taxonID	type	description	language	source
592EED1CFFE0FFA04D691FB4FB84A57F.taxon	materials_examined	Type Material. Holotype (Male): “ Pine forest 7 mi S., Mazamitla, Jalisco, MEXICO. 1 ­ XII­ 48 ” / “ H. B. Leech, Collection ” / “ HOLOTYPE, Enochrus (H.), plicatus, A. E. Z. Short. ” (USNM). Type Locality. Mazamitla, Jalisco State, Mexico.	en	Short, A. E. Z. (2005): Two new species of Enochrus Thomson, subgenus Hugoscottia Knisch, from Costa Rica and Mexico (Coleoptera: Hydrophilidae). Zootaxa 865: 1-7, DOI: 10.5281/zenodo.170859
592EED1CFFE0FFA04D691FB4FB84A57F.taxon	diagnosis	Diagnosis. Segments 2 and 3 of maxillary palpi bulbous and grooved; the punctation on pronotum and elytra relatively fine, the distinctly 4 ­ ridged mesosternal process, the hooked and acutely pointed parameres and non­emarginate anterior margin of the clypeus serve to distinguish this species from other species of Hugoscottia.	en	Short, A. E. Z. (2005): Two new species of Enochrus Thomson, subgenus Hugoscottia Knisch, from Costa Rica and Mexico (Coleoptera: Hydrophilidae). Zootaxa 865: 1-7, DOI: 10.5281/zenodo.170859
592EED1CFFE0FFA04D691FB4FB84A57F.taxon	description	Description. Total length 3.6 mm. Color. Elytra and pronotum yellow to light brown, however this specimen is slightly teneral. Maxillary palpi uniformly yellow. Clypeus darkened on central two­thirds. The remaining structures are generally pale. Punctation. General punctation of elytra moderately fine and distinct, distance between punctures 1.5 – 2.0 X the width of a puncture. Head and pronotal punctation slightly deeper and more distinct. Systematic punctures (sensu Hansen 1991) on elytra and pronotum slightly larger and more impressed than surrounding punctation, each puncture usually bearing a fine short seta. Head. Three segmented antennal club with apical segment about 1.5 X as long as penultimate segment. Maxillary palpi short, shorter than width of head anterior of the eyes; second segment as long as width of the mentum; segments 3 & 4 subequal in length; segments 2 & 3 bulbous; dorsal surface of segment 2 and ventral surface of segment 3 wide and flat and slightly curved, with the ability to fit against each other when the joint between them is folded. Mentum with moderately coarse, evenly distributed punctation, each puncture about as wide as deep; slightly depressed on anterior margin. Anterior margin of the clypeus entire, without emargination. Thorax. Prosternum with anterior third slightly depressed, with distinct median carina; posterior two­thirds sparsely pubescent and tectiform but not carinate. Mesosternal process consisting of a longitudinal, obtusely triangular crest with lateral carina on each side; the resulting four ridges meeting near the apex of the process; sides of carinae glabrous and with fine granular microsculpture. Mesosternum with elongate­oval glabrous area posteromedianly; glabrous area slightly more than half the total length of mesosternum. Hind femora pubescent on basal fourfifths. Elytra with a sutural stria in posterior two­thirds to three­fourths. Abdomen. Ventrites uniformly and densely pubescent. Fifth ventrite with shallow posteromedian emargination, distinctly wider than deep and lined with coarse yellow setae. Aedeagus (Fig. 1). Parameres slightly sinuate along outer margin, strongly sinuate along interior margin, with tips curved hooked inward, with an acute apex. Tip of median lobe not reaching apex of parameres; faintly bifid on dorsal side. Basal piece damaged but appearing constricted basally.	en	Short, A. E. Z. (2005): Two new species of Enochrus Thomson, subgenus Hugoscottia Knisch, from Costa Rica and Mexico (Coleoptera: Hydrophilidae). Zootaxa 865: 1-7, DOI: 10.5281/zenodo.170859
592EED1CFFE0FFA04D691FB4FB84A57F.taxon	distribution	Distribution. Jalisco State, Mexico.	en	Short, A. E. Z. (2005): Two new species of Enochrus Thomson, subgenus Hugoscottia Knisch, from Costa Rica and Mexico (Coleoptera: Hydrophilidae). Zootaxa 865: 1-7, DOI: 10.5281/zenodo.170859
592EED1CFFE0FFA04D691FB4FB84A57F.taxon	etymology	Etymology. Derived from the Latin plicatus, after the folding modifications of the maxillary palpi. Used here as a noun in apposition.	en	Short, A. E. Z. (2005): Two new species of Enochrus Thomson, subgenus Hugoscottia Knisch, from Costa Rica and Mexico (Coleoptera: Hydrophilidae). Zootaxa 865: 1-7, DOI: 10.5281/zenodo.170859
592EED1CFFE0FFA04D691FB4FB84A57F.taxon	discussion	Remarks. Other species of Enochrus (e. g. E. (Methydrus) punctipalpus) are known to have sexually dimorphic maxillary palpi in which only those of the male are modified. As only a single male is known of E. plicatus, it not yet known if this is true in this case. In his unpublished Ph. D. thesis, Gundersen (1967) records Enochrus (Enochrus) oviformis d’Orchymont, a Brazilian species, from Mexico. One of the four specimens he examined has the exact label data as the holotype of E. (H.) plicatus, and Gundersen’s illustration of the aedeagus and maxillary palpus leave little doubt that it is the same specimen, although it bears no label to demonstrate such a fact. There is no mention by Gundersen of the distinct mesosternal process that would unambiguously place this species in Hugoscottia. I here treat Gundersen’s (1967) Mexican records of E. oviformus as a misidentification of E. (H.) plicatu s. Gundersen’s subsequent 1977 and 1978 publications do not treat E. oviformis. The localities for the other specimens of ‘ E. oviformis ’ in Gundersen’s thesis are “ 15 – 20 mi. W. Jiquilpan ” in Jalisco State and “ Rio Metloc, nr. El Fortin ” in Veracruz State. I have not been able to locate or verify these other specimens are conspecific with E. (H.) plicatus, in part because no list of depositories was provided; they do not appear to be in the USNM collection.	en	Short, A. E. Z. (2005): Two new species of Enochrus Thomson, subgenus Hugoscottia Knisch, from Costa Rica and Mexico (Coleoptera: Hydrophilidae). Zootaxa 865: 1-7, DOI: 10.5281/zenodo.170859
592EED1CFFE2FFA74D691F7EF8D6A7FF.taxon	materials_examined	Type Material. Holotype (Male): “ COSTA RICA: Cartago Pr. / Tapanti National Park / pools along entrance road / A. E. Z. Short; 22 ­ vi­ 2003 ”, “ HOLOTYPE / Enochrus (H.) / talamanca / A. E. Z. Short. ” (INBio). Paratypes (35): COSTA RICA: Cartago Prov.: same data as holotype (AEZS 5, BMNH 1, ESMC 1, MCZ 1, NMW 1, USNM 1, INBio 10), Tapantí National Park: Quebrada Segunda, 1250 m elev., iii. 1992, G. Mora (AEZS 1, CUIC 1, INBio 8), same locality, viii. 1992 (INBio 1); Tapantí National Park, 1650 m elev., 18. ii. 1998, R. Anderson leg., “ cloud forest (dry) litter ” (CMNC 1, INBio 1, FSCA 1, AEZS 1). Type Locality. Tapantí National Park, Cartago Province, Costa Rica.	en	Short, A. E. Z. (2005): Two new species of Enochrus Thomson, subgenus Hugoscottia Knisch, from Costa Rica and Mexico (Coleoptera: Hydrophilidae). Zootaxa 865: 1-7, DOI: 10.5281/zenodo.170859
592EED1CFFE2FFA74D691F7EF8D6A7FF.taxon	diagnosis	Diagnosis. The dark brown coloration, generally coarse punctation, shape of the parameres (Fig. 2), and more subtle transverse ridges of the mesosternal process serve to distinguish this species from other species of Hugoscottia.	en	Short, A. E. Z. (2005): Two new species of Enochrus Thomson, subgenus Hugoscottia Knisch, from Costa Rica and Mexico (Coleoptera: Hydrophilidae). Zootaxa 865: 1-7, DOI: 10.5281/zenodo.170859
592EED1CFFE2FFA74D691F7EF8D6A7FF.taxon	description	Description. Total Length 3.0 – 3.5 mm. Color. Elytra and pronotum dark brown, slightly paler along lateral margins. Vertex and central third of clypeus dark brown; lateral margins of clypeus slightly paler. Maxillary palpi uniformly yellow. Venter brown to dark brown, with mesosternum and epipleurae paler. Femora brown to dark brown, paler distally. Punctation. General punctation on elytra moderately coarse, distance between punctures 2 – 3 X the width of a puncture, slightly denser on elytral disc. Head and pronotal punctation slightly denser, 1 – 2 X width of a puncture. Head. Three segmented antennal club with apical segment about 1.5 X as long as penultimate segment. Maxillary palpi short, shorter than width of head anterior to the eyes; second segment as long as width of the mentum; segments 3 & 4 subequal in length; segment 2 slightly thickened, especially in apical three fourths, but not bulbous. Mentum with moderately coarse, evenly distributed punctation, each puncture about as wide as deep; slightly depressed on anterior margin. Clypeo­labral suture nearly straight, the anterior margin of the clypeus slightly emarginate centrally. Thorax. Prosternum with anterior third slightly depressed, with distinct median carina; posterior two­thirds tectiform, sparsely pubescent. Elytra with sutural stria present in posterior two­thirds to three­fourths. Mesosternum with a longitudinal, elongate triangular crest bearing a transverse ridge on each side that reach the edge of the carina slightly posterior to the apex; sides of the carina glabrous, with fine granular microsculpture. Mesosternum with posteromedian elongate oval glabrous area about half the total length of the mesosternum. Hind femora pubescent on basal four­fifths. Protarsal claws of both sexes simple and unmodified. Abdomen. Ventrites uniformly and densely pubescent. Fifth ventrite with posteromedial emargination, distinctly wider than deep and lined with coarse yellow setae. Aedeagus (Fig. 2). Parameres sinuate, with tips moderately curved mediad and roundly hooked. Tip of median lobe almost reaching apex of parameres; appearing bifid on dorsal side. Basal piece narrowed and rounded in basal third.	en	Short, A. E. Z. (2005): Two new species of Enochrus Thomson, subgenus Hugoscottia Knisch, from Costa Rica and Mexico (Coleoptera: Hydrophilidae). Zootaxa 865: 1-7, DOI: 10.5281/zenodo.170859
592EED1CFFE2FFA74D691F7EF8D6A7FF.taxon	distribution	Distribution. Known only from Tapantí National Park, Costa Rica.	en	Short, A. E. Z. (2005): Two new species of Enochrus Thomson, subgenus Hugoscottia Knisch, from Costa Rica and Mexico (Coleoptera: Hydrophilidae). Zootaxa 865: 1-7, DOI: 10.5281/zenodo.170859
592EED1CFFE2FFA74D691F7EF8D6A7FF.taxon	etymology	Etymology. Named after the Cordillera de Talamanca, the mountain range in which the known specimens were found.	en	Short, A. E. Z. (2005): Two new species of Enochrus Thomson, subgenus Hugoscottia Knisch, from Costa Rica and Mexico (Coleoptera: Hydrophilidae). Zootaxa 865: 1-7, DOI: 10.5281/zenodo.170859
592EED1CFFE2FFA74D691F7EF8D6A7FF.taxon	discussion	Remarks. This species was collected from very shallow pools and mud puddles along the sides of a gravel road in Tapantí National park (Figs. 3 & 4). The beetles appeared to be living in and around clumps of emergent grasses and detritus. On one occasion, specimens were clinging to the submerged side of a dead Cecropia sp. leaf. The species has also been sifted from “ dry ” forest litter. This species was rare in collections, but after the microhabitat was discovered I was able to collect it rather easily. However, despite knowing the microhabitat preference, and sampling that microhabitat in many other areas, the species is still only known from one national park. Additional specimens collected in Tapantí National Park on 9 January 2004 but not listed as type material were preserved in 100 % ethanol and frozen for future molecular work by the author.	en	Short, A. E. Z. (2005): Two new species of Enochrus Thomson, subgenus Hugoscottia Knisch, from Costa Rica and Mexico (Coleoptera: Hydrophilidae). Zootaxa 865: 1-7, DOI: 10.5281/zenodo.170859
