identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
F36FA9FCCB2451AFA3249D9A43D0F7BD.text	F36FA9FCCB2451AFA3249D9A43D0F7BD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nipponogarypus Morikawa 1955	<div><p>Genus Nipponogarypus Morikawa, 1955</p><p>Type species.</p><p>Nipponogarypus enoshimaensis Morikawa, 1955, by original designation.</p><p>Diagnosis.</p><p>Nipponogarypus can be distinguished from other olpiid genera known to occur in East Asia as follows: from Beierolpium Heurtault, 1977, by trichobothrium st positioned distal to sb in Nipponogarypus and dorsal to sb in Beierolpium (Harvey 1988; Harvey and Leng 2008); from Euryolpium Redikorzev, 1938, by trichobothria it, isb, esb, and eb not clustered in Nipponogarypus but clustered in the latter. Nipponogarypus also shows similar characteristics to Olpium Koch, 1873, and Indolpium Hoff, 1945 . However, Nipponogarypus can be distinguished from Indolpium by the position of trichobothria st and isb. In Indolpium, trichobothrium isb is situated proximally to trichobohtorium st (Murthy and Ananthakrishnan 1977). However, in Nipponogarypus, trichobothrium isb is situated distally from st. Furthermore, Nipponogarypus an easily be distinguised from Olpium by the length of its venom ducts. Olpium has long venom ducts that extend to trichobothrium t, whereas Nipponogarypus ’ venom ducts only extend to half of trichobothrium t (Mahnert 1991; Nassirkhani 2015). Nipponogarypus is morphologically most similar to Olpiolum Beier, 1931, and both have trichobothrium est positioned in the middle of the fixed finger; ist positioned between est and isb; isb, esb, and eb grouped together; and sb positioned closer to b than st. However, Nipponogarypus differs from Olpiolum by tergal chaetotaxy (four to six setae on the middle tergites in Nipponogarypus, always six setae on the middle tergites in Olpiolum) and the number of pseudotactile seta (two setae present in Nipponogarypus, one seta in Olpiolum) (Muchmore 1986).</p><p>Remarks.</p><p>Subspecies are a rare concept in pseudoscorpion taxonomy since recognizable morphological divergences between populations are usually associated with morphological species. Unfortunately, Morikawa had the habit of designating subspecies (and subgenera) when morphological divergences were seen by him as too minor to warrant species- or genus status for any given taxon (e. g., Morikawa 1960). Pseudoscorpion taxonomy has advanced significantly in the past decades, and we are now aware that minor morphological divergences in cryptic lineages such as pseudoscorpions are generally indicative of species status (e. g., Hlebec et al. 2024; Muster et al. 2024). Following the recent example set by You et al. (2022), who elevated all subterranean subspecies of the genus Spelaeochthonius (family Pseudotyrannochthoniidae) in Japan and Korea to species status, we also elevate Morikawa’s subspecies of Nipponogarypus enoshimaensis to species rank. Unfortunately, this taxonomic act needs to be done without reexamining the primary types that are held at Ehime University but are difficult to access and in poor condition (slide-mounted specimens in dried and contracted Hoyer’s solution; see You et al. 2022). However, Morikawa’s diagnoses are clear and reiterated here: N. enoshimaensis okinoerabensis is elevated to species rank as N. okinoerabensis stat. nov., and this name refers to specimens from the Ryuku and Satsunan Islands that have relatively short body appendages (pedipalpal femur length 0.48 mm, pedipalpal patella length 0.45 mm) and two pseudotacticle hairs that are equal in size on the palpal femur. The subspecies N. enoshimaensis enoshimaensis sensu Morikawa (1960) actually refers to N. enoshimaensis sensu Morikawa (1955) and is here recognized in its original form, N. enoshimaensis . This is a rather widespread morphospecies with records from Honshu and Shikoku (Fig. 1) that has slightly longer body appendages than N. okinoerabensis (pedipalpal femur length 0.52–0.59 mm, pedipalpal patella length 0.57 mm) and unequal pseudotactile hairs (proximal hair smaller than distal hairs). A full description of both species is available in Morikawa (1960). Following this taxonomic act, the genus Nipponogarypus now contains three morphospecies that are found along the coastlines of the Korean Peninsula and Japan.</p></div>	https://treatment.plazi.org/id/F36FA9FCCB2451AFA3249D9A43D0F7BD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Jeong, Kyung-Hoon;Harms, Danilo;Yoo, Jung-Sun	Jeong, Kyung-Hoon, Harms, Danilo, Yoo, Jung-Sun (2024): The pseudoscorpion genus Nipponogarypus (Pseudoscorpiones, Olpiidae) found in seashore habitats in Japan and Korea. Zoosystematics and Evolution 100 (3): 1053-1060, DOI: 10.3897/zse.100.123213
D61B722AD0EE5A56BA6F007362EC9651.text	D61B722AD0EE5A56BA6F007362EC9651.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nipponogarypus seosanensis Jeong & Harms 2024	<div><p>Nipponogarypus seosanensis Jeong &amp; Harms sp. nov.</p><p>Type material.</p><p>Holotype. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.1746&amp;materialsCitation.latitude=36.601543" title="Search Plazi for locations around (long 128.1746/lat 36.601543)">Female</a> (NUHGIV 0000001225). Korea: Chungcheongnam-province: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.1746&amp;materialsCitation.latitude=36.601543" title="Search Plazi for locations around (long 128.1746/lat 36.601543)">Ganwoldo</a> 1 – gil, Buseok-myeon, Seosan-si, 36 ° 36 ' 5.56 " N, 128 ° 10 ' 28.54 " E, 03, Jul 2022, KH Jeong leg.</p><p>Paratypes: One female and two males (NUHGIV 0000001228), same data as holotype .</p><p>Etymology.</p><p>This species is named after the type locality, Seosan-si, Chungcheongnam-do, in South Korea.</p><p>Habitat.</p><p>This species was collected from cracks and fissures of moist rocks in the supralittoral zone, right next to the shoreline (Fig. 2).</p><p>Diagnosis.</p><p>This species is most similar to N. enoshimaensis by having similar L / W ratios in the pedipalpal femur (0.56–0.61 / 0.18 – 0.20 mm in N. seosanensis sp. nov. and 0.52–0.59 / 0.18 mm in N. enoshimaensis) and pedipalpal patella (0.53–0.59 / 0.21 – 0.23 mm in N. seosanensis sp. nov. and 0.57 / 0.22 mm in N. enoshimaensis). Both species can easily be distinguished by the number of marginal teeth on the fixed chelal finger (40–43 in N. seosanensis sp. nov., 50 in N. enoshimaensis). N. seosanensis sp. nov. can further be distinguished from N. okinoerabensis stat. nov. by having a longer pedipalpal femur (0.56–0.61 mm in N. seosanensis sp. nov. and 0.48 mm in N. okinoerabensis stat. nov.) and patella (0.53–0.59 mm in N. seosanensis sp. nov. and 0.45 mm in N. okinoerabensis stat. nov.).</p><p>Description.</p><p>Female, adult (holotype) (Fig. 3 A, B).</p><p>Color. Blackish-brown, glossy; the ventral surface darker than the end of the body appendages; the coxal region reddish-orange.</p><p>Cephalothorax (Fig. 4 A, H). Carapace 1.18 times longer than the broad; carapace sub-rectangular; four conspicuous eyes; two transverse furrows on the carapace; ten lyrifissures; first furrow situated in medial position on the carapace, second furrow near the posterior margin of the carapace; carapacal chaetotaxy 4–2: 22; setae short and acuminate. Pedipalpal coxa with 10 setae; coxal chaetotaxy 5: 5: 6: 13; one lyrifissure on coxa I – III, two lyrifissures on coxa IV.</p><p>Chelicera (Figs 4 C, 5 A). Cheliceral margin smooth; five setae on the cheliceral hand, one seta on the movable finger; galea long and shortly three-branched at the tip in both sexes; serrula exterior with 19 blades, rallum with three blades. Two lyrifissures on the hand.</p><p>Pedipalp (Figs 4 B, 5 B, C). Trochanter 1.60, femur 2.95, patella 2.46, chela 3.24 times longer than the broad, movable finger 0.99 times longer than the hand. Two pseudotactile setae on the femur. Sensory setae present between et and isb. b and sb on the basal of the finger; st on the middle of the t and b. ib, esb, and eb grouped at the base of the finger; est on the middle of et and esb; it and ib on the dorsal surface of the fixed finger, it on the middle of the finger, and ib on the basal of the finger; ist on the paraxial surface of the chela, between isb and est, but closer to isb; short venom ducts in both fingers, not extending past trichobothrium et.</p><p>Marginal teeth small and contiguous; triangular teeth on both fingers; fixed finger with 40, movable finger with 43 marginal teeth.</p><p>Legs (Figs 4 D, E, 5 D). Leg I: trochanter 1.56, femur 2.51, patella 1.80, tibia 3.06, metatarsus 2.08, tarsus 2.33 times longer than the broad; leg IV: trochanter 1.40, femur + patella 3.26, tibia 3.93, metatarsus 2.69, tarsus 2.98 times longer than the broad. Typical for the genus: femur and patella of leg I freely mobile; metatarsus of leg IV shorter than the tarsus; metatarsus of leg I longer than the tarsus; one each pseudotactile seta in the distal position of the tarsus; basal position of the metatarsus; arolium undivided, longer than claws, and simple.</p><p>Abdomen (Fig. 5 F). Pleural membrane striate; all tergites and sternites both undivided; all setae short and acuminate; tergal chaetotaxy 2: 4: 4: 4: 4: 4: 6: 5: 6: 4: 4: 2. Sternal chaetotaxy 10: 4: 6: 8: 6: 4: 6: 6: 4: 4: 2.</p><p>Dimensions (in mm). Body length: 1.93 cephalothorax: carapace 0.63 / 0.54, anterior eye 0.07, posterior eye 0.05; chelicera: total 0.22 / 0.12, movable finger 0.16; pedipalp: trochanter 0.31 / 0.19, femur 0.59 / 0.20, patella 0.58 / 0.23, chela 1.08 / 0.33, movable finger 0.53, hand 0.54; leg I: trochanter 0.15 / 0.10, femur 0.23 / 0.09, patella 0.18 / 0.10, tibia 0.21 / 0.07, metatarsus 0.10 / 0.05, tarsus 0.11 / 0.05; leg IV: trochanter 0.16 / 0.11, femur + patella 0.55 / 0.17, tibia 0.38 / 0.10, metatarsus 0.17 / 0.06, tarsus 0.16 / 0.05.</p><p>Variation (one female, paratype)</p><p>Cephalothorax. Carapace 1.16 times longer than broad.</p><p>Pedipalp. Trochanter 1.65, femur 3.13, patella 2.64, chela 3.29 times longer than broad, movable finger 0.88 times longer than hand. Fixed finger with 42, movable finger with 43 marginal teeth.</p><p>Legs. Leg I: trochanter 1.28, femur 3.29, tibia 3.34, metatarsus 2.38, tarsus 2.89 times longer than broad; leg IV: trochanter 1.50, femur + patella 3.06, tibia 3.08, metatarsus 2.25, tarsus 2.82 times longer than broad.</p><p>Abdomen. Tergal chaetotaxy 2: 4: 4: 4: 4: 6: 6: 6: 4: 2: 2. Sternal chaetotaxy 8: 4: 6: 6: 6: 6: 6: 4: 4: 4: 2.</p><p>Dimensions (in mm). Body length 1.90; cephalothorax: carapace 0.58 / 0.50, anterior eye 0.06, posterior eye 0.05; chelicera: total 0.20 / 0.12, movable finger 0.16; pedipalp: trochanter 0.33 / 0.20, femur 0.61 / 0.20, patella 0.59 / 0.22, chela 1.04 / 0.32, movable finger 0.49, hand 0.56; leg I: trochanter 0.14 / 0.11, femur 0.24 / 0.10, patella 0.19 / 0.10, tibia 0.25 / 0.08, metatarsus 0.12 / 0.05, tarsus 0.13 / 0.05; leg IV: trochanter 0.19 / 0.13, femur + patella 0.60 / 0.20, tibia 0.34 / 0.11, metatarsus 0.15 / 0.07, tarsus 0.18 / 0.06.</p><p>Variation (two males, paratype)</p><p>Cephalothorax. Carapace 1.20–1.23 times longer than broad.</p><p>Pedipalp. Trochanter 1.34–1.59, femur 3.07–3.13, patella 2.6, chela 3.05–3.21 times longer than the broad, movable finger 0.81–0.89 times longer than the hand. Fixed finger with 40–41, movable finger with 41–43 marginal teeth.</p><p>Legs. Leg I: trochanter 1.20–1.67, femur 2.08–2.45, patella 1.66–1.70, tibia 2.67–3.22, metatarsus 2.62–2.67, tarsus 2.86–3 times longer than broad; leg IV: trochanter 1.25–1.39, femur + patella 2.67–2.74, tibia 3.54–3.63, metatarsus 2.77–3.16, tarsus 2.5–2.7 times longer than broad.</p><p>Abdomen. Tergal chaetotaxy 2: 4: 4: 4: 4: 6: 6: 6: 6: 4: 4: 2. Sternal chaetotaxy 11: 6: 4: 6: 6: 6: 6: 5: 4: 4: 2.</p><p>Male genital sternites (Fig. 4 G). Nine lyrifissures on sternites II and III; 11 setae on the genital opening area; six setae on the sternite III.</p><p>Dimensions (in mm). Body length: 1.83–1.84; cephalothorax: carapace 0.61 / 0.50–0.51, anterior eye 0.06, posterior eye: 0.04–0.05; chelicera: total 0.20 / 0.10–0.11 movable finger 0.14; pedipalp: trochanter 0.25–0.32 / 0.19 – 0.20, femur 0.56–0.60 / 0.18 – 0.19, patella 0.53–0.57 / 0.21 – 0.22, chela 0.91–0.92 / 0.21 – 0.29, movable finger 0.41–0.43, z 0.49–0.50; leg I: trochanter 0.12 / 0.07–0.10, femur 0.18–0.24 / 0.09 – 0.10, patella 0.15–0.16 / 0.09 – 0.10, tibia 0.17–0.23 / 0.06 – 0.07, metatarsus 0.12–0.13 / 0.05, tarsus 0.13 / 0.04; leg IV: trochanter 0.17–0.18 / 0.13, femur + patella 0.51 / 0.19, tibia 0.37 / 0.10, metatarsus 0.15–0.16 / 0.05, tarsus 0.12–0.16 / 0.05 – 0.06.</p></div>	https://treatment.plazi.org/id/D61B722AD0EE5A56BA6F007362EC9651	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Jeong, Kyung-Hoon;Harms, Danilo;Yoo, Jung-Sun	Jeong, Kyung-Hoon, Harms, Danilo, Yoo, Jung-Sun (2024): The pseudoscorpion genus Nipponogarypus (Pseudoscorpiones, Olpiidae) found in seashore habitats in Japan and Korea. Zoosystematics and Evolution 100 (3): 1053-1060, DOI: 10.3897/zse.100.123213
