. XI. 1982, D. S. Horning, Jr., JSS 9147 (1 ♂, ANIC); 4 miles SW of Tarago, 35 ° 7 ’ S, 149 ° 36 ’ E, 20. III. 1966, Z. Liepa, JSS 9218 (1 ♂, ANIC); Culgoa National Park, 13.5 km SW Cawwell Homestead (CGN 1 M), 29 ° 7 ’ S, 146 ° 56 ’ E, Callitrus, 21. XII. 2009 – 30. I. 2010, B. Schiebaan, Malaise trap, CNC 576579 (1 ♂, QM); Callitris, 30. I – 18. V. 2010, QM Team, B. Shiebaan & R. Ohlsen, Malaise trap, CNC 576631, CNC 576635 (2 ♂, QM); Monga, near Braidwood, 35 ° 33 ’ S, 149 ° 56 ’ E, 13. II. 1966, Z. Liepa, JSS 9210 (1 ♂, ANIC); Yathong Nature Reserve near 8 Mile Tank, 32 ° 38 ’ S, 145 ° 41 ’ E, 205 m, field of flowers: open woodland, 3 – 25. X. 2003, C. Lambkin & N. Starick, Malaise trap, CNCD 3638 (1 ♂, CNC); Grafton Experimental Farm, 29 ° 40 ’ S, 152 ° 56 ’ E, 1. XII. 1960, M. F. Day, JSS 8642 (1 ♂, ANIC); Pearl Beach, near Woy Woy, 33 ° 33 ’ S, 151 ° 18 ’ E, 9 – 11. XII. 1988, M. J. Fletcher & J. A. Macdonald, Malaise trap, JSS 8742 (1 ♂, ASCU); Yathong Nature Reserve near Triangle Tank, 32 ° 38 ’ S, 145 ° 33 ’ E, 200 m, in sandy wash in low closed mallee, 3 – 25. X. 2003, C. Lambkin & N. Starick, Malaise trap, JSS 16039 (1 ♂, ANIC); Queensland: Brisbane, Department of Primary Industries, Indooroopilly site, 27 ° 30 ’ S, 152 ° 59 ’ E, II. 1977, Malaise trap, JSS 8936 (1 ♂, QDPC); Culgoa Floodplain National Park, 7 km WNW Toulby Gate (TOLB 6 M), 28 ° 57 ’ S, 146 ° 52 ’ E, 138 m, Callitris, 19. III – 20. V. 2010, C. Kelly & A. Coward, Malaise trap, CNC 575021, CNC 575024 (2 ♂, QM); Mount Coot-tha, 27 ° 29 ’ S, 152 ° 57 ’ E, 170 m, hilltop, 26. IV. 1998, C. Lambkin, hand collected, JSS 7739 – 41 (3 ♂, QM); Bundaberg, 24 ° 52 ’ S, 152 ° 21 ’ E, 15 – 28. II. 1972, H. Frauca, JSS 8617 (1 ♂, ANIC); Charleville, 26 ° 24 ’ S, 146 ° 14 ’ E, 15. II. 1970, A. L. Dyce, car net, JSS 8624 (1 ♂, ANIC); Western Australia: Charles Darwin Reserve, Wanarra Road, 29 ° 35 ’ S, 116 ° 60 ’ E, York Gums, Acacia, flowering herbs, 19 – 23. IX. 2009, Lambkin & Monteith, Malaise trap, CNC 575172 (1 ♂, QM); 158 km S Newman, 9 km N Kumarina Road House, 24 ° 38 ’ S, 117 ° 37 ’ E, 638 m, in wide sandy wash, 18 – 21. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 16409 (1 ♂, CNC); 171 km East of Marble Bar, 21 ° 17 ’ S, 121 ° 14 ’ E, 300 m, dry wash with flowers, 2 – 15. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15874 – 6 (2 ♂, 1 ♀, CNC); 47 km S Pardoo Road House on Shay Gap Road, 20 ° 23 ’ S, 120 ° 1 ’ E, 170 m, dry wash near Spinifex, 1 – 14. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15703 (1 ♀, CNC); 67 km SW Pardoo Road House on Shay Gap Road, 20 ° 28 ’ S, 120 ° 10 ’ E, 177 m, dry sandy ravine with flowers, 1 – 14. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15663 (1 ♂, CNC); 70 km S Newman on Great Northern Highway, 23 ° 54 ’ S, 119 ° 45 ’ E, 626 m, small wash with Eucalyptus, 6 – 18. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15879 (1 ♀, CNC); Karijini National Park, Juna Downs Road, 22 ° 40 ’ S, 118 ° 26 ’ E, 797 m; in open mulga with flowering Ptilotus, 19 – 25. IV. 2003, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16287, JSS 16289, JSS 16308 (2 ♂, 1 ♀, ANIC); 22 ° 44 ’ S, 118 ° 25 ’ E, 798 m, dry Turee Creek, grassy open Eucalyptus scrub, 25. IV – 14. V. 2003; 14 – 19. V. 2003, JSS 16096, JSS 16111 (2 ♀, ANIC); Mount Augustus National Park; south slope on Saddle Trail, 24 ° 18 ’ S, 116 ° 48 ’ E, 400 m, small dry wooded wash, 25. IV – 7. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15887 (1 ♂, CNC); Mount Augustus National Park, 24 ° 22 ’ S, 116 ° 50 ’ E, 427 m, wash on southern slope of mountain, 25. IV – 7. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15878 (1 ♀, CNC); base of “ The Governor ” on Great Northern Highway, 23 ° 3 ’ S, 118 ° 50 ’ E, 722 m, dry wash in wooded area, 23. IV – 6. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15689, JSS 15693, JSS 15698 (3 ♀, CNC); Victoria: Dartmouth Survey, Dart-Mitta River Junction, Locality L, 36 ° 31 ’ S, 147 ° 27 ’ E, 4. III. 1973, JSS 8760 (1 ♂, MVMA); Dartmouth Survey, Mitta Mitta River, Loc. EA, 36 ° 31 ’ S, 147 ° 27 ’ E, 9. II. 1973, JSS 8771 (1 ♂, MVMA); Mitcham, 37 ° 49 ’ S, 145 ° 12 ’ E, XII. 1982, C. Lai (D. Yu), Malaise trap, JSS 14387 (1 ♂, DEBU); Walhalla, 37 ° 57 ’ S, 146 ° 27 ’ E, 5. III. 1981, Z. Liepa, JSS 9177 (1 ♂, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF51ECEAD8D9EFF6FDA41694.taxon	distribution	Distribution: Australia (Australian Capital Territory, New South Wales, Queensland, Western Australia, Victoria) (Fig. 130). Notes: This is another common and widespread species that has never been recording hilltopping. Intraspecific genetic distance ranges from 0.0 – 1.5 %. It is genetically most similar to T. latistyla sp. nov. (8.8 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF54ECE6D8D9EFF6FD0E13D0.taxon	description	Figs 60 A – I, 131, 157 E	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF54ECE6D8D9EFF6FD0E13D0.taxon	diagnosis	Diagnosis: This species can be recognized by a distinct keel on the hind trochanter that bears 9 – 11 bristles; elongated surstyli in lateral view (Fig. 60 A, E); both surstyli bent towards sternite at the apex in lateral view (Fig. 60 C – D, 60 H – I); phallic guide with two small spines towards epandrium in lateral view (Fig. 60 C – D).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF54ECE6D8D9EFF6FD0E13D0.taxon	description	Description: MALE: Body length: 2.2 – 2.5 mm. Head. Flagellum brown and acuminate; face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose; eyes touching for distance equal to 1.3 – 2 times the length of ocellar triangle; occiput silvery pollinose, slightly less so on upper half. Thorax. Postpronotum pale, with 4 – 5 whitish bristles. Mesonotum (viewed obliquely from front) subshining black, anterior part indistinctly silver pollinose, slightly grayish also from the side. Scutellum silvery pollinose and without bristles. Dorsocentral bristles weakly developed, dark, longer on frontal part. Halter, knob pale, stem dark brown. Legs. Mid coxa with 2 – 3 spines. Trochanters and base of femora brown; femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees, basal 1 / 5 of tibiae yellow, tarsal segments yellow-brown, last segment dorsally darker. Ventroapical row of spines on fore femur missing, 4 – 6 short, black spines on mid femur; no spines on hind femur, only 12 – 14 short bristles posteriorly equally distributed, longer on distal half. Subapical (distal) spines on first four tibiae present (short). Hind trochanter with a keel bearing 9 – 11 distinct bristles. Distitarsi slightly flattened, first tarsomere slightly longer than 2 – 4 combined; ventral bristles yellow and scrub-like. Pulvilli as long as last tarsal segment. Wing. Length: 2.4 – 2.5 mm, LW: MWW = 2.0. Upper side of basal costal cell with one long distinct bristle. Fourth costal section 2.5 – 3 times as long as third costal section. Cross-vein r-m at middle or slightly proximal to middle of discal cell. Setulae on tegula absent. Abdomen. Viewed obliquely from front tergites brownish pollinose, tergite 1 silver-grey with 2 – 3 short brown lateral bristles (up to 0.1 mm), sides silvery pollinose on tergites 4 and 5 (the latter with larger spot). Dispersed short dark bristles on tergites present, the longest on tergite 5 and as long as width of hind tibia at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 as long as ST 8. Genitalia without dissection: ST 8 large, brown; membranous area long and broad, widest on dorsal part, vertical; surstyli relatively long, broad and flat. Genital capsule in dorsal view: epandrium dark brown; surstyli light brown. Epandrium longer than wide (MLE: MWE = 1.13). Both surstyli elongated, left longer than right one, left smoothly curved over right one or straight (Fig. 60 A, E). Genital capsule in ventral view: Subepandrial sclerite short to long; gonopods short or elongated towards surtsyli; phallus trifid, ejaculatory ducts short (Fig. 60 B, F). Genital capsule in lateral view: both surstyli elongated, bent towards sternite at the tips (Figs 60 C – D, 60 H – I). Phallic guide shortened with two small spines dorsally. Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 60 G) FEMALE: Body length (excluding antennae and head): 1.7 mm. Thorax. Postpronotal lobe light yellow, gray pollinose, with 1 – 2 light brown bristle along the upper margin (up to 0.01 mm). Prescutum and scutum, scutellum black, light brown pollinose in dorsocentral area and gray pollinose in prescutum. Pleura black, prosternum and metasternum brown. Wing. Length: 2.5 mm. LW: MWW = 3.3. Legs. Mid coxae with two long and one short dark anterior bristle. Hind trochanter with a row of short dark bristles ventrally. All femora bearing one wrinkled indentation in basal. Abdomen. Tergites 1 – 6 gray pollinose with scattered short dark bristles. Tergite 1 with 2 long and 2 – 3 short black lateral bristles. Ovipositor. Base of ovipositor dark, gray pollinose, piercer light brown. Viewed laterally (Fig. 157 E), base of piercer curved, piercer long and straight. LP: LB = 4.7. LDP: LPP = 3.0.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF54ECE6D8D9EFF6FD0E13D0.taxon	etymology	Etymology: The specific name is derived from the Greek word makros for long, which combined with stylus refers to the long surstyli of males of this species. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: Karijini National Park: Juna Downs Road (between drying pools in rocky Turee Creek), 22 ° 42 ’ S, 118 ° 25 ’ E, 789 m, 25. iv – 14. v. 2003, C. Lambkin, T. Weir, Malaise trap, CNCD 3861 (1 ♂, WAM); PARATYPES: AUSTRALIA: Western Australia: Karijini National Park: Juna Downs Road (between drying pools in rocky Turee Creek), 22 ° 42 ’ S, 118 ° 25 ’ E, 789 m, 25. iv – 14. v. 2003, C. Lambkin & T. Weir, Malaise trap, CNCD 3813, CNCD 3827 – 8, CNCD 3831, CNCD 3834, CNCD 3836, CNCD 3838, CNCD 3841 – 2, CNCD 3844, CNCD 3846, CNCD 3848 – 9, CNCD 3852, CNCD 3854 – 5, CNCD 3857 – 60, CNCD 3864 – 5 (4 ♂, 1 ♀, CNC; 14 ♂, 3 ♀, QM); Karijini National Park, Juna Downs Road, 22 ° 42 ’ S, 118 ° 25 ’ E, 789 m, between drying pools in rocky Turee Creek, 19 – 25. iv. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16257 (1 ♀, QM); 22 ° 44 ’ S, 118 ° 25 ’ E, 798 m, dry Turee Creek, grassy open Eucalyptus scrub, 25. iv – 14. v. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16094 (1 ♀, QM); Karijini National Park, Weano Gorge Road, 22 ° 22 ’ S, 118 ° 15 ’ E, ~ 775 m, on hilltop, open Eucalyptus, 25. iv – 14. v. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16137; JSS 16140 (2 ♂, ANIC); Millstream-Chichester National Park, Black Hill Pool, 21 ° 20 ’ S, 117 ° 15 ’ E, 176 m, over wide dry rocky creek bed, Eucalyptus, 7 – 12. v. 2003, C. Lambkin & D. Yeates, Malaise trap, JSS 16314 (1 ♂, ANIC); Millstream-Chichester Nat [ional] Park, Pipeline Road, 21 ° 36 ’ S, 117 ° 5 ’ E, 324 m, in Acacia Spinifex scrub, 2 – 11. V. 2003, C. Lambkin & D. Yeates, J. Recsei, Malaise trap, JSS 16153 (1 ♂, ANIC); Queensland: Mornington Island, 16 ° 33 ’ S, 139 ° 24 ’ E, 18. IV. 1983, J. F. Donaldson, D-Vac, JSS 8962 – 64; JSS 8966 (4 ♂, QDPC); Culgoa Floodplain National Park, 7 km Northwest of Toulby Gate (TOLB 7 M), 28 ° 57 ’ S, 146 ° 53 ’ E, 138 m, reedy swamp, 20. I – 19. III. 2010, C. Kelly & A. Coward, Malaise trap, CNC 575366 (1 ♂, QM); Culgoa Floodplain National Park, 7 km NNW Toulby Gate (CG 4 AM), 28 ° 56 ’ S, 146 ° 55 ’ E, 160 m, Gidgee, 20. i – 19. iii. 2010, C. Kelly, A. Coward, Malaise trap, CNC 576379, CNC 576382 – 3, CNC 576386 – 90 (6 ♂, QM; 2 ♂, USNM); Culgoa Floodplain National Park, 7 km Northwest of Toulby Gate (TOLB 7 M), 28 ° 57 ’ S, 146 ° 53 ’ E, 138 m, reedy swamp, 20. i – 19. iii. 2010, C. Kelly, A. Coward, Malaise trap, CNC 575373 – 6 (2 ♂, CNC; 2 ♂, QM); New South Wales: Culgoa National Park, 8 km West-Northwest of Cawwell Homestead, Diemunga Lagoon (CGN 2 M), 29 ° 3 ’ S, 146 ° 1 ’ E, 136 m, Coolibah, 21. XII. 2009 – 30. I. 2010, B. Schiebaan, Malaise trap, CNC 574749 (1 ♂, QM); Tasmania: 10 km NW by N St. Helens, 41 ° 15 ’ S, 148 ° 10 ’ E, 14. I. 1983, I. D. Naumann, J. C. Cardale, yellow pan trap, JSS 9264 (1 ♂, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF54ECE6D8D9EFF6FD0E13D0.taxon	distribution	Distribution: Australia (New South Wales, Queensland, Tasmania, Western Australia) (Fig. 131). Notes: Based on genitalia, there are two putative species in what we are treating as one species here. The COI sequences are overlapping and both species occur at the same locations on the same dates, so we have opted to be conservative and treat them as a single species. More research should be conducted on this potential species complex. Host data and other genetic markers would be useful to help determine if it is one or two species. One putative species has the dorsomedial margin of the surstyli curved in lateral view (Fig. 60 H – I) and includes CNC 576390, CNCD 3827, CNCD 3834, CNCD 3836, CNCD 3841, CNCD 3848, CNCD 3852, CNCD 3854 – 5, CNCD 3857, CNCD 3859 – 60, CNCD 3865, JSS 16094, JSS 16137 and JSS 16140. The second putative species has the dorsomedial margin of surstyli straight in lateral view and more elongated gonopods in ventral view (Fig. 60 A – D) which includes CNCD 3813, CNCD 3838, CNCD 3842, CNCD 3844, CNCD 3846, CNCD 3849, CNCD 3861, JSS 16314, CNC 576379, CNC 576382, CNC 576383, CNC 57638 – 9, CNC 575373 – 6, JSS 9262 – 4, JSS 8966 and JSS 9264. Genetic distance between sequenced specimens is low (0.0 to 0.7 % pairwise divergence). Intraspecific genetic distance ranges from 0.0 – 1.6 %. This species is genetically most similar to T. bulbosa sp. nov. and T. warrumbunglensis sp. nov. (3.2 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF5AECE4D8D9EC97FE1913EF.taxon	description	Figs 61 A – E, 130	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF5AECE4D8D9EC97FE1913EF.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a few short dark spines ventrobasally; elongated surstyli in dorsal view (Fig. 61 A); extended gonopods towards surstyli in ventral view (Fig. 61 B); phallic guide with some different-sized spines, longer one as wide as left surstylus in ventral view (Fig. 61 B); membranous sheath covered by small hairs.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF5AECE4D8D9EC97FE1913EF.taxon	description	Description: MALE: Body length: 2.4 – 2.5 mm. Head. Flagellum acuminate; yellow. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 4 – 5 distinct pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part silver pollinose; slightly grayish also from the side. Scutellum silver pollinose and with 4 – 7 weak bristles along the distal edge. Dorsocentral bristles developed, dark, longer on frontal part and numerous short dark bristles around postpronotum. Halter, knob pale, stem dark brown. Legs. Mid coxa with 2 bristles (one thicker). Trochanters and base of femora black, femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees, basal 1 / 5 of tibiae yellow (otherwise dark brown), tarsal segments yellow ventrally, yellow-brown dorsally, last segment dorsally darker. Hind trochanter with a few short dark spines (5 – 7) ventrobasally; ventrobasal spines absent on fore and mid femora. Ventroapical row of spines on fore femur missing, 4 – 6 very short, black spines on mid femur; no spines on hind femur, only 14 – 16 pale bristles posteriorly, more frequent on distal half (as long as 1 / 2 the width of hind tibia at the distal end); hind tibia slightly bent at middle in posterior view. Subapical (distal) spines on first four tibiae absent. Hind metatarsus flattened (1 and 2 especially), first tarsomere slightly longer than 2, both metatarsus with dorsal row of bristles along the edges but missing in the middle. Pulvilli as long as 2 / 3 of last tarsal segment. Wing. Length: 2.5 mm. Upper side of basal costal cell with 1 – 3 long distinct bristles. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m at middle of discal cell. 3 – 4 dark setulae on tegula. Abdomen. Viewed obliquely from front tergites subshining black, tergite 1 silvery grey, sides silvery pollinose on tergites 4 and 5 (the latter with larger spots). Dispersed short dark bristles on tergites present, the longest on tergite 5 and ST 8 and up to 0.5 – 07. times as long as width of hind tibia at base. Lateral bristles on first tergite present, 5 – 6 brown bristles up to as long as width of hind femur at distal tip. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1 – 1.2 times as long as ST 8. Genitalia without dissection: ST 8 medium sized, brownish, hairy; membranous area slit-like, long; epandrium brown; surstyli paler, appear slim. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.7). Surstyli rather asymmetrical, elongated, left slightly longer than right one (Fig. 61 A), right surstylus with a depression ventromedially before apex. Genital capsule in ventral view: subepandrial sclerite covered with some transverse streaks in upper margin near surstyli; gonopods extended towards surstyli, left longer than right one, hypandrium with distinct hypandrial apodeme (Fig. 61 B), phallic guide with 5 – 6 different-sized spines dorsolaterally (longer ones extended towards left surstylus), longer one is as wide as left surstylus, membranous sheath covered by small bristles. Genital capsule in lateral view: both surstyli curved towards sternite, moderately narrowed towards apical, right surstylus shorter and wider than left one (Fig. 61 D – E); ejaculatory apodeme flat, sperm pump elongate, linear (Fig. 61 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF5AECE4D8D9EC97FE1913EF.taxon	etymology	Etymology: This species is named in honour of Michael Mathieson. Michael introduced JHS to the type locality (Round Mountain) which proved to be an excellent location for sampling pipunculid diversity. Michael was also part of the team that collected both known specimens of this species. He has collected 9 % of the known species of Tomosvaryella (Supplementary file 2). Examined material: HOLOTYPE: AUSTRALIA. Queensland: Barakula State Forest No. 302, Summit Round Mountain, 26 ° 22 ’ S, 150 ° 58 ’ E, 8. i. 2000, J. & A. Skevington & M. Mathieson, JSS 7552 (1 ♂, QM); PARATYPE: AUSTRALIA: Queensland: same data as holotype, JSS 7553 (1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF5AECE4D8D9EC97FE1913EF.taxon	distribution	Distribution: Australia (Queensland) (Fig. 130). Notes: The only two known specimens of this species were caught hilltopping on Round Mountain. Intraspecific genetic distance is 0.0 %. This species is genetically most similar to T. powerae sp. nov. (2.3 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF58ECE2D8D9ECBFFD6F11FC.taxon	description	Figs 62 A – E, 130, 144 C	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF58ECE2D8D9ECBFFD6F11FC.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a small keel with 2 – 3 short dark spines (Fig. 144 C); small surstyli with two ventromedial lobes in dorsal view; both gonopods extended towards surstyli, right one with a distinct projection on inner margin; phallic guide with some dorso-dorsolateral spines in ventral view (Fig. 62 B); both surstyli straight in lateral view (Fig. 62 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF58ECE2D8D9ECBFFD6F11FC.taxon	description	Description: MALE: Body length: 2.4 mm. Head. Scape, pedicel dark brown; flagellum tapering, light brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2.1 times the length of ocellar triangle. Occiput silvery pollinose. Thorax. Postpronotum pale, with 2 – 3 long brown bristles. Mesonotum (viewed obliquely from front) silvery pollinose, with some short supra-alar bristles behind postpronotum lobes. Scutellum silvery brown pollinose and with 5 – 8 short dark bristles along distal edge. Halter, knob yellow, tip brownish, stem yellow, base dark brown. Legs. Mid coxa with 3 – 4 dark bristles. Trochanters brown, femora black, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments light brown. Hind trochanter with a small keel providing with 2 – 3 short dark spines (Fig. 144 C); ventrobasal spines absent on femora. Hind femur without ventroapical spines, except 8 – 10 equally spaced short bristles posteroventrally, longest up to 2 / 3 the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres not flattened, hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 2.5 mm. Upper side of basal costal cell with one brown bristles. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. Abdomen. Viewed obliquely from front tergites brown, tergite 1 brown pollinose. Lateral bristles on first tergite present, 8 – 10 long brown bristles up to as long as hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium rectangular-shaped (MLE: MWE = 1.1). Surstyli rather short, widened in the basal fourth, then constricted, broadened in the middle, both surstyli with a small lobe in dorsomedial margin, curved to each other at apex, left surstylus is longer (Fig. 62 A). Genital capsule in ventral view: both gonopods extended towards surstyli, pointed at apex, right one with a distinct projection on inner margin, gonopods unequal in height, left slightly longer; phallic guide with 3 – 6 short dorso- dorsolateral spines; two left ones longer than others; subepandrial sclerite distinct (Fig. 62 B); Genital capsule in lateral view: both surstyli straight (Fig. 62 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 62 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF58ECE2D8D9ECBFFD6F11FC.taxon	etymology	Etymology: This species is named after the paperbark trees (Melaleuca spp.) that dominate the floodplain in the otherwise treeless type locality. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Edgbaston National Reserve (EBM 1), 22 ° 44 ’ S, 145 ° 26 ’ E, 237 m, Melaleuca floodplain, 29. X – 15. XII. 2010, C. Lambkin et al., Malaise trap, CNC 596961 (1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF58ECE2D8D9ECBFFD6F11FC.taxon	distribution	Distribution: Australia (Queensland) (Fig. 130).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF5EECE0D8D9EEBBFE351168.taxon	description	Figs 63 A – F, 131	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF5EECE0D8D9EEBBFE351168.taxon	diagnosis	Diagnosis: This species can be recognized by the presence of four long, dark bristles on mid coxa; hind trochanter with 4 – 5 pale bristles in a row; elongated surstyli, bent outwards by 90 ° distally, with two small protrusions before apex at inner margin in dorsal view (Fig. 63 A); gonopods covered by short bristles (Fig. 63 B); phallic guide with a long spine dorsomedially in lateral view (Fig. 63 D – F).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF5EECE0D8D9EEBBFE351168.taxon	description	Description: MALE: Body length: 3.2 mm. Head. Flagellum long acuminate; yellow. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose; eyes touching for distance equal to 2 – 2.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 1 – 2 indistinct pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part slightly silver pollinose, very faintly grayish also from the side. Scutellum brownish pollinose and with 6 – 8 weak bristles along the distal edge. Dorsocentral bristles developed, short, somewhat longer on frontal half, numerous pale bristles around postpronotum. Halter, knob pale, stem brown. Legs. Mid coxa with four long, dark bristles. Trochanters and femora brown-black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees, basal 1 / 5 and distal extremity of tibia yellow (tibia otherwise brown), tarsal segments yellow, last segment brown. Fore and mid trochanters with 2 – 3 short bristles. Hind trochanter without a keel, but with a few pale bristles (4 – 5) in a row that looks like a half circle (flat dorsally). Ventroapical row of spines on fore femur missing; 6 – 8 short, black spines on mid femur; no spines on hind femur, only 14 – 16 pale bristles equally spaced posteriorly, uniform in length (up to as long as 0.4 – 0.5 times the width of hind tibia at the distal end). Subapical (distal) spines on first four tibiae present (short). Ventrobasal spines (2) present on fore and absent on mid femur. Hind metatarsus not flattened, metatarsus almost as long as 2 – 5 combined; last tarsomere unusually long almost as long as segments 3 – 4 combined. Pulvilli as long as the last tarsal segment, shorter on metatarsi. Wing. Length: 3.5 mm. Upper side of basal costal cell with one dark and short bristles. Fourth costal section 2 – 2.2 times as long as third costal section. Cross-vein r-m at middle of discal cell. Three distinct dark setulae on tegula. Abdomen. Viewed obliquely from front tergites subshining black with a greenish hint, tergite 1 silvery grey, sides silvery pollinose on tergites 5. Dispersed strong dark bristles on all tergites present, the longest on tergite 5 and up to as long as half the width of hind tibia at base, otherwise tergites covered with very faint, velvet-like bristles. Lateral bristles on first tergite present, 4 – 5 long pale bristles in a row up to 0.8 – 1 time as long as the width of hind femur at distal tip in lateral view. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 2.4 – 2.6 times as long as ST 8. Genitalia without dissection: ST 8 medium sized, wide, with occasional small bristles; membranous area elongated, broadening on dorsal part; epandrium and surstyli uniformly brown, surstyli appear to be long and with broad base. Genitalia. Genital capsule in dorsal view: epandrium square-shaped (MLE: MWE = 1.0). surstyli symmetrical, broadened at base, gradually slenderized before apical fourth, bent outwards by 90 ° distally, with two small protrusions shortly before apex at inner margin (Fig. 63 A). Genital capsule in ventral view: gonopods equal, extended towards surstyli, both gonopods covered by short bristles; lobes of hypandrium sclerotized; subepandrial sclerite shortened with some scattered short bristles in its middle (Fig. 63 B); Genital capsule in lateral view: both surstyli broadened at base and apex, constricted in middle (Fig. 63 D – E); phallic guide straight, with a long spine dorsomedially; phallus with three ejaculatory ducts (Fig. 63 F). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 63 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF5EECE0D8D9EEBBFE351168.taxon	etymology	Etymology: The name is in reference to the four long, dark bristles on the mid coxa and is derived from the Greek word merinx (bristle) and the Latin ater (black). Examined material: HOLOTYPE: AUSTRALIA: Queensland: Forest Station, Bulburin State Forest via Many Peaks, 24 ° 32 ’ S, 151 ° 34 ’ E, 600 m, long grass, wet sclerophyll forest, 12 – 15. iv. 1974, I. Naumann, JSS 8306 (1 ♂, QM); PARATYPE: AUSTRALIA: Queensland: Brisbane, 27 ° 28 ’ S, 153 ° 1 ’ E, 24. i. 1960, Haseler, JSS 8301 (1 ♂, QM); Northern Territory: McArthur River, 48 km SW by S of Borroloola, 15 ° 54 ’ S, 136 ° 40 ’ E, 14. IV. 1976, D. H. Colless, JSS 9221 – 22 (2 ♂, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF5EECE0D8D9EEBBFE351168.taxon	distribution	Distribution: Australia (Northern Territory, Queensland) (Fig. 131). Notes: One specimen was collected in wet sclerophyll forest and the Northern Territory specimen was found near the coast in what would presumably be wet, monsoon forest. Future searches for this species should likely focus on wet forest types.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF5CECFED8D9EE0FFA0012F0.taxon	description	Figs 64 A – E, 129	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF5CECFED8D9EE0FFA0012F0.taxon	diagnosis	Diagnosis: This species can be recognized by the triangle-shaped of the cerci in lateral view (Fig. 64 D – E); hind trochanter with a small keel and some short bristles ventromedially; small surstyli, both surstyli constricted in apical third at inner margin in dorsal view (Fig. 64 A); left gonopod slightly longer, right one wider, both gonopods with a short finger-like projection laterally; phallic guide with some different-sized spines dorso- dorsolaterally; subepandrial sclerite and hypandrial apodeme distinct (Fig. 64 B, D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF5CECFED8D9EE0FFA0012F0.taxon	description	Description: MALE: Body length: 3.2 – 3.5 mm. Head. Scape and pedicel dark brown. Pedicel with 2 – 3 upper bristles; flagellum light brown, gray pollinose and tapering. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2.5 – 3 times the length of ocellar triangle. Occiput grey pollinose, covered by short dark bristles. Thorax. Postpronotum light yellow, with 3 – 5 light brown bristles in upper margin. Mesonotum (viewed obliquely from front) silvery-brown pollinose, anterior part greyer; scutum with scattered bristles at anterior supra-alar area. Postalar callus dark brown with a batch of (3 – 5) pale bristles in its middle. Scutellum black, silvery pollinose with 10 – 12 short bristles along distal edge. Halter, knob pale, stem light brown. Legs. Mid coxa with 4 – 5 long dark bristles, two longer than others. Coxae, trochanters, femora dark brown, hind femur shining ventrally, knees and basal 1 / 4 of tibiae yellow (tibiae otherwise brown), tarsomeres yellow ventrally and light brown dorsally, last segment darker. Mid trochanter with two short bristles ventroapically. Hind trochanter silvery pollinose, with a small keel and 3 – 6 short bristles ventromedially; ventrobasal spines absent on fore and mid femur, two rows ventroapical spines present. Hind femur with two rows ventroapical spines. Tibiae with two rows of short black bristle on anterior and three rows on posterior side. Hind tarsomeres distinctly flattened (especially tarsomeres 1 – 4), hind metatarsus as long as 2 – 4 combined; posterior corners with distinct bristles on all tarsomeres. Pulvilli shorter than last tarsal segment. Wing. Length: 3.5 mm. Upper side of basal costal cell with one short dark brown bristle and 2 – 3 longer light brown bristles. Fourth costal section 1.5 – 2 times as long as third costal section. Cross-vein r-m at middle of discal cell. 3 – 4 short dark setulae on tegula. Abdomen. Tergites 1 silvery pollinose; tergite 2 – 5 grey-brown pollinose with scattered short dark bristles (more on tergite 5). Lateral bristles on first tergite present, 7 – 10 long brown bristles up to as long as hind femur’s width at base; sternite 2 with 3 – 6 small dark spines in the middle of posterior margin. Syntergosternite 8 enlarged, dark brown and grey pollinose. Membranous area long and broad in middle with a triangular edge close to cerci in lateral view; cerci triangle-shaped in lateral view. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.6). Surstyli asymmetrical, small, left surstylus longer than right one, right surstylus with a triangular edge at inner margin before apex, both surstyli constricted in apical third at inner margin (Fig. 64 A). Genital capsule in ventral view: gonopods rather unequal in height, left gonopod slightly longer, right one wider than left one, both gonopods with a short finger-like projection laterally; phallic guide with 2 – 3 different-sized spines dorso- dorsolaterally, one is longer than others; subepandrial sclerite and hypandrial apodeme distinct (Fig. 64 B); Genital capsule in lateral view: both surstyli widened, strongly curved towards sternite, left surstylus moderately narrowed after basal half (Fig. 64 D – E); ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 64 C). FEMALE: unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF5CECFED8D9EE0FFA0012F0.taxon	etymology	Etymology: This species is named after the type locality, Millstream-Chichester National Park. This park is particularly diverse for Tomosvaryella species with 17 species recorded, all of them undescribed until now. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: Millstream-Chichester Nat [ional] P [ark], Narrina Pool, 21 ° 21 ’ S, 117 ° 16 ’ E, 159 m, rocky creek bed, Eucalyptus, 7 – 12. V. 2003, C. Lambkin & D. Yeates, Malaise trap, JSS 16186 (1 ♂, WAM); PARATYPES: AUSTRALIA: Western Australia: Karijini National Park, Karijini Drive, 22 ° 34 ’ S, 118 ° 18 ’ E, 814 m, open Eucalyptus grassland, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 15581, JSS 15583, JSS 15584 (3 ♂, ANIC); 22 ° 34 ’ S, 118 ° 18 ’ E, 815 m, across dry gully, Acacia scrub, 19 – 25. IV. 2003, JSS 16157, (1 ♂, CNC); Millstream-Chichester National Park, Narrina Pool, 21 ° 21 ’ S, 117 ° 16 ’ E, 159 m, rocky creek bed, Eucalyptus, 7 – 12. V. 2003, C. Lambkin & D. Yeates, Malaise trap, JSS 16182 – 5, JSS 16187 (4 ♂, ANIC; 1 ♂, QM); beside bank of rocky creek bed, Eucalyptus, 27 – 30. IV. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16192 (1 ♂, ANIC); Millstream-Chichester National Park, Roebourne Road, 21 ° 26 ’ S, 117 ° 9 ’ E, 337 m, sandy creek, Eucalyptus Spinifex grassland, 7 – 12. V. 2003, C. Lambkin & D. Yeates, Malaise trap, JSS 15569, JSS 15571 – 2 (3 ♂, ANIC); 21 ° 20 ’ S, 117 ° 11 ’ E; 21 ° 26 ’ S, 117 ° 9 ’ E, 360 m; 337 m, Spinifex with scattered Acacia & Eucalyptus; sandy creek, Eucalyptus Spinifex grassland, 27. IV – 3. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16072 – 3, JSS 16198, JSS 16261 – 3 (6 ♂, ANIC); Millstream-Chichester National Park, Black Hill Pool, 21 ° 20 ’ S, 117 ° 15 ’ E, 176 m, across wide dry rocky creek bed, Eucalyptus, 30. IV – 3. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16043 (1 ♂, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF5CECFED8D9EE0FFA0012F0.taxon	distribution	Distribution: Australia (Western Australia) (Fig. 129). Notes: This species has only been found in Millstream-Chichester National Park and Karijini National Park in Western Australia. No specimens have been found hilltopping. Intraspecific genetic distance ranges from 0.0 % to 1.0 %. It is genetically most similar to T. tribula sp. nov. (0.8 – 2.5 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF42ECFFD8D9EDB7FCB5108C.taxon	description	Figs 65 A – E, 131, 144 D	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF42ECFFD8D9EDB7FCB5108C.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a keel covered by some short spines along its edge (Fig. 144 D); mid metatarsus flattened at its base, extended inward; elongated surstyli, curved to each other from middle to apex in dorsal view (Fig. 65 A); gonopods extended towards surstyli, subepandrial sclerite distinct with a fin-shaped projection in its center in ventral view (Fig. 65 B); right surstylus with hump-like edge dorsomedially in lateral view (Fig. 65 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF42ECFFD8D9EDB7FCB5108C.taxon	description	Description: MALE: Body length: 3.2 mm. Head. Flagellum tapering, pale at the apex, light brown at the base; scape and pedicel dark. Pedicel with two short dark upper bristles. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 1 – 2 short dark bristles. Mesonotum (viewed obliquely from front) brown pollinose, anterior part more grey, grayish also from the side. Scutellum silvery pollinose. Dorsocentral bristles short, dark, longer in frontal half, numerous pale bristles around postpronotum present. Halter, knob pale, tip brownish, stem brown. Legs. Mid coxa with 1 – 2 long dark bristles. Trochanters brown, femora black, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments brown. Hind trochanter with a keel covered by some short spines along its edge (Fig. 144 D). Ventrobasal spines absent on fore and absent on mid femur. Hind femur without ventroapical spines, except 3 – 6 short bristles posteroventrally. Subapical (distal) spines on first four tibiae present (short). Mid metatarsus flattened at its base, extended inward. Hind tarsomeres not flattened, hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 3.1 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 1 – 2 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites black, tergite 1 silvery brown pollinose, others brown pollinose. Lateral bristles on first tergite present, 4 – 6 long pale bristles up to as long as of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.8). Surstyli elongated, widened at the base with two small lobes ventrobasally. curved to each other at apex, left surstylus is longer than right one (Fig. 65 A). Genital capsule in ventral view: gonopods equal in height, extended towards surstyli, rounded at apex, gonopods with a small semicircular projection on inner margin; phallic guide unclear, perhaps detached; subepandrial sclerite distinct with a fin-shaped projection in its center (Fig. 65 B); Genital capsule in lateral view: both surstyli curved towards sternite, right surstylus with hump-like edge dorsomedially (Fig. 65 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 65 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF42ECFFD8D9EDB7FCB5108C.taxon	etymology	Etymology: From the Greek miltos, red earth and kentron, centre, in reference to the Red Centre (Alice Springs Region) of Australia where this specimen was collected. Examined material: HOLOTYPE: AUSTRALIA: Northern Territory: Waterhouse Range, 39 km SW by S of Alice Springs, 24 ° 1 ’ S, 133 ° 25 ’ E, 11. X. 1978, D. H. Colless, JSS 9239 (1 ♂, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF42ECFFD8D9EDB7FCB5108C.taxon	distribution	Distribution: Australia (Northern Territory) (Fig. 131).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF40ECFCD8D9EFF6FB2517E0.taxon	description	Figs 66 A – D, 124, 150 D	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF40ECFCD8D9EFF6FB2517E0.taxon	diagnosis	Diagnosis: This species can be recognized by the 8 – 10 pale long bristles along the distal half of the hind femur (Fig. 150 D); surstyli with five long bristles along dorsomedial margin in dorsal view (Fig. 66 A); gonopods broadened at the base and elongated towards surstyli in ventral view (Fig. 66 B); phallic guide with three to four spines dorsally in lateral view (Fig. 66 C – D).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF40ECFCD8D9EFF6FB2517E0.taxon	description	Description: MALE: Body length: 2.4 – 2.6 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose; eyes touching for distance equal to 1 – 1.5 times the length of ocellar triangle. Occiput silvery pollinose, including upper half. Thorax. Postpronotum pale, with 2 – 3 pale bristles. Mesonotum (viewed obliquely from front) subshining black, anterior part silver pollinose, slightly grayish also from the side. Scutellum silvery pollinose and without bristles. Dorsocentral bristles weakly developed, dark, longer on frontal part. Halter, knob pale, stem dark brown. Legs. Mid coxa with 2 – 3 dark anterior bristles on inner apical corner. Hind trochanter without a keel or projection, 4 – 5 indistinct bristles possible at ventral base. Femora brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees, basal 1 / 5 of tibiae yellow (otherwise pale brown), tarsal segments yellow, last segment dorsally brown. Ventroapical row of spines on fore femur missing, 8 – 10 short, black spines on mid femur, no spines on hind femur, only 8 – 10 pale long bristles posteriorly equally distributed on distal half, directed towards the distal end of femur (as long as width of hind tibia at the distal end) (Fig. 150 D). Subapical (distal) spines on first four tibiae present (short). Hind metatarsus flattened, first tarsomere slightly longer than 2 – 5 combined; ventral yellow scrub-like bristles. Pulvilli as long as 2 / 3 of last tarsal segment. Wing. Length: 2.5 – 2.6 mm. Upper side of basal costal cell with one long distinct bristle. Fourth costal section 2 – 3 times as long as third costal section. Cross-vein r-m slightly proximal to middle of discal cell. Setulae on tegula absent. Abdomen. Viewed obliquely from front tergites brownish pollinose, tergite 1 silvery grey, sides silvery pollinose on tergites 4 and 5 (the latter with side spots touching in the middle). Dispersed short dark bristles on tergites present, the longest on ST 8 and half as long as width of hind tibia at base. Lateral bristles on first tergite present, 2 – 3 brown bristles as long as width of hind femur at distal tip. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 as long as ST 8. Genitalia without dissection: ST 8 large, brown; membranous area round, positioned dorsally; epandrium paler, surstyli appear relatively short, tips often concealed. Genitalia. Genital capsule in dorsal view: epandrium and surstyli light brown. Epandrium longer than wide (MLE: MWE = 1.2). Surstyli rather symmetrical. Both surstyli triangle-shaped pointing tips at its apical inner corner, left slightly wider than right one (Fig. 66 A). Both surstyli with five long bristles along dorsomedial margin before apex. Genital capsule in ventral view: gonopods equal in height, broadened at the base and elongated towards surstyli (Fig. 66 B). Subepandrial sclerite elongated (Fig. 66 B). Genital capsule in lateral view: both surstyli slender and slightly curved in ventromedial margin (Fig. 66 C – D). Phallic guide with three to four spines dorsally before apex, phallus with three ejaculatory ducts. FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF40ECFCD8D9EFF6FB2517E0.taxon	etymology	Etymology: Named from the Greek minys for little, small or short and choma for bank or mound, in reference to the fact that all specimens were collected at the summit of a low rise (a hilltop of sorts). Examined material: HOLOTYPE: AUSTRALIA: Western Australia: ~ 3 km N [orth] of Denham, 25 ° 55 ’ S, 113 ° 32 ’ E, hilltop, 12. xii. 1999, J. Skevington, JSS 7363 (1 ♂, WAM); PARATYPES: Same data as holotype, CNC 944147 – 8, JSS 7331, JSS 7350, JSS 7352, JSS 7356 – 7, JSS 7370, JSS 7378 (3 ♂, CNC; 6 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF40ECFCD8D9EFF6FB2517E0.taxon	distribution	Distribution: Australia (Western Australia) (Fig. 124). Notes: All known specimens were collected from a low hilltop near the coast north of Dehnam, Western Australia. A small trail led through the arid shrublands to the hilltop where a bench with a roof over it afforded a view of Shark Bay and protection from the sun for those stopping to sit. The hill was barely longer than the surrounding land but together with this small structure it clearly offered an important landmark for mating insects. There were hundreds of flies and other insects using this simple structure as a hilltopping site. One hundred and four pipunculid specimens were collected here by JHS in a short period of time on 12 December 1999 (and included ten undescribed species of Tomosvaryella and Clistoabdominalis trochanteratus (Becker 1900 )).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF46ECFBD8D9EFF6FA8B1168.taxon	description	Figs 67 A – E, 129, 148 C, 152 C	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF46ECFBD8D9EFF6FA8B1168.taxon	diagnosis	Diagnosis: This species can be recognized by the tuft of short dark longer spines on the hind trochanter ventrobasally (Fig. 148 C); elongated surstyli in dorsal view (Fig. 67 A); elongated gonopods with an inwards projections (Fig. 67 B); hypandrium with two hairy membranous sheaths (Fig. 67 C – E); subepandrial sclerite with a small fin-shaped protrusion (Fig. 67 B); one of ejaculatory ducts with three spines (Fig. 67 C – E); phallic guide with some dorso-dorsolateral spines in lateral view (Fig. 67 C – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF46ECFBD8D9EFF6FA8B1168.taxon	description	Description: MALE: Body length: 2.3 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 3 – 4 pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum silvery pollinose and with 8 – 10 bristles along the distal edge. Dorsocentral bristles distinct, dark, somewhat longer along frontal edge, numerous dark bristles around postpronotum present. Halter, knob pale with a brownish tinge, stem brown. Legs. Mid coxa with 3 long dark bristles. Trochanters and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 4 of tibiae yellow (tibiae otherwise brown), tarsal segments brown, ventrally paler, last segment darker. Hind trochanter with tuft of short dark longer spines (7 – 10) ventrobasally in one group: “ echidna ” - like (Fig. 148 C); basal spines longer; ventrobasal spines (1) absent on fore and present on mid femur. Ventroapical row of spines on fore femur 2 – 3; 4 – 5 small, black spines on mid femur; hind femur without ventroapical spines, except 6 – 7 equally spaced bristles posteroventrally on distal half, longest up to 1 / 3 the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present (short). Hind tibia with a slight bending in posterior view. Hind metatarsus flattened, almost as long as 2 – 4 combined. Pulvilli shorter than last tarsal segment. Wing. Length: 3.1 mm. Upper side of basal costal cell with one long dark bristle. Fourth costal section 2.5 times as long as third costal section. Cross-vein r-m distal to middle of discal cell. 4 – 5 distinct brown setulae on tegula. Abdomen. Dissected. tergite 1 silvery grey. Lateral spines on first tergite present, 4 – 5 dark bristles up to 1 – 1.2 times as long as hind femur’s width at base. Postabdomen in dorsal view: Dissected. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.9). Surstyli symmetrical, elongated, right surstylus smaller than left one, right one with a small protrusion at inner margin before apex (Fig. 67 A). Genital capsule in ventral view: gonopods unequal in height, left slightly longer than right one, elongated towards surstyli with two small inward projection before apex, subepandrial sclerite rather wide with a small fin-shaped protrusion in its center; hypandrium with two hairy membranous sheaths extended towards phallus (Fig. 67 B). Genital capsule in lateral view: both surstyli curved towards sternite, right surstylus broadened in middle; phallus with three short ejaculatory ducts, one with three small spines, phallic guide with 6 – 7 dorso-dorsolateral spines (one is longer) (Fig. 67 C – E). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF46ECFBD8D9EFF6FA8B1168.taxon	etymology	Etymology: Named after Mount Moffatt Section of Carnarvon National Park. This is an incredibly diverse park for Diptera and 30 species of Tomosvaryella have been collected there, including the holotype for this species. Mount Moffatt is the centrepiece of this diversity and is without a doubt the best hilltop for concentrating Diptera in all of Australia. Two thousand four hundred and nine specimens of pipunculids have been collected in Mount Moffatt section of Carnarvon with 399 specimens on the summit of Mount Moffatt being the most collected in a single day (18 November 2014). Examined material: HOLOTYPE: AUSTRALIA: Queensland, Carnarvon National Park, Mount Moffatt Section, Mount Moffatt Summit, 25 ° 4 ’ S, 148 ° 3 ’ E, 1097 m, 20. I. 1998, hilltop, J. & A. Skevington & S. Winterton, JSS 2938 (1 ♂, QM); PARATYPE: AUSTRALIA: Queensland: 3 km NE of Mount Webb, 15 ° 3 ’ S, 145 ° 9 ’ E, 1. V. 1981, D. H. Colless, Malaise trap, JSS 218, JSS 8602 (2 ♂, ANIC); Cape York, Stewart Range, 14 ° 4 ’ S, 143 ° 41 ’ E, 500 m, 29. IV- 3. V. 1961, L. & M. Gressitt, CNCD 5554 (1 ♂, BPBM); Culgoa Floodplain National Park, 11.8 km N Headquarters (CG 2 AM), 28 ° 54 ’ S, 147 ° 8 ’ E, Brigalow, 20. I – 18. III. 2010, C. Kelly & A. Coward, Malaise trap, 19271, CNC 576686 (1 ♂, QM); Elizabeth Creek, Boodjamulla National Park, 18 ° 14 ’ S 138 ° 5 ’ E, 170 m, Bloodwood open forest, 18 – 22. IX. 2005, M. Mathieson & G. Smith, Malaise trap, 12398, JSS 50573 (1 ♂, QM); Mount Moffatt summit, 25 ° 4 ’ S, 148 ° 3 ’ E, 18. XI. 2014, J. H., A. M. & A. W. Skevington, CNC 374913 (1 ♂, CNC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF46ECFBD8D9EFF6FA8B1168.taxon	distribution	Distribution: Australia (Queensland) (Fig. 129). Notes: The third abdominal tergite of JSS 8602 is divided by a suture in the middle (Fig. 152 C) but the terminalia characters are similar to the holotype except that in the ventral view, the subepandrial sclerite has two finger-like projections basally close to gonopods. The different shape of the abdomen may be either an eclosion aberration or perhaps was triggered by a mutation or a parasitoid. There are no known adult pipunculid parasitoids.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF44ECF9D8D9EFF6FEDA15E8.taxon	description	Figs 68 A – E, 113, 152 A	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF44ECF9D8D9EFF6FEDA15E8.taxon	diagnosis	Diagnosis: This species can be recognized by a Y-shaped spine along the posterior margin of abdominal sternite 4 (Fig. 152 A); elongated surstyli, constricted in basal fourth, curved to each other, with some short bristles before apex in dorsal view (Fig. 68 A); gonopods extended towards surstyli, pointed at apex (Fig. 68 B); phallic guide with 3 – 4 short dorsolateral spines in ventral view.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF44ECF9D8D9EFF6FEDA15E8.taxon	description	Description: MALE: Body length: 2.5 mm. Head. Flagellum brown and tapering; brown pollinose, scape and pedicel dark. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching as long as the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 2 – 3 short pale bristles. Mesonotum black (viewed obliquely from front), silvery pollinose. Scutellum silvery pollinose with scattered short dark bristles. Halter, knob pale, tip brownish, stem brown. Legs. Mid coxa with 3 – 4 long dark bristles. Trochanters brown, femora black, silvery pollinose posteriorly; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments light brown. Hind trochanter without specific feature, grey pollinose with 4 – 6 short brown bristles ventroapically; ventrobasal spines present (1) on fore and mid femora. Hind femur without ventroapical spines. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres rather flattened (t 1 – 3), hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 2.4 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. Abdomen. Viewed obliquely from front tergites black, tergite 1 silvery grey. Dispersed strong dark bristles on all tergites present. Lateral bristles on first tergite present, 3 – 5 dark bristles up to as long as half of hind femur’s width at base; abdominal sternite 4 with Y shaped spine along posterior margin (Fig. 152 A). Genitalia without dissection: ST 8 large, round, globular in dorsal view, covered by some dark bristles; membranous area is slit-like; epandrium brown, surstyli brown. Genitalia. Genital capsule in dorsal view: epandrium as long as wide (MLE: MWE = 1.0). Surstyli elongated, wide at basal fourth, then strongly constricted, curved to each other at apex, left surstylus is longer, both with some distinct short bristles before apex (Fig. 68 A). Genital capsule in ventral view: both gonopods extended towards surstyli, pointed at apex; phallic guide with 3 – 4 short dorso- dorsolateral spines; subepandrial sclerite distinct (Fig. 68 B); Genital capsule in lateral view: both surstyli curved towards sternite (Fig. 68 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 68 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF44ECF9D8D9EFF6FEDA15E8.taxon	etymology	Etymology: This species is named in honour of Arturs Neboiss, who collected two of the three known specimens of this species as well as eight other species and 33 specimens of Australian Tomosvaryella. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Prince of Wales Isl [and], Torres Strait, 10 ° 41 ’ S, 142 ° 11 ’ E, 27 – 30. V. 1969, Neboiss, JSS 8782 (1 ♂, MVMA); PARATYPES: AUSTRALIA: Queensland: Prince of Wales Island, Torres Strait, 10 ° 41 ’ S, 142 ° 11 ’ E, 27 – 30. V. 1969, Neboiss, JSS 8779 (1 ♂, MVMA); Victoria: Dartmouth Survey, Dart-Mitta River Junction, Locality L, 36 ° 31 ’ S, 147 ° 27 ’ E, 4. III. 1973, JSS 8776 (1 ♂, MVMA).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF44ECF9D8D9EFF6FEDA15E8.taxon	distribution	Distribution: Australia (Queensland, Victoria) (Fig. 113). Notes: There is no variation in diagnostic characters between the specimens, so we are confident that all represent one species despite the strongly disjunct populations. We attempted to sequence the specimens, but they failed to amplify.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF45ECF6D8D9EA8FFDA417CC.taxon	description	Figs 69 A – E, 111, 145 F, 157 B	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF45ECF6D8D9EA8FFDA417CC.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter lacking a keel, but having a distinct spur at the distal end and a group of 10 – 12 dark short spines ventrobasally (reminiscent of the spiny fur of echidnas (Fig. 145 F); short subepandrial sclerite, covered with transverse streaks in ventral view (Fig. 69 B); phallic guide with three different-sized spines dorsally (Fig. 69 B); hypandrial apodeme divided into two projections distally in ventral view (Fig. 69 B); gonopods elongated with a small projection towards phallic guide in ventral view (Fig. 69 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF45ECF6D8D9EA8FFDA417CC.taxon	description	Description: MALE: Body length: 2.6 – 2.9 mm. Head. Flagellum acuminate, brown. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose; eyes touching for distance equal to 2 – 3 times the length of ocellar triangle. Occiput silvery pollinose, including upper half. Thorax. Postpronotum brownish, with 3 – 4 distinct pale bristles. Mesonotum (viewed obliquely from front) subshining black, anterior part silver pollinose, slightly grayish also from the side. Scutellum silvery pollinose and with a few indistinct pale bristles along the distal edge. Dorsocentral bristles developed, dark, longer on frontal part and numerous dark bristles around postpronotum. Halter, knob pale with a brownish tinge distally, stem dark brown. Legs. Mid coxa with 2 – 3 stronger bristles at inner apical corner. Trochanters and base of femora brown, femora brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees, basal 1 / 6 of tibiae yellow (otherwise brown), tarsal segments yellow-brown, last segment dorsally darker. Hind trochanter without keel, but distinct spur at the distal end and a group of 10 – 12 dark short spines ventrobasally (Fig. 145 F). Fore femur with two long spines ventrobasally and without ventroapical row of spines, mid femur with 4 – 6 short, black ventroapical spines and 1 – 2 long ventrobasal bristles, no spines on hind femur, only 6 – 8 pale bristles posteriorly equally distributed on distal ¼ (half as long as width of hind tibia at the distal end). Subapical (distal) spines on first four tibiae present (short); ventrobasal spines (2) present on fore and mid femora. Hind metatarsus flattened (all 5), first tarsomere as long as 2 – 3 combined; ventral bristles scrub-like, yellow. Pulvilli as long as 2 / 3 of last tarsal segment. Wing. Length: 3.0 – 3.2 mm. Upper side of basal costal cell with one long distinct bristle. Fourth costal section 4 – 5 times as long as third costal section (3 rd costal section very short). Cross-vein r-m at middle of discal cell. 5 – 6 dark bristles on tegula. Abdomen. Viewed obliquely from front tergites brownish pollinose, tergite 1 silvery grey, sides silvery pollinose on tergites 4 and 5 (the latter spots are larger). Dispersed short dark bristles on tergites present, the longest on ST 8 and up to twice as long as width of hind tibia at base. Lateral bristles on first tergite present, 8 – 10 black bristles up to twice as long as width of hind femur at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 as long as ST 8. Genitalia without dissection: ST 8 large, shiny black; membranous area roundish, mostly positioned posteriorly; epandrium paler, narrow; surstyli yellow. Genitalia. Genital capsule in dorsal view: epandrium dark brown, wider than long (MLE: MWE = 0.7). Surstyli light brown, rather symmetrical, elongated with broad base and inward bent tips, left surstylus longer than right one (Fig. 69 A). Genital capsule in ventral view: subepandrial sclerite short, covered with transverse streaks; gonopods elongated towards surstyli with a small projection towards phallic guide (Fig. 69 B), phallic guide with three different-sized spines dorsally (longer one closer to apex); hypandrial apodeme divided into two projections distally (Fig. 69 B). Genital capsule in lateral view: surstyli rather curved toward sternite (Fig. 69 D – E); phallus with three ejaculatory ducts, one bearing a small spine; ejaculatory apodeme flat, sperm pump elongated, linear (Fig. 69 C). FEMALE: Body length: 3.6 mm. As male except for the following characters. Frons, eyes separated, as wide as 1.2 – 1.5 times the width at antennae; completely silver-grey pollinose until up to 1 / 3 of the lower frons, shiny black in front of ocellar triangle, somewhat pollinose in the middle. Enlarged ommatidia silvery shining. Hind trochanter with two patch of spines, ventrobasal spines denser than ventroapical ones, without spur at the distal end. Pulvilli and claws about 1 – 1.2 times as long as last tarsal segment on metatarsi. Female abdomen with silver spots on sides of tergites 3 – 6. Base of ovipositor dark, gray pollinose with scattered bristles, piercer light brown. Viewed laterally, piercer straight, reaching middle of 3 rd segment. LP: LB = 2.0. LDP: LPP = 2.2. (Fig. 157 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF45ECF6D8D9EA8FFDA417CC.taxon	etymology	Etymology: Named in reference to the spiny native mammal, Short-beaked Echidna (Tachyglossus aculeatus (Shaw 1792 )), which comes to mind when looking at the dense, spiny area of the trochanter of males of this species. The Kunwinjku people of Western Arnhem Land have a lovely name for echidna: ngarrbek. We add the Greek ‘ iota’ to the ending of the name as it refers to anything small. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: ~ 3 km N [orth] of Denham, 25 ° 55 ’ S, 113 ° 32 ’ E, hilltop, 12. xii. 1999, J. Skevington, JSS 7379 (1 ♂, WAM); PARATYPES: AUSTRALIA: Western Australia: Karijini National Park, Weano Gorge Road, 22 ° 22 ’ S, 118 ° 15 ’ E; 775 m, on hilltop, open Eucalyptus, 25. iv – 14. v. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16151 (1 ♀ ANIC); 22 ° 24 ’ S, 118 ° 15 ’ E, 718 m, JSS 16219 (1 ♂ ANIC); Mount Magnet, Mount Waramboo, 28 ° 2 ’ S, 117 ° 49 ’ E, hilltop, 14. xii. 1999, J. Skevington, JSS 7435, JSS 7453, JSS 7464, JSS 7466, JSS 7470 – 1 (1 ♂, CNC; 5 ♂, QM); ~ 3 km North of Denham, 25 ° 55 ’ S, 113 ° 32 ’ E, ~ hilltop, 12. xii. 1999, J. Skevington, JSS 7318, JSS 7321, JSS 7323, JSS 7327, JSS 7329, JSS 7336, JSS 7347 – 8, JSS 7366 – 8, JSS 7372 – 3, JSS 7388 – 9, JSS 7394, JSS 7401, JSS 7404 – 5, JSS 7408 - 9, JSS 7483 (1 ♂, CNC; 20 ♂, QM; 1 ♂, USNM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF45ECF6D8D9EA8FFDA417CC.taxon	distribution	Distribution: Australia (Western Australia) (Fig. 111). Notes: All known specimens were collected at hilltops in Western Australia, most of them at one hilltop north of Denham. See the note for T. minychoma sp. nov. for more information on this remarkable little hilltop. Intraspecific genetic distance ranges 0.5 – 2 %. This species is genetically most similar to T. cona sp. nov. (5.3 – 10.2 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF4BECF2D8D9E871FC851544.taxon	description	Figs 70 A – E, 132, 151 B, 153 A	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF4BECF2D8D9E871FC851544.taxon	diagnosis	Diagnosis: This species can be recognized by hind trochanter with 8 – 12 short spines in a row; hind femur with 15 – 20 long bristles posteroventrally (Fig. 151 B); elongated surstyli in dorsal view (Fig. 70 A); phallic guide with 4 – 6 dorso-dorsolateral spines; one of ejaculatory ducts with 3 – 5 spines in ventral view (Fig. 70 D); elongated gonopods with two small triangular lobes on inner margin (Fig. 70 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF4BECF2D8D9E871FC851544.taxon	description	Redescription: MALE: Body length: 3.0 – 3.1 mm. Head. Flagellum acuminate; dark brown, grey-white pollinose. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for a distance of 9 facets. Occiput silvery pollinose, slightly less so on upper 1 / 3 with scattered short black bristle. Thorax. Postpronotum pale, with 4 – 5 long dark brown bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part greyer with scattered long bristles at anterior supra-alar area. Scutellum brown pollinose and with 6 – 8 short back bristles along the distal edge. Dorsocentral bristles distinct in two rows. Halter, knob pale, stem brown. Legs. Mid coxa with 2 – 3 strong, dark bristles. Trochanters and femora dark brown, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 5 of tibiae yellow-brown (tibiae otherwise dark brown), tarsal segments dark brown, ventrally paler, last segment darker. Hind trochanter grey pollinose with some short spines in a row; ventrobasal spines present on mid and absent on fore femur. Ventroapical row of spines on fore femur 3 – 4; 5 – 6 short dark spines on mid femur; hind femur without ventroapical spines, except 12 – 14 long bristles posteroventrally, bent towards distal tip of femur, longest up to 2 times the width of hind tibia at distal end (Fig. 151 B). Subapical spines on first four tibiae present (short). Hind tibia bent in posterior view. Hind metatarsus and tarsomeres 2 – 3 moderately flattened, metatarsi almost as long as 2 – 5 combined, with scrub-like bristles along posterior edge. Pulvilli shorter than last tarsal segment. Wing. Length: 3.0 mm. Upper side of basal costal cell with two dark bristles, one is longer. Fourth costal section 2 times as long as third costal section. Cross-vein r-m at middle of discal cell. Abdomen. Viewed obliquely from front tergites brown pollinose, tergite 1 silvery grey. Dispersed short dark bristles on all tergites present, longest on tergite 5 up to 1 / 3 of the width of hind tibia at distal end. Lateral bristles on first tergite present, 5 – 8 dark bristles up to as long as hind femur’s width at base. Postabdomen in dorsal view: T 6, S 7 invisible. Genitalia without dissection: ST 8 medium sized, ovate in dorsal view, pale brown and with some distinct bristles; membranous area ovate dorsally, narrow and approaching epandrium ventrally; epandrium brown, surstyli yellow. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.6). Both surstyli elongated, broadened in basal fourth, narrowed in middle, inclined apically to each other; left surstylus is longer and wider than right one (Fig. 70 A). Genital capsule in ventral view: gonopods elongated towards surstyli with two small triangular lobes on inner margin, equal in height; phallic guide with 4 – 6 dorso-dorsolateral spines, one is longer than others, subepandrial sclerite wide; phallus with three ejaculatory ducts, one with 3 – 5 spines (Fig. 70 B); Genital capsule in lateral view: left surstylus straight in basal two thirds, bent towards sternite in apical third; right surstylus curved towards sternite (Fig. 70 D – E). Ejaculatory apodeme tube-like, bent (Fig. 70 C). FEMALE: Body length: 2.9 mm. As male except for the following characters. Frons, eyes separated, as wide as 1.5 – 1.7 times the width at antennae; completely silver-grey pollinose, only shiny black directly in front of ocellar triangle, ca. 1 / 6 of the frons, very short transition zone; frons with a short ridge in front of the ocelli in the median line. Enlarged ommatidia, ocellar triangle shining. Pulvilli and claws about two times as long as last tarsal segment, smaller than metatarsi; Upper side of basal costal cell with one dark bristle and two shorter bristles. Female abdomen with silver spots on sides of tergite 3 – 6. Ovipositor. Straight in ventral view, robust and long, curved upwards in lateral view (yellow piercer, dark brown base), reaching distal end of 2 nd segment; base grey pollinose with occasional brown bristles. LP: LB = 2.6. LDP: LPP = 2.7. (Fig. 153 A). Examined material: Type material examined: PARATYPE: NEW ZEALAND: Cass, 6. XII. 1924, A. Tonnoir, 639, JSS 50572 (1 ♂, CMNZ); Other material examined: AUSTRALIA: Australian Capital Territory: Black Mountain Reserve, 35 ° 16 ’ S, 149 ° 6 ’ E, 4. I. 1980, Z. Liepa, JSS 9187 (1 ♂, ANIC); Black Mountain, 35 ° 16 ’ S, 149 ° 6 ’ E, 25. X. 1981, Z. Liepa, JSS 9188 (1 ♂, ANIC); Black Mountain, Canberra, 35 ° 16 ’ S, 149 ° 6 ’ E, 18 – 23. II. 1985, W. W. Middlekauff, flight interception trap, JSS 8455 (1 ♂, CAS); Queensland: Kroombit Tops National Park, hilltop, 24 ° 27 ’ N, 150 ° 56 ’ E, 21. XII. 2014, J. H., A. M. & A. W. Skevington, CNC 384418 (1 ♂, CNC); Mount Moffatt summit, 25 ° 4 ’ S, 148 ° 3 ’ E, 1097 m, 18. XI. 2014, J. H., A. M. & A. W. Skevington, CNC 374958, CNC 374981 (2 ♂, CNC); Brisbane, Mount Coot-tha, 27 ° 29 ’ S, 152 ° 57 ’ E, 170 m, hilltop, 7. III. 1998, J. & A. Skevington, hand collected, JSS 3633 (1 ♂, QM); Victoria: Wyperfeld NP, East Lookout, 35 ° 36 ’ S, 142 ° 7 ’ E, 21. X. 2014, 22. X. 2014, J. H., A. W., & A. M. Skevington, A. D. Young, A. Wyatt, & S. Namek, CNC 373276 – 81, CNC 373283 – 91, CNC 373395 – 401, CNC 373403, CNC 373405 – 7, CNC 373409 – 19, CNC 373421, CNC 373423, CNC 373425, CNC 373427 – 37, CNC 373440 – 56, CNC 373458 – 62, CNC 373541 (62 ♂, CNC; 10 ♂ QM; 2 ♂, USNM); Tasmania: 3 km south of Spreyton, 41 ° 15 ’ S, 146 ° 20 ’ E, dry sclerophyll forest, 21. XII. 0 – 9. I. 1999, J. & A. Skevington, Malaise trap, JSS 5057 (1 ♂, QM); Near Mertha Park, 41 ° 14 ’ S, 147 ° 11 ’ E, [in dry sclerophyll forest], 14. XII. 1998 – 9. I. 1999, J. & A. Skevington & S. Winterton, Malaise trap, JSS 5074 (1 ♀, QM); Victoria: Dart / Mitta River Junction, Locality L, 36 ° 31 ’ S, 147 ° 27 ’ E, 23 – 25. X. 1973, Dartmouth Survey, JSS 8749 (1 ♂, MVMA); Dartmouth Survey, Eight Mile Creek, Locality JA, 37 ° 13 ’ S, 146 ° 26 ’ E, 12. IV. 1973, JSS 8764 (1 ♂, MVMA); Warrandyte, 37 ° 44 ’ S, 145 ° 13 ’ E, 18. XI. 1928, G. F. Hill, JSS 8676 (1 ♂, ANIC); Hattah-Kulkyne NP, Warepil Lookout, 34 ° 43 ’ S, 142 ° 18 ’ E, 18. X. 2014, J. H., A. W., & A. M. Skevington, A. D. Young, A. Wyatt, & S. Namek, CNC 373059, CNC 373063, CNC 373153, CNC 373162, CNC 373170, CNC 373173 – 4, CNC 373181 (4 ♂, CNC; 4 ♂, USNM); Wyperfeld National Park, Eastern Lookout, 35 ° 36 ’ S, 142 ° 7 ’ E, 22. X. 2014, J. H., A. M. & A. W. Skevington, CNC 383849 – 70 (22 ♂, CNC); New South Wales: 14 km NW of Braidwood, 35 ° 26 ’ S, 149 ° 48 ’ E, 4. XI. 1975, D. H. Colless, JSS 8522 (1 ♂, ANIC); 8 km S Mendooran, 31 ° 50 ’ S, 149 ° 6 ’ E, 28. I. 1987, G. & A. Daniels, JSS 8912 (1 ♂, QM); Cudgegong River, 16 km N Mudgee, 32 ° 48 ’ S, 149 ° 49 ’ E, 30. XI. 1982, D. S. Horning, Jr., Malaise trap, JSS 9156 (1 ♂, ANIC); Eucumbene Dam, 36 ° 6 ’ S, 148 ° 36 ’ E, 18. XI. 1962, D. H. Colless, JSS 8665 (1 ♂, ANIC); Jindabyne, 36 ° 25 ’ S, 148 ° 37 ’ E, 21. II. 1969; 22. II. 1969, Neboiss, JSS 8780; JSS 8787 – 88 (3 ♂, MVMA); Royalla, 35 ° 31 ’ S 149 ° 9 ’ E, 29. III. 1961, D. H. Colless, JSS 8639 (1 ♂, ANIC); Warrumbungle National Park, Buckley’s Creek, 31 ° 17 ’ S, 149 ° 0 ’ E, 12 – 17. XII. 1995, Sharkey, CNCD 3594 (1 ♂, CNC); Warrumbungle National Park, 31 ° 14 ’ S 149 ° 1 ’ E, 19. X – 3. XI. 1997, J. Skevington & S. Winterton, Malaise trap, JSS 8438 (1 ♂, QM); Wombat, near Young, 34 ° 25 ’ S, 148 ° 15 ’ E, 13. XI. 1966, Z. Liepa, JSS 9211 (1 ♂, ANIC); South Australia: Vivonne Bay, Kangaroo Island, 35 ° 45 ’ S, 137 ° 37 ’ E, II. 1926, Museum Expedition, JSS 8722 (1 ♂, SAM); Western Australia: Charles Darwin Reserve, Edna Springs, 29 ° 30 ’ S, 117 ° 3 ’ E, Granite sheet, Acacia flowering herbs, 14 – 19. IX. 2009, Lambkin & Monteith, Malaise trap, 18309, CNC 591802 (1 ♂, QM); Fitzgerald River National Park, Old Ongerup Road near Hammersley River, 33 ° 45 ’ S, 119 ° 40 ’ E, ANIC Bulk Sample 2189, 219 m, mallee granite outcrop, 2 – 16. XI. 2003, J. Recsei & C. Lambkin, Malaise trap, ANIC Bulk Sample 2189, JSS 16210 (1 ♂, ANIC); Karara, 16.9 km Southeast of Bolada Camp, 29 ° 15 ’ S, 116 ° 41 ’ E, 312 m, Acacia woodland, flowering herbs, 18 – 24. IX. 2009, Lambkin & Monteith, Malaise trap, 18407, CNC 575347 (1 ♂, QM); Thomas River estuary, Esperance District, 33 ° 52 ’ S, 123 ° 1 ’ E, 4. XI. 1977; 8. XI. 1977, D. H. Colless, Malaise trap, JSS 9226, JSS 9231 (2 ♂, ANIC); Charles Darwin Reserve, Edna Springs, 29 ° 30 ’ S, 117 ° 3 ’ E, Granite sheet, Acacia, flowering herbs, 19 – 23. IX. 2009, Lambkin, Monteith, Malaise trap, 18439, CNC 575119 (1 ♂, QM); Lochada, 430 m S Bolada Camp, 29 ° 12 ’ S, 116 ° 31 ’ E, 288 m, Acacia Eremophila scrub, 15 – 16. IX. 2009, Lambkin, Monteith, Malaise trap, CNC 576392 (1 ♂, QM); Northern Territory: Kakadu National Park, Mirrai Lookout, 12 ° 52 ’ S, 132 ° 42 ’ E, hilltop, 27. IX. 1998, J. & A. Skevington, hand collected, JSS 4136 (1 ♂, QM); NEW ZEALAND: Goldney Saddle, 43 ° 16 ’ S, 171 ° 35 ’ E, 671 m, 1. II. 1962, E. G. White, sweep net, JSS 14524 – 5 (2 ♂, LUNZ); Lincoln College, 43 ° 39 ’ S, 172 ° 29 ’ E, ex. weeds, 9. I. 1969, M. G. McPherson, sweep net, JSS 14519 (1 ♂, LUNZ); NC, Arthur Pass National Park, Arthur Pass Village, 42 ° 57 ’ S, 171 ° 34 ’ E, 750 m, meadow, 5 – 7. III. 2000, H. Goulet, yellow pan trap, CNCD 137560 (1 ♂, CNC); Northland: Castaway Camp area, Great I., Three King Is., D. S. I. Q. Exp., 34 ° 9 ’ S, 172 ° 8 ’ E, 120 m, XI. 1973, bleached by alcohol, JSS 14130 (1 ♂, NZAC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF4BECF2D8D9E871FC851544.taxon	distribution	Distribution: Australia (Australian Capital Territory, New South Wales, Northern Territory, Queensland, South Australia, Tasmania, Victoria, Western Australia), New Zealand (North Island, Northland, South Island, Three Kings Island) (Fig. 132). Notes: Widespread in forested regions of Australia and New Zealand as far north as Kroombit Tops in Queensland. Intraspecific genetic distance is 0.2 %. This species is genetically most similar to T. velutina sp. nov. (3.9 – 6.0 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF4EECCED8D9EAE3FC6E108C.taxon	diagnosis	Diagnosis: This species can be recognized by short surstyli curved towards each other in dorsal view (Fig. 71 A); long subepandrial sclerite; elongated gonopods, right one with a triangular projection; distinct hypandrial apodeme in ventral view (Fig. 71 B); phallic guide with dense hairs dorsomedially in lateral view (Fig. 71 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF4EECCED8D9EAE3FC6E108C.taxon	description	Redescription: MALE: Body length: 2.5 – 2.8 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black, longer than usual; lower part distinctly silver pollinose, shorter than usual; eyes touching for distance equal to 1 – 1.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 3 – 4 pale bristles. Mesonotum (viewed obliquely from front) shiny brown, without greyish pollinosity. Scutellum shiny with 4 – 6 weak bristles along the distal edge. Dorsocentral bristles distinct, pale, somewhat longer along frontal edge, pale bristles around postpronotum absent. Halter, knob pale with dark brown spot at tip, stem brown. Legs. Mid coxa with 3 long dark bristles (one is shorter). Trochanters pale brown with some yellow distally, femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal ¼ of tibiae yellow (tibiae otherwise brown), tarsal segments yellow-brown, ventrally somewhat paler, last segment darker. Hind trochanter with no special features, in posterior view it looks like a half circle, straight on dorsal side; ventrobasal spines absent on fore and mid femur. Ventroapical row of spines on fore femur 3 – 4; 6 – 7 small, black spines on mid femur; hind femur without ventroapical spines, except 20 – 22 equally spaced bristles posteroventrally, longer on the distal half, longest at 2 / 3 from the base, in length up to the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present (short). Hind tibia slightly bent in posterior view. Hind metatarsus slightly flattened and almost as long as 2 – 5 combined and with scrub-like bristles ventrally. Pulvilli shorter than last tarsal segment. Wing. Length: 2.5 – 2.6 mm. Upper side of basal costal cell with one long brown bristle. Fourth costal section 1.5 – 1.7 times as long as third costal section (fourth section appears to be very short). Cross-vein r-m at proximal 1 / 3 of discal cell, sometimes more proximal. 2 – 3 distinct brown setulae on tegula. Abdomen. Viewed obliquely from front tergites shining brown, tergite 1 silvery grey, no silvery pollinose on tergites. Dispersed strong dark bristles on all tergites present, longest on tergite 5 up to 0.5 times the width of hind tibia at distal end. Lateral bristles on first tergite present, 2 – 3 weak bristles up to 0.4 – 0.5 times as long as width of hind femur at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1.8 – 2.2 times as long as ST 8. Genitalia without dissection: ST 8 medium sized rounded in dorsal view, brown and with the occasional distinct bristles, otherwise velvet-like coverage; membranous area round, directed posteriorly; epandrium brown (sometimes yellow distally), surstyli pale brown and short. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.7). Surstyli symmetrical and short, widened at base, moderately narrowed in middle, both surstyli inclined together (Fig. 71 A). Genital capsule in ventral view: phallic guide straight and slender, gonopods equal in height, elongated towards surstyli, right one with a triangular projection, subepandrial sclerite long, widened at base; hypandrial apodeme distinct (Fig. 71 B). Genital capsule in lateral view: right surstylus straight, bent towards sternite in apical fourth; left surstylus straight in ventral margin; both surstyli more broadened at middle; phallic guide pointed at apex with dense hairs dorsomedially (Fig. 71 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 71 C). FEMALE: Body length: 2.5 mm. As male except for the following characters. Frons, eyes separated, as wide as 1.5 times the width at antennae; completely silver-grey pollinose until up to 1 / 3 of the lower frons, shiny black in front of ocellar triangle, in the transition zone, pollinose area grades to shiny black; frons slightly depressed at the point where it is widest. Enlarged ommatidia silvery shining. Pulvilli and claws about 2 – 2.5 times as long as last tarsal segment, smaller on metatarsi. Female abdomen with no silver spots on sides of tergites. Ovipositor. Base of ovipositor dark brown with some scattered short dark bristles, piercer straight. LP: LB = 3.3. LDP: LPP = 3.5. (Fig. 154 B). Examined material: Type material examined: LECTOTYPE [HERE DESIGNATED]: AUSTRALIA: Queensland: Bundaberg, 24 ° 52 ’ S, 152 ° 21 ’ E, vi. 1904, CNCD 5532, (1 ♂, BPBM); PARALECTOTYPES: AUSTRALIA: same data as lectotype, BMNH (E) 964034, CNCD 5531, CNCD 5533 (1 ♂, BMNH; 1 ♂, 1 ♀, BPBM); North Queensland, 16 ° 54 ’ S, 145 ° 45 ’ E, JSS 4168 (1 ♂, ZMAN); Other material examined: AUSTRALIA: New South Wales: Cabbage Tree Creek, Clyde Mountain, 35 ° 35 ’ S, 150 ° 1 ’ E, 1. II. 1973, D. H. Colless, JSS 8553 (1 ♂, ANIC); Coocumbac Island Nature Reserve, Taree, 31 ° 54 ’ S, 152 ° 27 ’ E, ex. floodplain, rainforest, 21 – 30. XI. 1994, G. & T. Williams, [specimen badly faded], JSS 15984 (1 ♂, QM); Bruxner Park, 30 ° 14 ’ S, 153 ° 6 ’ E, 19. IV. 1970, D. H. Colless, “ site (1) & (2) see field notes ”, JSS 8673 (1 ♂, ANIC); Brunswick Heads, 28 ° 32 ’ S, 153 ° 33 ’ E, 22. v. 1966, Z. Liepa, JSS 9208 (1 ♂, ANIC); Coocumbac Island Nature Reserve, Taree, 31 ° 55 ’ S, 152 ° 27 ’ E, floodplain rainforest, 24 – 25. x. 1994, G. & T. Williams, JSS 8730 (1 ♂, AMS); Nelligen (tidal flat), 35 ° 38 ’ S, 150 ° 8 ’ E, 1. ii. 1973, D. H. Colless, JSS 8498 (1 ♂, ANIC); Pisgah Ridge near Glenbrook, Blue Mountains National Park, 33 ° 50 ’ S, 150 ° 33 ’ E, 210 m, 5. ii – 25. iii. 1988, G. R. Brown, Malaise trap, JSS 8747 (1 ♀, ASCU); Royal National Park, E waterfall rail. stat., 34 ° 9 ’ S, 151 ° 1 ’ E, diff. types forest, 12. x. 2002, leg. B. Merz / 10, JSS 14362 (1 ♂, MHNG); Sydney, Centennial Park, 33 ° 53 ’ S, 151 ° 13 ’ E, ex. eucalypt forest, 9. x. 1994, B. J. Sinclair, [badly eaten out by dermestids], JSS 9686 – 7 (2 ♂, CNC); Toukley, 33 ° 16 ’ S, 151 ° 32 ’ E, 18. i. 1963, Z. Liepa, JSS 9233 (1 ♂, ANIC); Northern Territory: 5 km NNW of Cahills Crossing, East Alligator River, 12 ° 5 ’ S, 132 ° 35 ’ E, 9. VI. 1973, D. H. Colless, JSS 8658 (1 ♂, ANIC); 22 km WSW Borroloola, 16 ° 8 ’ S, 136 ° 7 ’ E, 17. iv. 1976, D. H. Colless, at light, JSS 9212 (1 ♂, ANIC); Barramundi Creek, 13 ° 19 ’ S, 132 ° 26 ’ E, woodland, 13. v. 1998, R. Lynch, Malaise trap, JSS 7750 (1 ♂, QM); woodland, 14. v. 1998, R. Lynch, Non-Riparian Zone; Malaise trap 3, JSS 7904 (1 ♂, QM); Koolpin Crossing, [South Alligator River], 13 ° 33 ’ S, 132 ° 33 ’ E, riparian zone, 20. v. 1998, R. Lynch, Malaise trap 3, JSS 7906 (1 ♂, QM); Queensland: 1 km N of Eclectus, Iron Range, 12 ° 46 ’ S, 143 ° 17 ’ E, 20 m, 10. VII. 1997, G. and A. Daniels, JSS 50571 (1 ♂, QM); 2 km East Almaden, 17 ° 20 ’ S, 144 ° 41 ’ E, 22. IV. 1983, J. F. Donaldson & J. F. Grimshaw, D. vac, JSS 8958 (1 ♂, QDPC); 2 km SE Drillham, 26 ° 38 ’ S, 150 ° 0 ’ E, along creek, 8. X. 1974, I. D. Galloway, JSS 8940 (1 ♂, QDPC); 20 km E Mareeba, 17 ° 0 ’ S, 145 ° 26 ’ E, 15. IV. 1980, G. F. Hevel & J. A. Fortin, JSS 10522 (1 ♂, USNM); 3 km NE of Mount Webb, 15 ° 3 ’ S, 145 ° 9 ’ E, 30. IV. 1981, D. H. Colless, Malaise trap, JSS 8609 (1 ♂, ANIC); Acacia Ridge, 27 ° 35 ’ S, 153 ° 1 ’ E, 23. IX. 1962, E. B. Teh, JSS 103 (1 ♂, QM); Birthday Creek trail near Paluma, 18 ° 59 ’ S, 146 ° 9 ’ E, 774 m, 22. XI. 2014, J. H., A. M. & A. W. Skevington, CNC 374674 (1 ♂, CNC); Camp Mountain, SE Queensland, 27 ° 24 ’ S, 152 ° 53 ’ E, open sclerophyll gully, 27. XI – 7. XII. 1979, Marks, Malaise trap, JSS 8930 (1 ♂, QDPC); Carnarvon Station (CN 2 M 1), 24 ° 49 ’ S, 147 ° 45 ’ E, 757 m, brigalow on grassy hillside, 24. XI – 13. XII. 2010, C. Zwick, Malaise trap, CNC 575279 (1 ♂, QM); Maroochy Horticulture Research Station, Nambour, SE Queensland, 26 ° 37 ’ S, 152 ° 57 ’ E, 15 – 22. III. 1985, Malaise trap, JSS 8949 – 50 (2 ♂, QDPC); Prince of Wales Island, Torres Strait, 10 ° 41 ’ S, 142 ° 11 ’ E, 29. V. 1969, Neboiss, JSS 8784 (1 ♂, MVMA); Connor’s River, 99.7 km SWS Sarina, 22 ° 41 ’ S, 149 ° 7 ’ E, 5. vii. 1971, Z. Liepa, JSS 9161 (1 ♂, ANIC); Mount Coot-tha foothills, 27 ° 29 ’ S, 152 ° 57 ’ E, 26. vi. 1985, Z. Liepa, JSS 9203 – 4 (2 ♂, QM); Mount Glorious, Scrub Creek Road, Brisbane Forest Park, 27 ° 25 ’ S, 152 ° 50 ’ E, 9 – 16. i. 1998, N. Power, Malaise trap, JSS 7778 (1 ♂, QM); Pine Mountain, summit, central Queensland, 21 ° 45 ’ S, 148 ° 50 ’ E, 600 m, vine scrub, 5 – 6. x. 1999, C. J. Burwell, S. G. Evans, Malaise trap, JSS 9145 (1 ♂, QM); Proserpine, Lilly pool Site: XY 18, 16 ° 52 ’ S, 179 ° 53 ’ W, 30 m, 15 – 22. viii. 2007, J. Stanisic, Malaise trap, 15048, CNCD 159558 (1 ♀, CNC); West Burleigh, 28 ° 6 ’ S, 153 ° 26 ’ E, 4. v. 1970, Z. Liepa, JSS 9175 – 6 (2 ♂, ANIC); 4 km up Black Mountain Road via Kuranda, 16 ° 49 ’ S, 145 ° 38 ’ E, 14. IX – 12. X. 1982, Malaise trap, JSS 8968 (1 ♂, QDPC); Brisbane Forest Park, Enoggera Creek at Scrub Road Crossing, 27 ° 26 ’ S, 152 ° 50 ’ E, 200 m, 6 – 9. XI. 1995, M. E. Irwin, CNCD 147459 (1 ♂, CNC); Kuranda, 15 km NW Cairns, 16 ° 49 ’ S, 145 ° 38 ’ E, tropical rainforest and riverine habitat, 9. X. 1983, E. I. Schlinger & M. E. Irwin, JSS 3944 (1 ♂, INHS); Brisbane Forest Park, Scrub Creek, 27 ° 26 ’ S, 152 ° 50 ’ E, 28. IX – 15. X. 2002, J. Skevington, J. M. Cumming, Malaise trap, JSM 564, JSS 13115 (1 ♂, CNC); Lamington National Park, 28 ° 12 ’ S, 153 ° 10 ’ E, 25. V. 1962, J. Cribb, JSS 8250 (1 ♂, QM); Mareeba Shire, Kuranda, Russet Park, North Queensland, 16 ° 49 ’ S, 145 ° 38 ’ E, 460 m, 22. X. 1987; 28. X. 1987; 30. X. 1987, Thomas W. Davies, flight interception trap, Thomas W. Davies Collection donated to California Academy of Sciences 1987, JSS 8465, JSS 8467, JSS 8471, JSS 8473 (4 ♂, CAS); Nairana National Park (NR 2 M), 21 ° 34 ’ S, 146 ° 55 ’ E, 196 m, closed Acacia woodland, 10. XI – 7. XII. 2010, R. Raven, H. & D. Hanrahan, Malaise trap, CNC 576499, CNC 576504 (2 ♂, QM); near Teddington Weir S of Maryborough, 25 ° 39 ’ S, 152 ° 40 ’ E, 25. IX. 1994, G. & A. Daniels & C. J. Burwell, JSS 5139 (1 ♂, QM); Australian Capital Territory: Black Mountain, 35 ° 16 ’ S, 149 ° 6 ’ E, sclerophyll gully, 20 – 31. III. 1980, B. K. Cantrell, Malaise trap, JSS 8945 (1 ♂, QDPC); Victoria: Dartmouth Survey, Dart-Mitta River Junction, Locality L, 36 ° 31 ’ S, 147 ° 27 ’ E, 4. III. 1973, JSS 8765 (1 ♂, MVMA); INDONESIA: West Java: Bogor Botanical Gardens (Kebon Raja) [Kebun Raya], 6 ° 36 ’ S, 106 ° 48 ’ E, 3. VIII. 1964; 4. VIII. 1964, M. Delfinado, CNCD 5526 – 27 (1 ♂, 1 ♀, BPBM); Bogor, West Java, 6 ° 35 ’ S, 106 ° 48 ’ E, 22. VIII. 1964, J. L. Gressitt, CNCD 5525 (1 ♂, BPBM); Mussau, Boliu, 1 ° 32 ’ S, 149 ° 39 ’ E, 3. VI. 1962, Noona Dan Expedition 61 – 62, CNCD 5529 (1 ♂, BPBM); Mussau, Schadel Bay, 1 ° 30 ’ S, 149 ° 40 ’ E, 3. VI. 1962, Noona Dan Expedition 61 – 62, CNCD 5530 (1 ♂, BPBM); Yalom, 4 ° 25 ’ S, 151 ° 45 ’ E, 1000 m, 20. V. 1962, Malaise trap, Noona Dan Expedition 61 – 62, CNCD 5528 (1 ♂, BPBM); MALAYSIA: Sarawak: SW of Tapuh, Borneo, 1 ° 18 ’ N, 110 ° 25 ’ E, 4 – 9. VII. 1958, T. C. Maa, CNCD 5523 – 5524; JSS 8478 (3 ♀, BPBM); Sadong, Kampong, Tapuh, Borneo, 1 ° 10 ’ N, 110 ° 34 ’ E, 450 m, 4 – 9. VII. 1958, T. C. Maa, CNCD 5522; JSS 8477 (2 ♂, BPBM); NEW CALEDONIA: Noumea, 22 ° 16 ’ S, 166 ° 26 ’ E, 22 – 23. XI. 1963, R. Straatman, CNCD 5607 (1 ♂, BPBM); Tao, 20 ° 34 ’ S, 164 ° 48 ’ E, 10. II. 1963; 8 – 10. II. 1963, C. Yoshimoto and N. Krauss, Malaise trap, CNCD 253913; CNCD 5611 (2 ♂, BPBM); Yahoue, 22 ° 11 ’ S, 166 ° 30 ’ E, 20. II. 1963, C. Yoshimoto and N. Krauss, Malaise trap, CNCD 5615 (1 ♂, BPBM); PAPUA NEW GUINEA: East New Britain: Bainings Mountains: Raunsepna, 4 ° 26 ’ S, 151 ° 47 ’ E, 19 – 26. V. 1999, C. Mitparingi, Malaise trap, CNCD 29005; CNCD 28999; CNCD 29004; CNCD 29007 (4 ♂, CNC); West New Britain: Lindenhafen, S coast, 6 ° 15 ’ S, 150 ° 28 ’ E, 2 m, Cocoa grove, 23. IV. 1956, J. L. Gressitt, JSS 5521 (1 ♀, BPBM); Maprik, 3 ° 38 ’ S, 143 ° 3 ’ E, 21. III. 1964, D. H. Colless, JSS 9266 (1 ♂, ANIC); Woitape, Wharton Range, 19. X. 1963, D. K. McAlpine, JSS 8732 (1 ♂, AMS); PHILIPPINES: Palawan: Mantalingajan, Pinigisan, 8 ° 49 ’ N, 117 ° 40 ’ E, 600 m, outside forest, 9. IX. 1961, G. A. Samuelson, Malaise trap, Noona Dan Expedition 61 – 62, JSS 5520 (1 ♂, BPBM); THAILAND: Chanthaburi: Prew, 12 ° 37 ’ N, 102 ° 8 ’ E, 25 – 30. IV. 1958, T. C. Maa, CNCD 5517 (1 ♂, BPBM); Trang: Khaophappha, Khaochang, 7 ° 32 ’ N, 99 ° 36 ’ E, 200 to 400 m, 2. I. 1964, G. A. Samuelson, light trap, JSS 5518 – 19 (2 ♂, BPBM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF4EECCED8D9EAE3FC6E108C.taxon	distribution	Distribution: Australia (Australian Capital Territory, New South Wales, Northern Territory, Queensland, Victoria), Indonesia (Java), Malaysia (Sarawak), New Caledonia, Papua New Guinea (New Britain), Philippines (Palawan), Thailand) (Fig. 133). Notes: Based on terminalia characters, some specimens from Thailand (CNCD 5517, JSS 5518 – 19), Philippines (JSS 5520), Malaysia (CNCD 5522 – 4, JSS 8477 – 8), Indonesia (CNCD 5525 – 30), New Caledonia (CNCD 5607, CNCD 5611, CNCD 5615, CNCD 253913) and New Guinea (JSS 5521, CNCD 28999, CNCD 29004 – 5, CNCD 29007, JSS 9266) are very close to Australian T. nyctias. Among New Guinea specimens, the specimen JSS 8732 appears identical to the lectotype whereas Indonesian specimens are the most different. The latter present a few differential genitalia features, including the shape of surstyli in dorsal view, left surstylus in lateral view and the shape of gonopods. As these specimens from outside of Australia are too old to easily sequence, we leave them as T. nyctias until further study provides fresh specimens for DNA sequencing. Specimen CNCD 5532 is hereby nominated as the lectotype to fix and ensure consistent interpretation of the name. Tomosvaryella nyctias is genetically most similar to T. luma sp. nov. (8.7 – 11.1 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF72ECCED8D9EE2BFD5417A8.taxon	description	Figs 72 A – E, 111, 155 C	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF72ECCED8D9EE2BFD5417A8.taxon	diagnosis	Diagnosis: This species can be recognized by elongated surstyli that are moderately narrowed to the apex (Fig. 72 A); unequal gonopods, right gonopod shorter with two triangle-shaped projections on inner margin in ventral view (Fig. 72 B); distinct and long hypandrial apodeme (Fig. 72 B); phallic guide with 3 – 5 dorsolateral spines (Fig. 72 B, D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF72ECCED8D9EE2BFD5417A8.taxon	description	Description: MALE: Body length: 3.1 mm. Head. Flagellum short acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 2 – 3 pale bristles. Mesonotum (viewed obliquely from front) grey pollinose, anterior part more intensely pollinose, grayish also from the side. Scutellum silvery pollinose and with 6 – 8 bristles along the distal edge. Dorsocentral bristles short, dark, somewhat longer along frontal edge, numerous pale bristles around postpronotum present. Halter, knob pale, stem brown. Legs. Mid coxa with two dark bristles, one longer than the other. Trochanters and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments yellow-brown dorsally, ventrally yellow, last segment darker. Hind trochanter with a half circle (flat part dorsally), no distinct bristles; ventrobasal spines absent on fore and mid femur. Ventroapical row of spines on fore femur absent; 4 – 5 small, black, spines on mid femur; hind femur without ventroapical spines. Subapical (distal) spines on first four tibiae absent. Hind metatarsus distinctly flattened (tarsomeres 1 – 3 most flattened), first tarsomere slightly longer than 2 – 4 combined, with scrub-like bristles ventrally (tarsomeres 1 – 2 only). Pulvilli shorter than last tarsal segment. Wing. Length: 3.2 mm. Upper side of basal costal cell with one long dark bristle. Fourth costal section 2.2 times as long as third costal section. Cross-vein r-m distinctly distal to middle of discal cell. 5 – 6 distinct brown setulae on tegula. Abdomen. Dissected. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.4). Surstyli rather symmetrical, elongated, moderately narrowed to apex, left surstylus longer than right one, right surstylus bent to left one before apex (Fig. 72 A). Genital capsule in ventral view: gonopods unequal in height, left is longer, right is wider, right gonopod with two different-sized triangular projections on inner margin, left gonopod with one smaller triangular projection; distinct, long hypandrial apodeme; phallic guide with 3 – 5 dorso- dorsolateral spines, one is longer, distinct subepandrial sclerite (Fig. 72 B); Genital capsule in lateral view: both surstyli broadened at base, moderately narrowed to apex, curved towards sternite, phallic guide strong, pointed towards sternite at apex (Fig. 72 D – E); ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 72 C). FEMALE: Body length: 2 mm (without head). As male except for the following characters. Frons, eyes: NO HEAD. Pulvilli and claws about 2 – 2.8 times as long as last tarsal segment, smaller on metatarsi. Female abdomen with silver spots on sides of tergite 5 only. Ovipositor. Straight in ventral view, strongly bent (90 degrees at the base of piercer) in lateral view (yellow piercer, brown base), reaching proximal end of 5 th segment; base silvery pollinose with occasional bristles. LP: LB = 2.4. LDP: LPP = 4.0. (Fig. 155 C).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF72ECCED8D9EE2BFD5417A8.taxon	etymology	Etymology: This species is named in honour of Frank D. Parker who collected both known specimens with Mike Irwin. Frank was involved in the collection of 6 % of the specimens and 15 % of the species used in this paper (Supplementary file 2). Examined material: HOLOTYPE: AUSTRALIA: Western Australia: 47 km S Pardoo R [oa] d House on Shay Gap R [oa] d, 20 ° 23 ’ S, 120 ° 1 ’ E, 170 m, dry wash n [ea] r Spinifex, 1 – 14. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15702 (1 ♂, WAM); PARATYPE: AUSTRALIA: Western Australia: same data as holotype, JSS 15704 (1 ♀, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF72ECCED8D9EE2BFD5417A8.taxon	distribution	Distribution: Australia (Western Australia) (Fig. 111). Note: Intraspecific genetic distance is 0.0 %. This species is genetically most similar to T. quadrata sp. nov. (7.5 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF70ECCDD8D9EFF6FCAD14B0.taxon	description	Figs 73 A – E, 114, 150 C	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF70ECCDD8D9EFF6FCAD14B0.taxon	diagnosis	Diagnosis: This species can be recognized by the hind femur with a small triangular keel on the posterior side close to the base (Fig. 150 C); tiny surstyli, widened at base, curved in basal two thirds and straight in apical third in dorsal view (Fig. 73 A); gonopods extended towards surstyli, widened at the apex in ventral view (Fig. 73 B); phallic guide with 3 – 4 short dorsolateral spines in ventral view (Fig. 73 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF70ECCDD8D9EFF6FCAD14B0.taxon	description	Description: MALE: Body length: 2.4 mm. Head. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2.2 times the length of ocellar triangle. Occiput silvery pollinose. Thorax. Postpronotum pale, with 2 – 3 short pale bristles. Mesonotum (viewed obliquely from front) silvery pollinose, anterior part greyer. Scutellum silvery pollinose and with 6 – 8 short pale bristles along distal edge. Halter, knob and stem pale, base brownish. Legs. (Fore and mid legs missing beyond coxa). Mid coxa with 3 – 4 long brown bristles. Trochanters brown, hind femur black, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of hind tibia yellow (tibiae otherwise brown), tarsal segments bright yellow. Hind trochanter gray pollinose with scattered short pale bristles. Hind femur with ventroapical spines. Hind femur with small triangular keel (shark fin) on the posterior side close to base (Fig. 150 C). Hind tarsomeres not flattened, hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 2.5 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2 – 3 short brown setulae on tegula. Abdomen. Viewed obliquely from front tergites brown-black, except tergite 1 silvery grey. Lateral bristles on first tergite present, 6 – 8 dark bristles up to as long as ¾ of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.1). Surstyli small, widened at the base, then straight and bent to each other at the apex (Fig. 73 A). Genital capsule in ventral view: both gonopods extended towards surstyli, widened at apex, with a distinct projection on inner margin, pulled towards each other; phallic guide with 3 – 4 short dorsolateral spines, one longer than the other; subepandrial sclerite distinct with two edges in lateral sides (Fig. 73 B); Genital capsule in lateral view: both surstyli widened at the base, curved in basal two thirds, narrow and straight in apical third (Fig. 73 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 73 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF70ECCDD8D9EFF6FCAD14B0.taxon	etymology	Etymology: From the Latin parvus, little, in reference to the tiny surstyli. Examined material: HOLOTYPE: AUSTRALIA: New South Wales: Culgoa National Park, 8 km West-Northwest Cawwell Homestead, Diemunga Lagoon (CGN 2 M), 29 ° 3 ’ S, 146 ° 60 ’ E, Coolibah, 30. I – 18. V. 2010, C. Lambkin, R. Olsen & B. Shieban, Malaise trap, CNC 575122 (1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF70ECCDD8D9EFF6FCAD14B0.taxon	distribution	Distribution: Australia (New South Wales) (Fig. 114).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF71ECCAD8D9EA77FD4F1770.taxon	description	Figs 74 A – E, 134, 149 D, 155 D	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF71ECCAD8D9EA77FD4F1770.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a keel covered by short dark spines on the edge (Fig. 149 D); hind metatarsus distinctly flattened; elongated surstyli, left surstylus with a lobe on inner margin in dorsal view (Fig. 74 A); both gonopods with a triangular projection on inner margin; phallic guide with 3 – 5 dorsolateral spines (Fig. 74 B, D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF71ECCAD8D9EA77FD4F1770.taxon	description	Description: MALE: Body length: 2.9 – 3.1 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 3 – 4 pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum brownish pollinose and with 6 – 8 bristles along the distal edge. Dorsocentral bristles short, dark, somewhat longer along frontal edge, numerous pale bristles around postpronotum present. Halter, knob pale, stem brown. Legs. Mid coxa with 2 – 3 long dark bristles. Trochanters and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 4 of tibiae yellow (tibiae otherwise brown), tarsal segments yellow, ventrally paler, last segment brown. Hind trochanter with shiny edge (keel) at 45 degrees to the long axis of trochanter, with short dark spines on the edge (Fig. 149 D); weak ventrobasal spines (2) present on fore and missing on mid femur. Ventroapical row of spines on fore femur absent; 4 – 5 small, black spines on mid femur; hind femur without ventroapical spines, except 8 – 10 equally spaced bristles on distal half posteroventrally on a widened femur in the middle (almost like a small keel), longest up to 1.5 times the width of hind tibia at distal end (sometimes adpressed to femur). Subapical (distal) spines on first four tibiae present (short). Hind metatarsus distinctly flattened, as long as 2 – 5 combined and with strong bristles dorsally, scrub-like bristles ventrally (tarsomeres 1 – 2 only). Pulvilli shorter than last tarsal segment. Wing. Length: 3.0 mm. Upper side of basal costal cell with one long dark bristle. Fourth costal section 2 – 2.5 times as long as third costal section (Fig. 158 E). Cross-vein r-m at middle of discal cell (sometimes slightly distal to middle). 2 – 3 short brown setulae on tegula. Abdomen. Viewed obliquely from front tergites brownish pollinose, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (spot on t 5 larger). Dispersed strong dark bristles on all tergites present, longest on tergite 5 in length up to 1 / 4 the width of hind tibia at distal end. Lateral bristles on first tergite present, 4 – 6 short dark bristles up to as long as 2 / 3 of hind femur’s width at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 0.5 – 0.7 times as long as ST 8. Genitalia without dissection: ST 8 large, rounded in dorsal view, brown and with a few distinct bristles, otherwise with velvet-like coverage; membranous area with round part extended dorsally and narrow part approaching epandrium, mostly directed posteriorly; epandrium brown, surstyli yellow. Genitalia. Genital capsule in dorsal view: epandrium slightly as long as wide (MLE: MWE = 1.08). Surstyli elongate, left surstylus with a distinct lobe on inner margin in middle and longer than right one, right surstylus with a small denticle on inner margin after basal fourth, both surstyli strongly bent to each other at apex (Fig. 74 A). Genital capsule in ventral view: gonopods equal in height, extended towards surstyli, both gonopods with a triangular projection on inner margin; phallic guide with 3 – 6 dorso-dorsolateral spines, one is longer, distinct subepandrial sclerite (Fig. 74 B); Genital capsule in lateral view: both surstyli broadened at base, rather straight, right surstylus moderately narrowed to apex, left surstylus bent towards sternite before apex (Fig. 74 D – E); ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 74 C). FEMALE: Body length: 2.4 mm (without head). As male except for the following characters. Frons, eyes: NO HEAD. Pulvilli and claws about 2.5 – 3 times as long as last tarsal segment, smaller on metatarsi. Female abdomen with silver spots on sides of tergites 4 – 6. Ovipositor. Straight in ventral view, straight and robust in lateral view (yellow piercer, brown base), reaching middle of 3 rd segment; base silvery pollinose with dispersed bristles. LP: LB = 3.0. LDP: LPP = 5.0. (Fig. 155 D).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF71ECCAD8D9EA77FD4F1770.taxon	etymology	Etymology: From the Latin patulus, spread out, broad, in reference to the distinctly flattened hind metatarsus. Examined material: HOLOTYPE: AUSTRALIA: New South Wales: Warrumbungle National Park, 31 ° 14 ’ S, 149 ° 1 ’ E, 19. X – 3. XI. 1997, J. Skevington, & S. Winterton, Malaise trap, JSS 8439 (1 ♂, QM); PARATYPES: AUSTRALIA: New South Wales: Colo Heights, Putty Road, 33 ° 23 ’ S, 150 ° 45 ’ E, 6. II. 1968, D. H. Colless, JSS 8667 (1 ♂, ANIC); Culgoa National Park, 13.5 km SW Cawwell Homestead (CGN 1 M), 29 ° 7 ’ S, 146 ° 56 ’ E, Callitris, 30. I – 18. V. 2010, QM Team, B. Shiebaan & R. Ohlsen, Malaise trap, CNC 576617 (1 ♂, QM); Culgoa National Park, 8 km West-Northwest Cawwell Homestead, Diemunga Lagoon (CGN 2 M), 29 ° 3 ’ S, 146 ° 60 ’ E, Coolibah, 30. I – 18. V. 2010, C. Lambkin & R. Olsen, B. Shieban, Malaise trap, 19322, CNC 575139 (1 ♂, QM); Bundjalung National Park; near Evans Head, 29 ° 2 ’ S, 153 ° 26 ’ E, 8. I. 1997, C. L. Lambkin, Malaise trap, JSS 6068 (1 ♀, QM); Queensland: Carnarvon National Park, Mount Moffatt Section, The Tombs, 25 ° 5 ’ S, 147 ° 52 ’ E, 10. X. 2002, J. Skevington, hand collected, JSS 12635 (1 ♂, CNC); Carnarvon National Park, Mount Moffatt Summit, 25 ° 4 ’ S, 148 ° 3 ’ E, 1097 m, hilltop, 2. XII. 1997, J. Skevington & C. Lambkin, hand collected, JSS 1821 (1 ♂, QM); Carnarvon National Park, Mount Moffatt Summit, 25 ° 4 ’ S, 148 ° 3 ’ E, 1097 m, hilltop, 13. X. 2002, J. Skevington, JSM 466, CNC 484088 (1 ♀, CNC); Isla Gorge National Park, 25 ° 11 ’ S, 149 ° 58 ’ E, 320 m, hilltop near lookout, 13. XI. 1999, J. & A. Skevington, hand collected, JSS 7256 (1 ♂, QM); Western Australia: West of Norseman, 32 ° 11 ’ S, 121 ° 38 ’ E, Eucalyptus woodland, 1 – 17. XI. 2003, C. Lambkin & J. Recsei, Malaise trap, JSS 16212 (1 ♂, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF71ECCAD8D9EA77FD4F1770.taxon	distribution	Distribution: Australia (New South Wales, Queensland, Western Australia) (Fig. 134). Notes: Intraspecific genetic distance ranges from 0.3 % to 1.5 %. This species is genetically most similar to T. unda sp. nov. (1.6 – 8.3 % pairwise divergence) (Supplementary file 3). This is a forest species that is apparently widespread but rare. It has been found hilltopping.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF74ECC6D8D9EFF6FA1E1168.taxon	description	Figs 75 A – E, 132, 147 B, 154 A	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF74ECC6D8D9EFF6FA1E1168.taxon	diagnosis	Diagnosis: This species can be recognized by the blunt peg-like projection ventrobasally on the hind trochanter (Fig. 147 B); surstyli elongated, moderately tapering towards apex in dorsal view (Fig. 75 A); gonopods with a small projection towards surstyli in ventral view (Fig. 75 B); phallus with three short ejaculatory ducts, one with some small spines in lateral view (Fig. 75 E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF74ECC6D8D9EFF6FA1E1168.taxon	description	Description: MALE: Body length: 2.5 – 2.6 mm. Head. Flagellum long acuminate; yellow. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, without distinct pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part silver pollinose; slightly grayish also from the side. Scutellum brownish pollinose and with 4 – 6 well developed, short bristles along the distal edge. Dorsocentral bristles well developed, short spine-like, dark, uniform in length throughout, numerous dark bristles around postpronotum. Halter, knob pale, stem brown. Legs. Mid coxa with three short, pale bristles (one longer than the other). Trochanters yellow-brown, femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees, basal 1 / 4 and distal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments yellow, last segment brown dorsally. Ventroapical row of spines on fore femur missing; 5 – 7 short, black spines on mid femur; no spines on hind femur, only 10 – 12 pale bristles equally spaced posteriorly, somewhat longer on distal half (longest up to as long as 0.3 – 0.5 times the width of hind tibia at the distal end). Subapical spines on first four tibiae present, distal and short. Hind trochanter without a keel, but with a distinct, blunt peg ventrally at the distal end, 5 – 7 weak bristles anterior to the peg in a row (Fig. 147 B); ventrobasal spines absent on fore and mid femora. Hind metatarsus not flattened, slightly longer than 2 – 4 combined and with scrub-like bristles ventrally; last tarsomere long almost as long as tarsomeres 3 – 4 combined. Pulvilli as long as the last tarsal segment. Wing. Length: 2.7 mm. Upper side of basal costal cell with one long black bristle. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m distinctly distal to the middle of discal cell. 1 – 3 distinct dark setulae on tegula. Abdomen. Viewed obliquely from front tergites subshining black, tergite 1 silvery grey, sides silvery pollinose on tergites 4 (very small spots) and 5 (4 – 5 times larger spots). Dispersed strong dark bristles on all tergites present, the longest on tergite 5 and up to as long as width of hind tibia at base. Lateral bristles on first tergite present, 3 – 4 short brown bristles in a row up to 0.4 – 0.6 times as long as the width of hind femur at distal tip in lateral view. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 2 – 2.5 times as long as ST 8. Genitalia without dissection: ST 8 small, globular, silvery pollinose, with occasional small bristles; membranous area small, round, directed posteriorly; epandrium and surstyli covered with debris on the one undissected specimen. Genitalia. Genital capsule in dorsal view: epandrium small, wider than long (MLE: MWE = 0.7). surstyli symmetrical, elongated, broadened at base, gradually tapering and pointing towards each other at apex. Left surstylus slightly longer than right one (Fig. 75 A). Genital capsule in ventral view: gonopods equal, elongated with a small projection towards sternite; lobes of hypandrium sclerotized; subepandrial sclerite extended; hypandrium with slender hypandrial apodeme (Fig. 75 B); Genital capsule in lateral view: both surstyli gently curved toward sternite (Fig. 75 D – E); phallic guide straight, apically with pointed apex; phallus with three short ejaculatory ducts, one with 4 – 5 short spines. Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 75 C). FEMALE: Body length: 2.5 mm. As male except for the following characters. Frons, eyes separated, as wide as 1.5 times the width at antennae; completely silver-grey pollinose until up to 1 / 3 of the lower frons, shiny black in front of ocellar triangle, gradually changing from pollinose to shiny black in the transition zone; frons depressed at the point where it is widest. Enlarged ommatidia silvery shining. Pulvilli and claws about 2 – 2.5 times as long as last tarsal segment, 1.5 times on metatarsi. Female abdomen with silver spots on sides of tergites 3 – 6. Ovipositor. Straight in ventral view, straight in lateral view, reaching distal edge of 3 rd segment, somewhat down curved. LP: LB = 2.2. LDP: LPP = 3.0. (Fig. 154 A).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF74ECC6D8D9EFF6FA1E1168.taxon	etymology	Etymology: The name is from the Latin word paxillus for peg or small stake, in reference to the blunt peg-like projection on the hind trochanter. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Mica Creek, Mount Isa, 20 ° 49 ’ S, 139 ° 27 ’ E, 360 m, 7. iii. 2002, C. J. Burwell, JSS 13932 (1 ♂, QM); PARATYPES: AUSTRALIA: Queensland: Isla Gorge National Park, 25 ° 11 ’ S, 149 ° 58 ’ E, 320 m, hilltop near lookout, 13. XI. 1999, J. & A. Skevington, hand collected, JSS 7257 (1 ♂, QM); 2 km along Ridgepole Waterhole Road, 10 km ESE Musselbrook Resource Centre, Lawn Hill National Park, 18 ° 38 ’ S, 138 ° 12 ’ E, 220 m, 7. v. 1995, G. Daniels & M. A. Schneider, JSS 8294 (1 ♂, QM); Western Australia: Karijini National Park, Hamersley-Mount Bruce Road, 22 ° 34 ’ S, 118 ° 18 ’ E, 757 m, dry rocky creek bed, Eucalyptus grassland, 25. iv – 14. v. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16127 (1 ♀, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF74ECC6D8D9EFF6FA1E1168.taxon	distribution	Distribution: Australia (Queensland, Western Australia) (Fig. 132). Notes: This rarely encountered species is found mostly in the northern outback and should be expected in the Northern Territory. One specimen was collected hilltopping. Intraspecific genetic distance ranges from 0.3 % to 1.6 %. The species is genetically closest to T. trispina sp. nov. (12.1 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF7AECC4D8D9EE0FFCEA11FC.taxon	description	Figs 76 A – E, 132	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF7AECC4D8D9EE0FFCEA11FC.taxon	diagnosis	Diagnosis: This species can be recognized by the two stronger dark bristles and some soft short bristles on the hind trochanter; elongated surstyli with a protruding along inner margin in dorsal view (Fig. 76 A); unequal gonopods, left is longer than right one, right with an inward triangle-shaped projection on inner margin (Fig. 76 B); phallic guide with 6 – 7 dorsolateral spines; subepandrial sclerite distinct with dense spine-like bristles in ventral view (Fig. 76 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF7AECC4D8D9EE0FFCEA11FC.taxon	description	Description: MALE: Body length: 2.6 – 2.8 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 – 2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 2 – 3 pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more gray, grayish also from the side. Scutellum silvery brown pollinose and with 3 – 4 bristles along distal edge. Dorsocentral bristles indistinct, dark, more visible in frontal half, a few pale bristles around postpronotum present. Halter, knob pale, stem brown. Legs. Mid coxa with 1 – 2 long dark bristles (1 very long). Trochanters and femora brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 5 of tibiae yellow (tibiae otherwise brown), tarsal segments brown dorsally, yellow ventrally, last segment darker. Hind trochanter with slight keel, 2 stronger dark bristles, sometimes with soft short bristles as well; ventrobasal spines absent on fore and mid femur. Ventroapical row of spines on fore femur missing; 6 – 8 distinct, black spines on mid femur; hind femur without ventroapical spines, with 12 – 14 equally spaced bristles posteroventrally, longer and more frequent on distal half, longest up to 1 / 3 the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae absent. Hind tarsomeres slightly flattened, hind metatarsus as long as 2 – 5 combined. Pulvilli shorter than last tarsal segment. Wing. Length: 2.5 – 2.6 mm. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2 – 3 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites subshiny brown, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (spot on tergite 5 larger). Dispersed strong dark bristles on all tergites present, longest on tergite 5 in length up to 1 / 4 the width of hind tibia at distal end. Lateral bristles on first tergite present, 4 – 5 dark bristles up to as long as ¾ of hind femur’s width at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 2 – 2.3 times as long as ST 8. Genitalia without dissection: ST 8 small, short, rounded in dorsal view, brown and with a few distinct bristles, otherwise with velvet-like coverage; membranous rounded, mostly directed posterodorsally; epandrium brown, surstyli yellow. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.3). Surstyli elongated, symmetrical, left surstylus longer than right one, both surstyli with a narrow edge along inner margin, pointed at apex (Fig. 76 A). Genital capsule in ventral view: both gonopods extended towards surstyli, left is longer than right one, right one with an inward projection on inner margin; phallic guide with 6 – 7 dorso- dorsolateral spines, one spine longer than others; subepandrial sclerite distinct with condensed spine-like hairs (Fig. 76 B); Genital capsule in lateral view: left surstylus straight in basal two thirds, curved towards sternite apically, right surstylus slightly curved towards sternite (Fig. 76 D – E). Ejaculatory apodeme tube-like, bent in the middle (Fig. 76 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF7AECC4D8D9EE0FFCEA11FC.taxon	etymology	Etymology: This species is named after François Péron National Park in the hope that it will increase the profile of the terrestrial biodiversity of this park. The park is famous for Shark Bay and Monkey Mia, where tourists go to feed and interact with dolphins. The terrestrial habitats are largely ignored though as it is a harsh desert with apparently low diversity at first glance. JHS and Angela Skevington walked a short trail that provided an overlook of Shark Bay and the surrounding habitat and were stunned to find an abundance of insects hilltopping around a bench and canopy at the top of the small rise. Ten species of Tomosvaryella were collected in less than two hours at this location, all of them undescribed. Many have been found nowhere else. See more information about the habitat under T. minychoma. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: ~ 3 km N [orth] of Denham, 25 ° 55 ’ S, 113 ° 32 ’ E, hilltop, 12. XII. 1999, J. Skevington, JSS 7369 (1 ♂, WAM); PARATYPES: AUSTRALIA: same data as holotype, JSS 7320, JSS 7324, JSS 7338 – 9, JSS 7376, JSS 7386, JSS 7397 – 8, JSS 7402, JSS 7406 (1 ♂, CNC 9 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF7AECC4D8D9EE0FFCEA11FC.taxon	distribution	Distribution: Australia (Western Australia) (Fig. 132). Note. Intraspecific genetic distance ranges from 0.2 % to 0.5 %. This species is genetically similar to T. patula sp. nov. (7.6 – 8.5 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF78ECC2D8D9EEBBFF5312B8.taxon	description	Figs 77 A – E, 115, 150 B	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF78ECC2D8D9EEBBFF5312B8.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a slight keel covered by some short dark bristles; distal half of hind tibia distinctly flattened (Fig. 150 B); syntergosternite 8 covered by distinct bristles; elongated surstyli in dorsal view (Fig. 77 A); left gonopod longer than right one, both gonopods extended towards surstyli with a distinct projection on inner margin; phallic guide with 5 – 6 dorso- dorsolateral spines, one is downward and longer than others; subepandrial sclerite with a small fine-shaped edge in its center in ventral view (Fig. 77 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF78ECC2D8D9EEBBFF5312B8.taxon	description	Description: MALE: Body length: 2.4 – 2.5 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1 – 1.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 2 – 3 distinct pale bristles. Mesonotum (viewed obliquely from front) silvery pollinose, anterior part more grey, grayish also from the side. Scutellum silvery brown pollinose and with 4 – 6 bristles along distal edge. Dorsocentral bristles distinct, dark, mostly in frontal half, a few pale bristles around postpronotum present. Halter knob pale, stem brown. Legs. Mid coxa with 2 – 3 long dark bristles. Trochanters brown, femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 4 of tibia yellow (tibia otherwise brown), tarsal segments yellow, last segment darker. Hind trochanter with slight keel, and 7 – 10 short dark bristles, equally spaced; ventrobasal spines absent on fore and mid femur. Hind femur without ventroapical spines, except 10 – 12 short equally spaced bristles posteroventrally, longer on distal half, longest up to 1 / 3 the width of hind tibia at distal end. Subapical (distal) spines on fore tibia well developed. Distal half of hind tibia distinctly flattened. Hind tarsomeres distinctly flattened (especially tarsomeres 1 – 3), hind metatarsus as long as 2 – 4 combined; posterior corners with distinct bristles on all tarsomeres. Pulvilli shorter than last tarsal segment. Wing. Length: 2.5 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 1.5 – 2 times as long as third costal section. Cross-vein r-m at middle of discal cell. 4 – 5 short dark setulae on tegula. Abdomen. Viewed obliquely from anterior tergites subshiny brown-black, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (spot on t 5 larger). Dispersed strong dark bristles on all tergites present, longest on ST 8 posterior end, in length up to 1.5 times the width of hind tibia at distal end. Lateral bristles on first tergite present, 4 – 5 dark bristles up to as long as 3 / 4 hind femur’s width at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1.5 – 1.7 times as long as ST 8. Genitalia without dissection: ST 8 medium sized, rounded in dorsal view, brown and with the longest bristles of the abdomen; membranous area is rounded dorsally, narrow ventrally, mostly directed posterodorsally; epandrium brown, surstyli yellow. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.5). Surstyli symmetrical, elongated, left surstylus straight and longer than right one (not curved to right one), right one curved to left one apically (Fig. 77 A). Genital capsule in ventral view: gonopods unequal in height, left is longer than right one, both gonopods extended towards surstyli with a distinct projection on inner margin; phallic guide with 5 – 6 dorso- dorsolateral spines, one is downward and longer than others; subepandrial sclerite with a small fine-shaped edge in its center (Fig. 77 B); Genital capsule in lateral view: both surstyli moderately slender to apex, right surstylus curved to sternite, left surstylus straight in basal two thirds, distinctly curved towards sternite in apical third (Fig. 77 D – E). Ejaculatory apodeme tube-like, bent (Fig. 77 C). FEMALE: unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF78ECC2D8D9EEBBFF5312B8.taxon	etymology	Etymology: From the Greek petalos, broad, flat or outspread, in reference to the flattened distal half of the hind tibia. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: Millstream-Chichester Nat [ional] Park, Yarraloola Road, Fortescue River, 21 ° 38 ’ S, 117 ° 7 ’ E, rocky creek bed, drying pools, 28. IV – 3. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16215 (1 ♂, WAM); PARATYPES: AUSTRALIA: Australian Capital Territory: Black Mountain, 35 ° 16 ’ S, 149 ° 6 ’ E, 15 – 17. II. 1980; 27. II. 1980, D. H. Colless, Malaise trap, JSS 8505, JSS 8692 (1 ♂, ANIC; 1 ♂, CNC); New South Wales: 20 km E of Forbes, 33 ° 23 ’ S, 148 ° 1 ’ E, in Callitris forest, 9. IV. 1977, Z. Liepa, JSS 9178 (1 ♂, ANIC); Queensland: Prince of Wales Island, Torres Strait, 10 ° 41 ’ S, 142 ° 11 ’ E, 27 – 30. V. 1969, Neboiss, JSS 8795 (1 ♂, MVMA); Sunnybank, 27 ° 34 ’ S, 153 ° 3 ’ E, 9. III. 1957, Haseler, JSS 8279 (1 ♂, QM); 30 km north of Wrothan Park, 17 ° 0 ’ S, 144 ° 19 ’ E, 23. IV. 1983, J. F. Donaldson, JSS 8959 (1 ♂, QDPC); Western Australia: Heinsman Rock, Mount Ragged Track, 30 ° 7 ’ S, 123 ° 28 ’ E, 145 m, woodland near drying pools, 30. X – 18. XI. 2003, C. Lambkin & J. Recsei, Malaise trap, ANIC Bulk Sample 2179, JSS 16240 (1 ♂, ANIC); Karijini National Park, Banjima Road, 22 ° 24 ’ S, 118 ° 16 ’ E, 696 m, in muddy creek, burnt Eucalyptus grassland, 15 – 18. V. 2003, C. Lambkin, J. Recsei & N. Starick, Malaise trap, JSS 15970 (1 ♂, ANIC); Millstream-Chichester National Park, Black Hill Pool, 21 ° 20 ’ S, 117 ° 15 ’ E, 176 m, over wide dry rocky creek bed, Eucalyptus, 7 – 12. V. 2003, C. Lambkin & D. Yeates, Malaise trap, JSS 16316 (1 ♂, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF78ECC2D8D9EEBBFF5312B8.taxon	distribution	Distribution: Australia (Australian Capital Territory, New South Wales, Queensland, Western Australia) (Fig. 115).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF7EECC0D8D9EC7FFF641214.taxon	description	Figs 78 A – E, 115, 146 E, 151 E	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF7EECC0D8D9EC7FFF641214.taxon	diagnosis	Diagnosis: This species can be recognized by the two patches of spines on the hind trochanter (Fig. 146 E); long and condensed bristles on membranous area; surstyli elongated in dorsal view (Fig. 78 A); phallic guide with some different-sized spines (Fig. 78 B); one of ejaculatory ducts with some saw-like teeth (Fig. 78 B); both gonopods with a finger-like projection in ventral view (Fig. 78 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF7EECC0D8D9EC7FFF641214.taxon	description	Description: MALE: Body length: 3.0 – 3.1 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 – 1.8 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper half. Thorax. Postpronotum pale, with 4 – 5 distinct pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part silver pollinose, slightly grayish also from the side. Scutellum silver pollinose and with 5 – 7 weak bristles along the distal edge. Dorsocentral bristles well developed, dark, longer on frontal part and numerous dark bristles around postpronotum. Halter, knob pale, stem black. Legs. Mid coxa with 2 – 3 spines. Trochanters and femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees yellow, tibiae black, tarsal segments dark brown, only yellowish ventrally; fore metatarsus is transparent in ventral view. Hind trochanter with a few short dark spines (5 – 8) ventrobasally and (2 – 3) shorter ones ventroapically (Fig. 146 E). Ventroapical row of spines on fore femur missing, 4 – 6 very short, black spines on mid femur; no spines on hind femur, only 6 – 8 pale bristles posteriorly on distal half (as long as the width of hind tibia at the distal end); ventrobasal spines (2) present on fore and weaker on mid femur; hind tibia slightly bent at middle in posterior view. Subapical (distal) spines on first four tibiae present, very short. Hind metatarsus extremely flattened, spoon-like (extending posterior edge, widest at basal 1 / 3) (Fig. 151 E), second tarsomere slightly longer than 3 – 5. Pulvilli as long as last tarsal segment. Wing. Length: 3.0 mm. Upper side of basal costal cell with one dark and long bristles. Fourth costal section 1.5 – 2 times as long as third costal section, fourth section appears to be unusually short. Cross-vein r-m slightly distal to middle of discal cell. 4 – 5 dark setulae on tegula. Abdomen. Viewed obliquely from front tergites subshining black, tergite 1 silvery grey, sides silvery pollinose on tergites 4 and 5 (the latter with larger spots). Dispersed long dark bristles on tergites present, the longest on ST 8 and up to 3.5 – 4 times as long as width of hind tibia at base. Lateral bristles on first tergite present, 6 – 8 brown bristles in a patch up to as long as 1 – 1.2 times the width of hind femur at distal tip in lateral view. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1 – 1.5 times as long as syntergosternite 8 (ST 8). Genitalia without dissection: ST 8 large, black, hairy with longest bristles on posterior part; membranous area round, posteriorly directed, the longest bristles originate from membranous area; tip of epandrium brown; surstyli pale, appear elongated and uniformly broad. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.8). Surstyli asymmetrical, elongated, left longer than right one (Fig. 78 A). Genital capsule in ventral view: subepandrial sclerite covered with some transverse streaks in middle; both gonopods with a finger-like projection extended towards surstyli, left rather longer than right one (Fig. 78 B), phallic guide with 5 – 6 different-sized spines dorsolaterally (longer ones extended towards left surstylus); phallus with three ejaculatory ducts, one bears 10 – 11 saw-like teeth along its side. Genital capsule in lateral view: left surstylus straight in basal two thirds, bent towards sternite in apical one third; right surstylus curved towards sternite (Fig. 78 D – E); ejaculatory apodeme flat, sperm pump elongated, linear (Fig. 78 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF7EECC0D8D9EC7FFF641214.taxon	etymology	Etymology: The name planca is derived from the Latin plancus, flat-footed, in reference to the flattened hind metatarsus. Examined material: HOLOTYPE: AUSTRALIA: Queensland: 3.5 km SW by S Mount Baird, 15 ° 10 ’ S, 145 ° 7 ’ E, 3. v. 1981, D. H. Colless, Malaise trap, JSS 8576 (1 ♂, ANIC); PARATYPES: AUSTRALIA: Queensland: 20 km E Mareeba, 17 ° 0 ’ S, 145 ° 26 ’ E, 16. iv. 1980, G. F. Hevel & J. A. Fortin, JSS 10535 (1 ♂, USNM); 3.5 km SW by S Mount Baird, 15 ° 10 ’ S, 145 ° 7 ’ E, 3. v. 1981, D. H. Colless, Malaise trap, JSS 8577, JSS 8585 (1 ♂, ANIC; 1 ♂, QM); Edgbaston National Reserve (EBM 1), 22 ° 44 ’ S, 145 ° 26 ’ E, Melaleuca floodplain, 29. X – 15. XII. 2010, C. Lambkin et al., Malaise trap, 18528, CNC 596962 (1 ♂, CNC); Western Australia: Martin’s Well, West Kimberley, - 16.56604, 122.85010, 28. IV. 1977, D. H. Colless, Malaise trap, JSS 9223 (1 ♂, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF7EECC0D8D9EC7FFF641214.taxon	distribution	Distribution: Australia (Queensland, Western Australia) (Fig. 115). Notes: This species is mostly restricted to NE Queensland, but a disjunct record is from the West Kimberley region.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF7CECDED8D9EDD3FD0E1328.taxon	description	Figs 79 A – E, 134, 148 E, 154 F	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF7CECDED8D9EDD3FD0E1328.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a square keel and being covered with short bristles along the edge (Fig. 148 E); elongated surstyli in dorsal view (Fig. 79 A); gonopods elongated towards surstyli in ventral view (Fig. 79 B); long subepandrial sclerite with a long protrusion (Fig. 79 B); phallic guide with 6 – 7 dorsolateral spines (Fig. 79 E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF7CECDED8D9EDD3FD0E1328.taxon	description	Description: MALE: Body length: 2.5 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance 2 x length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 1 – 2 pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum brownish pollinose and with 5 – 7 bristles along the distal edge. Dorsocentral bristles weakly developed, dark, somewhat longer along frontal edge, numerous dark bristles around postpronotum. Halter, knob pale with brownish tinge, stem brown. Legs. Mid coxa with two long, dark bristles. Trochanters pale brown and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments yellow, last segment darker. Hind trochanter with a distinct, square keel (like a “ top hat ”) and with short bristles along the edge, longest basally (Fig. 148 E); one ventrobasal spine present on fore and absent on mid femur. Ventroapical row of spines on fore femur absent; 6 – 8 small, black spines on mid femur; hind femur without ventroapical spines, except 8 – 10 equally spaced bristles posteroventrally on distal half, longest up to ½ the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present. Hind tibia bent in posterior view. Hind tarsomeres flattened (especially metatarsus), metatarsus almost as long as 2 – 3 combined. Pulvilli shorter than last tarsal segment. Wing. Length: 3.0 mm. Upper side of basal costal cell with one long dark bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m slightly at middle of discal cell. 2 – 3 distinct brown setulae on tegula. Abdomen. Dissected. Tergite 1 silvery grey, Lateral bristles on first tergite present, 4 – 5 dark bristles up to as long as hind femur’s width at base. Postabdomen in dorsal view: Dissected. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.5). Surstyli elongated, right surstylus smaller than left one, right one with a small protrusion at inner margin before apex (Fig. 79 A). Genital capsule in ventral view: gonopods unequal in height, left slightly longer than right one, elongated towards surstyli, subepandrial sclerite long, rectangular-shaped, with a long protrusion, extended from its base to middle (Fig. 79 B). Genital capsule in lateral view: right surstylus rather straight, left surstylus curved towards sternite, phallic guide with 6 – 7 dorso-dorsolateral spines (one is longer) (Fig. 79 C – E). FEMALE: Body length: 2.5 mm. As male except for the following characters. Frons, eyes separated, as wide as 1.8 – 2 times the width at antennae; completely silver-grey pollinose until up to 1 / 3 of the lower frons, shiny black in front of ocellar triangle, gradual change in transition zone; frons depressed along upper 2 / 3 with a groove in the median line. Enlarged ommatidia silvery shining. Pulvilli and claws about 1 – 1.5 times as long as last tarsal segment, smaller on metatarsi. Female abdomen with silver spots on sides of tergites 2 – 6. Ovipositor. Straight in ventral view, straight and extra-long in lateral view, reaching distal base of 1 st segment; base brownish pollinose with occasional bristles. LP: LB = 3.3. LDP: LPP = 2.5. (Fig. 154 F).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF7CECDED8D9EDD3FD0E1328.taxon	etymology	Etymology: This species in named after Narelle Power. Narelle and her team collected the holotype and in her survey work on Bribie Island and Mount Glorious she collected a significant number of pipunculids that we have used in numerous revisions. She has collected 10 of the Tomosvaryella species used in this project (Supplementary file 2). Examined material: HOLOTYPE: AUSTRALIA: Queensland: Bribie Island, QDPI Fisheries Site, 27 ° 3 ’ S, 153 ° 11 ’ E, 5 – 12. IX. 1997, Heathland- Acacia regrowth, S. Winterton, N. Power & D. White, Malaise trap, JSS 8428 (1 ♂, QM); PARATYPES: AUSTRALIA: Northern Territory: Namatijira Drive, 20 km W Hugh River Crossing, 23 ° 49 ’ S, 133 ° 11 ’ E, 14 – 15. III. 2002, 760 m, C. J. Burwell, JSS 13934 (1 ♀, QM); 30 km NW by W of Alice Springs, 23 ° 42 ’ S, 133 ° 53 ’ E, 7. X. 1978, D. H. Colless, JSS 9240, JSS 9246 (2 ♂, ANIC); 32 km WNW of Alice Springs, 23 ° 42 ’ S, 133 ° 53 ’ E, 8. X. 1978, D. H. Colless, Malaise trap, JSS 9243 – 5 (3 ♂, ANIC); 53 km E by N of Alice Springs, 23 ° 42 ’ S, 133 ° 53 ’ E, 6. X. 1978, D. H. Colless, Malaise trap, JSS 9252 (1 ♂, ANIC); Roe Creek, 12 km SW by W Alice Springs, 23 ° 42 ’ S, 133 ° 53 ’ E, 10. X. 1978, D. H. Colless, Malaise trap, JSS 9229 (1 ♂, CNC); Queensland: Charleville, 26 ° 24 ’ S, 146 ° 14 ’ E, 15. II. 1970, A. L. Dyce, car net, ex. spirit via ethyl acetate, JSS 8620 (1 ♂, ANIC); Australian Capital Territory: Paddy’s River, 35 ° 27 ’ S, 148 ° 59 ’ E, 18. III. 1966, R. Pilfrey, JSS 9707 (1 ♂, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF7CECDED8D9EDD3FD0E1328.taxon	distribution	Distribution: Australia (Australian Capital Territory, Northern Territory, Queensland) (Fig. 134). Notes: The Australian Capital Territory specimen is disjunct, so it was checked again. It has the same morphological and terminalia characters as the holotype other than right surstylus in lateral view, which is narrower in the middle. This is typical variation found in other Tomosvaryella species, so it has been included in the type series. Intraspecific genetic distance is 0.3 %. This species is genetically similar to T. mathiesoni sp. nov. (2.3 – 2.9 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF62ECDFD8D9ECCFFCFD1110.taxon	description	Figs 80 A – E, 135	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF62ECDFD8D9ECCFFCFD1110.taxon	diagnosis	Diagnosis: This species can be recognized by the elongated surstyli, moderately narrowed towards the apex in dorsal view (Fig. 80 A); equal gonopods in height, extended towards surstyli; phallic guide with 2 – 4 different-sized spines dorsolaterally, subepandrial sclerite long, rounded in middle in ventral view (Fig. 80 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF62ECDFD8D9ECCFFCFD1110.taxon	description	Description: MALE: Body length: 2.0 mm (without abdomen). Head. Scape and pedicel dark brown. Pedicel with 2 – 4 upper bristles; flagellum brown and tapering. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput grey pollinose, covered by short brown bristles. Thorax. Mesonotum (viewed obliquely from front) silvery-brown pollinose; scutum with scattered bristles on anterior supra-alar area. Postalar callus dark brown with 2 – 3 short bristles in the middle. Scutellum silvery pollinose and with a few indistinct bristles along the distal edge. Halter, knob pale, stem light brown, base dark. Legs. Coxae, trochanters, femora dark, hind femur shining ventrally, knees and basal ¼ of tibiae yellow (tibiae otherwise dark brown), tarsomeres light brown. Hind trochanter silvery pollinose, with some short bristles in a row; ventrobasal spines absent on fore and mid femur. Hind femur with two rows of ventroapical spines. Hind tibia with two rows of short black bristles on anterior and posterior side. Hind tarsomeres moderately flattened, 5 th hind tarsomere as long as 3 – 4 combined; pulvilli shorter than 5 th tarsal segment. Wing. Length: 2.5 mm. Fourth costal section 1.5 – 2 times as long as third costal section. Cross-vein r-m at middle of discal cell. Abdomen. Tergites 1 silvery pollinose with 10 – 12 dark bristles. Abdomen dissected. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.4). Surstyli widened at the base, elongated, moderately narrowed before apex, pointed and bent to each other at the apex (Fig. 80 A). Genital capsule in ventral view: both gonopods equal in height, extended towards surstyli; phallic guide with 2 – 4 differently sized spines dorsolaterally; subepandrial sclerite long (Fig. 80 B); Genital capsule in lateral view: both surstyli widened at base, moderately curved towards sternite (Fig. 80 D – E); ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 80 C). FEMALE: unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF62ECDFD8D9ECCFFCFD1110.taxon	etymology	Etymology: The specific name is derived from the Latin word ‘ prolate’, meaning elongated, referring to the long surstyli. Examined material: HOLOTYPE: AUSTRALIA: South Australia: Flinder’s Ranges National Park, Brachina Gorge, Heysen Hilltop, 31 ° 20 ’ S, 138 ° 33 ’ E, 9. X. 1997, J. & A. Skevington & C. Lambkin, JSS 8392 (1 ♂, QM); PARATYPES: AUSTRALIA: Queensland: Culgoa Floodplain National Park, 7 km WNW Toulby Gate (TOLB 6 M), 28 ° 57 ’ S, 146 ° 52 ’ E, Callitris, 19. III – 20. V. 2010, C. Kelly & A. Coward, Malaise trap, 19280, CNC 575034 (1 ♂, QM); Mount Moffatt summit, 25 ° 4 ’ S, 148 ° 3 ’ E, 18. XI. 2014, J. H., A. M. & A. W. Skevington, CNC 374933 (1 ♂, CNC); Brisbane, Mount Coot-tha, 27 ° 29 ’ S, 152 ° 57 ’ E ,, 250 m, hilltop, 21. VIII. 2004, J. Skevington, hand collected, JSM 811 – 3; JSS 15388 – 90 (3 ♂, CNC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF62ECDFD8D9ECCFFCFD1110.taxon	distribution	Distribution: Australia (Queensland, South Australia) (Fig. 135). Notes: All but one of the specimens was captured hilltopping.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF60ECDDD8D9EFF6FCB2150C.taxon	description	Figs 81 A – E, 134	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF60ECDDD8D9EFF6FCB2150C.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of the surstyli, widened in the basal half and narrowed and bent towards the sternites in the middle in dorsal view (Fig. 81 A); extremely elongated hypandrium; gonopods extended towards surstyli, rounded at apex in ventral view (Fig. 81 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF60ECDDD8D9EFF6FCB2150C.taxon	description	Description: MALE: Body length: 2.5 mm. Head. Flagellum tapering; yellow-brown pollinose, scape and pedicel brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.1 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3, covered by scattered short dark bristles. Thorax. Postpronotum pale, with 2 long dark bristles. Mesonotum (viewed obliquely from front) brown, silvery pollinose with some long supra-alar bristles behind postpronotum lobes and two rows dorsocentral bristles. Scutellum brown, silvery pollinose with 4 – 6 long erect bristles (longer than dorsocentral bristles) along distal edge. Halter, knob, stem pale, base brown. Legs. Mid coxa with 3 – 4 long dark bristles. Trochanters brown, femora black, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments bright yellow. Fore and mid trochanters with three and two erect bristles, respectively. Hind trochanter gray pollinose without distinct projection; two distinct ventrobasal bristles present on fore and mid femora. Hind femur without ventroapical spines, except 5 – 8 equally spaced brown bristles posteroventrally, longest up to 2 / 3 the width of hind tibia at distal end. Short distal spines present on first four tibiae. Hind tarsomeres not flattened, hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 2.5 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 1 – 2 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brown-black, tergite 1 grey-brown pollinose. Dispersed short bristles on all tergites present. Lateral bristles on first tergite present, 4 – 5 dark bristles up to as long as ¾ of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: ST 8 longer than wide, epandrium longer than wide (MLE: MWE = 1.2). Surstyli widened in basal half, strongly narrowed in apical half, right one narrower and longer in apical half, both surstyli bent towards sternite by 90 ° in apical half (Fig. 81 A). Genital capsule in ventral view: both gonopods extended towards surstyli, rounded at apex, hypandrium extremely elongated; phallic guide strong and rather wide, with one small dorsal spine before apex (Fig. 81 B); Genital capsule in lateral view: both surstyli straight and wide in basal half, then narrowed and bent towards sternite in the middle (Fig. 81 D – E); ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 81 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF60ECDDD8D9EFF6FCB2150C.taxon	etymology	Etymology: From the Latin prolixus, stretched out long, in reference to the elongate hypandrium of this species. Examined material: Holotype: AUSTRALIA: New South Wales: Ledknapper National Reserve, 7.3 km ENE Beulah Headquarters (LDN 2 M), 29 ° 21 ’ S, 146 ° 13 ’ E, E. melanopholia, Spinifex, 10. XII. 2009 – 18. III. 2010, S. O’Sullivan & K. Taylor, Malaise trap, CNC 576520 (1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF60ECDDD8D9EFF6FCB2150C.taxon	distribution	Distribution: Australia (New South Wales) (Fig. 134).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF61ECDBD8D9EAABFB831590.taxon	description	Figs 82 A – E, 135, 156 D	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF61ECDBD8D9EAABFB831590.taxon	diagnosis	Diagnosis: This species can be recognized by the mid and hind femora bearing 8 – 15 short but distinct bristles posteriorly; surstyli triangle-shaped; narrowed in apical one third (Fig. 82 A); subepandrial sclerite short; gonopods wide, rounded at apex; ejaculatory ducts sinuous without any teeth in ventral view (Fig. 82 B); surstyli curved towards sternite in lateral view with same width from base to apex; phallic guide short, broadened at base with a hook-like projection in lateral view (Fig. 82 D – E); phallus with three ejaculatory ducts.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF61ECDBD8D9EAABFB831590.taxon	description	Redescription: MALE. Body length: 3.0 – 3.5 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black, lower part distinctly golden pollinose; eyes touching for distance equal to 2 – 2.5 x length of ocellar triangle. Occiput silvery pollinose, slightly less so on upper half. Thorax. Postpronotum pale brown, with 2 – 3 whitish bristles. Mesonotum (viewed obliquely from front) subshining black, anterior part indistinctly brown pollinose; slightly brownish also from the side. Scutellum with dorsocentral bristles weakly developed, dark, somewhat longer on frontal part. Halter, knob pale, stem dark brown. Legs. Mid coxa with 2 – 3 dark bristles. Trochanters and base of femora brown, femora dark brown, shining ventrally, femora silvery pollinose posteriorly except shiny hind femur; knees, basal 1 / 3 of tibiae yellow, tarsal segments pale brown, last segment dorsally darker. Ventroapical row of spines on fore femur tiny, 4 – 6, 8 – 10 short, black spines on mid femur; 6 – 8 spines on hind femur, mid and hind femora with 8 – 15 short but distinct bristles posteriorly. Subapical (distal) spines on first four tibiae absent. Hind trochanter without bristles on smoothly curved ventral side, ventrobasal spines absent on fore and mid femora. Hind tarsi slightly flattened, first tarsomere slightly shorter than 2 – 5 combined; ventral bristles scrub-like. Pulvilli as long as last tarsal segment. Wing. Length: 2.8 – 3.1 mm. Upper margin of basal costal cell with one short bristle. Fourth costal section 2 – 3 times as long as third costal section. Cross-vein r-m distinctly distal to middle of discal cell. Setulae on tegula absent. Abdomen. Viewed obliquely from front tergites golden-brownish pollinose, tergite 1 silvery grey pollinose with 6 – 8 short lateral bristles (up to 0.15 mm), sides faintly silvery on tergites 4 and 5 (the latter with larger spot, but weakly colored). Dispersed short dark bristles on tergites absent. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 twice as long as ST 8. Genitalia without dissection: ST 8 small, mainly shiny brown; membranous area lanceolate, shaped like a sunflower seed, vertical; surstyli relatively small, brown; hairy cerci visible without dissection in lateral view. Genitalia. Surstyli hairy, wide at base in ventral view, narrowing at 2 / 3 of the length and broadening at tip, both uniformly wide in lateral view and pointed at tip; epandrium short and wide; cerci symmetrically placed, small; ST 8 with a narrowing membranous area towards epandrium. Genital capsule in dorsal view: epandrium and surstyli dark brown. Epandrium wider than long (MLE: MWE = 0.6). Both surstyli broadened at base, triangle-shaped, narrowed in apical one third (Fig. 82 A). Genital capsule in ventral view: subepandrial sclerite short and narrow; gonopods symmetrical broad and rounded at apex; phallus trifid, ejaculatory ducts sinuous, without any teeth (Fig. 82 B). Genital capsule in lateral view: both surstyli gently curved to sternite, not narrowed at apex (Figs 82 D – E); phallic guide broadened with final thin hook-like projection, visible; ejaculatory apodeme flat; sperm pump elongated, linear (Fig. 82 C). FEMALE. Body length: 3.0 – 3.3 mm. As male except for the following characters. Frons, eyes separated, as wide as 1.2 – 1.5 times the width at antennae; completely silver-grey pollinose until up to ¾ of the frons, shiny black in front of ocellar triangle. Enlarged ommatidia silvery shining. Pulvilli and claws about 1 – 1.2 times as long as last tarsal segment. Female abdomen with silver spots on sides of tergites 3 – 6. Ovipositor (Fig. 156 D). Base dark brown with pale apical margin, piercer straight in ventral view, slightly up curved in lateral view. Examined material: LECTOTYPE [HERE DESIGNATED]: AUSTRALIA: Queensland: Cairns, [16 ° 55 ’ S, 145 ° 46 ’ E], 21. VIII. 1904, JSS 8479 (1 ♂, BPBM); PARALECTOTYPE: AUSTRALIA: same location as lectotype, VII. 1904, BMNH (E) 964035, JSS 8480 (1 ♀, BMNH; 1 ♀, BPBM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF61ECDBD8D9EAABFB831590.taxon	distribution	Distribution: Australia (Queensland) (Fig. 135). Notes: Based on the shape of the surstyli, this species is related to Tomosvaryella infundibula sp. nov. It differs by the shape of the ejaculatory ducts and surstyli in lateral view, with three long and sinuous ejaculatory ducts, without any teeth (Fig. 48 D – E); dorsomedial margin of surstyli bent towards sternites (Fig. 48 D – E); hind femur with bristles posteriorly. Perkins reared this species from a deltocephaline leafhopper, Hecalus sp. (Perkins 1905). Specimen JSS 8479 is hereby nominated as the lectotype to fix and ensure consistent interpretation of the name. Intraspecific genetic distance ranges from 0.0 % to 0.3 %. Tomosvaryella pseudophanes is genetically most similar to T. latistyla sp. nov. (5.6 – 8.8 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF67ECD4D8D9E957FD5F126C.taxon	description	Figs 83 A – E, 136, 152 F, 156 E	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF67ECD4D8D9E957FD5F126C.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter bearing 6 – 7 bristles ventrally; surstyli rectangular-shaped, pointing towards each other with distinct bristles on lateral distal corners directed laterally in dorsal view (Fig. 83 A), phallic guide with 3 – 4 dorsolateral spines, subepandrial sclerite long with fine transverse streaks (Fig. 83 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF67ECD4D8D9E957FD5F126C.taxon	description	Description: MALE: Body length: 2.2 – 2.4 mm. Head. Flagellum acuminate, yellow-brown. Face silvery pollinose. Frons, upper part shining black, lower part silvery pollinose; eyes touching for distance equal to 1.5 – 2 times ocellar triangle. Occiput silvery pollinose, slightly less so on upper half. Thorax. Postpronotum pale brown, with 2 – 3 whitish bristles. Mesonotum (viewed obliquely from front) subshining black, anterior part silvery pollinose, also silvery from the side. Scutellum grey pollinose, without bristles. Dorsocentral bristles weakly developed, pale, somewhat longer on frontal part. Halter, knob pale, stem dark black. Legs. Trochanters and base of femora brown, femora dark brown, shining ventrally (hind femur also posteriorly), knees, basal 1 / 6 of tibiae, tarsal segments yellow, last segment dorsally brown. Mid coxa with three strong bristles at inner apical corner. Ventroapical row of spines on fore femur missing, 4 – 6 very short, black spines on mid femur; no spines on hind femur, only 3 – 4 flattened thick bristles on keel. Subapical (distal) spines on first four tibiae absent. Hind trochanter with 6 – 7 short dark bristles on ventral side, basal spines absent (short pale basal bristles on fore and mid femur may be present). Hind tarsomeres slightly flattened, hind metatarsus slightly longer than 2 – 4 combined. Pulvilli slightly shorter than last tarsal segment. Wing. Length: 2.4 – 2.5 mm. Upper side of basal costal cell with one long distinct bristle (Fig. 152 F). Fourth costal section 2.5 – 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. Setulae on tegula absent. Abdomen. Viewed obliquely from front tergites brownish pollinose, hind margin silvery grey on tergite 1, sides completely silvery on tergites 4 and 5. Dispersed short dark bristles absent. Lateral bristles on first tergite: 2 – 3, pale brown, as long as width of hind femur at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 as long as ST 8. Genitalia without dissection: ST 8 appears large, mainly dark brown and forming a right-angle corner (90 degrees) in lateral view at the lower end of membranous area; membranous area slit-like, vertical, distal part of epandrium paler and surstylus tips whitish, surstyli relatively small. Genitalia. Genital capsule in dorsal view: surstyli rectangular-shaped with acute tips on apical inner corners, epandrium wider than long (Fig. 83 A), 0.5 times as long as ST 8; cerci symmetrically placed, small; ST 8 with slit-like membranous area. Genital capsule in ventral view: Subepandrial sclerite covered with horizontal transverse streaks; gonopods symmetrical; phallic guide with a membranous projection to the side; phallus trifid, branches short; phallic guide with 3 – 4 dorsolateral spines, ejaculatory apodeme flat; sperm pump elongated, linear (Fig. 83 B). Genital capsule in lateral view: both surstyli straight, bent before apex (Fig. 83 D – E). FEMALE: Body length: 2.2 – 2.6 mm. As male except for the following characters. Frons, eyes separated, as wide as 2 times the width at antennae; completely silver-grey pollinose until 1 / 6 – 1 / 4 from the antennae and along median line up to 2 / 3 of the frons, less so along the edges, smooth transition from grey pollinose to shiny black. Enlarged ommatidia silvery shining. Pulvilli and claws about 1 – 1.5 times as long as last tarsal segment on metatarsi. Ovipositor. Base of ovipositor dark brown with some scattered short dark bristles, piercer slightly down curved. LP: LB = 1.7. LDP: LPP = 3.0. (Fig. 156 E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF67ECD4D8D9E957FD5F126C.taxon	etymology	Etymology: Named after the Greek word pterygion for fin, in reference to the fin-shaped keel on the hind femur of males. Examined material: HOLOTYPE: AUSTRALIA: New South Wales: Yathong Nature Reserve near 8 mile tank, 32 ° 31 ’ S, 145 ° 37 ’ E, 218 m, across wide wash: field of flowers: open woodland, 4 – 25. x. 2003, C. Lambkin & N. Starick, Malaise trap, CNCD 3560 (1 ♂, QM); PARATYPES: AUSTRALIA: New South Wales: Culgoa National Park, 13.5 km SW Cawwell Homestead (CGN 1 M), 29 ° 7 ’ S, 146 ° 56 ’ E, Callitris, 30. I – 18. V. 2010, QM Team, B. Shiebaan & R. Ohlsen, Malaise trap, 19325, CNC 576619, CNC 576624, CNC 576626, CNC 576629 (4 ♂, QM); Culgoa National Park, 4.5 km WNW Cawwell Homestead, Claypan (CGN 3 M), 29 ° 3 ’ S, 147 ° 1 ’ E, 192 m, Gidgee, 30. I – 18. V. 2010, B. Schiebaan, Malaise trap, 17220, CNC 576533 – 35 (3 ♂, QM); Culgoa National Park, 8 km WNW Cawwell Homestead, Diemunga Lagoon (CGN 2 M), 29 ° 3 ’ S, 146 ° 60 ’ E, Coolibah, 22. XI – 21. XII. 2009, C. Lambkin, B. Shiebaan & N. Starick, Malaise trap, DB 9 (2 nd bottle), 19302, CNC 593683 – 84 (2 ♂, QM); Culgoa National Park, 8 km West – Northwest Cawwell Homestead, Diemunga Lagoon (CGN 2 M), 29 ° 3 ’ S, 146 ° 60 ’ E, Coolibah, 30. I – 18. V. 2010, C. Lambkin, R. Olsen & B. Shieban, Malaise trap, 19322, CNC 575125 (1 ♂, QM); Ledknapper National Reserve, 5 km N Beulah Homestead (LDN 1 M), 28 ° 19 ’ S, 147 ° 10 ’ E, Allocasuarina, 18. III – 15. V. 2010, S. O’Sullivan & C. Lambkin, Malaise trap, 12 DB, 19361, CNC 576716 (1 ♂, QM); Warrumbungles National Park, Woolshed at Wombelong Creek, 31 ° 16 ’ S, 148 ° 57 ’ E, 420 m, rocky valley, cedar forest, 12 – 16. XII. 1995, M. E. Irwin, Malaise trap, JSS 14028 (1 ♂, INHS); Culgoa National Park, 8 km WNW Cawwell Homestead, Diemunga Lagoon (CGN 2 M), 29 ° 3 ’ S, 146 ° 60 ’ E, Coolibah, 22. XI – 21. XII. 2009, C. Lambkin, B. Shiebaan & N. Starick, Malaise trap, CNC 593677 (1 ♂, CNC); Culgoa National Park, 13.5 km SW Cawwell Homestead (CGN 1 M), 29 ° 7 ’ S, 146 ° 56 ’ E, Callitrus, 21. XII. 2009 – 30. I. 2010, B. Schiebaan, Malaise trap, CNC 576570, CNC 576578, CNC 576580, CNC 576588 (4 ♂, QM); Blackbox, 22. XI – 21. XII. 2009, C. Lambkin, B. Shiebaan, N. Starick, Malaise trap, 11 DB, 19300, CNC 576655, CNC 576658, CNC 576664 (3 ♂, QM); Callitris, 30. I – 18. V. 2010, QM Team, B. Shiebaan & R. Ohlsen, Malaise trap, 19325, CNC 576606, CNC 576610, CNC 576639, CNC 576641 (4 ♂, QM); Culgoa National Park, 8 km West-Northwest Cawwell Homestead, Diemunga Lagoon (CGN 2 M), 29 ° 3 ’ S, 146 ° 60 ’ E, Coolibah, 30. I – 18. V. 2010, C. Lambkin, R. Olsen, B. Shieban, Malaise trap, 19322, CNC 575128, CNC 575132, CNC 575136 (3 ♂, QM); Billabong Creek, Conargo, 35 ° 19 ’ S, 145 ° 9 ’ E, 23 – 30. iv. 1978, Z. Liepa, JSS 9170 (1 ♂, ANIC); Culgoa National Park, 1 km North-Northwest Cawwell Homestead, Claypan (CGN 4 M), 29 ° 4 ’ S, 147 ° 3 ’ E, 136 m, Gidgee, 30. i. 2010, B. Schiebaan, Malaise trap, CNC 575350 (1 ♂, QM); Culgoa National Park, 8 km West-Northwest Cawwell Homestead, Diemunga Lagoon (CGN 2 M), 29 ° 3 ’ S, 146 ° 60 ’ E, Coolibah, 30. i – 18. v. 2010, C. Lambkin, R. Olsen & B. Shieban, Malaise trap, CNC 575142, CNC 575144, CNC 575147, CNC 575151 – 4 (7 ♂, QM); Culgoa National Park, 8 km West – Northwest of Cawwell Homestead, Diemunga Lagoon (CGN 2 M), 29 ° 3 ’ S, 146 ° 1 ’ E, 136 m, Coolibah, 21. xii. 2009 – 30. i. 2010, B. Schiebaan, Malaise trap, CNC 574755 – 6 (2 ♂, QM); Fowlers Gap Research Station, 31 ° 5 ’ S, 141 ° 42 ’ E, 29. xi – 2. xii. 1981, I. D. Naumann & J. C. Cardale, ex. ethanol via cellosolve-xylene, JSS 9234 – 6 (3 ♂, ANIC); Jerilderie, 35 ° 22 ’ S, 145 ° 43 ’ E, 28. xi. 1978, J. McGechan, JSS 8739 (1 ♂, ASCU); Mulwaree Ponds, Inveralochy Bridge, near Yass, 34 ° 44 ’ S 149 ° 45 ’ E, 27. ii. 1966, Z. Liepa, JSS 9238 (1 ♂, ANIC); Warrumbungle National Park, 31 ° 14 ’ S, 149 ° 1 ’ E, 19. x – 3. xi. 1997, J. Skevington & S. Winterton, Malaise trap, JSS 8436 (1 ♀, QM); Yathong Nature Reserve near Triangle Tank, 32 ° 38 ’ S, 145 ° 33 ’ E, 200 m, in sandy wash in low closed mallee, 3 – 25. x. 2003, C. Lambkin & N. Starick, Malaise trap, JSS 16038 (1 ♂, ANIC); Yathong Nature Reserve near 8 Mile Tank, 32 ° 38 ’ S, 145 ° 41 ’ E, ~ 205 m, field of flowers: open woodland, 3 – 25. x. 2003, C. Lambkin & N. Starick, Malaise trap; CNCD 3618, CNCD 3634 (2 ♂, QM); Yathong Nature Reserve near 8 mile tank, 32 ° 31 ’ S, 145 ° 37 ’ E, ~ 218 m, across wide wash: field of flowers: open woodland, 4 – 25. x. 2003, C. Lambkin & N. Starick, Malaise trap, CNCD 3559, CNCD 3563 (2 ♂, QM); Queensland: 2.5 km West – Northwest of Plevna Downs Homestead (PD 3 P 2), 26 ° 40 ’ S, 142 ° 35 ’ E, 133 m, Gidgee, 25. XI – 17. XII. 2008, R. Mackenzie, pitfall trap, 17309, CNC 575273 (1 ♂, QM); Culgoa Floodplain National Park, 11.8 km N Headquarters (CG 2 AM), 28 ° 54 ’ S, 147 ° 8 ’ E, 151 m, Brigalow, 20. I – 18. III. 2010, C. Kelly & A. Coward, Malaise trap, 19271, CNC 576676, CNC 576678 (2 ♂, QM); Elizabeth Creek, Boodjamulla NP, 18 ° 14 ’ S, 138 ° 5 ’ E, 170 m, Bloodwood open, 18 – 22. IV. 2005, M. Mathieson & G. Smith, Malaise trap, 12398, JSS 50564 (1 ♂, CNC); Kurumba, 17 ° 29 ’ S, 140 ° 53 ’ E, 18. IV. 1983, J. F. Donaldson, JSS 8957 (1 ♂, QDPC); Plevna Downs, 2.3 km SSW Arima (PD 2 M), 26 ° 32 ’ S, 142 ° 31 ’ E, 134 m, Gidgee, 25. XI – 17. XII. 2008, R. Mackenzie, Malaise trap, 17292, CNC 595264 (1 ♂, QM); Plevna Downs, 2.5 km WNW Homestead (PD 3), 26 ° 40 ’ S, 142 ° 35 ’ E, 133 m, Gidgee, 16. IX – 2. X. 2008, Starick, Lambkin & Mackenzie, Malaise trap, 16272, CNC 591861 (1 ♂, QM); 12.7 km South-Southeast of Plevna Downs Homestead (PD 8 M), 26 ° 47 ’ S, 142 ° 39 ’ E, 145 m, sandy creek bank, 25. XI – 18. XII. 2008, R. Mackenzie, Malaise trap, 17341, CNC 593802 (1 ♂, QM); Culgoa Floodplain National Park Headquarters, 11.8 km N, 28 ° 55 ’ S, 147 ° 8 ’ E, 150 m, Brigalow, 9. XII. 2007 – 16. I. 2008, N. Starick, C. Lambkin & R. Raven, Malaise trap, 15683, CNC 591849 (1 ♂, QM); Culgoa Floodplain National Park, 11.8 km N Headquarters (CG 2 AM), 28 ° 54 ’ S, 147 ° 8 ’ E, 151 m, Brigalow, 20. I – 18. III. 2010, C. Kelly, A. Coward, Malaise trap, 19271, CNC 576687 – 90, CNC 576700 (5 ♂, QM); 18. XI – 17. XII. 2009, C. Lambkin, A. Coward, C. Kelly & N. Starick, Malaise trap, CNC 576510, CNC 576512 (2 ♂, QM); Culgoa Floodplain National Park, 11.8 km North of Headquarters (CG 2 AM), 28 ° 54 ’ S, 147 ° 8 ’ E, 151 m, Brigalow Black box, 17. XII. 2009 – 20. I. 2010, C. Kelly & A. Coward, Malaise trap, CNC 574522 (1 ♂, QM); Currawinya National Park, 10 mile bore, 0.5 km Northwest, 28 ° 52 ’ S, 144 ° 28 ’ E, 130 m, Eucalyptus, 13. X – 15. XII. 2007, C. Lambkin & N. Starick, Malaise trap, CNC 575118 (1 ♂, QM); Currawinya National Park, 15.1 km Southwest of Headquarters (CW 9 M), 28 ° 55 ’ S, 144 ° 26 ’ E, 131 m, Eremophila, 12 – 26. IX. 2008, Lambkin, Starick, Townsend, Malaise trap, 17241, CNC 574663 (1 ♂, QM); Plevna Downs, Tompilly Hill base (PD 6 M), 26 ° 44 ’ S, 142 ° 39 ’ E, 187 m, Gidgee, 24. IV – 24. VII. 2008, G. Turner & R. Mackenzie, Malaise trap, CNC 595284 (1 ♂, QM); 2.5 km West-Northwest of Plevna Downs Homestead (PD 3 P 2), 26 ° 40 ’ S, 142 ° 35 ’ E, 133 m, Gidgee, 25. xi – 17. xii. 2008, R. Mackenzie, pitfall trap, CNC 575271 – 2 (2 ♂, QM); 200 km west Windorah, 25 ° 25 ’ S, 142 ° 39 ’ E, ~ Malaise in dry creek bed, 4. ix. 1997, S. Winterton, J & A. Skevington, JSS 8326 – 9 (3 ♂, 1 ♀, QM); 204 km west Windorah, 25 ° 42 ’ S, 140 ° 53 ’ E, Malaise in dry creek bed, 4. ix. 1997, S. Winterton, J. & A. Skevington, JSS 8338 (1 ♂, QM); 91.2 km west Windorah, 25 ° 22 ’ S, 141 ° 47 ’ E, Malaise in dry creek bed, 31. viii – 5. ix. 1997, S. Winterton, J. & A. Skevington, JSS 8342 (1 ♂, QM); Charleville, 26 ° 24 ’ S, 146 ° 14 ’ E, 15. ii. 1970, A. L. Dyce, car net, ex. spirit via ethyl acetate, JSS 8619, JSS 8623 (2 ♂, ANIC); Kroombit Tops National Park, hilltop, 24 ° 27 ’ N, 150 ° 56 ’ E, 22. xii. 2014, J. H., A. M. & A. W. Skevington, CNC 384505 – 8 (4 ♂, CNC); Mount Moffatt summit, 25 ° 4 ’ S, 148 ° 3 ’ E, 1097 m, 18. xi. 2014, J. H., A. M. & A. W. Skevington, CNC 374868, CNC 374963, CNC 375031, CNC 375036 (4 ♂, CNC); Plevna Downs, Tompilly Hill base (PD 6 M), 26 ° 44 ’ S, 142 ° 39 ’ E, 187 m, Gidgee, 24. vii – 17. ix. 2008, N. Starick, C. Lambkin & R. Mackenzie, Malaise trap, CNC 574713 (1 ♂, QM); Ridgepole Waterhole, 24 km ESE of Musselbrook Resource Centre, Lawn Hill National Park, 18 ° 40 ’ S, 138 ° 22 ’ E, 7. v. 1995, G. Daniels & M. A. Schneider, JSS 8300 (1 ♀, QM); South Australia: 6 km South Maynard’s Bore, Everard Pk. Station, 34 ° 57 ’ S, 138 ° 34 ’ E, by beating Grevillea, 6. XI. 1970, G. Gross, JSS 8708 (1 ♂, SAM); Leigh Creek, 30 ° 29 ’ S, 138 ° 25 ’ E, 15. IX. 1972, Z. Liepa, JSS 9206 (1 ♂, ANIC); Near Victory Well, Everard Park Station, 27 ° 3 ’ S, 132 ° 30 ’ E, 2 – 4. XI. 1970, G. Gross, Malaise trap, JSS 8710 (1 ♂, SAM); Gammon Ranges National Park, south end of Weetootla Gorge, 30 ° 29 ’ S, 139 ° 16 ’ E, 537 m, hilltop, 11. X. 1997, J. Skevington & C. Lambkin, [hand collected, thorax and 2 legs remaining, rest used for sequencing], JSS 1259 (1 ♂, QM); JSM 10829, JSS 1261 (1 ♂, QM); Flinder’s Ranges National Park, Brachina Gorge, Heysen Hilltop, 31 ° 20 ’ S, 138 ° 33 ’ E, 9. x. 1997, J. & A. Skevington & C. Lambkin, JSS 8379 – 81, JSS 8389 – 90, JSS 8393 (6 ♂, QM); Gammon Ranges National Park, Weetootla Gorge, 30 ° 28 ’ S, 139 ° 13 ’ E, 537 m, 13. x. 1997, J. Skevington & C. Lambkin, JSS 8370, JSS 8372 – 4 (4 ♂, QM); Gammon Ranges National Park, south end of Weetootla Gorge, 30 ° 29 ’ S, 139 ° 16 ’ E, 537 m, hilltop, 11. x. 1997, J. Skevington & C. Lambkin, hand collected, JSS 4425 (1 ♂, QM), hilltop, 11. x. 1997, J. Skevington & C. Lambkin, JSS 8345, JSS 8347 – 8, JSS 8352 – 3, JSS 8356 – 7, JSS 8361, JSS 8363 – 4, JSS 8366 – 7 (12 ♂, QM); Koonamore, inside KVR, 32 ° 3 ’ S 139 ° 23 ’ E, salt bush sweep, 18. ii. 1974, P. Greenslade, [genitalia missing; head damaged], JSS 8818 (1 ♂, SAM); Near Victory Well, Everard Park Station, 27 ° 3 ’ S, 132 ° 30 ’ E, 2 – 4. xi. 1970, G. Gross, Malaise trap, JSS 8713 (1 ♂, SAM); 2. xi. 1970, G. F. Gross, Malaise trap, JSS 8711 – 2 (2 ♂, SAM); Swan Reach Conservation Park, 34 ° 35 ’ S, 139 ° 29 ’ E, [in mallee], 29. ix – 4. x. 1997, S. Winterton, J. & A. Skevington & C. Lambkin, Malaise trap, JSS 8420 (1 ♂, QM); Western Australia: 70 km S of Newman; Great Northern Highway, 23 ° 54 ’ S, 119 ° 45 ’ E, 626 m, near trees in small wash, 6 – 18. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, CNCD 6902 (1 ♂, CNC); 48 km NW Carnarvon; Blowholes Road; sandy trail, 24 ° 35 ’ S, 113 ° 31 ’ E, dry vegetated coastal dunes, 27. iv – 11. v. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15827 (1 ♂, CNC); 11 km E Marble Bar at Brockman Creek, 21 ° 9 ’ S, 119 ° 52 ’ E, ~ 187 m, damp, sandy creek bed, 2 – 14. v. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15683, JSS 15685 (2 ♂, CNC); 137 km S Newman, 30 km W Kumarina Road House, 24 ° 28 ’ S, 119 ° 40 ’ E, 21 – 23. v. 2003, M. E. Irwin & F. D. Parker, Malaise trap, 19. iv. – 23. v. 2003, JSS 16417 – 9 (1 ♂, 2 ♀, CNC); 15 km N Auski Road House, 22 ° 16 ’ S, 118 ° 46 ’ E, 402 m, water hole in Acacia woodlands, 3 – 16. v. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15846 (1 ♀, CNC); 158 km S Newman, 9 km N Kumarina Road House, 24 ° 38 ’ S, 117 ° 37 ’ E, ~ 638 m, in wide sandy wash, 18 – 21. v. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 16406, JSS 16408, JSS 16410 – 4 (4 ♂, 2 ♀, CNC; 1 ♀, ANIC); wide sandy wash, 24. iv – 7. v. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15652, JSS 15705, JSS 15707 – 8, JSS 15710 – 4 (8 ♂, 1 ♀, CNC); ~ 638 m, wide sandy wash, 7 – 18. v. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15719, JSS 15721, JSS 15724 – 5, JSS 15727 – 31 (8 ♂, 1 ♀, CNC); 171 km east of Marble Bar, 21 ° 17 ’ S, 121 ° 14 ’ E, 300 m, dry wash with flowers, 2 – 15. v. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15873 (1 ♀, CNC); 30 km ESE Three Rivers Station, dry bed of Gascoyne River, 25 ° 14 ’ S, 118 ° 57 ’ E, 504 m, dry river bed, 24. iv – 7. v. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15651 (1 ♀, CNC); 47 km S Pardoo Road House on Shay Gap Road, 20 ° 23 ’ S 120 ° 1 ’ E, 170 m, dry wash near Spinifex, 1 – 14. v. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15701 (1 ♂, CNC); 70 km S Newman on Great Northern Highway, 23 ° 54 ’ S, 119 ° 45 ’ E, 626 m, small wash with Eucalyptus, 24. iv – 6. v. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15882 (1 ♀, CNC); Karijini National Park, Banjima Drive, 10 km W visitors center, 22 ° 29 ’ S, 118 ° 22 ’ E, 743 m, 4 – 16. v. 2003, M. E. Irwin & F. D. Parker, JSS 15823 (1 ♂, CNC); Karijini National Park, Hamersley-Mount Bruce Road, 22 ° 34 ’ S, 118 ° 18 ’ E, 757 m, dry rocky creek bed, Eucalyptus grassland, 25. iv – 14. v. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16119 – 20, JSS 16122 (3 ♂, ANIC); Karijini National Park, Juna Downs Road, 22 ° 40 ’ S, 118 ° 26 ’ E; 22 ° 42 ’ S, 118 ° 25 ’ E, ~ 797 m, 789 m, in open mulga with flowering Ptilotus; between drying pools in rocky Turee Creek, 19 – 25. iv. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16171, JSS 16255, JSS 16286, JSS 16288, JSS 16290 – 1, JSS 16293 – 4, JSS 16297, JSS 16299 (6 ♂, 4 ♀, ANIC); 22 ° 40 ’ S, 118 ° 26 ’ E, 22 ° 44 ’ S, 118 ° 25 ’ E, 797 m, 798 m, in open mulga with flowering Ptilotus, dry Turee Creek, grassy open Eucalyptus scrub, 25. iv – 14. v. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 15993, JSS 15997, JSS 16001, JSS 16004, JSS 16088, JSS 16095, JSS 16307, JSS 16309 – 11 (6 ♂, 4 ♀, ANIC); Karijini National Park, Karijini Drive, 22 ° 34 ’ S, 118 ° 18 ’ E, 814 m, open Eucalyptus grassland, 25. iv – 14. v. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 15585 (1 ♀, ANIC); Karijini National Park, Weano Gorge Road, 22 ° 22 ’ S, 118 ° 15 ’ E, 22 ° 24 ’ S, 118 ° 15 ’ E, 775 m, 718 m, on hilltop, in open Eucalyptus, 25. iv – 14. v. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16145, JSS 16218, JSS 16232 – 4, JSS 16237 – 9 (5 ♂, 3 ♀, ANIC); Karijini National Park: Juna Downs Road (between drying pools in rocky Turee Creek), 22 ° 42 ’ S, 118 ° 25 ’ E, 789 m, 25. iv – 14. v. 2003, C. Lambkin & T. Weir, Malaise trap, CNCD 3822 (1 ♂, CNC); Kimberley, Lennard River Crossing, Gibb River Road, 17 ° 23 ’ S, 124 ° 44 ’ E, ~ in riverine thicket, 14 – 28. vii. 1988, T. F. Houston, Malaise trap, JSS 8734 – 5 (1 ♂, 1 ♀, WAM); Lane Poole Reserve, 32 ° 49 ’ S, 116 ° 6 ’ E, 315 m, Banksia grandis / Jarrah woodland on bauxite ridge, 7 – 13. xi. 2003, C. Lambkin & J. Recsei, Malaise trap, JSS 16064 (1 ♂, ANIC); Mount Augustus National Park, south slope on Saddle Trail, 24 ° 18 ’ S, 116 ° 48 ’ E, 400 m, small dry wooded wash, 25. iv – 7. v. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15886 (1 ♀, CNC); Millstream-Chichester National Park, Black Hill Pool, 21 ° 20 ’ S, 117 ° 15 ’ E, 176 m, over wide dry rocky creek bed, Eucalyptus, 7 – 12. v. 2003, C. Lambkin & D. Yeates, Malaise trap, JSS 16313 (1 ♂, ANIC); Millstream, 21 ° 35 ’ S, 117 ° 4 ’ E, 10. iv. 1971, D. H. Colless, JSS 9198 – 9 (2 ♂, ANIC); 12. iv. 1971, D. H. Colless, JSS 9192 – 3, JSS 9200 (3 ♂, ANIC); ~ 3 km north of Denham, 25 ° 55 ’ S, 113 ° 32 ’ E, hilltop, 12. xii. 1999, J. Skevington, JSS 7319, JSS 7343, JSS 7360, JSS 7365, JSS 7380, JSS 7387, JSS 7396, JSS 7407 (8 ♂, QM); 23 km E. Marble Bar, 21 ° 11 ’ S, 119 ° 57 ’ E, 222 m, Sandy wash, 15. v. 2003, F. D. Parker, M. E. Irwin, pan trap, CSIRO – SCHLINGER PILBARA EXPEDITION 19. iv. – 23. v. 2003, CNCD 6951, CNCD 6953 – 4, CNCD 6956, CNCD 6958 – 9 (2 ♂, 4 ♀, ANIC); Lochada, 1.2 km East-Southeast of Bolada Camp, 29 ° 12 ’ S, 116 ° 32 ’ E, 282 m, open York Gum woodland, Hakea, 15 – 18. IX. 2009, Lambkin, Monteith, Malaise trap, CNC 595382 – 3 (2 ♂, CNC); Australia: Rhodes grass, 19. I. 1966, accession # D. P.; R 2367; DC. HC. NY. D. L., JSS 8971 (1 ♂, QDPC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF67ECD4D8D9E957FD5F126C.taxon	distribution	Distribution: Australia (New South Wales, Queensland, South Australia, Western Australia) (Fig. 136). Notes: This is one of the most common species of Tomosvaryella in Australia and is widespread, mostly in the outback. Based on the shape of the genitalia, this species is close to T. latipoda sp. nov.. It differs by having rectangular-shaped surstyli, its apex is wider and covered with dense long bristles in dorsal view (Fig. 83 A), but the surstyli of T. latipoda sp. nov. is narrowed at the apex with long bristles before the apex (Fig. 53 A). Intraspecific genetic distance ranges from 0.0 % to 2.5 %. This species is genetically most similar to T. kooncheraensis sp. nov. (1.1 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF68ECD5D8D9ED0BFC18108C.taxon	description	Figs 84 A – E, 137, 147 D	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF68ECD5D8D9ED0BFC18108C.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a distinct, square keel with tiny bristles along its edge (Fig. 147 D); elongate surstyli in dorsal view (Fig. 84 A); gonopods with an extended finger-like projection towards surstyli (Fig. 84 B); phallic guide with four spines.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF68ECD5D8D9ED0BFC18108C.taxon	description	Description: MALE: Body length: 3.1 mm. Head. Flagellum acuminate; yellow. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose with a golden tinge; eyes touching for distance equal to 1.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper half. Thorax. Postpronotum pale, with 4 – 5 extra-long, pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum brownish pollinose and with 12 – 14 bristles along the distal edge. Dorsocentral bristles well developed, somewhat longer along frontal edge, numerous long pale bristles around postpronotum. Halter, knob pale, stem brown. Legs. Mid coxa with 2 – 3 long dark bristles. Trochanters brown, femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees yellow, tibiae black, tarsomeres yellow-brown ventrally and brown dorsally, last segment same, brown. Hind trochanter with a distinct, square keel (like a “ top hat ”) and with tiny bristles along the edge, somewhat longer at the distal corner (Fig. 147 D); ventrobasal spines (1 small, 1 large) present on fore and mid femora. Ventroapical row of spines on fore femur absent; 5 – 6 tiny, black spines on mid femur; no spines on hind femur, only 14 – 16 long pale bristles equally spaced, longest at 2 / 3 from base (up to as long as 1 – 1.2 times the width of hind tibia at the distal end), somewhat pressed against the surface of the femur. Subapical (distal) spines on first four tibiae absent. Hind tarsi slightly flattened (1 – 3), hind metatarsus as long as 2 – 4 combined. Pulvilli as long as last tarsal segment. Wing. Length: 3.5 mm. Upper side of basal costal cell with one long dark bristle. Fourth costal section 2.5 times as long as third costal section. Cross-vein r-m distal to middle of discal cell. 2 – 3 distinct brown setulae on tegula. Abdomen. Dissected. tergite 1 silvery grey. Lateral bristles on first tergite present, 6 – 8 short pale bristles in a row up to 1 – 1.2 times as long as the width of hind femur at distal tip in lateral view. Postabdomen in dorsal view: Dissected. Genitalia. Genital capsule in dorsal view: epandrium and surstyli brown. Epandrium wider than long (MLE: MWE = 0.6). Surstyli symmetrical, broadened at base, elongated, slender, pointing at apex, left is wider in middle and longer than right one (Fig. 84 A). Genital capsule in ventral view: gonopods equal in height with a finger-like projection towards surstyli; hypandrium broad, subepandrial sclerite widened and covered with transverse streaks (Fig. 84 B); phallus with three short ejaculatory ducts; phallic guide with four spines dorso-dorsolaterally (the longest one twice as length as others). Genital capsule in lateral view: both surstyli gently curved towards sternite (Figs 84 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 84 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF68ECD5D8D9ED0BFC18108C.taxon	etymology	Etymology: From the Latin quadratus, four-cornered or square, in reference to the square keel on the hind trochanter. Examined material: HOLOTYPE: AUSTRALIA: New South Wales: Siding Springs Observatory summit, 31 ° 16 ’ S, 149 ° 4 ’ E, 17. x. 1997, J. Skevington & C. Lambkin, JSS 8395 (1 ♂, QM); PARATYPE: AUSTRALIA: Queensland: Mount Moffatt summit, 25 ° 4 ’ S, 148 ° 3 ’ E, 1097 m, 18. XI. 2014, J. H., A. M. & A. W. Skevington, CNC 374994 (1 ♂, CNC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF68ECD5D8D9ED0BFC18108C.taxon	distribution	Distribution: Australia (New South Wales, Queensland) (Fig. 137). Notes: Both specimens of this rare species were caught hilltopping. This species is genetically similar to T. novaezealandiae (1.7 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF6EECD3D8D9EFF6FCA815B4.taxon	description	Figs 85 A – E, 138	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF6EECD3D8D9EFF6FCA815B4.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli, broadened in the basal two thirds, and narrowed to the apex in dorsal view (Fig. 85 A); gonopods asymmetrical, both extended towards surstyli, right one flattened at apex, left one pointed at apex and triangle-shaped; phallic guide pointed towards right side at apex in ventral view (Fig. 85 B); abdomen covered by dense bristles.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF6EECD3D8D9EFF6FCA815B4.taxon	description	Description: MALE: Body length: 3.4 mm. Head. Scape dark, pedicel and flagellum brown, Flagellum tapering; yellow-brown pollinose. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to the length of ocellar triangle. Occiput silvery pollinose with scattered short dark and pale bristles. Thorax. Postpronotum brown, with 3 – 4 long pale bristles. Mesonotum (viewed obliquely from front) subshining black, silvery pollinose, with long bristles at supra-alar area in a row, extended to posterior and uniseriate rows of long intra-alar bristles. Scutellum black with 4 – 6 lateral bristles along distal edge. Halter, knob pale, tip brownish, stem and base dark. Legs. Mid coxa with 3 – 4 long dark bristles. Trochanters brown, femora black, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments light brown. Coxae grey pollinose with some longer pale bristles ventroapically, Mid trochanter with two or three dark bristles posteroventrally Hind trochanter without any feature, grey pollinose with scattered short bristles ventroapically, 1 – 2 ventrobasal spines present on fore and mid femora. Hind femur with 6 – 8 ventroapical spines, with two rows of postero- and anteroventral bristles and two rows postero- and anterodorsal bristles, longest up to 2 / 3 the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres not flattened, hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Hyaline brown. Length: 3.5 mm. Upper side of basal costal cell with one long dark, one shorter dark and three pale bristles. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. Three short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites subshining brown, tergite 1 grey pollinose with 10 – 12 long dark lateral bristles up to as long as of hind femur’s width at base. All tergites covered by strong dark bristles. Genitalia. Genital capsule in dorsal view: epandrium as long as wide (MLE: MWE = 1.0). Surstyli rather symmetrical, broadened in basal two thirds, then narrowed to apex (Fig. 85 A). Genital capsule in ventral view: gonopods asymmetrical, both extended towards surstyli, right one flattened at apex, rather quadratic- shape, left one pointed at apex and triangle-shaped; phallic guide pointed towards right side at apex; subepandrial sclerite distinct (Fig. 85 B); Genital capsule in lateral view: both surstyli widened at the base, moderately narrowed at the middle, then curved to sternite (Fig. 85 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 85 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF6EECD3D8D9EFF6FCA815B4.taxon	etymology	Etymology: From the Latin reburrus, one with bristling hair, in reference to the densely bristled abdomen. Examined material: HOLOTYPE: AUSTRALIA: Queensland: St. Lucia, 27 ° 30 ’ S, 153 ° 0 ’ E, 1. iii. 1954 [date recorded as 1 - 3 - 54], S. Sekiton, JSS 8304 (1 ♂, QM); PARATYPE: AUSTRALIA: Queensland: Brisbane, 27 ° 28 ’ S, 153 ° 1 ’ E, 26. VI. 1964, G. B. Monteith, JSS 9704 (1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF6EECD3D8D9EFF6FCA815B4.taxon	distribution	Distribution: Australia (Queensland) (Fig. 138). Notes: Based on the terminalia characters this species is close to T. seticosta sp. nov. but the shape of surstyli in lateral view and the shape of the gonopods are different.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF6FECD0D8D9E973FD0E17A8.taxon	description	Figs 86 A – E, 139, 147 E, 158 A	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF6FECD0D8D9E973FD0E17A8.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having velvet-like bristles and some erect bristles ventrally (Fig. 147 E); elongate surstyli in dorsal view, left surstylus pointed towards outer margin (Fig. 86 A); phallus with three ejaculatory ducts, one covered by dense teeth at base (Fig. 86 D – E); gonopods widened in ventral view (Fig. 86 B); ejaculatory apodeme spade-shaped (Fig. 86 C).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF6FECD0D8D9E973FD0E17A8.taxon	description	Description: MALE: Body length: 2.9 – 3.1 mm. Head. Flagellum short acuminate; yellow. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose with a golden tinge; eyes touching for distance equal to 1.8 – 2.2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 3 – 4 extra short, pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum brownish pollinose and with 7 – 8 weak bristles along the distal edge. Dorsocentral bristles indistinct, very short and pale, somewhat longer along frontal edge, a few very short pale bristles around postpronotum. Halter, knob pale with brownish tinge, stem brown. Legs. Mid coxa with 1 – 5 short, dark bristles. Trochanters and femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 5 of tibiae yellow, extending to basal 1 / 3 along posterior edge (tibia otherwise black), tarsal segments dark brown, ventrally somewhat paler, last segment same dark brown. Hind trochanter without a keel, only velvet-like bristles ventrally with some (6 – 7) slightly longer, erect bristles, equally spaced (Fig. 147 E). Ventroapical row of 4 – 5 very weak spines on fore femur present; 5 – 6 small, black spines on mid femur; 5 – 6 short, dark spines on distal half on hind femur. Subapical (distal) spines on first four tibiae absent. Ventrobasal spines absent on fore and mid femora. Hind tarsi slightly flattened, metatarsus almost as long as 2 – 5 combined; stronger, peg-like dark spines on metatarsus laterally along anterior edge. Pulvilli as long as last tarsal segment. Wing. Length: 3.5 – 3.6 mm. Upper side of basal costal cell with 1 – 3 short dark bristle. Fourth costal section 3 – 3.5 times as long as third costal section (section 3 appears short). Cross-vein r-m distinctly distal to middle of discal cell. 2 – 3 distinct brown setulae on tegula. Abdomen. Viewed obliquely from front tergites subshining black with a green tinge, tergite 1 silvery grey, sides silvery pollinose on tergite 5 (very small). Dispersed strong dark bristles on all tergites absent. Lateral bristles on first tergite absent, only 3 – 4 indistinct, pale bristles. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1.6 – 1.8 times as long as ST 8. Genitalia without dissection: ST 8 medium sized and rounded, brown and with the occasional indistinct bristles, otherwise velvet-like covering of ST 8; membranous area drop-shaped, directed posteroventrally; epandrium brown, cerci pale, surstyli brown, elongated, uniformly broad. Genitalia. Genital capsule in dorsal view: epandrium and surstyli brown. Epandrium wider than long (MLE: MWE = 0.5). Surstyli rather symmetrical, elongated, covered by dense short bristles; right surstylus wider than left one in middle, gently curved to left one; left surstylus constricted before apex, pointed towards outer margin (Fig. 86 A). Genital capsule in ventral view: gonopods widened, equal in height; hypandrium broad and sclerotized, subepandrial sclerite small (Fig. 86 B); phallus with three distinct long sinuous ejaculatory ducts, one of ejaculatory ducts covered by dense teeth at base. Genital capsule in lateral view: both surstyli broadened in middle and bent towards sternite; phallic guide broad, pointed at apex (Fig. 86 D – E). Ejaculatory apodeme spade-shaped (Fig. 86 C). FEMALE: Body length: 2.8 – 2.9 mm. As male except for the following characters. Frons, eyes separated, as wide as 1.2 times the width at antennae; completely silver-grey pollinose until up to 1 / 3 of the lower frons, shiny black in front of ocellar triangle, gradually changing from pollinose to shiny black in the transition zone; frons depressed at the point where it is widest. Enlarged ommatidia silvery shining. Pulvilli and claws about 1.2 – 1.5 times as long as last tarsal segment, also on hind tarsi. Female abdomen with silver spots on sides of tergite 6 only. Ovipositor. Base of ovipositor dark brown with some gray pollinosity. Viewed laterally (Fig. 158 A), piercer straight. LP: LB = 2.1. LDP: LPP = 2.0. (Fig. 158 A)	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF6FECD0D8D9E973FD0E17A8.taxon	etymology	Etymology: The species is named scapania from the Greek skapane, for spade or hoe, in reference to the spade-shaped ejaculatory apodeme. Examined material: HOLOTYPE: AUSTRALIA: Queensland: 3.5 km SW by S Mount Baird, 15 ° 10 ’ S, 145 ° 7 ’ E, 3. v. 1981, D. H. Colless, Malaise trap, JSS 8583 (1 ♂, ANIC); PARATYPES: AUSTRALIA: Queensland: 10 miles South Bowen, 20 ° 8 ’ S, 148 ° 15 ’ E, 26. ix. 1950, E. F. Riek, JSS 8634 (1 ♂, ANIC); 2 km S of Horseshoe lookout, Blackdown land, 23 ° 50 ’ S, 149 ° 3 ’ E, 23 – 24. iv. 1981, D. H. Colless, JSS 8614 (1 ♂, ANIC); Carnarvon National Park, Mount Moffatt section, 25 ° 4 ’ S, 148 ° 1 ’ E, 26. xi. 1997, J. Skevington & C. Lambkin, Malaise trap, JSS 2300 (1 ♂, QM); Double Island Point, 25 ° 56 ’ S, 150 ° 11 ’ E, 110 m, 28. ix. 2002, S. A. Marshall, debu 210268 (1 ♂, DEBU); Proserpine, Thompson Creek; XY 15, 20 ° 31 ’ S, 148 ° 34 ’ E, 30 m, 3 – 10. v. 2007, J. Stanisic, 15028, CNCD 159594 – 5 (1 ♂, 1 ♀, CNC); Proserpine, Deadman Creek, XY 17, 20 ° 30 ’ S, 148 ° 34 ’ E, 21 m, 3. x – 10. v. 2007, J. Stanisic, 15030, CNCD 159288 – 93 (4 ♂, 2 ♀, QM); Tibrogargan, 26 ° 56 ’ S, 152 ° 57 ’ E, 1961, P. R. Webb, JSS 8262 (1 ♂, QM); Mareeba, 17 ° 0 ’ S, 145 ° 26 ’ E, 8. II. 1997, [Howden], flight interception trap, JSS 14388 (1 ♂, DEBU); Tinaroo Falls Dam, North Queensland, 17 ° 11 ’ S, 145 ° 34 ’ E, open savannah, 27. IV. 1967, D. H. Colless, JSS 138 (1 ♂, ANIC); Northern Territory: Arnhem Land, Maningrida, 12 ° 3 ’ S, 134 ° 13 ’ E, 5 m, 18. III. 1961; 19. III. 1961, J. L. & M. Gressitt, Malaise trap, CNCD 5565 – 66 (2 ♂, BPBM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF6FECD0D8D9E973FD0E17A8.taxon	distribution	Distribution: Australia (Northern Territory, Queensland) (Fig. 139). Notes: This is a forest species that occurs mostly along the coast from Brisbane to Darwin. Intraspecific genetic distance ranges from 0.0 % to 1.1 %. It is genetically closest to T. pseudophanes and T. quadrata sp. nov. (11.3 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF6DECAFD8D9E876FF63149C.taxon	description	Figs 87 A – E, 138	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF6DECAFD8D9E876FF63149C.taxon	diagnosis	Diagnosis: This species can be recognized by symmetrical and elongated surstyli in dorsal view (Fig. 87 A); gonopods extended towards surstyli, pointed at apex, right gonopod with a distinct projection on inner margin; phallic guide with 3 – 6 dorso- dorsolateral spines; large subepandrial sclerite with a small wrinkled edge in its center in ventral view (Fig. 87 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF6DECAFD8D9E876FF63149C.taxon	description	Description: MALE: Body length: 2.0 mm (without abdomen). Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, without bristles. Mesonotum (viewed obliquely from front) silvery pollinose, anterior part more grey, grayish also from the side. Scutellum silvery brown pollinose and with 4 – 6 bristles along distal edge. Dorsocentral bristles short, dark, longer in frontal half, numerous pale bristles around postpronotum present. Halter, knob pale, stem brown. Legs. Mid coxa with 5 – 6 long dark bristles. Trochanters brown, femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 5 of tibiae yellow (tibiae otherwise brown), tarsal segments yellow-brown dorsally, yellow ventrally. Hind trochanter covered with scattered short bristles. Hind femur without ventroapical spines, except 8 – 10 equally spaced bristles posteroventrally, longer on distal half, longest up to 2 / 3 the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae absent. Hind trochanter without keel, and 4 – 5 short spines, mostly proximally; ventrobasal spines absent on fore and mid femur. Hind tarsomeres slightly flattened, hind metatarsus as long as 2 – 4 combined, all tarsomeres with erect dark bristles dorsally. Pulvilli shorter than last tarsal segment. Wing. Length: 2.7 mm. Upper side of basal costal cell with two long dark brown bristles. Fourth costal section 2 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2 – 3 short dark setulae on tegula. Abdomen. Dissected. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.6). Surstyli symmetrical and elongated, curved to each other at apex, left surstylus longer than right one (Fig. 87 A). Genital capsule in ventral view: gonopods equal in height, extended towards surstyli, pointed at apex, right gonopod with a distinct projection on inner margin; phallic guide with 3 – 6 dorsolateral spines, one is longer than others; subepandrial sclerite large, with a small wrinkled edge in its center, hypandrial apodeme distinct (Fig. 87 B); Genital capsule in lateral view: both surstyli moderately narrowed to apex and curved towards sternite (Fig. 87 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 87 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF6DECAFD8D9E876FF63149C.taxon	etymology	Etymology: From the Latin seta, bristle and the Greek pteron, wing, in reference to the bristles on basal costal cell of the wing. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: Millstream-Chichester Nat [ional] P [ark], Roebourne Road, 21 ° 26 ’ S, 117 ° 9 ’ E, 337 m, sandy creek, Eucalyptus Spinifex grassland, 7 – 12. V. 2003, C. Lambkin & D. Yeates, Malaise trap, JSS 15574 (1 ♂, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF6DECAFD8D9E876FF63149C.taxon	distribution	Distribution: Australia (Western Australia) (Fig. 138). Note: This species is genetically closest to T. cona sp. nov. (3.6 – 9.5 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF13ECACD8D9EA5BFE191694.taxon	description	Figs 88 A – E, 135, 158 C	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF13ECACD8D9EA5BFE191694.taxon	diagnosis	Diagnosis: This species can be recognized by distinct spines along the posterior margin of sternites 2 – 4; hind trochanter with a small keel and some short bristles ventromedially; both gonopods with a long finger-like projection emerged from their bottom (Fig. 88 A); phallic guide long, pointed at apex, widened at the middle with two longer spines in its middle and some short, scattered spines mostly basally (Fig. 88 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF13ECACD8D9EA5BFE191694.taxon	description	Description: MALE: Body length: 3.5 – 3.6 mm. Head. Scape and pedicel dark brown. Pedicel with three upper bristles; flagellum light brown, gray pollinose and tapering. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 – 2 times the length of ocellar triangle. Occiput covered by short dark bristles. Thorax. Postpronotum light yellow, with 2 – 4 distinct light brown bristles in upper margin. Mesonotum (viewed obliquely from front) silvery pollinose, anterior part grey; scutum with scattered bristles on anterior supra-alar area. Postalar callus light brown with 3 – 4 brown bristles on upper margin. Scutellum black, silvery pollinose with 10 – 15 short bristles along distal edge. Halter, knob pale, stem light brown. Legs. Mid coxa with 4 – 6 dark bristles, two longer than others. Coxae, trochanters, femora dark brown, hind femur shining ventrally, knees and basal 1 / 4 of tibiae yellow (tibiae otherwise brown), tarsal segments yellow ventrally and light brown dorsally, last segment darker. Fore trochanter with two distinct bristles ventroapically. Hind trochanter silvery pollinose, with a small keel and 3 – 6 short bristles ventromedially; ventrobasal spines absent on fore and mid femur, two rows of ventroapical spines present. Hind femur with two rows ventroapical spines. Tibiae with two rows of short black bristle on anterior and three rows on posterior side. Hind tarsomeres distinctly flattened (especially tarsomeres 1 – 3), hind metatarsus as long as 2 – 4 combined; posterior corners with distinct bristles on all tarsomeres. Pulvilli shorter than last tarsal segment. Wing. Length: 3.5 mm. Upper side of basal costal cell with one dark brown bristle. Fourth costal section 1.5 – 2 times as long as third costal section. Cross-vein r-m at middle of discal cell. 5 – 8 short dark setulae on tegula. Abdomen. Tergites 1, 5 silvery pollinose; tergite 4 with two grey spots laterally, otherwise brown pollinose. Lateral bristles on first tergite present, 6 – 10 long brown bristles up to as long as hind femur’s width at base; sternite 2 with 3 – 6 small dark spines in the middle of posterior margin; sternite 3 – 4 with two distinct dark spines along posterior margin. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.8). Surstyli symmetrical, rather straight and rectangular-shaped, left surstylus slightly wider than right one, right surstylus curved to left one at apex (Fig. 88 A). Genital capsule in ventral view: gonopods rather unequal in height, left gonopod longer, right one wider than left one, both gonopods with long finger-like projection emerged from the base, left one extended towards surstyli, right one bent along inner margin of right gonopod; phallic guide long with two different-sized spines laterally, left spine longer and lower than right one; subepandrial sclerite wide and distinct (Fig. 88 B); Genital capsule in lateral view: both surstyli widened, left surstylus gently curved towards sternite; right surstylus straight in apical two thirds with a small projection in its basal towards hypandrium; phallic guide long, pointed at apex, widened at the middle with two longer spines centrally and some short scattered spines mostly at its base (Fig. 88 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 88 C). FEMALE: Body length: 3.2 mm. As male except for the following characters. Frons, eyes separated, as wide as the width at antennae; completely silver-grey pollinose on lower 1 / 3, shiny black only around directly ocellar triangle, smooth change in transition zone (just before ocellar triangle) to shiny black. Enlarged ommatidia silvery shining. Pulvilli and claws about 1.2 – 1.5 times as long as last tarsal segment. Tergites 3 – 6 and somewhat tergite 2 brown pollinose in its middle; Ovipositor. Straight in ventral view, straight, in lateral view (yellow piercer, dark brown base), reaching sternite 2; base silvery brown pollinose. LP: LB = 2.3. LDP: LPP = 2.0. (Fig. 158 C).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF13ECACD8D9EA5BFE191694.taxon	etymology	Etymology: From the Latin seta, bristle, in reference to the distinct spines along the posterior margin of sternites 2 – 4. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: Karijini National Park, Juna Downs Road, 22 ° 43 ’ S, 118 ° 25 ’ E, 799 m, near rock outcrop, Eucalyptus Eremophila scrub, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16047 (1 ♂, WAM); PARATYPES: AUSTRALIA: Queensland: 2 km along Ridgepole Waterhole Road, 10 km ESE Musselbrook Resource Centre, Lawn Hill National Park, 18 ° 38 ’ S, 138 ° 12 ’ E, 220 m, 10. V. 1995, G. Daniels & M. A. Schneider, JSS 8295 (1 ♀, QM); Western Australia: 35 km N Marble Bar, 20 ° 55 ’ S, 119 ° 59 ’ E, 160 m, 14. V. 2003, F. D. Parker & M. E. Irwin, hand netted in burned area from Eucalyptus and Abutilon dioicum (Malvaceae) flowers; CSIRO-SCHLINGER PILBARA EXPEDITION, April 19 – May 23, 2003, JSS 15751 (1 ♂, CNC); Karijini National Park, Hamersley-Mount Bruce Road, 22 ° 37 ’ S, 118 ° 19 ’ E, 762 m, open mallee with Spinifex, 14 – 19. V. 2003, C. Lambkin & J. Recsei, Malaise trap, JSS 15973 (1 ♂, ANIC); 22 ° 34 ’ S, 118 ° 18 ’ E, 757 m, dry rocky creek bed, Eucalyptus grassland, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16123 (1 ♀, ANIC); Karijini National Park, Juna Downs Road, 22 ° 44 ’ S, 118 ° 25 ’ E, 798 m, dry Turee Creek, grassy open Eucalyptus scrub, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16102 (1 ♀, ANIC); Karijini National Park: Juna Downs Road (between drying pools in rocky Turee Creek), 22 ° 42 ’ S, 118 ° 25 ’ E, 789 m, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, CNCD 3867 (1 ♀, CNC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF13ECACD8D9EA5BFE191694.taxon	distribution	Distribution: Australia (Queensland, Western Australia) (Fig. 135). Notes: This is a desert species found from NW Queensland to West Central Western Australia. Intraspecific genetic distance ranges from 0.0 % to 3.1 %. Based on the genetic divergence this may be a species complex and needs more investigation. It is genetically closest to T. millstreamensis sp. nov. (4.8 – 6.0 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF16ECA8D8D9EFF6FCA912DC.taxon	description	Figs 89 A – E, 139, 155 B	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF16ECA8D8D9EFF6FCA912DC.taxon	diagnosis	Diagnosis: This species can be recognized by the upper side of the basal costal cell having two long, brown bristles and 3 – 4 short, light brown bristles; symmetrical surstyli, broadened basally, moderately narrowed to apex in dorsal view (Fig. 89 A); gonopods with two different different-sized finger-like projections towards surstyli (Fig. 89 B); hypandrial apodeme distinct; subepandrial sclerite with a long protrusion along its middle in ventral view (Fig. 89 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF16ECA8D8D9EFF6FCA912DC.taxon	description	Description: MALE: Body length: 2.8 – 3.2 mm. Head. Flagellum short acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3, but it is silver all along the edge of the eyes. Thorax. Postpronotum pale, with 8 – 10 pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side and the posterior edge (2 small patches). Scutellum silvery pollinose and with 6 – 8 weak bristles along the distal edge. Dorsocentral bristles distinct, dark, somewhat longer along frontal edge, numerous dark bristles around postpronotum present. Halter, knob pale with brownish tinge, stem brown. Legs. Mid coxa with 4 – 5 long brown bristles. Trochanters and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 4 of tibiae yellow (tibiae otherwise brown), tarsal segments yellow-brown, ventrally paler, last segment darker. Hind trochanter with a half circle, no distinct bristles, half circle with flat part dorsally; two ventrobasal spines present on fore femur but absent on mid femur. Ventroapical row of spines on fore femur 5 – 7; 8 – 10 small, black spines on mid femur; hind femur without ventroapical spines, except 18 – 20 equally spaced bristles posteroventrally, longer on distal half, longest up to 1 / 2 the width of hind tibia at distal end, outer edge (lateral side) with 10 – 12 stronger dark spines in the middle line. Subapical (distal) spines on first four tibiae absent. Hind metatarsus moderately flattened and almost as long as 2 – 5 combined, with scrub-like bristles ventrally, strong spine-like bristles dorsally on all tarsi. Pulvilli shorter than last tarsal segment. Wing. Length: 3.5 – 3.6 mm. Upper side of basal costal cell with two long brown bristles and 3 – 4 short light brown bristles. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m at middle of discal cell. 4 – 5 distinct brown setulae on tegula. Abdomen. Viewed obliquely from front tergites subshining black, tergite 1 silvery grey, silvery pollinose lateral spots on tergite 5. Dispersed strong dark bristles on all tergites absent. Lateral spines on first tergite present, 5 – 7 short dark bristles up to 0.6 times as long as width of hind femur at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 3 times as long as ST 8. Genitalia without dissection: ST 8 small, rounded in dorsal view, brown and without distinct bristles, but with velvet-like coverage; membranous area round, mostly directed posteroventrally; epandrium brown, surstyli yellow and brown. Genitalia. Genital capsule in dorsal view: epandrium slightly longer than wide (MLE: MWE = 1.07). Surstyli rather symmetrical, broadened in basal two thirds, then narrowed to apex (Fig. 89 A). Genital capsule in ventral view: gonopods small with two finger-like projections towards surstyli, right is longer than left one, subepandrial sclerite with a protrusion along its middle; hypandrial apodeme prolapsed phallic guide strong (Fig. 89 B); Genital capsule in lateral view: both surstyli widened, curved towards sternite (Fig. 89 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 89 C). FEMALE: Body length: 3.1 – 3.2 mm. As male except for the following characters. Frons, eyes separated, as wide as 1.2 – 1.4 times the width at antennae; completely silver-grey pollinose, only shiny black directly in front of ocellar triangle, ca. 1 / 6 of the frons, very short transition zone; frons with a short ridge in front of the ocelli in the median line. Enlarged ommatidia silvery shining. Pulvilli and claws about 2 – 2.5 times as long as last tarsal segment, smaller on hind tarsomeres. Female abdomen with silver spots on sides of tergite 6. Ovipositor. Straight in ventral view, robust, curved towards sternite apically in lateral view (yellow piercer, brown base), reaching distal end of 3 rd segment; base brownish pollinose with occasional bristles. LP: LB = 3.5. LDP: LPP = 2.5. (Fig. 155 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF16ECA8D8D9EFF6FCA912DC.taxon	etymology	Etymology: The name refers to the bristles found on the basal costal cell and comes from the Latin seta, bristle. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Ridgepole Waterhole, 24 km ESE of Musselbrook Resource Centre, Lawn Hill Nat [ional] P [ar] k, 18 ° 40 ’ S, 138 ° 22 ’ E, 180 m, 13. V. 1995, G. Daniels, M. A. Schneider, JSS 8283 (1 ♂, QM); PARATYPES: AUSTRALIA: Queensland: Bulburin State Forest, 24 ° 32 ’ S, 151 ° 34 ’ E, 27 – 29. V. 1960, F. A. Perkins, JSS 8257 (1 ♂, QM); 200 km west Windorah, 25 ° 25 ’ S, 142 ° 39 ’ E, Malaise trap in dry creek bed, 4. IX. 1997, S. Winterton, J. & A. Skevington, JSS 8325 (1 ♀, QM); Ridgepole Waterhole, 24 km ESE of Musselbrook Resource Centre, Lawn Hill National Park, 18 ° 40 ’ S, 138 ° 22 ’ E, 180 m, 5. V. 1995, G. Daniels & M. A. Schneider, JSS 8299 (1 ♀, QM); Carnarvon Station near Homestead, 24 ° 48 ’ S, 147 ° 45 ’ E, 750 m, Eucalyptus woodland, 12. X. 2014, Lambkin, Wright, J. & K. Wilson, hand net, CNC 576417 (1 ♂, CNC); Western Australia: Karijini National Park, Juna Downs Road, 22 ° 40 ’ S, 118 ° 26 ’ E, 797 m, in open mulga with flowering Ptilotus, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 15991 (1 ♂, ANIC); New South Wales: Culgoa National Park, 8 km West-Northwest Cawwell Homestead, Diemunga Lagoon (CGN 2 M), 29 ° 3 ’ S, 146 ° 60 ’ E, Coolibah, 30. I – 18. V. 2010, C. Lambkin, R. Olsen & B. Shieban, Malaise trap, CNC 575120 (1 ♂, CNC); Northern Territory: Darwin, 12 ° 27 ’ S, 130 ° 50 ’ E, 6 – 9. XII. 1963; 8 – 9. XII. 1963, J. Sedlacek, CNCD 5571, CNCD 5575 (2 ♂, BPBM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF16ECA8D8D9EFF6FCA912DC.taxon	distribution	Distribution: Australia (New South Wales, Northern Territory, Queensland, Western Australia) (Fig. 139). Notes: This species is widespread but rarely collected.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF14ECA6D8D9ED9BFCF41328.taxon	description	Figs 90 A – E, 140, 157 F	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF14ECA6D8D9ED9BFCF41328.taxon	diagnosis	Diagnosis: This species can be recognized by 15 – 20 dark bristles on the hind trochanter ventrally; fore and mid trochanters with 2 – 3 short dark bristles; hind femur with extra-long bristles; phallic guide with a distinct sheath; phallus with a dagger-like projection at the base of ejaculatory ducts in ventral view (Fig. 90 B); small and triangle-shaped surstyli in dorsal view (Fig. 90 A); both gonopods widened in ventral view (Fig. 90 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF14ECA6D8D9ED9BFCF41328.taxon	description	Description: MALE: Body length: 2.6 – 2.8 mm. Head. Scape, pedicel and arista dark brown. Pedicel with 2 – 3 short upper bristles; flagellum brown, gray pollinose and tapering. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.2 times the length of ocellar triangle. Occiput grey pollinose. Thorax. Postpronotum light yellow, with 1 – 2 light brown bristles in upper margin. Mesonotum (viewed obliquely from front) silvery-brown pollinose, anterior part grey; scutum with scattered dark bristles on anterior supra-alar area, extended to posterior. Postalar callus dark brown with 3 – 4 short light brown bristles in the middle. Scutellum silvery pollinose and with a few (3 – 6) short dark bristles along the distal edge. Halter, knob pale, stem light brown, base dark. Legs. Fore coxa with a short dark bristle; mid coxa with 3 – 4 dark bristles, 1 – 2 bristles longer than others. Coxae, trochanters, femora dark, hind femur shining ventrally, knees and basal 1 / 3 of tibiae yellow (tibiae otherwise dark brown), tarsomeres light brown (last tarsomere rather brown dorsally). Fore and mid trochanter with 2 – 3 short dark bristles; hind trochanter silvery pollinose covered by 15 – 20 dark bristles ventrally; two ventrobasal spines present on fore and absent on mid femur. Hind femur with two rows long bristles ventrally and extra long bristles posteroventrally. Hind tibia with two rows of short black bristles on anterior and posterior side. Hind tarsomeres not flattened, covered by short brown bristles dorsally and brown bristles ventrally, hind metatarsus as long as 2 – 4 combined; pulvilli shorter than last tarsal segment. Wing. Length: 2.6 mm. Upper side of basal costal cell with one dark brown bristle. Fourth costal section 1.5 – 2 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2 – 3 short dark setulae on tegula. Abdomen. Tergites 1 silvery pollinose with 7 – 10 long brown lateral bristles; tergites 2 – 5 covered by scattered dark bristles. Syntergosternite 8 enlarged, dark brown and grey-brown pollinose. Membranous area long and broad in middle. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.8). Surstyli small (somewhat rectangular-shaped), widened at base, flattened at apex; left surstylus slightly longer (Fig. 90 A). Genital capsule in ventral view: both gonopods widened, right wider than left one (Fig. 90 B); phallic guide broad, pointed at apex; subepandrial sclerite small and rounded in the middle (Fig. 90 B); phallus with a dagger-like projection at the base of ejaculatory ducts (Fig. 90 B); Genital capsule in lateral view: both surstyli widened, dorsal margin straight, ventral margin curved, right surstylus with a small triangle-shaped projection in ventrobasally, phallic guide with a distinct sheath as long as phallic guide (Fig. 90 D – E); ejaculatory apodeme tube-like, bent, with a long bulb in its middle (Fig. 90 C). FEMALE: Body length: 2.6 mm. As male except for the following characters. Frons, eyes separated; completely silver-grey pollinose on lower 1 / 3, shiny black only around ocellar triangle, deep depression just before ocellar triangle. Enlarged ommatidia silvery shining; pulvilli and claws longer than last tarsal segment. Tergite 1 with 11 lateral bristles. Ovipositor. Straight in ventral view, long and straight in lateral view (piercer brown), reaching sternite 1; basal two thirds silvery brown pollinose. LP: LB = 3.5. LDP: LPP = 5.0. (Fig. 157 F).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF14ECA6D8D9ED9BFCF41328.taxon	etymology	Etymology: The specific name is derived from the Latin word ‘ sicula’, meaning dagger, referring to the dagger-like projection on the phallus. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: Karijini National Park, Juna Downs Road, 22 ° 42 ’ S, 118 ° 25 ’ E, 789 m, between drying pools in rocky Turee Creek, 19 – 25. IV. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16256 (1 ♂, WAM); PARATYPES: AUSTRALIA: Western Australia: Karijini National Park, Juna Downs Road, 22 ° 42 ’ S, 118 ° 25 ’ E, 789 m, between drying pools in rocky Turee Creek, 19 – 25. IV. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16165, JSS 16169, JSS 16173, JSS 16259 (2 ♂, 2 ♀, ANIC); Karijini National Park, Weano Gorge Road, 22 ° 23 ’ S, 118 ° 15 ’ E, 711 m, beside drying pool bank in grassland, 25. IV – 15. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 15978 (1 ♂, ANIC); Karijini National Park: Juna Downs Road (between drying pools in rocky Turee Creek), 22 ° 42 ’ S, 118 ° 25 ’ E, 789 m, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, CNCD 3826, CNCD 3829, CNCD 3851 (1 ♂, 2 ♀, QM); Queensland: Carnarvon National Park, Mount Moffatt section, near Kennifs Lookout (MM 1 M), 24 ° 55 ’ S, 148 ° 0 ’ E, 859 m, open Eucalyptus woodland, 13. XI – 13. XII. 2010, Reeves, Sternberg, Spinaze, Malaise trap, CNC 592058 (1 ♂, CNC); 1 km E Mt. Molloy, 16 ° 41 ’ S, 145 ° 20 ’ E, 15. IV. 1980, G. F. Hevel & J. A. Fortin, JSS 10526 (1 ♂, USNM); 3.5 km SW by S Mount Baird, 15 ° 10 ’ S, 145 ° 7 ’ E, 5. V. 1981, D. H. Colless, Malaise trap, JSS 8573, JSS 8575 (2 ♂, ANIC); Carnarvon, National Park, Marlong Plain, 24 ° 57 ’ S, 147 ° 58 ’ E, grassland, 13. X. 2002, S. Boucher, sweep net, LEM _ 0016171 (1 ♂, LEM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF14ECA6D8D9ED9BFCF41328.taxon	distribution	Distribution: Australia (Queensland, Western Australia) (Fig. 140). Notes: Despite the disjunct populations, we could not find any variation in external morphological and terminalia characters, so we consider it to be one widespread species. We could not successfully DNA barcode the Queensland specimens so future work should focus on testing the species concept by barcoding material from the eastern part of the range. Intraspecific genetic distance ranges from 0.0 % to 0.3 %. This species is genetically closest to T. bulbosa sp. nov. (8.3 – 12.1 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF1AECA4D8D9ECCFFA8F134C.taxon	description	Figs 91 A – E, 141, 150 A	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF1AECA4D8D9ECCFFA8F134C.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter with a keel and 6 – 7 dark spines (Fig. 150 A); slender and elongated surstyli in dorsal view (Fig. 91 A); widened gonopods, pointed at apex, with a distinct projection on inner margin, their projections with an upwards small finger-like process in the middle (Fig. 91 B); phallic guide with 3 – 4 short dorso- dorsolateral spines; subepandrial sclerite widened, with a fin-shaped projection in its center (Fig. 91 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF1AECA4D8D9ECCFFA8F134C.taxon	description	Description: MALE: Body length: 3.1 – 3.1 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 3 – 4 short pale bristles. Mesonotum (viewed obliquely from front) silvery pollinose, anterior part more grey, greyish also from the side. Scutellum silvery brown pollinose and with 4 – 6 bristles along distal edge. Dorsocentral bristles short, dark, longer in frontal half, numerous pale bristles around postpronotum present. Halter knob pale, tip brownish, stem brown. Legs. Mid coxa with 3 – 4 long dark bristles. Trochanters brown, femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments bright yellow. Hind trochanter with 45 - degree keel, and 6 – 7 dark spines, longest proximally (Fig. 150 A); one ventrobasal spine present on fore femur, absent on mid femur. Hind femur without ventroapical spines, except 8 – 10 equally spaced bristles posteroventrally, longer on distal half, longest up to 2 / 3 the width of hind tibia at distal end. Short, subapical (distal) spines on fore- and mid tibiae present. Hind tarsomeres distinctly flattened (t 1 – 3), hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 3.2 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 4 – 5 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites subshiny brown-black, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (spot on tergite 5 larger). Dispersed strong dark bristles on all tergites present, longest on tergite 5 in length up to 1 / 3 the width of hind tibia at distal end. Lateral bristles on first tergite present, 4 – 5 dark bristles up to as long as ¾ of hind femur’s width at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 0.6 – 0.7 times as long as ST 8. Genitalia without dissection: ST 8 large, round, globular in dorsal view, brown and with distinct bristles, otherwise with velvet-like coverage; membranous area is slit-like, rounded, mostly directed posteriorly; epandrium brown, surstyli brown. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.2). Surstyli slender and elongated, curved towards midline at apex, left surstylus is longer and its apex is wider than right one (Fig. 91 A). Genital capsule in ventral view: both gonopods widened at base, extended towards surstyli, pointed at apex, gonopods with a distinct projection on inner margin, their projections with a small finger-like process in the middle; phallic guide with 3 – 4 short dorsolateral spines; subepandrial sclerite widened, with a fin-shaped projection in its center, its left margin covered with condensed short hairs (Fig. 91 B); Genital capsule in lateral view: both surstyli curved towards sternite, left one more widened, with hump-like edge dorsomedially (Fig. 91 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 91 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF1AECA4D8D9ECCFFA8F134C.taxon	etymology	Etymology: This species is named after one of the dominant arid Australian grasses and is associated with Spinifex dominated grassland. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: Millstream-Chichester Nat [ional] P [ark], Roebourne Road, 21 ° 26 ’ S, 117 ° 9 ’ E, 337 m, sandy creek, Eucalyptus Spinifex grassland, 27. IV – 3. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16200 (1 ♂, WAM); PARATYPES: AUSTRALIA: Western Australia: 48 km NW Carnarvon; Blowholes Road; sandy trail, 24 ° 35 ’ S, 113 ° 31 ’ E, 7 m, dry vegetated coastal dunes, 27. IV – 11. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15826, JSS 15829 (1 ♂, CNC; 1 ♂, QM); Badgingarra National Park, 40 km east of Cervantes, 30 ° 24 ’ S, 115 ° 25 ’ E, 30. X. 1987, M. E. Irwin & E. I. Schlinger, JSS 8459 (1 ♂, CAS).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF1AECA4D8D9ECCFFA8F134C.taxon	distribution	Distribution: Australia (Western Australia) (Fig. 141). Note: Based on the DNA sequencing, this species is genetically similar to T. latistyla sp. nov. (4.0 – 5.4 % pairwise divergence). Intraspecific genetic distance ranges from 0.0 % to 1.3 % (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF18ECA2D8D9ECEBFD0E1328.taxon	description	Figs 92 A – E, 140, 146 C	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF18ECA2D8D9ECEBFD0E1328.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a patch of short dark bristles at the base and a small group of shorter bristles at distal end (Fig. 146 C); hind femur with 22 – 24 pale long bristles posteriorly on distal half (Fig. 146 C); subepandrial sclerite covered with transverse streaks and a row of finlike projection in its middle in ventral view (Fig. 92 B); phallic guide with six different-sized spines dorsolaterally.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF18ECA2D8D9ECEBFD0E1328.taxon	description	Description: MALE: Body length: 3.3 – 3.4 mm. Head. Flagellum acuminate, yellow-brown. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose; eyes touching for distance equal to 2.5 – 3 times the length of ocellar triangle. Occiput silvery pollinose, more brownish on upper half. Thorax. Postpronotum brownish pale, with 3 – 4 distinct pale bristles. Mesonotum (viewed obliquely from front) subshining black, anterior part silver pollinose, slightly grayish also from the side. Scutellum silvery pollinose and with 12 short pale bristles along the distal edge. Dorsocentral bristles developed, dark, longer on frontal part and numerous dark bristles around postpronotum. Halter, knob pale with a brownish tinge distally, stem dark brown. Legs. Mid coxa with two bristles. Trochanters and base of femora black, femora black, shining ventrally, silvery pollinose except hind femur; knees, tibiae brown with basal 1 / 6 yellow, tarsal segments yellow ventrally, yellow-brown dorsally, last segment darker dorsally. Hind trochanter without keel, short dark bristles (7 – 9, longest at base, shortening distally) at the base emerging in a group (“ echidna ” - like) and distal end with a small group of shorter bristles (Fig. 146 C); two ventrobasal bristles on fore femur (one is longer). Ventroapical row of spines on fore femur missing, 4 – 6 very short, black spines on mid femur; no spines on hind femur, only 22 – 24 pale bristles posteriorly equally distributed, and longer on distal half (as long as 1.5 – 2 times the width of hind tibia at the distal end) (Fig. 146 C); hind tibia bent at middle in posterior view. Subapical (distal) spines on fore- and mid-tibiae present. Hind metatarsus not flattened, first tarsomere slightly longer than 2 – 4 combined; ventral bristles scrub-like, yellow. Pulvilli as long as or slightly longer than last tarsal segment. Wing. Length: 3.1 – 3.2 mm. Upper side of basal costal cell with 1 – 3 long distinct bristles. Fourth costal section 2.5 – 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 5 – 6 dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brownish pollinose, tergite 1 silvery grey, sides silvery pollinose on tergites 4 and 5 (the latter with larger spots), slightly silver pollinose on tergites 2 and 3. Dispersed short dark bristles on tergites present, the longest on tergite 5 and up to 1.5 times as long as width of hind tibia at base. Lateral bristles on first tergite present, 5 – 6 brown bristles up to as long as width of hind femur at distal tip. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1 – 1.5 times as long as ST 8. Genitalia without dissection: ST 8 medium sized, subshining; membranous area elongated, widening at top; epandrium paler; yellow surstyli appear long and flat. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.2). Surstyli light brown, rather symmetrical with broad base and finger-like apex, left slightly broader and longer than right one; both tips bent inward apically (Fig. 92 A). Genital capsule in ventral view: subepandrial sclerite covered with transverse streaks, in its middle with a row of small fin-like projection embedded (like spinal column); gonopods elongated close to apex of subepandrial sclerite, with two small projections (Fig. 92 B), phallic guide with six different-sized spines dorsolaterally (two longer ones closer to apex). Genital capsule in lateral view: both surstyli curved towards sternite, gradually narrowed to apex (Fig. 92 D – E); phallus with three ejaculatory ducts; ejaculatory apodeme flat, sperm pump elongated, linear (Fig. 92 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF18ECA2D8D9ECEBFD0E1328.taxon	etymology	Etymology: From the Latin word spina for thorn and various for different in reference to the variously sized spines on the phallic guide. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: ~ 3 km N [orth] of Denham, 25 ° 55 ’ S, 113 ° 32 ’ E, hilltop, 12. xii. 1999, J. Skevington, JSS 7374 (1 ♂, WAM); PARATYPES: AUSTRALIA: Western Australia: ~ 3 km north of Denham, 25 ° 55 ’ S, 113 ° 32 ’ E, hilltop, 12. xii. 1999, J. Skevington, JSS 7325, JSS 7362, JSS 7390, JSS 7400 (1 ♂, CNC; 3 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF18ECA2D8D9ECEBFD0E1328.taxon	distribution	Distribution: Australia (Western Australia) (Fig. 140). Notes: Most of the specimens of this species were collected on the small hilltop north of Denham, Western Australia. See the note under T. minychoma sp. nov. for more information on this amazing locality. Intraspecific genetic distance ranges from 0.2 % to 1.0 %. This species is genetically most similar to T. velutina sp. nov. (5.7 – 8.2 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF1EECA3D8D9ECCFFD6F1110.taxon	description	Figs 93 A – E, 140	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF1EECA3D8D9ECCFFD6F1110.taxon	diagnosis	Diagnosis: This species can be recognized by the elongated surstyli, the left surstylus with a small edge along ventromedial margin (Fig. 93 A); both gonopods with a finger-like projection towards surstyli; subepandrial sclerite with an elongated suture in its middle; phallic guide with 3 – 4 short dorso- dorsolateral spines in ventral view (Fig. 93 B); left surstylus bent towards sternite in apical fourth, right one rather straight in lateral view (Fig. 93 D – E)	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF1EECA3D8D9ECCFFD6F1110.taxon	description	Description: MALE: Body length: 2.4 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2.2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 3 – 4 long pale bristles. Mesonotum (viewed obliquely from front) silvery pollinose, anterior part more grey, grayish also from the side. Scutellum silvery brown pollinose and with 2 – 4 short bristles along distal edge. Halter, knob pale, tip brownish, stem brown. Legs. Mid coxa with 3 – 4 long dark bristles. Trochanters light brown, femora dark brown, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments light brown. Hind trochanter with 2 – 4 short bristles anteroventrally. Hind femur without ventroapical spines. Subapical (distal) spines on fore tibiae present (short). Hind tarsomeres not flattened, hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 2.4 mm. Upper side of basal costal cell with one long brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 4 – 5 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brown, tergite 1 silvery grey. Lateral bristles on first tergite present, 2 – 4 dark bristles up to as long as ¾ of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.1). Surstyli elongate, curved to each other at apex, left surstylus is longer, with a small edge along ventromedial margin (Fig. 93 A). Genital capsule in ventral view: both gonopods with a finger-like projection towards surstyli; subepandrial sclerite with an elongated suture in its middle like spinal cord; phallic guide with 3 – 4 short dorso- dorsolateral spines, one is longer (Fig. 93 B); Genital capsule in lateral view: left surstylus bent towards sternite in apical fourth, right one straighter (Fig. 93 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 93 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF1EECA3D8D9ECCFFD6F1110.taxon	etymology	Etymology: This species is named in honour of Noel Starick, one of the collectors of the only known specimen. Noel has collected or contributed to the collection of 3 % of the specimens and 17 % of the species used in this revision (Supplementary file 2). Examined material: HOLOTYPE: AUSTRALIA: Queensland: Culgoa Floodplain National Park, 11.8 km N Headquarters (CG 2 AM), 28 ° 54 ’ S, 147 ° 8 ’ E, 151 m, Brigalow, 18. XI – 17. XII. 2009, C. Lambkin, A. Coward, C. Kelly & N. Starick, Malaise trap, CNC 576508 (1 ♂, CNC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF1EECA3D8D9ECCFFD6F1110.taxon	distribution	Distribution: Australia (Queensland) (Fig. 140).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF1CECBFD8D9EFF6FAA013BC.taxon	description	Figs 94 A – D, 142, 156 F	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF1CECBFD8D9EFF6FAA013BC.taxon	diagnosis	Diagnosis: This species can be recognized by the asymmetrical surstyli, with the right surstylus widened in the basal half in dorsal view (fig. 94 A), both with a distinct an upwards horn-like projection in the ventromedial margin in lateral view (Fig. 94 C – D); gonopods and subepandrial sclerite small (Fig. 94 B); hypandrium wide; phallus strong, one side covered by dense short forked hairs from its middle to apex in lateral view (Fig. 94 C).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF1CECBFD8D9EFF6FAA013BC.taxon	description	Redescription: MALE: Body length: 2.8 – 3.1 mm. Head. Scape, pedicel and arista dark brown. Pedicel with 2 – 3 bristles located on upper side of pedicel; flagellum brown, lighter at apex, gray pollinose and tapering. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal the length of ocellar triangle. Occiput grey pollinose. Thorax. Postpronotum light yellow, with 1 – 2 long, light brown, and 2 – 5 short pale bristles on upper margin. Mesonotum (viewed obliquely from front) silvery-brown pollinose; scutum with scattered dark bristles at anterior supra-alar area, extending to posterior and uniseriate rows of intra-alar bristles. Postalar callus dark brown with 1 – 3 short light brown bristles in the middle. Scutellum silvery pollinose and with 8 – 10 short dark bristles along the distal edge. Halter, knob pale, stem light brown, base dark. Legs. Mid coxa with 2 – 3 dark bristles. Coxae, trochanters, femora dark, hind femur shining ventrally, knees and basal 1 / 3 of tibiae yellow (tibiae otherwise dark brown), tarsomeres light brown ventrally, dark brown dorsally. Fore and mid trochanter with 2 – 3 short dark bristles; hind trochanter silvery pollinose covered by some scattered tiny dark bristles ventrally; ventrobasal spines absent on fore mid femur. Hind femur with two rows of short spines ventrally. Hind tibia with two rows of short brown bristles on anterior and posterior side. Hind tarsomeres slightly flattened on one side, covered by short light brown bristles dorsally, hind metatarsus as long as 2 – 4 combined; pulvilli shorter than last tarsal segment. Wing. Length: 3.0 mm. Upper side of basal costal cell with 1 – 2 long, dark brown bristles and two long, light brown bristles inside basal costal cell. Fourth costal section 1.5 – 2 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2 – 3 short dark setulae on tegula. Abdomen. Tergites 1 silvery pollinose with 5 – 7 different-sized lateral bristles; tergite 2 – 5 brown pollinose, covered by scattered short dark bristles. Syntergosternite 8 enlarged, dark brown and grey-brown pollinose. Membranous area long, approaching epandrium. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.8), with a depression in its middle (Fig. 94 A). Surstyli asymmetrical, widened at base, equal in height, left surstylus rectangular-shaped, right surstylus widened in basal half, then narrowed to apex (Fig. 94 A). Genital capsule in ventral view: both gonopods small, widened and rounded at apex right wider than left one (Fig. 94 B); phallic guide short and broad; subepandrial sclerite small and wide, right side with a distinct wrinkle, phallus strong with one ejaculatory duct, hypandrium broadened, hypandrial apodeme small but distinct (Fig. 94 B); Genital capsule in lateral view: both surstyli rather straight, with a distinct horn-like projection in ventromedial margin, then narrowed to apex, base of right surstylus wider than left one; phallus strong and sinuate, one side covered by condensed furcate? hairs from its middle to apex (Fig. 94 C – D). FEMALE: Body length: 2.5 mm. As male except for the following characters. Frons, eyes separated; completely silver-grey pollinose on lower 1 / 3, wider in middle, shiny black only around ocellar triangle, deep depression just before ocellar triangle. Enlarged ommatidia silvery shining; pulvilli and claws longer than last tarsal segment. Tergites 2 – 5 with lateral gray spots. Ovipositor. Straight in ventral view, straight in lateral view (piercer brown), reaching sternite 2; basal two thirds silvery brown pollinose. LP: LB = 2.0. LDP: LPP = 2.8. (Fig. 156 F). Examined material: LECTOTYPE [HERE DESIGNATED]: AUSTRALIA: Queensland: Bundaberg, 24 ° 52 ’ S, 152 ° 21 ’ E, 20. VIII. 1919, F. Muir, CNCD 5542 (1 ♂, BPBM); PARALECTOTYPES: same data as lectotype, IX. 1904, BMNH (E) 964036 – 41, CNCD 5537 – 8, CNCD 5540, CNCD 5543 (6 ♂, 4 ♀, BMNH; 2 ♂, 2 ♀, BPBM); Cairns, 16 ° 55 ’ S, 145 ° 46 ’ E, VII. 1904, JSS 8486 – 7 (1 ♂, 1 ♀, BPBM); Childers, 25 ° 14 ’ S, 152 ° 16 ’ E, 30. X. 1904, [this specimen clearly reared by Perkins but no host data given. Two dates on label, first is “ Oct. 15 04 ” - likely the date of pupariation. “ Oct. 30 04 ” is likely the date of emergence], CNCD 5541 (1 ♂, BPBM); Mid. Queensland, 23 ° 7 ’ S 144 ° 8 ’ E, JSS 4180 – 2 (2 ♂, 1 ♀, ZMAN); 20 ° 55 ’ S, 142 ° 42 ’ E, JSS 4183 (1 ♂, ZMAN); Other Material Examined: AUSTRALIA: Queensland: 2 km SE of Coop’s Corner, end of power line road, 26 ° 3 ’ S, 153 ° 3 ’ E, vegetation, 8. X. 2002, S. Boucher, sweep net, LEM _ 0016183 (1 ♂, LEM); Carnarvon Nat. Park, Ranger Station, 25 ° 1 ’ S, 147 ° 57 ’ E, near campsite in morning, 11. X. 2002, S. Boucher, sweep net, LEM _ 0016209 (1 ♂, LEM); Carnarvon National Park, The Tombs, 25 ° 5 ’ S, 147 ° 52 ’ E, vegetation, 10. X. 2002, S. Boucher, sweep net, LEM _ 0016195 (1 ♂, LEM); Carnarvon, National Park Ranger Station, 25 ° 1 ’ S, 147 ° 57 ’ E, near campsite in morning, 11. X. 2002, S. Boucher, sweep net, LEM _ 0016208 (1 ♂, LEM); Emu Creek Crossing, East Petford, 17 ° 21 ’ S, 144 ° 56 ’ E, grass, 22. IV. 1983, J. F. Grimshaw, sweep net, JSS 8944 (1 ♂, QDPC); Karumba, 12 ° 28 ’ S, 130 ° 47 ’ E, 18. IV. 1983, J. F. Donaldson, JSS 8943 (1 ♂, QDPC); Mount Webb National Park, 15 ° 4 ’ S, 145 ° 7 ’ E, 30. IX. 1980, D. H. Colless, Malaise trap, JSS 223 (1 ♂, ANIC); Brampton Island, east of Mackay, 20 ° 48 ’ S, 149 ° 16 ’ E, 17. IX. 1963, D. E. Havenstein, JSS 8656 (1 ♂, ANIC); Brisbane, 27 ° 28 ’ S, 153 ° 1 ’ E, 11. IV. 1962, B. Wilson, JSS 8259 (1 ♂, QM); 23. III. 1951, S. Barker, JSS 8302 (1 ♂, QM); Culgoa Floodplain National Park, 11.8 km N Headquarters (CG 2 AM), 28 ° 54 ’ S, 147 ° 8 ’ E, 151 m, Brigalow, 18. XI – 17. XII. 2009, C. Lambkin, A. Coward, C. Kelly, N. Starick, Malaise trap, 19255, CNC 576509 (1, CNC); Currawinya National Park, 4 km NW Headquarters (CW 3), 28 ° 49 ’ S, 144 ° 28 ’ E, Mulga, 15. XII. 2007 – 8. II. 2008, Lambkin, Townsend, Starick, Malaise trap, CNC 576537 (1 ♂, CNC); Dunwich, North Stradbroke Island, 27 ° 35 ’ S, 153 ° 37 ’ E, 22. VIII. 1958, S. S. Sekhon, JSS 8264 (1 ♂, QM); Mount Crosby, 27 ° 33 ’ S, 152 ° 48 ’ E, 15. X. 1970, J. Wilson, JSS 9741 (1 ♂, QM); McKinlay district, NW Queensland, 25 ° 55 ’ S, 144 ° 36 ’ E, 17 – 19. VIII. 1963, T. E. Woodward, JSS 8267 (1 ♂, QM); Normanton, 17 ° 40 ’ S, 141 ° 4 ’ E, swept near swamp, 19. V. 1976, I. R. Bock, JSS 8552 (1 ♂, ANIC); Plevna Downs, Tompilly Hill base (PD 6 M), 26 ° 44 ’ S, 142 ° 39 ’ E, 187 m, Gidgee, 24. IV – 24. VII. 2008, G. Turner, R. Mackenzie, Malaise trap, CNC 595283 (1 ♂, CNC); Ridgepole Waterhole, 24 km ESE of Musselbrook Resource Centre, Lawn Hill National Park, 18 ° 40 ’ S, 138 ° 22 ’ E, 180 m, 1. V. 1995, G. Daniels, M. A. Schneider, JSS 8296 (1 ♂, QM); Rosevale Area, SE Queensland, 27 ° 52 ’ S, 152 ° 29 ’ E, 16. III. 1975, B. K. Cantrell, JSS 8933 (1 ♂, QDPC); Toowong, 27 ° 28 ’ S, 152 ° 59 ’ E, 30. III. 1952, DJT, JSS 8280 (1 ♂, QM); Babinda, 17 ° 20 ’ S, 145 ° 56 ’ E, IV. 1919, F. X. Williams, CNCD 5539 (1 ♂, BPBM); Meringa, 17 ° 5 ’ S, 145 ° 47 ’ E, IX. 1919, F. Muir, CNCD 5536 (1 ♀, BPBM); Carnarvon Station, near Conglomerate Spring (SSS 2), 24 ° 50 ’ S, 147 ° 47 ’ E, 839 m, dry creek bed in gully, 8 – 14. X. 2014, C. Lambkin, Malaise trap, CNC 591900 (1 ♂, CNC); Carnarvon National Park, Marlong Plain, 24 ° 57 ’ S, 147 ° 58 ’ E, grassland, 13. X. 2002, S. Boucher, sweeping, CNCD 253910 (1 ♂, LEM); Carnarvon National Park, Mount Moffatt, Angelina Creek, 25 ° 4 ’ S, 148 ° 3 ’ E, 675 m, creek bed, over creek, 9 – 10. X. 2002, S. A. Marshall, Malaise trap, debu 210512 (1 ♂, DEBU); Dauan Island, North Queensland, 9 ° 25 ’ S, 142 ° 32 ’ E, 28. III. 1984, J. W. Turner, JSS 8953, JSS 8955 – 6 (3 ♂, QDPC); Edward Allison Park, Eatons Hill, Brisbane, 27 ° 20 ’ S, 152 ° 57 ’ E, 10. XI. 1995, S. G. Evans, JSS 12613 (1 ♂, QM); Elizabeth Creek, Boodjamulla National Park, 18 ° 14 ’ S, 138 ° 5 ’ E, 170 m, Bloodwood open forest, 18 – 22. IV. 2005, M. Mathieson, G. Smith, Malaise trap, JSS 19190 (1 ♀, QM); Eungella Range, 52 miles W Mackay, 21 ° 9 ’ S, 148 ° 24 ’ E, 15. IX. 1963, D. E. Havenstein, JSS 8646 – 52 (7 ♂, ANIC); Peawaddy Gorge, Mount Moffatt National Park, 24 ° 56 ’ S, 148 ° 4 ’ E, 1100 m, 26. II. 1996, C. J. Burwell & S. Evans, JSS 9744 (1 ♂, QM); Scrub Road, Brisbane Forest Park, 27 ° 25 ’ S, 152 ° 50 ’ E, 21 – 28. XI. 1997, S. Winterton, N. Power & D. White, Malaise trap, JSS 8847 (1 ♂, QM); Western Australia: 8 km S of Cape Bertholet, 17 ° 15 ’ S, 122 ° 10 ’ E, 18. IV. 1977, D. H. Colless, JSS 9256 (1 ♂, ANIC); Derby, 17 ° 23 ’ S, 123 ° 40 ’ E, 27. I. 1967, G. Beamish, JSS 8736 (1 ♂, WAM); Karijini National Park, Juna Downs Road, 22 ° 40 ’ S, 118 ° 26 ’ E, 797 m, in open mulga with flowering Ptilotus, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, ANIC [Bulk Sample] 2046, JSS 15987 (1 ♂, ANIC); 48 km NW Carnarvon; Blowholes Road; sandy trail, 24 ° 35 ’ S, 113 ° 31 ’ E, 7 m, dry vegetated coastal dunes, 27. IV – 11. V. 2003, M. E. Irwin, F. D. Parker, Malaise trap, JSS 15831 (1 ♀, CNC); Crawley, 31 ° 59 ’ S, 115 ° 49 ’ E, 2. IV. 1924, K. R. Norris, JSS 8644 (1 ♂, ANIC); Karijini National Park, Hamersley-Mount Bruce Road, 22 ° 34 ’ S, 118 ° 18 ’ E, 757 m, dry rocky creek bed, Eucalyptus grassland, 25. IV – 14. V. 2003, C. Lambkin, T. Weir, Malaise trap, ANIC [Bulk Sample] 2056, JSS 16121 (1 ♂, ANIC); Millstream-Chichester National Park, Roebourne Road, 21 ° 26 ’ S, 117 ° 9 ’ E, 337 m, sandy creek, Eucalyptus Spinifex grassland, 7 – 12. V. 2003, C. Lambkin, D. Yeates, Malaise trap, JSS 15573 (1 ♂, ANIC); Karijini National Park, Juna Downs Road, 22 ° 44 ’ S, 118 ° 25 ’ E, 798 m, dry Turee Creek, grassy open Eucalyptus scrub, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, 2145, JSS 19186 (1 ♀, ANIC); Karijini National Park, Weano Gorge Road, 22 ° 21 ’ S, 118 ° 15 ’ E; 22 ° 23 ’ S, 118 ° 15 ’ E, 695 m; 711 m, grassy dry creek, Eucalyptus Acacia scrub; beside drying pool bank in grassland, 25. IV – 15. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 15981, JSS 16080, JSS 16082 (1 ♂, 2 ♀, ANIC); Mount Augustus National Park; south slope on Saddle Trail, 24 ° 18 ’ S, 116 ° 48 ’ E, 400 m, small dry wooded wash, 25. IV – 7. V. 2003, M. E. Irwin, F. D. Parker, Malaise trap, JSS 15884 (1 ♀, CNC); Mount Augustus National Park, 24 ° 22 ’ S, 116 ° 50 ’ E, 427 m, wash on southern slope of mountain, 25. IV – 7. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15877 (1 ♀, CNC); New South Wales: 23 km SE Tamworth, 31 ° 13 ’ S, 151 ° 7 ’ E, 26. XI. 1982, D. S. Horning, Jr., JSS 9151 (1 ♂, ANIC); 25 km NW Coonamble, 30 ° 49 ’ S, 148 ° 12 ’ E, 17. VI. 1976, Z. Liepa, JSS 9169 (1 ♂, ANIC); 27 km N Dubbo, 32 ° 1 ’ S, 148 ° 37 ’ E, 17. VI. 1976, Z. Liepa, JSS 9205 (1 ♂, ANIC); Ledknapper National Reserve, 25 km NE Beulah Homestead, near old Gerara Springs (LDN 6 M), 29 ° 16 ’ S, 146 ° 23 ’ E, Mulga, 10. XII. 2009 – 18. III. 2010, S. O’Sullivan, K. Taylor, Malaise trap, CNC 576646, CNC 576649 (2 ♂, CNC); Culgoa National Park, 13.5 km SW Cawwell Homestead (CGN 1 M), 29 ° 7 ’ S, 146 ° 56 ’ E, Callitris, 30. I – 18. V. 2010, QM Team, B. Shiebaan & R. Ohlsen, Malaise trap, CNC 576620 (1 ♂, CNC); Northern Territory: McArthur River, 14 km S by W of Cape Crawford, 15 ° 54 ’ S, 136 ° 40 ’ E, 12. IV. 1976, D. H. Colless, JSS 9220 (1 ♂, ANIC); 25 miles SW of Auvergne homestead, 15 ° 56 ’ S, 129 ° 44 ’ E, 28. V. 1968, M. Mendum, JSS 8554 (1 ♂, ANIC); Casurina Beach, Darwin, 12 ° 27 ’ S, 130 ° 50 ’ E, 22. X. 1972, D. H. Colless, JSS 8527 (1 ♂, ANIC); Koongarra, 15 km E of Mount Cahill, 12 ° 52 ’ S, 132 ° 51 ’ E, 6 – 9. III. 1973, D. H. Colless, JSS 8547 (1 ♂, ANIC); Roper River, 14 ° 44 ’ S, 135 ° 23 ’ E ,, N. B. Tindale, JSS 8720 – 1 (2 ♂, SAM). NEW CALEDONIA: Anse Vata, 22 ° 18 ’ S, 166 ° 27 ’ E, 23. X. 1958, C. R. Joyce, CNCD 5613 (1 ♂, BPBM); Chesterfield Islands, 19 ° 52 ’ S, 158 ° 19 ’ E, IX. 1957, F. Cohic, CNCD 5600 (1 ♂, BPBM); Noumea, 22 ° 16 ’ S, 166 ° 26 ’ E, 200 m, III. 1973, N. L. H. Krauss, CNCD 5609 (1 ♂, BPBM); 22 – 23. XI. 1963, R. Straatman, CNCD 5606, CNCD 5608 (2 ♂, BPBM); PHILIPPINES: Mountain Province: Abatan, Buguias, 60 km S of Bontoc, 16 ° 43 ’ N, 120 ° 49 ’ E, 2000 m, 11. V. 1964; 17 – 19. V. 1964; 21 – 27. V. 1964; 28. IV. 1964; 30. V. 1964, H. M. Torrevillas, light trap; CNCD 5544 – 9 (5 ♂, 1 ♀, BPBM); SOLOMON ISLANDS: Guadalcanal: Honiara, 9 ° 27 ’ S, 159 ° 58 ’ E, 0 to 100 m, X. 1970, N. L. H. Krauss, CNCD 5627 – 30 (4 ♂, BPBM); Malaita: Antong Java Atoll, Peku, 10 m, 17. XII. 1972, N. L. H. Krauss, CNCD 5620 (1 ♂, BPBM); NDAI Island, Bethlehem, 7 ° 53 ’ S, 160 ° 37 ’ E, 10 m, XII. 1972, N. L. H. Krauss, CNCD 5619 (1 ♂, BPBM); Gizo Island, Gizo, 8 ° 6 ’ S, 156 ° 50 ’ E, 0 to 200 m, XII. 1975, N. L. H. Krauss, CNCD 5624 – 5 (2 ♂, BPBM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF1CECBFD8D9EFF6FAA013BC.taxon	distribution	Distribution: Australia (New South Wales, Northern Territory, Queensland, Western Australia), New Caledonia, Philippines (Mountain Province), Solomon Islands (Guadalcanal, Malaita) (Fig. 142). Notes: This species is widespread in Australia north of Dubbo in the east and Perth in the west. It extends through the Solomons, New Caledonia and New Guinea to the northern Philippines. It is predominantly a forest species and has never been found hilltopping. Perkins (1905) notes that one specimen was reared from a ‘ jassid’. This is a general name for a leafhopper. Specimen CNCD 5542 is hereby nominated as the lectotype to fix and ensure consistent interpretation of the name. Intraspecific genetic distance ranges from 0.4 % to 2.4 %. Tomosvaryella synadelpha is genetically closest to T. cona sp. nov. (8.9 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF03ECBDD8D9EB7BFC4311FC.taxon	description	Figs 95 A – E, 141, 145 B	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF03ECBDD8D9EB7BFC4311FC.taxon	diagnosis	Diagnosis: This species can be recognized by a ventroapical keel on the hind trochanter (Fig. 145 B); elongated surstyli, left surstylus club-shaped, right one with a triangular lobes ventroapically in dorsal view (Fig. 95 A); gonopods elongated, left little longer and wider than right one with a distinct projection on inner margin (Fig. 95 B); phallic guide with 3 – 4 short dorsolateral spines in ventral view.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF03ECBDD8D9EB7BFC4311FC.taxon	description	Description: MALE: Body length: 2.5 mm. Head. Flagellum acuminate; yellow-brown, scape and pedicel dark brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 3 – 4 dark bristles. Mesonotum (viewed obliquely from front) silvery brown pollinose. Dorsocentral bristles short, numerous dark bristles around postpronotum present. Scutellum silvery brown pollinose and with 4 – 6 scattered bristles. Halter, stem and knob pale, tip brownish, base brown. Legs. Mid coxa with 3 – 4 long dark bristles. Trochanters brown, femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments bright yellow completely. Hind trochanter with a keel ventroapically; 1 – 2 ventrobasal short bristles present on fore mid femur. Hind femur without ventroapical spines, except 8 – 10 equally spaced short bristles posteroventrally, longer on distal half, longest up to 1 / 3 the width of hind tibia at distal end. Short subapical spines on fore- and mid tibiae present. Hind tarsomeres distinctly flattened, hind metatarsus almost as long as 2 – 4 combined, all tarsomeres with erect bristles dorsally. Pulvilli shorter than last tarsal segment. Wing. Length: 2.5 mm. Upper side of basal costal cell with one short dark brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2 – 3 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites subshiny brown, tergite 1 silvery brown grey pollinose. Dispersed short dark bristles on all tergites present. Lateral bristles on first tergite present, 6 – 8 dark bristles up to as long as hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.5). Surstyli elongate, left surstylus broadened on apical half, apex club-shaped; right one with a triangular lobes ventroapically, left surstylus is longer than right one (Fig. 95 A). Genital capsule in ventral view: gonopods unequal, left is little longer and wider than right one, elongated towards surstyli, both gonopods with a distinct projection on inner margin; phallic guide with 3 – 4 short dorso- dorsolateral spines; subepandrial sclerite long (Fig. 95 B); Genital capsule in lateral view: both surstyli curved towards sternite, rather widened at basal half, narrowed at apical half (Fig. 95 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 95 C). FEMALE: Unknown. Etymology: From the Latin tescum, a desert, in reference to the desert habitat of this species. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Culgoa Floodplain National Park, 7 km Northwest of Toulby Gate (TOLB 7 M), 28 ° 57 ’ S, 146 ° 53 ’ E, 138 m, reedy swamp, 20. I – 19. III. 2010, C. Kelly & A. Coward, Malaise trap, CNC 575371 (1 ♂, QM); PARATYPE: AUSTRALIA: Western Australia: 171 km E Marble Bar, 21 ° 17 ’ S, 121 ° 14 ’ E, 2 – 15. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, CNCD 11620 (1 ♂, CNC). Distribution: Australia (Queensland, Western Australia) (Fig. 141).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF01ECBBD8D9EEBBFD541110.taxon	description	Figs 96 A – E, 139, 146 F	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF01ECBBD8D9EEBBFD541110.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a keel with tiny dark spines ventrobasally (Fig. 146 F); some long posterior bristles in apical half of hind femur; elongate surstyli in dorsal view (Fig. 96 A); gonopods with four projections in ventral view (Fig. 96 B); hypandrium with two small projections towards gonopod (Fig. 96 B); phallic guide with six dorsolateral spines (Fig. 96 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF01ECBBD8D9EEBBFD541110.taxon	description	Description: MALE. Body length: 2.6 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose; eyes touching for distance equal to 1 – 1.3 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 6 – 7 distinct pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part silver pollinose, slightly grayish also from the side. Scutellum silver pollinose and with 6 – 8 weak bristles along the distal edge. Dorsocentral bristles well developed, dark, longer on frontal part and numerous dark bristles around postpronotum. Halter, knob pale, stem brown. Legs. Mid coxa with two spines. Trochanters and femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 5 of tibiae yellow (tibiae otherwise brown), tarsal segments yellow, last segment darker dorsally. Hind trochanter with a keel provided with tiny dark spines (3 – 8) in a group ventrobasally (Fig. 146 F). Ventroapical row of spines on fore femur absent, 4 – 6 very short, black spines on mid femur; no spines on hind femur, hind femur with 14 – 16 pale, long bristles equally spaced posteriorly, longer on distal half (longest up to as long as 1.2 – 1.4 times the width of hind tibia at the distal end). Subapical (distal) spines on fore and mid tibiae present (very short). Hind metatarsus slightly flattened in middle, extended along the posterior edge, with dorsal bristles, missing in the middle, arranged in rows along the edges; hind metatarsus and second tarsomere with scrub-like? bristles ventrally. Pulvilli as long as last tarsal segment. Wing. Length: 3.0 mm. Upper side of basal costal cell with one dark and long bristles. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m at middle of discal cell. 8 – 9 dark setulae on tegula. Abdomen. Viewed obliquely from front tergites subshining black, tergite 1 and anterior half of tergite 2 silvery grey, sides silvery pollinose on tergites 4 and 5 (the latter with only slightly larger spots). Dispersed long dark bristles on tergites present, the longest on tergite 5 and up to 0.5 – 0.8 times as long as width of hind tibia at base. Lateral bristles on first tergite present, 6 – 8 brown bristles in a row up to as long as the width of hind femur at distal tip in lateral view. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1 – 1.2 times as long as ST 8. Genitalia without dissection: ST 8 medium sized, brown-black, with occasional small bristles; membranous area elongated, widening on upper part, which is directed dorsally; epandrium brown-black; surstyli yellow, appear to be elongated and slim. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.6). Surstyli elongate, slender and symmetrical. Left surstylus slightly longer than right surstylus. Both surstyli with broad base and tips curving inward (Fig. 96 A). Genital capsule in ventral view: subepandrial sclerite wider in middle, gonopods rounded at the apex with two long finger-like projections towards surstyli and two projections extended towards phallus (Fig. 96 B); lobes of hypandrium sclerotized and broadened in middle with two small projections towards gonopod (Fig. 96 B). Phallus strong and straight, with three short ejaculatory ducts; phallic guide with six spines dorsolaterally (three left spines longer than others) (Fig. 96 B). Genital capsule in lateral view: left surstylus straight in basal half and curved towards sternite in apical half; right surstylus slightly curved towards sternite in apical half (Fig. 96 D – E). Ejaculatory apodeme tube-like with a bulb in its middle (Fig. 96 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF01ECBBD8D9EEBBFD541110.taxon	etymology	Etymology: The name is derived from the Greek tetra, four and the Latin lobus, an elongated projection or protuberance, in reference to the four projections on the male gonopods. Examined material: HOLOTYPE: AUSTRALIA: New South Wales: Yathong Nature Res [erve] n [ea] r Triangle Tank, 32 ° 38 ’ S, 145 ° 32 ’ E, 200 m, in sandy wash in low closed mallee, 3 – 25. x. 2003, C. Lambkin & N. Starick, Malaise trap, JSS 16037 (1 ♂, ANIC); PARATYPE: AUSTRALIA: same data as holotype, JSS 16036 (1 ♂, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF01ECBBD8D9EEBBFD541110.taxon	distribution	Distribution: Australia (New South Wales) (Fig. 139). Notes: The shape of the surstyli is similar to T. biflocca sp. nov., elongated and slender in dorsal view. It differs by the hind trochanter having a keel ventrobasally; gonopods rounded at the apex and the subepandrial sclerite without a vertical protrusion in middle in ventral view (Fig. 96 B), without membranous sheath (Fig. 96 B). Intraspecific genetic distance is 0.3 %. Tomosvaryella tetraloba sp. nov. is genetically closest to T. quadrata sp. nov. (4.4 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF07ECB9D8D9EED7FAC11214.taxon	description	Figs 97 A – E, 143, 145 D, 152 E, 157 D	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF07ECB9D8D9EED7FAC11214.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a keel and a patch of short bristles (Fig. 145 D); sternite 2 – 4 with two distinct dark spines in the middle of posterior margin (Fig. 152 E); small surstyli in dorsal view (Fig. 97 A); widened gonopods (Fig. 97 B); hypandrium with two hairy finger-like membranous sheaths on posterior margin; phallic guide with 3 – 6 different-sized spines dorso- dorsolaterally (Fig. 97 D); subepandrial sclerite and hypandrial apodeme wide and distinct in ventral view (Fig. 97 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF07ECB9D8D9EED7FAC11214.taxon	description	Description: MALE: Body length: 3.7 – 3.8 mm. Thorax. Postpronotum light yellow, with 5 – 7 brown bristles in upper margin. Mesonotum (viewed obliquely from front) silvery-brown pollinose, anterior part greyer; scutum with scattered bristles at anterior supra-alar area. Postalar callus dark brown with three dark bristles in the middle. Scutellum silvery pollinose and with a few indistinct pale bristles along the distal edge. Halter, knob pale, stem light brown, base dark. Legs. Mid coxa with 2 – 5 dark bristles, 1 – 2 bristles longer than others. Coxae, trochanters, femora dark, hind femur shining ventrally, knees and basal 1 / 4 of tibiae yellow (tibiae otherwise dark brown), tarsomeres light brown ventrally and dark dorsally. Mid trochanter with 1 – 2 short bristles ventroapically. Hind trochanter silvery pollinose, with a keel and batch (3 – 6) of short bristles ventroapically (Fig. 145 D); ventrobasal spines absent on fore and mid femur. Hind femur with two rows of ventroapical spines. Hind tibia with two rows of short black bristles on both anterior and posterior side. Hind tarsomeres distinctly flattened, covered by dark bristles dorsally and brown bristles ventrally, last hind tarsomere not rounded as long as 3 – 4 combined; pulvilli shorter than last tarsal segment. Wing. Length: 3.2 mm. Upper side of basal costal cell with one dark brown bristle. Fourth costal section 1.5 – 2 times as long as third costal section. Cross-vein r-m at middle of discal cell. 4 – 6 short dark setulae on tegula. Abdomen. Tergite 1 silvery pollinose with 6 – 8 dark bristles; tergites 2 – 5 covered by scattered brown bristles. Sternite 2 – 3 with two small dark spines in the middle of posterior margin; sternite 4 with two triangle-shaped spines along posterior margin (Fig. 152 E). Syntergosternite 8 enlarged, dark brown and grey-brown pollinose. Membranous area long and broad in middle. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.8). Surstyli small, rather triangle-shaped, slightly narrowed at apex; left surstylus somewhat longer, slightly bent towards the right side at apex (Fig. 97 A). Genital capsule in ventral view: both gonopods widened, equal in height; hypandrium widened with two hairy finger-like membranous sheaths in upper margin; phallic guide with 3 – 6 different-sized spines dorsolaterally, one is longer than others; subepandrial sclerite wide and distinct; hypandrial apodeme wide and distinct, located between hypandrium (Fig. 97 B); Genital capsule in lateral view: both surstyli widened at base, triangle-shaped with a small downward projection at the base (Fig. 97 D – E); ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 97 C). FEMALE: Body length: 3.0 – 3.1 mm. As male except for the following characters. Frons, eyes separated, as wide as the width at antennae; completely silver-grey pollinose on lower 1 / 3, shiny black only around ocellar triangle, smooth depression just before ocellar triangle. Enlarged ommatidia silvery shining; pulvilli and claws about 1.2 – 1.5 times as long as last tarsal segment. Tergites 4 – 6 with two grey lateral spots. Abdominal sternites without spines. Ovipositor. Straight in ventral view, rather straight, in lateral view (yellow piercer, dark brown base), reaching sternite 1; base silvery brown pollinose. LP: LB = 2.5. LDP: LPP = 2.0. (Fig. 157 D).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF07ECB9D8D9EED7FAC11214.taxon	etymology	Etymology: The specific name is derived from the Latin word ‘ tribulus’, meaning thorny, referring to spines on the abdominal sternites. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: 82 km S j [un] ct [ion] Karijini Dr [ive] on Great Northern H [igh] w [a] y, 23 ° 7 ’ S, 119 ° 6 ’ E, 694 m, wash with pools, 23. IV – 6. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15675 (1 ♂, WAM); PARATYPES: AUSTRALIA: Western Australia: 67 km SW Pardoo Road House on Shay Gap Road, 20 ° 28 ’ S, 120 ° 10 ’ E, 177 m, dry sandy ravine with flowers, 1 – 14. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15667 (1 ♂, CNC); 82 km S junction Karijini Drive on Great Northern Highway, 23 ° 7 ’ S, 119 ° 6 ’ E, 694 m, wash with pools, 23. IV – 6. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15672 (1 ♂, QM); Karijini National Park, Hamersley-Mount Bruce Road, 22 ° 34 ’ S, 118 ° 18 ’ E, 757 m, dry rocky creek bed, Eucalyptus grassland, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16125 (1 ♀, ANIC); Karijini National Park, Karijini Drive, 22 ° 34 ’ S, 118 ° 18 ’ E, 815 m; 814 m, across dry gully, Acacia scrub; open Eucalyptus grassland, 19 – 25. IV. 2003; 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 15579 – 80, JSS 15586, JSS 16158 (2 ♂, 2 ♀, ANIC); Karijini National Park, Weano Gorge Road, 22 ° 24 ’ S, 118 ° 15 ’ E, 718 m, in open Eucalyptus, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16220, JSS 16236 (2 ♀, ANIC); Millstream-Chichester National Park, Roebourne Road, 21 ° 26 ’ S, 117 ° 9 ’ E, 337 m, sandy creek, Eucalyptus Spinifex grassland, 27. IV – 3. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16076 – 8 (3 ♀, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF07ECB9D8D9EED7FAC11214.taxon	distribution	Distribution: Australia (Western Australia) (Fig. 143). Notes: This apparent West Australian endemic is restricted to the west central region of the state. Intraspecific genetic distance ranges from 0.3 % to 2.4 %. It is genetically most similar to T. millstreamensis sp. nov. (0.3 – 1.0 % pairwise divergence) (Supplementary file 3). These two species show very low genetic distance, so it is possible that these are a single species with polymorphic external characters. More likely, they are young species and the mtDNA has simply not yet diverged. Future genetic and ecological work are required to test their taxonomic status.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF05ECB7D8D9EDD3FD5F1398.taxon	description	Figs 98 A – E, 141, 146 B	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF05ECB7D8D9EDD3FD5F1398.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having some dense short bristles ventrally (Fig. 146 B); elongated surstyli in dorsal view (Fig. 98 A); distinct hypandrial apodeme in lateral view (Fig. 98 D – E); three spines on phallic guide and two short spines on one of ejaculatory ducts in lateral view.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF05ECB7D8D9EDD3FD5F1398.taxon	description	Description: MALE: Body length: 2.6 – 3.0 mm. Head. Flagellum acuminate, brown. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose; eyes touching for distance equal to 2 – 2.5 times the length of ocellar triangle. Occiput silvery pollinose, more brownish on upper half. Thorax. Postpronotum pale, with 4 – 5 distinct pale bristles. Mesonotum (viewed obliquely from front) subshining black, anterior part silver pollinose, slightly grayish from the side. Scutellum silvery pollinose with 12 distinct brown bristles along the distal edge. Dorsocentral bristles developed, dark, longer on frontal part and numerous dark bristles around postpronotum. Halter; knob pale with a brownish tinge distally, stem dark brown. Legs. Mid coxa with 2 – 3 bristles. Trochanters and base of femora black, femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees, basal 1 / 5 of tibiae yellow (otherwise brown), tarsal segments yellow ventrally, yellow-brown dorsally, last segment dorsally darker. Hind trochanter gray pollinose, without keel, some short dark spines (7 – 9) at the base, otherwise covered with dense short bristles ventrally (Fig. 146 B). Ventroapical row of spines on fore femur missing, one ventrobasal spine present on fore and absent on mid femur; 4 – 6 very short, black spines on mid femur; no spines on hind femur, 20 – 22 pale bristles posteriorly, equally distributed, and longer on distal 1 / 3 (as long as 2 / 3 the width of hind tibia at the distal end). Subapical spines on first four tibiae present, distal and medium-sized. Hind metatarsus not flattened, first tarsomere slightly longer than 2 – 4 combined, with short and dense bristles ventrally, yellow, dorsal bristles on all tarsi semi-erect and dark. Pulvilli as long as last tarsal segment. Wing. Length: 2.7 – 2.8 mm. Upper side of basal costal cell with one long distinct bristle. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m slightly proximal to middle of discal cell. 6 – 8 dark setulae on tegula. Abdomen. Viewed obliquely from front tergites subshining black, tergite 1 silvery grey, sides silvery pollinose on tergites 4 and 5 (the latter with larger spots). Dispersed short dark bristles on tergites present, the longest on tergite 5 and up to half as long as width of hind tibia at base. Lateral bristles on first tergite present, 3 – 4 brown bristles up to as long as width of hind femur. Postabdomen in dorsal view; T 6, S 7 invisible; T 5 1.5 times as long as ST 8. Genitalia without dissection: ST 8 medium sized, subshiny black as tergite 5; membranous area roundish, cone-shaped; epandrium paler, narrow; yellow surstyli long and flat. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.07). Surstyli light brown, rather symmetrical, elongated, left surstylus longer than right one, its tip curved toward right one in apical third (Fig. 98 A). Genital capsule in ventral view: subepandrial sclerite wide, extended beneath gonopods, covered with transverse streaks; gonopods elongated, with a small upward projection (Fig. 98 B), phallic guide with 3 – 4 different-sized spines dorsally (longer one closer to apex). Genital capsule in lateral view: left surstylus smoothly curved towards sternite, broader than right one; right surstylus smoothly curved towards sternite (Fig. 98 D – E); phallus with three ejaculatory ducts, one with two small spines; hypandrial apodeme distinct and long (Fig. 98 D – E); ejaculatory apodeme flat, sperm pump elongated, linear (Fig. 98 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF05ECB7D8D9EDD3FD5F1398.taxon	etymology	Etymology: From the Latin tres, three and spina, spine, in reference to the three spines found on the phallic guide. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Brisbane, Mt. Coot-tha, 27 ° 29 ’ S, 152 ° 57 ’ E, 170 m, hilltop, 7. xii. 1997, J. & A. Skevington, hand collected, JSS 2352 (1 ♂, QM); PARATYPES: AUSTRALIA: Queensland: 14.6 km east of Lakeland Downs, 15 ° 46 ’ S, 144 ° 57 ’ E, 270 m, hilltop in open, dry savanna (rangeland for cattle), 8. vi. 1997, J. & A. Skevington, JSS 8853 – 4 (2 ♂, QM); Barakula State Forest No. 302, Summit Round Mountain, 26 ° 22 ’ S, 150 ° 58 ’ E, 8. i. 2000, J. & A. Skevington & M. Mathieson, JSS 7556 (1 ♂, QM); Brisbane, Mount Coot-tha, 27 ° 29 ’ S, 152 ° 57 ’ E, 170 m, hilltop, 16. ix. 1997, J. Skevington, hand collected, JSS 3800 (1 ♂, CNC); ~ 170 m, hilltop, 25. ix. 1997, J. & A. Skevington, hand collected, JSS 3793 – 4 (2 ♂, QM); 26. x. 1997, J. Skevington, hand collected, JSS 3795, JSS 3797 (2 ♂, QM); 27. i. 1998, J. & A. Skevington, hand collected, JSS 3083 (1 ♂, QM); Brisbane, Mt. Coot-tha, 27 ° 29 ’ S, 152 ° 57 ’ E, ~ 170 m, hilltop, 7. xii. 1997, J. & A. Skevington, hand collected, JSS 2333, JSS 2335 – 6, JSS 2339 – 40, JSS 2351 (1 ♂, ANIC, 5 ♂, QM); 8. xi. 1997, J. & A. Skevington, hand collected, 9: 45 am – 10: 15 am, 10: 15 am – 10: 45 am, 9: 15 am – 9: 45 am, JSS 978, JSS 987, JSS 990, JSS 997 – 8 (5 ♂, QM); 250 m, hilltop, 21. VIII. 2004, J. Skevington, hand collected, JSM 780, JSM 809, JSS 15358, JSS 15386 (2 ♂, CNC); Carnarvon National Park, Mount Moffatt Summit, 25 ° 4 ’ S, 148 ° 3 ’ E, 1097 m, hilltop, 29. xi. 1997, J. Skevington & C. Lambkin, hand collected, JSS 1608 (1 ♂, QM); Scrub Road, Brisbane Forest Park, 27 ° 25 ’ S, 152 ° 50 ’ E, 3 – 10. x. 1997, S. Winterton, N. Power & D. White, Malaise trap, JSS 8855 (1 ♂, QM); New South Wales: Honeysuckle Creek, 35 ° 35 ’ S, 149 ° 0 ’ E, 11 – 22. IV. 1985, I. Naumann & J. Cardale, Malaise trap, JSS 9237 (1 ♂, ANIC); Western Australia: Karijini National Park, Weano Gorge Road, 22 ° 24 ’ S, 118 ° 15 ’ E, ANIC Bulk Sample 2042, 718 m, in open Eucalyptus, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, ANIC Bulk Sample 2042, JSS 16229 (1 ♂, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF05ECB7D8D9EDD3FD5F1398.taxon	distribution	Distribution: Australia (New South Wales, Queensland, Western Australia). Notes: This species can be differentiated from T. atraspina sp. nov. by having bristles (not spines) on the hind trochanter; having one ventrobasal bristle on the fore femur (two bristles on T. atraspina sp. nov.); and a different shape of the left surstylus – in T. atraspina sp. nov. the left surstylus is bent towards the sternite in the apical third (Fig. 9 C). All but three of the known specimens of this species have been collected on hilltops. Intraspecific genetic distance ranges from 0.0 % to 4.2 %. Tomosvaryella trispina sp. nov. is genetically closest to T. quadrata sp. nov. (6.5 % pairwise divergence) (Supplementary file 3). The large intraspecific variation suggests that this may be a species complex and more investigation is required.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF0BECB5D8D9EB5FFC401214.taxon	description	Figs 99 A – E, 136	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF0BECB5D8D9EB5FFC401214.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a large, fin-shaped keel ventroapically and a small tuber-shaped protrusion ventrobasally involving two small spines at its apex; hind tibia broadened distinctly in apical half; long surstyli, widened in apical fourth in dorsal view (Fig. 99 A); gonopods unequal, hypandrium long in ventral view (Fig. 99 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF0BECB5D8D9EB5FFC401214.taxon	description	Description: MALE: Body length: 2.5 – 2.6 mm. Head. Scape, pedicel, flagellum and arista dark, flagellum tapering, mostly gray pollinose. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.3 times the length of ocellar triangle. Occiput silvery-brown pollinose. Thorax. Postpronotum pale, with 4 long pale bristles. Mesonotum (viewed obliquely from front) silvery-brown pollinose with some long supra-alar bristles behind postpronotum lobes. Scutellum silvery-brown pollinose with 8 – 12 short dark bristles along distal edge. Halter, knob yellow, stem and base brown. Legs. Mid coxa with 3 – 4 long dark bristles. Trochanters brown, femora black, silvery pollinose posteriorly except shiny hind femur; tibiae brown except knees and basal 1 / 6 of tibiae yellow, tarsal segments light brown. Hind trochanter with large fin-shaped keel ventroapically and a small protrusion ventrobasally involving two small spines at its apex. Two ventrobasal spines present on fore- and 3 – 5 shorter ones on mid femur. Hind femur with a small triangle-shaped keel ventrobasally, without ventroapical spines except 8 – 10 equally spaced bristles in two rows posteroventrally. Subapical (distal) spines on first four tibiae present (short). Hind tibia broadened distinctly in apical half covered by yellow erected bristles ventrobasally. Hind metatarsus slightly flattened, almost as long as 2 – 4 combined; all tarsomeres with erect bristles dorsally. Pulvilli covered by short yellow bristles. Wing. Length: 2.7 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 1 – 2 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brown-black, tergite 1 silvery grey. Dispersed short brown bristles on all tergites present. Lateral bristles on first tergite present, 8 – 10 dark bristles up to as long as ¾ of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium obviously longer than wide (MLE: MWE = 1.5). Membranous area rounded. Surstyli elongated, as long as epandrium width, both surstyli slightly broadened in apical fourth and curved to each other at apex (Fig. 99 A). Genital capsule in ventral view: gonopods unequal in height, extended towards surstyli, hypandrium long; phallic guide long; subepandrial sclerite distinct (Fig. 99 B); Genital capsule in lateral view: both surstyli straight in basal three fourths, bent 90 ° towards sternite (Fig. 99 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 99 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF0BECB5D8D9EB5FFC401214.taxon	etymology	Etymology: The name tuber is Latin for a swelling or bulb and refers to the bulbous protrusion on the hind trochanter. Examined material: HOLOTYPE: AUSTRALIA: Queensland: Tinaroo Falls Dam, North Queensland, 17 ° 11 ’ S, 145 ° 34 ’ E, open savannah, 27. IV. 1967, D. H. Colless, JSS 8528 (1 ♂, ANIC); PARATYPES: AUSTRALIA: New South Wales: Shoalhaven Bdg. near Braidwood, 35 ° 26 ’ S, 149 ° 48 ’ E, 9. I. 1968, D. H. Colless, JSS 8526 (1 ♂, ANIC); Queensland: Southport, 27 ° 59 ’ S, 153 ° 24 ’ E, 29. V. 1968, J. W. Boyes, JSS 9702 (1 ♂, CNC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF0BECB5D8D9EB5FFC401214.taxon	distribution	Distribution: Australia (New South Wales, Queensland) (Fig. 136).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF09ECB3D8D9EDD3FD4F11D8.taxon	description	Figs 100 A – E, 137, 156 B	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF09ECB3D8D9EDD3FD4F11D8.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a small keel covered by some short bristles; slender surstyli in dorsal view (Fig. 100 A); elongated gonopods, right gonopod with a distinct projection on inner margin; phallic guide with 5 – 6 dorso- dorsolateral spines; subepandrial sclerite with a central, small, fin-shaped ridge in ventral view (Fig. 100 B); phallic guide with a hook-like projection ventrally in lateral view (Fig. 100 D).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF09ECB3D8D9EDD3FD4F11D8.taxon	description	Description: MALE: Body length: 2.6 – 2.8 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for 2 x length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 2 – 3 distinct pale bristles. Mesonotum (viewed obliquely from front) silvery pollinose, anterior part more grey, greyish also from the side. Scutellum silvery brown pollinose and without bristles along distal edge. Dorsocentral bristles long, dark, mostly in frontal half, as long as ¾ width of hind femur at base, pale bristles around postpronotum absent. Halter, knob pale, stem brown. Legs. Mid coxa with 2 – 3 long dark bristles. Trochanters brown, femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 4 of tibia yellow (tibia otherwise brown), tarsal segments yellow-brown dorsally, yellow ventrally, last segment darker. Hind trochanter with small keel, and 6 – 7 short dark bristles, equally spaced; ventrobasal spines absent on fore and mid femur. Hind femur without ventroapical spines, without long bristles posteroventrally. Subapical (distal) spines on first four tibiae well developed (longer than usual). Hind tarsomeres slightly flattened (especially hind metatarsus on distal end), and as long as 2 – 5 combined; posterior corners with distinct bristles on all hind tarsomeres. Pulvilli shorter than last tarsal segment. Wing. Length: 2.6 – 2.7 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2 – 3 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites subshiny brown-black, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5, larger on tergite 5. Dispersed strong dark bristles on all tergites present, longest on tergite 5; length up to 1 / 2 the width of hind tibia at distal end. Lateral bristles on first tergite present, 4 – 6 dark bristles up to as long as hind femur’s width at base. Postabdomen in dorsal view; T 6, S 7 invisible, T 5 0.6 – 0.8 times as long as ST 8. Genitalia without dissection: ST 8 large, rounded in dorsal view, brown and with distinct bristles, otherwise with velvet-like coverage; membranous area is rounded dorsally, narrow ventrally, mostly directed posteriorly; epandrium brown, surstyli yellow. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.6). Surstyli symmetrical, slender and elongated, curved to each other at apex, left surstylus longer than right one (Fig. 100 A). Genital capsule in ventral view: gonopods equal in height, extended towards surstyli, pointed at apex, right gonopod with a distinct projection on inner margin; phallic guide with 5 – 6 dorso- dorsolateral spines, three are longer than others; subepandrial sclerite large, with a small, central, fin-shaped ridge, hypandrial apodeme distinct (Fig. 100 B); Genital capsule in lateral view: both surstyli straight in basal three fourths and bent towards sternite apically, phallic guide with a hook-like projection ventrally before apex (Fig. 100 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 100 C). FEMALE: Body length: 2.2 mm. As male except for the following characters. Frons, eyes separated, as wide as 2 times the width at antennae; completely silver-grey pollinose on lower 1 / 3, shiny black only around ocellar triangle, smooth change in transition zone (just before ocellar triangle) to shiny black. Enlarged ommatidia silvery shining. Pulvilli and claws about 1.5 – 2.5 times as long as last tarsal segment, smaller on metatarsi. Female abdomen with silver spots on sides of tergites 4 – 6. Ovipositor. Straight in ventral view; straight, dagger-like in lateral view (yellow piercer, brown base), reaching base of abdomen; base silvery brown pollinose with numerous short dark bristles. LP: LB = 3.0. LDP: LPP = 4.0. (Fig. 156 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF09ECB3D8D9EDD3FD4F11D8.taxon	etymology	Etymology: From the Latin uncus, hook, barb or angle, in reference to the hook-like phallic guide projection. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: Karijini National Park: Juna Downs Road (between drying pools in rocky Turee Creek), 22 ° 42 ’ S, 118 ° 25 ’ E, 789 m, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, CNCD 3863 (1 ♂, WAM); PARATYPES: AUSTRALIA: New South Wales: 12 miles NW Milton, 35 ° 18 ’ S, 150 ° 26 ’ E, 488 m, 5. XI. 1968, D. H. Colless, JSS 8671 (1 ♂, ANIC); Culgoa National Park, 8 km West-Northwest Cawwell Homestead, Diemunga Lagoon (CGN 2 M), 29 ° 3 ’ S, 146 ° 60 ’ E, Coolibah, 30. I – 18. V. 2010, C. Lambkin, R. Olsen, B. Shieban, Malaise trap, CNC 575140 (1 ♂, QM); Porters Dam Road 16 km NW of Milton, 35 ° 19 ’ S, 150 ° 26 ’ E, 9. II. 1974, Z. Liepa, JSS 9157 (1 ♂, ANIC); Rotary Lookout 23 km NW of Milton, 35 ° 19 ’ S, 150 ° 26 ’ E, 9. II. 1974, Z. Liepa, JSS 9162 (1 ♂, ANIC); Sawpit Creek, Kosciusko National Park, 36 ° 30 ’ S, 148 ° 16 ’ E, 11. III. 1978, Z. Liepa, JSS 9171 (1 ♂, ANIC); Ledknapper National Reserve, 25.1 km Northeast of Beulah Homestead, near old Gerara Springs (LDN 6 M), 29 ° 16 ’ S, 146 ° 23 ’ E, Mulga, 10. XII. 2009 – 18. III. 2010, S. O’Sullivan & K. Taylor, Malaise trap, CNC 574628 (1 ♂, QM); Queensland: Culgoa Floodplain National Park, 7 km NNW Toulby Gate (CG 4 AM), 28 ° 56 ’ S, 146 ° 55 ’ E, 160 m, Gidgee, 20. I – 19. III. 2010, C. Kelly & A. Coward, Malaise trap, CNC 576381, CNC 576391 (2 ♂, QM); 17. XII. 2009 – 20. I. 2010, Kelly & Coward, Malaise trap, CNC 591829 – 30, CNC 591832, CNC 591835 (4 ♂, QM); Culgoa Floodplain National Park, 7 km North-Northwest of Toulby Gate (CG 4 AM), 28 ° 56 ’ S, 146 ° 55 ’ E, 160 m, Gidgee, 19. III – 20. V. 2010, C. Kelly & A. Coward, Malaise trap, CNC 575358 (1 ♂, QM); Culgoa Floodplain National Park, 7 km Northwest of Toulby Gate (TOLB 7 M), 28 ° 57 ’ S, 146 ° 53 ’ E, 138 m, reedy swamp, 20. I – 19. III. 2010, C. Kelly & A. Coward, Malaise trap, CNC 575364 (1 ♂, QM); Culgoa Floodplain National Park, 7 km WNW Toulby Gate (TOLB 6 M), 28 ° 57 ’ S, 146 ° 52 ’ E, 138 m, Callitris, 19. III – 20. V. 2010, C. Kelly & A. Coward, Malaise trap, CNC 575019 (1 ♂, QM); Australian Capital Territory: Nass Road, S of Tharwa, 35 ° 31 ’ S, 149 ° 4 ’ E, 28. XI. 1965, Z. Liepa, JSS 9219 (1 ♂, ANIC); Queensland: 14 km W by N Hope Vale Mission, 15 ° 16 ’ S, 144 ° 59 ’ E, 10. X. 1980, D. H. Colless, Malaise trap, JSS 8564 (1 ♂, ANIC); 2 km S of Horseshoe lookout, Blackdown land, 23 ° 50 ’ S, 149 ° 3 ’ E, 23 – 24. IV. 1981, D. H. Colless, JSS 8615 (1 ♂, ANIC); Culgoa Floodplain National Park, 11.8 km N Headquarters (CG 2 AM), 28 ° 54 ’ S, 147 ° 8 ’ E, 151 m, Brigalow, 20. I – 18. III. 2010, C. Kelly & A. Coward, Malaise trap, CNC 576673 (1 ♂, QM); Culgoa Floodplain National Park, 7 km NW Toulby Gate (TOLB 7 M), 28 ° 57 ’ S, 146 ° 53 ’ E, 138 m, Coolibah, reedy swamp, 17. XII. 2009 – 20. I. 2010, C. Kelly & A. Coward, Malaise trap, CNC 575106, CNC 575108 – 9 (3 ♂, QM); Culgoa Floodplain National Park, 7 km North-Northwest of Toulby Gate (CG 4 AM), 28 ° 56 ’ S, 146 ° 55 ’ E, 160 m, Gidgee, 17. XII. 2009 – 20. I. 2010, Kelly & Coward, Malaise trap, CNC 575354 – 6 (3 ♂, QM); Culgoa Floodplain National Park, 7 km Northwest of Toulby Gate (TOLB 7 M), 28 ° 57 ’ S, 146 ° 53 ’ E, 138 m, reedy swamp, 20. I – 19. III. 2010, C. Kelly & A. Coward, Malaise trap, CNC 575368 (1 ♂, QM); Culgoa Floodplain National Park, 7 km WNW Toulby Gate (TOLB 6 M), 28 ° 57 ’ S, 146 ° 52 ’ E, 138 m, Callitris, 19. III – 20. V. 2010, C. Kelly & A. Coward, Malaise trap, CNC 575027 (1 ♂, QM); Tasmania: 10 km NW by N St. Helens, 41 ° 15 ’ S, 148 ° 10 ’ E, 14. I. 1983, I. D. Naumann, J. C. Cardale, yellow pan trap, JSS 9255, JSS 9265 (2 ♂, ANIC); Victoria: 12 km SE Merrijig, Howqua River, 37 ° 12 ’ S, 146 ° 22 ’ E, 30. XI. 1971, Neboiss, JSS 8817 (1 ♂, MVMA); 17 km SE of Merrijig, 8 mile Creek off Howqua River, 37 ° 13 ’ S, 146 ° 24 ’ E, 1. XII. 1971, Neboiss, JSS 8814 (1 ♂, MVMA); Dartmouth Survey, Eight Mile Creek, Locality JA, 37 ° 13 ’ S, 146 ° 26 ’ E, 12. IV. 1973, JSS 8767 (1 ♂, MVMA); Western Australia: 80 km S Pardoo Road House on Shay Gap Road, 20 ° 31 ’ S, 120 ° 8 ’ E, 120 m, near spring with bog, 1 – 14. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15680 (1 ♀, CNC); Cape Range National Park; Milyering Ranger House, 22 ° 2 ’ S, 113 ° 56 ’ E, 22 m, below cliff near Ficus, 28. IV – 11. V. 2003, M. E. Irwin & F. D. Parker, Malaise trap, JSS 15657 (1 ♂, CNC); Karijini National Park, Juna Downs Road, 22 ° 42 ’ S, 118 ° 25 ’ E; 22 ° 43 ’ S, 118 ° 25 ’ E, 789 m; 799 m, between drying pools in rocky Turee Creek; near rock outcrop, Eucalyptus Eremophila scrub, 14 – 19. V. 2003; 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16027, JSS 16061 (1 ♂, 1 ♀, ANIC); Karijini National Park: Juna Downs Road (between drying pools in rocky Turee Creek), 22 ° 42 ’ S, 118 ° 25 ’ E, 789 m, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, CNCD 3817, CNCD 3819, CNCD 3837, CNCD 3868 – 9 (1 ♂, CNC; 4 ♂, QM); Millstream-Chichester National Park, Snappy Gum Drive, 21 ° 34 ’ S, 117 ° 4 ’ E, 329 m, across head of deep rocky gully, 27. IV – 2. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16023 (1 ♂, ANIC); Millstream-Chichester National Park, Snappy Gum Drive, 21 ° 34 ’ S, 117 ° 4 ’ E, 329 m, over deep rocky gully, 2 – 11. V. 2003, C. Lambkin, D. Yeates & J. Recsei, Malaise trap, JSS 16181 (1 ♂, ANIC); Newmanns Rocks, 50 km W of Balladonia Motel, 32 ° 6 ’ S, 123 ° 10 ’ E, 31. XII. 1986, G. and A. Daniels, JSS 8914 (1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF09ECB3D8D9EDD3FD4F11D8.taxon	distribution	Distribution: Australia (Australian Capital Territory, New South Wales, Queensland, Tasmania, Western Australia) (Fig. 137). Notes: Tomosvaryella uncinata sp. nov. is widespread in forested parts of Australia and has not been found hilltopping. Intraspecific genetic distance ranges from 0.0 %. It is genetically most similar to T. cona sp. nov. (2.6 – 4.8 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF0CEC8ED8D9EFF6FB6D1408.taxon	description	Figs 101 A – D, 128, 153 F	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF0CEC8ED8D9EFF6FB6D1408.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having dense short bristles; elongated surstyli, strongly bent at apex (Fig. 101 A); phallic guide with 3 – 6 dorsolateral spines, one is sinuous and longer (Fig. 101 B – D); elongated gonopods towards surstyli in ventral view (Fig. 101 B).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF0CEC8ED8D9EFF6FB6D1408.taxon	description	Description: MALE: Body length: 3.0 – 3.1 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 – 2.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 3 – 4 pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum brownish pollinose and with 6 – 8 bristles along the distal edge. Dorsocentral bristles distinct, dark, somewhat longer along frontal edge, numerous pale bristles around postpronotum present. Halter, knob pale, stem brown. Legs. Mid coxa with 2 – 4 long pale bristles. Trochanters and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments yellow, ventrally paler, last segment brown. Hind trochanter with dense short bristles and no distinct bristles; ventrobasal spines (2) present on fore and weaker on mid femur. Ventroapical row of spines on fore femur absent; 4 – 5 small, black spines on mid femur; hind femur without ventroapical spines, except 12 – 14 equally spaced bristles posteroventrally on a slight keel, longer on distal half, longest up to the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present (short). Hind metatarsus moderately flattened, almost as long as 2 – 3 combined, with dense short bristles ventrally (t 1 only). Pulvilli shorter than last tarsal segment. Wing. Length: 3.0 mm. Upper side of basal costal cell with one long dark bristle. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m slightly distal to middle of discal cell. 2 – 3 distinct brown setulae on tegula. Abdomen. Viewed obliquely from front tergites subshiny black, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (spot on t 5 larger). Dispersed strong dark bristles on all tergites present, longest on tergite 5 in length up to 1 / 3 the width of hind tibia at distal end. Lateral bristles on first tergite present, 4 – 6 short dark bristles up to as long as hind femur’s width at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1.5 – 1.7 times as long as ST 8. Genitalia without dissection: ST 8 medium sized, rounded-off square in dorsal view, brown and with a few distinct bristles, otherwise with velvet-like coverage; membranous area round, mostly directed posteriorly; epandrium brown, surstyli yellowish and long. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.1). Surstyli rather symmetrical, elongated, moderately narrowed before apex, strongly bent to each other at apex, right surstylus with small depression in middle on inner margin (Fig. 101 A). Genital capsule in ventral view: gonopods elongated towards surstyli, right one inclined to inner side, subepandrial sclerite widened distally; distinct hypandrial apodeme; phallic guide with 3 – 6 dorso-dorsolateral spines, one is sinuous and longer (Fig. 101 B); Genital capsule in lateral view: both surstyli curved towards sternite, broadened at basal half, moderately narrowed to apex, phallic guide strong, pointed towards surstyli at apex (Fig. 101 C – D). FEMALE: Body length: 2.7 – 2.8 mm. As male except for the following characters. Frons, eyes separated, as wide as 1.5 – 1.8 times the width at antennae; completely silver-grey pollinose on lower 1 / 3, shiny black in front of ocellar triangle, smooth change in transition zone to shiny black. Enlarged ommatidia silvery shining. Pulvilli and claws about 1 – 2 times as long as last tarsal segment, smaller on metatarsi. Female abdomen with silver spots on sides of tergites 4 – 6. Ovipositor. Straight in ventral view, straight and robust in lateral view (yellow piercer, brown base), reaching proximal end of 4 th segment; base brownish silvery pollinose with occasional bristles. LP: LB = 2.4. LDP: LPP = 2.5. (Fig. 153 F).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF0CEC8ED8D9EFF6FB6D1408.taxon	etymology	Etymology: The Latin word unda means wave and is in reference to the long, sinuous spine on the phallic guide. Examined material: HOLOTYPE: AUSTRALIA: Western Australia: Karijini National Park: Juna Downs Road (between drying pools in rocky Turee Creek), 22 ° 42 ’ S, 118 ° 25 ’ E, 789 m, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, CNCD 3835 (1 ♂, WAM); PARATYPES: AUSTRALIA: Queensland: 12 km E of Heathlands homestead, 11 ° 45 ’ S, 142 ° 41 ’ E, 17. III. 1992, G. Daniels & M. A. Schneider, JSS 8322 (1 ♀, QM); 2 km along Ridgepole Waterhole Road, 10 km ESE Musselbrook Resource Centre, Lawn Hill National Park, 18 ° 38 ’ S, 138 ° 12 ’ E, 220 m, 12. V. 1995, G. Daniels & M. A. Schneider, JSS 8293 (1 ♂, QM); Bamaga-Captain Billy Creek Road Junction, 16 km NE Heathlands Homestead, 11 ° 41 ’ S, 142 ° 42 ’ E, 16. III. 1992, G. Daniels & M. A. Schneider, JSS 8269 (1 ♂, QM); Bertie Creek (pump), 1 km SE Heathlands Homestead, Cape York Peninsula, 11 ° 45 ’ S, 142 ° 35 ’ E, 16. III. 1992, G. Daniels & M. A. Schneider, JSS 8272 (1 ♂, QM); Brisbane, Mount Coot-tha, 27 ° 29 ’ S, 152 ° 57 ’ E, 170 m, hilltop, 19. IV. 1998, 13. IV. 1997, 16. IX. 1997, 26. X. 1997, J. Skevington, hand collected, 9: 15 – 9: 45 am, 9: 45 – 10: 15 am, JSS 3840, JSS 3887, JSS 3791, JSS 3796, JSS 3798, JSS 3801 – 3, JSS 3891 (9 ♂, QM); 170 m, hilltop, 16. VIII. 1997, J. & A. Skevington & C. Lambkin, hand collected, JSS 3947 – 9 (3 ♂, CNC); 170 m, hilltop, 15. V. 1999, 8. II. 1998, 25. IX. 1997, 27. I. 1998, J. & A. Skevington, hand collected, JSS 3059, JSS 3072, JSS 3075, JSS 3077, JSS 3082, JSS 3084, JSS 3086, JSS 3089 – 91, JSS 3093, JSS 3238, JSS 3243 – 7, JSS 3249 – 52, JSS 3792, JSS 5774, JSS 5776 – 7, JSS 5780, JSS 5782 – 4 (29 ♂, QM); Brisbane, Mt. Coot-tha, 27 ° 29 ’ S, 152 ° 57 ’ E, 170 m, hilltop, 16. VIII. 1997, J. & A. Skevington & C. Lambkin, hand collected, JSS 735 – 6 (2 ♂, USNM); 170 m, hilltop, 7. XII. 1997, 8. XI. 1997, J. & A. Skevington, hand collected, JSS 957, JSS 962, JSS 1008, JSS 1032, JSS 2329, JSS 2332, JSS 2341, JSS 2350 (8 ♂, QM); Carnarvon National Park, Hilltop north of Tombs, 25 ° 5 ’ S, 147 ° 52 ’ E, 26. XI. 1997, J. Skevington & C. Lambkin, hand collected, JSS 2212 (1 ♂, QM); Carnarvon National Park, Mount Moffatt Section, 25 ° 4 ’ S, 148 ° 3 ’ E, 760 m, in bottom land, 25. XI. 1995, S. D. Gaimari, Malaise trap, JSS 8836 (1 ♀, INHS); Carnarvon National Park, Mount Moffatt Section, 25 ° 2 ’ S, 147 ° 58 ’ E, 27. XI – 2. XII. 1997, J. Skevington & C. Lambkin, Malaise trap, JSS 8414 (1 ♂, QM); Western Australia: Karijini National Park, Hamersley-Mount Bruce Road, 22 ° 34 ’ S, 118 ° 18 ’ E, 757 m, dry rocky creek bed, Eucalyptus grassland, 25. IV – 14. V. 2003, C. Lambkin, T. Weir, Malaise trap, JSS 15967, JSS 16115, JSS 16126 (2 ♂, 1 ♀, ANIC); Karijini National Park, Juna Downs Road, 22 ° 44 ’ S, 118 ° 25 ’ E, 798 m, dry Turee Creek, grassy open Eucalyptus scrub, 14 – 19. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16112 (1 ♀, ANIC); Karijini National Park, Weano Gorge Road, 22 ° 21 ’ S, 118 ° 15 ’ E, 695 m, grassy dry creek, Eucalyptus Acacia scrub, 20 – 25. IV. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 16035 (1 ♂, ANIC); Karijini National Park, off Karijini Drive, 22 ° 35 ’ S, 118 ° 17 ’ E, 724 m, dry wash: closed Acacia scrub, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, JSS 15590 (1 ♂, ANIC); Karijini National Park, Juna Downs Road (between drying pools in rocky Turee Creek), 22 ° 42 ’ S, 118 ° 25 ’ E, 789 m, 25. IV – 14. V. 2003, C. Lambkin & T. Weir, Malaise trap, CNCD 3845 (1 ♀, CNC); Junee State Forest; 9 NW of Marlborough Z 55; 705229, 7476480, 22 ° 48 ’ S, 149 ° 0 ’ E, 4. X. 2002, M. Mathieson, JSM 504, CNC 484120 (1 ♂, CNC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF0CEC8ED8D9EFF6FB6D1408.taxon	distribution	Distribution: Australia (Queensland, Western Australia) (Fig. 128). Notes: This species is commonly found hilltopping in Eucalyptus forest and is particularly common at Mount Coot-tha in Brisbane. Intraspecific genetic distance ranges from 0.0 % to 0.8 %. It is genetically most similar to T. patula sp. nov. and T. dolicholoba sp. nov. (4.0 – 4.4 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF32EC8CD8D9EBAFFC9411D8.taxon	description	Figs 102 A – E, 137	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF32EC8CD8D9EBAFFC9411D8.taxon	diagnosis	Diagnosis: This species can be recognized by the hind femur having 10 – 15 equally spaced bristles posteroventrally; hind trochanter with a keel; surstyli straight in dorsal and lateral views (Fig. 102 A, D – E); gonopods triangle-shaped; hypandrium large, with two small protrusions towards gonopods ventroapically (Fig. 102 B), phallus with three spines emerging before ejaculatory ducts; phallic guide with 3 – 6 dorsolateral spines of different size.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF32EC8CD8D9EBAFFC9411D8.taxon	description	Description: MALE: Body length: 3.1 mm. Head. Scape and arista dark, pedicel brown, flagellum light brown and tapering, tips lighter. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 3. Thorax. Postpronotum pale, with 3 – 4 short pale bristles. Mesonotum (viewed obliquely from front) silvery pollinose, anterior part greyer. Dorsocentral bristles short, dark, longer in frontal half, numerous pale bristles around postpronotum present. Scutellum silvery brown pollinose and with 4 – 8 short dark bristles along distal edge. Halter, knob pale, stem brown with 3 – 4 short bristles, base dark. Legs. Mid coxa with 4 – 5 long dark bristles. Trochanters brown, femora black, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments light brown. Hind trochanter with a keel; hind femur without ventroapical spines, except 10 – 15 equally spaced bristles posteroventrally, longer on distal half, longest up to the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres flattened (t 1 – 3), hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 3.2 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 1 – 2 tiny setulae on tegula. Abdomen. Viewed obliquely from front tergites black, tergite 1 silvery grey, Lateral bristles on first tergite present, 2 – 4 dark bristles up to as long as half of hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.7), more widened at its base. Both surstyli straight, left surstylus longer, right one with pointed projection before apex on inner margin (Fig. 102 A). Genital capsule in ventral view: both gonopods widened at base, extended towards surstyli, triangle-shaped; hypandrium large, the height is twice the gonopods height with two small protrusions towards gonopods ventroapically, phallus trifid with three spines emerging before ejaculatory ducts; phallic guide with 3 – 6 different-sized dorsolateral spines, one is longer and bent downwards at its tip; subepandrial sclerite small (Fig. 102 B); Genital capsule in lateral view: both surstyli straight (Fig. 102 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 102 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF32EC8CD8D9EBAFFC9411D8.taxon	etymology	Etymology: From the Latin vastus, desert, in reference to the species being found in the vast desert of central Australia. Examined material: HOLOTYPE: AUSTRALIA: South Australia: N [ea] r Victory Well, Everard P [ar] k St [atio] n, 27 ° 3 ’ S, 132 ° 30 ’ E, 5. XI. 1970, G. F. Gross, Malaise trap, JSS 8723 (1 ♂, SAM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF32EC8CD8D9EBAFFC9411D8.taxon	distribution	Distribution: Australia (South Australia) (Fig. 137).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF30EC8AD8D9EE9FFDA4134C.taxon	description	Figs 103 A – F, 143, 154 D	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF30EC8AD8D9EE9FFDA4134C.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a few tiny dark pegs and velvet-like pile; elongated surstyli in dorsal view (Fig. 103 A); gonopods elongated towards surstyli in ventral view (Fig. 103 B); one of the ejaculatory ducts with 4 – 6 small spines in lateral view (Fig. 103 D – F); phallic guide with 6 – 8 dorsolateral spines in lateral view (Fig. 103 F).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF30EC8AD8D9EE9FFDA4134C.taxon	description	Description: MALE: Body length: 2.6 – 2.7 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1 / 5. Thorax. Postpronotum pale, with 3 – 4 pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum silvery pollinose and with 8 – 10 bristles along the distal edge. Dorsocentral bristles distinct, dark, somewhat longer along frontal edge, numerous dark bristles around postpronotum present. Halter, knob pale, stem brown. Legs. Mid coxa with 1 – 2 long dark bristles. Trochanters and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae yellow (tibiae otherwise brown), tarsal segments yellow-brown, ventrally paler, last segment darker. Hind trochanter with few dark tiny pegs (5 – 8, not always visible) in a line at the base ventrally, velvet-like coverage; ventrobasal spines (1) present on fore and mid femora. Ventroapical row of spines on fore femur 2 – 3; 4 – 5 small, black spines on mid femur; hind femur without ventroapical spines, except 8 – 10 equally spaced bristles posteroventrally on distal half, longest up to 1.5 times the width of hind tibia at distal end. Subapical (distal) spines on first four tibiae absent. Hind metatarsus slightly flattened and, almost as long as 2 – 3 combined. Pulvilli shorter than last tarsal segment. Wing. Length: 3.0 – 3.1 mm. Upper side of basal costal cell with one long brown bristle. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2 – 3 distinct brown setulae on tegula. Abdomen. Viewed obliquely from front tergites brownish pollinose, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (larger on t 5). Dispersed strong dark bristles on all tergites present, longest on tergite 5 up to the width of hind tibia at distal end. Lateral bristles on first tergite present, 4 – 6 dark bristles up to 1.2 – 1.4 times as long as hind femur’s width at base. Postabdomen in dorsal view: T 6, S 7 invisible; T 5 1.2 – 1.4 times as long as ST 8. Genitalia without dissection: ST 8 medium sized, square in dorsal view, brown and with distinct bristles, otherwise covered with dense short bristles; membranous area cone-shaped; epandrium brown, surstyli yellow, long. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.0). Surstyli symmetrical, elongate, right surstylus smaller than left one, left inclined to right one at apex (Fig. 103 A). Genital capsule in ventral view: gonopods unequal in height, left longer than right one, elongated towards surstyli, subepandrial sclerite wide, covered with scattered short hair (Fig. 103 B). Genital capsule in lateral view: both surstyli curved towards sternite, phallus with three short ejaculatory ducts, one with 4 – 6 small spines, phallic guide with 6 – 8 dorso-dorsolateral spines (one is longer) (Fig. 103 D – F). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 103 C). FEMALE: Body length: 2.2 – 2.3 mm. As male except for the following characters. Frons, eyes separated, as wide as 1.6 – 1.8 times the width at antennae; completely silver-grey pollinose until up to 1 / 3 of the lower frons, shiny black in front of ocellar triangle, no transition zone; frons depressed along upper 2 / 3 with a groove in the median line. Enlarged ommatidia silvery shining. Pulvilli and claws about 2 – 2.5 times as long as last tarsal segment, smaller on metatarsi. Female abdomen with silver spots on sides of tergites 4 – 6. Ovipositor. Piercer straight in ventral view; long, curved upwards in lateral view, thin and reaching distal end of 2 nd segment; base brownish pollinose without bristles. LP: LB = 4.0. LDP: LPP = 3.0 (Fig. 154 D).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF30EC8AD8D9EE9FFDA4134C.taxon	etymology	Etymology: This species is named after the velvet-like covering of the hind trochanter (from the Latin velutinus, velvety). Examined material: HOLOTYPE: AUSTRALIA: South Australia: Flinders Ranges National Park, 4 km s [outh] Wilpeena Pound, 31 ° 33 ’ S, 138 ° 39 ’ E hilltop, 8. X. 1997, J. & A. Skevington, JSS 8394 (1 ♂, QM); PARATYPES: AUSTRALIA: Queensland: Brisbane Forest Park, 27 ° 26 ’ S, 152 ° 51 ’ E 200 m, 9 – 14. XI. 1995, M. E. Irwin, CNCD 136754, (1 ♂, CNC); Carnarvon National Park, Mount Moffatt Summit, 25 ° 4 ’ S, 148 ° 3 ’ E, hilltop, 2. XII. 1997, J. Skevington & C. Lambkin, JSS 1822, (1 ♂, QM); Mount Moffatt National Park, 25 ° 10 ’ S, 147 ° 50 ’ E, ~ 900 m, 10 – 13. II. 2004, S. & J. Wright, Malaise trap, JSS 19187, JSS 19189 (2 ♀, QM); Aldinga Sellicks Beach Resort, 25. III – 3. IV. 1987, J. A. Forrest, E. G. Matthews, Malaise trap, JSS 8822 (1 ♂, SAM); Gammon Ranges National Park, 30 ° 25 ’ S, 139 ° 14 ’ E, 12. X. 1997, J. Skevington, S. Winterton & C. Lambkin, Malaise trap, JSS 8323 (1 ♂, QM); Sandy Creek Conservation Park, 8 km northeast Gawler, 34 ° 36 ’ S, 138 °, 3. X. 1997, S. Winterton, J. & A. Skevington & C. Lambkin, Malaise trap, JSS 8401 (1 ♂, QM); Western Australia: 8 km E Salmon Gums, 32 ° 58 ’ S, 121 ° 42 ’ E, 6. I. 1987, G. & A. Daniels, JSS 8916 (1 ♂, QM); Fitzgerald River National Park, Old Ongerup Road near Hammersley River, 33 ° 45 ’ S, 119 ° 40 ’ E, 2 – 16. XI. 2003, J. Recsei & C. Lambkin, Malaise trap, ANIC Bulk Sample 2189, JSS 16211 (1 ♂, ANIC); Mount Magnet, Mount Waramboo, 28 ° 2 ’ S, 117 ° 49 ’ E, hilltop, 14. XII. 1999, J. Skevington, JSS 7463 (1 ♂, QM); Pine Hill, Cape Arid National Park, Balladonia Road, 33 ° 18 ’ S, 123 ° 22 ’ E, 31. X – 18. XI. 2003, C. Lambkin & J. Recsei, Malaise trap, ANIC Bulk Sample 2182, JSS 15592 (1 ♂, ANIC); South Australia: Wyperfeld NP, East Lookout, 35 ° 36 ’ S, 142 ° 7 ’ E, 22. X. 2014, J. H., A. W., & A. M. Skevington, A. D. Young, A. Wyatt, & S. Namek, JSM 6043; CCDB- 27867, CNC 384991 (1 ♂, CNC); Victoria: Hattah-Kulkyne NP, Warepil Lookout, 34 ° 43 ’ S, 142 ° 18 ’ E, 18. X. 2014, A. D. Young, A. Wyatt, & S. Namek, J. H., A. W., & A. M. Skevington, JSM 6109; CCDB- 27176, CNC 385057 (1 ♂, CNC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF30EC8AD8D9EE9FFDA4134C.taxon	distribution	Distribution: Australia (Queensland, South Australia, Victoria, Western Australia) (Fig. 143). Notes: This is a widespread but uncommon forest species that is sometimes found hilltopping. Intraspecific genetic distance ranges from 0.0 % to 5.6 %. It is genetically closest to T. trispina sp. nov. (5.8 – 8.6 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF36EC88D8D9ECEBFD0E1214.taxon	description	Figs 104 A – E, 138	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF36EC88D8D9ECEBFD0E1214.taxon	diagnosis	Diagnosis: This species can be recognized by the long epandrium and elongated and somewhat rectangular-shaped surstyli in dorsal view, both bent towards each other before the apex (Fig. 104 A); long and pointed gonopods in ventral view (Fig. 104 B); both surstyli straight in basal two thirds and bent towards sternite in lateral view (Fig 104 D – E); phallic guide with 4 – 6 spines dorsolaterally, one spine longer than others and bent toward gonopods at the apex (Fig. 104 C).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF36EC88D8D9ECEBFD0E1214.taxon	description	Description: MALE. Body length: 2.1 mm. Head. (holotype without head) Scape and pedicel dark. Flagellum tapering; light brown at the apex. Face silvery pollinose. Frons, upper part black, lower part distinctly silver pollinose; eyes touching for distance equal to 1 – 1.2 times the length of ocellar triangle. Occiput silvery pollinose. Thorax. Postpronotum light yellow, with 1 – 3 dark bristles. Mesonotum (viewed obliquely from front) grey pollinose. Scutellum silver pollinose and with 10 – 12 weak bristles along the distal edge. Dorsocentral and supra-alar bristles well developed, dark, longer on frontal part and numerous dark bristles around postpronotum. Halter, knob pale with two short bristles, stem and base dark. Legs. Mid coxa with 2 – 3 spines. All parts of legs dark brown except knees and basal 1 / 5 of tibiae yellow (tibiae otherwise brown), tarsal segments yellow. Hind trochanter covered by strong and short bristles. Ventroapical row of spines on fore femur missing, hind femur with 8 – 12 subapical spines. All tarsomeres with erect dark bristles dorsally. Pulvilli as long as last tarsal segment. Wing. Length: 2.3 mm. Upper side of basal costal cell with one dark and long bristles. Fourth costal section 2 – 2.5 times as long as third costal section. Cross-vein r-m at middle of discal cell. 6 – 8 dark setulae on tegula. Abdomen. Viewed obliquely from front tergites subshining black, tergite 1 and tergite 2 silvery grey. Lateral bristles on first tergite present, 6 – 8 brown bristles in a row up to as long as the width of hind femur at distal tip in lateral view. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.3). Surstyli elongate and symmetrical. Left surstylus slightly longer than right surstylus. Both surstyli finger-like shaped with inward tips (Fig. 104 A). Genital capsule in ventral view: subepandrial sclerite long, gonopods elongated towards surstyli, pointed at the apex with two small triangular projections before apex towards phallus (Fig. 104 B). Genital capsule in lateral view: phallus straight, with three short ejaculatory ducts; phallic guide with 4 – 6 spines dorsolaterally (one spine longer than others and bent toward gonopods at the apex) (Fig. 104 C). Right and left surstylus straight in basal two thirds and bent towards sternite in apical third (Fig. 104 D – E). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF36EC88D8D9ECEBFD0E1214.taxon	etymology	Etymology. This species is named after Warrumbungle National Park, the location where the holotype was collected and an important natural area that protects a wealth of insect diversity. Examined material: HOLOTYPE: AUSTRALIA: New South Wales: Warrumbungle National Park, 31 ° 14 ’ S, 149 ° 1 ’ E, 19. X – 3. XI. 1997, J. Skevington & S. Winterton, Malaise trap, JSS 8437 (1 ♂, QM); PARATYPES: AUSTRALIA: New South Wales: Ledknapper National Reserve, 7.3 km ENE Beulah Headquarters (LDN 2 M), 29 ° 21 ’ S, 146 ° 13 ’ E, Eucalyptus melanopholia, Spinifex, 10. XII. 2009 – 18. III. 2010, S. O’Sullivan & K. Taylor, Malaise trap, CNC 576514, CNC 576518 (2 ♂, CNC, QM); Queensland: Carnarvon National Park, Carnarvon Gorge, Consuelo land, 24 ° 57 ’ S, 148 ° 6 ’ E, 12. X. 2002, S. Boucher, sweeping, LEM _ 0016167 (1 ♂, LEM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF36EC88D8D9ECEBFD0E1214.taxon	distribution	Distribution: Australia (New South Wales, Queensland) (Fig. 138). Notes: The genitalia feature of specimen LEM _ 0016167 are similar to other specimens but there is a distinct keel on its hind trochanter. None of the other specimens have this keel. Tomosvareylla warrumbunglensis sp. nov. appears to be a dry sclerophyll forest species. It is genetically most similar to T. macrostyla sp. nov. (2.4 – 3.5 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF34EC86D8D9EDD3FCEA1110.taxon	description	Figs 105 A – E, 143	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF34EC86D8D9EDD3FCEA1110.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having dense short bristles and a few pegs along a slight ridge on the ventral side; elongated surstyli in dorsal view (Fig. 105 A); elongated gonopods, tapering at apex with small projection on inner margin before apex; subepandrial sclerite with some wrinkled in its middle and rounded outer margin in ventral view (Fig. 105 B); phallic guide with 5 – 6 dorso-dorsolateral spines (Fig. 105 D).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF34EC86D8D9EDD3FCEA1110.taxon	description	Description: MALE: Body length: 2.6 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for 1.5 x length of ocellar triangle. Occiput silvery pollinose. Thorax. Postpronotum pale, with 0 – 1 pale bristles. Mesonotum (viewed obliquely from front) silvery pollinose, anterior part more grey, grayish also from the side. Scutellum silvery pollinose and without bristles along the distal edge. Dorsocentral bristles weakly developed, dark, somewhat longer along frontal edge, numerous dark bristles around postpronotum absent. Halter, knob pale with brownish tinge, stem brown. Legs. Mid coxa with 3 – 4 strong, dark brown bristles. Trochanters pale brown (yellow band posteroventrally) and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 4 of tibiae yellow (tibiae otherwise brown), tarsal segments yellow, last segment darker. Hind trochanter with velvet-like bristles, posteriorly a tiny shiny spot on the inside; a few pegs along a slight edge on the ventral side; ventrobasal spine absent on fore and mid femur. Ventroapical row of weak spines on fore femur 3 – 4; 6 – 8 small, black spines on mid femur; hind femur without ventroapical spines; 14 – 15 equally spaced bristles posteroventrally, longer on distal half, longest up to 1 / 4 the width of hind tibia at distal end. Short, subapical spines on fore- and mid tibiae distally. Hind tibia slightly bent in posterior view. Metatarsi slightly flattened, first tarsomere elongated, almost as long as 2 – 5 combined. Pulvilli shorter than last tarsal segment. Wing. Length: 2.7 mm. Upper side of basal costal cell with one short dark bristle. Fourth costal section 2.5 times as long as third costal section. Cross-vein r-m at 3 / 5 of discal cell. 2 – 3 short brown setulae on tegula. Abdomen. Dissected. tergite 1 silvery grey, lateral bristles on first tergite present, 3 – 4 dark bristles up to 2 / 3 as long as hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.5). Surstyli elongated, right surstylus smaller than left one and curved towards left one (Fig. 105 A). Genital capsule in ventral view: gonopods elongated towards surstyli, tapering at apex with small finger-like projection located at inner margin before apex, unequal in height, left slightly longer than right one, subepandrial sclerite with some wrinkled in its middle and rounded outer margin (Fig. 105 B). Genital capsule in lateral view: right surstylus rather straight, left surstylus curved towards sternite, phallic guide with 5 – 6 dorsolateral spines (one is longer) (Fig. 105 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 105 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF34EC86D8D9EDD3FCEA1110.taxon	etymology	Etymology: This species is dedicated to Shaun Winterton, one of the collectors of the holotype. Shaun and JHS did their doctorates together in Brisbane and did a lot of collecting during that time. Many of Shaun’s specimens have been critical to pipunculid revisions. He collected 23 % of the species and 4 % of the specimens used in this revision (Supplementary file 2). Examined material: HOLOTYPE: AUSTRALIA: Queensland: 91.2 km west Windorah, 25 ° 22 ’ S, 141 ° 47 ’ E, Malaise trap in dry creek bed, 31. VIII – 5. IX. 1997, S. Winterton, J. & A. Skevington, JSS 8344 (1 ♂, QM); PARATYPE: AUSTRALIA: Northern Territory: Koongarra, 15 km E of Mount Cahill, 12 ° 52 ’ S, 132 ° 51 ’ E, 6 – 9. III. 1973, D. H. Colless, JSS 8548 (1 ♂, ANIC).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF34EC86D8D9EDD3FCEA1110.taxon	distribution	Distribution: Australia (Northern Territory, Queensland) (Fig. 143). Notes: The terminalia characters of the paratype specimen, JSS 8548, are similar to the holotype but the hind trochanter has only with a few pegs along its edge. It is conceivable that they are different species, but more material would be needed to support this. Certainly, the habitats are different with the holotype having been captured in a desert environment and the paratype being found in monsoon forest. This species is genetically closest to T. bulbosa sp. nov. (2.3 – 3.8 % pairwise divergence) (Supplementary file 3).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF3AEC86D8D9EED7FB4F1738.taxon	description	Figs 106 A – E, 136, 144 E	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF3AEC86D8D9EED7FB4F1738.taxon	diagnosis	Diagnosis: This species can be recognized by the hind trochanter having a keel covered by small teeth along its edge (Fig. 144 E); hind femur with two rows bristles posteroventrally and posterodorsally (Fig. 144 E); surstyli elongated, outer margin sinuous in dorsal view (Fig. 106 A); gonopods equal in height, extended towards surstyli (Fig. 106 B); phallic guide with 3 – 4 short dorsolateral spines; both surstyli boot-shaped in lateral view (Fig. 106 D – E).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF3AEC86D8D9EED7FB4F1738.taxon	description	Description: MALE: Body length: 3.0 mm. Head. Scape and pedicel dark; flagellum brown, grey pollinose, long tapering. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput silvery pollinose. Thorax. Postpronotum pale with 3 – 4 different-sized pale bristles. Mesonotum, postalar callus and scutellum brown pollinose, otherwise shiny, somewhat grey pollinose. Mesonotum with two rows of short, dark dorsocentral bristles, some longer bristles in upper margin behind postpronotum. Scutellum with some scattered short dark bristles. Halter, knob pale, tip brownish, stem dark. Legs. Mid coxa with 3 – 4 long dark bristles. Trochanters brown, femora black, silvery pollinose posteriorly except shiny hind femur; knees and basal 1 / 6 of tibiae light brown (tibiae otherwise dark), tarsal segments light brown. Hind trochanter with a keel, covered by small teeth along its edge (Fig. 144 E); two ventrobasal spines present on fore and absent on mid femur. Hind femur without ventroapical spines, except 8 – 10 equally spaced long bristles posteroventrally, longer on distal half, longest up to 2 / 3 the width of hind tibia at distal end and shorter bristles posterodorsally (Fig. 144 E). Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres distinctly flattened, hind metatarsus almost as long as 2 – 4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 3.0 mm. Upper side of basal costal cell with one long pale bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2 – 3 short dark setulae on tegula. Abdomen. First tergite grey pollinose with 6 – 8 different-sized lateral bristles up to as long as ¾ of hind femur width at base. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.7). Surstyli elongated, outer margin sinuous, rather widened at base, constricted in the middle, then broadened, as wide as the base, pointed at apex, left surstylus longer than right one (Fig. 106 A). Genital capsule in ventral view: gonopods equal in height, extended towards surstyli, pointed at apex, both gonopods with a triangular projection on inner margin; phallic guide with 3 – 4 short dorsolateral spines; subepandrial sclerite widened in the middle (Fig. 106 B); Genital capsule in lateral view: both surstyli boot shaped, straight in basal two thirds, broadened and strongly bent towards sternite in apical third (Fig. 106 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 106 C). FEMALE: Unknown.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF3AEC86D8D9EED7FB4F1738.taxon	etymology	Etymology: This species is named after the type locality. Six of the species included in this revision were collected in Wyperfeld National Park. This is the third largest park in Victoria and it protects a significant portion of the Victorian mallee diversity. Examined material: HOLOTYPE: AUSTRALIA: Victoria: Wyperfeld NP, East Lookout, 35 ° 36 ’ S, 142 ° 7 ’ E, 22. X. 2014, J. H., A. W., & A. M. Skevington, A. D. Young, A. Wyatt, & S. Namek, CNC 373426 (1 ♂, QM).	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
715487A7FF3AEC86D8D9EED7FB4F1738.taxon	distribution	Distribution: Australia (Victoria) (Fig. 136). Notes: This is another species that is only known from a single specimen found hilltopping.	en	Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H., Kelso, Scott (2023): Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species. Zootaxa 5599 (1): 1-271, DOI: 10.11646/zootaxa.5599.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
