taxonID	type	description	language	source
6B7C7B58FFACFFB2FF3CCDFEC98AFCD0.taxon	materials_examined	Type species Platycoelus depressus Blanchard 1853: 25 = Platycaelus Blanchard, 1843; Plate 2 image 10. Emberson (1993) lists dates of publication of individual plates. = Platycoelus Blanchard, 1853	en	Will, Kipling (2015): A taxonomic review, new species and a key to species of Platycoelus Blanchard, 1843 (Coleoptera: Carabidae: Pterostichini). Zootaxa 4034 (2): 291-308, DOI: 10.11646/zootaxa.4034.2.3
6B7C7B58FFACFFB2FF3CCDFEC98AFCD0.taxon	description	Description. Head. Clypeo-ocular sulci not impressed, punctiform or more elongate, straight, very broadly and very shallowly impressed; mentum moderately or very shallowly emarginate, sides divergent, paramedial pits small or large, deeply or very deeply impressed; median tooth bifid or broadly emarginate; paraglossae small, without elongate setae at apex; ligular sclerite with two setae on apical margin; maxillary palpifer with one basal seta; antennae filiform, with three basal segments glabrous. Thorax. Pronotum quadrate or transverse, width across base slightly broader than or equal to apex, two marginal setae; pro-, meso- and metasterna glabrous; proepisternum with or without very scattered punctulae; elytra free along suture, border at base complete, interrupted or effaced, nine distinctly impressed complete striae or striae 1 – 7 not impressed on elytral disc, apicolateral plica visible, parascutellar stria long, distinctly impressed, connected or not connected to stria 1, angular base of stria 1 present, parascutellar punctues at base of stria 2, no discal punctures, intervals flat or slightly convex; flight wing full length or very short; anterior tarsi of male with three basal segments expanded, ventrally squamous. Abdomen. Ventrites 3 – 6 without sulci; aedeagus ostium dorsal, median lobe oriented left side up in repose; parameres attenuate with long narrow apex or broad and apically pointed, both nearly of equal length, left wider than right; female reproductive tract (Fig. 9) with dorsolateral bursal lobe, elongate spermatheca broadly attached laterally at base of bursal lobe or spermatheca relatively narrow, very elongate and attached apically on dorsal lobe, with appended gland attached near base of spermatheca, spermatheca without digitiform diverticulum near base, without spermathecal gland duct diverticulum. Differential diagnosis. The following combination of character states distinguishes members of the genus from other pterostichines in the Australian region: Antennae pubescent from antennomere 4; mental tooth entire; stipes with a single setae near base; elytra usually with eight more or less impressed striae; when impressed the parascutellar stria is present and intervals are of more or less equal width; abdominal ventrites without transverse sulci; and the length between hind margin of mesocoxa and anterior margin of metacoxa equal to or greater than diameter of mesocoxa. The male genitalia and female reproductive tract share similarities that may prove synapomorphic with Delinius Westwood, 1864, Euryabax Fauvel, 1903, and Sphodrosomus Perroud and Montrouzier, 1864. Nomenclatural note. Bousquet (2002), overturned the spelling that was uniformly in use " Platycoelus, " to " Platycaelus, " based on what appears in the plate caption initially published by Blanchard (1843, see Emberson 1993). However, Bousquet's action was contrary to the 2000 Code (ICZN 1999), which states explicitly: " 33.2.3.1. when an unjustified emendation is in prevailing usage and is attributed to the original author and date it is deemed to be a justified emendation. " Therefore Platycoelus, as of the year 2000, was and remains the correct Codecompliant spelling as found in the catalogs by Lorenz (1998, 2005). Subsequent publications that used Platycaelus (Will & Kavanaugh 2012; Will 2011) following the spelling used by Bousquet (2002), are incorrect.	en	Will, Kipling (2015): A taxonomic review, new species and a key to species of Platycoelus Blanchard, 1843 (Coleoptera: Carabidae: Pterostichini). Zootaxa 4034 (2): 291-308, DOI: 10.11646/zootaxa.4034.2.3
6B7C7B58FFAFFFB2FF3CCAF0CF1BF81A.taxon	materials_examined	Type locality. Queensland, Southwood National Park. Vicinity of 27 ° 50.2 " S 150 ° 06.7 " E.	en	Will, Kipling (2015): A taxonomic review, new species and a key to species of Platycoelus Blanchard, 1843 (Coleoptera: Carabidae: Pterostichini). Zootaxa 4034 (2): 291-308, DOI: 10.11646/zootaxa.4034.2.3
6B7C7B58FFAFFFB2FF3CCAF0CF1BF81A.taxon	description	Description. Dorsal habitus (Fig. 1 C). Size. Overall length (sbl) 16.2 mm; greatest width over elytra 6.8 mm. Color. Dorsal and ventral surfaces black; legs, mouthparts, and antennae black. Luster. Dorsally and ventrally moderately shiny. Iridescence. Elytra with obvious spectral iridescence; pronotum slightly iridescent; ventral surface of body without iridescence. Head. Dorsal microsculpture with microlines not visible at 50 x magnification. Frons punctulate, more densely and coarsely near base of head. Clypeal-ocular sulci represented by broad, shallow poorly defined impressions. Ocular ratio 1.62; eyes large, rounded. Labrum very slightly emarginate with the medial four setae broadly distributed, width from the outermost medial seta to the lateral seta subequal, slightly wider than width between medial setae. Mentum long, deeply emarginate prominent lateral lobes, with one pair small, round, deep pits; median tooth prominent, bifid; one pair of setae positioned laterad of median tooth. Gula narrow, width at middle about as wide width of mentum emargination, anterior tentorial pits small, punctiform. Antennae, overall length long, antennomeres 10 – 11 extended beyond pronotal base, antennomeres 5 – 11 elongate. Thorax. Pronotum slightly transverse, sides evenly and shallowly rounded from apex to base. Marginal bead continuous from apex to base; basal margin bordered; anterior angles scarcely produced; hind angles nearly right angled and rounded; inner basal impression well impressed, broad, linear; outer impression shallow, round impressions. Seta at hind angle touches basal bead and one pore width from lateral bead. Microsculpture of disc not visible at 50 x magnification, very shallowly and sparsely punctulate, much denser near hind angles. Elytral striae complete, sharply impressed. Elytral microsculpture hardly visible at 50 x, formed as transverse mesh of microlines. Intervals with dense micropunctulae throughout. Profemur in males and females unmodified. Metacoxal sulcus straight, extended to lateral end of coxa. Prosternal process at apex rounded and not margined. Sterna with micropunctulae throughout. Abdomen. Abdominal ventrites smooth. Male genitalia. Aedeagus (Fig. 5 A, B) abruptly curved ventrally in apical quarter.	en	Will, Kipling (2015): A taxonomic review, new species and a key to species of Platycoelus Blanchard, 1843 (Coleoptera: Carabidae: Pterostichini). Zootaxa 4034 (2): 291-308, DOI: 10.11646/zootaxa.4034.2.3
6B7C7B58FFAFFFB2FF3CCAF0CF1BF81A.taxon	etymology	Etymology. The specific epithet derived from the Brigalow (Acacia harpophylla) forest at the type locality, is a Latinized noun, nominative case. This habitat type has largely been cleared and habitat loss may be a contributing factor in the apparent rarity of this beetle.	en	Will, Kipling (2015): A taxonomic review, new species and a key to species of Platycoelus Blanchard, 1843 (Coleoptera: Carabidae: Pterostichini). Zootaxa 4034 (2): 291-308, DOI: 10.11646/zootaxa.4034.2.3
6B7C7B58FFAEFFB4FF3CCF60CE82FB00.taxon	materials_examined	Type locality. Queensland, Iron Range National Park, vicinity of 12 ° 42 ' 50 " S / 143 ° 17 ' 14 " E.	en	Will, Kipling (2015): A taxonomic review, new species and a key to species of Platycoelus Blanchard, 1843 (Coleoptera: Carabidae: Pterostichini). Zootaxa 4034 (2): 291-308, DOI: 10.11646/zootaxa.4034.2.3
6B7C7B58FFAEFFB4FF3CCF60CE82FB00.taxon	description	Description. Dorsal habitus (Fig. 1 E). Size. Overall length (sbl) 9.6 – 13.5 mm; greatest width over elytra 3.5 – 4.2 mm. Color. Dorsal and ventral surfaces black. Legs, mouthparts, and antennae slightly paler piceous to rufopiceous. Luster. Dorsally and ventrally very shiny. Iridescence. Elytra and ventral surface of body with strong spectral iridescence. Head. Dorsal microsculpture with microlines very shallow, scarcely visible at 50 x magnification; sculpticells isodiametric. Frons smooth, impunctate. Clypeal-ocular sulci represented by a punctiform impression near clypeus. Ocular ratio 1.30 – 1.70; eyes large size, rounded. Labrum slightly emarginate with the medial four setae narrowly distributed, width from outermost medial seta to lateral seta subequal to width across all four medial setae. Mentum short, shallowly emarginate, lateral lobes little produced, with one pair very large, round, very deep pits; median tooth short, very broad, shallowly emarginate; one pair of setae below median tooth. Gula moderately wide, width at middle about one half the width of mentum, anterior tentorial pits very large. Antennae, overall length long, antennomeres 10 – 11 extended to beyond base, antennomeres 5 – 11 elongate. Thorax. Pronotum slightly transverse, broadly and distinctly reflexed near hind angles, sides evenly rounded from apex to base or slightly sinuate from about one fifth of the pronotum length forward of base. Sinuation of lateral margins variable, but not markedly sinuate. Marginal bead continuous from apex to about one fifth pronotal length from base or nearly extended to base; basal margin not bordered; anterior angles scarcely produced; hind angles obtuse and rounded; inner basal impression absent or a shallow linear impression; outer impression absent. Seta at hind angle set about 4 x pore widths forward of hind angles. Microsculpture of disc not visible at 50 x magnification. Elytral striae complete or striae 5 – 7 shallow or not impressed in basal 2 / 3 and / or all striae shallow or scarcely impressed in apical half of disc; all striae well impressed and impunctate near apex and distinctly crenulate in basal third to half. Elytral microsculpture hardly visible at 50 x, formed as transverse mesh of microlines. Profemur in males with deep, broad transverse sulcus (Fig. 10). Female profemur unmodified. Metacoxal sulcus straight, extended to lateral end of coxa. Prosternal process at apex rounded and not margined. Sterna smooth throughout. Abdomen. Abdominal ventrites smooth, glabrous. Male genitalia. Aedeagus (Fig. 5 E, F) abruptly curved ventrally in apical quarter.	en	Will, Kipling (2015): A taxonomic review, new species and a key to species of Platycoelus Blanchard, 1843 (Coleoptera: Carabidae: Pterostichini). Zootaxa 4034 (2): 291-308, DOI: 10.11646/zootaxa.4034.2.3
6B7C7B58FFAEFFB4FF3CCF60CE82FB00.taxon	etymology	Etymology. The specific epithet is a Latinized noun in the genitive case from the name Chong Hee Will, my wife, whose generous support enabled the collection of the bulk of the type material, DNA quality material, and specimens of many other Australian carabids in addition to this new species.	en	Will, Kipling (2015): A taxonomic review, new species and a key to species of Platycoelus Blanchard, 1843 (Coleoptera: Carabidae: Pterostichini). Zootaxa 4034 (2): 291-308, DOI: 10.11646/zootaxa.4034.2.3
6B7C7B58FFA9FFB5FF3CCC0DCCAAFD88.taxon	materials_examined	Type locality. Papua New Guinea, Sandaun Province, Aitape. In the vicinity of 3 ° 8 ' S 142 ° 21 ' E.	en	Will, Kipling (2015): A taxonomic review, new species and a key to species of Platycoelus Blanchard, 1843 (Coleoptera: Carabidae: Pterostichini). Zootaxa 4034 (2): 291-308, DOI: 10.11646/zootaxa.4034.2.3
6B7C7B58FFA9FFB5FF3CCC0DCCAAFD88.taxon	description	Description. Dorsal habitus (Fig. 2 A). Size. Overall length (sbl) 8.8 mm, greatest width over elytra 3.2 mm. Color. Dorsal and ventral surfaces black. Legs, mouthparts, and antennae piceous. Luster. Dorsally and ventrally shiny. Iridescence. Elytra and ventral surface of body with pronounced spectral iridescence. Head. Dorsal microsculpture not visible at 50 x magnification. Frons with dense, irregularly placed and variously sized punctulae. Clypeal-ocular sulci represented by a broad, very shallow impression. Ocular ratio 1.53; eyes large size, rounded. Labrum very slightly emarginate with the medial four setae moderately widely distributed, width from the outermost medial seta to lateral seta one half width across all four medial setae. Mentum short, shallowly emarginate, lateral lobes little produced, with one pair very large, round, very deep pits; median tooth short, very broad, shallowly emarginate; one pair of setae below median tooth. Gula very wide, width at middle about two thirds the width of mentum, anterior tentorial pits small, deep. Antennae, damaged so overall length unknown, antennomeres 5 – 9 elongate. Thorax. Pronotum slightly transverse, sides very shallowly rounded from apex to just anterad base, then straightening and very slightly sinuate to base. Marginal bead continuous from apex to base; basal margin not bordered; anterior angles not produced; hind angles obtusely angled, rounded; posterior impression not present, lateral area with a very broad, flat region to lateral margin; this region not or scarcely delimited medially, medially convex, not reflexed near hind angles. Seta at hind angle set at hind angles, touches lateral bead. Microsculpture of disc not visible at 50 x magnification. Elytral striae complete, all striae very shallowly impressed and minutely crenulate, except near apex. Elytral microsculpture hardly visible at 50 x, formed as an irregular or nearly isodiametric mesh of microlines. Intervals 1 – 7 with micropunctulae; 8 – 9 and across apex of elytron punctures denser and sculpturing more pronounced and irregular. Profemur in male and female unmodified. Metacoxal sulcus straight, extended to lateral end of coxa. Prosternal process at apex rounded and not margined. Sterna smooth throughout. Abdomen. Abdominal ventrites smooth, glabrous. Male genitalia. Aedeagus (Fig. 5 I, J) sharply flexed ventrally at tip, which is bluntly rounded.	en	Will, Kipling (2015): A taxonomic review, new species and a key to species of Platycoelus Blanchard, 1843 (Coleoptera: Carabidae: Pterostichini). Zootaxa 4034 (2): 291-308, DOI: 10.11646/zootaxa.4034.2.3
6B7C7B58FFA9FFB5FF3CCC0DCCAAFD88.taxon	etymology	Etymology. The specific epithet is a Latinized noun, nominative case, based on the name of the messenger of the Greek pantheon, Hermes sp. nov., in allusion to his winged helm and boots and the fully developed flight wings characteristic of this species.	en	Will, Kipling (2015): A taxonomic review, new species and a key to species of Platycoelus Blanchard, 1843 (Coleoptera: Carabidae: Pterostichini). Zootaxa 4034 (2): 291-308, DOI: 10.11646/zootaxa.4034.2.3
6B7C7B58FFABFFB6FF3CC9A1CC77F855.taxon	materials_examined	Type locality. Kowanyama, Queensland. Estimated coordinates: 15 ° 28 ' 44 " S 141 ° 15 ' 30 " W.	en	Will, Kipling (2015): A taxonomic review, new species and a key to species of Platycoelus Blanchard, 1843 (Coleoptera: Carabidae: Pterostichini). Zootaxa 4034 (2): 291-308, DOI: 10.11646/zootaxa.4034.2.3
6B7C7B58FFABFFB6FF3CC9A1CC77F855.taxon	description	Description. Dorsal habitus (Fig. 3 A). Size. Overall length (sbl) 12.8 – 15.5 mm, greatest width over elytra 3.8 – 4.3 mm. Color. Dorsal and ventral surfaces black. Legs, mouthparts, and antennae piceous. Luster. Dorsally and ventrally shiny. Iridescence. Elytra and ventral surface of body with pronounced spectral iridescence. Head. Dorsal microsculpture not visible at 50 x magnification. Frons with dense, irregularly placed fine punctulae. Clypealocular sulci linear, very shallowly and broadly impressed near clypeus. Ocular ratio 1.81 – 1.83, eyes large, prominent. Labrum truncate with setae equally distributed along width. Mentum long, deeply emarginate prominent lateral lobes, with one pair small, round, deep pits; median tooth prominent, broadly emarginate-bifid; one pair of setae positioned laterad of median tooth. Gula narrow, width at middle about as wide width of mentum emargination, anterior tentorial pits small, punctiform. Antennae, overall length long, antennomeres 9 – 11 reaching beyond pronotal base, antennomeres 5 – 11 elongate. Thorax. Pronotum transverse, sides evenly rounded from apex to base, without sinuation; base wider than width across anterior angles. Marginal bead wide, continuous from apex to base, basal margin bordered along lateral third, anterior angles not produced, hind angles rounded, slightly obtuse, inner basal impression shallowly impressed, broad, linear; outer impression shallow, elliptical impressions. Seta near hind angle in contact with or just apical of basal bead and one pore width from lateral bead. Microsculpture of disc not visible at 50 x magnification. Elytral striae complete, sharply impressed. Elytral microsculpture hardly visible at 50 x, formed as transverse mesh of microlines. Intervals with dense micropunctulae throughout. Profemur in males and females unmodified. Metacoxal sulcus straight, reaching lateral end of coxa. Prosternal process at apex rounded and not margined. Sterna with micropunctulae throughout. Abdomen. Abdominal ventrites punctate. Male genitalia. Aedeagus (Fig. 6 G, H) abruptly curved ventrally in apical quarter.	en	Will, Kipling (2015): A taxonomic review, new species and a key to species of Platycoelus Blanchard, 1843 (Coleoptera: Carabidae: Pterostichini). Zootaxa 4034 (2): 291-308, DOI: 10.11646/zootaxa.4034.2.3
6B7C7B58FFABFFB6FF3CC9A1CC77F855.taxon	etymology	Etymology. A Latinized noun in the nominative case from the name Orion Will, my son and my field assistant on the trip where the paratype male specimen of this species, and many other important carabid specimens, were collected.	en	Will, Kipling (2015): A taxonomic review, new species and a key to species of Platycoelus Blanchard, 1843 (Coleoptera: Carabidae: Pterostichini). Zootaxa 4034 (2): 291-308, DOI: 10.11646/zootaxa.4034.2.3
6B7C7B58FFAAFFBCFF3CC9A1CFF1FABF.taxon	materials_examined	Type locality. Queensland, Camp Milo, Cooloola National Park. Estimated coordinates 25 ° 59 ' 45 " S 153 ° 4 ' 14 " E.	en	Will, Kipling (2015): A taxonomic review, new species and a key to species of Platycoelus Blanchard, 1843 (Coleoptera: Carabidae: Pterostichini). Zootaxa 4034 (2): 291-308, DOI: 10.11646/zootaxa.4034.2.3
6B7C7B58FFAAFFBCFF3CC9A1CFF1FABF.taxon	description	Description. Dorsal habitus (Fig. 4 B). Size. Overall length (sbl) 13.2 – 15.7 mm, greatest width over elytra 4.1 – 4.9 mm. Color. Dorsal and ventral surfaces black or piceous. Legs, mouthparts, and antennae piceous. Luster. Dorsally and ventrally moderately shiny. Iridescence. Elytra with obvious spectral iridescence; pronotum without iridescence; ventral surface of body slightly iridescent. Head. Dorsal microsculpture with microlines not visible at 50 x magnification. Frons minutely and sparsely punctulate, more densely near base of head. Clypeal-ocular sulci impressed, linear, parallel near clypeus and divergent on frons. Ocular ratio 1.64 – 1.75, eyes large, prominent. Labrum truncate with setae equally distributed along width. Mentum long, deeply emarginate prominent lateral lobes, with one pair small, round, deep pits; median tooth prominent, bifid; one pair of setae positioned laterad of median tooth. Gula narrow, width at middle about as wide width of mentum emargination, anterior tentorial pits small, punctiform. Antennae, overall length long, antennomeres 10 – 11 reaching beyond pronotal base, antennomeres 5 – 11 elongate. Thorax. Pronotum slightly transverse, nearly quadrate; sides evenly and very shallowly rounded from apex to base, without sinuation; base wider than width across anterior angles. Marginal bead wide, continuous from apex to base; basal margin bordered, sometimes interrupted medially, anterior angles not produced, hind angles rounded, very slightly obtuse and nearly right angled, inner basal impression shallowly impressed, linear; outer impression very shallow, broad and not well defined onto the slightly reflexed basolateral region. Seta near hind angle in contact with or just apical of basal bead and two pore widths from lateral bead. Microsculpture of disc not visible at 50 x magnification. Elytral striae complete, sharply impressed. Elytral microsculpture hardly visible at 50 x, formed as transverse mesh of microlines. Profemur in males and females unmodified. Metacoxal sulcus straight, reaching lateral end of coxa. Prosternal process at apex rounded and not margined. Sterna smooth throughout or with very shallow micropunctulae laterally. Abdomen. Abdominal ventrites smooth. Male genitalia. Aedeagus (Fig. 7 C, D) very slightly curved ventrally in apical quarter.	en	Will, Kipling (2015): A taxonomic review, new species and a key to species of Platycoelus Blanchard, 1843 (Coleoptera: Carabidae: Pterostichini). Zootaxa 4034 (2): 291-308, DOI: 10.11646/zootaxa.4034.2.3
6B7C7B58FFAAFFBCFF3CC9A1CFF1FABF.taxon	etymology	Etymology. The specific epithet is a the Latin word for smooth or polished. In allusion to the smooth proepisternum characteristic of this species.	en	Will, Kipling (2015): A taxonomic review, new species and a key to species of Platycoelus Blanchard, 1843 (Coleoptera: Carabidae: Pterostichini). Zootaxa 4034 (2): 291-308, DOI: 10.11646/zootaxa.4034.2.3
