taxonID	type	description	language	source
607F1804682E5F3179D0FDA3FC88FEA3.taxon	materials_examined	Type series. Holotype: male (29 - kas – IBSS), Russia, Khabarovsk Territory, Komsomolsky District, Komsomolsk – on – Amur – Nikolaevsk – on – Amur road, near road bridge, the Amur River basin, the Axyan River, N 50 ° 46.266 ’; E 138 ° 05.471 ’; depth 60 cm; substrates: stones, pebbles. 0 1.07.2005, coll. K. A. Semenchenko, D. A. Sidorov.	en	Tuzovskij, Petr V., Semenchenko, Ksenia A. (2009): New water mites species of the genus Feltria Koenike (Acariformes, Feltriidae) from the Asian Russia. Zootaxa 2124: 38-50, DOI: 10.5281/zenodo.188186
607F1804682E5F3179D0FDA3FC88FEA3.taxon	description	Description. Male. Body flat and oval. Dorsal shield very large, covering almost all dorsal surface, supplied with five pairs of setae: Oi, Oe, Sci, Li, Si (Fig. 1). Setae Fp, Hi, Oi, Oe, Sci, Li, Si without glandularia, other dorsal idiosomal setae located on smooth interscutal membrane around the dorsal shield. Setae Fch, Vi, Ve, He, Sce and Le provided by glandularia and located on small separate platelets. Setae Fp and Hi situated on very small sclerites. Anterior and posterior margins of dorsal shield convex, with lateral margins more or less parallel to each other. All lyriform organs placed on dorsum: i 1 on platelets, bearing setae Vi, i 2 between setae Ve and Hi, i 3 behind setae He, i 4 behind setae Sce, and i 5 behind setae Le. Leg coxae incorporated into four groups, situated closely but not fused to each other, and occupied about two thirds of venter surface (Fig. 2). Second coxae not extending to midline, posterior apodemes of first coxae short and posterolaterally directed. Medial pair of setae (Pe) located between fourth coxae and genital plate, lacking glandularia and situated on posterior extensions of fourth coxae; lateral pair of setae (Se) with glandularia and situated on posterior extensions of fourth coxae too. Genital plate large and separate from posterior extensions of fourth coxae; acetabula 18 on one side and 19 on the other side of genital plate. Ventrolateral platelets free and extending between fourth coxae and genital plate. Gonoporus located near anterior end of genital plate. Pregenital sclerite very small, located between posteromedial corners of fourth coxae. Excretory pore, setae Pi and setae Ci with glandularia situated free on smooth interscutal membrane behind genital plate. Pedipalp slender (Fig. 3). Trochanter relatively short, with single dorsodistal seta. Femur with three thick proximal and two thin dorsodistal setae, ventrally with proximoventrally directed projection, bearing rather numerous sharp-pointed spines (Fig. 4). Genu with two thin dorsodistal setae. Tibia very long and narrow with two unequal ventral setae and one short distolateral spine. Surface of femur and genu with cell-shaped reticulation, surface of tibia with strips. First four segments of all legs provided with thick setae (Fig. 5). Tarsus of leg III straight and slightly expanded in a median portion, with small flat ventrolateral ledge, bearing 6 – 8 short modified setae with flat or rounded tips and one relatively long thin seta (Fig. 6 – 7). Claws (Fig. 8) with three dents: large median, short and thin internal and external ones. Ventral margin of claw plate concave. Measurements (n = 1). Length of body 455, width 318; length of dorsal shield 378, width 252; length of seta Fch 30; length of genital plate 155, width 270; length of pedipalpal segments (P 1 – 5): 36, 66, 63, 123, 27; length of coxae I – IV 312; length of leg segments: I — 45, 60, 60, 78, 102, 96; II — 45, 63, 54, 72, 102, 90; III — 50, 48, 60, 78, 110, 102; IV — 78, 75, 72, 93, 120, 96. Differential diagnosis. The new species is similar to the European species Feltria denticulata Angelier, 1949 and to the North American subspecies F. echinopalpis echinopalpis Cook, 1963 and F. echinopalpis projecta Cook, 1970. All coxal groups in F. echinopalpis echinopalpis are fused without trace of a suture line, the tarsus of leg III is expanded with six pointed modified setae (Cook 1963); in contrast, in F. aculeata sp. n. all coxal groups are separated, the tarsus of leg III is slightly expanded with 6 – 8 blunt setae. In F. echinopalpis projecta the excretory pore plate and the setae Ci with associated glandularia are fused with the genital plate, the tarsus of leg III is expanded and curved with five partially fused short setae (Cook 1970); in F. aculeata sp. n. the excretory pore platelets and the setae Ci with associated glandularia are not fused with the genital plate, the tarsus of leg III straight and slightly expanded with 6 – 8 separated short setae. F. aculeata sp. n. differs from F. denticulata in the following characters (character states of F. denticulata are indicated in parentheses from Gledhill 1983): the anterior coxal groups are separated (fused to each other), the medial pair of setae (Pe) between the fourth coxae and the genital field without glandularia (with glandularia), the lateral pair of setae (Se) with glandularia is fused with the posterior extension of the coxae IV (lying free on smooth membrane between the fourth coxae and the genital field), the excretory pore and the setae Ci with glandularia are lying free (fused with the genital field), the tarsus of leg III is slightly expanded with 6 – 8 blunt spines (well expanded with 4 – 5 pointed spines). It should be noted that the new subspecies has more common features with the North American subspecies F. echinopalpis echinopalpis and F. echinopalpis projecta, than with the European species F. denticulata. However, it can be distinguished from all these species by the following features: the tarsus of leg III slightly expanded; the excretory pore and associated setae (Pi and Ci) with glandularia are situated free on smooth interscutal membrane behind the genital plate.	en	Tuzovskij, Petr V., Semenchenko, Ksenia A. (2009): New water mites species of the genus Feltria Koenike (Acariformes, Feltriidae) from the Asian Russia. Zootaxa 2124: 38-50, DOI: 10.5281/zenodo.188186
607F1804682E5F3179D0FDA3FC88FEA3.taxon	etymology	Etymology. Named for its proximoventrally directed projection on the femur of pedipalp, bearing sharppointed spines (from lat. “ aculeáta ” – spiny). Habitat. Running waters.	en	Tuzovskij, Petr V., Semenchenko, Ksenia A. (2009): New water mites species of the genus Feltria Koenike (Acariformes, Feltriidae) from the Asian Russia. Zootaxa 2124: 38-50, DOI: 10.5281/zenodo.188186
607F1804682E5F3179D0FDA3FC88FEA3.taxon	distribution	Distribution. Khabarovsk Territory, Far East of Russia.	en	Tuzovskij, Petr V., Semenchenko, Ksenia A. (2009): New water mites species of the genus Feltria Koenike (Acariformes, Feltriidae) from the Asian Russia. Zootaxa 2124: 38-50, DOI: 10.5281/zenodo.188186
607F1804682B5F3579D0FF36FCFDFEA3.taxon	materials_examined	Type series. Holotype: female (9493 – IBIW), Russia, Primory Territory, Khasansky District, “ Kedrovaya Pad National Nature Biosphere Reserve ”, Sea of Japan basin, the Kedrovaya River, N 43 ° 06.056 ’; E 131 ° 33.310 ’; depth 18 – 30 cm; substrates: stones, pebbles. 25.11.1993, coll. T. M. Tiunova. Paratypes: 10 females, 1 male (9490 - 9496 – IBIW); 2 females (27 - 28 - kas – IBSS). Paratypes with same locality and date as holotype.	en	Tuzovskij, Petr V., Semenchenko, Ksenia A. (2009): New water mites species of the genus Feltria Koenike (Acariformes, Feltriidae) from the Asian Russia. Zootaxa 2124: 38-50, DOI: 10.5281/zenodo.188186
607F1804682B5F3579D0FF36FCFDFEA3.taxon	description	Description. Female. Body flat. Dorsum with large shield and seven pairs of platelets (Fig. 9). Median portion of frontal margin straight. Dorsal shield with four pairs of setae: Oi, Oe, Sci and Li. Setae Fp, Oi and Pi without glandularia, other idiosomal setae associated with glandularia. Dorsal shield with two median furrows and three pairs of small lateral depressions; these furrows various in shape and usually fused to each other caudally. Setae Fch, Fp, Vi, Ve and first pair of lyriform organs (i 1) located on narrow anterior plates; setae Hi, He, Sce and Le placed on separate lateral sclerits; second pair of lyriform organs (i 2) fused with sclerits, bearing setae Hi. Setae Si placed on relatively large triangular posterior plates. One pair of small platelets located anterolaterally to triangular plates. Three pairs of lyriform organs (i 3 – i 5) usually lying free on smooth membrane, but i 4 can be fused with platelets, bearing setae Sce. Anterior coxal groups fused medially, but suture line between coxae I present; posterior coxal groups close together but not fused medially (Fig. 10). Posterior apodemes of first coxae short and directed more or less laterally. Second coxae not extending to midline. Acetabular plates fused with each other, but trace of a suture line present, its anterior margins forming blunt angle; acetabula very numerous and located mainly in their lateral parts. Gonoporus located in posterior half of acetabular plates and surrounded by a frame, bearing three pairs of anterior setae and one pair of posterior setae. Median portions of acetabular plates fused with posterior extensions of fourth coxae, but trace of a suture line between them present. Pregenital sclerite very small, located between posteromedial corners of posterior extensions of fourth coxae and anterior margin of acetabular plates. Setae Pe and Se and associated glandularia fused with posterior extensions of fourth coxae. Excretory pore platelets free, setae Pi and setae Ci fused with acetabular plate. Capitulum with short anchoral projection. Pedipalp moderately slender (Fig. 11). Trochanter relatively short, with single dorsodistal seta. Femur expanded, with two proximal and three thick dorsodistal setae, its ventral margin convex. Genu with one proximal and two dorsodistal setae. Tibia long and tapering distally, with two unequal ventral setae and one short distolateral spine. Surface of femur and genu with cell-shaped reticulation, surface of tibia with strips. First four segments of all legs usually with thick setae (Fig. 12). Claws with three dents: large median, short internal and external ones, but internal claw thicker than external one. Ventral margin of claw plate concave. Measurements (n = 5). Length of body 510 – 540, width 390 – 430; length of dorsal shield 420 – 435, width 310 – 340; length of seta Fch 18 – 24; length of genital plate 190 – 210, width 360 – 395; length of capitulum 90 – 96; length of pedipalpal segments (P 1 – 5): 30 – 36, 65 – 72, 48 – 60, 118 – 125, 18 – 27; length of coxae I – IV 360 – 395; length of leg segments: I — 42 – 54, 54 – 65, 60 - 64, 69 – 72, 70 – 85, 75 – 90; II — 48 – 54, 57 – 66, 52 – 55, 65 – 68, 82 – 90, 85 – 95; III — 60 – 72, 48 – 66, 54 – 66, 75 – 90, 100 – 110, 95 – 102; IV — 70 – 95, 65 – 75, 78 – 85, 105 – 115, 120 – 126, 78 – 100. Male. Dorsum with large shield and five pairs of platelets (Fig. 15). Median portion of frontal margin convex. Dorsal shield with five pairs of setae: Oi, Oe, Sci, Li and Si; setae Oi thin, other setae thick. Setae Fp, Oi and Pi without glandularia, other idiosomal setae associated with glandularia. Dorsal shield with two pairs of median furrows and three pairs of small lateral depressions; furrows of anterior and posterior pairs fused to each other distally. Setae Fch, Fp, Vi, Ve and first pair of lyriform organs (i 1) located on long, narrow anterior platelets; setae Hi and i 2 placed on first lateral platelets, setae He, Sce and Le placed on separated lateral platelets too. Fourth pairs of lyriform organs (i 4) located on plates, bearing setae Sce; i 3 and i 5 lying free on smooth membrane. Anterior coxal groups fused medially, but suture line between coxae I present; posterior coxal groups close together but not fused medially (Fig. 16). Posterior apodemes of first coxae moderately long and directed more or less laterally. Second coxae not extending to midline. Anterior margin of acetabular plates forming blunt angle; acetabula numerous. Gonoporus small and narrow, located in anterior half of acetabular plates, with two pairs of perigenital setae. Median portion of acetabular plate fused with posterior extensions of fourth coxae, but trace of suture line between them present. Pregenital sclerite very small, located between posteromedial corners of posterior extensions of fourth coxae and anterior margin of acetabular plate. Setae Pe and Se and associated glandularia fused with posterior extensions of fourth coxae. Excretory pore platelet fused with acetabular plate. Setae Pi and setae Ci not fused with acetabular plate. Capitulum with short posterior projection. Pedipalp similar to that of female, but femur with proximoventral protrusion (Fig. 13). Tibia of leg III (Fig. 14) with five unequal setae, tarsus with large ventrolateral protrusion, bearing six pointed setae. Measurements (n = 1). Length of body 485, width 355; length of dorsal shield 438, width 288; length of seta Fch 24; length of genital plate 198, width 270; length of capitulum 95; length of pedipalpal segments (P 1 – 5): 30, 69, 57, 120,?; length of coxae I – IV 385; length of leg segments: I — 50,?,?,?,?,?; II — 54, 60, 57, 69, 102,?; III — 65, 60, 78, 84, 120, 90; IV — 85, 72, 78, 102, 126, 102. Differential diagnosis. The combination of the following characters separate F. ussuriensis sp. n. from all other species of the subgenus Feltria: the dorsal shield with longitudinal medial furrows and three pairs of the lateral depressions, long pedipalpal tibia, the female’s acetabular plates are fused to each other, and the gonoporus is surrounded by a frame and located in the posterior half of the genital field.	en	Tuzovskij, Petr V., Semenchenko, Ksenia A. (2009): New water mites species of the genus Feltria Koenike (Acariformes, Feltriidae) from the Asian Russia. Zootaxa 2124: 38-50, DOI: 10.5281/zenodo.188186
607F1804682B5F3579D0FF36FCFDFEA3.taxon	etymology	Etymology. The species epithet ussuriensis is derived from the former historical name of the region (Ussuri region) where it was collected. Habitat. Running waters.	en	Tuzovskij, Petr V., Semenchenko, Ksenia A. (2009): New water mites species of the genus Feltria Koenike (Acariformes, Feltriidae) from the Asian Russia. Zootaxa 2124: 38-50, DOI: 10.5281/zenodo.188186
607F1804682B5F3579D0FF36FCFDFEA3.taxon	distribution	Distribution. Primory Territory, Far East of Russia.	en	Tuzovskij, Petr V., Semenchenko, Ksenia A. (2009): New water mites species of the genus Feltria Koenike (Acariformes, Feltriidae) from the Asian Russia. Zootaxa 2124: 38-50, DOI: 10.5281/zenodo.188186
607F180468285F3979D0FA7AFB1AFD07.taxon	materials_examined	Type series. Holotype: female (9498 – IBIW), Russia, Khabarovsk Territory, Chegdomynsky District, the Bureya River basin, the Mylny spring, N 51 ° 01.480 ’; E 132 ° 24.481 ’; depth 20 cm; substrates: boulders, stones, pebbles, moss. 22.09.2005, coll. D. A. Sidorov. Paratypes: 1 female (30 - kas – IBSS), 1 male (9498 – IBIW) with same locality and date as holotype; 1 male (31 - kas – IBSS) Russia, Sakha Republic (Yakutia), Neryngrynsky District, the Lena River basin, headwater of the Bolshoy Duray River, N 57 ° 12.237 ’; E 124 ° 48.314 ’; depth 40 cm; substrates: stones, small-grained sand. 0 2.08.2006, coll. K. A. Semenchenko, D. A. Sidorov; 1 male (9499 – IBIW), Russia, Sakha Republic (Yakutia), Nyzhnekolymsky ulus, Letyatkin Cape, the Kolyma River basin, unnamed spring. 22.08.2008, coll. H. Abé.	en	Tuzovskij, Petr V., Semenchenko, Ksenia A. (2009): New water mites species of the genus Feltria Koenike (Acariformes, Feltriidae) from the Asian Russia. Zootaxa 2124: 38-50, DOI: 10.5281/zenodo.188186
607F180468285F3979D0FA7AFB1AFD07.taxon	description	Description. Female. Dorsum with central shield and four pairs of platelets (Fig. 17). Dorsal shield relatively small (ratio length / width = 1.07 – 1.16), cross-shaped, with well developed lateral protrusion and bearing two pairs of setae: Oi and Sci. Setae Fch, Fp, Vi, Ve, Hi, He, Sce, Li, Le, Ci and Pi located free on smooth integument; setae Oe placed on first pair of lateral plates, setae Si on fourth pair of lateral plates. Setae Fp, Oi and Pi without glandularia, other idiosomal setae associated with glandularia. Setae Fch, Oi and Sci thick, other idiosomal setae thin. First and second pairs of lateral plates nearly equal in size, forth pair of lateral plates much larger than third pair of lateral ones. First pair of lyriform organs (i 1) placed on platelets, bearing setae and glandularia Vi, other lyriform organs (i 2 – i 5) lying free on smooth integument along lateral portions of body. Excretory pore plate slightly broader than long, located dorsally. Setae Pi and Ci separated. All coxal groups separated (Fig. 18). Posterior apodemes of first coxae short and directed laterally. Second coxae not extending to midline. Posterior extensions of fourth coxae slightly developed. Acetabular plates separated, with 26 – 33 genital acetabula and three thin median setae (two proximal and one distal) on each side. Acetabular plates with distinctly developed posteromedial corners. Pregenital sclerite small, located between posteromedial corners of posterior extensions of fourth coxae and platelets, bearing setae Pe. Anterior genital sclerite slightly broader than long. Setae Pe and Se with associated glandularia lying free on smooth integument between posterior extensions of fourth coxae and genital area. Ventrolateral plates narrow, transverse and lying free between fourth coxae and acetabular plates. Pedipalp (Fig. 19) rather slender. Trochanter relatively short, with single dorsodistal seta. Femur slightly expanded, with three proximal and three thick dorsodistal setae, its ventral margin is straight. Genu with one proximal and two dorsodistal setae. Tibia ventrally with small ventrolateral projection near middle, bearing proximal seta, distal seta without tubercle located more distal. Measurements (n = 2). Length of body 450 – 475, width 395 – 435; length of dorsal shield 265 – 303, width 228 – 283; length of seta Fch 54; length of genital plates 115 – 119, width 132 – 163; length of capitulum 90; length of pedipalpal segments (P 1 – 5): 25 – 27, 36 – 44, 36 – 41, 60 – 71, 37 – 39; length of coxae I – IV 180 – 264; length of leg segments: I — 27 – 36, 37 – 54, 42 – 44, 60 – 65, 72 – 82, 78 – 92; II — 34 – 36, 38 – 51, 47 – 48, 60 – 64, 78 – 85, 81 – 95; III — 40 – 42, 41 – 48, 54 – 66, 69 – 75, 90 – 98, 90 – 102; IV — 69 – 75, 51 – 54, 60 – 64, 90 – 98, 108 – 122, 93 – 114. Male. Body flat and almost rounded (ratio length / width without coxae 1.02). Dorsum with large shield (Fig. 25). Anterior margin convex. Dorsal shield with five pairs of setae: Oi, Oe, Sci, Li and Si; setae Oi without glandularia, other setae associated with glandularia; setae Oe thicker than other setae on this shield. Dorsal shield slightly elongated, ratio length / width = 1.2 – 1.3. Setae Fch long and thicker than other dorsal setae. Setae Vi and first pair of lyriform organs located on common platelets; other setae and lyriform organs (i 2 – i 5) lying free on smooth integument. Setae Hi thicker than He. All coxal groups separated (Fig. 26). Posterior apodemes of first coxae moderately long and directed more or less laterally. Second coxae not extending to midline. Anterior margin of acetabular plate convex; 27 – 32 acetabula on each side. Gonoporus small, narrow, located near anterior margin of acetabular plates, with 2 – 6 perigenital setae. Pregenital sclerite small, located between posteromedial corners of posterior extensions of fourth coxae and anterior margin of acetabular plate. Setae Pe and Se lying free between posterior extensions of fourth coxae and anterior margin of acetabular plate. Ventrolateral plates narrow, transverse and lying free between fourth coxae and acetabular plate. Excretory pore platelets, setae Pi and Ci not fused with acetabular plate (Fig. 20). Pedipalp relatively short (Fig. 21). Trochanter with single dorsodistal seta. Femur expanded with three proximal and three dorsodistal setae, its ventral margin straight. Genu with one proximal and two dorsodistal setae. Tibia ventrally with well developed, curved, sharply-pointed projection near middle, much smaller setal tubercle located more distal, and with one thin dorsodistal seta. Tibia of leg III (Fig. 22) with four unequal thick distal setae and several thin setae; tarsus with moderately developed ventrolateral protrusion, bearing three short blunt setae (Fig. 23). Leg claws (Fig. 24) with three dents: large median, short and thin internal and external ones. Ventral margin of claw plate straight or slightly concave. Measurements (n = 2). Length of body without coxae 310 – 330, width 285 – 310; length of dorsal shield 280 – 290, width 215 – 235; length of seta Fch 50; length of genital plate 108 – 125, width 204 – 228; length of capitulum 90 – 95; length of pedipalpal segments (P 1 – 5): 21 – 24, 47 – 49, 36 – 39, 60 – 63, 36 – 39; length of coxae I – IV 240 – 245; length of leg segments: I — 30 – 35, 42 – 48, 39 – 42, 54 – 60, 65 – 72, 72 – 78; II — 30 – 38, 39 – 42, 36 – 39, 54 – 57, 65 – 72, 78 – 81; III — 42 – 45, 36 – 39, 45 – 48, 60 – 63, 83 – 85, 78 – 84; IV — 70 – 73, 48 – 51, 54 – 60, 78 – 81,100 – 105, 95 – 96. Differential diagnosis. The present subspecies is closely related to F. cornuta cornuta Walter, 1927 but differs from it in the following characters (character states of F. cornuta cornuta are indicated in parentheses from Bader 1976): in female, the dorsal shield cross-shaped with well developed lateral protrusion (oval without lateral protrusion or its slightly developed); length and width of the dorsal shield almost subequal in female, ratio L / W = 1.07 – 1.16 (elongated, ratio L / W = 1.42 – 1.52; 1.28 according to Lundblad (1956); the anterior genital sclerite transverse (longitudinal). It should be noted that in female of F. c o r n u t a cornuta in Lundblad’s description (Lundblad 1956) the platelets, bearing setae Oe, are fused with the dorsal shield, however this specimen is probably aberrant.	en	Tuzovskij, Petr V., Semenchenko, Ksenia A. (2009): New water mites species of the genus Feltria Koenike (Acariformes, Feltriidae) from the Asian Russia. Zootaxa 2124: 38-50, DOI: 10.5281/zenodo.188186
607F180468285F3979D0FA7AFB1AFD07.taxon	etymology	Etymology. The subspecies is named after the country where it was collected (Russia). Habitat. Running waters.	en	Tuzovskij, Petr V., Semenchenko, Ksenia A. (2009): New water mites species of the genus Feltria Koenike (Acariformes, Feltriidae) from the Asian Russia. Zootaxa 2124: 38-50, DOI: 10.5281/zenodo.188186
607F180468285F3979D0FA7AFB1AFD07.taxon	distribution	Distribution. Khabarovsk Territory (Far East of Russia) and Sakha Republic (Siberia).	en	Tuzovskij, Petr V., Semenchenko, Ksenia A. (2009): New water mites species of the genus Feltria Koenike (Acariformes, Feltriidae) from the Asian Russia. Zootaxa 2124: 38-50, DOI: 10.5281/zenodo.188186
