taxonID	type	description	language	source
19BC8AAFDFF20FDC31D92A8811AF4C62.taxon	diagnosis	Diagnosis. Catharylla species have snow white to creamy white wings and short labial palpi. They can be separated from other Argyriini by the presence on the forewing of median and subterminal thin transverse lines, slightly curved, convex on costal 1 / 3. The labial palpi are also shorter in comparison to those of Vaxi. The highly variable male genitalia do not show any synapomorphy or generic diagnostic character. In females, a possible synapomorphy is the strongly reduced anterior and posterior apophyses of abdominal segments VIII and IX, but this is shared with some Crambini and a few other Crambinae (see Landry 1995).	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
19BC8AAFDFF20FDC31D92A8811AF4C62.taxon	distribution	Distribution. The genus is restricted to the Neotropical Region, from Costa Rica to Santa Catarina, Brazil, from sea level to 1300 m (Figs 43 - 46).	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
EF70CAC8C957F4C5822419A050DCF3EA.taxon	diagnosis	Diagnosis. On the forewing (Fig. 1), the seven, thin, marginal dark brown dashes with the most tornal two shaped like spots will separate this species from the others. In male genitalia (Fig. 11), the strongly sclerotized double costal arm of the valva with the ventral arm tubular is a distinctive character. In female genitalia, the best diagnostic character is the sclerotized projection latero-ventrally on sternite VIII (Fig. 34).	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
EF70CAC8C957F4C5822419A050DCF3EA.taxon	description	Description. Male (n = 17) (Fig. 1): Antenna brown with light ochreous scales; with patch of dark brown scales at base. Maxillary palpus ringed with brown at base and half of length, white tipped. Labial palpus: 1.4 - 1.6 mm long; ochreous, slightly lighter basally, ringed with dark brown at 2 / 3, white tipped. Thorax slightly ochreous at collar. Foreleg coxa white; femur ochreous, dark brown dorsally; tibia and tarsomeres ochreous, distally ringed with brown. Midleg and hindleg light ochreous; tibia-femur joint brown on midleg; tarsomeres II-V brown to dark brown on upperside, with white ringed tips. Abdomen dull white to greyish brown. Forewing length: 9.0 - 10.5 mm; snow white, with yellow ochreous to brown costal margin, partially disrupted when meeting transverse lines; median line yellow ochreous; subterminal line yellow ochreous to brown, forming small triangular spot on costal margin; subapical triangle on costal margin ochreous; outer margin slightly ochreous with five dark brown dashes regularly spaced or sometimes forming faintly continuous line, and one cubital and one anal spots, with cubital spot slightly displaced toward base; fringes brass colored; underside dull white to light ochreous along costal margin, with marginal dashes pronounced. Hindwing snow white, veins slightly ochreous, with shiny aspect; marginal line thin, brown, pronounced up to CuA 1, then shiny white; fringes white; underside white, with same margin as on recto. Tympanal organs (n = 8): Tympanic pockets extending slightly beyond transverse ridge, rounded. Tympanic drum elongate, more or less oval, postero-laterally extended beyond transverse ridge. Male genitalia (n = 8) (Figs 11, 12): Uncus slightly down-curved, about 3 / 5 length of tegumen arms, with few setae laterally; tip pointed; uncus arms not separated at base, forming low bump medio-ventrally. Gnathos arms joining at half their length; distal half with short, rounded, dorsal projection at base; directed upward subapically at about 50 ° angle; slightly shorter than uncus and thinner. Tegumen pedunculi progressively widening toward uncus; dorsal connection of tegumen about 1 / 3 length of pedunculi; ventral margin straight; dorsal margin slightly convex, bare. Cucculus moderately wide, narrowing in distal 1 / 4; costal arm of valva double, bare, about as long as cucculus, joined to cucculus until 3 / 5 of its length; ventral arm thin, tubular, strongly sclerotized, slightly curved inward, apex directed upward, narrowed, pointed; dorsal arm broader, slightly shorter than ventral arm, straight, apically rounded, less thickly sclerotized than ventral arm. Vinculum enlarging latero-dorsally, ventrally narrow; saccus short, rounded. Juxta triangular, apically broadly rounded, slightly curved downward, basally projected into two large lateral lobes. Phallus almost straight, with slightly upturned sclerotized apex; vesica covered with microspicules barely visible, with one large, curved, pointed cornutus. Female (n = 10): Labial palpi: 1.1 - 1.4 mm long. Forewing length: 11 - 15 mm; frenulum triple. Female genitalia (n = 6) (Fig. 34): Papillae anales slightly projected ventrally and dorsally, dorsally forming prominent sclerotized rounded bulge. Posterior apophyses widened basally, 0.35 - 0.5 x length of papillae anales. Segment VIII circular in cross section, enlarging progressively toward papillae. Tergite VIII narrow, about 2 / 5 length of sternite VIII, with short setae along posterior edge. Anterior apophyses wide at base, about 0.1 x length of papillae. Sterigma with thin slightly sclerotized membrane covered with minute spicules dorsad of ostium bursae, with posterior margin slightly indented; with sclerotized projection laterad from sternite VIII antero-ventrally, with tip bifid, longer part directed downward, shorter part lateral, curved posterad. Basal part of ductus bursae ventrally sclerotized, looping and narrow, progressively widening toward corpus bursae. Corpus bursae poorly differenciated from ductus, twice as long as wide, without signum.	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
EF70CAC8C957F4C5822419A050DCF3EA.taxon	distribution	Distribution. The species occurs in lowlands in the three Guianas and Brazil (Fig. 45).	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
EF70CAC8C957F4C5822419A050DCF3EA.taxon	etymology	Etymology. Bijuga comes from the Latin bijugus, a, um which means " yoked together, double ", in reference to the bifid costal arm of the male genitalia.	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
4DFD6E9EE74A8777069BDF702ED4C9AC.taxon	diagnosis	Diagnosis. From Catharylla gigantea, Catharylla chelicerata differs in having the male costal arm hook shaped, longer, and thinner than in Catharylla gigantea, and the juxta is strongly downcurved, apically conical whereas it is long, almost straight, without apical conical projection downward in Catharylla gigantea. In female genitalia the sterigma forms a strongly sclerotized symmetrical structure made of two asymmetrical bell-shaped cavities, opened anterad in Catharylla chelicerata whereas it forms a pair of shallow pockets opened posterad in Catharylla gigantea.	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
4DFD6E9EE74A8777069BDF702ED4C9AC.taxon	description	Description. Male (n = 21) (Figs 2, 9): Head with ochreous to brown chaetosemata. Antenna greyish brown with light brown scales, with patch of brown scales at base. Maxillary palpus ochreous to dark brown, lightly ringed with dark brown at 2 / 3, white tipped. Labial palpus: 1.3 - 2.0 mm long; ochreous, basally white, tip of segment II light greyish-brown; white tipped. Thorax with dark brown patch at collar. Foreleg coxa white; femur white, ashen brown dorsally, tibia and tarsomeres ochreous, distally ringed with dark brown. Midleg femur white, tibia ashen brown basally, tarsomeres ochreous, brown to ashen brown on upperside, with white ringed tips. Hindleg white, tarsomere I ochreous; II-V brown on upperside, with white ringed tips. Abdomen dull white to light ochreous. Forewing length: 10.5 - 15.0 mm; costal band wide, brown from base to apex; median and subterminal transverse lines faded brown, sometimes completely faded; dark brown spots on apical margin forming more or less continuous line; fringes brass colored; underside white with costal margin brown; outer margin with somewhat triangular spots. Hindwing snow white, with marginal spots between veins; fringes white; underside silvery white with marginal spots pronounced. Tympanal organs (n = 9): Transverse ridge almost straight medially. Tympanic pockets conical, extending slightly beyond transverse ridge. Tympanic bridge lightly sclerotized, dorsal base of praecinctorium sclerotized. Tympanic drums elongate, bean shaped, posteriorly reaching transverse ridge or slightly beyond. Male genitalia (n = 9) (Figs 13, 14): Uncus straight, of about 4 / 5 length of tegumen arms, dorso-ventrally compressed, with setae dorsally and laterally; apex truncated, slightly rounded, tip with short projection pointing posterad, ventrally convex, sometimes with median bump. Gnathos arms joining at 1 / 5 of length, regularly hook shaped, forming angle of about 100 ° with axis of basal arms, about 1 / 4 longer than uncus. Tegumen arms narrow at base, enlarging progressively toward dorsum to 2 x basal width, projected dorsally with bump at connection, connecting at distal 1 / 6. Cucculus densely setose, slightly directed upward on distal 1 / 3, apically truncated; basal 2 / 3 of costa of valva dorso-ventrally and laterally widened; costal arm hook-shaped, strongly sclerotized, directed upward at about 45 ° from costal arm base. Juxta triangular, curved downward, tip rounded and sac-like, basal lateral lobes curved ventrally. Saccus short, curved upward medially. Phallus narrow, S-shaped; vesica covered with tiny spicules, with one large, curved, pointed cornutus apically, preceded by string of 13 - 14 smaller cornuti increasing in size toward apex. Female (n = 4): Labial palpi: 1.8 - 2.2 mm long. Forewing: 15 - 19.5 mm. Frenulum quadruple. Female genitalia (n = 3) (Fig. 35): Papillae anales dorsally strongly produced posterad, with rounded bulge dorso-apically; ventrally slightly produced. Posterior apophyses 0.3 - 0.45 x length of papillae. Tergite VIII about half of length of sternite VIII. Anterior apophyses about 0.1 x length of papillae anales, slightly wider and rounded apically. Sternite VIII narrowing ventrally, densely covered with spinules, slightly connected medially at lamella antevaginalis; lamella antevaginalis slightly projected downward. Sterigma forming strongly sclerotized ventro-laterally symmetrical structure made of two asymmetrical bell-shaped cavities in ventral view, opened anterad, with dorsal lobe longer, expanding upward, slightly indented along latero- anterior margin; covered with minute punctuation. Ventro-basal section of ductus bursae tongue shaped, strongly sclerotized; ductus bursae long, ventrally sclerotized, widened and looped in basal half; enlarging progressively into corpus bursae. Corpus bursae egg-shaped with one signum.	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
4DFD6E9EE74A8777069BDF702ED4C9AC.taxon	distribution	Distribution. The species was found in French Guiana and Brazil (Amazonas) (Fig. 43).	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
4DFD6E9EE74A8777069BDF702ED4C9AC.taxon	etymology	Etymology. " Chelicerata " refers to the shape of the costal arms of the male valva, which look like mygalomorph chelicerae.	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
AFBC37122FE0A716CDB27F98E3B8A7AA.taxon	diagnosis	Diagnosis. From Catharylla chelicerata, Catharylla gigantea differs in having the male costal arm shorter, basally wide and tooth shaped while it is long, narrow throughout and hook shaped in Catharylla chelicerata. The juxta is long, tongue shaped, almost straight, and apically rounded, whereas it is downcurved and apically conical in Catharylla chelicerata. In female genitalia, the sterigma forms a pair of shallow pockets opened posterad whereas in Catharylla chelicerata the sterigma forms a strongly sclerotized symmetrical structure made of two asymmetrical bell-shaped cavities opened anterad.	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
AFBC37122FE0A716CDB27F98E3B8A7AA.taxon	description	Description. Male (n = 6) (Fig. 3): Head with ochreous chaetosemata. Antenna brown with light brown scales, with patch of dark brown scales at base. Maxillary palpus brown with dark brown spot at half of length, white tipped. Labial palpus: 1.6 - 2.4 mm long; ochreous to brown ochreous, basally white, with patch of dark brown scales at half of length, white tipped. Thorax with some brown at collar. Foreleg coxa white, femur white, ashen brown dorsally; tibia and tarsomeres brown-ochreous, distally ringed with dark brown. Midleg white with tibia-femur joint and base of tibia ashen; tarsomeres ochreous to brown ochreous with upperside brown to dark brown, white tipped. Hindleg white with tarsomeres II-V ochreous to brown ochreous, upperside brown, with white tips. Forewing length: 13.5 - 14.5 mm; snow white with wide brown to dark brown costal line from base to apex; median and subterminal transverse lines faded brown; dark brown spots on termen forming more or less continuous line; fringes brass colored; underside white, with costal margin brown ochreous, outer margin with subtriangular spots. Hindwing snow white; marginal spots dark brown between R 5, M 1, M 2, M 3, and CuA 1; fringe white; underside snow white, with same spots as on upperside. Tympanal organs (n = 5): Transverse ridge medially convex. Tympanic pockets extending slightly beyond transverse ridge, rounded. Tympanic bridge lightly sclerotized, dorsal base of praecinctorium sclerotized. Tympanic drums elongate, bean shaped. Male genitalia (n = 5) (Figs 15, 16): Uncus straight, about 3 / 4 length of tegumen arms, dorso-ventrally flattened, dorsally convex, ventral margin convex in basal half, concave in distal half; basally and laterally setose; apex slightly rounded, medially with short projection pointing postero-ventrally. Gnathos arms joining at 1 / 5, about 1 / 4 longer than uncus, regularly curved. Tegumen arms narrow at base, widening regularly to reach 1.5 x basal width dorsally, with connection at distal 1 / 6. Cucculus densely setose, broad at base, slightly widening and truncate at apex; costal arm of valva basally wide, short, tooth shaped, slightly curved inward. Juxta long, tongue shaped, almost straight, apically rounded, with basal lateral lobes curved ventrally. Saccus short, curved upward. Phallus narrow, S-shaped; vesica covered with tiny spicules, with string of 14 small cornuti increasing in size toward apex, with apical cornutus up to 5 x length of previous one. Female (n = 2): Labial palpi: 2.5 - 3.1 mm long. Forewing length: 17.5 - 22 mm; frenulum triple. Female genitalia (n = 2) (Fig. 36): Papillae anales ventrally projected. Posterior apophyses about 0.35 x length of papillae anales, narrow. Segment VIII narrowing ventrally, densely covered with spinules; narrow connection at lamella antevaginalis; lamella antevaginalis slightly projected downward. Sterigma forming pair of shallow pockets opened posterad at base of segment VIII. Anterior apophyses about 0.08 x length of papillae anales, of medium width, basally wide. Ductus bursae wide, as long as twice segment VIII, regularly enlarging into corpus bursae. Corpus bursae with one rounded signum.	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
AFBC37122FE0A716CDB27F98E3B8A7AA.taxon	distribution	Distribution. Catharylla gigantea has been found in French Guiana, Guyana, and Brazil (Amazonas) (Fig. 43).	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
AFBC37122FE0A716CDB27F98E3B8A7AA.taxon	etymology	Etymology. The name comes from the Latin giganteus, a, um meaning very large.	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
18ED9374AA903199FE961C3AF9ADED84.taxon	diagnosis	Diagnosis. From Catharylla serrabonita and Catharylla coronata, Catharylla tenellus can be separated by the median transverse line, which is faintly convex towards costa, whereas it is more strongly convex in Catharylla coronata and Catharylla serrabonita. The male genitalia provide the best diagnostic characters. The most obvious refers to the transtilla, which forms a pair of short, narrow sclerotized arms with pointed tips, projecting posterad, with, in between, a pair of brushes directed medio-ventrally, whereas it forms a pair of arms pointing posterad with a string of spines ventrally in Catharylla serrabonita and Catharylla coronata. In female genitalia, the anterior angle of sternite VIII is directed downward into a more or less rounded projection covered with short spinules of same length, whereas it is projected anterad in Catharylla serrabonita, and it is not projected in Catharylla coronata.	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
18ED9374AA903199FE961C3AF9ADED84.taxon	distribution	Distribution. The species is known from Brazil in the Atlantic Forest (Bahia, Minas Gerais, Parana, Rio de Janeiro, Sao Paulo) (Fig. 45). Notes. The species was described from " one female collected in Brazil, near Rio de Janeiro ". Hence, the lectotype designated by a label by S. Bleszynski is not warranted. This designation is presumably based on the fact that Zeller (1863: 50) mentions a pair deposited in the Vienna Museum. The association of sexes in this species is not 100 % certain. Specimens from Porto Seguro, Brazil show a divergence of 3.34 % in COI barcode sequences with the specimen from Ubatuba, Brazil. In morphology, differences in male genitalia are also observed: in the specimens from Bertioga, Caraca, Sao Paulo and Ubatuba the costal arm of the valva is wide and 1 / 3 of the length of the cucullus, almost reaching its tip, and the dorsal edge at base is slightly produced (Fig. 27). In the specimens from Porto Seguro, the costal arm is about 1 / 5 the length of the cucullus, relatively narrow, and the dorsal edge is slightly produced at base (Fig. 29). Another form, from Caraca, Minas Gerais (Fig. 28) was also found. No differences were found in the female genitalia. We feel that specimens and data are currently lacking to conclude that possibly more than one taxon should be recognized under Catharylla tenellus, or that there is indeed a deep divergence in the COI barcode between populations of this species.	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
C575A3423350B2EB56556E02A23B7179.taxon	diagnosis	Diagnosis. From Catharylla serrabonita and Catharylla tenellus, Catharylla coronata can be separated with characters of the male genitalia: the uncus is apically bifid and grooved on distal 1 / 5 in Catharylla coronata whereas it is only indented medially at apex in Catharylla serrabonita and Catharylla tenellus; the costal arm of the valva is short and the apex is curved inward in Catharylla coronata whereas the costal arm is longer and points postero-dorsally in the other two species; the transtilla forms a pair of sclerotized arms slightly bent inward distally, ventrally with a row of short spines increasing in size from base to apex whereas it forms a pair of short, narrow sclerotized arms with pointed tips, projecting posterad, and with a pair of brushes directed medio-ventrally in Catharylla tenellus and a pair of sclerotized arms strongly bent inward on distal 1 / 4 and with a string of long spines of same length medially along it in Catharylla serrabonita; the juxta is shorter than in Catharylla tenellus, and regularly narrowing toward apex whereas it is strongly narrowing on distal 1 / 4 in Catharylla serrabonita; the ventral projections of the juxta form a pair of shallow pockets whereas they are bell-shaped in Catharylla serrabonita and thumb-like in Catharylla tenellus; the vesica has a row of 6 - 7 cornuti in Catharylla coronata whereas it does not show any cornuti in Catharylla serrabonita and Catharylla tenellus. In the female genitalia of Catharylla coronata, the anterior angle of sternite VIII is not projected whereas it is rounded, projected anterad and covered with short spinules in Catharylla serrabonita, and projected downward in Catharylla tenellus. The anterior apophyses are quadrangular, anvil shaped whereas they are spine like in the other two species.	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
C575A3423350B2EB56556E02A23B7179.taxon	description	Description. Male (n = 21) (Fig. 5): Head white with ochreous chaetosemata. Antenna brown, with whitish ochreous scales and patch of brown scales at base. Maxillary palpi light ochreous to ochreous, white tipped. Labial palpi: 1.6 - 1.85 mm long; light ochreous, white tipped. Thorax white, with ochreous patch at collar. Foreleg coxa white; femur white, dorsally dark brown; tibia and tarsomeres ochreous, distally ringed with brown; midleg and hindleg white to light ochreous, tarsomeres II-V ochreous, upperside brown, with white ringed tips. Forewing length: 10 - 13 mm; costal margin line thin, light ochreous, apically faded; median transverse line light ochreous, concave on costal half, more or less disrupted; subterminal transverse line ochreous, curving toward base on costal half; R 5 vein faintly marked apically with ochreous; outer margin ochreous with 7 pronounced dark brown spots more or less triangular between veins, sometimes connecting; fringes brass colored; underside white ochreous to ochreous, costal margin basally brown; outer margin with pronounced spots. Hindwing white to creamy white, usually with marginal brown spots between Sc + R 1, Rs, M 1, M 2, M 3, CuA 1 and CuA 2, forming more or less continuous line; fringes white; underside light ochreous, with dark brown marginal spots pronounced. Tympanal organs (n = 7): Transverse ridge more or less regularly rounded. Tympanic pocket extending faintly beyond transverse ridge, rounded. Tympanic drum glomerular, not reaching transverse ridge. Male genitalia (n = 7) (Figs 19, 20): Uncus about 3 / 4 length of tegumen arms, downcurved; uncus arms basally with ventro-lateral tuft of setae; dorsal furrow pronounced medially with row of few setae on each side; thin, bifid on distal 1 / 5, slightly grooved, with apex slightly pointed; with shallow cavity ventro-apically. Gnathos arms connecting at 1 / 3 of length; shaft slightly downcurved, with apex pointing upward. Tegumen arms enlarging progressively toward uncus; tegumen connection about 1 / 3 arms length. Costa of valva basally narrow, with quadrangular projection, apically narrowing into arm pointing posterad with short tip curved inward; cucullus curved upward in distal 1 / 3, with apex rounded. Juxta triangular, regularly narrowing toward apex with shallow pockets projected ventro-laterally; with baso-lateral angles curved upward. Transtilla modified into two arms projecting posterad, slightly curved inward in distal 1 / 4, with longitudinal string of short spines ventrally at base, medially along arms, and at apex, increasing in size from base to apex in factor of about 1 to 4 - 5. Phallus almost straight, apex dorsally triangular; vesica basally covered with tiny spicules, microspicules barely visible all along vesica, also with row of 5 - 6 straight, short spine-like cornuti wider at their base. Female (n = 4): Labial palpi: 1.6 - 2.2 mm long. Forewing length 14 - 16 mm. Frenulum triple. Female genitalia (n = 4) (Fig. 38): Papillae anales straight, thick. Posterior apophyses 0.3 - 0.5 x length of papillae anales, wide at base, about half of length of papillae. Intersegmental membrane between segment VIII and IX covered with microspines. Sternite VIII laterally about 1 / 3 longer than tergite VIII. Sternite VIII formed by 2 lobes regularly narrowing downward into triangle, not connected ventrally, densely covered with spinules, with spinules longer ventrally. Anterior apophyses about 0.05 x length of papillae anales, quadrangular, anvil shaped. Anterior margin of sternite VIII latero-dorsally strongly sclerotized, thicker; posterior margin with dorsal line of setae. Sterigma membranous, covered with spinules. Ductus bursae regularly enlarging into corpus bursae, basally directed downward. Corpus bursae more or less rounded, faintly delimited from ductus bursae, with one oval signum.	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
C575A3423350B2EB56556E02A23B7179.taxon	distribution	Distribution. The species occurs in Brazil in the following states: Bahia, Espirito Santo, Parana, Rio de Janeiro, Santa Catarina, Sao Paulo (Fig. 45).	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
C575A3423350B2EB56556E02A23B7179.taxon	etymology	Etymology. The name comes from the latin coronatus, a, um: crowned, referring to the longitudinal string of short spines of the transtilla in the male genitalia.	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
B2474C9B6617ED1E9BF1ACA677F29A5C.taxon	diagnosis	Diagnosis. From Catharylla coronata and Catharylla tenellus, Catharylla serrabonita can be separated by the zigzagging median transverse line with the short triangular dent at CuA 2 and the pronounced creamy color of the hindwing. The male genitalia provide the best discriminant characters: in Catharylla serrabonita, the transtilla forms a pair of sclerotized arms bent inward in distal 1 / 4 and with a string of long spines of same length medially along it, whereas it forms a pair of short, narrow sclerotized arms with pointed tips projecting posterad, and with a pair of brushes directed medio-ventrally in Catharylla tenellus, and two sclerotized arms slightly bent inward distally, with a row of short spines increasing in size from base to apex in Catharylla coronata, and the juxta is apically narrow and pointed whereas it is triangular and regularly narrowed in Catharylla coronata and Catharylla tenellus. In female genitalia, the anterior angle of sternite VIII is projected anterad into a rounded protrusion covered with short spinules in Catharylla serrabonita, whereas it is projected downward in Catharylla tenellus and it is not projected in Catharylla coronata.	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
B2474C9B6617ED1E9BF1ACA677F29A5C.taxon	description	Description. Male (n = 21) (Fig. 6): Head white with ochreous chaetosemata. Antenna brown with whitish-ochreous scales and patch of brown scales at base. Maxillary palpus light ochreous, with patches of dark brown scales at 1 / 3 and 2 / 3, white tipped. Labial palpus: 1.7 - 2.5 mm long; light ochreous, white tipped. Thorax white, with ochreous patch at collar. Foreleg coxa white; femur white, dorsally dark brown; tibia and tarsomeres ochreous, distally ringed with brown. Midleg and hindleg white to light ochreous; tarsomeres II-V ochreous, brown on upperside, with white ringed tips. Forewing length: 10 - 14 mm; costal line ochreous; median transverse line ochreous to brown, zigzagging with short brown pronounced spot at M 1 and short triangular dent at CuA 2; subterminal transverse line ochreous to brown, regularly curved up to CuA 2, then curved again; R 5 faintly marked apically with ochreous; outer margin ochreous with 7 more or less triangular and connected dark brown spots between veins; fringes brass colored; underside ochreous, outer margin with pronounced spots. Hindwing cream-coloured, usually with more or less connected marginal brown spots between Sc + R 1, Rs, M 1, M 2, M 3, CuA 1 and CuA 2; fringes white; underside light ochreous, with marginal spots pronounced. Tympanal organs (n = 4): Transverse ridge more or less rounded, medially slightly flattened. Tympanic pocket extending faintly beyond transverse ridge, rounded. Tympanic drum glomerular, not reaching transverse ridge. Male genitalia (n = 4) (Figs 21 - 26): Uncus about as long as tegumen arms, downcurved; uncus arms connecting basally, with ventro-lateral tuft of setae at base; dorsal furrow pronounced medially with row of few hairs on each side; apex rounded, slightly indented medially, slightly convex ventro-apically. Gnathos arms connecting at 1 / 3; main shaft slightly downcurved with apex pointing upward. Tegumen arms regularly enlarging toward apex, connection at about 4 / 5 length of arms. Costa with apically rounded arm pointing postero-dorsally; cucullus curved upward in distal 1 / 3, with apex rounded. Juxta triangular, narrowing in distal 1 / 4 with bell-shaped ventro-lateral projections, regularly curved with apex horizontally straightened; baso-lateral angles curved upward. Transtilla with two very large sclerotized arms projecting posterad, bent inward in apical 1 / 4, with longitudinal string of long spines medially. Phallus slightly S-shaped, with apex dorsally sclerotized; vesica covered with microspicules, without cornuti. Female (n = 1): Labial palpi: 1.9 mm long. Forewing length: 14 mm. Frenulum triple. Female genitalia (n = 1) (Fig. 39): Papillae anales straight, thick. Posterior apophyses 0.4 x length of papillae anales, narrow, wider at base. Intersegmental membrane between segment VIII and IX covered with microspines. Sternite VIII laterally about 5 / 3 length of tergite VIII; posterior margin of tergite VIII with line of setae; sternite VIII forming 2 triangular lobes regularly narrowing downward, not connected, densely covered with short spinules of same length; anterior angle of sternite VIII slightly projected anterad, rounded, covered with short spinules of same length. Anterior apophyses 0.03 x length of papillae anales. Sterigma membranous, covered with spinules. Ductus bursae about 3 x length of corpus bursae, narrow, basally directed downward and then bent upward. Corpus bursae elongate, ovoid, with one tiny signum.	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
B2474C9B6617ED1E9BF1ACA677F29A5C.taxon	distribution	Distribution. The species occurs in Brazil (Bahia, Espirito Santo) (Figs 45 & 46).	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
B2474C9B6617ED1E9BF1ACA677F29A5C.taxon	etymology	Etymology. The name comes from that of the Serra Bonita Reserve founded by Vitor O. Becker and Clemira de Souza. It is managed by Instituto Uiracu in the State of Bahia, Brazil.	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
B6C5D5999FE042B1A94EB5B1179641C2.taxon	diagnosis	Diagnosis. The best discriminant characters externally between the two species of the mayrabonillae group are the shape of the forewing outer margin, which is slightly produced apically in Catharylla mayrabonillae and not produced in Catharylla paulella, and the forewing median transverse line with two strongly pronounced spots at 1 / 3 and 2 / 3 in Catharylla paulella, whereas these spots are lacking in Catharylla mayrabonillae. The hindwing of Catharylla mayrabonillae has a faded subterminal transverse line on costal half whereas the hindwing of Catharylla paulella lacks this marking. In male genitalia, the heavily sclerotized sacculus bears a dorso-lateral sclerotized string of short spines on distal 1 / 4 whereas the two processes of the costa are S-shaped in Catharylla paulella, and the apex of the phallus is trifid, rounded medially, shortly triangular laterally, whereas it is simply rounded in Catharylla paulella. In female genitalia, the sterigma forms double rounded cavities with a mustachio-shape arrangement of short spines in ventral view, and the ductus bursae is wide, progressively widening toward corpus in Catharylla mayrabonillae, whereas the sterigma forms a pair of shallow rounded pockets on each side of middle and the ductus bursae is narrow, with the rounded corpus bursae clearly differentiated from it in Catharylla paulella.	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
B6C5D5999FE042B1A94EB5B1179641C2.taxon	description	Description. Male (n = 17) (Fig. 7): Head with light ochreous chaetosemata. Antenna brown, with white scales dorsally and patch of dark brown scales at base. Maxillary palpus light ochreous, ringed with dark brown at 2 / 3; white tipped. Labial palpus: 1 - 1.4 mm; white, with patch of dark brown scales at 1 / 3 and 2 / 3 laterally. Thorax with patch of light ochreous scales at collar. Foreleg coxa whitish brown, femur white, dorsally ashen brown, tibia and tarsomeres ochreous, distally ringed with dark brown. Midleg femur white, tibia light ochreous, basally brown, tarsomeres II-V ochreous with tips ringed white. Hindleg white, except tarsomeres, as in midleg. Abdomen dull white. Forewing length: 7.5 - 8.5 mm; with apex slightly produced; costal line thin, ochreous or white in basal half, white in apical half; median transverse line ochreous, slightly undulated; subterminal transverse line ochreous; transverse lines enlarging into brown spot on costal margin with ochreous bar on costa following subterminal transverse line; terminal sector with light ochreous between veins, margin with thin, dark brown line from apex to CuA 1, with two dark brown spots in cubital sector, with spot between CuA 1 and CuA 2 slightly displaced toward base; fringes brass colored; underside light ochreous with some brownish scales, with thin brown margin. Hindwing white with thin transverse subterminal line faded ochreous, in continuity with forewing median transverse line; outer margin line pronounced, dark brown; underside dull white with thin faded brown margin; fringes white. Tympanal organs (n = 7): Transverse ridge regularly rounded, medially slightly flattened. Tympanic pockets broadly rounded, extended widely beyond transverse ridge, connected medially at base of praecinctorium. Tympanic drum bean-shaped, elongated, extended beyond tympanic pockets. Male genitalia (n = 7) (Figs 30, 31): Uncus thick and wide, about 2 / 5 length of tegumen arms, densely setose, with shortly projecting apex dorsally rounded. Gnathos reaching about 1 / 4 longer than uncus; arms wide, joining at 2 / 5 of length; distal 2 / 5 at angle of about 85 °. Tegumen almost regularly narrow, joined in last 1 / 4. Cucculus narrow, shorter than sacculus, apically rounded; sacculus greatly enlarged, thickly sclerotized, directed upward, then apically straight, slightly narrowing toward apex, laterally with string of short spines and 2 - 3 longer basal spines pointing downward; costal arm of valva directed upward, located at about 1 / 3 of costal margin, thin, strongly sclerotized, slightly curved. Vinculum arms narrow; saccus short and wide, tongue shaped, projecting posterad apically. Juxta elongate, distal 1 / 4 narrowed with rounded tip; wide base with ear-like lobes laterally and baso-lateral angle projected anterad. Phallus slightly bent sideways in distal 1 / 4, with trifid sclerotized apex rounded medially and shortly triangular laterally; vesica basally covered with tiny spicules, microspicules barely visible all along, with long spine-like, down-curved cornutus of about 2 / 5 length of phallus. Female (n = 37): Labial palpi length: 1.1 - 1.3 mm. Forewing length: 9.5 - 10.5 mm; frenulum triple. Female genitalia (n = 16) (Fig. 40): Papillae anales strongly curved in lateral view; sclerotized line along papillae expanding ventrally into triangle. Posterior apophyses 0.35 - 0.45 x length of papillae anales. Tergite VIII about 1 / 3 length of sternite VIII; postero-dorsal margin with few setae of moderate length; anterior apophyses 0.03 - 0.1 x papillae anales; sternite VIII with patches of minute setae antero-ventrally on each side of bare median band. Sterigma forming double rounded cavities with mustachio-shaped arrangement of short spines (in ventral view); remaining cavity wall with tiny spines. Ductus bursae short and wide, enlarged near middle; partly sclerotized on right side of enlargement and posterior section. Corpus bursae circular to elongate, about as long as tergite VII; single signum faintly pronounced.	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
B6C5D5999FE042B1A94EB5B1179641C2.taxon	distribution	Distribution. The species has been found so far in Panama, Costa Rica, Colombia, Venezuela, Guyana, Suriname, French Guiana, Ecuador, Peru and Brazil (Acre, Amazonas, Distrito Federal, Para, Rondonia) (Fig. 44). It is the most widespread species of Catharylla and the only one so far found in Central America and in Venezuela, Columbia, Ecuador and Peru.	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
B6C5D5999FE042B1A94EB5B1179641C2.taxon	etymology	Etymology. Catharylla mayrabonillae is named in honor of Ms. Mayra Bonilla of San Jose, Costa Rica, in recognition of her artistic portrayal of the biodiversity and ecosystems of Costa Rica and her many years of support for the existence of the rain forest in Area de Conservacion Guanacaste.	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
7FEE41A39E782EDF569CC181B49FB278.taxon	diagnosis	Diagnosis. This species can be easily separated from the other Catharylla species by the forewing median transverse line with two strongly pronounced spots at 1 / 3 and 2 / 3. The forewing is also sparkled with dark brown scales, which is unique in the genus. In male genitalia, the two S-like projections of the costal arm of the valva discriminate this species from the other species of Catharylla. In female genitalia, the sterigma forms a pair of shallow rounded pockets on each side of middle, and the ductus bursae is narrow, with the rounded corpus bursae clearly differentiated from it in Catharylla paulella, whereas it forms double rounded cavities with a mustachio-shape arrangement of short spines in ventral view, and the ductus bursae is wide, progressively widening toward corpus in Catharylla mayrabonillae.	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
7FEE41A39E782EDF569CC181B49FB278.taxon	distribution	Distribution. The species has been found in Brazil (Federal District, Maranhao, Mato Grosso, Para, Sao Paulo) and in Bolivia (Fig. 44).	en	Leger, Theo, Landry, Bernard, Nuss, Matthias, Mally, Richard (2014): Systematics of the Neotropical genus Catharylla Zeller (Lepidoptera, Pyralidae s. l., Crambinae). ZooKeys 375: 15-73, DOI: http://dx.doi.org/10.3897/zookeys.375.6222, URL: http://dx.doi.org/10.3897/zookeys.375.6222
