taxonID	type	description	language	source
841A744242F69D699F9D6C78B48B6D28.taxon	materials_examined	Additional material examined. Two specimens, both from Colombia. One female labeled " P'to. Berrio / Ant. Colomb " (FMNH) and one male labeled " Colombia " (HNHM).	en	Smith, Aaron D., Sanchez, Lucio A. (2015): Revision of the West Indian Wattius Kaszab (Tenebrionidae, Toxicini, Eudysantina) with lectotype designations for Pascoe's South American species. ZooKeys 537: 111-130, DOI: http://dx.doi.org/10.3897/zookeys.537.6115, URL: http://dx.doi.org/10.3897/zookeys.537.6115
841A744242F69D699F9D6C78B48B6D28.taxon	diagnosis	Diagnosis. Wattius asperulus can be separated from Wattius cucullatus and the West Indian members of the genus based on the following character combination: Frontoclypeal suture strongly incised, frons shallowly depressed anterior to eyes and raised into near vertical ridge above eyes to cranial apex; pronotal horn strongly produced, apex expanded and spatulate in males, prosternal process raised behind coxae; wings fully developed, meso- and metacoxae separated by more than mesocoxal width; femora lacking smooth rounded callosities; outer margins of tibia flat with two indistinct rows of callosities, apical spine present on all tibia in males, all male femora lacking rounded protuberances.	en	Smith, Aaron D., Sanchez, Lucio A. (2015): Revision of the West Indian Wattius Kaszab (Tenebrionidae, Toxicini, Eudysantina) with lectotype designations for Pascoe's South American species. ZooKeys 537: 111-130, DOI: http://dx.doi.org/10.3897/zookeys.537.6115, URL: http://dx.doi.org/10.3897/zookeys.537.6115
841A744242F69D699F9D6C78B48B6D28.taxon	discussion	Discussion. Wattius asperulus was synonymized under Wattius cucullatus by Champion (1886). Champion's rationale was that the separation between Wattius cucullatus and Wattius asperulus could not be maintained based on the variability displayed in the specimens, which ranged from Mexico to Argentina, available to him. However, a large series of BMNH specimens identified as Wattius cucullatus by Champion, and corresponding to many of the localities listed in the Biologia Centrali-Americana, represents an estimated seven Wattius species that are only now being described (Smith in prep.).	en	Smith, Aaron D., Sanchez, Lucio A. (2015): Revision of the West Indian Wattius Kaszab (Tenebrionidae, Toxicini, Eudysantina) with lectotype designations for Pascoe's South American species. ZooKeys 537: 111-130, DOI: http://dx.doi.org/10.3897/zookeys.537.6115, URL: http://dx.doi.org/10.3897/zookeys.537.6115
D998B8BC07A84C47A55903DE2345D0C6.taxon	materials_examined	Additional material examined. Thirteen specimens. Six labeled " BRAZIL: Sta. Catarina / Nova Teutonia / [dates from Jan. 1975, Nov. 1977, and Mar. 1977] / F. Plaumann " (CASC - 3 ♂, 2 ♀, EMEC - 1 ♂). Five labeled " Lanca / St. Cath. Brazil / Oct. 1944 " (AMNH - 1 ♂, 4 ♀). One female labeled " Cauna / S. Cath., Brazil / Dec. 1948 " (AMNH). One female labeled " Rio Vermelho / S. Cath., Brazil / I. 1946 " (AMNH).	en	Smith, Aaron D., Sanchez, Lucio A. (2015): Revision of the West Indian Wattius Kaszab (Tenebrionidae, Toxicini, Eudysantina) with lectotype designations for Pascoe's South American species. ZooKeys 537: 111-130, DOI: http://dx.doi.org/10.3897/zookeys.537.6115, URL: http://dx.doi.org/10.3897/zookeys.537.6115
D998B8BC07A84C47A55903DE2345D0C6.taxon	diagnosis	Diagnosis. Wattius cucullatus can be separated from all other known members of the genus based on the following character combination: Frontoclypeal suture strongly incised, frons shallowly depressed anterior to eyes; pronotal horn strongly produced, apex weakly bifurcated in males, prosternal process deflexed behind coxae, rarely with small projecting tubercle near apex; flight wings fully developed, meso- and metacoxae separated by more than mesocoxal width; femora lacking smooth rounded callosities; outer margins of tibia flat with two indistinct rows of callosities, apical spine absent on all tibia in males, all male femora with rounded protuberance on anterior edge of basal half.	en	Smith, Aaron D., Sanchez, Lucio A. (2015): Revision of the West Indian Wattius Kaszab (Tenebrionidae, Toxicini, Eudysantina) with lectotype designations for Pascoe's South American species. ZooKeys 537: 111-130, DOI: http://dx.doi.org/10.3897/zookeys.537.6115, URL: http://dx.doi.org/10.3897/zookeys.537.6115
0C972CD225A7A7738B4B9C587C04E603.taxon	diagnosis	Diagnosis. Wattius andersoni can be separated from the other West Indian members of the genus based on the following character combination: apterous, meso- and metacoxae separated by less than mesocoxal width; pronotal horn reduced, barely projecting past medial anterior margin of pronotum.	en	Smith, Aaron D., Sanchez, Lucio A. (2015): Revision of the West Indian Wattius Kaszab (Tenebrionidae, Toxicini, Eudysantina) with lectotype designations for Pascoe's South American species. ZooKeys 537: 111-130, DOI: http://dx.doi.org/10.3897/zookeys.537.6115, URL: http://dx.doi.org/10.3897/zookeys.537.6115
0C972CD225A7A7738B4B9C587C04E603.taxon	description	Description (Male). Length 5.4 - 5.7 mm, width 2.5 - 2.7 mm (n = 2 specimens). Body, excepting antennae, eyes, underside of head, scutellum, tarsi, and coxae generally coated with thin shellac, often capturing debris on surface. Color ferruginous to black. Head: Frons and clypeus with dense foveae, shallow to absent on clypeus, each fovea with one decumbent scale-like setae near center. Sharp setose tubercle with minute pit at apex present above eye, setae curved towards tubercle apex. Frontoclypeal suture distinct, deeply impressed; clypeus with sharp lip along anterior margin, margin straight. Epistoma between eye and clypeus raised, with one or more tubercles. Deep impression present around eye from epistoma to apex. Eye reniform; emarginate at epistoma anteriorly, lobes subequal in size, with smooth triangular callus posterior to dorsal lobe on head. Labrum with transverse medial ridge, long golden setae present from ridge to anterior margin on dorsal surface, margin straight with lateral setae on vertical surface. Mandible bifid at apex; maxillary palp four segmented, apical segment securiform; mentum trapezoidal, widest at anterior margin, faint medial longitudinal ridge present, more defined in anterior half. Antenna with distinct three segmented club, club lighter than preceding segments and tomentose, antennomeres 10 and 11 partially fused but with sinus clearly visible; antennomere 3 approximately 1.7 x length of antennomere 4, antennomeres 4 - 8 subequal in length. Prothorax: Pronotal disc convex, widest near middle; densely foveate, each fovea with one decumbent scale-like setae near center; moderately tuberculate, each tubercle bearing apical minute pit and covered in scale-like setae curved towards apex; anterior fourth of pronotum with short stout medial horn, horn margin straight; posterior fourth of pronotum with slight medial depression near scutellum; lat eral margin distinct and crenulate; anterior apices produced and acute, posterior apices acute, not projecting. Hypomeron densely deeply foveate, each fovea with one decumbent scale-like setae. Prosternum anterior to coxa short, less than length of coxal cavity, medially depressed well below height of prosternal process; prosternal process raised between coxa, apex subacute, projecting behind coxa. Pterothorax: Apterous. Elytron gradually widening to posterior third, before sharply sloping and tapering caudad; stria weakly indicated by deep elongate oval to rounded punctures, interstria with somewhat regularly spaced tubercles and decumbent scale-like setae, tubercle structure as described for those on head and pronotum; 4 th, 7 th, and 10 th interstria with tubercles forming short costae near elytral base. Scutellum glabrous and impunctate, conspicuously lacking shellac coating compared to elytron and pronotum, ~ 1.6 x as wide as long, U-shaped. Mesoventrite short, anteriorly weakly emarginate behind prosternal process, mesocoxal cavities open. Metaventrite short, separating meso- and metacoxal cavities by less than mesocoxal cavity length. All ventrites on the pterothorax with shallow indistinct punctures, often obscured by shellac, and decembent scale-like setae. Legs: Mesotrocantin exposed; femora lacking spines or other protrusions, sculpturing finely transversely rugose, with decumbent scale-like setae emerging from shallow folds; tibia clothed in decumbent scale-like setae, outer margins with indistinct rows of elongate smooth callosities, inner apical margin with socketed spurs greatly reduced to absent at base of acute weakly curved spine, small patch of golden setae present near anterior apex of protibia; tarsal formula 5 - 5 - 4, venter of distal tarsomere on all legs with sparse golden setae, venter of all other tarsomeres clothed with dense long golden setae. Abdomen: Ventrites weakly longitudinally rugose, clothed in sparse decumbent scale-like setae; abdominal intercoxal process broader than prosternal process, anterior margin straight; intersegmental membranes concealed; ventrite 5 lacking submarginal groove; abdominal defensive reservoirs present; sternite viii weakly sclerotized and setose, deeply medially emarginate, emargination V-shaped; parameres fused, sharply acuminate to apex and curved ventrad. Female. Unknown. Based on an examination of other species in the genus, the female is likely to be very similar to the male, except lacking apical spines on the tibia and emargination on sternite viii.	en	Smith, Aaron D., Sanchez, Lucio A. (2015): Revision of the West Indian Wattius Kaszab (Tenebrionidae, Toxicini, Eudysantina) with lectotype designations for Pascoe's South American species. ZooKeys 537: 111-130, DOI: http://dx.doi.org/10.3897/zookeys.537.6115, URL: http://dx.doi.org/10.3897/zookeys.537.6115
0C972CD225A7A7738B4B9C587C04E603.taxon	distribution	Distribution. Both specimens were found above 750 m in elevation in the Parque Natural Topes de Collantes, Cuba.	en	Smith, Aaron D., Sanchez, Lucio A. (2015): Revision of the West Indian Wattius Kaszab (Tenebrionidae, Toxicini, Eudysantina) with lectotype designations for Pascoe's South American species. ZooKeys 537: 111-130, DOI: http://dx.doi.org/10.3897/zookeys.537.6115, URL: http://dx.doi.org/10.3897/zookeys.537.6115
0C972CD225A7A7738B4B9C587C04E603.taxon	etymology	Etymology. The species epithet honors Robert S. Anderson, weevil expert, avid field researcher, and collector of the holotype.	en	Smith, Aaron D., Sanchez, Lucio A. (2015): Revision of the West Indian Wattius Kaszab (Tenebrionidae, Toxicini, Eudysantina) with lectotype designations for Pascoe's South American species. ZooKeys 537: 111-130, DOI: http://dx.doi.org/10.3897/zookeys.537.6115, URL: http://dx.doi.org/10.3897/zookeys.537.6115
3F9848D6C789C43109F7A69B06F08C45.taxon	diagnosis	Diagnosis. Wattius emmabaconae can be separated from the other West Indian members of the genus based on the following character combination: flight wings fully developed, meso- and metacoxae separated by more than mesocoxal width; pronotal horn strongly produced, apex strongly expanded and bifurcate in males; femora with smooth rounded callosities; outer margins of tibia with two distinct rows of elongate smooth callosities, males lacking apical spine.	en	Smith, Aaron D., Sanchez, Lucio A. (2015): Revision of the West Indian Wattius Kaszab (Tenebrionidae, Toxicini, Eudysantina) with lectotype designations for Pascoe's South American species. ZooKeys 537: 111-130, DOI: http://dx.doi.org/10.3897/zookeys.537.6115, URL: http://dx.doi.org/10.3897/zookeys.537.6115
3F9848D6C789C43109F7A69B06F08C45.taxon	description	Description (Male). Length 8.0 - 8.4 mm, width 2.8 - 3.3 mm (n = 2 specimens). Body, excepting antennae, eyes, underside of head, scutellum, tarsi, and coxae generally coated with thin shellac, often capturing debris on surface. Color ferruginous to black. Head: Frons and clypeus with dense shallow foveae, shallow to absent on clypeus, each fovea with one decumbent scale-like setae near center. Sharp setose tubercle with minute pit at apex present above eye, setae curved towards tubercle apex; one or more similar tubercles present between apex of eye and frontoclypeal margin. Frontoclypeal suture distinct, deeply impressed; clypeus with sharp lip along anterior margin, margin straight. Epistoma between eye and clypeus raised, with one or more sharp tubercles. Deep impression pre sent around eye from epistoma to apex. Eye reniform; emarginate at epistoma anteriorly, ventral lobe larger than dorsal, with micro-granulate triangular callus posterior to middle of eye. Labrum with transverse medial ridge, long golden setae present from ridge to anterior margin on dorsal surface, margin straight with setae on vertical surface. Mandible bifid at apex; maxillary palp four segmented, apical segment securiform; mentum trapezoidal, widest at anterior margin, medial longitudinal ridge present, forming anteriorly facing tooth near front margin. Antenna with distinct three segmented club, club lighter than preceding segments and tomentose, antennomeres 10 and 11 fused, with sinus visible near lateral edges, segment 9 darker at base and lightening towards apex and lateral margins; antennomere 3 approximately 1.2 x length of antennomere 4, antennomeres 4 - 8 subequal in length. Prothorax: Pronotal disc weakly convex, widest anterior to middle; densely, nearly confluently, shallowly foveate, each fovea with one decumbent scale-like setae near center; moderately tuberculate, forming irregular V-shaped pattern from near scutellum to anterior fourth, each tubercle bearing apical minute pit and covered in scale-like setae curved towards apex; anterior fourth of pronotum giving rise to raised medial horn, horn gradually sloping towards head, strongly expanded and bifid in apical third of length; posterior fourth of pronotum with slight medial depression, lacking tubercles, near scutellum; lateral margin distinct and crenulate; anterior apices strongly produced and acute, posterior apices acute, not projecting. Hypomeron densely shallowly foveate, each fovea with one decumbent scale-like setae. Prosternum anterior to coxa approximately as long as coxal cavity, medially nearly level with prosternal process; prosternal process raised between coxa, apex acute, projecting behind coxa. Pterothorax: Wings fully developed. Elytron parallel sided to posterior fourth, before sharply sloping and tapering caudad; stria weakly indicated by deep rounded punctures, interstria with somewhat regularly spaced tubercles and decumbent scale-like setae, tubercle structure as described for those on head and pronotum; 4 th, 7 th, and 10 th interstria with tubercles forming weak costae, tubercles between 4 th and 7 th interstria and elytron suture occasionally forming irregular transverse costae. Scutellum glabrous and impunctate, width approximately 1.4 x length, nearly V-shaped. Mesoventrite short, sparsely setose, anteriorly weakly emarginate behind prosternal process with submedial rows of rounded tubercles anterior to mesocoxal cavities, mesocoxal cavities open. Metaventrite long, separating meso- and metacoxal cavities by more than mesocoxal cavity length, sparsely setose with decumbent scale-like setae, impunctate. All other ventrites on the pterothorax micro-granulate, often obscured by shellac, with decembent scale-like setae. Legs: Mesotrocantin exposed; femora lacking spines or other protrusions, sculpturing finely transversely rugose with irregular smooth callosities on distal laterad half, decumbent scale-like setae emerging from shallow folds throughout; tibia clothed in decumbent scale-like setae, outer margins with two distinct rows of elongate smooth callosities, inner apical margin with socketed spurs and apical spine vestigial to absent, small patch of golden setae present near anterior apex of all tibia; tarsal formula 5 - 5 - 4, venter of distal tarsomere on all legs with sparse golden setae, venter of all other tarsomeres clothed with dense long golden setae. Abdomen: Ventrites smooth, clothed in sparse decumbent scale-like setae; abdominal intercoxal process subequal in width to prosternal process, anterior margin with small medial projection; intersegmental membranes concealed; ventrite 5 lacking submarginal groove; abdominal defensive reservoirs present; sternite viii weakly sclerotized and setose, deeply medially emarginate, emargination V-shaped; parameres fused, weakly acuminate to apex and curved ventrad. Female. Similar to male, horn very weakly expanded and bifid at apex.	en	Smith, Aaron D., Sanchez, Lucio A. (2015): Revision of the West Indian Wattius Kaszab (Tenebrionidae, Toxicini, Eudysantina) with lectotype designations for Pascoe's South American species. ZooKeys 537: 111-130, DOI: http://dx.doi.org/10.3897/zookeys.537.6115, URL: http://dx.doi.org/10.3897/zookeys.537.6115
3F9848D6C789C43109F7A69B06F08C45.taxon	distribution	Distribution. Known from only two localities in the Dominican Republic on Hispaniola.	en	Smith, Aaron D., Sanchez, Lucio A. (2015): Revision of the West Indian Wattius Kaszab (Tenebrionidae, Toxicini, Eudysantina) with lectotype designations for Pascoe's South American species. ZooKeys 537: 111-130, DOI: http://dx.doi.org/10.3897/zookeys.537.6115, URL: http://dx.doi.org/10.3897/zookeys.537.6115
3F9848D6C789C43109F7A69B06F08C45.taxon	etymology	Etymology. The specific epithet is in honor of Emma C. Bacon and was chosen by her loving partner Christiaan Harden, a generous contributor to the authors' ongoing biodiversity studies.	en	Smith, Aaron D., Sanchez, Lucio A. (2015): Revision of the West Indian Wattius Kaszab (Tenebrionidae, Toxicini, Eudysantina) with lectotype designations for Pascoe's South American species. ZooKeys 537: 111-130, DOI: http://dx.doi.org/10.3897/zookeys.537.6115, URL: http://dx.doi.org/10.3897/zookeys.537.6115
E7BE396D7BDB797A9DFD34CE84FB09FD.taxon	diagnosis	Diagnosis. Wattius viatorus can be separated from the other West Indian members of the genus based on the following character combination: flight wings fully devel oped, meso- and metacoxae separated by more than mesocoxal width; pronotal horn strongly produced, apex expanded and bifurcated in males; femora lacking smooth rounded callosities; outer margins of tibia lacking distinct rows callosities, apical spine present on all tibia in males.	en	Smith, Aaron D., Sanchez, Lucio A. (2015): Revision of the West Indian Wattius Kaszab (Tenebrionidae, Toxicini, Eudysantina) with lectotype designations for Pascoe's South American species. ZooKeys 537: 111-130, DOI: http://dx.doi.org/10.3897/zookeys.537.6115, URL: http://dx.doi.org/10.3897/zookeys.537.6115
E7BE396D7BDB797A9DFD34CE84FB09FD.taxon	description	Description (Male). Length 4.4 - 6.4 mm, width 1.8 - 2.6 mm (n = 44 specimens). Body, excepting antennae, eyes, underside of head, scutellum, tarsi, and coxae generally coated with thin shellac, often capturing debris on surface. Color ferruginous to black. Head: Frons and clypeus with dense deep foveae, somewhat shallower on clypeus, each fovea with one decumbent scale-like setae near center. Rounded setose tubercle lacking minute pit at apex present above eye, setae curved towards tubercle apex; tubercles absent between apex of eye and frontoclypeal margin. Frontoclypeal suture distinct, deeply impressed; clypeus with sharp lip along anterior margin, margin straight. Epistoma between eye and clypeus raised, rarely with one or two low tuber cles weakly indicated. Deep impression present around eye from epistoma to apex. Eye reniform; emarginate at epistoma anteriorly, ventral lobe larger than dorsal, with micro-granulate and punctate triangular callus posterior to middle of eye. Labrum with transverse medial ridge, long golden setae present from ridge to anterior margin on dorsal surface, margin straight with setae on vertical surface. Mandible bifid at apex; maxillary palp four segmented, apical segment securiform; mentum trapezoidal, widest at anterior margin, medial longitudinal ridge present, strongest near anterior margin. Antenna with distinct three segmented club, club lighter than preceding segments and tomentose, antennomeres 10 and 11 fused, with sinus visible near lateral edges; antennomere 3 approximately 1.3 x length of antennomere 4, antennomeres 4 - 8 subequal in length. Prothorax: Pronotal disc weakly convex, widest anterior to middle; densely, nearly confluently, deeply foveate, each fovea with one decumbent scale-like setae; densely tuberculate submedially, each tubercle bearing apical minute pit and covered in scale- like setae curved towards apex; anterior fourth of pronotum giving rise to raised medial horn, horn gradually sloping towards head, strongly expanded and either distinctly bifid or weakly medially emarginate in apical third of length; posterior fourth of pronotum with slight medial depression, lacking tubercles, near scutellum; lateral margin distinct and crenulate; anterior apices strongly produced and acute, posterior apices acute, not projecting. Hypomeron densely deeply foveate, each fovea with one decumbent scale-like setae. Prosternum anterior to coxa approximately as long as coxal cavity, medially nearly level with prosternal process; prosternal process raised between coxa, apex acute, projecting behind coxa. Pterothorax: Wings fully developed. Elytron parallel sided to posterior fourth, before sharply sloping and tapering caudad; stria weakly indicated by deep rounded punctures, interstria with somewhat regularly spaced tubercles and decumbent scale-like setae, tubercle structure as described for those on head and pronotum; 4 th, 7 th, and 10 th interstria with tubercles forming weak costae, tubercles between 4 th and 7 th interstria and elytron suture occasionally forming irregular transverse costae. Scutellum glabrous and impunctate, ~ 1.4 x wider than long, U- to approximately pentagonal in shape. Mesoventrite short, sparsely setose, distinctly emarginate behind prosternal process and forming submedial ridges anterior to mesocoxal cavities, mesocoxal cavities open. Metaventrite long, separating meso- and metacoxal cavities by more than mesocoxal cavity length, transversely rugose, sparsely setose with decumbent scale-like setae, moderately shallowly punctate around base of setae. All other ventrites on the pterothorax micro-granulate, often obscured by shellac, with decembent scale-like setae. Legs: Mesotrocantin exposed; femora lacking spines or other protrusions, sculpturing finely transversely rugose, lacking callosities, decumbent scale-like setae emerging from shallow folds throughout; tibia clothed in decumbent scale-like setae, outer margins lacking distinct rows of elongate smooth callosities, inner apical margin with socketed spurs vestigial to absent at base of small apical spine, patch of golden setae present on apical spines of all tibia; tarsal formula 5 - 5 - 4, venter of distal tarsomere on all legs with sparse golden setae, venter of all other tarsomeres clothed with dense long golden setae. Abdomen: Ventrites clothed in sparse decumbent scale-like setae, base of setae set in moderately sized punctures; abdominal intercoxal process wider than prosternal process, anterior margin straight to weakly rounded; intersegmental membranes concealed; ventrite 5 lacking submarginal groove; abdominal defensive reservoirs present; sternite viii weakly sclerotized and setose, deeply medially emarginate, emargination V-shaped; parameres fused, sharply acuminate to apex and weakly curved ventrad. Female. Similar to male, but lacking apical tibial spine and horn not as strongly expanded and / or bifid at apex.	en	Smith, Aaron D., Sanchez, Lucio A. (2015): Revision of the West Indian Wattius Kaszab (Tenebrionidae, Toxicini, Eudysantina) with lectotype designations for Pascoe's South American species. ZooKeys 537: 111-130, DOI: http://dx.doi.org/10.3897/zookeys.537.6115, URL: http://dx.doi.org/10.3897/zookeys.537.6115
E7BE396D7BDB797A9DFD34CE84FB09FD.taxon	distribution	Distribution. Cuba, Bahamas: South Bimini and North Andros Islands. Label data indicates that specimens have been collected between sea level and 20 meters in elevation.	en	Smith, Aaron D., Sanchez, Lucio A. (2015): Revision of the West Indian Wattius Kaszab (Tenebrionidae, Toxicini, Eudysantina) with lectotype designations for Pascoe's South American species. ZooKeys 537: 111-130, DOI: http://dx.doi.org/10.3897/zookeys.537.6115, URL: http://dx.doi.org/10.3897/zookeys.537.6115
E7BE396D7BDB797A9DFD34CE84FB09FD.taxon	etymology	Etymology. The species epithet is a noun in apposition from the Latin viator, meaning traveler or tourist, due to the distribution of the species on multiple islands considered to be vacation destinations.	en	Smith, Aaron D., Sanchez, Lucio A. (2015): Revision of the West Indian Wattius Kaszab (Tenebrionidae, Toxicini, Eudysantina) with lectotype designations for Pascoe's South American species. ZooKeys 537: 111-130, DOI: http://dx.doi.org/10.3897/zookeys.537.6115, URL: http://dx.doi.org/10.3897/zookeys.537.6115
