taxonID	type	description	language	source
634287B5E74B807AE180F8ADFC29F93C.taxon	description	(Figs 1 – 2, 7 – 12, 35 – 36)	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E74B807AE180F8ADFC29F93C.taxon	diagnosis	Diagnosis. A medium-sized voriine tachinid. Prosternum and proepisternum bare; frons at most 1 / 4 of head width in male, 1 / 3.5 in female; one outwardly directed orbital seta; parafacial with a row of strong setae and with at most a few hairs on upper portion; antenna with postpedicel about two times as long as pedicel in male and female, arista bare, 2 nd aristal segment at most 2 times as long as wide; costal spine distinct; 1 st and 2 nd costal sectors setulose dorsally and ventrally; vein R 4 + 5 with 4 – 5 fine setulae from base to 2 / 5 way to r-m crossvein; wing cell r 4 + 5 open.	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E74B807AE180F8ADFC29F93C.taxon	description	Description. Body length: 6.1 – 6.8 mm. Male. Head black, with grayish pruinosity, face more whitish and occiput darker; frontal vitta black, appearing silvery brownish when seen from front; antenna and palpus black. Frons at narrowest point about 0.25 of head width (0.67 – 0.69 of single eye width); frontal vitta widened anteriorly, about 4 / 5 as wide as fronto-orbital plate near middle; parafacial slightly swollen, narrowed below, 0.60 – 0.65 as wide as postpedicel at middle in horizontal position (0.45 – 0.55 in profile); frontal length slightly more than facial length in profile (4: 3.5); face well excavated, lower margin not warped forward, about 1 / 2 as wide as long between vibrissae; facial ridge not visible in profile; gena 0.33 – 0.35 of eye height in horizontal position (0.22 – 0.25 in profile); occiput noticeably bulged. Inner vertical seta strong, slightly curved inward, about 4 / 5 as long as eye height; outer vertical seta indistinct from long fine occipital setae; ocellar setae fine hair-like, subequal in length to postvertical seta; 1 pair of postvertical setae, cruciate apically; 2 postocellar setae; 1 outwardly directed orbital seta, about 2 / 3 as long as inner vertical setae; 6 – 7 frontal setae, upper setae stronger and lowest seta inserted nearly level with middle of pedicel; a row of 5 – 7 rather long hairs on fronto-orbital plate outside row of frontal setae; parafacial with a row of 7 – 9 strong, down-curved setae increasing in length toward lower portion; vibrissa nearly level with lower margin of face; facial ridge hidden behind parafacial, with 2 – 3 short setae just above vibrissa; genal dilation with dense long hairs; occiput with dense long black hairs, pale yellowish hairs confined on ventromedian portion. Antenna short and rather wide, falling only slightly short of lower margin of face; postpedicel about 2 times as long as wide and 3.5 times as long as pedicel; arista thickened on basal 1 / 2, 2 nd aristomere about 2 times as long as wide. Palpus slender, weakly swollen apically, about 2 / 3 as long as postpedicel. Eye with dense pale yellowish hairs. Thorax black in ground color, scutum and scutellum with thin grayish pruinosity, pruinosity denser on postpronotal lobe, notopleural area and pleura; 2 very broad outer and 2 narrow inner longitudinal vittae rather distinct on presutural area and postsutural scutum. Five postpronotal setae, basal 3 setae set in a triangle; 2 presutural and 3 postsutural acrostichal setae; 3 presutural and 3 postsutural dorsocentral setae; 3 postsutural intra-alar setae; 3 supra-alar setae, 1 st seta nearly as long as 3 rd seta and longer than notopleural setae; prosternum, proepisternum, katepimeron and anatergite bare; 3 katepisternal setae; anepimeral seta fine but well developed, almost extending to posterior margin of upper calypter; scutellum with a pair of short, suberect discal setae and 3 strong marginal setae, lateral seta absent; discal setae about 5 / 6 as long as scutellum; basal setae about 1.5 times as long as scutellum, subapical setae about 2 times as long as scutellum, apical setae about 1.7 times as long as scutellum; distance between bases of subapical scutellar setae about 2 times that between basal and subapical setae. Wing hyaline, weakly tinged with pale yellow especially on basal area; tegula black, basicosta dark brown; calypter translucent pale yellowish brown; halter yellowish brown; 1 st and 2 nd costal sectors setulose above and below, sometimes basal portion of 3 rd costal sector also setulose; costagial seta (seta (e) arising from base of wing between basicosta and costagial break) strong, extending well over humeral crossvein; costal spine about 2 times as long as costal setulae; 2 nd costal sector about 1 / 3 as long as 3 rd and 4 / 5 of 4 th; vein R 4 + 5 with several setulae from base to 1 / 3 – 2 / 5 way to r-m crossvein; vein M from dm-cu crossvein to bend about 1.3 times distance between bend and wing margin; last section of vein CuA 1 about 3 / 4 as long as dm-cu crossvein; crossvein CuA 2 well developed; vein CuA 2 + A 1 obliterated before wing margin. Legs black, pulvilli pale yellowish; fore tibia with 3 preapical dorsal setae, anterodorsal seta longest, a row of 5 – 6 anterodorsal and 2 posterior setae; mid tibia with 2 – 3 anterodorsal, 2 posterodorsal and 1 – 2 ventral setae; hind tibia with a row of irregular-sized, strong anterodorsal, 2 – 3 posterodorsal and 3 – 4 posteroventral setae, and with 2 preapical dorsal and 1 apical anteroventral setae; claws and pulvilli slightly shorter than 5 th tarsomere. Abdomen black in ground color, dorsum with very thin grayish pruinosity on syntergite 1 + 2, rather thin pale yellowish pruinosity on anterior 1 / 2 – 1 / 3 of 3 rd tergite and entire 4 th and 5 th tergites, pruinosity with tessellate appearance and sometimes thinner on posterior portion of 4 th tergite; venter evenly covered with pale yellowish gray pruinosity on syntergite 1 + 2 to 5 th tergite. Hairs on dorsum fine dense and somewhat recumbent on syntergite 1 + 2 and 3 rd tergite, rather sparse and suberect on 4 th tergite and still sparse and erect on 5 th tergite; syntergite 1 + 2 excavated to anterior 1 / 2, with a pair of strong median marginal setae and 2 – 3 lateral marginal setae; 3 rd tergite with a pair of rather short median discal setae, 2 rather fine lateral discal setae, a pair of strong median marginal setae and 3 – 4 strong lateral marginal setae; 4 th tergite with a pair of strong median discal setae, 2 – 3 lateral discal setae and a row of strong marginal setae; 5 th tergite with regular rows of strong discal and marginal setae; 5 th sternite with a long and narrow postero-median cleft occupying almost posterior 3 / 4 of sternite, median membranous plate short and wide, posterior lobe with many short hairs on anterior portion and several long hairs on posterior margin. Terminalia: Cerci in dorsal view weakly narrowed from middle to apical 1 / 4, apex slightly widened and rounded, in lateral view weakly curved and pointed ventrally at apex; surstylus in lateral view narrowed to apex, with fine short hairs; pregonite flat; postgonite long, as long as basiphallus plus epiphallus; distiphallus long, with ventromedian membrane, dorsal sclerite flat and extending to posterior 4 / 5 of distiphallus, ventral sclerite widened apically in lateral view; ejaculatory apodeme rather large. Female. Differing from male as follows: Frons at narrowest point 0.32 – 0.33 of head width (0.95 – 1.00 of eye width); 2 subequally long proclinate orbital setae, slightly longer than reclinate orbital seta; parafacial about 0.33 as wide as postpedicel at middle in profile (0.5 in horizontal position); gena 0.33 – 0.35 of eye height in horizontal position (0.25 – 0.27 in profile); antenna falling short of lower margin of face by about 1 / 2 length of pedicel; postpedicel about 2 times as long as wide and about 3 times as long as pedicel; 2 nd aristomere as long as wide. First and 2 nd costal sectors of wing with only 1 – 2 fine hairs or bare above; claws and pulvilli at most 1 / 2 as long as 5 th tarsomere. Abdominal dorsum with more broadly and densely tessellate grayish pruinosity. Terminalia: Sixth tergite entire, semicircularly convex on anteromedian dorsal portion, with a row of strong setae on posterior margin; 6 th sternite nearly as long as 6 th tergite; 6 th spiracle on anteroventral portion of 6 th tergite, 7 th spiracle on membrane just behind posterosubmedian portion of 6 th tergite; 7 th tergite narrowly separated mid-dorsally into quadrate hemitergites, nearly as long as 6 th tergite, with a row of some strong setae dorsally and several fine setae ventrally on posterior margin; 8 th tergite reduced to a small, nearly triangular hemitergite, without hairs; 8 th sternite rather well expanded posteromedially, with many minute hairs on posterior portion; epiproct absent; hypoproct broad, semicircular. Puparium (Figs 35 – 36). Length 5.7 – 6.7 mm, width 2.3 – 2.7 mm. Reddish brown; surface with very minute wrinkles, especially on last two segments; rather weakly narrowed to posterior portion; posterior spiracles black, raised above obliquely on posterior portion of puparium, each spiracle divided transversely into two tubercles with a shallow median groove on which a median cicatrix (button) is present, each tubercle with 4 sinuate spiracular slits.	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E74B807AE180F8ADFC29F93C.taxon	etymology	Etymology. The specific name refers to the short antenna of this species, from the Latin adjective brevis (short) plus the Latin noun cornu (antenna) and adjectival suffix is (with).	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E74B807AE180F8ADFC29F93C.taxon	materials_examined	Type material. Holotype male *: Japan, Hokkaido / Tomakomai, Toma- / komai Experimental Forest // 10. vi. 2014, (emerged) / ex Erannis golda / M. Libra & T. Abe (TOC 7818) (HUM). Paratypes *: 1 male, same data as holotype except date, 30. v. 2014 (TOC 02699); 3 males, 2 females, same data as holotype [TOC 06975 (1 male, 1 female); TOC 07468 (1 male); TOC 07820 (1 female); TOC 08207 (1 male)]; 1 female, same data as holotype except date, 28. v. 2014 (TOC 03837); 1 male, same as preceding, 9. vi. 2014 (TOC 08423); 1 female, same as preceding, 19. vi. 2014, (TOC 10077); 1 female same data as preceding except host, ex Orthosia odiosa (TOC 10082) (KUM, HUM, SMNS, USNM); 1 male, 1 female, Japan, Honshu, Kanayama, Masutomi, 26. v. 1975, A. Nakanishi & J. Emoto, Malaise trap (KUM) Additional material *. 1 male (shrunken), 10. vi. 2014, same data as holotype (TOC 07820); 1 puparium (not emerged), 19. vi. 2014, same data as holotype except host, ex Orthosia odiosa (TOC 10082); 1 male (shrunken), same as preceding except date and host, 30. v. 2015, ex Catocala lara, M. Libra & T. Abe (TOC 22656) (HUM). Other parasitoids *. 1 hymenopteran ex B. brevicornis (puparium) parasitic on Erannis golda (pupa), 19. vi. 2014, M. Libra & T. Abe (TOC 07887).	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E74B807AE180F8ADFC29F93C.taxon	biology_ecology	Hosts. Lepidoptera, Geometridae: Erannis golda Djakonov. Noctuidae: Catocala lara Bremer, Orthosia odiosa (Butler).	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E74B807AE180F8ADFC29F93C.taxon	discussion	Remarks. The genus Blepharomyia Brauer & Bergenstamm is known from four species in the Palearctic Region and two in the northern Nearctic. The Nearctic species have characteristically yellowish tibiae (Coquillett 1897: 53, Curran 1927: 149), in contrast to the Palearctic ones, which all have black tibiae. Only one endemic species, B. foliacea Mesnil, 1975 (Figs 3 – 4), was previously known from Japan. The new species is similar to the European B. pagana (Meigen, 1824) (Figs 5 – 6), but differs from it in having a less apically widened postpedicel and a narrower frons and narrower gena in the male. These three species can be distinguished as follows:	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E742807BE180F941FD32FD04.taxon	description	(Figs 26, 37 – 38)	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E742807BE180F941FD32FD04.taxon	diagnosis	Diagnosis. A medium to large-sized exoristine tachinid. Eye hairy; parafacial bare; facial ridge with setae on lower 3 / 4 or more; antenna large, 2 nd aristal segment at most 2 times as long as wide; 3 presutural and 4 postsutural dorsocentral setae; wing veins bare except for base of vein R 4 + 5 with 2 – 3 short and fine setulae; wing vein M with a rather weak wrinkle at bend; abdomen without discal setae even on 5 th tergite; mid coxa of female with 5 – 6 rather short and strong spines; male cerci broad on basal 4 / 5 and strongly narrowed to apex.	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E742807BE180F941FD32FD04.taxon	description	Description of puparium (Figs 37 – 38). Length ca. 9.0 – 9.2 mm, width 0.49 – 0.50 mm. Brown, barrel-shaped; surface with small wrinkles; posterior spiracles shining dark brown, nearly flat on posterior surface of posterior surface, rather circular with inner margin weakly curved; cicatrix (button) nearly at middle of spiracular plate; spiracular slits absent, but many tiny spots recognizable on spiracular plate. Specimens examined *. 1 female, Japan Hokkaido, Tomakomai, Tomakomai Experimental Forest, 7. vii. 2014 (emerged), ex Himeropteryx miraculosa, M. Libra & T. Abe (TOC 11405) (HUM); 1 male, same data as preceding except date, 10. vii. 2015 (TOC 11737) (KUM).	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E742807BE180F941FD32FD04.taxon	biology_ecology	Host. Lepidoptera, Notodontidae: Himeropteryx miraculosa Staudinger.	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E742807BE180F941FD32FD04.taxon	discussion	Remarks. Members of this genus are oviparous. This species was described by Mesnil (1967: 43) from Sapporo, Hokkaido, Japan and later recorded from Amur, Russia (Ziegler & Shima 1969). The female has peculiar peg-like spines on the ventral surface of the mid coxa, which may assist with oviposition on host caterpillars. This species appears to be a solitary parasitoid; it pupates within the dead host caterpillar and ecloses from it. The present record is the first reported host for this species.	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E743807BE180FD29FDB0F884.taxon	description	(Fig. 27)	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E743807BE180FD29FDB0F884.taxon	diagnosis	Diagnosis. A large goniine tachinid. Eye bare; only 1 reclinate orbital seta; frons narrow in male, about 0.2 of head width, 0.25 in female; prosternum with a pair of strong setae and many long hairs on both sides; proepisternum bare; 3 postpronotal setae arranged in a nearly straight line; 3 presutural and 4 postsutural dorsocentral setae; lateral scutellar setae double; apical scutellar setae subequal in length to scutellum, crossed horizontally; antenna and palpus dark brown to black in male, sometimes antennal pedicel and apex of palpus weakly and narrowly reddish brown in female; abdomen broad, without median discal setae on 3 rd and 4 th tergites, male with well-developed sexual hair-patches on venter of 4 th tergite; 5 th tergite at most 3 / 5 as long as 4 th tergite.	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E743807BE180FD29FDB0F884.taxon	description	Description of puparium. Length 9.2 – 10.5 mm, width 4.4 – 5.6 mm. Reddish brown, barrel-shaped and weakly widened on posterior 1 / 3; band of minute spines on anterior 1 / 3 of each tergite; spiracular plates black, each with three sinuate spiracular slits; terminal projection (“ Terminalhöker ” of Ziegler 1998) distinct between and below the spiracular plates. Very similar to the puparium of B. schineri Mesnil illustrated by Ziegler (1998: fig. 98), but spiracular plates closer together and spiracular slits more weakly sinuate. Specimens examined *. 1 female, Japan Hokkaido, Tomakomai, Tomakomai Experimental Forest, 24. vii. 2014 (emerged), ex Marumba jankowskii (TOC 11183); 1 male, 1 female, same data as preceding except date, 25. vii. 2014 (TOC 10852); 2 males, 2 females, same data as preceding, 25. vii. 2014 (TOC 10416) (KUM, HUM, USNM).	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E743807BE180FD29FDB0F884.taxon	biology_ecology	Host. Lepidoptera, Sphingidae: Marumba jankowskii (Oberthür).	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E743807BE180FD29FDB0F884.taxon	discussion	Remarks. Species of Blepharipa Rondani are difficult to identify because of their very similar general appearance and only slight differences in the male and female terminalia. This species was described from Japan (Hokkaido) and was recorded from Xizang (Tibet), China by Chao & Shi (1982: 271). However, Chao et al. (1998) did not record it in “ Flies of China ”. It is probable that this species occurs elsewhere in China, but Xizang must be considered a doubtful record. This is a large tachinid that is micro-ovolarviparous. The female lays microtype eggs (fully incubated with a hard chorion, nearly 0.25 – 0.30 mm in length) on the foliage. This is one of the most common species in Japan, but a host had not been previously recorded. According to the present rearing records, this species is gregarious and pupates outside the dead host pupa.	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E743807EE180F91BFE76FE59.taxon	description	(Figs 13 – 17, 28 – 29, 39 – 40)	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E743807EE180F91BFE76FE59.taxon	diagnosis	Diagnosis. A small to medium-sized grayish goniine tachinid. Head dichoptic; eye haired; 2 reclinate orbital setae; antenna with postpedicel elongate, about 7 times as long as pedicel in male, 4.5 times in female; frontal length in profile about 0.7 of facial length; 3 presutural and 3 postsutural dorsocentral setae; frons at narrowest point 0.34 – 0.35 of head width in male, 0.38 – 0.39 in female; gena 0.22 – 0.24 of eye height in profile; puparium with posterior spiracles raised well above posterior surface.	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E743807EE180F91BFE76FE59.taxon	description	Description. Body length: 4.5 – 5.0 mm. Male. Head with dense whitish pruinosity, fronto-orbital plate pale yellowish and dark gray posteriorly, occiput grayish; frontal vitta velvety brown; antenna brown-black; palpus yellow, narrowly darkened basally. Frons at narrowest point 0.34 – 0.35 of head width (1.10 – 1.25 of single eye width); frontal vitta nearly parallel-sided, subequal in width to fronto-orbital plate; frontal length about 0.7 of facial length in profile; parafacial weakly bulged in profile, about 1 / 2 as wide as postpedicel at middle height in horizontal position (about 2 / 5 in profile); face well excavated, lower margin not warped, about 2 / 5 as wide as long at level of vibrissa; gena 0.31 – 0.33 of eye height in horizontal position (0.22 – 0.24 in profile); occiput weakly bulged. Inner vertical setae parallel, about 3 / 4 as long as eye height; 2 reclinate orbital setae, anterior seta stronger and about 1 / 2 as long as inner vertical seta; ocellar setae slightly longer than anterior reclinate orbital seta; 5 – 6 frontal setae, lowest seta nearly level with base of arista; fronto-orbital plate with fine and rather sparse hairs, not descending below lowest frontal seta; parafacial bare; facial ridge with only a few setulae just above vibrissa; vibrissa inserted level with lower margin of face; occiput with 2 – 3 rows of fine black setae behind postocular setae and whitish pile. Antenna long, postpedicel about 4 times as long as wide and 7 times as long as pedicel; arista thickened on basal 1 / 3, 2 nd aristomere only slightly longer than wide. Palpus slender, only slightly bulged apically. Thorax black in ground color, with pale yellowish gray pruinosity; scutum with 5 broad longitudinal vittae, of which median 3 vittae sometimes fused forming a broad median vitta, outer vitta about 2 times as wide as pruinose portion between inner and outer vittae. Prosternum bare; postpronotal lobe with 4 strong setae, 3 basal setae set in a straight line; 3 presutural and 3 postsutural acrostichal setae; 3 presutural and 3 postsutural dorsocentral setae; 1 presutural and 3 postsutural intra-alar setae; 3 supra-alar setae; proepisternum, katepimeron and anatergite bare; 3 katepisternal setae; scutellum with rather dense, fine, erect short hairs, a pair of fine, erect discal setae set closely together, 4 marginal setae, basal and subapical setae subequal in length and about 2 times as long as scutellum, lateral seta subequal in length to scutellum, apical setae very fine, hair-like and parallel to each other. Wing hyaline, tegula black, basicosta dark brown; calypter whitish; halter pale yellowish brown; costal spine about 2 times as long as costal setae; 2 nd costal sector haired below; vein M from dm-cu crossvein to bend about 2 times distance between bend and wing margin; dm-cu crossvein weakly concave, slightly longer than last section of vein CuA 1. Legs black, hind tibia sometimes slightly paler on median portion; pulvilli pale yellowish; fore tibia with 2 posterior setae; mid tibia with 1 anterodorsal, 2 posterodorsal and 1 ventral setae; hind tibia with a sparsely set row of anterodorsal setae, submedian seta stronger, 1 strong posterodorsal and 2 short ventral setae, and with 2 preapical dorsal and 1 apical anteroventral setae, no apical posteroventral seta; fore claws and pulvilli subequal in length to 5 th tarsomere. Abdomen black in ground color, with pale yellowish gray pruinosity on anterior 1 / 3 – 1 / 4 of 3 rd tergite, 4 / 5 of 4 th tergite and entire 5 th tergite, pruinosity thinner on median and posterior portions of 3 rd tergite and with somewhat shifting appearance on 4 th and 5 th tergites; venter with evenly grayish pruinosity. Hairs on dorsum rather sparse, short, fine and suberect, longer on lateral portions and fine and erect on 5 th tergite; syntergite 1 + 2 excavated to posterior margin, with a pair of rather fine median marginal setae and 2 lateral marginal setae; 3 rd tergite with a pair of rather fine median discal setae, 2 fine lateral discal setae, a pair of strong median marginal setae and 2 strong lateral marginal setae; 4 th tergite with a pair of strong median discal setae, 1 – 2 strong lateral discal setae and a row of strong marginal setae; 5 th tergite with regular rows of strong discal setae and marginal setae. Terminalia: Cerci in dorsal view narrowed from base to basal 1 / 3 and parallel-sided to apical 1 / 4, then weakly narrowed to apex, apical portion narrowly separated on distal 1 / 4, in lateral view weakly narrowed to rounded apex; surstylus in lateral view with dorsal and ventral margins parallel from base to apical 1 / 2, then dorsal margin curved ventrally to rather rounded apex, apical portion with 3 short and strong spinules. Female. Differing from male as follows: Fronto-orbital plate slightly paler, with pale yellowish brown pruinosity; frons at narrowest point 0.38 – 0.39 of head width (1.1 – 1.3 of eye width); inner vertical seta subequal in length to eye height; outer vertical seta about 1 / 2 as long as inner vertical seta; 2 subequally long proclinate orbital setae, nearly as long as outer vertical seta; 4 frontal setae; parafacial about 5 / 7 as wide as postpedicel at mid height in horizontal position (2 / 3 in profile); frontal length about 0.75 of facial length in profile; postpedicel about 4.5 times as long as pedicel; no apical scutellar seta; claws and pulvilli shorter than 5 th tarsomere. Terminalia: Sixth tergite narrowly separated mid dorsally into hemitergite, only slightly shorter than 6 th sternite, with several strong setae marginally; 7 th tergite longer than 6 th tergite; 7 th abdominal spiracle situated in posteromedian portion of 6 th tergite; 7 th tergite medially separated into two hemitergites, long quadrate in form, longer than 6 th tergite; 8 th tergite well developed, nearly quadrate, about 1 / 4 as long as 7 th tergite, without hairs; 8 th sternite small, triangular; epiproct absent. Puparium (Figs 39 – 40). Length 3.3 – 5.2 mm, width 1.7 – 2.3 mm. Brown in color; narrow band of very fine wrinkles or minute spinules on anterior 1 / 3 – 1 / 5 of each segment; spiracular plate black, nearly circular, distinctly raised from pupal surface; 15 spiracular slits raised and surrounding central cicatrix (button); no terminal projection.	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E743807EE180F91BFE76FE59.taxon	etymology	Etymology. This species is named for its horse-faced (long-faced) head, from the Latin nouns equus (horse) plus facies (face).	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E743807EE180F91BFE76FE59.taxon	materials_examined	Type material. Holotype male *: Japan, Hokkaido / Tomakomai, Toma- / komai Experimental Forest // ex 10. vi. 2015 (emerged) / ex Archips nigricaudana / M. Libra & T. Abe (TOC 24977) (HUM). Paratypes *: 1 male, same data as holotype except date and host, 31. v. 2015 (emerged), ex Archips crataegana (TOC 21835) (USNM); 1 female, same data as preceding, 23. v. 2015 (emerged), ex Epinotia exquisitana (TOC 20837) (KUM). Other specimens examined * (all in HUM): 1 specimen (body mostly broken, sex not determined; head, legs and abdomen missing, with exuvia of puparium), same data as holotype except date and host, 4. vi. 2015 (emerged), ex Faristenia geminisignella (TOC 23368); 1 female (head and left fore leg missing), same data as preceding, 26. v. 2014, ex Pseudohedya gradana (TOC 02882).	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E743807EE180F91BFE76FE59.taxon	biology_ecology	Hosts. Lepidoptera, Gelechiidae: Faristenia geminisignella Ponomarenko. Tortricidae: Archips crataegana (Hübner); Archips nigricauda Walshingham; Epinotia exquisitana Christoph; Pseudohedya gradana Christoph.	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E743807EE180F91BFE76FE59.taxon	discussion	Remarks. This is a micro-oviparous tachinid, laying fully incubated minute eggs with a hard chorion (about 0.18 mm long) on the foliage. The female is probably attracted to damaged or partially eaten leaves and lays eggs on them. Only one larva successfully develops within the host and pupates inside the dead host pupa. This species is characteristic in its horse-faced appearance, long antenna (Fig. 29) and bare prosternum. The puparium has also characteristically expanded posterior spiracles. Cyzenis albicans (Fallén, 1810) is also recorded from Hokkaido, Japan (Mesnil & Pschorn-Walcher 1963). The new species resembles C. albicans, but may be distinguished from it by its narrower gena, bare prosternum and absence of proclinate or lateroclinate outer orbital setae in the male. The posterior spiracles of the puparium are raised above the surrounding surface as in C. albicans (see O’Hara & Cooper 1992: 793, figs 24 – 26), but the spiracular slits are evenly arranged around the cicatrix, situated near the center (Fig. 40).	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E746807CE180FC2DFA08FCCA.taxon	description	(Figs 18 – 23, 41 – 42)	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E746807CE180FC2DFA08FCCA.taxon	diagnosis	Diagnosis. Frons at narrowest point 0.30 – 0.33 of head width in male, 0.32 – 0.33 in female; frontal length in profile about 0.9 of facial length; 4 postsutural dorsocentral setae; postpedicel about 5.5 times as long as pedicel in male, 4 times in female; gena 0.3 of eye height in profile; posterior spiracles of puparium only slightly raised above surface of puparium.	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E746807CE180FC2DFA08FCCA.taxon	description	Description. Body length: 5.2 – 7.3 mm. [Resembling the preceding species and differing from it as follows:] Male. Head with dense yellow to golden yellow pruinosity, face and facial ridge whitish and genal dilation and occiput grayish; frons at narrowest point 0.30 – 0.33 of head width (0.95 – 1.00 of single eye width); parafacial weakly bulged in profile, subequal in width to postpedicel in horizontal position at mid height (only slightly narrower in profile); frontal length about 0.9 of facial length in profile; face about 4 / 9 as wide as long at level of vibrissa; gena 0.37 – 0.40 of eye height in horizontal position (0.30 – 0.32 in profile); inner vertical seta about 2 / 3 as long as eye height; 2 reclinate orbital setae, anterior seta stronger and about 3 / 4 as long as inner vertical seta; fronto-orbital plate with fine and rather dense hairs; facial ridge with short setulae on at most lower 2 / 5; antenna with postpedicel about 3.5 times as long as wide and 5.5 times as long as pedicel; arista thickened on basal 1 / 2, 2 nd aristomere only slightly longer than wide; palpus slender, only slightly bulged apically, about 3 / 5 as long as postpedicel. Thorax: prosternum with a pair of rather long and strong setae and 2 – 3 pairs of fine, short hairs; 3 presutural and 4 postsutural dorsocentral setae; basal scutellar seta about 5 / 6 as long as subapical seta, and about 1.6 times as long as scutellum, lateral seta about 1.5 times as long as scutellum; very fine, hair-like apical setae parallel to each other or divergent; wing with 2 nd costal sector bare or with at most 3 fine setulae below; vein M from dm-cu crossvein to bend about 1.6 times distance between bend and wing margin; dm-cu crossvein rather well concave below, subequal in length to last section of vein CuA 1; legs with fore claws and pulvilli slightly shorter than 5 th tarsomere. Abdominal dorsum with pale yellowish gray pruinosity on 3 rd to 5 th tergites, pruinosity with shifting appearance in direction of light, posterior margin of 3 rd and 4 th tergites weakly darkened; hairs on dorsum rather sparse, short, fine and suberect, longer on median and lateral portions and fine and erect on 5 th tergite. Terminalia [differing from the preceding species as follows]: Cerci in dorsal view more widened at base; surstylus in lateral view curved posteriorly near apex, dorsal margin curved ventrally in apical 1 / 4, apical portion with 3 – 4 fine setulae; epiphallus long. Female. Differing from male as follows: Frons at the narrowest point 0.32 – 0.33 of head width (0.9 – 1.1 of eye width); outer vertical seta about 1 / 2 as long as inner vertical seta; 2 subequally long proclinate orbital setae, nearly as long as anterior reclinate orbital seta; 4 – 5 frontal setae; parafacial about 1.25 times as wide as postpedicel at mid height in horizontal position (about as wide as in profile); frons in profile about as long as facial length; postpedicel of antenna about 3.5 times as long as wide and 4 times as long as pedicel; no apical scutellar setae; claws and pulvilli distinctly shorter than 5 th tarsomere. Terminalia: 6 th tergite rather short, with several long hairs on posterior margin; 7 th spiracle on posteroventral portion of 7 th tergite; 7 th tergite reduced to rather small hemitergite, slightly shorter than 6 th tergite; 8 th tergite small and short; 8 th sternite reduced to two small sclerites. Puparium. Length 5.0 – 6.2 mm, width 2.2 – 2.9 mm. Similar to the preceding species, but slightly larger; posterior spiracles only slightly raised above surface of puparium, slightly more widely separated than in the preceding species; 11 – 12 spiracular slits encircling cicatrix (button).	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E746807CE180FC2DFA08FCCA.taxon	etymology	Etymology. The specific epithet refers to the four postsutural dorsocentral setae on the thorax, from the Latin nouns tetra (four) plus seta (bristle) and the suffix osus (nature of).	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E746807CE180FC2DFA08FCCA.taxon	materials_examined	Type material. Holotype male *: Japan, Hokkaido / Tomakomai, Toma- / komai Experimental Forest // ex 10. vi. 2014 (emerged) / ex Operophtera relegate / M. Libra & T. Abe (TO: 10564) (HUM). Paratypes *: 2 males, 3 females, same data as holotype except date, 2. vi. 2015 [(TOC 22078 (1 male); TOC 22205 (1 female); TOC 22624 (1 male); TOC 22379 (1 female); TOC 23643 (1 female)]; 7 males 6 females, same data as holotype except date and host, 27. v. 2014, 10. vi. 2014, 5. v. 2015, 2. vi. 2015, 6. vii. 2015, ex Operophtera brunnea [TOC 03549 (1 male); TOC 04707 (1 male); TOC 07916 (1 female); TOC 06623 (1 female); TOC 20740 (1 male); TOC 21253 (1 female); TOC 21370 (1 female); TOC 21576 (1 male); TOC 23155 (1 female); TOC 23211 (1 female); TOC 23147 (1 male); TOC 23743 (1 male); TOC 23944 (1 male)]; 4 males 2 females, same data as preceding, 25. v. 2015, 2. vi. 2015, 10. vi. 2015, ex Lomographa simplicior [(TOC 20608 (1 male); TOC 21969 (1 male); TOC 22843 (1 female); TOC 22934 (1 male); TOC 24486 (1 male); TOC 24499, (1 female)] (HUM, USNM, SMNS); 1 male, Akita Pref., Oga Pen., Mt. Shinzan, 4. v. 1989, S. Fukushi; 1 male, Saitama Pref., Yorii, Mt. Kanetsukido, 11. iv. 1989, K. Hara; 1 male, Fukuoka Pref., Mt. Kusenbu, 15. iv. 1988, H. Shima (KUM); 8 males, Fukuoka Pref., Mt. Hikosan, 1000 m, 19. iv. 1938, K. Yasumatsu (ELKU). Other specimens examined *. 1 male (shrunken), same data as holotype except date and host, 25. v. 2015, ex Operophtera brunnea (TOC 04426) (HUM).	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E746807CE180FC2DFA08FCCA.taxon	biology_ecology	Hosts. Lepidoptera, Geometridae: Lomographa simplicior (Butler), Operophtera brunnea Nakajima, O. relegata Prout.	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E746807CE180FC2DFA08FCCA.taxon	discussion	Remarks. This species is a solitary parasitoid, developing in the host and pupating within the dead host caterpillar. The hosts are geometrid larvae instead of the leaf rolling gelechiids or tortricids parasitized by Cyzenis equifacialis sp. nov., which all occurs sympatrically and synchronically in this forest. The three Japanese species of Cyzenis Robineau-Desvoidy can be distinguished by the following key:	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E7448063E180F829FD7DFD90.taxon	description	(Figs 24 – 25, 32 – 34, 43 – 44)	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E7448063E180F829FD7DFD90.taxon	diagnosis	Diagnosis. A dark-colored species. Male head nearly holoptic, inner vertical setae indistinct (Fig. 34), dichoptic in female, inner vertical setae strong and decussate, outer vertical setae distinct in both sexes; eye densely haired; facial ridge bare except for several short setae above vibrissa; parafacial with 1 – 2 rows of strong setae; prosternum with several fine hairs on both sides; 3 postpronotal setae in a straight line; 3 presutural and 3 postsutural dorsocentral setae; 2 katepisternal setae; 3 pairs of marginal scutellar setae, apical setae absent; costagial seta strong, extending beyond humeral crossvein; 2 nd costal sector bare below; hind tibia with 3 preapical dorsal setae; abdomen with discal setae, sometimes also on syntergite 1 + 2. Body length: 5.1 – 5.9 mm. [The female and puparium are described here for the first time.]	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E7448063E180F829FD7DFD90.taxon	description	Description of female and puparium. Female (Figs 32 – 34). Head dichoptic, with whitish gray pruinosity, upper fronto – orbital and anterodorsal portion of parafacial dark gray; frons wide, at narrowest point 0.32 – 0.34 of head width (0.9 – 1.1 of eye width); frontal vitta about 1 / 2 as wide as fronto-orbital plate at middle; inner vertical setae strong and cruciate, about as long as eye height; outer vertical seta well developed, about 3 / 4 as long as inner vertical seta; outwardly directed prevertical seta slightly shorter than outer vertical seta; 2 subequally long proclinate orbital setae, subequal in length to outer vertical seta; 5 – 7 strong frontal setae, lowest seta nearly level with apex of scape; fronto-orbital plate with several fine hairs outside row of frontal setae; face 0.65 – 0.68 as long as frons in profile; parafacial about 0.83 width of postpedicel at mid height in horizontal position (0.6 – 0.7 in profile), with a row of 5 – 6 down-curved setae which increase in length below; gena 0.32 – 0.34 of eye height in horizontal position (0.26 – 0.29 in profile); postpedicel of antenna about 2 times as long as wide and 2.7 – 3.0 times as long as pedicel; claws and pulvilli of legs shorter than 5 th tarsomere, fore claws and pulvilli about 1 / 2 as long as 5 th tarsomere; abdominal dorsum with rather thin, whitish pruinosity on anterior 1 / 4 – 1 / 5 of 3 rd to 5 th tergites. Terminalia (Figs 24 – 25): Sixth tergite broadly divided mid-dorsally into 2 hemitergites, without hairs; 6 th spiracle on membrane close to anteroventral portion of 6 th hemitergite; 6 th sternite about 1 / 3 as long as 5 th sternite, of equilateral triangle shape, with a row of some rather strong setae and rather long hairs on posterior margin; 7 th spiracle on membrane close to posteromedian portion of 6 th hemitergite; 7 th and 8 th tergites absent; 7 th sternite nearly triangular, about 2 / 3 as long as 6 th sternite, with a row of fine hairs; 8 th sternite with posterior margin weakly rounded, well narrowed to anterior margin, slightly smaller than 7 th sternite; epiproct small, divided medially, each half with a few hairs posteriorly; three spermathecae. Puparium (Figs 43 – 44). Length 4.7 – 6.0 mm, width 1.7 – 2.5 mm. Brown, posterior margin rather weakly narrowed, surface smooth; posterior spiracles black and small, situated close to each other on raised tubercle of posterior portion; 1 – 2 small and short spiracular slits at apex.	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E7448063E180F829FD7DFD90.taxon	materials_examined	Material examined *. 13 males, 2 females, Japan, Hokkaido, Tomakomai Experimental Forest, Tomakomai, 24 – 25 and 27. v. 2014, 2. vi. 2014, 14. vi. 2014, 9. vi. 2015, ex Rhopobota naevana [TOC 01522 (abdomen and legs missing) (1 male); TOC 02020 (1 male); TOC 02091 (1 male); TOC 02513 (1 male); TOC 02531 (1 male); TOC 02609 (1 male); TOC 02743 (1 male); TOC 02773 (1 female); TOC 03024 (1 female); TOC 03274 (1 male); TOC 05048 (1 male); TOC 06325 (1 male); TOC 09197 (1 male); TOC 09605 (1 male); TOC 24483 (1 male)], M. Libra & T. Abe; 1 female (head, legs and abdomen missing, thorax broken), same data as preceding except date and host, 31. v. 2015, ex Rhopobota sp. (TOC 01627); 1 female, same data as preceding except date and host, 3. v. 2015, ex Rhopobata sp. (TOC 23302); 1 female (head missing), same data as preceding except date and host, 31. v. 2015, ex Pseudohedya gradana (TOC 22257); 1 female, same data as preceding except date and host, 24. v. 2014, ex Archips crataegana (TOC 00033), (HUM, KUM, USNM); 1 puparium, 24. vi. 2014, ex Rhopobota naevana (TOC 03480) (HUM); 1 male, Hokkaido, Sapporo, Jozankei, 30. v. 1965, S. Takano (CNC); 2 females, Hokkaido, Asahikawa City, 30. i. 1967, ex Zeiraphera rufimitrata truncata, K. Kamijo; 1 female, same locality and host as preceding, ii. 1969, K. Kamijo (KUM).	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E7448063E180F829FD7DFD90.taxon	biology_ecology	Hosts. Lepidoptera, Tortricidae: Archips crataegana (Hübner); Pseudohedya gradana (Christoph); Rhopobota naevana (Hübner); Rhopobota sp.; Zeiraphera rufimitrata truncata Oku (Shima 1999).	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
634287B5E7448063E180F829FD7DFD90.taxon	discussion	Remarks. This species was well described by Mesnil (1963) based on a male from Khabarovsk, Russia. Mesnil & Shima (1978) recorded the male also from Japan and described and illustrated the male terminalia. The genus Eulasiona Townsend has been traditionally classified in the subfamily Dexiinae (Mesnil 1966, as Eulasionina of Voriini; Herting 1984, as Voriini; O’Hara & Wood 2004, as Voriini), mainly because of the structure of the phallus. However, the molecular analysis of Stireman et al. (2019) placed it in the subfamily Tachininae, close to Phytomyptera Rondani and Graphogaster Rondani. They considered the hinged structure of the phallus in Eulasiona to have independently arisen in this genus and in the Dexiinae. The curvature of the phallus in Eulasiona is weak (not angled but rounded) and it lacks an epiphallus (see Mesnil & Shima 1978), whereas the female terminalia lack the 7 th and 8 th abdominal tergites and sternites; so far as we know, the 8 th abdominal tergite and sternite are never lacking in the Dexiinae. Given the sum of the existing evidence, Eulasiona is moved from the subfamily Dexiinae to an unplaced position within the subfamily Tachininae. This species is a solitary parasitoid and the female deposits fully incubated eggs close to or on the host. Fully developed larvae appear to kill the host while it is still in the larval stage and pupate within the dead host or very close to it. This behavior is inferred from finding the host’s head capsules, fragments of the host’s body or hairs attached to the tachinid puparia in this study. Shima (1999) recorded a tortricid species as a host of E. zimini, and here we add four additional tortricid species as new hosts.	en	Shima, Hiroshi, Abe, Tomokazu, Libra, Martin (2021): Tachinid flies (Diptera: Tachinidae) reared from deciduous plant-feeding lepidopteran larvae at Hokkaido University Tomakomai Forest (Japan), with descriptions of three new species. Zootaxa 5060 (2): 275-295, DOI: 10.11646/zootaxa.5060.2.8
