taxonID	type	description	language	source
D9AE2F5E87C0FB6DB7A89CDD5D6CD51A.taxon	diagnosis	Diagnosis. The following characteristics pertain to preserved specimens unless otherwise noted. A medium-sized Anomaloglossus differing from other species in the genus by the following combination of characters: (1) mean SVL in males 18.53 mm (18.15 - 18.86 mm, n = 3), mean SVL in females 19.15 mm (17.66 - 21.26, n = 5); (2) skin on dorsum shagreened, venter smooth; (3) tympanic annulus visible anteroventrally; (4) Fingers I and II subequal in length, FI = FII when fingers adpressed; (5) tip of Finger IV not surpassing the base of the distal subarticular tubercle on Finger III when fingers adpressed; (6) distal subarticular tubercle on Finger III and IV present; (7) Finger III swollen in males (conspicuous pre- and postaxial swelling in breeding males); (8) fringes on fingers absent; (9) toes basally webbed, fringes on toes absent; (10) tarsal keel well defined, slightly tubercle-like and weakly curved at proximal end; (11) black arm gland absent, glandular supracarpal pad present in both sexes (larger and more glandular in males); (12) cloacal tubercles absent; (13) pale paracloacal mark present; (14) in life, thin dorsolateral stripe present, from tip of snout to tip of urostyle (not visible, or only barely distinguishable in preservative); (15) ventrolateral stripe absent, but presence of irregular white blotches on the lower flank; (16) oblique lateral stripe absent; (17) sexual dichromatism in throat colour pattern: throat heavily pigmented with melanophores in males (dark brown to black in life), immaculate cream in females (yellowish-orange in life); (18) sexual dichromatism in ventral colour pattern: belly pigmented with melanophores in males, immaculate cream in females; (19) in life, iris metallic reddish bronze with fine dark brown reticulation; (20) large intestine extensively pigmented; (21) testes cream, unpigmented; (22) mature oocytes partly pigmented; (23) median lingual process small, longer than wide, tapered; (24) maxillary teeth present, small.	en	Kok, Philippe J. R., Nicolai, Michael P. J., Lathrop, Amy, MacCulloch, Ross D. (2018): Anomaloglossusmeansi sp. n., a new Pantepui species of the Anomaloglossusbeebei group (Anura, Aromobatidae). ZooKeys 759: 99-116, DOI: http://dx.doi.org/10.3897/zookeys.759.24742, URL: http://dx.doi.org/10.3897/zookeys.759.24742
D9AE2F5E87C0FB6DB7A89CDD5D6CD51A.taxon	description	Description of the holotype. Adult male (ROM 43896; Figure 3), 18.58 mm SVL, in suboptimal state of preservation (extensive ventral incisions, dorsal skin locally damaged); body robust; head as wide as long, HL 99.7 % of HW, HW 32 % of SVL; dorsal skin shagreened; ventral skin smooth; snout moderately long, SL 47 % of HL, 128 % of EL, round in dorsal view, protruding in lateral view, extending past lower jaw; nares located close to tip of snout, directed posterolaterally, visible from front, barely visible in dorsal and ventral views, EN 28 % of HL, 77 % of ED, EN 60 % of SL, TSL 49 % of SL; posterior rim of naris bordered posteriorly by an inconspicuous crescent-shaped ridge; IND 40 % of HW; canthus rostralis rounded; loreal region concave; IOD 104 % of EL, longer than upper eyelid; postrictal tubercles low and inconspicuous; tympanic membrane inconspicuous, round, concealed posterodorsally by a diffuse supratympanic swelling; tympanic annulus visible anteroventrally, TYM 52 % of EL; choanae small, circular, located anterolaterally. Maxillary teeth present, small. Tongue longer than wide, free posteriorly, with rounded margin, small median lingual process longer than wide, tapered. Vocal slits bilateral, large, extending from edge of tongue to angle of jaw. Forelimb swollen, robust, 94 % of FAL. Ulnar fold absent, metacarpal ridge absent; swollen, glandular supracarpal pad present, heavily pigmented with melanophores; hand moderate in size, 24 % of SVL, 75 % of HW; relative length of fingers III> II = I = IV; pre- and postaxial swelling on third finger (i. e. Finger III swollen); fingers without fringes; tip of Finger IV not reaching distal subarticular tubercle on Finger III when fingers adpressed; finger discs expanded, wider than long, about 1.4 times width of digit; width of disc on Finger III 0.52 mm; palmar tubercle large, egg shaped, 0.72 mm (larger than Finger III disc), thenar tubercle smaller, elliptical; one or two round to ovoid subarticular tubercles (one each on Fingers I and II, two each on Fingers III and IV, with distal tubercle on Finger IV less conspicuous). Hind limb robust, moderately long, with heel of adpressed leg reaching posterior corner of eye; skin granular with no cloacal tubercles discernible (but this could be an artefact of preservation); TL 46 % of SVL, heels not in contact when hind limbs are flexed at right angle to sagittal plane of body; FL 38 % of SVL; relative length of adpressed toes IV> III> V> II> I; Toe I very short, its tip barely reaching the base of subarticular tubercle of Toe II when adpressed; toe discs larger than width of toes; disc on Toe I only slightly larger than width of digit; width of disc on Toe IV 0.67 mm; toes basally webbed, lateral fringes absent; one to three round to ovoid subarticular tubercles (one each on Toes I and II, two each on Toes III and V, and three on Toe IV, with distal tubercle on Toe IV the smallest and least conspicuous). Inner metatarsal tubercle protuberant elliptical, 0.47 mm in length, outer metatarsal tubercle round, protuberant, pigmented, 0.35 mm in diameter. No medial metatarsal tubercle discernible. Tarsal keel slightly tubercle-like and weakly curved at proximal end, extending distally to preaxial edge of Toe I. Metatarsal fold not visible.	en	Kok, Philippe J. R., Nicolai, Michael P. J., Lathrop, Amy, MacCulloch, Ross D. (2018): Anomaloglossusmeansi sp. n., a new Pantepui species of the Anomaloglossusbeebei group (Anura, Aromobatidae). ZooKeys 759: 99-116, DOI: http://dx.doi.org/10.3897/zookeys.759.24742, URL: http://dx.doi.org/10.3897/zookeys.759.24742
D9AE2F5E87C0FB6DB7A89CDD5D6CD51A.taxon	description	Measurements of holotype (in mm). SVL = 18.58; HL = 5.91; HW = 5.93; IOD = 2.26; EN = 1.68; SL = 2.77; TSL = 1.35; EL = 2.16; TYM = 1.12; IND = 2.39; HAND I = 3.06; HAND II = 3.2; HAND III = 4.44; HAND IV = 3.09; WFD = 0.52; FAL = 3.81; THL = 7.80; TIL = 8.61; TAL = 4.48; FL = 7.08; WTD = 0.67.	en	Kok, Philippe J. R., Nicolai, Michael P. J., Lathrop, Amy, MacCulloch, Ross D. (2018): Anomaloglossusmeansi sp. n., a new Pantepui species of the Anomaloglossusbeebei group (Anura, Aromobatidae). ZooKeys 759: 99-116, DOI: http://dx.doi.org/10.3897/zookeys.759.24742, URL: http://dx.doi.org/10.3897/zookeys.759.24742
D9AE2F5E87C0FB6DB7A89CDD5D6CD51A.taxon	distribution	Distribution and natural history. The only localities documented for the new species are depicted in Figure 2. Specimens were collected in cloud forest (Figure 5), on the ground or low vegetation. Most were collected after nightfall, although one adult and one juvenile were collected during daylight. Specimens were collected on mountain flanks, not summits; at 1490 m on Ayanganna, and at 1234 m, 1371 m and 1411 m on Wokomung. The majority of specimens (eight) were collected at 1234 m on Wokomung. Fewer were collected at higher elevations; only one each at 1490 m on Ayanganna, 1371 m and 1411 m on Wokomung. This may have been because of habitat differences; high-canopy open forest at lower elevation and dense, low-canopy vegetation at higher elevations (see Discussion).	en	Kok, Philippe J. R., Nicolai, Michael P. J., Lathrop, Amy, MacCulloch, Ross D. (2018): Anomaloglossusmeansi sp. n., a new Pantepui species of the Anomaloglossusbeebei group (Anura, Aromobatidae). ZooKeys 759: 99-116, DOI: http://dx.doi.org/10.3897/zookeys.759.24742, URL: http://dx.doi.org/10.3897/zookeys.759.24742
D9AE2F5E87C0FB6DB7A89CDD5D6CD51A.taxon	etymology	Etymology. It is a great pleasure to name this new species after our friend and colleague D. Bruce Means, indefatigable explorer of the " islands in the sky ", and who collected one specimen of the new species and contributed with photographs and data. Thanks to his extensive fieldwork, Bruce Means greatly contributed to our understanding of the distribution, ecology, and taxonomy of Pantepui amphibians and reptiles. The specific epithet should be treated as a noun in the genitive case.	en	Kok, Philippe J. R., Nicolai, Michael P. J., Lathrop, Amy, MacCulloch, Ross D. (2018): Anomaloglossusmeansi sp. n., a new Pantepui species of the Anomaloglossusbeebei group (Anura, Aromobatidae). ZooKeys 759: 99-116, DOI: http://dx.doi.org/10.3897/zookeys.759.24742, URL: http://dx.doi.org/10.3897/zookeys.759.24742
D9AE2F5E87C0FB6DB7A89CDD5D6CD51A.taxon	discussion	Discussion. Although Ayanganna and Wokomung are close neighbours, the habitats on their slopes are not exactly similar. The slopes of Ayanganna are a series of relatively flat poorly drained plateaus alternating with steeper slopes. Collecting activities were concentrated on the plateaus, where the vegetation consists of dense, low-canopy high-tepui forest, with a dense understory of woody shrubs and large terrestrial bromeliads (MacCulloch and Lathrop 2009). The slopes of Wokomung have no large flat plateaus. Habitat at the collecting sites consists of well-drained slopes covered in lower montane cloud forest with some epiphytes and medium density understory, including scattered terrestrial bromeliads. Streams were common on the slopes (MacCulloch et al. 2006). The majority of specimens were found in this habitat, and this may indicate that Anomaloglossus meansi sp. n. prefers this to other habitat types; or is a reflection of collecting effort in these habitats. Species in the Anomaloglossus beebei group are currently only known from east of the Rio Caroni, in the Eastern Pantepui District. Anomaloglossus rufulus is restricted to the highlands of the eastern part of the Chimanta Massif in Venezuela, whereas A. kaiei has a rather large distribution in the uplands of the Pakaraima Mountains of Guyana (Figure 2). The sister species A. roraima and A. beebei are allopatric, A. roraima being restricted to the highlands of the eastern tip of the Eastern Tepui Chain, whereas A. beebei is reported further to the east in the uplands of Kaieteur National Park, the Wokomung Massif and Mount Ayanganna (Figure 2). A similar spatial distribution is detected in the sister species A. praderioi and A. meansi sp. n., which are also allopatric, with A. praderioi reported from the uplands of the Eastern Tepui Chain, whereas A. meansi sp. n. is only known further to the east in the uplands of Mount Wokomung and Mount Ayanganna. As mentioned above, virtually nothing is known about the ecology of A. meansi sp. n. Based on its phylogenetic position it is likely this species has an exotrophic, lentic tadpole (Figure 1). Comprehensive ecological data are crucial for the assessment of species conservation status, but these assessments are known for a few species only in the Pantepui region and there is a high risk that population declines remain unnoticed in such remote areas. Two additional phylogenetically distinct species of Anomaloglossus remain to be described in the megacephalus group (see Grant et al. 2017; the authors, in progress), but several locally restricted Anomaloglossus species probably await discovery (Vacher et al. 2017).	en	Kok, Philippe J. R., Nicolai, Michael P. J., Lathrop, Amy, MacCulloch, Ross D. (2018): Anomaloglossusmeansi sp. n., a new Pantepui species of the Anomaloglossusbeebei group (Anura, Aromobatidae). ZooKeys 759: 99-116, DOI: http://dx.doi.org/10.3897/zookeys.759.24742, URL: http://dx.doi.org/10.3897/zookeys.759.24742
