taxonID	type	description	language	source
653787B2FF81FFB8FEA0A26EFB631BC8.taxon	materials_examined	Type species. Stratiotes setosus Thomson, 1899, junior subjective synonym of Pagurus setosus H. Milne Edwards, 1848.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF81FFB8FEA0A26EFB631BC8.taxon	diagnosis	Diagnosis. Twelve pairs of bi­ or quadriserial gills; no pleurobranch on fifth or eighth thoracic somites. Shield well calcified, with rostrum well developed or reduced. Ocular acicles well developed or reduced, simple, bi­ or multidenticulate. Antennal peduncle with supranumerary segmentation; acicles elongate, spinose or spinulose. Mandible with unarmed mesial margin, palp often indistinctly 3 ­ segmented. Maxillule with external lobe of endopod prominently recurved. Maxilla with elongate endopod. First maxilliped with well developed epipod. Second maxilliped without distinguishing characters. Third maxilliped with basal segments approximate or somewhat separated; crista dentata usually well developed; accessory tooth absent; merus usually with 1 or more spines; carpus, propodus and dactyl usually unarmed. Chelipeds equal, subequal or unequal, with left considerably larger than right; similar or dissimilar in armament; claws corneous, fingers opening in horizontal or oblique plane. Fourth pereopods subchelate or weakly semichelate, with or without preungual process developed at base of claw. Fifth pereopods chelate. Males with paired gonopores; first and second pleonal somites each with pair of pleopods modified as gonopods; unpaired, unequally biramous left pleopods on pleomeres 3 – 4, pleopod 5 with external ramus well developed, internal ramus rudimentary. Females with paired gonopores; first pleonal somite with pair of modified pleopods; following 3 pleomeres with unequally biramous left pleopods; fifth pleopod as in male; brood pouch usually well developed, occasionally entirely absent. Uropods asymmetrical. Telson with lateral incisions; posterior portion divided by median cleft into 2 subequal to markedly unequal lobes.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF81FFB8FEA0A26EFB631BC8.taxon	discussion	Remarks. Thomson (1899) established the genus Stratiotes for Pagurus setosus Filhol 1885, and mentioned that it appeared to be allied to Diogenes because of the unequal chelipeds (left much larger than right) but differed in the absence of the intercalary rostral process. Forest and McLaughlin (2000) discussed the status of Stratiotes and mentioned the opinions on this genus expressed by several authors such as Alcock (1905) and Thompson (1930). As the gill number was not examined, these authors were not aware that the type species, Stratiotes setosus, which is in fact synonymous with Paguristes setosus, had only 12 pairs of gills (absence of the pleurobranch on the fifth and eighth thoracic somites). For more than a century the genus Stratiotes was considered a junior synonym of Paguristes. Now that it has been confirmed that Paguristes setosus (H Milne Edwards) possesses 12 pairs of gills, the genus Stratiotes is reinstated. Among the species included in the genus Paguristes s. l in the collection of the Muséum national d’Histoire naturelle, 13 species have been found to have 12 pairs of gills. Therefore, they are transferred, herein, to the genus Stratiotes (Table 1). Paguristes hispidus A. Milne Edwards & Bouvier, 1892 MNHN Pg 2927 Stratiotes hispidus n. comb.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF87FFB7FEA0A23BFBF21C58.taxon	materials_examined	Material examined. 2 males, 1.7, 2.3 mm, Kai Islands, Siboga, stn 260, 05 ° 36.5 ' S, 132 ° 55.2 ' E, 90 m, 16 – 18 Dec 1899 (ZMA).	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF87FFB7FEA0A23BFBF21C58.taxon	description	Description. Biserial phyllobranchiae. Shield (Fig. 1) as long or longer than broad; lateral margins sloping or weakly terraced, spinulose, or with prominent spine. Rostrum broadly triangular, not overreaching level of lateral projections. Lateral projections triangular, with marginal or submarginal small spine. Branchiostegites each with row of very small spines on dorsal margin in distal half. Ocular peduncles slender to moderately stout, 0.6 – 0.8 length of shield; corneal diameter 0.2 – 0.4 of peduncular length; ocular acicles elongate, very slender, terminating acutely, often with moderately long, plumose setae. Antennular peduncles overreaching distal margins of corneas by 0.2 – 0.3 length of ultimate segments; basal segment with acute spine dorsolaterally. Antennal peduncles reaching nearly to or slightly overreaching distal margins of corneas; fifth and fourth segments unarmed; third segment unarmed or with spine at ventrodistal angle; second segment with dorsolateral distal angle produced, terminating in prominent simple or bifid spine, subdistal spine frequently present on lateral margin, dorsomesial distal angle with small spine; first segment unarmed. Antennal acicle reaching to or slightly beyond proximal margin of ultimate peduncular segment, terminating in prominent bifid spine, mesial margin with 2 – 4 spines, lateral margin usually with 1 or 2 small spines. Antennal flagellum longer than carapace, each article with long, paired setae ventrally. Chelipeds unequal and dissimilar, left cheliped much larger; carpus and chela of both articulating obliquely with merus, putting dorsal surfaces in lateral positions relative to body axis. Left cheliped (Fig. 2 a – c) with row of minute spinules or granules on ventromesial margin of ischium. Merus with 1 spine at dorsodistal margin, dorsal margin with spines or spinulose protuberances distally, row of low, sometimes spinulose protuberances and tufts of plumose setae proximally; lateral face with scattered small tubercles, at least in lower half, and tufts of plumose setae in dorsal half, distal margin with row of small spines in dorsal half, ventrolateral margin with row of often corneous­tipped spines; ventromesial margin with row of much more prominent, corneous­tipped spines, mesial face glabrous; ventral surface with moderately long plumose setae. Carpus with row of conical, sometimes corneous­tipped spines on upper margin; outer surface with numerous, but not closely­packed, smaller spines, forming longitudinal row in upper half, distal margin with row of small conical spines, 1 prominent spine at lower distal angle; inner surface with scattered low tubercles, lower distal margin with few plumose setae. Palm with single to double row of conical or tuberculate spines on upper margin and additional row adjacent to upper margin; outer surface with few individual, conical or tuberculate spines proximally and 2 or 3 longitudinal rows of clusters of 2 to several conical or elongate blunt or spiniform tubercles, lower row extending onto fixed finger on weakly elevated broad ridge; lower margin with 1 – 3 irregular rows of tubercles and tufts of moderately long, plumose setae; inner and lower surfaces with scattered low protuberances and sparse tufts of plumose setae. Dactyl approximately equal to length of palm; upper surface with irregular double row of conical or tuberculate spines and tufts of moderately long, plumose setae; outer surface with prominent longitudinal sulcus, often flanked dorsally and / or ventrally with few small tubercles; inner surface with row of small, conical spines and tufts of plumose setae dorsally, row of low protuberances and tufts of plumose setae ventrally; cutting edge with row of small, calcareous teeth, terminating in small calcareous claw; no hiatus between dactyl and fixed finger. Right cheliped (Fig. 2 d, e) with row of widely­spaced spinules or tubercles on ventromesial margin of ischium. Merus with 1 prominent spine at dorsodistal margin and slightly smaller subdistal spine, both partially obscured by long, plumose setae, remainder of dorsal margin with row of tiny spinules or small spines concealed by long, plumose setae; mesial face unarmed, ventromesial margin with long, plumose setae not obscuring row of small spines; lateral face with few scattered spinules, ventrolateral margin with few long, plumose setae and row of very small spines or spinules; ventral surface unarmed. Carpus with row of spines on upper margin increasing in size distally and partially obscured by long, plumose setae; outer face with row of small spines in upper half and 1 to several spinules in lower half; inner surface with few scattered setae. Palm with row of 3 – 5 moderately small spines on upper margin; outer surface convex, with 4 or 5 irregular longitudinal rows of small spines partially obscured by long plumose setae, particularly in upper half, lower with 2 rows extending nearly to tip of fixed finger; inner surface of palm and fixed finger with widely­spaced tufts of short setae; cutting edge with row of very small calcareous teeth, terminating in corneous claw and with 1 or 2 adjacent corneous spines on lower margin. Dactyl with row of small spines on upper margin, at least partially obscured by long, plumose setae; outer surface with irregular short row of spinules accompanied by plumose setae; inner surface with few short setae; cutting edge with central longitudinal groove and few corneous teeth distally, terminating in small corneous claw. Second and third pereopods (Fig. 3 a – c) with left third dissimilar. Ischia of second pereopods each with 1 or 2 spinules on ventromesial margin; ischium of right third unarmed. Meri of second pereopods each with dorsal row of spinules and long plumose setae, ventral margins each with row of small spines; merus of third right with spinules or spinulose protuberances and long setae on dorsal margin, ventral margin with long, plumose setae and small spine at ventrolateral distal angle. Carpi each with dorsal row of spines partially concealed by long, plumose setae; spines of third right smallest. Propodi of second pereopods each with dorsal row of small spines and ventral row of spiniform setae, all obscured by tufts of long, plumose setae; propodus of third right with only plumose setae dorsally and ventrally. Dactyls of second and right third slightly longer than propodi; dorsal surfaces of each with row of very small spiniform setae obscured by long plumose setae; ventral margins each with row of small spiniform setae, also concealed by tufts of long, plumose setae; mesial and lateral faces of dactyl of second pereopods each with faint longitudinal sulcus; mesial and lateral faces of right third convex. Left third pereopod (Fig. 3 b) with ischium unarmed. Merus with row of very small spines or spinules on dorsal surface concealed by long, plumose setae; ventrolateral distal angle with small spine, ventral margin with long plumose setae. Carpus with small spine at dorsodistal margin and row of spinules on dorsal surface; lateral face with faint longitudinal sulcus in dorsal half. Propodus with minutely spinulose dorsal surface and long, plumose setae; lateral face with broad longitudinal sulcus flanked dorsally by row of very small spines and ventrally by row of blunt or subacute tooth­like spines; ventral surface with long plumose setae and row of tiny spinules, at least proximally. Dactyl longer than propodus; dorsal surface with 1 or 2 irregular rows of minute spinules and long, plumose setae; lateral face with broad longitudinal sulcus, flanked dorsally by row of minute spinules and ventrally by row of acute and / or blunt tooth­like spines, ventral surface spinulose, at least proximally, but armature frequently masked by long, plumose setae; mesial face with faint longitudinal sulcus. Fourth pereopod (Fig. 3 d) with small preungual process. Male first pleopods (Fig. 3 e, f) robust; inferior lamella obtusely triangular, distal margin with row of small hook­like spines. External lobe elongate, separated from internal lobe by deep narrow cleft; internal lobe broad. No brood pouch developed in ovigerous females, but tuft of long, plumose setae present posterior to egg­bearing fourth pleopod (McLaughlin, pers. comm.). Telson (Fig. 3 g) with posterior lobes asymmetrical, right smaller; right terminal margin with few setae and 2 to several spinules, not extending onto lateral margin, left with setae and 6 or more stronger spinules, often extending onto lateral margin. Color. Antennae and ambulatory legs annulated with narrow reddish bands. Chelae bright red, tips of fingers of larger hand white, body and limbs scantly clothed with brownish setae (after Whitelegge 1900). Habitat. Reported from sand and pebble bottom in New South Wales. In southwestern Australian, Morgan (1989) recorded this species from soft substrates, usually associated with seagrass meadows. McCulloch (1913) reported that the shells of this species were almost always accompanied by polyzoan colonies; however, Morgan (1989) did not find such an association in specimens collected in Western Australia. In Indonesia this species was collected from sand substrate and shell fragments.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF87FFB7FEA0A23BFBF21C58.taxon	distribution	Distribution. New South Wales, Western Australia, and now from the Kai Islands, Indonesia; subtidal to 90 m.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF87FFB7FEA0A23BFBF21C58.taxon	discussion	Remarks. The description provided above is based in part on the two males collected during the Siboga Expedition, and in part on information provided on the type series by P. McLaughlin. The grossly unequal chelipeds with the tuberculate armature of the left chela and the dissimilar third pereopod set this species apart from the other species now assigned to Stratiotes. Until its present recognition, S. tuberculatus n. comb. was believed to be endemic to Tasmania, southern and western Australia at depths up to 50 m (Morgan 1989). Its presence in the Kai Islands represents both range and depth extensions.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF89FFB3FEA0A283FD361B98.taxon	materials_examined	Material examined. Holotype: male, 9.0 mm, Snellius II, stn. 104, north­east of Komodo Island, 08 ° 25.3 ' S, 119 ° 36.2 ' E, 140 – 150 m, 20 Sept 1984 (MZB).	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF89FFB3FEA0A283FD361B98.taxon	description	Description. Biserial phyllobranchiae. Shield (Fig. 4 a) longer than broad; lateral margins sloping, dorsal surface rugose, with low elevation on either side of midline and short transverse row of small spines and tuft of setae. Rostrum broadly triangular, distinctly overreaching lateral projections, terminating in acute spine obscured by dense setae. Lateral projections triangular, with acute marginal spine. Ocular peduncles slender, 0.6 length of shield; corneal diameter 0.2 of peduncular length; ocular acicles small, distinctly separated basally, terminating acutely, with moderately long, plumose setae. Antennular peduncles, when fully extended, reaching bases of corneas; basal segment with distal acute spine dorsomesially and dorsolaterally, and 1 median spine dorsolaterally. Antennal peduncles reaching nearly to or slightly overreaching half length of ocular peduncles; fifth segment unarmed; fourth segment with dorsodistal spine; third segment with ventromesial distal angle strongly produced terminating in strong spine; second segment with dorsolateral distal angle produced, terminating in prominent simple or bifid spine, subdistal spine frequently present on lateral margin, dorsomesial distal angle with strong spine; first segment unarmed. Antennal acicle reaching to or slightly beyond distal margin of ultimate peduncular segment, terminating in prominent bifid spine, mesial margin with strong proximal spine, lateral margin with 2 spines. Antennal flagellum 2.5 times shorter than carapace, consisting of about 17 articles, each article with short, sparse setae. Basis of third maxilliped with 2 distoventral spines; ventral margin of ischium with distal spine; merus with 4 small spines on ventral margin. Chelipeds (Fig. 4 b, c) subequal, armature similar. Ischia each with row of minute spinules or granules on ventromesial margin. Meri each with 1 spine at dorsodistal margin, dorsal margin with 2 distal transverse rows of spines extending mesially and laterally; ventromesial margins each with row of strong spines and tuft of plumose setae; ventrolateral margins each with row of less prominent spines and dense plumose setae; mesial faces glabrous, lateral faces with scattered small tubercles. Carpi each with row of 4 conical spines on dorsomesial margin; dorsal surface with 2 irregular rows of spines, first row adjacent to dorsomesial margin, consisting of moderately small spines or tubercles, second row with more numerous and moderately larger spines; tuft of moderately dense setae on dorsal surface. Palms half length of carpi. Dorsomesial margins each with 3 strong, occasionally corneous­tipped spines; dorsal surfaces each with several irregular rows of large, occasionally corneous­tipped spines, becoming closely­spaced on fixed finger; ventral surfaces each with single row of large spines medially, not extending to fixed finger; cutting edge with small calcareous teeth, terminating in moderately large corneous claw. Dactyls approximately 3 times length of palms; each with row of large, corneous­tipped spines on dorsomesial margin; dorsal surface with rows of small to moderately large spines, one strong spine proximally; mesial surface with large corneous, widely­spaced spines and tufts of setae; cutting edge with small calcareous teeth, terminating in small corneous claw; no hiatus between dactyl and fixed finger. Tufts of dense setae on entire surface of palm, fixed finger and dactyl concealing spines and tubercles. Left and right pereopods similar. Ischia of second pereopods (Fig. 5 a, b) each with row of spinules on ventromesial margin. Meri each with row of small spines on ventrolateral margin, stronger row of spines on ventromesial margins. Carpi approximately 0.8 length of meri, each with dorsal row of 6 irregular, strong spines; lateral faces convex, each with longitudinal sulcus near dorsal margin accompanied by 2 distal spines. Propodi slightly longer than carpi, each with dorsal row of 8 irregular, moderately large spines, dorsodistal margin with small corneous spines; lateral surfaces convex, each with row of protuberances accompanied by tufts of setae near dorsal and ventral margins, row of tufts of shorter setae in midline; mesial surface adjacent to ventral margin with longitudinal row of clusters of 2 spines, each cluster with tuft of setae, low protuberance accompanied by long setae near dorsal margin; ventral margins each with denticles. Dactyls 1.4 length of propodi, dorsal margins each with row of corneous spines proximally; mesial surfaces each with row of small spiniform setae near dorsal and ventral margins; ventral margins each with row of small spiniform setae, stronger distally. Mesial and lateral faces of dactyls each with faint longitudinal sulcus proximally. All surfaces and particularly dorsal and ventral margins partially or completely obscured by long and dense plumose setae. Third pereopod (Fig. 5 c, d) with ischia unarmed. Meri each with row of very small spines or spinules on dorsal surface; ventromesial distal angle each with small spine, ventromesial and ventrolateral margins each with row of spines. Carpi slightly shorter than meri, each with small irregular spines on dorsal margins; lateral surfaces convex, each with longitudinal sulcus near dorsal margin. Propodi each with minutely spinulose dorsal and ventral surfaces; lateral faces with protuberances accompanied by long setae, shallow longitudinal sulcus accompanied by tufts of setae in midline; mesial surfaces each with 2 rows of corneous­ tipped spines: 1 row near ventral margin composed of clusters of 2 – 3 small corneous­tipped tubercles with tufts of setae on each cluster and 1 other row of simple spines near dorsal margin, concealed at least partially by dense setae. Dactyls almost 1.5 longer than propodi; dorsal margins each with row of minute spinules; mesial faces each with irregular rows of spiniform setae; lateral and mesial faces each with narrow longitudinal sulcus proximally; ventral margins each with spiniform setae. All surfaces and particularly dorsal and ventral margins, partially or completely obscured by long and dense plumose setae. Fourth pereopod (fig. 4 e) with preungual process. Male first pleopods (Fig. 5 e, f) robust; inferior lamella obtusely triangular, distal margin with several rows of hook­like spines. External lobe elongate, tip curve, separated from internal lobe by deep narrow cleft; internal lobe broad. Female unknown. Telson (Fig. 4 d) with posterior lobes asymmetrical, separated by narrow cleft; right lobe smaller, terminal margins of left and right lobes with row of tiny spinules extending onto proximal half of lateral margins.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF89FFB3FEA0A283FD361B98.taxon	etymology	Etymology. The specific name is derived from Latin brevi, meaning short, and antennae, alluding to the short antennal peduncles. Habitat. The single specimen was collected from substrate of calcareous block with sponges, epifaunal tunicates and scleractinians.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF89FFB3FEA0A283FD361B98.taxon	distribution	Distribution. Komodo Island, 140 – 150 m.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF89FFB3FEA0A283FD361B98.taxon	discussion	Remarks. Stratiotes breviantennatus sp. nov. is easily distinguished from other species of the genus by the shape of the telson, which has row of small spinules almost the entire length of the terminal margins. The short and almost glabrous antennal flagella also separate this species from its congeners.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF8DFFAFFEA0A443FD0D1A78.taxon	materials_examined	Material examined. Holotype: male, 4.6 mm, without right third pereopod, northeast of Binongko Island, Snellius II, stn 37, 05 ° 54.2 ' S, 123 ° 57.0 ' E, 350 – 500 m, 10 Sep 1984 (MZB). Paratypes: 2 females, 3.0, 4.0 mm, 1 ovig. female, 3.5 mm, southeast of Karang Kaledupa, Snellius II, stn 24, 5 ° 57 ' S, 123 ° 49 ' E, 350 – 500 m, 0 9 Sep 1984 (MZB); 1 male, 3. 5 mm, CORINDON, stn 229, 00 ° 02.2 ' N, 119 ° 49.8 ' E, 4 Nov 1980, 445 – 411 m (MNHN); 1 male, 6.5 mm, Galathea, stn. 500, 07 ° 34 ' S, 132 ° 44 ' E, 390 m, 25 Sep 1951.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF8DFFAFFEA0A443FD0D1A78.taxon	description	Description. Biserial phyllobranchiae. Shield (Fig. 6 a) 1.2 longer than broad; lateral margins sloping, short transverse row of small spines and tuft of setae on dorsal surface. Rostrum triangularly obtuse, reaching distal 0.3 of ocular acicles, about same length as lateral projections, terminating in acute spine. Lateral projections triangular, with acute marginal spine. Ocular peduncles stout, 0.5 length of shield; slightly inflated basally and in corneal region; corneal diameter 0.3 of peduncular length; ocular acicles small, distinctly separated basally, terminating in simple spine. Antennular peduncles, when fully extended, overreaching corneas by entire length of distal segment; basal segment with distal acute spine dorsomesially and dorsolaterally and median spine dorsolaterally. Antennal peduncles approximately same length or slightly longer than ocular peduncles; fifth segment unarmed; fourth segment with small distolateral spine; third segment with ventromesial distal angle strongly produced, terminating in strong spine; second segment with dorsolateral distal angle produced, terminating in prominent bifid spine, subdistal spine on lateral margin, strong spine on distal margin; first segment unarmed. Antennal acicles reaching distal margin of ultimate peduncular segment, terminating in prominent bifid spine, mesial margin with 3 – 4 strong spines, lateral margin with 2 – 3 strong distal spines. Antennal flagellum same length or slightly longer than shield, consisting of about 20 articles, proximal articles with short, sparse setae, distal articles with longer and denser setae. Basis of third maxilliped with 1 distoventral spine; merus with 3 – 4 small spines on ventral margin. Chelipeds (Figs. 7 a – c) subequal, armature similar. Ventromesial margins of ischia each with row of small spines. Meri each with denticles on dorsal margin; ventromesial margins each with row of strong spines, less prominent spines on ventrolateral margins and sparse setae, mesial faces glabrous; lateral faces with scattered small tubercles near dorsal and ventral margins. Carpi 0.6 length of meri, row of 3 – 4 corneous­tipped spines on each dorsomesial margin; dorsal surface with longitudinal rows of moderately small spines near dorsomesial and dorsolateral margins; long and sparse setae on dorsal surface. Palms 0.6 length of carpi, dorsomesial margins each with 3 strong, corneous­tipped spines; dorsal surfaces each with several irregular rows of small, occasionally corneous­tipped spines, becoming closely­spaced on fixed finger; ventral surfaces each with single row of tubercles medially, extending onto fixed finger, tuft of setae at base of each tubercle; cutting edge with small calcareous teeth in proximal 0.7, corneous teeth in distal 0.3 terminating in corneous claw; dorsal surface with sparse long setae. Dactyls approximately twice as long as palms; cutting edge with small calcareous teeth in proximal halves, corneous teeth in distal halves, terminating in small corneous claws. Dorsomesial margins each with row of small, occasionally corneous­tipped spines; dorsal surface with scattered small spines and tufts of setae; mesial surfaces densely covered with small corneous spines, sparsely setose; no hiatus between dactyl and fixed finger. Second pereopods (Fig. 7 f) slightly shorter than third, left and right second pereopods with similar armature. Dorsal margins of meri unarmed, ventromesial margins each with row of minute spines. Carpi approximately 0.7 length of meri, each with dorsal row of 6 irregular, occasionally corneous­tipped spines; lateral faces convex, each with longitudinal sulcus near dorsal margin. Propodi slightly longer than carpi, each with dorsal row of 9 large corneous­tipped spines; mesial surfaces each with row of setae near dorsal margins. Dactyls 1.5 length of propodi, weakly curved, dorsal margins each with row of small corneous­tipped spines, becoming smaller and spiniform distally, obscured by tufts of setae; mesial surfaces with scattered setae; ventral margins each with row of small spiniform setae. Meri of third pereopods (Figs. 7 d, e) with long setae on dorsal and ventral margins. Carpi 0.8 length of meri, each with small subdistal spine on dorsal margin; lateral surfaces convex, each with longitudinal sulcus near dorsal margin; long setae on dorsal and ventral margins. Propodi unarmed, long setae on dorsal and ventral margins, mesial faces with long, rather dense setae. Dactyls almost 1.6 length of propodi; each dorsal margin with long setae; row of spiniform setae and long simple setae on dorsomesial margin and mesial face; ventral margin with spiniform setae and long simple setae. Fourth pereopod with preungual process (Fig. 6 b). Male first pleopods (Fig. 6 d, e) slender; inferior lamella rounded, distal margin with row of hook­like spines. External lobe elongate, tip slightly curved, separated from internal lobe by narrow cleft; internal lobe broad. Female brood pouch small, triangular, with sparse, short setae. Telson (Fig. 6 c) with asymmetrical posterior lobes, left lobe larger than right; cleft separating two lobes narrow and deep. Terminal margins of left and right lobes each with 6 corneous­tipped spinules, closely­spaced near cleft.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF8DFFAFFEA0A443FD0D1A78.taxon	etymology	Etymology. This species is dedicated to the late Michèle de Saint Laurent. Her help at the beginning of this work is greatly appreciated. Habitat. The species was collected from substrate with calcareous stones, dead corals and sponges.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF8DFFAFFEA0A443FD0D1A78.taxon	distribution	Distribution. Southeast of Sulawesi and Macassar Strait, Indonesia	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF8DFFAFFEA0A443FD0D1A78.taxon	discussion	Remarks. The form of male first pleopod varies in relation to the size of the individual: the distal margin of inferior lamella in the small male paratype (SL 3.5 mm) was without a row of hook­like spines, while in holotype and other paratype the row of hook­like spines was well developed. Stratiotes micheleae n. sp. resembles S. ngochoae n. sp., subsequently described in this paper, in having stout ocular peduncles and dense spinules on the mesial surfaces of dactyls of the chelipeds. However, the more significant differences, include simple instead of bifid ocular acicles; ventral margins of the meri of the second and third pereopods that are smooth or armed only with tiny spinules ventromesially rather than bearing stronger spines. Additionally, the terminal margin of the telson of S. micheleae is armed with six small spines while in S. ngochoae it is armed with four to 10 large spines. The form of male first pleopod of specimen of the same small size of both species is notably different. In S micheleae the distal margin of inferior lamella is without a row of hook­like spines, while in S. ngochoae the hook­like spines formed several rows on the distal margin and dorsal surface of inferior lamella.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF91FFABFEA0A4E3FE5E19C0.taxon	materials_examined	Material examined. Holotype: male, 3. 8 mm, Taruna bay, Sangir Besar (Great Sangir) Island, Siboga, stn. 127, 45 m, 20 – 21 Jul 1899 (ZMA). Paratype: 1 male, 4.9 mm, type locality (ZMA)	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF91FFABFEA0A4E3FE5E19C0.taxon	description	Description. Biserial phyllobranchiae. Shield (Fig. 8 a) 1.2 longer than broad; lateral margins sloping, short transverse row of small spines and tufts of setae on dorsal surface. Rostrum broadly triangular, distinctly overreaching lateral projections, terminating in acute spine. Lateral projections triangular, with acute marginal spine. Ocular peduncles stout, left slightly longer than right, 0.7 – 0.8 length of shield; swollen proximally, corneas less inflated; corneal diameter 0.3 of peduncular length; ocular acicles small, distinctly separated basally, terminating in bifid spine. Antennular peduncles, when fully extended, overreaching corneas; basal segment with distal acute spine dorsomesially and dorsolaterally, and median spine dorsolaterally. Antennal peduncles reaching nearly to bases of corneas; fifth and fourth segments unarmed; third segment with ventromesial distal angle strongly produced terminating in strong spine; second segment with dorsolateral distal angle produced, terminating in prominent bifid spine, subdistal spine on lateral margin, strong spine on distal margin; first segment unarmed. Antennal acicle not reaching distal margin of ultimate peduncular segment, terminating in prominent bifid spine, mesial margin with 2 strong proximal spines, lateral margin with 2 strong distal spines. Antennal flagellum slightly shorter than carapace, consisting of 15 – 17 articles, each article with short, sparse setae. Basis of third maxilliped with 2 distoventral spines; merus with 5 small spines on ventral margin. Chelipeds subequal (Fig. 9 a – d), armature similar. Ventromesial margins of ischia each with row of moderately strong spines. Meri each with row of strong spines on dorsal margin, weaker spines proximally; ventromesial margins each with row of strong spines, less prominent spines on ventrolateral margins, partially obscured by dense plumose setae; mesial faces glabrous, lateral faces with scattered small tubercles. Carpi each with row of corneous­tipped spines on dorsomesial margin; dorsal surfaces each with 1 row of moderately small spines near dorsomesial margin, second row of moderately larger spines medially; small spines on distal margin and tufts of setae. Palms 0.7 length of carpi, dorsomesial margins each with 3 strong, corneous­tipped spines; dorsal surfaces with several irregular rows of large, occasionally corneous­tipped spines, becoming closely­spaced on fixed finger; ventral surfaces each with single row of corneous spines medially, extending to fixed finger; cutting edge with small calcareous teeth terminating in moderately large corneous claw; dorsal and lateral surfaces of palm and fixed finger covered with long setae. Dactyls approximately twice length of palms; dorsomesial margins each with row of small, corneous­tipped spines; dorsal surfaces with scattered small spines, tufts of setae on entire dorsal surfaces; mesial surfaces each covered with small corneous spines, more or less arranged in traverse rows, accompanied by tufts of setae; cutting edge with small calcareous teeth, terminating in small corneous claw, no hiatus between dactyl and fixed finger. Left and right second and third pereopods similar. Ischia of second pereopods (Fig. 9 e, f) each with distal spines on ventromesial and dorsal margins. Meri each with row of small spines on ventrolateral margin, stronger row of spines on ventromesial margin; long setae on dorsal and ventral margins. Carpi approximately 0.7 length of meri, each with dorsal row of 9 or 10 irregular strong spines; lateral faces convex, each with longitudinal sulcus near dorsal margin accompanied by 2 distal spines and long setae. Propodi slightly longer than carpi, each with dorsal row of 8 large corneous­tipped spines; ventral margins each with denticles; mesial surfaces each with row of setae near ventral margins. Dactyls 1.2 length of propodi, dorsal margins each with row of corneous spines and tufts of setae; mesial surfaces each with row of small spiniform setae near ventral margins and scattered setae; ventral margins each with row of small spiniform setae, stronger distally. Third pereopods (Figs. 9 g, h) each with distal spine on ventromesial margin of ischium. Meri each with row of spines or spinules on dorsal margin; ventromesial margins each with row of spines and long setae; ventrolateral margins each with 2 spinules medially. Carpi slightly shorter than meri, each with irregular spines on dorsal margin, distal spine strongest; lateral surfaces convex, each with longitudinal sulcus near dorsal margin accompanied by 2 distal spines and long setae. Propodi each with minutely spinulose dorsal surface; lateral faces glabrous; mesial surfaces each with transverse striae consisting of 2 or 3 small corneous­tipped spines, concealed at least partially by dense setae. Dactyls almost 1.4 longer than propodi; dorsal margins each with row of minute spinules and long setae; mesial faces each with irregular rows of spiniform setae and long simple setae; lateral and mesial faces each with narrow longitudinal sulcus proximally; ventral margins each with spiniform setae and long, simple setae. Fourth pereopods with preungual process (Fig. 9 i). Male first pleopods (Figs. 8 d, e) robust; inferior lamella rounded, distal margin with several rows of hook­like spines. External lobe elongate, tip slightly curved, separated from internal lobe by shallow cleft; internal lobe rounded, broad. Female unknown. Telson with posterior lobes asymmetrical, separated by wide cleft; right lobe much smaller (holotype) (Fig. 8 b); right lobe only slightly smaller than left (paratype) (Fig. 8 c). Terminal margin of left lobe with 10 spines (holotype) or 7 strong spines (paratype); right lobes with row of 4 (holotype) or 6 (paratype) strong corneous spines, extending onto lateral margins.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF91FFABFEA0A4E3FE5E19C0.taxon	etymology	Etymology. Dedicated to Dr Nguyen Ngoc­Ho who has contributed so much to our knowledge of the anomuran fauna. Habitat. The specimens were collected from fine sand substrate.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF91FFABFEA0A4E3FE5E19C0.taxon	distribution	Distribution. North of Sulawesi, Indonesia.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF91FFABFEA0A4E3FE5E19C0.taxon	discussion	Remarks. The dissimilarity between left and right lobes of the telson of the holotype seems to be an anomaly. The telson of the paratype is considered the normal form. S. ngochoae n. sp. most closely resembles S. breviantennatus n. sp. in the general shape of the chelipeds and the first male pleopods, and in the armature of the pereopods. Differences between these two species are apparent in the armature of the mesial faces of the dactyls of the chelipeds and the form and spination of the telson. In S. breviantennatus the mesial faces of the dactyls of the chelipeds are armed with a few moderately large, corneous spines, whereas in S. ngochoae they are covered with dense, small corneous spines. The telson of S. breviantennatus is armed with row of tiny spinules on the terminal margins of both lobes and three larger spines on the lateral margin of the left lobe. In S. ngochoae, the telson has larger, corneous­tipped spines extending onto the lateral margins. Furthermore, the ocular acicles of S. ngochoae are bifid, whereas in S. breviantennatus they are simple.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF9AFFA4FEA0A05CFB021AD7.taxon	materials_examined	Type species. Pseudopaguristes janetkae McLaughlin, 2002, by original designation.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF9AFFA4FEA0A05CFB021AD7.taxon	diagnosis	Diagnosis. Eight pairs of functional phyllobranchiate gills (no pleurobranchs on thoracic somites 5 and 8); arthrobranchs of third maxillipeds and chelipeds markedly reduced or absent; remaining gills bi­ or quadriserial. Rostrum usually well developed. Ocular acicles well developed, widely separated. Antennal peduncles with supernumerary segmentation; acicles moderately long, spinose. Antennal flagella of short to medium length, slightly shorter to slightly longer than carapace. Mandible with smooth medial margin; palp 3 ­ segmented. Endopod of maxillule with well developed, recurved external lobe. First maxilliped with setose epipod. Third maxilliped with crista dentata consisting of row of quite small teeth, no accessory tooth. Chelipeds subequal or unequal, sometimes dimorphic, dissimilar or similar; claws corneous, fingers opening in horizontal or oblique plane. Fourth pereopods with or without preungual process at base of dactylar claw. Fifth pereopods chelate. Males with single or paired gonopores; unpaired or paired pleopods modified as gonopods on first and second pleomeres; unpaired, unequally biramous left pleopods on pleomeres 3 and 4, pleopod 5 uniramous. Females with paired gonopores or single gonopore on coxa of left third pereopod; paired, modified, uniramous pleopods on pleonal somite 1; following 3 pleomeres with unpaired, unequally biramous left pleopods; pleopod 5 with external ramus well developed, internal ramus rudimentary; brood pouch absent or present, sometimes represented by dense setae. Uropods markedly asymmetrical. Telson with lateral incisions; posterior lobes slightly to markedly asymmetrical.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF9AFFA4FEA0A05CFB021AD7.taxon	discussion	Remarks. Three species in the collection of the Muséum national d’Histoire naturelle, but not present in the Indonesian material have eight pairs of gills and are herein transferred to Pseudopaguristes: Paguristes calliopsis Forest & de Saint Laurent, 1968 (MNHN Pg 2379), Paguristes invisisacculus McLaughlin & Provenzano, 1974 (MNHN Pg 1854) and Paguristes laurentae Morgan & Forest, 1991 (MNHN Pg 4489).	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF9BFFA6FEA0A36BFE111DF8.taxon	materials_examined	Material examined. 2 males 4.5 and 3.5 mm, Teluk Kotania, Seram Island, Maluku, 6 – 10 m, 23 Jul 1997 (MZB); 1 male 3.5 mm, 1 female 3.4 mm, Morela, Ambon Island, Maluku, 6 – 10 m, 16 Jul 1997 (RCO).	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF9BFFA6FEA0A36BFE111DF8.taxon	diagnosis	Diagnosis. Quadriserial phyllobranchiae. Shield longer than broad; rostrum triangular, longer than lateral projections, reaching almost to distal tips of ocular acicles. Ocular peduncles swollen proximally; slightly shorter than shield; longer than antennal and antennular peduncles; corneas small; ocular acicles with 2 – 4 terminal spines. Antennal flagella slightly longer than shield, composed of about 20 articles with short, sparse setae. Male chelipeds unequal, right larger than left; cutting edges of dactyl and fixed finger with narrow hiatus, each with row of small teeth. Female chelipeds equal. Ambulatory legs exceeding chelipeds by half length of dactyls; sparse setae on dorsal and ventral margins; dactyls same length as propodi. Fourth pereopod with preungual process. First and second male pleopod and male gonopores paired. Female with gonopore only on coxa of left pereopod; brood pouch absent. Telson asymmetrical, left slightly larger than right, with minute spines on terminal margins. Color in life. “ Shield uniform white or pale orange. Ocular peduncles and corneas solid bright orange. Antennules uniform orange, flagellum transparent. Antennal flagella alternately transparent and red or purple. Chelipeds bright orange; merus with red or dark purple spot on lateral and ventral faces, 2 spots on mesial face; carpus and chela each with similar marking on dorsal, lateral and mesial faces. Pereopods 2 and 3 bright orange, with lateral red patch on merus and carpus ” (Haig & Ball 1988).	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF9BFFA6FEA0A36BFE111DF8.taxon	materials_examined	Type locality. Ternate, Maluku, Indonesia. Habitat. Coral reef.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF9BFFA6FEA0A36BFE111DF8.taxon	distribution	Distribution. At present, P. kuekenthali is known only from Vanuatu, and the Maluku waters of Indonesia.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF9BFFA6FEA0A36BFE111DF8.taxon	discussion	Remarks. Pseudopaguristes kuekenthali was redescribed and figured by Haig & Ball (1988) as Paguristes kuekenthali from the specimens collected by the Alpha Helix Expedition. Neither De Man (1902) nor Haig & Ball (1988) discussed the dissimilarities of male chelipeds. The Alpha Helix specimens could not be found in the National Institute of Oceanology (now Research Centre for Oceanography) Jakarta, Indonesia, but the material used in this study are from the same locality, i. e. Ceram Island. The specimens agree well with the redescription of the species by Haig & Ball (1988). Therefore the description of P. kuekenthali can be supplemented by adding that the male has unequal chelipeds, the right much larger than left, whereas the female has equal chelipeds. Pseudopaguristes kuekenthali, P. janetkae and P. bollandi share several characters such as the larger right cheliped in males, ocular acicles that terminate in several spines, and ocular peduncles that are longer than the antennal and antennular peduncles. These three species differ, however, in coloration, P. kuekenthali is generally bright orange, P. janetkae has red chelipeds and red and yellow or cream pereopods, and P. bollandi is uniformly red.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF98FFA7FEA0A263FEF01B98.taxon	materials_examined	Material examined. 2 females 1.5, 3.5 mm, 1 ovig. female 1.5 mm, Kolorai Island, north Maluku, intertidal, 16 Oct 1993 (MZB); 4 males 1.8 – 2.8 mm, 1 female 1.8 mm, Babar Island, southeast Maluku, intertidal, 11 Oct 1996; 5 males 1.8 – 2. 2 mm, 1 female 1.8 mm, 2 ovig. females 1.8, 2.5 mm, Morela, Ambon Island, Maluku, intertidal, 20 Feb 1998, (RCO).	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF98FFA7FEA0A263FEF01B98.taxon	diagnosis	Diagnosis. Biserial phyllobranchiae. Shield longer than broad, covered by plumose setae, denser on lateral borders; rostrum broadly triangular, same length or longer than lateral projections. Ocular peduncles slightly shorter than length of shield, longer than antennular and antennal peduncles; swollen proximally, corneas only slightly inflated. Ocular acicles slender distally, simple, heavily setose, separated basally. Antennal flagella shorter than shield, scarcely setose. Chelipeds subequal, densely setose, concealing surface tubercles; hiatus between dactyl and fixed finger. Second and third pereopods longer than chelipeds by entire length of dactyls; dactyls same length as propodi. Dense plumose setae on dorsal and ventral margin of pereopods. Male first and second pleopods and gonopore of male on right side only. Gonopore on coxa of third left pereopod in female; brood pouch absent. Telson asymmetrical, left larger than right, with spines on terminal margins. Color in life. “ Carapace mottled white and salmon or brown. Ocular peduncles longitudinally striped purple and white or purple and orange, stripes continuing onto cornea. Antennular peduncles with basal segment brown or purple, terminal segment blue or green; flagellum orange. Antennal flagella alternately banded brown and transparent. Chelipeds with merus mottled dark brownish purple through white; carpus and chela fairly uniform orange tan on white. Pereopods 2 and 3 with orange tan or brown longitudinal stripes on white background; merus and carpus sometimes mottled brown and white. Setation white ” (Haig & Ball, 1988).	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF98FFA7FEA0A263FEF01B98.taxon	materials_examined	Type locality. Port Essington, Australia. Habitat. This species is known from shallow waters on soft sandy bottoms, silt, rubbles, rocky shore or associated with coral (Morgan 1987, Haig & Ball 1988, Rahayu & Komai 2000, McLaughlin 2002 a). The material for this study was collected in intertidal areas on mud and sand substrates.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF98FFA7FEA0A263FEF01B98.taxon	distribution	Distribution. Thailand (Phuket), Philippines, Singapore, Indonesia (Maluku) and Australia.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF98FFA7FEA0A263FEF01B98.taxon	discussion	Remarks. When describing Paguristes monoporus, Morgan (1987) indicated that the species might warrant separation from Paguristes sensu stricto by the absence of one male gonopore. The absence of the gonopore is a variable character in P. monoporus (the male gonopore was observed on the right side only in some specimens of this study), but its removal from Paguristes is justified on other grounds. Only eight pairs biserial gills are present, the unpaired first and second male pleopods are located on the right side, and the left male pleopods in P. monoporus are unequally biramous. Therefore, P. monoporus is transferred to Pseudopaguristes. Whilst other species of the genus Pseudopaguristes have bright colors and sparse setae, the only remarkable color of P. monoporus is in the red stripes on the ocular peduncles and the bright blue antennular peduncles. Setae on the chelipeds and ambulatory legs are very dense and very often covered with mud owing to the intertidal sandy mud habitat (1 – 3 m).	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF99FFA3FEA0A443FA851CB0.taxon	materials_examined	Material examined. Holotype: 1 male, 3.5 mm, southeast of Karang Kaledupa, Snellius II, stn. 24, 05 ° 57 ' S, 123 ° 49 ' E, 350 – 500 m, 9 Sep 1984 (MZB). Paratypes: 1 male 3.5 mm, KARUBAR, stn. DW 14, 05 ° 18 ' S, 132 ° 38 ' E, 24 Oct 1991, 245 – 246 m (MNHN Pg 5933); 1 male 2.0 mm, 1 female 1.4 mm, KARUBAR, stn. DW 18, 05 ° 18 ' S, 133 ° 01 ' E, 205 – 212 m, 24 Oct 1991 (MNHN); 1 female 1.3 mm, KARUBAR, stn. DW 50, 07 ° 59 ' S, 133 ° 02 ' E, 184 – 186 m, 29 Oct 1991 (MNHN); 2 males 1.3, 1.5 mm, Madura Bay, Siboga, stn. 51, 69 – 91 m, 19 Apr 1899 (ZMA).	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF99FFA3FEA0A443FA851CB0.taxon	description	Description. Biserial phyllobranchiae. Shield (Fig. 10 a) slightly longer than broad; several tubercles anterolaterally; dorsal surface slightly convex, with scattered short setae. Rostrum broadly triangular, rounded at apex, slightly produced beyond base of ocular acicles; shorter than generally pointed lateral projections. Ocular peduncles stout, approximately 0.6 length of shield, swollen proximally. Corneas slightly dilated, diameter of corneas 0.1 – 0.2 length of ocular peduncles. Ocular acicles moderately large, separated basally, distal parts triangular and blunt at apices, mesial borders convex. Subquadrate calcified interocular lobe with 3 – 4 rounded spinules anteriorly. Antennular peduncles exceeding ocular peduncles by 0.7 – 0. 8 of ultimate segments; basal segment with small spine distomesially and distolaterally. Antennal peduncles reaching midlength of corneas; fifth segment with small spine mesially; fourth and third segments strongly produced at dorsodistal angle, terminating acutely; second segment with dorsolateral distal angle produced, terminating in pointed spine; dorsomesial distal angle with small spine; first segment short, unarmed. Antennal flagellum about 0.6 length of shield, consisting of 9 to 10 articles; each article with few short setae. Third maxilliped with small spine on ventrodistal margin of ischium; ventral margin of merus with 4 spines and 1 spine distodorsally; carpus, propodus and dactyl unarmed. Chelipeds subequal (Figs. 10 b – e), armature generally similar. Ischia each with row of small spines on ventromesial margin. Meri each with row of spines on dorsal margin, stronger distally, 1 distal transverse row of spines extending mesially and laterally; lateral faces with sparse minute tubercles, mesial faces smooth; ventromesial margins each with row of rather strong spines distally, minute spines proximally; ventrolateral margins each with row of weak spines. Carpi slightly more than half length of meri, dorsomesial margins each with 4 strong spines, dorsal surfaces each with 1 or 2 irregular rows of small, widely­spaced spines and few tufts of setae. Palms approximately 1.5 length of carpi, dorsomesial margins each with 4 strong conical spines, dorsal surfaces relatively smooth proximally, strong spines or conical tubercles distally, tending to form 3 longitudinal rows of widely­spaced spines, 2 rows continued to fixed finger, becoming smaller and closelyspaced conical tubercles and with few tufts of setae; mesial faces each with two prominent spines adjacent to ventral margin; cutting edge of fixed finger with large teeth, terminating in corneous claw. Dactyls approximately 1.5 length of palms; dorsomesial margins each with row of spinules, dorsal surfaces each with 2 irregular longitudinal rows of widelyspaced tubercles, mesial surfaces each with 4 tubercles in distal half medially, longitudinal row of tuft of setae near ventral margin, several scattered tubercles near dorsal margin; cutting edge with row of large calcareous teeth proximally, smaller teeth distally, terminating in small corneous claw; dactyl and fixed finger with small hiatus. All surface covered with sparse setae. Second and third pereopods slender, longer than chelipeds; similar from left to right. Meri of second pereopods (Fig. 11 a) with ventral margins each bearing 2 – 3 small spines distally. Carpi approximately 0.7 – 0.8 length of meri, each with 5 irregular spines on dorsal margins, lateral faces each with shallow longitudinal groove. Propodi considerably longer than carpi; each with row of 6 (left) or 8 (right) strong spines on dorsal margin subdistally and tufts of long setae; ventral margins with row of spinules (left) or smooth (right) and sparse setae; lateral faces each with shallow longitudinal groove and tufts of setae medially. Dactyls very slightly curved, 1.6 length of propodi, each terminating in strong corneous claw; ventral margins each with row of corneous spines distally, long setae along each dorsal and ventral margin. Third pereopod (Fig. 11 b, c) with meri unarmed (left) or with small spinules on dorsal margin (right). Carpi 0.8 length of meri, each with strong spine at dorsodistal angle and tufts of plumose setae along each dorsal and ventral margin. Propodi considerably longer than carpi, unarmed; lateral faces each with shallow longitudinal groove medially, dorsal and ventral margins with tufts of plumose setae. Dactyls 1.4 length of propodi, each terminating in strong corneous claw; dorsal and ventral margins each with row of corneous spines distally; tufts of long setae on each dorsal and ventral margin; mesial surfaces each with longitudinal row of corneous spines on distal half medially, and sparse setae. Fourth pereopod (Fig. 10 f) without preungual process; propodi each with 2 rows of corneous scales in propodal rasp. Male first pleopods (Fig. 10 g) small; inferior lamella rounded, distal margin with row of hook­like spines. External lobe with triangular tip, separated from internal lobe by shallow cleft; internal lobe rounded, curved towards inferior lamella. Unpaired left pleopods 3 to 5 uniramous. Female with paired gonopores; brood pouch absent. Telson (Fig. 10 h) subquadrate, left lobe slightly longer than right. Each lobe of holotype armed with 4 (left) or 5 (right) strong spines. Lobes in paratype each armed with 3 or 5 (left) and 3 or 4 (right) small spines.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF99FFA3FEA0A443FA851CB0.taxon	etymology	Etymology. From latin gracilis meaning slender, reflecting the slenderness of the second and third pereopods. Habitat. This species was collected from substrate with calcareous stones, dead coral and sponges.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF99FFA3FEA0A443FA851CB0.taxon	distribution	Distribution. South Sulawesi, Madura Bay and Kai Island, Indonesia.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF99FFA3FEA0A443FA851CB0.taxon	discussion	Remarks. Morphological character variation in P. gracilis appears to size related. The holotype is the largest specimen in the collection. In smaller individuals, the calcified interocular lobe is armed with subacute spinules anteriorly; the antennular peduncles are shorter; the antennal acicles have more spinules; and the chelipeds are more robust. This species resembles Pseudoguristes laurentae (Morgan & Forest, 1991) in general shape and spination of the chelipeds and ambulatory legs and in the form of the ocular peduncles, which are swollen proximally. The shape and spination of the telson separates the two species. P. gracilis bears four or five spines on the terminal margin of each lobe while in P. laurentae the terminal margin of each lobe is armed only with two or three strong spines. The presence of a subquadrate interocular lobe and two prominent spines on the mesial face of palm in P. gracilis also immediately distinguishes it from P. l a u re n t a e.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF9DFF9DFEA0A2ABFDB71CB8.taxon	materials_examined	Material examined. 3 males 2.5 – 3.7 mm, 1 female 3.5 mm (without abdomen), Pidjol Bay, Lombok Island, Siboga, stn. 33, 24 – 25 Mar 1899, 22 m (ZMA); 1 male 2.3 mm, Samalona, Macassar, Th. Mortensen Expedition, 29 Jun 1922, 25 m (ZMUC)	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF9DFF9DFEA0A2ABFDB71CB8.taxon	diagnosis	Diagnosis. Biserial phyllobranchiae. Shield (12 a) longer than broad, rostrum broadly triangular, same length as lateral projections, with acute apex. Ocular peduncles inflated proximally, weakly inflated distally, shorter than shield, longer than antennular and antennal peduncles. Ocular acicles elongate, with two large spines distally. Subquadrate calcified interocular lobe with 3 – 5 rounded or subacute spinules anteriorly. Antennal flagella shorter than carapace. Chelipeds (Fig. 12 b, c) subequal, right slightly larger than left, dorsal and mesial surfaces of dactyl with closely­spaced strong pointed tubercles; dactyl and fixed finger with distinct hiatus. Sparse setae on chelipeds. Pereopods (Fig. 12 d – f) similar from left to right. Dactyls of second and third pereopods 1.5 length of propodi; carpi of second pereopods each with 6 dorsal spines; carpi of third pereopods each with single dorsodistal spine. Fourth pereopod (12 g) without preungual process. Male first pleopod reduced (Fig. 12 h, i), male second pleopod absent, unpaired left pleopods 3 to 5 uniramous. Female with single left gonopore, brood pouch large. Telson (Fig. 12 j) subquadrate, left and right lobes about the same length, each armed with 4 – 6 spines, some corneous­tipped.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF9DFF9DFEA0A2ABFDB71CB8.taxon	materials_examined	Type locality. Off Manila, Philippines. Habitat. Reported from blue mud (Henderson 1888). The material in this study was collected from mud and coral.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF9DFF9DFEA0A2ABFDB71CB8.taxon	distribution	Distribution. Oman, Maldives; Red and Arabian Seas, Malaysia, Philippines, and Lombok Island and Macassar in Indonesia.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FF9DFF9DFEA0A2ABFDB71CB8.taxon	discussion	Remarks. McLaughlin & Clark (1997) redescribed the holotype of this species as Paguristes hians. They did not confirm the number of gills. The specimens in this collection agree well with the redescription, and examination of gills revealed that the species possesses eight pairs of biserial phyllobranchiae. Paguristes hians is thus transferred to Pseudopaguristes. Pseudopaguristes hians n. comb. shares one character in particular with P. gracilis: the lobe between ocular acicles is calcified and armed with several subacute or blunt spinules anteriorly.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FFA3FF99FEA0A294FD0D1DA0.taxon	materials_examined	Material examined. Holotype: male 6.0 mm, Siboga, stn. 12, 07 ° 15 ' S, 115 ° 15.6 ' E, 289 m, 14 Mar 1899 (ZMA). Paratypes: 3 females 3.4 – 5.5 mm, same locality as holotype (ZMA); 1 ovig. female 3.0 mm, Th. Mortensen Expedition 1899 ­ 1930, stn. 15, 7 ° 29 ' S, 114 ° 49 ' E, 240 m, 10 Apr 1929 (ZMUC).	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FFA3FF99FEA0A294FD0D1DA0.taxon	description	Description. Biserial phyllobranchiae. Shield (Fig. 13 a) slightly longer than broad; dorsal surface with few spinules laterally, lateral margins sloping. Rostrum broadly triangular, weakly projecting with obtuse apex, not overreaching bases of ocular acicles. Lateral projections longer than rostrum, triangular, terminating in 1 or 2 acute spines. Ocular peduncles long and slender, 0.7 – 0.8 length of shield, weakly inflated proximally; corneas not dilated, diameter 0.1 – 0.2 of peduncular length. Ocular acicles moderately large, subquadrate, with 5 – 6 terminal spines or spinules, long plumose setae at apices. Antennular peduncles exceeding ocular peduncles by half length of ultimate segment. Mesial and lateral margins of proximal segments each with strong distal spine, weaker median spine on lateral margin. Antennal peduncles reaching 0.7 – 0.8 length of ocular peduncles. Fifth and fourth segments unarmed; third segment strongly projecting ventrally, armed with 3 strong distal spines; second segment with dorsolateral distal angle produced, terminating in prominent simple or bifid spine, dorsomesial distal angle weakly produced, subdistal spine on lateral or dorsal margins; first segment with small spine on lateral margin. Antennal acicles large, reaching approximately 0.7 of fifth segment; with bifid terminal spine, mesial margin with 5 – 6 spines, lateral margin with 2 larger spines. Antennal flagellum moderately short, about the same length as shield, consisting of about 20 articles; each article with short, sparse setae. Third maxilliped with 2 strong spines and 1 smaller spine on ventrodistal margin of ischium; ventral margin of merus with 6 spines; carpus, propodus and dactyl unarmed. Chelipeds (Fig. 14 a – c) subequal, armature generally similar. Coxae each with row of small spines on ventromesial margin. Dorsal margins of meri each with row of spines, stronger distally, subdistal transverse row of spines extending mesially and laterally; lateral faces each with sparse tubercles, mesial faces smooth; ventromesial and ventrolateral margins each with row of spines and plumose setae. Carpi slightly more than half length of meri; dorsomesial margins each with 5 strong, blunt spines, dorsal surface near dorsomesial margin with strong subdistal spine followed by row of small tubercles and tuft of stiff setae, second row of small tubercles medially obscured by tufts of stiff setae. Palms approximately same length as carpi; dorsomesial margins each with 5 large spines; dorsal surfaces with more or less longitudinal rows of small tuberculate spines, becoming larger and closely­spaced on fixed fingers; tuft of long and short stiff setae on each tubercles; mesial surfaces with few small tubercles. Cutting edges of fixed fingers each with large teeth, terminating in corneous claws. Dactyls approximately same length as palms; dorsomesial margins each with rows of strong spines and few sparse tufts of setae, dorsal surface with few strong, blunt spines, few sparse tufts of long setae proximally and tufts of short setae distally; mesial surfaces covered with small conical, corneous­tipped tubercles and few tufts of setae; cutting edge with row of large calcareous teeth, terminating in rather long corneous claw, dactyl and fixed finger with distinct hiatus Second and third pereopods slightly longer than chelipeds; similar from left to right. Meri of second pereopods (Fig. 14 d) with ventral margins each bearing small spines distally. Carpi with 7 (left) or 8 (right) irregular spines on dorsal margins, 1 spine located dorsomesially. Propodi considerably longer than carpi, each with row of 7 (left) or 8 (right) strong spines and tufts of long setae on dorsal margin subdistally; ventral margins each with row of spinules and tufts of setae; lateral faces with longitudinal row of setae medially, mesial faces with scattered tufts of setae. Dactyls 1.3 length of propodi, each terminating in strong corneous claw; dorsal margins each with double rows of long setae; ventral margins each with row of corneous spines and long setae; lateral faces with tufts of setae on distal halves; mesial faces with tuft setae on proximal halves. Meri of third pereopods (Figs. 14 e, f) with dorsal margins each bearing row of spines and long plumose setae, ventral margins each with long plumose setae. Carpi 0.9 length of meri, each with strong subdistal spine on dorsal margin, long setae on dorsal margin and tuft of setae on lateral surface medially. Propodi considerably longer than carpi, dorsal margins with double rows of setae, lateral faces with tufts of setae medially, ventral margins with long setae. Dactyls 1.4 length of propodi, each terminating in strong corneous claw; dorsal margins each with double rows of long setae; ventral margins each with row of corneous spines and long setae. Fourth pereopod (Fig. 13 b) without preungual process. Male first pleopods robust (Figs. 13 d, e); inferior lamella elongate, distal margin rounded with curved lobe. External lobe slightly curved at tip, separated from internal lobe by deep narrow cleft; internal lobe broadly elongate. Unpaired left pleopods 3 to 5 uniramous. Female with single left gonopore, brood pouch large. Telson (Fig. 13 c) subquadrate, left and right lobe about same length, separated by shallow, wide V shaped median cleft; each lobe armed with spiniform setae on terminal margin, not extending onto lateral margin.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FFA3FF99FEA0A294FD0D1DA0.taxon	etymology	Etymology. From latin asper meaning rough, reflecting the surfaces of pereopods that are spinulose and rough. Habitat. Collected from sand, mud and shell fragment substrate.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FFA3FF99FEA0A294FD0D1DA0.taxon	distribution	Distribution. Bali Sea, Indonesia.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
653787B2FFA3FF99FEA0A294FD0D1DA0.taxon	discussion	Remarks. The females examined are smaller than the male holotype. Several, possibly size­related differences, such as stouter ocular peduncles with slightly convex lateral margins, and the denser, longer setae on chelipeds and pereopods. The telson and the first male pleopods of this species are remarkably distinctive: the posterior lobes of the telson are subquadrate and separated from each other by wide, shallow Vshaped median cleft, the terminal margins each are armed with spiniform setae. The distal margin of the inferior lamella of the first male pleopod forms a lobe that curves toward exterior instead of having the usual hook­like spines. These two characters will easily separate P. a s p e r from its congeners.	en	Rahayu, Dwi Listyo (2005): Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa 831: 1-42, DOI: 10.5281/zenodo.170654
