taxonID	type	description	language	source
F36FA9FCCB2451AFA3249D9A43D0F7BD.taxon	type_taxon	Type species. Nipponogarypus enoshimaensis Morikawa, 1955, by original designation.	en	Jeong, Kyung-Hoon, Harms, Danilo, Yoo, Jung-Sun (2024): The pseudoscorpion genus Nipponogarypus (Pseudoscorpiones, Olpiidae) found in seashore habitats in Japan and Korea. Zoosystematics and Evolution 100 (3): 1053-1060, DOI: 10.3897/zse.100.123213
F36FA9FCCB2451AFA3249D9A43D0F7BD.taxon	diagnosis	Diagnosis. Nipponogarypus can be distinguished from other olpiid genera known to occur in East Asia as follows: from Beierolpium Heurtault, 1977, by trichobothrium st positioned distal to sb in Nipponogarypus and dorsal to sb in Beierolpium (Harvey 1988; Harvey and Leng 2008); from Euryolpium Redikorzev, 1938, by trichobothria it, isb, esb, and eb not clustered in Nipponogarypus but clustered in the latter. Nipponogarypus also shows similar characteristics to Olpium Koch, 1873, and Indolpium Hoff, 1945. However, Nipponogarypus can be distinguished from Indolpium by the position of trichobothria st and isb. In Indolpium, trichobothrium isb is situated proximally to trichobohtorium st (Murthy and Ananthakrishnan 1977). However, in Nipponogarypus, trichobothrium isb is situated distally from st. Furthermore, Nipponogarypus an easily be distinguised from Olpium by the length of its venom ducts. Olpium has long venom ducts that extend to trichobothrium t, whereas Nipponogarypus ’ venom ducts only extend to half of trichobothrium t (Mahnert 1991; Nassirkhani 2015). Nipponogarypus is morphologically most similar to Olpiolum Beier, 1931, and both have trichobothrium est positioned in the middle of the fixed finger; ist positioned between est and isb; isb, esb, and eb grouped together; and sb positioned closer to b than st. However, Nipponogarypus differs from Olpiolum by tergal chaetotaxy (four to six setae on the middle tergites in Nipponogarypus, always six setae on the middle tergites in Olpiolum) and the number of pseudotactile seta (two setae present in Nipponogarypus, one seta in Olpiolum) (Muchmore 1986).	en	Jeong, Kyung-Hoon, Harms, Danilo, Yoo, Jung-Sun (2024): The pseudoscorpion genus Nipponogarypus (Pseudoscorpiones, Olpiidae) found in seashore habitats in Japan and Korea. Zoosystematics and Evolution 100 (3): 1053-1060, DOI: 10.3897/zse.100.123213
D61B722AD0EE5A56BA6F007362EC9651.taxon	etymology	Etymology. This species is named after the type locality, Seosan-si, Chungcheongnam-do, in South Korea.	en	Jeong, Kyung-Hoon, Harms, Danilo, Yoo, Jung-Sun (2024): The pseudoscorpion genus Nipponogarypus (Pseudoscorpiones, Olpiidae) found in seashore habitats in Japan and Korea. Zoosystematics and Evolution 100 (3): 1053-1060, DOI: 10.3897/zse.100.123213
D61B722AD0EE5A56BA6F007362EC9651.taxon	diagnosis	Diagnosis. This species is most similar to N. enoshimaensis by having similar L / W ratios in the pedipalpal femur (0.56 – 0.61 / 0.18 – 0.20 mm in N. seosanensis sp. nov. and 0.52 – 0.59 / 0.18 mm in N. enoshimaensis) and pedipalpal patella (0.53 – 0.59 / 0.21 – 0.23 mm in N. seosanensis sp. nov. and 0.57 / 0.22 mm in N. enoshimaensis). Both species can easily be distinguished by the number of marginal teeth on the fixed chelal finger (40 – 43 in N. seosanensis sp. nov., 50 in N. enoshimaensis). N. seosanensis sp. nov. can further be distinguished from N. okinoerabensis stat. nov. by having a longer pedipalpal femur (0.56 – 0.61 mm in N. seosanensis sp. nov. and 0.48 mm in N. okinoerabensis stat. nov.) and patella (0.53 – 0.59 mm in N. seosanensis sp. nov. and 0.45 mm in N. okinoerabensis stat. nov.).	en	Jeong, Kyung-Hoon, Harms, Danilo, Yoo, Jung-Sun (2024): The pseudoscorpion genus Nipponogarypus (Pseudoscorpiones, Olpiidae) found in seashore habitats in Japan and Korea. Zoosystematics and Evolution 100 (3): 1053-1060, DOI: 10.3897/zse.100.123213
D61B722AD0EE5A56BA6F007362EC9651.taxon	description	Description. Female, adult (holotype) (Fig. 3 A, B). Color. Blackish-brown, glossy; the ventral surface darker than the end of the body appendages; the coxal region reddish-orange. Cephalothorax (Fig. 4 A, H). Carapace 1.18 times longer than the broad; carapace sub-rectangular; four conspicuous eyes; two transverse furrows on the carapace; ten lyrifissures; first furrow situated in medial position on the carapace, second furrow near the posterior margin of the carapace; carapacal chaetotaxy 4 – 2: 22; setae short and acuminate. Pedipalpal coxa with 10 setae; coxal chaetotaxy 5: 5: 6: 13; one lyrifissure on coxa I – III, two lyrifissures on coxa IV. Chelicera (Figs 4 C, 5 A). Cheliceral margin smooth; five setae on the cheliceral hand, one seta on the movable finger; galea long and shortly three-branched at the tip in both sexes; serrula exterior with 19 blades, rallum with three blades. Two lyrifissures on the hand. Pedipalp (Figs 4 B, 5 B, C). Trochanter 1.60, femur 2.95, patella 2.46, chela 3.24 times longer than the broad, movable finger 0.99 times longer than the hand. Two pseudotactile setae on the femur. Sensory setae present between et and isb. b and sb on the basal of the finger; st on the middle of the t and b. ib, esb, and eb grouped at the base of the finger; est on the middle of et and esb; it and ib on the dorsal surface of the fixed finger, it on the middle of the finger, and ib on the basal of the finger; ist on the paraxial surface of the chela, between isb and est, but closer to isb; short venom ducts in both fingers, not extending past trichobothrium et. Marginal teeth small and contiguous; triangular teeth on both fingers; fixed finger with 40, movable finger with 43 marginal teeth. Legs (Figs 4 D, E, 5 D). Leg I: trochanter 1.56, femur 2.51, patella 1.80, tibia 3.06, metatarsus 2.08, tarsus 2.33 times longer than the broad; leg IV: trochanter 1.40, femur + patella 3.26, tibia 3.93, metatarsus 2.69, tarsus 2.98 times longer than the broad. Typical for the genus: femur and patella of leg I freely mobile; metatarsus of leg IV shorter than the tarsus; metatarsus of leg I longer than the tarsus; one each pseudotactile seta in the distal position of the tarsus; basal position of the metatarsus; arolium undivided, longer than claws, and simple. Abdomen (Fig. 5 F). Pleural membrane striate; all tergites and sternites both undivided; all setae short and acuminate; tergal chaetotaxy 2: 4: 4: 4: 4: 4: 6: 5: 6: 4: 4: 2. Sternal chaetotaxy 10: 4: 6: 8: 6: 4: 6: 6: 4: 4: 2. Dimensions (in mm). Body length: 1.93 cephalothorax: carapace 0.63 / 0.54, anterior eye 0.07, posterior eye 0.05; chelicera: total 0.22 / 0.12, movable finger 0.16; pedipalp: trochanter 0.31 / 0.19, femur 0.59 / 0.20, patella 0.58 / 0.23, chela 1.08 / 0.33, movable finger 0.53, hand 0.54; leg I: trochanter 0.15 / 0.10, femur 0.23 / 0.09, patella 0.18 / 0.10, tibia 0.21 / 0.07, metatarsus 0.10 / 0.05, tarsus 0.11 / 0.05; leg IV: trochanter 0.16 / 0.11, femur + patella 0.55 / 0.17, tibia 0.38 / 0.10, metatarsus 0.17 / 0.06, tarsus 0.16 / 0.05. Variation (one female, paratype) Cephalothorax. Carapace 1.16 times longer than broad. Pedipalp. Trochanter 1.65, femur 3.13, patella 2.64, chela 3.29 times longer than broad, movable finger 0.88 times longer than hand. Fixed finger with 42, movable finger with 43 marginal teeth. Legs. Leg I: trochanter 1.28, femur 3.29, tibia 3.34, metatarsus 2.38, tarsus 2.89 times longer than broad; leg IV: trochanter 1.50, femur + patella 3.06, tibia 3.08, metatarsus 2.25, tarsus 2.82 times longer than broad. Abdomen. Tergal chaetotaxy 2: 4: 4: 4: 4: 6: 6: 6: 4: 2: 2. Sternal chaetotaxy 8: 4: 6: 6: 6: 6: 6: 4: 4: 4: 2. Dimensions (in mm). Body length 1.90; cephalothorax: carapace 0.58 / 0.50, anterior eye 0.06, posterior eye 0.05; chelicera: total 0.20 / 0.12, movable finger 0.16; pedipalp: trochanter 0.33 / 0.20, femur 0.61 / 0.20, patella 0.59 / 0.22, chela 1.04 / 0.32, movable finger 0.49, hand 0.56; leg I: trochanter 0.14 / 0.11, femur 0.24 / 0.10, patella 0.19 / 0.10, tibia 0.25 / 0.08, metatarsus 0.12 / 0.05, tarsus 0.13 / 0.05; leg IV: trochanter 0.19 / 0.13, femur + patella 0.60 / 0.20, tibia 0.34 / 0.11, metatarsus 0.15 / 0.07, tarsus 0.18 / 0.06. Variation (two males, paratype) Cephalothorax. Carapace 1.20 – 1.23 times longer than broad. Pedipalp. Trochanter 1.34 – 1.59, femur 3.07 – 3.13, patella 2.6, chela 3.05 – 3.21 times longer than the broad, movable finger 0.81 – 0.89 times longer than the hand. Fixed finger with 40 – 41, movable finger with 41 – 43 marginal teeth. Legs. Leg I: trochanter 1.20 – 1.67, femur 2.08 – 2.45, patella 1.66 – 1.70, tibia 2.67 – 3.22, metatarsus 2.62 – 2.67, tarsus 2.86 – 3 times longer than broad; leg IV: trochanter 1.25 – 1.39, femur + patella 2.67 – 2.74, tibia 3.54 – 3.63, metatarsus 2.77 – 3.16, tarsus 2.5 – 2.7 times longer than broad. Abdomen. Tergal chaetotaxy 2: 4: 4: 4: 4: 6: 6: 6: 6: 4: 4: 2. Sternal chaetotaxy 11: 6: 4: 6: 6: 6: 6: 5: 4: 4: 2. Male genital sternites (Fig. 4 G). Nine lyrifissures on sternites II and III; 11 setae on the genital opening area; six setae on the sternite III. Dimensions (in mm). Body length: 1.83 – 1.84; cephalothorax: carapace 0.61 / 0.50 – 0.51, anterior eye 0.06, posterior eye: 0.04 – 0.05; chelicera: total 0.20 / 0.10 – 0.11 movable finger 0.14; pedipalp: trochanter 0.25 – 0.32 / 0.19 – 0.20, femur 0.56 – 0.60 / 0.18 – 0.19, patella 0.53 – 0.57 / 0.21 – 0.22, chela 0.91 – 0.92 / 0.21 – 0.29, movable finger 0.41 – 0.43, z 0.49 – 0.50; leg I: trochanter 0.12 / 0.07 – 0.10, femur 0.18 – 0.24 / 0.09 – 0.10, patella 0.15 – 0.16 / 0.09 – 0.10, tibia 0.17 – 0.23 / 0.06 – 0.07, metatarsus 0.12 – 0.13 / 0.05, tarsus 0.13 / 0.04; leg IV: trochanter 0.17 – 0.18 / 0.13, femur + patella 0.51 / 0.19, tibia 0.37 / 0.10, metatarsus 0.15 – 0.16 / 0.05, tarsus 0.12 – 0.16 / 0.05 – 0.06.	en	Jeong, Kyung-Hoon, Harms, Danilo, Yoo, Jung-Sun (2024): The pseudoscorpion genus Nipponogarypus (Pseudoscorpiones, Olpiidae) found in seashore habitats in Japan and Korea. Zoosystematics and Evolution 100 (3): 1053-1060, DOI: 10.3897/zse.100.123213
