identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
665A87B4797CFF8FFF54FD32FE7E5145.text	665A87B4797CFF8FFF54FD32FE7E5145.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Holodentata	<div><p>Holodentata gen. nov.</p> <p>Type species: Paramunna gaussi Vanhöffen, 1914</p> <p>Species included: Holodentata gaussi (Vanhöffen, 1914), comb. nov.; H. caeca sp. nov.; and H. triangulata sp. nov.</p> <p>Diagnosis. Head with eyestalks, with or without ommatida. Lateral margins of eyestalks, pereonites, pleotelson and coxal plates surrounded with denticles. Article 3 of the antenna short and with strong denticles. Mandibular palp absent. Article 2 of maxilliped palp longest. Coxal plates visible in dorsal view in all the pereonites. Pereopod 1 carpus oval with 2 robust setae on ventral margin. Pereopods 2–7 without supplementary claw. Uropod biramous without protopod.</p> <p>Etymology. The genus name is combined from Greek holos meaning entirely and Latin dentatus meaning toothed, alluding to the denticles of the lateral margins of the pereonites, pleotelson and coxal plates.</p> <p>Remarks. The most striking character of the genus Holodentata is the fact that the second article of the maxillipedal palp is the longest article. This feature has never been reported before in any other genus of the family Paramunnidae. Four other genera of Paramunnidae show coxal plates process-like, viz., Antennulosignum Nordenstam, 1933; Austrogonium Menzies &amp; George, 1972; Bathygonium Kussakin &amp; Vasina, 1984 and Pleurosignum Vanhöffen, 1914. However, Holodentata can easily be distinguished from those genera, as all the coxal plates possess denticles. Some members of the genus Pleurogonium Sars G.O., 1883 also have coxal plates process-like with denticles, but they differ from Holodentata in the absence of eyestalks and pleotelson laterally smooth.</p> <p>Geographic distribution. Species of the genus Holodentata have only been found in the Southern Ocean; H. gaussi was originally described from Wilkes Land (385 m), H. caeca sp. nov. was collected in the Powell Basin (1584 m), Weddell Sea and H. triangulata sp. nov. was found at different stations around Cape Hallett (84–353 m), Ross Sea.</p> </div>	https://treatment.plazi.org/id/665A87B4797CFF8FFF54FD32FE7E5145	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Doti, Brenda Lía;Choudhury, Madhumita;Brandt, Angelika	Doti, Brenda Lía, Choudhury, Madhumita, Brandt, Angelika (2009): Holodentata gen. nov. (Isopoda: Asellota: Paramunnidae) with a description of two new species: H. caeca and H. triangulata from the Southern Ocean *. Zootaxa 2096 (1): 395-412, DOI: 10.11646/zootaxa.2096.1.24, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2096.1.24
665A87B4797CFF85FF54F91BFA8E50F0.text	665A87B4797CFF85FF54F91BFA8E50F0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Holodentata caeca Doti, Choudhury & Brandt 2009	<div><p>Holodentata caeca sp. nov.</p> <p>(Figs. 1–5, 10D)</p> <p>Holotype: 1 ovigerous ♀ (1.6 mm), ZMH –41970, Weddell Sea; Station 133–2, (62°46.73’S, 53°02.57’W), depth 1584 m, 16 March 2005, RV Polarstern.</p> <p>Paratypes: same locality as holotype: 25 brooding ♀ (1.2–1.5 mm), 87 preparatory ♀ (1.2–1.4 mm), 81 adult ♂♂ (0.9–1.4 mm) and 5 juveniles (0.8–0.9 mm) (ZMH –41971).</p> <p>Diagnosis. Head anterior lobe rounded and curving upward in lateral view. Eyestalk long axis points laterally, without ommatidia. Lateral margin of pereonites 1–3 expanded into a subquadrate projection, 4 rounded and 5–7 produced into a single processes. Coxal plates produced into processes.</p> <p>Description of ovigerous female (habitus description based on the holotype ZMH–41970, description of appendages on the paratype ZMH–41971). Length: 1.6 mm (Fig. 1A, B). Body width 0.6 length, widest at pereonite 3. Head width 2.2 length; anterior lobe rounded and curving upward in lateral view, margin with small denticles (broken off in the specimen illustrated, Fig. 1C). Apex of eyestalks denticulated, long axis pointing laterally, without ommatidia. Lateral margin of pereonites 1–3 expanded into a subquadrate projection, 4 rounded and 5–7 produced into single processes. Pereonite 1 about as long as pereonite 2, pereonite 2 &lt;3&gt; 4&gt; 5&lt;6 = 7 = free pleonite. Pereonite 5 shortest and pereonite 3 longest. Coxal plates produced into processes and denticulated, visible in dorsal view in all pereonites. Pleotelson width 1.2 length, marginally with 21–23 denticles on each side, apex pointed.</p> <p>Antennula (Fig. 2B), article 1 largest, with 1 simple seta and 5 robust denticles on one side; article 2 0.7 length of article 1, with 4 broom and 3 simple setae; article 3 shorter than article 2, with 1 simple seta, article 4 shortest with 1 broom seta, article 5 slightly longer than article 6 without setation; article 6 with 4 simple setae and 1 aesthetasc.</p> <p>Antenna (Fig. 2A), article 1 without setation, article 2 with 1 simple seta, article 3 with 5 robust denticles on each side and 3 denticles on distal margin, and 4 simple setae; article 4 shortest with 2 simple setae; article 5 subequal in length to article 6 with 1 broom and 2 simple setae; article 6 with 3 broom and 2 simple setae; flagellum with 8 articles, each article with numerous setae.</p> <p>Right mandible (Fig. 2C), incisor process with 5 blunt cusps (proximal one quadrate); spine row with 5 serrate setae; molar process with 1 row of teeth and 1 seta on distal edge. Left mandible (Fig. 2D) as right except for: incisor process with a 3-cusped lacinia mobilis and spine row with 4 serrate setae; molar process without setation.</p> <p>Maxillula (Fig. 2E), lateral lobe with 1 simple seta close to distal margin, 11 cuspidate setae distally (2 of these setulated) and 5 simple setae on lateral margin; mesial lobe with 4 large setulated setae distally and 4 slender simple setae on lateral margin.</p> <p>Maxilla (Fig. 2F), lateral and middle lobe protruding distomedially, with 1 pectinated and 3 simple setae distally; mesial lobe with 2 pectinated, 1 setulated and 5 simple (with pore-bearing tip) setae on distal margin, 1 setulated seta and numerous simple slender setae on mesial margin.</p> <p>Maxilliped (Fig. 3A), endite with 2 coupling hooks, distal margin with 3 simple and 2 setulated setae, ventral surface with 2 fan setae, dorsal with 3 setulated setae (see detail drawing). Epipod ovate, width 0.5 of length.</p> <p>Pereopod 1 (Fig. 3B), basis longest article with 3 simple setae. Ischium 0.6 basis length, with 3 simple setae. Merus cup-shaped, with 6 simple setae and 2 cuticular combs. Carpus oval, 0.7 ischium length; dorsal margin with 2 simple setae distally; ventral margin with 2 robust and 5 simple setae and anterior surface with 1 cuticular comb. Propodus 0.9 ischium length, dorsal margin with 3 simple setae, ventral margin with 7 simple setae, anterior surface with 5 cuticular combs and 1 simple seta. Dactylus with 2 simple setae near distal end and 2 simple setae between unguis and supplementary claw, unguis slightly shorter than dactylus, supplementary claw 0.7 unguis length.</p> <p>Pereopods 2–7 (Figs. 3C, D; 4). Bases with 2–4 simple setae. Ischia with 3–4 simple setae. Meri with 3–5 simple setae. Carpi with 2 simple setae at half length of article, and 3–5 simple setae and 1 broom seta on distal end, carpus of pereopod 7 with 6 cuticular combs. Propodi with 2–3 simple setae at half length of article, and 3–4 simple setae and 1 broom seta on distal end. Dactyli with 4–6 simple setae, unguis slightly longer than dactylus, supplementary claw absent.</p> <p>Operculum (Fig. 5A) ovoid and pointed distomedially, width 0.9 length; lateral margins with several simple setae (many of these broken).</p> <p>Pleopod 3 (Fig. 5B), endopod width 0.6 length, with 3 plumose setae distally. Exopod with 2 articles, distal one with 1 distal simple seta, which extends beyond the tips of the endopod setae.</p> <p>Pleopod 4 (Fig. 5C), exopod reaching half length of endopod (endopod folded in the specimen illustrated).</p> <p>Pleopod 5 (Fig. 5D) width 0.5 length.</p> <p>Uropod (Fig. 5E) exopod 0.2 endopod length, with 2 simple setae distally; endopod with 3 simple setae subapically and 5 broom setae distally.</p> <p>Description of adult male (paratype ZMH–41971). Pleopod 1 (Fig. 5F) maximum width 0.6 length; lateral lobes at 0.7 of its length from proximal end, each one with 6 simple setae; ventral surface with 4 simple setae (2 of these close to lateral margin).</p> <p>Pleopod 2 (Fig. 5G), protopod width 0.4 length, with 18 simple setae on lateral margin. Endopod slightly surpassing protopod. Exopod bilobed.</p> <p>Remarks. H. caeca sp. nov. is most similar to H. triangulata sp. nov.; the main differences between these two species are discussed in the remarks section of the latter one.</p> <p>Distribution. Only known from type locality.</p> <p>Etymology. The species name is Latin caeca meaning blind and refers to the absence of ommatidia.</p> </div>	https://treatment.plazi.org/id/665A87B4797CFF85FF54F91BFA8E50F0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Doti, Brenda Lía;Choudhury, Madhumita;Brandt, Angelika	Doti, Brenda Lía, Choudhury, Madhumita, Brandt, Angelika (2009): Holodentata gen. nov. (Isopoda: Asellota: Paramunnidae) with a description of two new species: H. caeca and H. triangulata from the Southern Ocean *. Zootaxa 2096 (1): 395-412, DOI: 10.11646/zootaxa.2096.1.24, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2096.1.24
665A87B47977FF82FF54FF15FEB45328.text	665A87B47977FF82FF54FF15FEB45328.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Holodentata triangulata Doti, Choudhury & Brandt 2009	<div><p>Holodentata triangulata sp. nov.</p> <p>(Figs. 6–9, 10A–C)</p> <p>Holotype: 1 adult ♀ (ZMH –41972); Ross Sea, RV Italica, station H out 4, 72°18.5’S, 170°26.8’E, 235 m depth, 12 Feb 2004.</p> <p>Paratypes: same locality as holotype: 25 brooding ♀♀ (1.4–1.8 mm), 26 preparatory ♀♀ (1–1.7 mm), 12 adult ♂♂ (1.1–1.5 mm) and 7 juveniles (0.9–1 mm); (ZMH –41973). Station H out 2, 72°17.5’S, 170°29.4’E, 353 m depth, 11 Feb 2004: 7 brooding ♀♀ (1.5–1.7 mm), 5 preparatory ♀♀ (1.1–1.5 mm), 3 juvenile ♀♀ (0.9–1 mm) and 3 adult ♂♂ (1.1–1.2 mm). Station H in 3, 72°17.0’S, 170°13.1’E, 316 m depth, 16 Feb 2004: 3 preparatory ♀♀ (1–1.4 mm) and 2 adult ♂♂ (1.4–1.5 mm). Station H in 4, 72°17.1’S, 170°14.0’E, 196 m depth, 16 Feb 2004: 5 brooding ♀♀ (1.3–1.7 mm), 9 preparatory ♀♀ (1–1.6 mm), 2 adult ♂♂ (1.2–1.3mm) and 7 juveniles (0.8–1 mm). Station H in 5, 72°17.2’S, 170°17.9’E, 84 m depth, 16 Feb 2004: 4 preparatory ♀♀ (1.3–1.4 mm), 1 juvenile ♀ (0.9 mm) and 2 adult ♂♂ (0.9–1.1 mm). Station H out 1, 72°14.8’ S, 170°15.2’E, 542 m depth, 9 Feb 2009: 4 preparatory ♀♀ (1.2–1.6 mm) and 1 juvenile ♀ (1 mm).</p> <p>Diagnosis. Head anterior lobe triangular with blunt apex, and curving upward in lateral view. Eyestalk long axis pointing laterally, with 4 ommatidia. Lateral margin of pereonites 1–3 expanded into a subquadrate projection, 4 rounded and 5–7 produced into single processes. Coxal plates produced into processes. Pereopod 7, carpus with 1 robust seta distoventrally.</p> <p>Description of brooding female (habitus description based on the holotype ZMH–41972, description of appendages on the paratype ZMH–41973). Length: 1.9 mm (Fig. 6A, B). Body width 0.6 length, widest at pereonite 3. Head width 2.4 length; anterior lobe triangular with blunt apex, curving upward in lateral view, marginally denticulated (Figs. 6C, 10B). Eyestalks long axis pointing laterally, with 4 ommatidia. Lateral margin of pereonites 1–3 expanded into a subquadrate projection, 4 rounded and 5–7 produced into single processes. Total length of all pereonites and free pleonite combined subequal to pleotelson length. Coxal plates produced into processes and denticulated, visible in dorsal view in all pereonites. Pleotelson width 1.2 length, marginally with 20–25 denticles on each side, apex truncate.</p> <p>Antennula (Fig. 7A), article 1 largest, with 1 broom seta, and 3 and 1 robust denticles on each side; article 2 0.6 length of article 1, with 2 broom and 3 simple setae; article 3 0.7 length of article 2 and subequal in length to article 5 (and 6); article 4 shortest, with 1 simple seta; article 6 with 3 simple setae and 1 aesthetasc.</p> <p>Antenna (Figs. 7B, 10C), articles 1 and 2 glabrous; article 3 short distally broadened, with 2 simple setae and several robust denticles; article 4 with 1 simple seta; article 5 subequal in length to article 6, with 1 simple seta; article 6 with 3 broom and 3 simple setae; flagellum with 9 articles, each article with numerous setae.</p> <p>Right mandible (Fig. 7C), incisor process with 4 blunt cusps (proximal one quadrate); spine row with 5 serrate setae; molar process with 1 row of teeth and 1 serrate seta on distal edge. Left mandible (Fig. 7C detail) as right except for: incisor process with a 4-cusped lacinia mobilis (1 of these cusps minute) and spine row with 4 serrate setae; molar process without setation.</p> <p>Maxillula (Fig. 7D), lateral lobe with 1 simple seta close to distal margin and 12 cuspidate setae distally (2 of these setulated), mesial lobe with 5 setulated setae distally and 4 slender simple setae.</p> <p>Maxilla (Fig. 7E), lateral and middle lobe protruding distomedially, with 1 pectinated and 3 simple setae distally; mesial lobe with 2 pectinated, 1 setulated and 6 simple (with pore-bearing tip) setae on distal margin, 1 setulated seta and numerous slender simple setae on mesial margin.</p> <p>Maxilliped (Fig. 7F), endite with 2 coupling hooks, distal margin with 4 setulated setae, ventral surface with 2 fan setae and 1 setulated seta, dorsal surface with 3 setulated setae (see detail). Epipod ovate, width 0.6 of length.</p> <p>Pereopod 1 (Fig. 8A, B), basis longest article with 3 simple setae. Ischium 0.5 basis length, with 2 simple setae and 1 cuticular comb. Merus cup-shaped, with 5 simple setae and 2 cuticular combs. Carpus oval, 0.8 ischium length; dorsal margin with 1 simple seta distally; ventral margin with 2 robust and 6 simple setae, anterior surface with 1 cuticular comb. Propodus 0.8 ischium length, dorsal margin with 4 simple setae (1 at half length article and 3 distal), ventral margin with 7 simple setae, anterior surface with 1 simple seta and 4 cuticular combs. Dactylus with 3 simple setae near distal end and 2 simple setae between unguis and supplementary claw, unguis slightly shorter than dactylus, supplementary claw 0.8 unguis length.</p> <p>Pereopods 2–7 (Figs. 8C–E, 9A–C). Bases with 2–3 simple setae. Ischia with 2–4 simple setae. Meri with</p> <p>3–5 simple setae. Carpi with 2 simple setae at half length article, and 3–6 simple setae and 1 broom seta on distal end; carpus of pereopod 7 with 1 robust seta distoventrally and some cuticular combs. Propodi with 2–3 simple setae half way along the length of article, 2–4 simple setae and 1 broom seta on distal end; propodus of pereopod 7 with some cuticular combs. Dactyli with 4–5 simple setae, unguis slightly longer than dactylus, supplementary claw absent.</p> <p>Operculum (Fig. 9D) ovoid and acuminating distomedially, width 0.9 length; lateral margins with several simple setae (many of these broken).</p> <p>Pleopod 3 (Fig. 9E), endopod width 0.6 length, with 3 plumose setae distally. Exopod with 2 articles; distal one with 3 minute setae and 1 distal simple seta, which extends towards the tips of the endopod setae.</p> <p>Pleopod 4 (Fig. 9F), endopod width 0.5 length, exopod reaching half length of endopod.</p> <p>Pleopod 5 (Fig. 9G) width 0.5 length.</p> <p>Uropod (Fig. 9H) exopod 0.3 endopod length, with 2 simple setae distally; endopod with 3 simple setae subapically and 5 broom setae distally.</p> <p>Description of adult male (paratype ZMH–41973). Pleopod 1 (Fig. 9I) maximum width 0.6 length; lateral lobes at 0.7 of its length from proximal end, each one with 6–7 simple setae; distal projection with 2 simple setae; ventral surface with 4 simple setae (2 of these close to lateral margins).</p> <p>Pleopod 2 (Fig. 9J), protopod width 0.4 length, with 17 simple setae on lateral margin. Endopod slightly surpassing protopod. Exopod bilobed.</p> <p>Remarks. H. triangulata is most similar to H. caeca, but it can be easily distinguished from the latter by having (features dealing with H. caeca are in parentheses): Head anterior lobe triangular (anterior lobe rounded), eyestalks with ommatidia (blind), antenna article 3 with denticles on one side (article 3 with denticles on both margins), carpus pereopod 7 with one distal robust seta on ventral margin (robust seta absent).</p> <p>Distribution. Only known from type locality.</p> <p>Etymology. The epithet is derived from Latin triangulus meaning triangular alluding to the triangular shape of the head.</p></div> 	https://treatment.plazi.org/id/665A87B47977FF82FF54FF15FEB45328	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Doti, Brenda Lía;Choudhury, Madhumita;Brandt, Angelika	Doti, Brenda Lía, Choudhury, Madhumita, Brandt, Angelika (2009): Holodentata gen. nov. (Isopoda: Asellota: Paramunnidae) with a description of two new species: H. caeca and H. triangulata from the Southern Ocean *. Zootaxa 2096 (1): 395-412, DOI: 10.11646/zootaxa.2096.1.24, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2096.1.24
