taxonID	type	description	language	source
6B7E007DFFBFF072D8E8FAF59F65BC12.taxon	diagnosis	Diagnosis. Recognized by the following characters: Head, thorax, and abdomen black to brownish black. Antennae dark brown; segments III and IV pale yellow. Rostrum dark brown; all of segment III and basal segment of IV pale yellow. Hemelytra dark brown with clear central area. Legs dark brown; fore – and mid tibiae and tarsi pale yellow; hind legs entirely dark brown with pale yellow tarsi. Copulatory tube located mesally at sternum VII and and somewhat separated from base of ovipositor.	en	Yamada, Kazutaka, Hirowatari, Toshiya, Susila, Wayan, Okajima, Shûji (2007): The flower bug genus Montandoniola Poppius (Hemiptera: Heteroptera: Anthocoridae) from Indonesia, with descriptions of two new species. Zootaxa 1604: 37-45, DOI: 10.5281/zenodo.178743
6B7E007DFFBFF072D8E8FAF59F65BC12.taxon	materials_examined	Material examined. INDONESIA: 1 Ψ (Fig. 1), Java, Yogyakarta, Gadjah Mada Univ., 17. viii. 2005, T. Ishikawa.	en	Yamada, Kazutaka, Hirowatari, Toshiya, Susila, Wayan, Okajima, Shûji (2007): The flower bug genus Montandoniola Poppius (Hemiptera: Heteroptera: Anthocoridae) from Indonesia, with descriptions of two new species. Zootaxa 1604: 37-45, DOI: 10.5281/zenodo.178743
6B7E007DFFBFF072D8E8FAF59F65BC12.taxon	distribution	Distribution. Europe, North and Central Africa, Asia (Israel, India, Indonesia, Philippines, China, Japan), Australia, Micronesia, U. S. A.	en	Yamada, Kazutaka, Hirowatari, Toshiya, Susila, Wayan, Okajima, Shûji (2007): The flower bug genus Montandoniola Poppius (Hemiptera: Heteroptera: Anthocoridae) from Indonesia, with descriptions of two new species. Zootaxa 1604: 37-45, DOI: 10.5281/zenodo.178743
6B7E007DFFBFF072D8E8FDF79967BFA4.taxon	discussion	Comments. In addition to generic characters described by Poppius (1909), Postle et al. (2001) indicate that Montandoniola is defined by the following characters: Body> 2.0 mm, cylindrical; head in front of eyes longer than diameter of eye; hind wing with two or three veins, hamus remote from junction of Cu and m – cu. The two Indonesian new species described below also possess these characters, except for their relatively small body size (1.90 – 2.20 mm in M. ishikawai; 1.50 – 1.80 mm in M. bellatula). The two new species were collected from the flowers of evergreen trees, Macaranga spp. (Euphorbiaceae) by sweeping, together with many tiny thrips (Ishikawa, pers. comm.). Montandoniola moraguesi is a well – known biological control agent of the Cuban laurel thrips, Gynaikothrips ficorum (Marchal) (e. g., Tawfik & Nagui 1965; Lattin 2000). However, this anthocorid has been reported to feed on more than 20 species of gall – forming thrips from a wide variety of host plants (Muraleedharan & Ananthakrishnan 1978; Dobbs & Boyd 2006). Therefore, it is inferred that the species of Montandoniola are predators on thrips. The two new Indonesian species also possibly prey on thrips in the flowers of Macaranga spp.	en	Yamada, Kazutaka, Hirowatari, Toshiya, Susila, Wayan, Okajima, Shûji (2007): The flower bug genus Montandoniola Poppius (Hemiptera: Heteroptera: Anthocoridae) from Indonesia, with descriptions of two new species. Zootaxa 1604: 37-45, DOI: 10.5281/zenodo.178743
6B7E007DFFBCF076D8E8F98F9E3EBB94.taxon	diagnosis	Diagnosis. Recognized by the following characters: Body (Fig. 2) much slenderer than congeners. Antennal segment III (Figs. 2, 6, 7) black except for pale yellow base and apex. Hemelytra (Fig. 2) black to brownish black except for off – white on outer half of clavi, almost all of embolium, and part along embolial suture in corium; membrane semi – transparent, centrally with blackish stripes. Pygophore (Fig. 13) turbinate; flagellum (Figs. 13 – 15) basally bifurcate, nearly straight. Copulatory tube (Fig. 16) noticeably elongate, bulbous at apical 1 / 3.	en	Yamada, Kazutaka, Hirowatari, Toshiya, Susila, Wayan, Okajima, Shûji (2007): The flower bug genus Montandoniola Poppius (Hemiptera: Heteroptera: Anthocoridae) from Indonesia, with descriptions of two new species. Zootaxa 1604: 37-45, DOI: 10.5281/zenodo.178743
6B7E007DFFBCF076D8E8F98F9E3EBB94.taxon	description	Description. Coloration. Body (Fig. 2) generally black, shiny on dorsal and ventral surfaces. Head, thorax, and abdomen uniformly black. Eyes and ocelli reddish black. Antennae (Figs. 2, 6, 7) black to brownish black except for pale yellow base and apex of segment III in male, entirely pale yellow segment III in female; segment IV apically tinged with reddish black. Rostrum uniformly black. Hemelytra (Fig. 2) black to brownish black except for off – white on outer half of clavi, almost of embolium, and part along embolial suture in corium; membrane semi – transparent, medially with blackish stripe. Legs generally black; fore tibia pale yellow except for black at base; mid – and hind tibiae mostly black; tarsi pale yellow, gradually darkened toward apex. Structure. Body (Fig. 2) elongate, much slenderer than the other congeners. Head (Figs. 2, 5) elliptical, smooth on dorsal surface, about 1.2 times as long as head width across eyes, sparsely clothed with silky, short, recumbent setae; pairs of long, erect setae laterally on tylus, on anterior part of inner margin of eye, and between eye and ocellus; ante-ocular region about 1.2 times as long as length of eye in dorsal view; vertex about 2.0 times as wide as eye in dorsal view; postocular portion distinctly long, not constricted; eyes with short setae; ocelli small. Antennae (Figs. 6, 7) densely clothed with short recumbent and long suberect setae; long suberect setae slightly longer than or as long as width of segment; segment I almost reaching apex of head, sparsely clothed with short setae; segment II cylindrical in male, slightly thickened toward apex, as long as head width across eyes; segment III approximately 0.57 times as long as segment II; segment IV weakly flattened, approximately 1.4 times as long as segment III. Rostrum stout, not reaching fore coxae, sparsely clothed with silky, short, erect setae; segment II thicker than the other segments; segment III about 4.0 times as long as segment II; segment IV about 0.6 times as long as segment III. Pronotum (Figs. 2, 5) trapezoidal, highly polished, impunctate, sparsely distributed with silky, short, recumbent setae, bearing four distinct corner setae and a pair of long erect setae at anterior angle of callus; anterior margin nearly straight, slightly longer than mesal length; lateral margin weakly curved and strongly carinate at anterior 2 / 3; posterior margin shallowly concave, approximately 2.2 times as wide as anterior margin; collar long, weakly rugose, with sparse, silky, short setae; posterior half of pronotum weakly depressed, with transverse rugae. Scutellum (Fig. 2) levigate, extremely swollen on anterior half and depressed on posterior half, with a pair of long, erect setae on basal 1 / 4 of lateral margin. Hemelytra (Fig. 2) narrowed toward apex, impunctate, with poorly distributed, silky, short, recumbent setae; apical part of corium approximately 2.5 times as wide as that of embolium; cuneal margin about 0.75 times as long as embolial margin; cuneus with short setae at apex. Ostiolar peritreme (Fig. 11) accompanied with narrow, smooth, shining area anterior to gutter and broad alutaceous area posterior to gutter; outer margin of ostiolar peritreme posterolaterally a little raised above level of surrounding evaporative area; smooth shining area slightly broader than that of M. bellatula; evaporative area subtriangular. Legs densely covered with yellow, recumbent setae; fore tibia slightly thickened toward apex, furnished with a row of 18 – 20 short teeth along entire ventral surface in male; hind tibiae weakly depressed; hind coxae separated. Abdomen clothed with rows of short setae on ventral surface of each segments; seventh and eighth sterna laterally with pairs of long, suberect setae; scissure on abdominal tergite reaching end of posterior margin of third segment. Male genitalia (Figs. 13 – 15): Pygophore (Fig. 13) turbinate, posteroventrally covered with 8 – 10 long and stout setae; paramere (Figs. 14, 15) broadened in general shape; cone (Figs. 14, 15) wide, acute at apex in dorsal view, abruptly elevated at apex in lateral view; flagellum (Figs. 14, 15) basally bifurcate, long, nearly straight, tapering, approximately 1.5 times as long as width of cone. Female genitalia (Fig. 16): Copulatory tube (Fig. 16) mesally located at sternum VII, very close to base of ovipositor, elongate, accompanied by strongly sclerotized basal part leading into membranous and blossom – like apex; strongly sclerotized basal part bulbous at anterior 1 / 3. Measurements [% (n = 8) / Ψ (n = 2)]. Body length 1.90 – 2.20 / 2.10 – 2.15; head length 0.35 – 0.38 / 0.38 – 0.40; head width across eyes 0.28 – 0.30 / 0.28 – 0.30; vertex width 0.16 – 0.18 / 0.18 – 0.19; width between ocelli 0.11 – 0.13 / 0.13 – 0.14; length of antennal segments I – IV: 0.10 – 0.12 / 0.11 – 0.12, 0.26 – 0.28 / 0.27, 0.15 – 0.16 / 0.15 – 0.16, 0.20 – 0.24 / 0.22 – 0.25; length of last three rostral segments II – IV: 0.06 – 0.09 / 0.07 – 0.09, 0.27 – 0.29 / 0.27 – 0.29, 0.17 – 0.19 / 0.20; anterior pronotal width 0.26 – 0.29 / 0.29 – 0.31; mesal pronotal length 0.23 – 0.29 / 0.28 – 0.30; basal pronotal width 0.60 – 0.66 / 0.68 – 0.69; length of embolial margin 0.53 – 0.58 / 0.58; length of cuneal margin 0.41 – 0.43 / 0.44 – 0.46; maximum width across hemelytra 0.63 – 0.68 / 0.69 – 0.70.	en	Yamada, Kazutaka, Hirowatari, Toshiya, Susila, Wayan, Okajima, Shûji (2007): The flower bug genus Montandoniola Poppius (Hemiptera: Heteroptera: Anthocoridae) from Indonesia, with descriptions of two new species. Zootaxa 1604: 37-45, DOI: 10.5281/zenodo.178743
6B7E007DFFBCF076D8E8F98F9E3EBB94.taxon	materials_examined	Holotype. % (with glass slide No. OPU – KY 163 for genitalia), “ INDONESIA, Bali, Buleleng, Melanting, S 08 ° 10 ’ 56.7 ’’ E 114 ° 40 ’ 55.7 ’’, 150 m alt., 17. viii. 2006, T. Ishikawa leg. ”. Paratypes. INDONESIA: 7 % (Figs. 2, 5 – 7), 2 Ψ (Figs. 11, 16), Bali, Buleleng, Melanting, S 08 ° 10 ’ 56.7 ’’ E 114 ° 40 ’ 55.7 ’’, 150 m alt., 16 – 17. viii. 2006, T. Ishikawa.	en	Yamada, Kazutaka, Hirowatari, Toshiya, Susila, Wayan, Okajima, Shûji (2007): The flower bug genus Montandoniola Poppius (Hemiptera: Heteroptera: Anthocoridae) from Indonesia, with descriptions of two new species. Zootaxa 1604: 37-45, DOI: 10.5281/zenodo.178743
6B7E007DFFBCF076D8E8F98F9E3EBB94.taxon	distribution	Distribution. Indonesia (Bali Island).	en	Yamada, Kazutaka, Hirowatari, Toshiya, Susila, Wayan, Okajima, Shûji (2007): The flower bug genus Montandoniola Poppius (Hemiptera: Heteroptera: Anthocoridae) from Indonesia, with descriptions of two new species. Zootaxa 1604: 37-45, DOI: 10.5281/zenodo.178743
6B7E007DFFBCF076D8E8F98F9E3EBB94.taxon	etymology	Etymology. The species epithet, a noun in the genitive, is dedicated to Dr. Tadashi Ishikawa, who collected the type specimens of this new species.	en	Yamada, Kazutaka, Hirowatari, Toshiya, Susila, Wayan, Okajima, Shûji (2007): The flower bug genus Montandoniola Poppius (Hemiptera: Heteroptera: Anthocoridae) from Indonesia, with descriptions of two new species. Zootaxa 1604: 37-45, DOI: 10.5281/zenodo.178743
6B7E007DFFBCF076D8E8F98F9E3EBB94.taxon	discussion	Remarks. This new species is most similar in general aspect to M. moraguesi, from which it can be distinguished easily by the male antennal segment III being black to brownish black except for pale yellow base and apex (Fig. 6) (in M. moraguesi, uniformly pale yellow); the rostrum being uniformly black (vs. segment III pale yellow); the corium being black to brownish black except for off – white along embolial suture (Fig. 2) (vs. mostly clear); cone being wide (Fig. 14) (vs. much narrower); and the flagellum being basally bifurcate and nearly straight (Figs. 14, 15) (vs. not bifurcated and roundly curved). This new species also resembles M. sawtellense in having similar coloration of male antennal segment III and a basally bifurcated flagellum. However, M. ishikawai is readily separable from M. sawtellense by the male fore tibiae being furnished with a row of 18 – 20 short teeth on whole of ventral surface (in M. sawtellense, 15 – 16 long teeth); the femora without teeth (vs. 5 – 7 long teeth); the flagellum being nearly straight (Fig. 14) (vs. slightly curved); and the copulatory tube being elongate and bulbous at anterior 1 / 3 of sclerotized basal part (Fig. 16) (vs. weakly sclerotized cylinder merging into longer, slightly convoluted tubule).	en	Yamada, Kazutaka, Hirowatari, Toshiya, Susila, Wayan, Okajima, Shûji (2007): The flower bug genus Montandoniola Poppius (Hemiptera: Heteroptera: Anthocoridae) from Indonesia, with descriptions of two new species. Zootaxa 1604: 37-45, DOI: 10.5281/zenodo.178743
6B7E007DFFBBF074D8E8FCF69E22BDEC.taxon	diagnosis	Diagnosis. Recognized by the following characters: Body (Figs. 3, 4) smaller and wider than its congeners. Antennal segment II about 0.6 times as long as head width across eyes. Hemelytra (Figs. 3, 4) mostly off – white; inner margin of clavi, base and apex of embolium, and cuneus black to brownish black; membrane semitransparent, with blackish patch on apical half. Pygophore (Fig. 17) swollen at left lateral margin; flagellum (Figs. 18, 19) apically curved. Copulatory tube (Fig. 20) noticeably long, straight.	en	Yamada, Kazutaka, Hirowatari, Toshiya, Susila, Wayan, Okajima, Shûji (2007): The flower bug genus Montandoniola Poppius (Hemiptera: Heteroptera: Anthocoridae) from Indonesia, with descriptions of two new species. Zootaxa 1604: 37-45, DOI: 10.5281/zenodo.178743
6B7E007DFFBBF074D8E8FCF69E22BDEC.taxon	description	Description. Coloration. Body (Figs. 3, 4) generally black, shiny on dorsal and ventral surfaces. Head, thorax, and abdomen uniformly black. Eyes and ocelli reddish black. Antenna (Figs. 3, 4, 9, 10) black to brownish black except for pale yellow segment III in female; segment IV reddish brown. Rostrum entirely black. Hemelytra (Figs. 3, 4) mostly off – white; inner margin of clavi, base and apex of embolium, and cuneus black to brownish black; membrane semitransparent, with blackish patch on apical half. Legs generally black; fore tibiae pale yellow except for fuscous at base; apex of tarsi and base of claws tinged with dark brown. Structure. Body (Figs. 3, 4) oblong oval, smaller and wider than the other congeners. Head (Figs. 3, 4, 8) smooth on dorsal surface, slightly shorter than head width across eyes, sparsely covered with silky, short, recumbent setae; pairs of long, erect setae present on side of tylus, anterior part of inner margin of eye, and between eye and ocellus; anteocular portion about 1.2 times as long as length of eye in dorsal view; vertex approximately 2.3 times as wide as eye in dorsal view; postocular portion slightly constricted, shorter than that of M. ishikawai; eyes poorly setose; ocelli small. Antennae (Figs. 9, 10) densely covered with short recumbent and long suberect setae; long suberect setae slightly longer than or as long as width of the segment; segment I just reaching apex of head; segment II clublike in male, about 0.6 times as long as head width across eyes; segment III about 0.8 times as long as segment II; segment IV weakly flattened, subequal to or slightly longer than segment III. Rostrum stout, nearly reaching fore coxae, with sparsely distributed, silky, short, erect setae; segment II thicker than the other segments; segment III longer than three times of segment II; segment IV approximately 0.8 times as long as segment III. Pronotum (Figs. 3, 4, 8) wider trapezoidal, highly polished, impunctate, sparsely clothed with silky, short, recumbent setae, bearing four distinct corner setae and a pair of long erect setae at anterior angle of callus; anterior margin nearly straight, about 1.3 times as long as mesal length; lateral margin nearly straight and strongly carinate at anterior 2 / 3; posterior margin shallowly concave, twice as wide as anterior margin in male, about 2.2 times as wide as anterior margin in female; collar long, weakly rugose, with short setae; callus swollen; posterior half of pronotum weakly depressed, transversely rugose. Scutellum (Figs. 3, 4) almost smooth, extremely swollen on anterior half and depressed on posterior half, with a pair of long, erect setae on basal 1 / 4 of lateral margin. Hemelytra (Figs. 3, 4) subparallel-sided, impunctate, sparsely covered with silky, short, recumbent setae; apical part of corium twice as wide as that of embolium; cuneal margin 0.43 times as long as embolial margin; cuneus with short setae at apex. Ostiolar peritreme (Fig. 12) accompanied with narrow, smooth, shining area anterior to gutter and broad a lutaceous area posterior to gutter; smooth shining area gradually acute toward anteriorly; evaporative area narrower than that of M. ishikawai. Legs densely covered with yellow, recumbent setae; hind tibiae weakly depressed. Abdomen covered with rows of short setae on ventral surface of each segment; seventh and eighth sterna laterally with pairs of long, suberect setae; scissure on abdominal tergite reaching end of posterior margin of third segment. Male genitalia (Figs. 17 – 19): Pygophore (Fig. 17) swollen along left lateral margin in dorsal view, posteroventrally covered with 6 – 9 long and stout setae; cone narrower than that of M. ishikawai, tapering at apex, elevated at apex in lateral view; flagellum (Figs. 18, 19) apically curved, about 1.2 times as long as width of cone. Female genitalia (Fig. 20): Copulatory tube (Fig. 20) mesally located at sternum VII, a little separated to base of ovipositor, noticeably long, slender, straight, accompanied with strongly sclerotized basal part leading into membranous and elongate apex. Measurements [% (n = 2) / Ψ (n = 2)]. Body length 1.50 – 1.70 / 1.76 – 1.80; head length 0.26 – 0.29 / 0.31; head width across eyes 0.31 / 0.31 – 0.33; vertex width 0.16 – 0.18 / 0.19; width between ocelli 0.11 – 0.13 / 0.14; length of antennal segments I – IV: 0.08 – 0.09 / 0.10 – 0.11, 0.18 – 0.19 / 0.20, 0.15 / 0.16, 0.16 – 0.17 / 0.18; length of last three rostral segments II – IV: 0.06 – 0.07 / 0.09, 0.20 / 0.22, 0.15 – 0.16 / 0.17; anterior pronotal width 0.30 – 0.31 / 0.30 – 0.31; mesal pronotal length 0.23 – 0.25 / 0.26; basal pronotal width 0.60 – 0.63 / 0.68 – 0.69; length of embolial margin 0.53 – 0.54 / 0.63; length of cuneal margin 0.23 – 0.24 / 0.28; maximum width across hemelytra 0.63 / 0.67 – 0.70.	en	Yamada, Kazutaka, Hirowatari, Toshiya, Susila, Wayan, Okajima, Shûji (2007): The flower bug genus Montandoniola Poppius (Hemiptera: Heteroptera: Anthocoridae) from Indonesia, with descriptions of two new species. Zootaxa 1604: 37-45, DOI: 10.5281/zenodo.178743
6B7E007DFFBBF074D8E8FCF69E22BDEC.taxon	materials_examined	Holotype. % (with glass slide No. OPU – KY 166 for genitalia), “ INDONESIA, Bali, Buleleng, Melanting, S 08 ° 10 ’ 56.7 ’’ E 114 ° 40 ’ 55.7 ’’, 150 m alt., 17. viii. 2006, T. Ishikawa leg. ”. Paratypes. INDONESIA: 1 %, 2 Ψ (Figs. 4, 10, 12, 20), Bali, Buleleng, Melanting, S 08 ° 10 ’ 56.7 ’’ E 114 ° 40 ’ 55.7 ’’, 150 m alt., 17. viii. 2006, T. Ishikawa.	en	Yamada, Kazutaka, Hirowatari, Toshiya, Susila, Wayan, Okajima, Shûji (2007): The flower bug genus Montandoniola Poppius (Hemiptera: Heteroptera: Anthocoridae) from Indonesia, with descriptions of two new species. Zootaxa 1604: 37-45, DOI: 10.5281/zenodo.178743
6B7E007DFFBBF074D8E8FCF69E22BDEC.taxon	distribution	Distribution. Indonesia (Bali Island).	en	Yamada, Kazutaka, Hirowatari, Toshiya, Susila, Wayan, Okajima, Shûji (2007): The flower bug genus Montandoniola Poppius (Hemiptera: Heteroptera: Anthocoridae) from Indonesia, with descriptions of two new species. Zootaxa 1604: 37-45, DOI: 10.5281/zenodo.178743
6B7E007DFFBBF074D8E8FCF69E22BDEC.taxon	etymology	Etymology. From the Latin bellatulus (= pretty, beautiful), referring to the small beautiful body of this new species; an adjective.	en	Yamada, Kazutaka, Hirowatari, Toshiya, Susila, Wayan, Okajima, Shûji (2007): The flower bug genus Montandoniola Poppius (Hemiptera: Heteroptera: Anthocoridae) from Indonesia, with descriptions of two new species. Zootaxa 1604: 37-45, DOI: 10.5281/zenodo.178743
6B7E007DFFBBF074D8E8FCF69E22BDEC.taxon	discussion	Remarks. This new species differs from its congeners by the much smaller body (Figs. 3, 4); antennal segment II being about 0.6 times as long as head width across eyes (Fig. 8); the apically curved flagellum (Figs. 18); and the noticeably long, slender, and straight copulatory tube (Fig. 20). In having mostly off-white hemelytra, this new species is similar to M. sawtellense. However, it is easily distinguished from the latter by the fore tibiae being pale yellow except for fuscous at base (in M. sawtellense, uniformly dark brown); the flagellum not being bifurcated (Figs. 18, 19) (vs. bifurcated); and the noticeably long, slender, and straight copulatory tube (Fig. 20) (vs. longer, slightly convoluted).	en	Yamada, Kazutaka, Hirowatari, Toshiya, Susila, Wayan, Okajima, Shûji (2007): The flower bug genus Montandoniola Poppius (Hemiptera: Heteroptera: Anthocoridae) from Indonesia, with descriptions of two new species. Zootaxa 1604: 37-45, DOI: 10.5281/zenodo.178743
