taxonID	type	description	language	source
498A376BDEE35B40A8B6677DCF2A7CD9.taxon	description	Figures 17 B, F, 21 C	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
498A376BDEE35B40A8B6677DCF2A7CD9.taxon	distribution	Distribution and habitat. Mostly mesic habitats in Morocco, Europe and the northwestern Middle East. In the Levant, limited to mountains above 1300 m in north and central Lebanon (Wood et al. 2020).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
E0302F3CFAAA5D1789C88CB32B4AA551.taxon	description	Figures 13 C, 16 A, G, 21 B	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
E0302F3CFAAA5D1789C88CB32B4AA551.taxon	distribution	Distribution and habitat. Shrublands, sub-alpine and semi-arid habitats in Cyprus, all countries of the Levant, and southern Turkey.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
48EC22F4C95C55C0AE6541DF32B982C0.taxon	description	Figures 3, 15 B, K, 20 L	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
48EC22F4C95C55C0AE6541DF32B982C0.taxon	etymology	Etymology. Named after the Arabic name of Mount Hermon, Jabal al-Shaykh (‘ Mountain of the Sheikh’). The species epithet is a noun in apposition.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
48EC22F4C95C55C0AE6541DF32B982C0.taxon	diagnosis	Diagnosis. Within the Andrena minutula species group, A. alshaykh belongs to the species around A. rugulosa Stöckhert, characterized by a smooth clypeus and punctate terga, which include also A. lindbergella Pittioni and A. libanica Wood sp. nov. It is closest to A. lindbergella Pittioni from Cyprus, and until the discovery of the distinct male of A. lindbergella, they were considered conspecific. The female differs from A. lindbergella in the tergal discs which are more strongly shagreened and more obscurely punctured (Fig. 15 K, L), and in the brighter terminal fringe. The male is easily differentiated by the genital capsule, in which the gonostyli suddenly converge medially, producing distinct kinks in the inner and outer margins (uniformly curving in A. lindbergella), and the penis valves are broader (Fig. 20 L, N). In addition, the scutum is more sparsely punctured, and the body size is smaller. Both A. alshaykh and A. lindbergella are closely related to A. rugulosa Stöckhert (absent from the Levant) and A. libanica Wood sp. nov., but differ in the smoother scutum and more weakly punctured terga (Fig. 15 J – L). In addition, the male of A. rugulosa does not have dorsal gonocoxite lobes, and the male of A. libanica does not have kinks in the inner and outer margin of the gonostyli (Fig. 20 L – N).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
48EC22F4C95C55C0AE6541DF32B982C0.taxon	description	Description. FEMALE. Body length: 6.5 – 7.5 mm. Integumental colour: Body and legs black. Anterior side of flagellum greyish-black to slightly reddish. Apical tarsomeres brown. Wings weakly infuscate, veins brown, stigma brown peripherally, reddish centrally (Fig. 3 A). Tergal marginal zones brown near apical margin (Fig. 3 D). — Pubescence: Body hair mostly short to medium-lengthed, minutely plumose, white to brown (Fig. 3 A). Clypeus with sparse medium-lengthed whitish hairs (Fig. 15 B). Area around antennal sockets with dense medium-lengthed white hair. Scape with dense short to medium white hair. Vertex with medium-lengthed erect whitish-golden hair. Genal area with dense short hair, whitish-golden dorsally, white ventrally. Facial fovea with dense, minute white to slightly yellowish hair, colour depending on angle of view (Fig. 3 B, C). Periphery of scutum, scutellum and metanotum with moderately dense, erect, short to medium, golden-brown hair. Scutal disc with sparse and thin, medium-lengthed, golden-brown hair (Fig. 3 C). Mesepisternum with sparse, long white hair. Propodeal corbicula incomplete, dorsal fringe consisting of dense and long, distinctly plumose golden hair. Corbicular surface with sparse and long, simple golden hairs. Leg hair short, whitish to brownish. Flocculus incomplete, white. Femoral scopa composed of long, simple to coarsely plumose golden hair. Tibial scopa weakly developed, composed of medium-lengthed, simple whitish – brownish hair (Fig. 3 A). Tergal discs essentially hairless. Tergal marginal zones 2 – 4 with distinct, strongly interrupted apicolateral white hair bands, slightly protruding onto following tergal discs. Terminal fringe light brown (Fig. 3 A, D). — Head: 1.2 times broader than long. Mandibles bidentate, weakly crossed. Galea finely shagreened. Labral process narrowly trapezoidal to triangular. Malar area undeveloped. Clypeus moderately arched, apically shiny and more or less smooth, basally gradually more shagreened; punctation distinct, distance between punctures 1 – 3 puncture diameters, puncture density decreasing medioapically, an impunctate midline is not indicated (Fig. 15 B). Lower part of paraocular area smooth and shiny, strongly and densely punctured. Supraclypeal area and lateral parts of frons with strong longitudinal striations, interspersed with strong and dense, fine punctures. Centre of frons with weaker and finer oblique striations, medial carina not reaching frontal ocellus, punctation similar. Flagellomere 1 1.0 – 1.1 times as long as 2 + 3 (Fig. 3 B). Facial foveae shallow, elongate and more or less uniformly narrow, extending from level of lower end of lateral ocellus to slightly above clypeus base, fovea width equals ⅓ antennocular distance. Distance of fovea from lateral ocellus equals 1.8 ocellus diameters. Vertex moderately carinate. Ocelloccipital distance slightly shorter than 1 ocellus diameter (Fig. 3 B, C). — Mesosoma: Dorsolateral angle of pronotum not elevated. Anterior ⅓ of scutum finely shagreened, disc shiny and more or less smooth. Scutal punctation dense and strong, distance between punctures 0.5 – 1 puncture diameters. Scutellum smooth, punctation similar (Fig. 3 C). Anterior part of mesepisternum finely rugose-areolate, posterior part shagreened. Propodeal corbicula finely reticulated, with a few very large crater-shaped punctures around hair bases. Basal half of propodeal triangle very coarsely rugose-areolate, apical half and flanking propodeal regions much more finely so (Fig. 3 C). Hind pretarsal claw with strong inner tooth. Recurrent vein 1 meets submarginal cell 2 more or less near its middle. Nervulus interstitial (Fig. 3 A). — Metasoma: Tergal discs shagreened, with moderately strong, irregular punctation, distance between punctures 1 – 2 puncture diameters at densest parts, punctation much finer on tergum 4. Tergal marginal zones broad, on tergum 4 covering about ¾ tergum length medially, marginal zones shagreened except near apex, very weakly and finely punctured (Fig. 3 D). Pygidial plate simple, central area not elevated. — Male. Body length: 5.5 – 6 mm. — Integumental colour: Body and legs black. Flagellum dark brown to black. Apical tarsomeres brown. Wings hyaline, veins brown, stigma light brown centrally (Fig. 3 E). Tergal marginal zones yellowish-brown (Fig. 3 H). — Pubescence: Body hair mostly minutely plumose, brightly coloured (Fig. 3 E). Face with medium to long white hair, dense on lower half and on scape, sparser on upper half. Vertex with medium-lengthed erect whitish hair. Genal area with dense white hair, becoming longer ventrally (Fig. 3 E – G). Scutum, scutellum and metanotum with sparse, medium to long, whitish to yellowish erect hairs. Mesepisternum with dense, very long white hair. Propodeum with sparse, long white hair (Fig. 3 E, G). Legs with short to medium, white to golden hair (Fig. 3 E). Tergal discs with minute inconspicuous white hair. Tergal marginal zones 2 – 5 with strongly interrupted apicolateral white hair bands. Terminal fringe whitish (Fig. 3 E, H). — Head: Labral process very narrowly trapezoidal, apical half thickened, apical margin strongly concave. Clypeus weakly arched, smooth and shiny except near base, distinctly punctured, distance between punctures about 1 puncture diameter, without impunctate midline, clypeal sculpture hidden by dense pubescence. Flagellomere 1 slightly shorter than 2 + 3, 2 shorter than 3 (Fig. 3 F). Ocelloccipital distance about 1 ocellus diameter (Fig. 3 G). Rest of head as in female. — Mesosoma: Scutum and scutellum very shiny, superficially shagreened, occasionally smooth centrally, distinctly punctured, distance between punctures 1 – 3 puncture diameters (Fig. 3 G). Nervulus interstitial to antefurcal (Fig. 3 E). Rest of mesosoma as in female. — Metasoma: Similar to female, but tergal disc punctation finer and more uniform, distance between punctures about 1.5 puncture diameters. Tergal marginal zones narrower than in female (Fig. 3 H). — Genitalia and hidden sterna: Dorsal gonocoxite lobes developed, converging apically, more or less rounded. Penis valves moderately broad basally, gradually tapering up to mid-length, hereafter uniformly narrow. Gonostyli finger-shaped, with a distinct kink in mid-length, where they are slightly bent inwards. Sternum 8 simple, columnar, slightly broadening apically, apical process undeveloped (Fig. 20 L).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
48EC22F4C95C55C0AE6541DF32B982C0.taxon	distribution	Distribution and habitat. Sub-alpine habitats in northern Israel (Mt. Hermon) and Lebanon, likely also Syria, above 1900 m. Previously reported from Israel and Lebanon as A. lindbergella (Pisanty et al. 2018; Wood et al. 2020; Boustani et al. 2021).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
57481728E09F56EA8DD106840D2BCB95.taxon	description	Figures 4, 18 B, H, 21 I	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
57481728E09F56EA8DD106840D2BCB95.taxon	etymology	Etymology. Named after the Greek goddess Aphrodite, who is strongly associated with the island of Cyprus.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
57481728E09F56EA8DD106840D2BCB95.taxon	diagnosis	Diagnosis. Within the Andrena minutula species group, A. aphroditae belongs to the species around A. spreta Pérez, which are characterized in the female by a clypeus which is non-rugose, weakly domed and partly to fully shagreened, a scutum which is shagreened to partly smooth, with fine, relatively sparse punctures, and terga which are mostly shagreened and impunctate. In the Levant and Cyprus, these include A. hebraica Pisanty & Wood sp. nov., A. minutuloides Perkins, A. spreta, A. tiaretta Warncke, and A. tkalcui Gusenleitner & Schwarz. Female Andrena aphroditae are distinguished from A. hebraica and A. tiaretta by the scutum and scutellum which are usually somewhat shiny and distinctly punctured (completely dull in A. hebraica; dull and weakly punctured in A. tiaretta; Figs 17 J, 18 H), and the terga which are uniformly shagreened (tergum 3 and often also 2 distinctly shinier apically vs. basally in A. hebraica). From Cypriot populations of A. spreta Pérez, female A. aphroditae can be distinguished by the clypeus which is more strongly domed and usually completely dull (Fig. 18 B; usually smooth apically in A. spreta), dark flagellum (Fig. 4 B; often reddish distally in A. spreta), usually shiny scutellum (usually dull in A. spreta; Figs 17 K, 18 H), brown stigma (yellowish in A. spreta; Figs 4 A, 17 M), and dark brown terminal fringe (Fig. 4 A, D; golden in Cypriot A. spreta). Continental populations of A. spreta are much more morphologically variable, but on average they differ from A. aphroditae by the same characters, most of all in the clypeus and stigma. Compared to A. minutuloides, female A. aphroditae has weaker and much finer punctation on the clypeus (Fig. 18 B, C), as well as somewhat weaker, finer and denser punctation on the scutum (Fig. 18 H, I). Separation from female A. tkalcui is easily done by the much darker colouration of the flagellum and wing veins (Fig. 17 L), as well as the more strongly shagreened scutum (Figs 17 H, 18 H) and weaker tergal apical hair bands. From the sympatric A. cervina Warncke, female A. aphroditae can be distinguished by the clypeus which is densely and finely punctured, often with hints of transverse striation on the basal half (more sparsely and coarsely punctured, without any transverse striation in A. cervina; Figs 18 A, B), and the scutum and scutellum, which are usually somewhat shiny, distinctly and not very densely punctured (completely dull, superficially and very densely punctured in A. cervina; Figs 18 G – H). The male genitalia (Fig. 21 I) provides good separation from all the above related species: the gonocoxites of A. spreta and A. tkalcui possess a distinct dorsal lobe (Fig. 20 F, J), the gonostyli of A. hebraica and A. tiaretta strongly converge apically (Fig. 21 G, J), the gonostyli of A. cervina have a strong inward kink near the apex (Fig. 20 O), and the penis valves of A. minutuloides and A. tiaretta are broader (Fig. 21 F, G). The males are further easily differentiated from A. spreta by the much smoother scutum and scutellum (Fig. 4 G) and the darker flagellum and stigma (Figs 4 E, F, 17 M).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
57481728E09F56EA8DD106840D2BCB95.taxon	description	Description. Female. Body length: 5 – 6.5 mm. Integumental colour: Body and legs black. Flagellum black, distal flagellomeres increasingly covered with minute greyish setae. Distal tarsomeres black to dark brown. Wings weakly infuscate, veins dark brown, stigma centrally dark to light brown (Fig. 4 A). Tergal marginal zones black (Fig. 4 D). — Pubescence: Clypeus with moderately dense, semi-erect, short and thin whitish hairs, underlying cuticle visible. Apex of clypeus with two small strands of long brown hair. Supraclypeal area with short, erect whitish hairs. Paraocular area, scape and area around antennal sockets with moderately dense, erect medium-lengthed white hair (Fig. 4 B). Facial fovea in dorsal view with light brown hairs on upper part of fovea, white hairs on lower half (Fig. 4 C). Frons with sparse, short to medium, erect, mixed black and white hairs. Ocellar triangle with few erect medium-lengthed black hairs. Preoccipital ridge with erect, short to long, whitish to brownish hairs. Genal area with few short black hairs behind upper margin of compound eye, elsewhere with semi-erect to erect whitish hairs, short on dorsal part, medium-lengthed on ventral part (Fig. 4 A, C). Discs of scutum and scutellum with sparse, inconspicuous minute to short whitish hairs, with few scattered longer erect hairs. Periphery of scutum with moderately dense, erect short whitish hairs. Posterior margin of scutellum with few erect, medium to long, whitish to brown hairs (Fig. 4 C). Mesepisternum with long white hairs. Propodeal corbicula incomplete, posterodorsal fringe composed of long white plumose hairs, corbicular surface with few long, simple white hairs. Leg hair white to light brown, flocculus incomplete, white, tibial scopa composed of simple hairs, brownish posterobasally, elsewhere whitish (Fig. 4 A). Tergal discs 1 – 2 centrally essentially hairless, 3 – 4 gradually with minute inconspicuous whitish hairs. Lateral parts of tergal discs with sparse short white hair. Tergal marginal zones 2 – 4 with distinct lateral bands of white hair extending onto following tergal discs, strong and dense on terga 2 – 3, weaker on tergum 4. Terminal fringe brownish, with few flanking white hairs (Fig. 4 D). — Head: 1.25 times broader than long. Mandibles bidentate, moderately crossed. Galea very finely shagreened (Fig. 4 B). Labral process trapezoidal, not much broader than long. Clypeus moderately domed, basal half fully shagreened and matt, often with hints of fine transverse striations, apical half fully shagreened and matt to partly shagreened and weakly shiny. Clypeus punctation moderately strong, distance between punctures 0.5 – 1.5 puncture diameters, without impunctate midline (Figs 4 B, 18 B). Frons and upper part of paraocular area densely longitudinally striated, essentially impunctate (Fig. 4 B, C). Flagellomere 1 0.9 – 1.0 times as long as 2 + 3, 2 almost as long as 3 (Fig. 4 B). Facial fovea moderately broad above, here 0.4 – 0.5 times as broad as antennocular distance, weakly tapering below, extending from level of lower end of lateral ocellus to base of clypeus, lower ⅔ separated from compound eye by narrow smooth cuticular strip (Fig. 4 B, C). Distance of fovea from lateral ocellus about 1.5 ocellus diameters. Ocelloccipital distance about ¾ ocellus diameter. Preoccipital ridge weakly to moderately carinate (Fig. 4 C). — Mesosoma: Dorsolateral angle of pronotum not elevated. Scutum largely shagreened, usually with some shiny, partly smooth areas centrally, punctation fine, distance between punctures 0.5 – 1.5 puncture diameters. Scutellum similar but usually shinier, often completely smooth (Figs 4 C, 18 H). Mesepisternum finely alveolate, anterior part overlayed with oblique punctures. Surface of propodeal corbicula reticulated, with sparse, coarse punctures around hair bases. Posterolateral part of propodeum finely rugose-areolate. Propodeal triangle weakly delineated, horizontal part strongly radially rugose near base, elsewhere rugose-areolate, vertical part very finely areolated (Fig. 4 C). Hind pretarsal claw with small inner tooth. Recurrent vein 1 meets submarginal cell 2 at its middle or slightly proximal to it. Nervulus interstitial (Fig. 4 A). — Metasoma: Tergal discs impunctate, 1 – 2 and usually 3 completely shagreened and matt, 4 and occasionally 3 slightly shinier and more weakly shagreened. Tergal marginal zones weakly depressed, 2 – 3 centrally occupying 0.4 – 0.5 of tergal length, shagreening as on discs, much weaker on tergum 4 (Fig. 4 D). Pygidial plate triangular, completely flat, smooth to superficially shagreened. — Male. Body length: 5 – 5.5 mm. — Integumental colour: As in female, but tergal marginal zones somewhat brownish (Fig. 4 E, H). — Pubescence: Clypeus with moderately dense, semi-erect, short to medium white hair. Paraocular area with short to medium erect hair, black near compound eye, whitish near antennal sockets. Scape with medium-lengthed white hairs (Fig. 4 F). Frons with short to medium, erect dark hairs. Preoccipital ridge with short to long erect white hairs. Genal area with erect black hair behind upper margin of compound eye, elsewhere with white hair, short dorsally, gradually long ventrally (Fig. 4 E – G). Scutum and scutellum with sparse short whitish hair, some medium to long erect hairs appear mostly on periphery (Fig. 4 G). Mesepisternum with very long white hairs (Fig. 4 E). Propodeum with long white hairs (Fig. 4 E, G). Femora and tibiae with white hair, tarsi with brownish-white hair (Fig. 4 E). Base of tergal disc 1 and lateral parts of all tergal discs with sparse, short to medium white hair. Rest of tergal discs with minute inconspicuous whitish hair. Tergal marginal zones 2 – 4 with small lateral bands of white hair, extending onto following discs, very weak on tergum 4. Terminal fringe brownish-white (Fig. 4 H). — Head: 1.25 times broader than long. Labral process much broader than long, apical margin usually concave. Clypeus moderately domed, very shiny and almost completely smooth, punctation strong and dense, distance between punctures 0.5 – 1.5 puncture diameters. Flagellomere 1 about as long as 3, much longer than 2 (Fig. 4 F). Ocelloccipital distance about 1.1 ocellus diameters. Preoccipital ridge moderately carinate (Fig. 4 G). — Mesosoma: Scutum shiny, superficially shagreened to smooth, punctation fine and sparse, distance between punctures 1 – 3 puncture diameters. Scutellum similar, but smoother and more sparsely punctured. Propodeal triangle strongly rugose-areolate on horizontal part, very finely so on vertical part, usually becoming radially rugose near base (Fig. 4 G). Hind pretarsal claw strongly bifurcated. Rest of mesosoma as in female. — Metasoma: As in female (Fig. 4 H). — Genitalia and hidden sterna: Gonocoxites without dorsal lobe or with only hint thereof. Gonostyli simple, finger-shaped, uniformly broad, blade flattened, rounded apically. Penis valves moderately narrow basally, basal ⅕ of visible area parallel-sided, the following ⅖ tapering apically, distal ⅖ narrow (Fig. 21 I). Sternum 8 columnar, broadening apically, apical margin blunted.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
57481728E09F56EA8DD106840D2BCB95.taxon	distribution	Distribution and habitat. Endemic to Mediterranean shrublands in Cyprus.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
2ADC3469FF4D530B9C9446D867025D60.taxon	diagnosis	Diagnosis. As a unified group, Andrena (Micrandrena) are too morphologically diverse to diagnose with unique defining characters, especially with regard to other members of the highly diverse clade of small-sized Andrena distributed primarily in the Old World (clades 23 – 24 in Pisanty et al. 2022 a), which includes also Aciandrena, Aenandrena, Fuscandrena, Graecandrena, Parandrenella, and the A. janthina species group. All Micrandrena are relatively small-sized, and all Palaearctic species lack bright facial markings in both sexes. They generally also lack most of the distinct modifications which characterize many other Andrena subgenera, such as elongated mouthparts, enlarged vertex and gena, carinate pronotum, complete propodeal corbicula, sloping profile of propodeum, carinate or toothed hind femur, modified hind tibial spur, plumose scopa, raised area on pygidial plate, or strong male pygidial plate. Beyond these simple generalizations, to provide a clear diagnosis against similar subgenera and species groups, it is necessary to divide Micrandrena into several morphological groups and diagnose each one of them separately, as specified below. As reference groups, the following discussion will focus on the related subgenera Aciandrena (as presently circumscribed), Fuscandrena, Graecandrena and the Andrena janthina group, which are the main taxa most often confused with Micrandrena. For the sake of the present discussion and for utility within the fauna of the Levant and Cyprus, we divide the Palaearctic members of A. (Micrandrena) into the following groups, which to our best understanding, correspond to distinct clades in phylogenomic analysis (Pisanty et al. 2022 a; Bossert et al. in prep.): 1. Andrena longibarbis group – former subgenus Distandrena. 2. Andrena minutula group – this is the old concept of subgenus Micrandrena sensu Warncke in the Palaearctic but excluding the species around A. oedicnema Warncke and A. proxima (Kirby). This group contains the bulk of the species diversity of the subgenus. It corresponds to Schmid-Egger & Scheuchl’s minutula - and nana - groups combined (Schmid-Egger & Scheuchl 1997). We see no justification to distinguish the nana group here, as preliminary molecular data does not support its monophyly, and the minutula / nana group division is not clearly applicable outside Central Europe. 3. Andrena oedicnema group – A. oedicnema Warncke and the closely related A. cedricola Wood. 4. Other unassigned, peculiar taxa, including the well-known Andrena proxima group, here represented by a single species only. The Andrena longibarbis group can be well characterized against other Andrena subgenera by the combination of: 1. A clypeus that is completely flat or almost so, often with longitudinal striations (Fig. 13 E – I); 2. Supraclypeal plate often longitudinally striated; 3. Facial foveae extremely long and narrow, usually extending below the antennal sockets, with the lower ½ – ⅔ almost linear (Fig. 13 D); 4. Propodeal triangle entirely reticulated, essentially lacking any rugae, even basally (Fig. 13 A); 5. Scutum and terga strongly shagreened and impunctate to weakly punctured (Fig. 13 J – P); 6. Hind leg pretarsal claw usually bidentate. This species group is most similar to members of subgenus Aciandrena and the A. janthina species group, but in these reference taxa the clypeus is not always flat, the clypeus and supraclypeal area are never longitudinally striated, the foveae are shorter and more drop-shaped, the body shagreening is usually weaker and finer, and the hind leg pretarsal claw is usually unidentate. The highly diverse Andrena minutula group is more variable morphologically compared to the A. longibarbis group, but it is characterized most of all by a propodeal triangle that is finely but strongly rugose to rugose-areolate, at least on the basal half (Fig. 13 C). Many members of the A. minutula group also have strong integumental sculpturing such as deep punctures or strong shagreening (Figs 15 – 18). Both these traits set this group apart from the related subgenera Aciandrena, Fuscandrena and Graecandrena as well as the A. janthina species group. In these reference groups, the propodeal triangle is more reticulated, with the rugosity absent or weaker and more confined to the basal margin; the integumental sculpturing is generally weaker, with punctures and shagreening usually appearing shallower; and in some species the males have yellow facial markings and / or reduced gonostyli, but these are more difficult to generalize. There are exceptions to most of the above criteria, and caution should be exercised when excluding similar subgenera. Most of the remaining taxa of A. (Micrandrena) in the treatment below can also be diagnosed against other subgenera based on the same criteria mentioned above. Hence, most of the characters of the Andrena minutula group apply also to the A. oedicnema group and to A. proxima; A. extenuata Wood also possesses a similar strongly rugose propodeal triangle. The A. oedicnema group is further characterized by mirror-smooth, mostly impunctate terga (Fig. 14 P), and A. proxima possesses an almost unique, star-shaped wrinkling of the surface of the propodeal corbicula (Fig. 14 M). The three remaining species are more challenging to diagnose, as their propodeal triangles are more weakly or narrowly rugose, as in Graecandrena (Fig. 13 B), however all possess unusual characteristics which are rare among the related subgenera: A. dividicincta Pisanty has very strong tergal hairbands which are strictly limited to the tergal sides (Fig. 14 O); A. pandosa Warncke has broad facial foveae (Fig. 14 K), a protuberant, medially flattened clypeus in which the apicolateral corners are distinctly elevated, and a strongly notched labral process (Fig. 14 B); and A. yelkouan Warncke has an entirely flat clypeus (Fig. 14 A) as well as strongly contrasting dull scutum versus shiny scutellum (Fig. 14 G). It is important to note that several species of the A. minutula group occur in two generations which are morphologically distinct. This includes A. alfkenella Perkins, A. alfkenelloides Warncke, A. chananaea Pisanty & Wood sp. nov., A. minutula, and A. minutuloides Perkins. As a general rule, the second generations possess a smoother cuticle and brighter pubescence (particularly the male facial pubescence), which are most likely adaptations for the warmer ambient temperature and stronger solar radiation associated with late season (Ostwald et al. 2025).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
157CFFF7A77C5CCA8025EA0A695FC0F8.taxon	description	Figures 5, 16 D, J, 21 E	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
157CFFF7A77C5CCA8025EA0A695FC0F8.taxon	etymology	Etymology. Nominative feminine singular form of the Latin adjective asper, meaning rough, uneven, coarse, in reference to the species’ integumental sculpturing.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
157CFFF7A77C5CCA8025EA0A695FC0F8.taxon	diagnosis	Diagnosis. The female of Andrena aspera stands out among Levantine Micrandrena species by the combination of a labral process which is often weakly emarginate apically, distinctly domed clypeus with transverse striations and / or oblique punctures (Fig. 16 D), dull, strongly roughened scutum with dense crater-like punctures (Figs 5 C, 16 J), and dull, shagreened terga which are subtly punctate apicolaterally (Fig. 5 D). It is closest to species such as A. simontornyella Noskiewicz, A. rugothorace Warncke and A. lunaris Pisanty & Wood, but in all these species the scutum is more finely sculptured, with denser, finer and smoother punctation. Similar differences apply to the clypeus, which is more finely rugose in A. simontornyella, very densely, finely rugose in A. rugothorace, and non-rugose in A. lunaris. Among Levantine Micrandrena, the male of Andrena aspera is characterized by the combination of a domed, very densely punctured clypeus covered by dense long white hair (Fig. 5 F), scutum with some crater-punctures centrally (Fig. 5 G), and penis valves which slightly broaden immediately above the visible base (Fig. 21 E). It is very similar to A. rugothorace, but in the latter species the clypeus is more strongly transversely striated, and the gonocoxites have distinct dorsal lobes (Fig. 20 R).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
157CFFF7A77C5CCA8025EA0A695FC0F8.taxon	description	Description. Female. Body length: 6.5 – 7.5 mm. — Integumental colour: Body and legs black. Distal flagellomeres brown to black (Fig. 5 A). Wings weakly infuscate, veins brown, stigma centrally light brown (Fig. 5 E). Tergal marginal zones black basally, more brownish apically (Fig. 5 D). — Pubescence: Head and mesosoma with distinctly plumose hair of varying brightness (Fig. 5 A – C). Lower half of face, up to level of antennal sockets, with dense, moderately long, semi-erect brownish-white hair (Fig. 5 B). Paraocular area usually with erect black hairs near facial fovea, elsewhere with brownish-white hairs (Fig. 5 A, B). Scape with brownish-white hairs (Fig. 5 B). Hair on upper half of facial fovea distinctly brown, on lower half brown to whitish, depending on angle of view. Frons and ocellar triangle with sparse golden hairs, often interspersed with black hairs (Fig. 5 B, C). Preoccipital ridge with long, erect golden hairs. Genal area behind upper margin of compound eye with few short black hairs, elsewhere with short to medium, golden to whitish hairs (Fig. 5 A – C). Scutum, scutellum and metanotum with moderately dense, erect, golden-brown hairs of varying length, their density increasing peripherally, underlying cuticle visible (Fig. 5 A, C). Mesepisternum with dense, long whitish hairs, becoming slightly golden dorsally. Propodeal corbicula incomplete, dorsoposterior fringe whitish, corbicular surface with sparse, long simple whitish hairs. Leg hair mostly whitish; flocculus incomplete, white; tibial scopal hairs simple, whitish (Fig. 5 A). Terga centrally with inconspicuous minute hair, laterally with sparse medium-lengthed whitish hair, creating weak, broadly interrupted hair bands on marginal zones 2 – 3, and a very weak and sparse continuous hair band on marginal zone 4. Terminal fringe golden to light brown, flanked by few long white hairs (Fig. 5 A, D). — Head: 1.2 times broader than long. Mandibles bidentate, moderately crossed. Galea finely shagreened (Fig. 5 B). Labral process rectangular, moderately broad, apical margin weakly concave. Clypeus distinctly domed, fully shagreened and matt, punctation oblique, distance between punctures 0.5 – 1 puncture diameter, without impunctate midline, punctures appearing connected by subtle transverse striation (Fig. 16 D). Lower part of paraocular area with strongly oblique punctures merging into striation. Supraclypeal area finely striated. Flagellomere 1 1.0 – 1.1 times as long as 2 + 3, 2 slightly shorter than 3 (Fig. 5 B). Frons and upper part of paraocular area rugose-areolate, sculpturing becoming coarser centrally. Facial foveae of moderate width, upper half slightly broader, here 0.5 times as broad as antennocular distance, fovea extending from level of lower end of lateral ocellus to base of clypeus or slightly below, separated from compound eye by very narrow cuticular strip which slightly broadens on the lower half. Frons with complete, shiny medial carina, lower side of frontal ocellus surrounded by polished cuticular area (Fig. 5 B, C). Lateral ocelli connected posteriorly by a transverse furrow. Distance of fovea from lateral ocellus about 2 ocellus diameters (Fig. 5 B, C). Ocelloccipital distance about 1 ocellus diameter. Vertex moderately carinate (Fig. 5 C). — Mesosoma: Dorsolateral angle of pronotum not elevated. Scutum entirely with strong granular shagreening, punctation fine, dense and strong, distance between punctures 0.5 – 1 puncture diameters, punctation often appearing oblique especially on anterior part, with some punctures displaying raised margins (crater-like punctures). Scutellum similar but with straight punctation, often somewhat shinier and more sparsely punctured (Figs 5 C, 16 J). Mesepisternum finely alveolate, overlayed by strongly oblique, dense punctation. Surface of propodeal corbicula finely alveolate, with a few large punctures around hair bases. Posterior part of propodeum strongly differentiated into horizontal vs. vertical regions. Posterolateral area of propodeum rugose-areolate. Propodeal triangle weakly demarcated, horizontal part rugose-areolate, often more radially rugose near base, vertical part finely alveolate (Fig. 5 C). Hind pretarsal claw with small inner tooth. Recurrent vein 1 meets submarginal cell 2 proximal to its middle. Nervulus antefurcal to interstitial (Fig. 5 E). — Metasoma: Tergal discs finely shagreened, basolaterally with raised oblique punctures, elsewhere more or less impunctate. Tergal marginal zones shagreened and impunctate, moderately arched, 2 – 3 centrally occupying about half the tergum length, 2 hardly depressed, the following gradually more so (Fig. 5 D). Pygidial plate normally developed. — Male. Body length: 5.5 – 6.5 mm. — Integumental colour: As in female (Fig. 5 E). — Pubescence: Clypeus and lower part of paraocular area with dense and long, semi-erect whitish plumose hairs. Supraclypeal area, scape and area around antennal sockets with medium to long erect whitish hairs. Dorsolateral part of paraocular area bordering compound eyes with erect black hairs (Fig. 5 E, F). Frons and ocellar triangle with sparse erect hairs of varying brightness. Preoccipital ridge with long, erect whitish to golden hairs (Fig. 5 F, G). Dorsal part of genal area with short black hair, central and ventral parts with white hair, becoming longer ventrally (Fig. 5 E, F). Scutum, scutellum and metanotum with medium to long erect whitish hair, denser peripherally, underlying cuticle visible (Fig. 5 E, G). Mesepisternum with very long, erect plumose white hair (Fig. 5 E). Propodeum with long, plumose white hair (Fig. 5 E, G). Leg hair mostly white, tarsal hair often with some weak golden reflections (Fig. 5 E). Tergal discs laterally with moderately dense, short to medium whitish hair, densest and longest on tergum 2, hair becoming short and inconspicuous towards centre of disc. Tergal marginal zones 2 – 3 with weak and sparse, broadly interrupted white hair bands; 4 – 5 with continuous, very weak and sparse whitish hair bands. Terminal fringe whitish (Fig. 5 H). — Head: 1.25 times broader than long. Labral process moderately broad, rectangular, apical margin usually concave. Clypeus domed, entire surface with almost confluent honeycomb-like punctation, distance between punctures less than 0.5 puncture diameter, clypeal sculpture hidden by dense pubescence. Flagellomere 1 longer than 3 but shorter than 2 + 3, 2 distinctly shorter than 3 (Fig. 5 F). Upper part of paraocular area longitudinally striated. Frons finely rugose-areolated. Ocelloccipital distance about 1.4 ocellus diameters. Preoccipital ridge carinate (Fig. 5 G). — Mesosoma: As in female, but scutum usually more sparsely punctured, distance between punctures 0.5 – 2 puncture diameters (Fig. 5 G). Hind pretarsal claw bifurcated. — Metasoma: Tergal discs with strong granular shagreening, centrally with only hint of minute punctures, laterally slightly more distinctly punctate, though here partly obscured by dense overlying pubescence. Tergal marginal zones moderately depressed, finely reticularly shagreened, impunctate (Fig. 5 H). — Genitalia and hidden sterna: Apical margin of gonocoxite very slightly protruding medially, weakly rounded to almost truncate. Gonostyli broad and elongate, blade flattened, inner margin slightly broadening and curving dorsally at about 0.4 gonostylus length. Penis valves narrow, slightly but distinctly broadening close to visible base (Fig. 21 E). Sternum 8 columnar, distinctly broadening apically, apical margin rounded.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
157CFFF7A77C5CCA8025EA0A695FC0F8.taxon	distribution	Distribution and habitat. Mediterranean shrublands in Israel, the West Bank and Lebanon. Previously reported from Lebanon as A. simontornyella ssp. corpana (Wood et al. 2020; Boustani et al. 2021).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
5F360FCD123B5F89BBCF24F4CC09B9A3.taxon	description	Figures 6, 15 C, E, 20 H	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
5F360FCD123B5F89BBCF24F4CC09B9A3.taxon	diagnosis	Diagnosis of male. The male of A. calandra, similarly to the female, is easily distinguished from other members of the A. minutula group by the combination of a small body size, strong orange colouration of the anterior of the flagellum, polished and mirror-smooth scutum (Fig. 6 C), and distinctly punctured tergal discs (Fig. 6 D). It is most similar to A. tkalcui Gusenleitner & Schwarz, but clearly differs in the genitalia which lack a strong dorsal gonocoxite lobe (Fig. 20 F, H), the eighth sternum which is apically truncate (strongly emarginate in A. tkalcui, Fig. 20 W), and the clearly punctate tergal discs. In addition, the scutum of A. tkalcui is often partly shagreened centrally, and only occasionally mirror-smooth.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
5F360FCD123B5F89BBCF24F4CC09B9A3.taxon	description	Description of male. Body length: 4.5 – 5.5 mm. — Integumental colour: Head and mesosoma black (Fig. 6 A). Flagellum posteriorly black to reddish-brown, anteriorly flagellomere 1 black to reddish-black, the following flagellomeres reddish to orange. Legs black to dark brown, apical tarsomeres golden to light brown. Wings hyaline, vein R of forewing dark brown, internal veins golden to slightly brownish, stigma brownish-golden peripherally, yellow centrally (Fig. 6 A). Metasoma usually black, occasionally partly to fully reddish-brown. Tergal marginal zones usually as dark as adjacent tergal discs, occasionally more brownish near apex (Fig. 6 D). — Pubescence: Almost completely white (Fig. 6 A). Lower ⅔ of face, including clypeus, supraclypeal and paraocular areas, scape and lower part of frons, with moderately dense, erect to semi-erect, short to medium white hairs (Fig. 6 A, B). Upper part of frons and ocellar triangle with sparse, short to medium white hairs. Preoccipital ridge with short to medium erect white hairs. Genal area with moderately dense white hair, short dorsally, medium-lengthed ventrally (Fig. 6 A – C). Discs of scutum and scutellum with sparse, short to medium erect white hair. Metanotum and periphery of scutum and scutellum with moderately dense, medium-lengthed erect white hair. Mesepisternum and propodeum with long white hair (Fig. 6 A, C). Leg hair almost completely white, foretarsal hair with weak golden reflections (Fig. 6 A). Base of tergal disc 1 and lateral parts of all terga with sparse, short to medium white hair, rest of tergal discs with minute inconspicuous white hair (Fig. 6 D). Tergal marginal zones 2 – 5 with weak bands of white hair, strongly interrupted on 2 – 3, continuous on 4 – 5. Terminal fringe whitish (Fig. 6 A, D). — Head: 1.3 times broader than long. Mandibles bidentate, moderately crossed. Galea very finely shagreened. Labral process subquadrate to weakly rectangular, apical margin concave (Fig. 6 B). Clypeus weakly domed to almost flat, very shiny and more or less completely smooth, occasionally with hints of transverse striation on basal half, punctation strong and dense, distance between punctures 0.5 – 1 puncture diameters. Flagellomere 0.8 – 1.0 times as long as broad, 0.9 – 1.1 times as long as 3, 1.1 – 1.4 times as long as 2. Frons honeycomb-areolated, with dense punctures embedded within the rugosity. Ocelloccipital distance about 1.2 ocellus diameters (Fig. 6 C). Preoccipital ridge weakly carinate. — Mesosoma: Dorsolateral angle of pronotum not elevated. Anterior margin of scutum shagreened, rest of scutum and scutellum extremely shiny and polished-smooth, strongly punctured, distance between punctures 0.5 – 2 puncture diameters (Fig. 6 C). Mesepisternum honeycomb-areolated, with dense punctures embedded within the rugosity. Anteroventral corner of propodeum reticulate, rest of propodeum rugose-areolate. Propodeal triangle occasionally delineated by partial carina, basally radially rugose to rugose-areolate, similar to flanking areas, apically very finely areolated (Fig. 6 C). Recurrent vein 1 meeting submarginal cell 2 slightly proximal to its middle. Nervulus antefurcal to interstitial (Fig. 6 A). — Metasoma: Sculpturing of tergal discs highly variable, basally more or less shagreened and matt, occasionally with fine transverse striation, apically shagreened and matt to completely smooth and shiny. Tergal discs with distinct, dense fine punctation, distance between punctures about 1 puncture diameter. Tergal marginal zones moderately depressed, superficially shagreened to smooth, impunctate, 2 – 3 centrally occupying about 0.3 and 0.5 of tergal length, respectively (Fig. 6 D). — Genitalia and hidden sterna: Gonocoxites with small, weakly rounded dorsal lobe. Gonostyli simple, finger-shaped, more or less uniformly broad, rounded apically. Penis valves moderately broad basally, parallel-sided close to visible base, then tapering apically (Fig. 20 H). Sternum 8 columnar, apical process broadened, apical margin more or less blunt.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
5F360FCD123B5F89BBCF24F4CC09B9A3.taxon	distribution	Distribution and habitat. Shrublands and semi-arid habitats in south-central Turkey and the the Levant * (Northern Israel *, Jordan *).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
FB2B97620BD85CEF8D4B58B2F91399F3.taxon	description	Figures 14 F, J, 19 C, P	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
FB2B97620BD85CEF8D4B58B2F91399F3.taxon	distribution	Distribution and habitat. High altitudes in the Levant (northern Israel, Lebanon, Syria) and Turkey, above 900 m. (Pisanty et al. 2022 b).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
E46964E1EC295571923F594FF85CB534.taxon	description	Figures 18 A, G, 20 O	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
E46964E1EC295571923F594FF85CB534.taxon	distribution	Distribution and habitat. Endemic to Mediterranean shrublands in Cyprus and the Levant (Israel, Lebanon *) (Pisanty et al. 2022 b). Previously reported from Lebanon as A. tiaretta (Wood et al. 2020; Boustani et al. 2021).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
5B62CAB13C6354ACB2CD9CEB7B295715.taxon	description	Figures 7, 18 F, L, 21 A	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
5B62CAB13C6354ACB2CD9CEB7B295715.taxon	etymology	Etymology. Nominative feminine singular form of the Latin adjective chananaeus (= Canaanite).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
5B62CAB13C6354ACB2CD9CEB7B295715.taxon	diagnosis	Diagnosis. Within the Andrena minutula species group, the female of A. chananaea is characterized by the combination of a shagreened, strongly punctate clypeus, often with a strong impunctate midline (Fig. 18 F), densely and strongly punctate scutum (Figs 7 C, 18 L), and shagreened tergal discs with at most obscure punctation (Fig. 7 D). These characters apply equally well to A. minutula (Kirby) itself, and we could not identify any good morphological character that separates females of the two taxa. Within the Levantine Micrandrena fauna, the female of A. chananaea most closely resembles A. sillata Warncke and A. alfkenelloides cardalia Warncke. Compared to A. sillata, A. chananaea has stronger and coarser clypeal puncturing (Fig. 18 E, F), and slightly sparser and coarser scutal punctation (Fig. 18 K, L). Compared to A. a. cardalia, it has a more evenly sculptured clypeus (very densely punctate and transversely striated basally, partly smooth medioapically in A. a. cardalia; Figs 16 A, 18 F) and a dull, microreticulate, and very weakly punctate tergal disc 1 (Fig. 7 D; usually smoother and more distinctly punctate in A. a. cardalia, especially in the second generation). The male of A. chananaea is very hard to diagnose among members of the A. minutula group, due to its extreme morphological variability, both among and within the two seasonal generations. The main consistent characters are a short flagellomere 1, clypeus and scutum which are densely punctured and usually at least partly shagreened, terga which are highly variable but at least weakly punctate, and simple genitalia. The male is closest to A. alfkenella Perkins and A. alfkenelloides cardalia, but exhibits subtle differences in the shape of the genitalia, with the gonostyli only slightly narrowed basally (inner margin of gonostylus more strongly narrowing basally in A. alfkenella and A. a. cardalia), and the penis valves distinctly broader basally vs. apically (more uniformly narrow in the comparison species) (Fig. 21 A – C). In general overview, the genitalia of A. chananaea appear somewhat more robust and enlarged. In addition, males of A. chananaea usually possess a longer flagellomere 1 when compared to A. a. cardalia, and tergal discs that are more strongly shagreened and more weakly and finely punctured when compared to both A. alfkenella and A. a. cardalia, but all these traits are far too variable among individuals of these species to constitute reliable differentiating characters. The male of A. chananaea may also resemble A. minutuloides, but its scutal punctation is denser, and its penis valves are narrower basally (Fig. 21 A, F).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
5B62CAB13C6354ACB2CD9CEB7B295715.taxon	description	Description. First generation female. Body length: 5.5 – 6 mm. — Integumental colour: Body and legs black. Flagellum black, partly covered by greyish scales especially on distal flagellomeres. Tarsi black to brown. Wings weakly infuscate, veins and stigma brown (Fig. 7 A). Tergal marginal zones black, apical margin black to brownish (Fig. 7 D). — Pubescence: Face mostly with light brownish to golden, moderately dense hair, on clypeus semi-erect, short and thin, on paraocular area erect and medium-lengthed, on supraclypeal area and scape erect and short, on frons and preoccipital ridge erect, short to long, on genal area semi-erect to erect, short to medium (Fig. 7 A – C). Few inconspicuous short dark hairs occur on ocellar triangle and behind upper margin of compound eyes. Hair of facial fovea brownish on upper ⅓ of fovea, more whitish below (Fig. 7 B, C). Scutum with brownish hair, centrally short to minute, interspersed with few long erect hairs, underlying cuticle visible, peripherally short to medium and erect. Metanotum and periphery of scutellum with erect, short to long brownish hair (Fig. 7 C). Mesepisternum with long whitish hair (Fig. 7 A). Propodeal corbicula incomplete, posterodorsal fringe composed of long plumose whitish hair, corbicular surface with few long, simple whitish hairs. Leg hair mostly whitish to golden, flocculus incomplete, white, tibial scopal hair simple, whitish, becoming brown near posterobasal margin of tibia (Fig. 7 A). Majority of tergal discs 1 – 2 essentially hairless, 3 – 4 with minute inconspicuous bright hair, tergal discs laterally with few sparse short to medium white hairs. Tergal marginal zones 2 – 4 with prominent, broadly interrupted white hair bands, consisting of very sparse medium-lengthed hairs emerging from the basal rim of the marginal zone, and denser short hairs emerging near its apex. Tergal marginal zone 4 with very weak continuous band of very sparse medium-lengthed whitish hairs emerging from its basal rim. Terminal fringe light brown, flanked by few white hairs (Fig. 7 D). — Head: 1.2 times broader than long. Mandibles bidentate, moderately crossed. Galea finely shagreened (Fig. 7 B). Labral process trapezoidal, not much broader than long. Clypeus domed, entirely shagreened and matt except for an occasionally shinier apical margin, punctation strong, distance between punctures 1 – 2 puncture diameters, an impunctate midline is usually weakly to strongly developed (Figs 7 B, 18 F). Ventrolateral part of paraocular area very smooth, strongly and densely punctured. Frons and rest of paraocular area strongly longitudinally striated, with coarse punctures embedded between the striations (Fig. 7 B, C). Flagellomere 1 0.9 – 1 times as long as 2 + 3, 2 slightly shorter than 3 (Fig. 7 B). Facial fovea dorsally moderately narrow, here about 0.4 times as broad as antennocular distance, tapering ventrally, extending from level of lower end of lateral ocellus to slightly above base of clypeus, lower ⅔ separated from compound eye by smooth cuticular strip. Frons with distinct medial carina, dorsally usually ending with a smooth broadened flat area under the medial ocellus. Distance of fovea from lateral ocellus about 2 ocellus diameters (Fig. 7 B, C). Lateral ocelli not connected posteriorly by trough-like furrow. Ocelloccipital distance roughly 1 ocellus diameter. Preoccipital ridge weakly carinate (Fig. 7 C). — Mesosoma: Dorsolateral angle of pronotum not elevated. Scutum usually strongly shagreened and matt, rarely weakly to strongly shiny and partly smooth, punctation strong and dense, distance between punctures <0.5 to 1 puncture diameter. Scutellum similar, anterior part often shiny and somewhat smooth (Figs 7 C, 18 L). Mesepisternum posteriorly finely alveolate, anteriorly gradually strongly and densely, obliquely punctured. Surface of propodeal corbicula reticulated, with few large punctures around hair bases. Posterior part of propodeum strongly differentiated into horizontal vs. vertical parts. Posterolateral area of propodeum rugose-areolate. Propodeal triangle delineated from flanking areas by change in sculpture, not surrounded by carina, horizontal part coarsely radially rugose, vertical part very finely areolated (Fig. 7 C). Hind pretarsal claw with small inner tooth. Recurrent vein 1 meeting submarginal cell 2 more or less at its middle. Nervulus distinctly antefurcal to almost interstitial (Fig. 7 A). — Metasoma: Tergal discs 1 – 2 strongly shagreened and matt, with hints of fine punctation blended into the shagreening; 3 and especially 4 often more weakly shagreened, 3 with stronger hints of fine, dense punctation. Tergal marginal zones impunctate, weakly depressed, moderately arched, 2 – 3 centrally occupying 0.4 – 0.7 of tergal length, 1 – 2 distinctly shagreened and matt, the following gradually more weakly shagreened (Fig. 7 D). Pygidial plate normally developed. — Second generation female. Body length: 5 – 6.5 mm. — Integumental colour: As in first generation, but flagellum often brown to reddish-brown anteriorly. — Pubescence: Similar to first generation, but overall somewhat sparser, shorter and brighter, without dark hairs on ocellar triangle and upper margin of compound eye. — Head: Similar to first generation, but impunctate midline of clypeus usually absent or poorly developed. — Mesosoma: As in first generation, but scutum and scutellum nearly always strongly shiny, partly to fully smooth at least on disc. Sculpturing of horizontal part of propodeal triangle slightly finer, more rugose-areolate. — Metasoma: As in first generation. — First generation male. Body length: 5 – 6 mm. — Integumental colour: Body and legs black. Flagellum black, partly covered by greyish scales especially on distal flagellomeres. Tarsi black to brown. Wings weakly infuscate, veins and stigma brown (Fig. 7 E). Tergal marginal zones black to golden-brown (Fig. 7 H). — Pubescence: Face, including clypeus, supraclypeal and paraocular areas, scape and frons, with moderately dense, semi-erect to erect, medium to long black hair. A few whitish hairs can often be noted near apex of clypeus and on scape, exact colour dependent on angle of view. Preoccipital ridge with long erect whitish to brownish hairs. Dorsal part of genal area with short to medium erect black hairs, ventral part with long erect whitish hairs (Fig. 7 E – G). Scutum, scutellum and metanotum with long erect hairs, mostly whitish to brownish, a few dark, these hairs sparse on scutum, here interspersed with shorter inconspicuous hairs, dense on metanotum and periphery of scutellum. Mesepisternum with very long erect hairs, whitish on ventral part, usually black and rarely whitish on dorsal part (Fig. 7 E, G). Propodeum with long hairs, mostly whitish, some dark hairs often occur near base of wing. Leg hair mostly white, some weak golden reflections appear mostly on tarsi (Fig. 7 E). Base of tergum 1 and lateral areas of all terga with sparse, short to medium white hair; rest of tergal discs with minute inconspicuous whitish hair. Tergal marginal zones 2 – 3 with weak, broadly interrupted white hair bands, concentrated mostly near apex of marginal zone. Tergal marginal zone 4 with extremely weak hints of similar hair bands. Terminal fringe whitish (Fig. 7 H). — Head: 1.2 times broader than long (Fig. 7 F). Mandibles bidentate, moderately crossed. Galea finely shagreened. Labral process trapezoidal to subquadrate, not much broader than long, apical half usually elevated, apical margin concave. Clypeus domed, centrally often somewhat flattened, basally strongly shagreened and matt with some transverse striations, apically usually shiny and weakly shagreened, rarely strongly shagreened and matt. Clypeus punctation strong and dense, distance between punctures <0.5 to 1 puncture diameter, puncture density decreasing medioapically, often creating a distinct impunctate midline at least on apical half. Lower part of paraocular area shiny and strongly punctured, upper part with narrow strip of fine, dense punctation along inner margin of compound eye, elsewhere strongly longitudinally shagreened to rugose-areolate, some embedded punctures can occasionally be discerned. Flagellomere 1 1.1 – 1.2 times longer than broad, usually 1.45 – 1.8 times longer than 2 and 1.05 – 1.2 times longer than 3, rarely slightly shorter (Fig. 7 F). Frons strongly rugose-areolate, with some embedded punctures especially near centre. Ocelloccipital distance 1 ocellus diameter. Preoccipital ridge moderately carinate (Fig. 7 G). — Mesosoma: Dorsolateral angle of pronotum not elevated. Scutum fully shagreened and matt, strongly punctured, distance between punctures 0.5 – 1.5 puncture diameters, some punctures with elevated margins, especially on anterior half of scutum. Scutellum similar, often somewhat smoother centrally, punctures without elevated margins (Fig. 7 G). Mesepisternum with fine alveolation, overlayed by sparse coarse oblique punctures. Anterolateral part of propodeum ventrally reticulated, dorsally gradually rugose-areolated. Posterolateral part of propodeum strongly, finely rugose-areolate. Horizontal part of propodeal triangle strongly, coarsely radially rugose to rugose-areolate, vertical part very finely areolated (Fig. 7 G). Recurrent vein 1 meeting submarginal cell 2 more or less near its middle. Nervulus antefurcal to almost interstitial (Fig. 7 E). — Metasoma: Tergal discs exhibiting high variability in sculpture, usually shagreened basally and weakly shagreened to smooth apically, proximal terga with stronger shagreening. Very fine punctation is discernible especially on the smoother areas, distance between punctures 1 – 2 puncture diameters. Tergal marginal zones not to weakly depressed, very shiny, smooth to very superficially shagreened, essentially impunctate (Fig. 7 H). — Genitalia and hidden sterna: Gonocoxites apically truncate, without dorsal lobe. Gonostyli simple, finger-shaped, very slightly broadening apically, blade flattened, apical margin rounded. Penis valves moderately narrow, basal ¼ of visible area parallel-sided, the following ¼ strongly tapering apically, distal half very narrow (Fig. 21 A). Sternum 8 columnar, apical process somewhat broadened, apical margin rounded to truncate. — Second generation male. Body length: 5 – 5.5 mm. — Integumental colour: As in first generation. Pubescence: Only a single individual is available to us with intact hair, hence the description in this section should be considered preliminary. Clypeus with sparse short whitish hairs. Paraocular area, scape and area around antennal sockets with short to medium, mixed bright and dark hairs. Preoccipital ridge with medium-lengthed erect whitish hairs. Genal area with whitish hairs, short dorsally, medium-lengthed ventrally. Hairs of scutum, scutellum and metanotum completely whitish. Rest of pubescence as in first generation. — Head: Clypeus at least on apical ⅔ very shiny, partly to fully smooth. Rest of head as in first generation. — Mesosoma: Scutum and scutellum shiny, weakly shagreened to completely smooth, very strongly punctured. Mesepisternum with fine alveolation overlayed by coarse regular (not oblique) punctures with raised margins. Rest of mesosoma as in first generation. Metasoma: Tergal discs strongly shiny, partly shagreened to fully smooth, punctation strong, fine to moderately sized, distance between punctures 1 – 2 puncture diameters. Tergal marginal zones as in first generation. — Genitalia and hidden sterna: As in first generation.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
5B62CAB13C6354ACB2CD9CEB7B295715.taxon	distribution	Distribution and habitat. Shrubland and mesic habitats in all countries of the Levant. Previously reported from Lebanon as A. minutula (Wood et al. 2020; Boustani et al. 2021).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
A09A1B2E93075FF1836C5BD28FF52F0D.taxon	description	Figures 15 F, 20 I	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
A09A1B2E93075FF1836C5BD28FF52F0D.taxon	distribution	Distribution and habitat. Endemic to the southern Levant, in scrublands and semi-arid habitats (northern Israel, Jordan; Pisanty et al. 2022 b).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
2443EF0ACBFC56A198D214FCADAC2F10.taxon	description	Figures 13 G, N, 19 G	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
2443EF0ACBFC56A198D214FCADAC2F10.taxon	distribution	Distribution and habitat. Coastal habitats along the south Mediterranean. In the Levant, limited to Israel’s coastal plain, likely extending into Lebanon (Warncke 1969, as A. longibarbis Pérez; Pisanty et al. 2022 b).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
5B3D7D06DDB95DECB713DA5C8A87915E.taxon	description	Figures 14 D, O, 19 N	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
5B3D7D06DDB95DECB713DA5C8A87915E.taxon	distribution	Distribution and habitat. Endemic to Mediterranean shrublands in the Levant (northern Israel, likely also Lebanon and Syria).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
18A795175B5158C3A0C7C2C7655DE6AC.taxon	description	Figures 15 I, 20 U	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
18A795175B5158C3A0C7C2C7655DE6AC.taxon	distribution	Distribution and habitat. Broadly west and central Palaearctic (Gusenleitner & Schwarz 2002). Newly reported from the Levant * (Jordan *, Lebanon *).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
ED895AA0B54D58248806AB27E45D7FAE.taxon	description	Figures 14 L, 20 E	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
ED895AA0B54D58248806AB27E45D7FAE.taxon	distribution	Distribution and habitat. High elevations above 1400 m in Syria and Iran.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
F9ABA8117F6D504DA63C45ADD6FCE802.taxon	description	Figures 8, 16 B, H, L, 20 K	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
F9ABA8117F6D504DA63C45ADD6FCE802.taxon	etymology	Etymology. Named after the Israeli coleopterist Ariel Leib Leonid (a. k. a. Laibale) Friedman, who collected most of the type series, as well as many other rare and unusual Hymenoptera.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
F9ABA8117F6D504DA63C45ADD6FCE802.taxon	diagnosis	Diagnosis. Andrena friedmani is distinguished from other Levantine Micrandrena by the combination of small body size, finely transversely striated clypeus in the female (Figs 8 B, 16 B), dark flagellum (Fig. 8 A, B), polished-smooth scutum (Figs 8 C, 16 H), and partly smooth, finely punctate tergum 1 (Figs 8 D, 16 L). Andrena tkalcui Gusenleitner & Schwarz and A. calandra Warncke are also small-sized with a shiny scutum, but easily differ by the flagellum which is anteriorly orange. In addition, in A. tkalcui, the female clypeus is not striated (Fig. 17 C), the scutum is often partly shagreened centrally (Fig. 17 H), tergum 1 is shagreened and impunctate, the dorsal gonocoxite lobe is rounded apically (Fig. 20 F), and sternum 8 is apically emarginate (Fig. 20 W). In the female of A. calandra, tergum 1 is shagreened and impunctate, whereas in the male, all terga are more strongly punctured (Fig. 6 D), and the genitalia lacks a strong dorsal gonocoxite lobe (Fig. 20 H).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
F9ABA8117F6D504DA63C45ADD6FCE802.taxon	description	Description. Female. Body length: 5.5 – 6 mm. — Integumental colour: Body black. Flagellum dark brown, slightly tinted reddish. Legs brown to black. Wings slightly infuscate, veins golden to brown, stigma golden centrally (Fig. 8 A). Tergal marginal zones black basally, more brownish apically (Fig. 8 D). — Pubescence: Clypeus with rather sparse, medium-lengthed, semi-erect white hair. Supraclypeal area with short erect white hair. Paraocular area and scape with dense, medium-lengthed erect white hair (Fig. 8 A, B). Frons and ocellar triangle with sparse, medium-lengthed white hair. Preoccipital ridge with long, erect whitish hair (Fig. 8 A – C). Lower of half of facial fovea with whitish hair, upper half with brownish to whitish hair, depending on angle of view (Fig. 8 C). Scutum, scutellum and metanotum with erect whitish hair of varying length, longer and thicker peripherally, mostly minute and inconspicuous centrally, underlying cuticle visible (Fig. 8 A, C). Mesepisternum with very long white hair (Fig. 8 A). Propodeal corbicula incomplete, dorsoposterior fringe with long white hair, corbicular surface with sparse and long, simple white hairs. Leg hair mostly whitish; flocculus incomplete, white; tibial scopa composed of white simple hairs (Fig. 8 A). Tergal discs centrally with minute inconspicuous hair, laterally with small patches of sparse, short white hair. Tergal marginal zones 2 – 3 with broadly interrupted, weak bands of white hair; 4 with very weak, almost continuous band of extremely sparse whitish hair. Terminal fringe yellowish- to brownish-white (Fig. 8 A, D). — Head: 1.2 times broader than long. Galea very finely shagreened (Fig. 8 B). Labral process trapezoidal, apical margin blunt (Fig. 16 B). Clypeus moderately domed, basal half matt and finely transversely striated, apical half gradually more finely shagreened, becoming partly shiny near apex. Clypeus punctation of moderate strength and size, distance between punctures 1 – 2 puncture diameters (Figs 8 B, 16 B). Upper part of paraocular area longitudinally striated. Flagellomere 1 about 1.2 times longer than 2 + 3, 2 slightly shorter than 3 (Fig. 8 A, B). Frons finely, strongly rugose-areolate, with complete medial carina, flanked by very fine areolation almost resembling dense punctures (Fig. 8 B). Facial fovea moderately broad and very shallow above, 0.4 times as broad as antennocular distance, deeper and somewhat narrower below, extending from level of lower end of lateral ocellus to slightly above base of clypeus, fovea almost adjacent to compound eye, lower half occasionally separated by very narrow, shiny cuticular strip. Distance of fovea from lateral ocellus about 2 ocellus diameters (Fig. 8 B, C). Ocelloccipital distance about 0.8 ocellus diameter. Preoccipital ridge moderately carinate (Fig. 8 C). — Mesosoma: Dorsolateral angle of pronotum not elevated. Scutum and scutellum very shiny, almost completely smooth, strongly and densely punctured, distance between punctures 0.5 – 1 puncture diameter (Figs 8 C, 16 H). Posterior part of mesepisternum finely alveolate, anterior part strongly and densely, obliquely punctured. Surface of propodeal corbicula finely alveolate, with large punctures around hair bases, surrounded by weak star-shaped wrinkles. Posterior part of propodeum strongly differentiated into horizontal vs. vertical regions, strongly rugose-areolated. Propodeal triangle poorly differentiated from flanking regions, basal half very coarsely rugose-areolated, becoming more radially rugose near basal margin, apical half gradually very finely sculptured (Fig. 8 C). Hind pretarsal claw with distinct inner tooth. Recurrent vein 1 meeting submarginal cell 2 close to its middle. Nervulus more or less interstitial (Fig. 8 A). — Metasoma: Tergal disc 1 shiny and more or less smooth, with fine, irregular punctation becoming dense apically (Figs 8 D, 16 L). Tergal discs 2 – 4 strongly shagreened and matt, essentially impunctate. Tergal marginal zones weakly depressed, weakly arched, 2 – 3 centrally occupying 0.4 – 0.5 of tergal length, cuticle shiny, mostly shagreened, apically often partly smooth, punctation absent (Fig. 8 D). Pygidial plate normally developed. — Male. Body length: 5 – 5.5 mm. — Integumental colour: Body and legs black. Flagellum black to brown. Apical tarsomeres black to brown. Wings almost hyaline, veins golden to light brown, stigma golden centrally (Fig. 8 E). Tergal marginal zones black basally, dark brown to reddish-brown apically (Fig. 8 H). — Pubescence: Completely white. Clypeus with dense long hair, underlying cuticle partly concealed. Paraocular area with moderately dense, medium-lengthed hair. Scape and area around antennal sockets with dense long hair (Fig. 8 E, F). Frons and ocellar triangle with moderately dense, medium-lengthed erect hairs. Preoccipital ridge with long erect hairs. Genal area with medium-lengthed hair dorsally, long hair ventrally (Fig. 8 E – G). Scutum, scutellum and metanotum with sparse, short to long erect hair, longest and most prominent peripherally, underlying cuticle visible (Fig. 8 G). Mesepisternum with very long hair (Fig. 8 E). Propodeum with long hair. Terga centrally with inconspicuous short thin hair, laterally with moderately dense medium-lengthed hair, on marginal zones 2 – 4 creating distinct, broadly interrupted hair bands (Fig. 8 E, H). — Head: 1.25 times broader than long (Fig. 8 F). Labral process broad trapezoidal, apical margin concave. Clypeus domed, very shiny, strongly and very densely punctured, distance between punctures 0 – 1 puncture diameters, without impunctate midline. Lower part of paraocular area smooth and densely punctate, upper part with a narrow, finely punctured strip near inner margin of compound eye, elsewhere strongly longitudinally striated. Flagellomere 1 0.9 times shorter than 2 + 3, 2 distinctly shorter than 3 (Fig. 8 F). Frons strongly rugose-areolate, punctures blended within the rugosity can occasionally be discerned. Ocelloccipital distance about 1 ocellus diameter. Preoccipital ridge moderately carinate (Fig. 8 G). — Mesosoma: Dorsolateral angle of pronotum not elevated. Scutum very shiny, partly shagreened on anterior margin, elsewhere weakly roughened to almost completely polished, punctation dense and strong, distance between punctures 0.5 – 1.5 puncture diameters. Scutellum completely polished, similarly punctured (Fig. 8 G). Mesepisternum finely alveolate, overlayed by dense, shallow oblique punctures creating an almost honeycomb-like shallow rugosity. Anteroventral edge of propodeum reticularly shagreened, rest of propodeum coarsely areolated. Propodeal triangle strongly and coarsely sculptured, basal part radially rugose, apical part rugose-areolate (Fig. 8 G). Recurrent vein 1 meets submarginal cell 2 close to its middle or proximal to it. Nervulus interstitial to very slightly postfurcal. — Metasoma: Tergal disc 1 shiny and more or less smooth basally, weakly to strongly shagreened apically, punctation fine, distance between punctures 0.5 – 2 puncture diameters. Tergal discs 2 – 4 matt, strongly granularly shagreened, apically occasionally partly smooth, punctation dense, usually hardly discernible from underlying shagreening, distance between punctures about 1 puncture diameter. Tergal marginal zones weakly to strongly shagreened basally, gradually smooth apically, impunctate, 2 – 4 distinctly depressed (Fig. 8 H). — Genitalia and hidden sterna: Gonocoxites with strong, elongate and pointed, parallel-sided dorsal lobes. Gonostyli simply built, elongate, finger-shaped, blade flattened. Penis valves narrow and elongate, very slightly broadening close to visible base, hereafter gradually tapering apically (Fig. 20 K). Sternum 8 columnar, broadening apically, apical margin blunt-ended.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
F9ABA8117F6D504DA63C45ADD6FCE802.taxon	distribution	Distribution and habitat. Endemic to scrublands in the southwestern Levant (northern Israel, West Bank).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
D3A6F624E27E57DFB7F96E95A81ADAA5.taxon	description	Figures 13 J, K, 19 I	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
D3A6F624E27E57DFB7F96E95A81ADAA5.taxon	distribution	Distribution and habitat. Desert habitats in north Africa and the Levant (southern Israel, Jordan *, Syria *, Pisanty et al. 2018).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
F37F4D204ADA511CB97FD2524DF4650E.taxon	description	Figures 9, 17 E, J, 21 J	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
F37F4D204ADA511CB97FD2524DF4650E.taxon	etymology	Etymology. Nominative feminine singular form of the Latin adjective “ hebraicus ” (= Hebrew).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
F37F4D204ADA511CB97FD2524DF4650E.taxon	diagnosis	Diagnosis. Within the Andrena minutula species group, A. hebraica belongs to the species around A. spreta Pérez, which are characterized in the female by a clypeus which is non-rugose, weakly domed and partly to fully shagreened, a scutum which is shagreened to partly smooth, with fine, relatively sparse punctures, and terga which are mostly shagreened and impunctate. In the Levant and Cyprus, these include A. aphroditae sp. nov., A. minutuloides, A. spreta, A. tiaretta and A. tkalcui. Andrena hebraica differs from all the above species by the sculpturing of tergal disc 3 and to a lesser extent 2, which are apically shiny, instead of completely shagreened and matt. This difference is difficult to illustrate by photographs, but is clearly visible to the eye under magnification. In addition, the clypeus is more clearly domed and more strongly and extensively shagreened compared to A. spreta and A. tiaretta (Figs 8 B, 17 E), the clypeus is more finely punctured and the scutum more densely punctured compared to A. minutuloides, and the flagella and wing veins are darker compared to A. tkalcui. Within the Levantine and Cypriot members of the A. minutula species group, the male of A. hebraica is characterized by a flagellomere 1 which is subequal to slightly longer than 3, clypeus and scutum which are partly to fully shagreened and densely punctured, tergal discs 2 and 3 which are more strongly shagreened basally vs. apically, and characteristic genitalia lacking dorsal gonoxocite lobes and with gonostyli that converge apically. The genitalia are almost identical to those of A. povolnyi Warncke stat. nov. from Afghanistan, but they clearly differ from other similar species in the region of study as follows: lack of dorsal gonocoxite lobes (present in A. cervina and A. spreta), gonostyli more strongly converging apically compared to A. spreta and A. aphroditae, gonostyli slenderer compared to A. aphroditae, gonostyli shorter and penis valves shorter and narrower compared to A. tiaretta (Figs 20 J, 21 G, I, J).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
F37F4D204ADA511CB97FD2524DF4650E.taxon	description	Description. Female. Body length: 5.5 – 6.5 mm. — Integumental colour: Body and legs predominantly black. Flagellum black, partly covered by grayish setae, distal flagellomeres occasionally brownish. Apical tarsomeres brown to black. Wings weakly infuscate, veins brown, stigma brownish centrally (Fig. 9 A). Tergal marginal zones black, apical margins black to brown (Fig. 9 D). — Pubescence: Body and leg hair mostly brightly coloured. Clypeus with sparse, short and thin, semi-erect white hairs. Supraclypeal area with short whitish hairs. Paraocular area and scape with dense, medium-lengthed whitish hairs (Fig. 9 A – C). Frons with sparse, medium-lengthed whitish hairs. Facial fovea in dorsal view with brownish hair on upper ⅓ of fovea, white hair on lower ⅔ (Fig. 9 B, C). Preoccipital ridge with long erect whitish hair. Genal area with whitish hair, short on dorsal part, medium to long on ventral part (Fig. 9 A – C). Scutum, scutellum and metanotum with moderately dense, erect whitish to yellowish hair, centrally minute to short, with some scattered longer hairs, underlying cuticle visible, peripherally short to medium-lengthed (Fig. 9 A, C). Mesepisternum with long whitish hair (Fig. 9 A). Propodeal corbicula incomplete, posterodorsal fringe with long whitish plumose hairs, corbicular surface with sparse, long simple whitish hairs. Leg hair mostly whitish, some golden reflections appear especially on tarsi. Flocculus incomplete, white. Tibial scopal hairs simple, whitish, becoming brownish near posterobasal edge of tibia (Fig. 9 A). Tergal discs centrally with sparse, inconspicuous minute thin hair, laterally with sparse short to medium white hair. Tergal marginal zones 2 – 4 with distinct bands of white hair, moderately dense and broadly interrupted on 2 – 3, sparser but almost continuous on 4. Terminal fringe golden to brownish, flanked by some long white hairs (Fig. 9 A, D). — Head: 1.2 times broader than long. Mandibles bidentate, moderately crossed. Galea finely shagreened (Fig. 9 B). Labral process fairly large, trapezoidal, not much broader than long. Clypeus weakly domed, surface entirely finely shagreened and matt, except for an occasional shinier apical margin, punctation moderately strong, distance between punctures 1 – 2 puncture diameters, an impunctate midline is not clearly discernible (Figs 9 B, 17 E). Supraclypeal area finely longitudinally striated. Lowermost part of paraocular area smooth and shiny, densely and finely punctured. Frons and rest of paraocular area strongly longitudinally striated, some interspersed punctures can occasionally be discerned on frons. Flagellomere 1 as long as 2 + 3, 2 about as long as 3 (Fig. 9 B). Facial foveae narrow and elongate, extending from level of lower end of lateral ocellus to slightly above base of clypeus, upper part 0.4 – 0.5 times as broad as antennocular distance, lower ⅔ distinctly narrowed, here separated from compound eye by distinct strip of smooth cuticle. Distance of fovea from lateral ocellus about 2 ocellus diameters (Fig. 9 B, C). Lateral ocelli connected posteriorly by trough-shaped horizontal furrow. Ocelloccipital distance about 0.9 ocellus diameter. Preoccipital ridge weakly to distinctly carinate (Fig. 9 C). — Mesosoma: Dorsolateral angle of pronotum not elevated. Entire scutum strongly shagreened and matt, densely punctured, distance between punctures 0.5 – 1.5 puncture diameters, puncture strength varying among individuals. Scutellum similar, occasionally somewhat shinier (Figs 9 C, 17 J). Mesepisternum strongly, finely reticulated, anterior part overlayed with dense, strong oblique punctures. Surface of propodeal corbicula finely reticulated, with few large punctures around hair bases. Posterior part of propodeum strongly differentiated into horizonal vs. vertical regions. Propodeal triangle delineated by lateral carinae, horizontal part strongly and coarsely rugose, radially rugose near base, rugose-areolate further down, vertical part very finely areolated (Fig. 9 C). Posterolateral part of propodeum strongly, finely rugose-areolate. Hind pretarsal claw with inner tooth. Recurrent vein 1 meets submarginal cell 2 at its middle or slightly proximal to it. Nervulus interstitial to weakly antefurcal (Fig. 9 A). — Metasoma: Tergal disc 1, basal part of disc 2 and occasionally of 3 strongly shagreened and matt, impunctate. Tergal disc 4, apical part of tergal disc 2 and at least apical part of disc 3 shiny, partly to fully smooth, the tight shagreening of basal areas here gradually disintegrating into irregular, dense fine punctation, distance between punctures up to 1 puncture diameter. Tergal marginal zones weakly depressed, impunctate, 2 – 3 centrally occupying 0.4 – 0.5 of tergum length, 1 – 2 distinctly shagreened, the following gradually more weakly so (Fig. 9 D). Pygidial plate normally developed. — Male. Body length: 5 – 6 mm. — Integumental colour: As in female (Fig. 9 E, H). — Pubescence: Clypeus with moderately dense, medium-lengthed, semi-erect white hairs, underlying cuticle visible (Fig. 9 E, F). Supraclypeal area with short erect whitish hair. Paraocular area with moderately dense, medium-lengthed erect hair, whitish to black on outer part bordering compound eye, whitish on inner part. Frons and paraocular area with sparse, erect short to medium hair of varying brightness, usually with predominantly dark hairs on dorsal parts (Fig. 9 F, G). Preoccipital ridge with long erect whitish hairs. Genal area often with few short to medium erect dark hairs around upper margin of compound eye, elsewhere with whitish hairs, short dorsally, gradually long ventrally (Fig. 9 E – G). Scutum, scutellum and metanotum with moderately dense erect whitish hair, centrally with mixed short and long hairs, underlying cuticle visible, peripherally medium to long. Mesepisternum and propodeum with very long white hairs (Fig. 9 E, G). Leg hair mostly white, with some subtle golden reflections mostly on tarsi (Fig. 9 E). Tergal discs centrally with minute inconspicuous bright hair, laterally and on base of disc 1 with moderately dense, medium-lengthed white hair. Tergal marginal zones 2 – 4 with strong, broadly interrupted bands of white hair, slightly extending onto following discs. Tergal marginal zone 5 with very sparse, continuous band of long whitish hair. Terminal fringe whitish (Fig. 9 E, H). — Head: About 1.2 times broader than long (Fig. 9 F). Mandibles bidentate, moderately crossed. Galea finely shagreened. Labral process trapezoidal to almost rectangular, apical margin concave. Clypeus moderately domed, strongly shagreened and matt basally, shiny and weakly shagreened apically, extent of shagreening highly variable among individuals, punctation moderately dense, distance between punctures 0.5 – 1.5 puncture diameters, without impunctate midline. Outer part of paraocular area bordering compound eye smooth and finely punctured. Supraclypeal area, frons and inner part of paraocular area longitudinally striated, occasionally with hints of punctures blended into the rugosity. Flagellomere 1 1.1 – 1.3 times longer than 3, distinctly shorter than 2 + 3, 2 distinctly shorter than 3 (Fig. 9 F). Ocelloccipital distance about 1 ocellus diameter. Preoccipital ridge moderately carinate (Fig. 9 G). — Mesosoma: Dorsolateral angle of pronotum not elevated. Scutum and scutellum usually strongly shagreened and matt, rarely weakly shagreened and shiny, punctation distinct, distance between punctures 0.5 – 1.5 puncture diameters (Fig. 9 G). Mesepisternum finely alveolated, overlayed by weak oblique punctures. Anterolateral part of propodeum ventrally reticulated, dorsally gradually rugose-areolate. Posterior part of propodeum rugose-areolate, triangle weakly delineated, radially rugose near base, coarsely rugose-areolate further down, very finely so on vertical part (Fig. 9 G). Recurrent vein 1 meets submarginal cell 2 at its middle or slightly proximal to it. Nervulus interstitial to weakly antefurcal (Fig. 9 E). — Metasoma: As in female (Fig. 9 H). — Genitalia and hidden sterna: Gonocoxites apically truncate, without dorsal lobe. Gonostyli more or less finger-shaped, inner and outer margins with slight kink at mid-length, distal part more strongly converging apically. Penis valves moderately narrow basally, gradually tapering apically (Fig. 21 J). Sternum 8 columnar, broadening apically, apical margin blunt to slightly rounded.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
F37F4D204ADA511CB97FD2524DF4650E.taxon	distribution	Distribution and habitat. Mediterranean shrublands in Israel, the West Bank, Jordan and Lebanon. Mostly absent from the coastal plain, where it is replaced by A. tiaretta.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
5C72F9BBF30D5FA799801F7D97E7DBE6.taxon	description	Figures 13 H, O	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
5C72F9BBF30D5FA799801F7D97E7DBE6.taxon	distribution	Distribution and habitat. Endemic to arid habitats in the southern Levant (southern Israel, West Bank, Jordan).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
C6B058CA6A13546FBFCB265E046D02C9.taxon	description	Figures 2, 13 F, 19 L	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
C6B058CA6A13546FBFCB265E046D02C9.taxon	etymology	Etymology. Named after the Israeli entomologist Jehoshua Kugler (1916 – 2007).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
C6B058CA6A13546FBFCB265E046D02C9.taxon	diagnosis	Diagnosis. Within the Andrena longibarbis species group, A. kugleri belongs to the species around A. mariana Warncke, which are characterized by a largely non-rugose clypeus. Among these species, A. kugleri can be diagnosed by the combination of the relatively narrow female labral process (Fig. 13 F; broader in A. abjecta Pérez, A. leptura Warncke stat. nov. and A. mica Warncke), the flat, apically smooth, non-rugose clypeus (Figs 2 B, F, 13 F; weakly domed in A. abjecta, A. mariana and A. tenostra Warncke, more extensively shagreened in A. obsoleta Pérez and male A. mica, less so in A. rubecula Warncke, basally rugose in A. leptura and A. mica), reddish-brown flagellum (Fig. 2; more orange-lightened in A. govinda Warncke, A. mariana and female A. rubecula, dark in male A. alma Warncke), male flagellomere proportions (Fig. 2 F; flagellomere 1 longer in A. obsoleta), normally developed male gena (broadened in A. alma and A. rubecula), weak pronotal angle (stronger in A. rubecula and male A. alma), dull scutellum (Fig. 2 C, G; shinier in A. govinda, A. rubecula and A. tenostra), impunctate tergal discs (punctate in female A. alma), reddish-brown tergal marginal zones (Fig. 2 D, H; distinctly orange in A. mariana, somewhat more orange-lightened in female A. rubecula), relatively narrow tergal hair bands (Fig. 2 D, H; broader in A. govinda), and relatively narrow penis valves (Fig. 19 L; broader in A. alma and A. leptura).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
C6B058CA6A13546FBFCB265E046D02C9.taxon	description	Description. FEMALE. Body length: 6 – 7 mm. — Integumental colour: Body black. Flagellum brown anteriorly, occasionally with weak reddish hue. Legs black to brown. Wings hyaline, veins and stigma golden (Fig. 2 A). Tergal marginal zones reddish-yellow basally, yellow to whitish apically (Fig. 2 D). — Pubescence: Body hair relatively sparse, mostly minutely plumose, white (Fig. 2 A). Face and genal area with short to medium white hair, densest on paraocular area and scape. Facial fovea with dense, minute whitish hair. Vertex with long erect white hair (Fig. 2 B, C). Dorsal mesosomal surfaces with sparse erect white hair, short on scutum, longer on scutellum and metanotum (Fig. 2 C). Mesepisternum with moderately dense, long white hair (Fig. 2 A). Propodeal corbicula incomplete, dorsoposterior fringe composed of long, plumose white hairs. Corbicular surface with sparse, simple long white hairs. Leg hair white (Fig. 2 A). Flocculus incomplete. Femoral and tibial scopae well-developed, femoral scopa composed of moderately long, curved plumose hairs dorsally and straight long hairs ventrally, tibial scopa composed of simple hairs (Fig. 2 A). Tergal discs 1 – 2 hairless centrally, with sparse inconspicuous short white hair laterally. Tergal discs 3 – 4 entirely with sparse, inconspicuous minute white hair, slightly longer laterally. Tergal marginal zones 2 – 4 with distinct narrow bands of dense white hair, strongly interrupted on tergum 2, almost continuous on 3, continuous on 4. Terminal fringe whitish-golden (Fig. 2 D). — Head: 1.2 times broader than long. Mandibles bidentate, moderately crossed (Fig. 2 B). Galea finely shagreened. Labral process small, subquadrate, as long as broad or longer. Clypeus entirely flat to minutely domed, smooth apically and finely shagreened basally, lacking striations, punctation distinct, distance between punctures 1 – 2 puncture diameters, an impunctate midline is absent or very weakly indicated (Figs 2 B, 13 F). Paraocular and supraclypeal areas and frons finely longitudinally striated, punctation absent except on lower paraocular area (Fig. 2 B). Facial fovea extremely narrow and linear in lower half, broader in upper ⅓ – ½, here about ⅓ as broad as antennocular distance, fovea extending from level of middle – lower end of lateral ocellus to lower end of antennal socket, separated from compound eye by very narrow strip of smooth cuticle. Flagellomere 1 as long as 2 + 3, 2 slightly shorter than 3 (Fig. 2 B). Distance of fovea from lateral ocellus 1.8 ocellus diameters. Vertex distinctly carinate. Ocelloccipital distance equals 0.7 ocellus diameter (Fig. 2 C). — Mesosoma: Dorsolateral angle of pronotum weakly developed. Scutum and scutellum entirely shagreened and dull, scutum with extremely shallow, hardly discernible punctation, distance between punctures 1 – 2 puncture diameters (Fig. 2 C). Mesepisternum and propodeum finely reticulated. Base of propodeal triangle more coarsely reticulated than apical part and flanking propodeal regions, occasionally slightly rugose-areolate at basal margin (Fig. 2 C). Hind pretarsal claw with small inner tooth. Recurrent vein 1 meeting submarginal cell 2 near its middle. Nervulus usually strongly antefurcal, rarely weakly antefurcal to interstitial (Fig. 2 A). — Metasoma: Tergal discs strongly, finely shagreened, impunctate. Tergal marginal zones weakly depressed, weakly broadening medially, shagreened basally and smooth apically, impunctate (Fig. 2 D). Pygidial plate with distinctly elevated central zone. — MALE. Body length: 5.5 – 7 mm. — Integumental colour: As in female (Fig. 2 E). — Pubescence: Body hair longer and denser than in female, mostly minutely plumose, white (Fig. 2 E). Face, vertex and gena with moderately dense, long white hairs. A few short dark hairs occur near centre of inner margin of compound eye (Fig. 2 F). Dorsal mesosomal surfaces with long erect white hair, densest on metanotum and scutellum, sparse on scutal disc (Fig. 2 E, G). Mesepisternum with moderately dense, very long white hairs. Propodeum with long white hairs. Leg hair white to yellowish (Fig. 2 E). Tergal discs with short to minute inconspicuous bright hair, longer laterally and on base of tergum 1. Tergal marginal zones 2 – 4 with weak narrow bands of white hair, strongly interrupted on terga 2 – 3, more weakly so on 4. Terminal fringe whitish-golden (Fig. 2 H). — Head: Labral process quadrate, apical margin concave to blunt-ended. Clypeus entirely flat, apical ⅔ extremely smooth. Flagellomere 1 about 1.2 times as long as 3, 3 distinctly longer than 2 (Fig. 2 F). Ocelloccipital distance about 1 ocellus diameter (Fig. 2 G). Genal area 0.95 – 1.05 times as broad as compound eye. Rest of head as in female. — Mesosoma: As in female (Fig. 2 G). — Metasoma: As in female (Fig. 2 H). — Genitalia and hidden sterna: Dorsal gonocoxite lobes developed, converging apically, pointed. Gonostyli finger-shaped, elongate, weakly broadening apically, inner margin with very small concavity in mid-length. Penis valves moderately broad basally, basal ⅔ gradually tapering apically (Fig. 19 L). Sternum 8 simple, columnar, narrow, apical process broadened, apical margin often weakly emarginate.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
C6B058CA6A13546FBFCB265E046D02C9.taxon	distribution	Distribution and habitat. In a broad sense, Andrena kugleri is endemic to Israel, the West Bank and southern Lebanon. The principal distribution is in desert areas of southern Israel and the West Bank, while the species is absent from most of the Mediterranean region of Israel. However, a disjunct population exists on both sides of the Israeli-Lebanese border, where annual precipitation exceeds 600 mm. This population is known only from females, and the single available barcode (from Lebanon) is 13 % different from the main population. Males should be located and examined to conclude whether this population represents a separate taxon. The species is also known from only two localities in the central coastal plain, one of these is the old botanical garden in southern Tel Aviv, where a great proportion of the type material has originated. A single specimen with a label from Turkey in Warncke’s handwriting is of dubious origin, as noted by Warncke himself. More DNA barcoding is needed to better assess the uniformity of this taxon across its range.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
C1CA1AD8F78C5033826EBEA659DD4B9C.taxon	distribution	Distribution and habitat. Desert habitats in the south-east Mediterranean. Newly reported from the Levant *, in southern Israel *.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
68E0A0BA7C2F52E68C28CEEB8A946129.taxon	description	Figures 10, 15 J, 20 A, M	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
68E0A0BA7C2F52E68C28CEEB8A946129.taxon	etymology	Etymology. From the Latin name for Mount Lebanon, “ Libanus ”, with the feminine suffix - ica, thus “ libanica ” meaning “ of Mount Lebanon ”, given the observed geographic distribution of the species.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
68E0A0BA7C2F52E68C28CEEB8A946129.taxon	diagnosis	Diagnosis. Andrena libanica is a typical member of the A. minutula species group with the propodeal triangle entirely rugose (Fig. 10 C, G). Within this group, it belongs to the species with more or less the apical half of the clypeus polished and shining (Fig. 10 B, F), the facial foveae relatively narrow along their entire length (equaling the diameter of a flagellum, Fig. 10 B), the head overall relatively elongate, around 1.1 times wider than long (Fig. 10 B, F), and the terga distinctly punctate. These are the species around A. rugulosa Stöckhert, which in the Levant and Cyprus are found in mountainous regions usually above 1200 m and include A. libanica, A. alshaykh Pisanty sp. nov. and A. lindbergella Pittioni. The female of Andrena libanica can be separated from all three comparison species due to the punctation of the terga, which have the punctures of the tergal discs separated by <0.5 – 0.5 puncture diameters, these punctures clearly defined against the underlying sculpture (Figs 10 D, 15 J). In A. rugulosa, the tergal punctures are slightly sparser, separated by 0.5 – 1 puncture diameters, but obscure and difficult to see, disappearing into the underlying sculpture. In A. alshaykh and A. lindbergella, the tergal punctation is sporadic, with punctures separated by 0.5 – 3 puncture diameters (Fig. 15 K, L). As in the females, the males can be recognised as close to A. alshaykh and A. lindbergella due to the clearly punctate terga combined with the clypeus smooth and strongly punctured in the apical ½ (Figs 10 F, H, 20 A). Males of A. rugulosa are very similar, but in this species the terga are slightly more obscurely punctate, with punctures separated by 0.5 – 1 puncture diameters and not so strongly differentiated from the underlying shagreen (in A. libanica with punctures clearly distinct against the shagreened and weakly shining terga, punctures separated by <0.5 – 0.5 puncture diameters, Figs 10 H, 20 A), and the scutal punctation slightly disappears into the strongly microreticulate and dull scutum (in A. libanica with the scutum more finely microreticulate, weakly shining, punctures remaining distinct across the entire disc, Fig. 10 G). Male A. libanica can be separated from A. alshaykh and A. lindbergella due to the tergal punctation and genital capsule. The genital capsule has the outer margin of the gonostyli straight, without a kink in the inner or outer margins (in A. alshaykh with a noticeable kink in the inner and outer margins of the gonostyli; Fig. 20 L, M), and the terga are strongly and densely punctate, with punctures separated by <0.5 – 0.5 puncture diameters (in A. lindbergella with the tergal punctures slightly but noticeably sparser, separated by 0.5 – 1 puncture diameters; Fig. 20 A, B).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
68E0A0BA7C2F52E68C28CEEB8A946129.taxon	description	Description. Female. Body length: 6 – 7 mm. — Integumental colour: Body black. Flagellomeres 3 – 10 ventrally slightly lightened by presence of greyish scales. Legs black, apical tarsomeres slightly lightened orange-brown. Wings hyaline, stigma dark brown, venation dark orange (Fig. 10 A). Terga including marginal areas uniformly dark (Fig. 10 D). — Pubescence: Body hair relatively sparse. Face and gena with short whitish hairs, becoming light brown on vertex. Facial foveae filled with light brown hairs (Fig. 10 A – C). Dorsal mesosomal surfaces with short chestnut to dark brown hairs, these densely plumose, almost subsquamous (Fig. 10 A, C). Mesepisternum with longer white plumose hairs (Fig. 10 A). Propodeal corbicula incomplete, dorsoposterior fringe composed of moderately long plumose brownish hairs. Corbicular surface with abundant long light brown hairs. Leg hair whitish to light brown. Flocculus incomplete. Femoral and tibial scopae well-developed, femoral scopa composed of simple white hairs, tibial scopa composed of simple hairs, white ventrally, brown dorsally, tibial scopa thus bicoloured (Fig. 10 A). Tergal discs with very short pale hairs, visible only in profile. Marginal areas of terga 2 – 4 with apical hair fringes laterally, hairs emerging from base of marginal area and apical rim, thus apical fringes are formed from two rows of hairs; apical fringes widely interrupted, not forming complete bands. Terminal fringe dark brown (Fig. 10 A, D). — Head: 1.1 times broader than long (Fig. 10 B). Mandibles bidentate, moderately crossing apically. Labral process small, narrowly trapezoidal, slightly longer than wide to as long as wide. Clypeus weakly domed, smooth to polished in apical ¼ to ½, finely shagreened in basal ½, punctation distinct, punctures separated by 0.5 – 3 puncture diameters, densest basally (Fig. 10 B). Facial foveae narrow, occupying ¼ space between compound eye and lateral ocellus, uniformly narrow, equaling width of flagellum, ventrally extending to just below lower margin of antennal insertions, very slightly separated from inner margin of compound eye. Flagellomere 1 slightly exceeding 2 + 3, shorter than 2 + 3 + 4. Distance of fovea from lateral ocellus 2.2 ocellus diameters (Fig. 10 B, C). Ocelloccipital distance equals 0.6 ocellus diameter. Vertex distinctly angulate but not carinate (Fig. 10 C). — Mesosoma: Dorsolateral angle of pronotum rounded. Scutum and scutellum finely shagreened and shining, scutum densely and deeply punctate, punctures separated by 0.5 – 1 puncture diameters (Fig. 10 C). Mesepisternum finely reticulate. Propodeum with dense network of raised reticulation, propodeal triangle lacking lateral carinae, internally with dense network of strongly raised rugae with shining interspaces (Fig. 10 C). Hind pretarsal claw with small inner tooth. Recurrent vein 1 meeting submarginal cell 2 near its middle. Nervulus interstitial (Fig. 10 A). — Metasoma: Tergal discs finely shagreened, weakly shining, densely and deeply punctate, punctures separated by <0.5 – 0.5 puncture diameters. Punctures on disc of tergum 1 most coarse, almost confluent, becoming slightly finer on subsequent terga, punctures not extending onto marginal areas. Tergal margins slightly depressed (Fig. 10 D). Pygidial plate rounded, surface flat, with granular shagreen. — Male. Body length: 5 – 6 mm. — Integumental colour: As in female (Fig. 10 E, H). — Pubescence: Body hair longer and denser than in female, but more finely and less densely plumose, all pubescence white. Face, gena, and vertex with long white hairs, longest equaling length of scape (Fig. 10 E – G). Dorsal mesosomal surfaces with long erect white hair, long hairs present also on mesepisternum and propodeum (Fig. 10 E, G). Leg hair white (Fig. 10 E). Tergal discs with very short pale hairs, visible only in profile. Marginal areas of terga 2 – 4 with apical hair fringes laterally, hairs emerging from base of marginal area and apical rim, thus apical fringes are formed from two rows of hairs; apical fringes widely interrupted, not forming complete bands. Terminal fringe whitish-brown (Fig. 10 H). — Head: Clypeus structurally as in female, with clypeal shagreen uniformly weak, clypeus weakly shining over entire surface. Process of labrum very small, trapezoidal, slightly broader than long. Flagellomere 1 exceeding length of 2, shorter than 2 + 3; flagellomere 2 short, subsquare (Fig. 10 F). Ocelloccipital distance subequal to diameter of lateral ocellus (Fig. 10 G). — Mesosoma: As in female, with exception of slightly stronger scutal sculpture, becoming dull (Fig. 10 G). — Metasoma: As in female (Fig. 10 H). — Genitalia and hidden sterna: Dorsal gonocoxal lobes weakly but distinctly developed, forming short points. Gonostyli in dorsal view depressed below level of gonocoxites, dorsal surface subtly concave basally, apically becoming flattened and spatulate, inner margin slightly raised. Penis valves moderately broad basally, occupying ½ space between gonostyli, narrowing subapically to narrow point (Fig. 20 M). Sternum 8 simple, columnar, apex rounded to truncate, ventral surface covered with spreading fan of pale golden hairs.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
68E0A0BA7C2F52E68C28CEEB8A946129.taxon	distribution	Distribution. Andrena libanica is known from remnant cedar forest habitats in the central part of the Mount Lebanon mountain chain. It has been collected between 1726 – 1885 m above sea level, making it a high altitude species. Previously reported from Lebanon as A. rugulosa (Boustani et al. 2021)	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
4FAA824491BB59D49A012589392505E2.taxon	description	Figures 11, 15 L, 20 B, N	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
4FAA824491BB59D49A012589392505E2.taxon	diagnosis	Diagnosis of male. Within the Andrena minutula species group, A. lindbergella belongs to the species around A. rugulosa Stöckhert, characterized by a smooth clypeus and punctate terga, which include also A. alshaykh Pisanty sp. nov. and A. libanica Wood sp. nov. The male is easily differentiated from A. alshaykh by the genital capsule, in which the gonostyli are uniformly curving (suddenly converging medially, producing distinct kinks in the inner and outer margins in A. alshaykh), and the penis valves are narrower (Fig. 20 L, N). In addition, the scutum is more densely punctured, and the body size is larger. Both A. alshaykh and A. lindbergella are closely related to A. rugulosa Stöckhert (absent from the Levant) and A. libanica Wood sp. nov., but differ in the smoother scutum and more weakly punctured terga (Fig. 15 J – L). In addition, the male of A. rugulosa does not have dorsal gonocoxite lobes.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
4FAA824491BB59D49A012589392505E2.taxon	description	Description of male. Body length: 6.5 mm. — Integumental colour: Body and legs black. Flagellum fully black, overlayed with minute grey setae. Wings weakly infuscate, veins and stigma dark brown (Fig. 11 A). Tergal marginal zones black, occasionally brownish apically (Fig. 11 D). — Pubescence: Clypeus with dense and long semi-erect white hairs, underlying cuticle partly obscured. Paraocular area ventrally with dense and long, semi-erect to erect whitish hairs, dorsally with short to medium, mostly black hairs. Supraclypeal area with medium-lengthed erect hairs, mostly whitish, a few black. Scape with dense medium to long hairs, mostly white, a few black. Frons with sparse, semi-erect medium-lengthed black hairs (Fig. 11 A, B). Preoccipital ridge with short to long erect whitish hairs. Upper margin of compound eye with sparse short black hair. Genal area with dense whitish hair, medium-lengthed dorsally, long ventrally (Fig. 11 A – C). Scutum, scutellum and metanotum with long erect hair, mostly yellowish-whitish, sparse centrally, denser peripherally, scutellum with few short erect black hairs. Mesepisternum and propodeum with very long white hairs (Fig. 11 A, C). Leg hair mostly whitish, partly brownish on tarsi (Fig. 11 A). Base of tergal disc 1 and lateral parts of all tergal discs with sparse, short to medium white hair, rest of tergal discs with minute inconspicuous whitish hair. Tergal marginal zones 2 – 4 with distinct, lateral white hair bands, 5 with very weak and sparse, continuous whitish hair band. Terminal fringe brownish, flanked by whitish hairs (Fig. 11 A, D). — Head: 1.2 times broader than long (Fig. 11 B). Mandibles bidentate, moderately crossed. Galea very finely shagreened. Labral process trapezoidal, apical part slightly elevated, apical margin concave. Clypeus domed, very shiny, basally very superficially shagreened, apically smooth, punctation strong and dense, density decreasing medioapically, distance between punctures <0.5 – 1 puncture diameter, without impunctate midline. Flagellomere 1 1.3 times longer than 3, 1.8 times longer than 2 (Fig. 11 B). Frons rugose-areolate, with clearly discernible embedded fine punctures. Ocelloccipital distance 1 ocellus diameter. Preoccipital ridge weakly carinate (Fig. 11 C). — Mesosoma: Dorsolateral angle of pronotum not elevated. Scutum and scutellum fully shagreened, slightly shiny, punctation fine, strong and dense, distance between punctures 0.5 – 1 puncture diameters on scutum, slightly sparser on scutellum (Fig. 11 C). Mesepisternum finely alveolate, overlayed by shallow oblique punctures. Anteroventral corner of propodeum reticulated, rest of propodeum rugose-areolate, propodeal triangle basally slightly more coarsely sculptured compared to flanking areas, apically very finely areolated (Fig. 11 C). Recurrent vein 1 meets submarginal cell 2 near its middle. Nervulus slightly antefurcal (Fig. 11 A). — Metasoma: Tergal discs 1 – 3 shagreened, distinctly finely punctured, distance between punctures 1 puncture diameter. Tergal discs 4 – 6 with similar, weaker sculpturing. Tergal marginal zones shiny, very superficially shagreened, with occasional extremely fine, inconspicuous punctures (Figs 11 D, 20 B). — Genitalia and hidden sterna: Gonocoxites with small, partly rounded dorsal lobes. Gonostyli simple, finger-shaped, slightly broadening apically, blade flattened, apical margin rounded. Penis valves moderately broad basally, parallel-sided close to visible base, then tapering apically (Fig. 20 N). Sternum 8 columnar, apical process as broad as stem, apical margin rounded.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
4FAA824491BB59D49A012589392505E2.taxon	distribution	Distribution and habitat. Endemic to shrubland and sub-alpine habitats of the Troodos range in Cyprus, usually above 1200 m. Previous reports from the Levant (Pisanty et al. 2018; Wood et al. 2020) are reinterpreted as Andrena alshaykh sp. nov.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
AC6CE757081C591EA543B6373A4E4FD9.taxon	distribution	Distribution and habitat. Shrublands, warm grasslands, and semi-deserts in the southern Mediterranean and the Iberian Peninsula. In the Levant, limited to southern Israel and the West Bank * (Pisanty et al. 2018).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
4757F80CBD365AE698D96194B7835DD5.taxon	description	Figures 17 G, N, 20 S	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
4757F80CBD365AE698D96194B7835DD5.taxon	distribution	Distribution and habitat. Endemic to Mediterranean shrublands throughout the Levant (Israel, West Bank, Jordan, Lebanon *, Syria).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
2867203A181251C58B37FB596E92213B.taxon	description	Figures 15 H, 20 V	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
2867203A181251C58B37FB596E92213B.taxon	distribution	Distribution and habitat. High elevations throughout the northern Middle East. In the Levant, occurring in northern Israel, Lebanon and Syria (Pisanty et al. 2018; Wood & Monfared 2022; Wood et al. 2024 b).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
98BCA389FD1D5C7392E18D4DBDBD7D0B.taxon	description	Figures 15 G, 20 C, T	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
98BCA389FD1D5C7392E18D4DBDBD7D0B.taxon	distribution	Distribution and habitat. Grasslands and shrublands in south-eastern Europe, Turkey and the Levant (northern Israel, Jordan *, Lebanon, Syria *; Pisanty et al. 2018; Wood et al. 2020; IUCN 2024).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
92F5990B6EE756A49ECDB5F11961E34A.taxon	distribution	Distribution and habitat. South Mediterranean (Wood 2023 a). Newly recorded from the Levant * (Israel *, Jordan *). Previous accounts from the Levant (Warncke 1969) refer to Andrena kugleri sp. nov.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
12CA0E8091835128AB523714B1E37E0A.taxon	description	Figures 18 C, I, 21 F	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
12CA0E8091835128AB523714B1E37E0A.taxon	distribution	Distribution and habitat. Broadly West and Central Palaearctic, in diverse habitats. In the Levant, A single female was recorded from northern Lebanon, at 1600 m (Wood et al. 2020).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
A7DE1575AA7B5398A5C1CB97E7874578.taxon	description	Figures 14 E, I, P, 19 B, O	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
A7DE1575AA7B5398A5C1CB97E7874578.taxon	distribution	Distribution and habitat. Mediterranean shrublands in Greece and the northern Middle East, including all countries of the Levant (Wood et al. 2024 b).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
74F283FC8F7F5B13A9D171DBFBB321F3.taxon	description	Figures 16 F, K, 21 K	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
74F283FC8F7F5B13A9D171DBFBB321F3.taxon	distribution	Distribution and habitat. Mediterranean shrublands in Greece, Turkey and all countries of the Levant (Israel, West Bank, Jordan *, Lebanon *, Syria *; Pisanty et al. 2018; IUCN 2024).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
CC0AC95713165FE1A2BE2F33ADD9A7AD.taxon	description	Figures 14 B, K, 19 D, F	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
CC0AC95713165FE1A2BE2F33ADD9A7AD.taxon	distribution	Distribution and habitat. South Mediterranean. In the Levant, occurring in semi-arid to desert habitats in Israel, the West Bank and Jordan (Wood 2023 b).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
0E6F125654F6564F874AF7A5BAFE0AE4.taxon	description	Figures 12, 17 D, I, 20 P	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
0E6F125654F6564F874AF7A5BAFE0AE4.taxon	etymology	Etymology. Nominative feminine singular form of the Latin adjective phoenicius (= Phoenician).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
0E6F125654F6564F874AF7A5BAFE0AE4.taxon	diagnosis	Diagnosis. Within the Andrena minutula species group, the female of A. phoenicia is characterized by the combination of a labral process which is large and trapezoidal, a clypeus which is apically protuberant and strongly domed, strongly shagreened throughout and finely, sparsely punctured, a scutum which is shagreened, finely and densely punctured, and terga which are shagreened and impunctate. The male is characterized most clearly by the genitalia which possess large rounded dorsal gonocoxite lobes and gonostyli with a strong inward king near the apex. The species is most closely related to A. stolida Warncke and A. simontornyella Noskiewicz, which share similar genitalia. The female differs in the larger labral process (Figs 17 D, 18 D), the clypeus which is transversely arched and apically protuberant (domed and not apically protuberant in A. stolida and A. simontornyella), and the moderately dull scutum (centrally shiny in A. stolida, completely dull and more weakly punctured in A. simontornyella; Figs 12 C, 17 I, 18 J). The male differs in the larger dorsal gonocoxite lobes which, when viewed together, are broader than the width of the visible base of the penis valves (slightly narrower than this width in the comparison species; Fig. 20 P, Q).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
0E6F125654F6564F874AF7A5BAFE0AE4.taxon	description	Description. Female. Body length: 6 – 6.5 mm. — Integumental colour: Body and legs black. Flagellum dark, distal flagellomeres often with weak reddish hue anteriorly. Apical tarsomeres brown. Wings weakly infuscate, veins and stigma brown (Fig. 12 A). Tergal marginal zones black almost to apex (Fig. 12 D). — Pubescence: Body hair relatively sparse, mostly short to medium-lengthed, brightly coloured. Clypeus with sparse, short and thin, whitish plumose hairs. Paraocular area and frons with moderately dense, medium-lengthed plumose whitish hairs (Fig. 12 A, B). Facial foveae with dense, minute whitish to brownish hairs, colour changing with angle of view (Fig. 12 B, C). Vertex with sparse, erect brownish hairs of varying length. Genal area with sparse short whitish hair (Fig. 12 A – C). Discs of scutum and scutellum with moderately dense, inconspicuous thin brownish-golden hairs, the majority very short, with few medium-lengthed scattered in between. Periphery of scutum and scutellum with narrow band of short and thick, golden plumose hairs (Fig. 12 A, C). Mesepisternum with moderately dense, long whitish plumose hairs. Propodeal corbicula incomplete, dorsoposterior fringe composed of long whitish plumose hairs, corbicular surface with few, long whitish simple hairs. Leg hair whitish to brownish. Flocculus incomplete, white. Femoral and tibial scopae well-developed, whitish, tibial scopal hairs simple (Fig. 12 A). Metasomal surface almost hairless, tergal discs centrally with sparse, minute inconspicuous whitish hairs, laterally with few short whitish hairs. Tergal marginal zones with few medium-lengthed white hairs basolaterally, tergal hair bands absent. Terminal fringe golden to light brown centrally, laterally flanked by few white hairs (Fig. 12 A, D). — Head: 1.15 times broader than long. Mandibles bidentate, moderately crossed. Galea finely shagreened. Labral process broad and large, shiny, trapezoidal. Clypeus transversely arched, fully shagreened and matt, non-rugose or at most with hint of very weak, fine transverse striation on basal half, punctation fine and shallow, distance between punctures 1 – 2 puncture diameters, an impunctate midline is weakly indicated on basal ¾ (Figs 12 B, 17 D). Malar area undeveloped. Supraclypeal area longitudinally striated. Paraocular area strongly longitudinally striated, becoming punctate near apex. Flagellomere 1 slightly longer than 2 + 3, 3 slightly longer than 2 (Fig. 12 B). Facial fovea about half as broad as antennocular distance above, weakly tapering below, extending from level of lower end of lateral ocellus to base of clypeus or slightly above, separated from compound eye by narrow strip of smooth cuticle. Distance of fovea from lateral ocellus 1.7 ocellus diameters. Frons coarsely, obliquely striated to rugose-areolate below, more finely and longitudinally so above, medial carina almost reaching frontal ocellus (Fig. 12 B, C). Vertex with shallow but distinct groove behind lateral ocelli, dorsal preoccipital ridge moderately carinate. Ocelloccipital distance equals 1 ocellus diameter (Fig. 12 C). — Mesosoma: Dorsolateral angle of pronotum not elevated. Scutum and scutellum entirely finely shagreened, finely punctured, distance between punctures 0.5 – 1.5 puncture diameters (Fig. 12 C, 15 F). Mesepisternum anteriorly with oblique punctures often resembling star-shaped wrinkles, posteriorly shagreened. Propodeal corbicula finely reticulate, with few large crater-shaped punctures around hair bases. Posterior part of propodeum rugose-areolate, propodeal triangle weakly delineated, with areolation gradually finer apically (Fig. 12 C). Hind pretarsal claw with small inner tooth. Recurrent vein 1 reaching submarginal cell 2 at its middle or slightly proximal to it. Nervulus weakly to distinctly antefurcal (Fig. 12 A). — Metasoma: Tergal discs fully shagreened, impunctate. Tergal marginal zones 2 – 4 arched, 2 fully shagreened, hardly depressed, the following gradually smoother and more strongly depressed (Fig. 12 D). Pygidial plate simple, without elevated central zone. — Male. Body length: 5.5 – 6.5 mm. — Integumental colour: As in female, but flagellum entirely black to dark brown, without reddish hue (Fig. 12 E, H). — Pubescence: Body hair mostly white. Clypeus with dense long plumose hairs, partly obscuring underlying cuticle. Rest of face and scape with moderately dense white hair of varying length (Fig. 12 E, F). Vertex and gena with whitish erect hair, fully white and very long ventrally (Fig. 12 E – G). Periphery of scutum and scutellum with moderately dense whitish hair, disc of scutum with minute, inconspicuous erect whitish hair. Mesepisternum and propodeum with very long, white plumose hair (Fig. 12 E, G). Leg hair white (Fig. 12 E). Tergal discs centrally with minute inconspicuous hair, laterally with short whitish hair. Tergal marginal zones with white hairs basolaterally, forming very weak lateral hair bands. Terminal fringe white (Fig. 12 E, H). — Head: 1.2 times broader than long (Fig. 12 F). Mandibles bidentate, moderately crossed. Galea finely shagreened. Labral process rectangular, smooth. Clypeus arched, densely but shallowly punctured, distance between punctures 0 – 1 puncture diameters, without impunctate midline, clypeal surface roughened and slightly rugose by elevated puncture margins. Malar area undeveloped. Supraclypeal area longitudinally striated. Paraocular area densely, finely punctate near compound eye, strongly longitudinally striated centrally. Flagellomere 1 about as long as 2 + 3, 3 longer than 2 (Fig. 12 F). Frons coarsely, obliquely striated to rugose-areolate below, finely areolate above. Vertex with shallow but distinct groove behind lateral ocelli, dorsal preoccipital ridge carinate. Ocelloccipital distance equals 1 ocellus diameter (Fig. 12 G). — Mesosoma: Scutum dull, fully, granularly shagreened, densely and finely punctured, distance between punctures 0 – 1 puncture diameters, some punctures with raised margins (crater-like). Scutellum similar but without crater-like punctures, surface occasionally smoother (Fig. 12 G). Hind pretarsal claw bifurcated. Nervulus interstitial to antefurcal. Rest of mesosoma as in female. — Metasoma: Tergal discs 1 – 2 fully shagreened, essentially impunctate, 3 and especially 4 often smoother and more weakly shagreened, with hint of very fine punctation. Tergal marginal zones 2 – 4 weakly arched, moderately depressed, impunctate, 2 more or less fully shagreened, the following gradually smoother (Fig. 12 H). — Genitalia and hidden sterna: Dorsal gonocoxite lobes well-developed, broad, mostly rounded but with weak apicolateral points. Gonostyli thick, outer margins with strong inward curve on distal half, apexes rounded. Penis valves broad basally, basal 2 / 3 tapering apically (Fig. 20 P). Sternum 8 simple, columnar, apical process broadened, apical margin weakly but distinctly emarginate, resembling a fishtail.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
0E6F125654F6564F874AF7A5BAFE0AE4.taxon	distribution	Distribution and habitat. Mesic habitats in Israel and Syria, likely also Lebanon.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
2B3EF7C362D85D8C83DFFCB31D53FDB6.taxon	description	Figures 14 C, H, M, 19 A, M	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
2B3EF7C362D85D8C83DFFCB31D53FDB6.taxon	distribution	Distribution and habitat. Broadly across the Western Palaearctic, but excluding the most Atlantic parts, mostly in mesic habitats (McLaughlin et al. 2022). In the Levant *, limited to mountains in Lebanon *.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
B6635A764E75562094BEC6094E44AC34.taxon	description	Figures 16 C, I, 20 R	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
B6635A764E75562094BEC6094E44AC34.taxon	distribution	Distribution and habitat. Shrubland habitats in the north-east Mediterranean. In the Levant, limited to northern Israel, Lebanon and Syria * (Pisanty et al. 2018; Wood et al. 2020; IUCN 2024).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
DCD06FC96BC559499501B4C86BA9BCC9.taxon	description	Figures 18 E, K, 21 D	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
DCD06FC96BC559499501B4C86BA9BCC9.taxon	distribution	Distribution and habitat. As currently understood in a narrow sense (not considering ssp. histrionica), limited to shrublands and montane habitats in all countries of the Levant, as well as south-western Turkey and the island of Rhodes (Wood 2024).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
479DF3FDF7885EC9862AA249F7F8E95D.taxon	description	Figures 17 K, M, 20 J	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
479DF3FDF7885EC9862AA249F7F8E95D.taxon	distribution	Distribution and habitat. Circum-Mediterranean (excluding the Balkan Peninsula), including all the Levant and Cyprus, in coastal, shrubland and semi-desert habitats.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
AA48EB953F8D59EC849AD7BDE1BB5E5F.taxon	description	Figures 16 E, 18 D, J, 20 Q	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
AA48EB953F8D59EC849AD7BDE1BB5E5F.taxon	distribution	Distribution and habitat. Mediterranean shrublands in Turkey and the Levant * (Israel *, Jordan *, Lebanon *, Syria *).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
3ADBF16A035C5726BBA6788C4904E9A4.taxon	description	Figures 17 A, 20 G	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
3ADBF16A035C5726BBA6788C4904E9A4.taxon	distribution	Distribution and habitat. Endemic to desert habitats in central Syria.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
31B2C76A396754378DB33A1909DA3570.taxon	description	Figure 21 G	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
31B2C76A396754378DB33A1909DA3570.taxon	distribution	Distribution and habitat. South Mediterranean. In the Levant, limited to sandy and hamra soils along Israel’s coastal plain. Reports from Lebanon, Syria and Iran (Warncke 1974 b; Grace 2010; Kratochwil 2015; Wood et al. 2020; Boustani et al. 2021) are refuted (see below).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
5682FA46B291514FA484AD008B12E128.taxon	description	Figures 17 C, H, L, 20 F, W	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
5682FA46B291514FA484AD008B12E128.taxon	distribution	Distribution and habitat. Shrubland and semi-desert habitats in the northern Middle East, including all countries of the Levant (Israel, West Bank, Jordan, Lebanon *, Syria).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
D2A9833E9F5B5ADD9AE2549D955A5864.taxon	description	Figures 15 A, D, 21 H	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
D2A9833E9F5B5ADD9AE2549D955A5864.taxon	distribution	Distribution and habitat. Eastern Europe, Turkey and the Levant (northern Israel and Lebanon; Pisanty et al. 2018; Wood et al. 2020). In Europe associated with steppe habitats (IUCN 2024); in the Levant, limited to mesic habitats mostly at high elevations.	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
F520E58C900A5A4F9DCCCCD25A302241.taxon	description	Figures 13 B, 14 A, G, N, 20 D	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
F520E58C900A5A4F9DCCCCD25A302241.taxon	distribution	Distribution and habitat. Northern Middle East. Records from Central Asia are refuted – these originate from a misinterpretation by Osytschnjuk et al. (2008), who describe the male as having a yellow clypeus. In the Levant, present in Mediterranean shrubland, semi-desert and desert habitats in Israel, the West Bank, Jordan and Syria (Wood & Monfared 2022).	en	Pisanty, Gideon, Santerre, Rémi, Martin, Teresa, Cardinal, Sophie, Wood, Thomas J. (2025): Integrated taxonomic revision of the mining bee subgenus Andrena (Micrandrena) (Hymenoptera: Andrenidae) in the Levant and Cyprus. Arthropod Systematics & Phylogeny 83: 573-655, DOI: 10.3897/asp.83.e166986
