identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
596FA515FFA0FFBA6D96C29CFDA4FC07.text	596FA515FFA0FFBA6D96C29CFDA4FC07.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cleroidea Latreille 1802	<div><p>Superfamily Cleroidea Latreille, 1802</p><p>In light of the different views on the systematics of groups within Cleroidea (see e.g. Lawrence &amp; Leschen 2010) and with aim of explanation of the systematic position of new tribe, the key for identification of Melyrid lineage families combined with the identification key for subfamilies and tribes within Melyridae (according to Majer 1987 with modifications) is provided below.</p></div>	https://treatment.plazi.org/id/596FA515FFA0FFBA6D96C29CFDA4FC07	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tshernyshev, Sergei E.;Alekseev, Vitalii;Bukejs, Andris;Vasilenko, Dmitry V.;Legalov, Andrej A.;Perkovsky, Evgeny E.	Tshernyshev, Sergei E., Alekseev, Vitalii, Bukejs, Andris, Vasilenko, Dmitry V., Legalov, Andrej A., Perkovsky, Evgeny E. (2025): First record of the family Melyridae (Coleoptera: Cleroidea) in late Eocene European ambers with description of a new tribe, a new genus and seven new species. Zootaxa 5696 (4): 542-566, DOI: 10.11646/zootaxa.5696.4.5, URL: https://doi.org/10.11646/zootaxa.5696.4.5
596FA515FFA0FFBB6D96C6BEFA71FE7B.text	596FA515FFA0FFBB6D96C6BEFA71FE7B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Melyridae Leach 1815	<div><p>Key to subfamilies and tribes of Melyridae</p><p>1. Surface of body covered with double pubescence, short adpressed and long semi-erect or erect hairs. Ventral sides of tarsomeres with short and sharp setae (Astylinae Majer, 1987)........................................................... 2</p><p>- Surface of body covered with adpressed or semi-erect pubescence, lacking erect hairs. Tarsomeres lacking sharp setae, bare or pubescent ventrally ( Melyrinae Leach, 1815)............................................................... 3</p><p>2. Body short, parallel and suboval, slightly convex, penultimate antennomeres transverse. Legs short and strong.......................................................................................... Arthrobrachini Majer, 1987</p><p>- Body elongate, evenly widened posteriorly, slightly flattened, penultimate antennomeres longitudinal or equilateral, never transverse. Legs long and slender............................................................ Astylini Pic, 1929</p><p>3. Elytra evenly covered with distinct adpressed hairs, lacking carinae. Tarsomeres pubescent ventrally..... Cerallini Pic, 1929</p><p>- Elytra covered with extremely short semi-erect dense pubescence, canaliculate. Tarsomeres bare or with strong setae on ventral side................................................................................................ 4</p><p>4. Penultimate antennomeres distinctly triangular with sharp distal edges. Elytral punctation irregular. Lateral sides of pronotum with singular or double thin margination, not vertical. Surface of pronotum with small irregular punctation. Tarsomeres bare or with sparse short semi-erect hairs in periphery of tarsomere............................. Melyrini Kiesenwetter, 1863</p><p>- Penultimate antennomeres oval-triangular with rounded edges. Elytral with three regular rows of punctures between longitudinal carinae. Pronotum with vertical lateral sides bordered with strong carinae. Surface of pronotum with strongly pronounced foveolate punctation. Tarsomeres with strong semi-erect dark setae on ventral side................. Amelyrini trib. nov.</p></div>	https://treatment.plazi.org/id/596FA515FFA0FFBB6D96C6BEFA71FE7B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tshernyshev, Sergei E.;Alekseev, Vitalii;Bukejs, Andris;Vasilenko, Dmitry V.;Legalov, Andrej A.;Perkovsky, Evgeny E.	Tshernyshev, Sergei E., Alekseev, Vitalii, Bukejs, Andris, Vasilenko, Dmitry V., Legalov, Andrej A., Perkovsky, Evgeny E. (2025): First record of the family Melyridae (Coleoptera: Cleroidea) in late Eocene European ambers with description of a new tribe, a new genus and seven new species. Zootaxa 5696 (4): 542-566, DOI: 10.11646/zootaxa.5696.4.5, URL: https://doi.org/10.11646/zootaxa.5696.4.5
596FA515FFA1FFBB6D96C21BFE01FAD1.text	596FA515FFA1FFBB6D96C21BFE01FAD1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amelyrini Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov	<div><p>Tribe Amelyrini Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov &amp; Perkovsky, trib. nov.</p><p>urn:lsid:zoobank.org:act: 152FEE2D-4A73-4D44-AC9A-0A246AC84986</p><p>Type genus: Amelyris Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov &amp; Perkovsky, gen. nov., by present designation</p><p>Diagnosis. Small beetles, 2.5–3.8 mm in length, characterized by the following combination of features: (1) head prognathous, small, rounded; (2) pronotum with lateral carinae; (3) elytra with longitudinal carinae and three regular rows of punctures between them, whereas humeri exhibiting an additional fine carina; (4) abdominal ventrites 1 and 2 connate; (5) antennal club friable (symmetrical or asymmetrical), composed of finely articulated segments; and (6) tarsi with tarsomere 1 distinctly reduced.</p><p>Differential diagnosis. Representatives of Amelyrini trib. nov. are superficially similar to the species of Melyris or Falsomelyris ( Melyrini), but can be distinguished in smaller body size (2.5–3.8 mm vs. 7–9 mm in Melyrini), in prognathous convex head (in contrast to orthognathous non-convex head in Melyrini), in pronotum with lateral carinae (vs. lateral margins lacking carinae or bearing a single longitudinal carina near the margin, e.g., in Melyris rufomarginatus), in elytral intercarinal spaces with three regular puncture rows (vs. irregular 3–4 rows in Melyrini), in abdominal ventrites 1 and 2 connate (in contrast to freely articulated in Melyrini), and in humeri with a distinct carina (vs. absent in Melyrini).</p><p>Etymology. The tribe Amelyrini trib. nov. is named after the nominative genus Amelyris gen. nov., whose etymology is presented below.</p></div>	https://treatment.plazi.org/id/596FA515FFA1FFBB6D96C21BFE01FAD1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tshernyshev, Sergei E.;Alekseev, Vitalii;Bukejs, Andris;Vasilenko, Dmitry V.;Legalov, Andrej A.;Perkovsky, Evgeny E.	Tshernyshev, Sergei E., Alekseev, Vitalii, Bukejs, Andris, Vasilenko, Dmitry V., Legalov, Andrej A., Perkovsky, Evgeny E. (2025): First record of the family Melyridae (Coleoptera: Cleroidea) in late Eocene European ambers with description of a new tribe, a new genus and seven new species. Zootaxa 5696 (4): 542-566, DOI: 10.11646/zootaxa.5696.4.5, URL: https://doi.org/10.11646/zootaxa.5696.4.5
596FA515FFA1FFBC6D96C502FC0CFDAF.text	596FA515FFA1FFBC6D96C502FC0CFDAF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amelyris Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov	<div><p>Genus Amelyris Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov &amp; Perkovsky, gen. nov.</p><p>urn:lsid:zoobank.org:act: B034B11D-84DC-4E76-9277-1281C2256731</p><p>Type species: Amelyris (Amelyris) sveneldi Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov &amp; Perkovsky, sp. nov., by present designation</p><p>Description. Body moderately small (2.5–3.8 mm long), elongate, slender, parallel, and flattened dorsally. Head small, round, slightly convex, as wide as pronotum, not sculptured, finely punctured, covered with fine recumbent hairs. Antennae 11-segmented, attached near lower edge of eye and near clypeus; terminal antennomeres forming loose club, symmetrical in Amelyris subgen. nov. and asymmetrical in Asymelyris subgen. nov.</p><p>Pronotum convex, elongate, slightly narrowed anteriorly, with acute angles and lateral side with typical doubled carinae; surface uniformly punctured with large punctures, microsculpture distinct; pubescence small, curved and sparse; disc clearly impressed at posterior and anterior angles.</p><p>Elytra distinctly carinate, elongate, subparallel, slightly convex just below middle, evenly narrowed and rounded posteriorly; apices remaining separate near suture; humeri small, distinct, with small, short, curved carina; epipleura broad and distinct along elytra from base to apex; each elytron with four carinae; interspace between carinae with three regular rows of sparse distinct punctures and developed microsculpture. Elytral pubescence short, semi-erect, situated mainly on carinae. Suture distinctly marginate, with even margin in Amelyris (Amelyris) subgen. nov. and thickened apically in Amelyris (Asymelyris) subgen. nov. Metathoracic wings well developed.</p><p>Legs short, simple. Femora of all legs flattened and widened, not curved. Tibiae straight, evenly flattened, with short, black, strong, erect setae. Tarsi 5-segmented. Tarsomeres strongly compressed, with short, black, erect hairs on the periphery of the pedal part; tarsomere 1 about 1.5–2.5× shorter than tarsomere 2.</p><p>Ventral side moderately convex, weakly shiny, finely and densely punctate, bare or with fine dense adpressed hairs. Pygidium (apical tergite) and apical sternite simple, narrow, with tufts of several long, strong black bristles on outer margins of lateral sides in nominative subgenus of Amelyris, Amelyris subgen. nov., and lacking bristles in the subgenus Asymelyris subgen. nov.</p><p>Etymology. The new genus-group name is composed of the Greek prefix ἀ- / ἀν-, i.e. ‘not’ (the ‘alpha privative’) and Melyris, the nominative genus name for the family Melyridae . Gender masculine.</p><p>Composition. The new genus, Amelyris gen. nov., is divided into two subgenera ( Amelyris subgen. nov. and Asymelyris subgen. nov.) and includes seven species known from inclusions in late Eocene European ambers: five species from Baltic amber, one species from Rovno amber and one species from Danish amber. Identification key for all species of the genus is provided after subgeneric descriptions below.</p></div>	https://treatment.plazi.org/id/596FA515FFA1FFBC6D96C502FC0CFDAF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tshernyshev, Sergei E.;Alekseev, Vitalii;Bukejs, Andris;Vasilenko, Dmitry V.;Legalov, Andrej A.;Perkovsky, Evgeny E.	Tshernyshev, Sergei E., Alekseev, Vitalii, Bukejs, Andris, Vasilenko, Dmitry V., Legalov, Andrej A., Perkovsky, Evgeny E. (2025): First record of the family Melyridae (Coleoptera: Cleroidea) in late Eocene European ambers with description of a new tribe, a new genus and seven new species. Zootaxa 5696 (4): 542-566, DOI: 10.11646/zootaxa.5696.4.5, URL: https://doi.org/10.11646/zootaxa.5696.4.5
596FA515FFA6FFBC6D96C2D4FE94FBB3.text	596FA515FFA6FFBC6D96C2D4FE94FBB3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	(Amelyris) Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov	<div><p>Subgenus Amelyris Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov &amp; Perkovsky, subgen. nov.</p><p>urn:lsid:zoobank.org:act: CB7BEFAD-2CA2-41BB-8721-0AF614825926</p><p>Type species: Amelyris (Amelyris) sveneldi Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov &amp; Perkovsky, sp. nov., by present designation</p><p>Diagnosis. Antennal club symmetrical; sutural margin of equal width along entire elytron; third elytral carina from suture without tendency to bend or merge with second carina at elytral apex; pygidium (apical tergite); and last visible abdominal ventrite (apical sternite) with tufts of several long, strong black bristles on outer corners of margins.</p><p>Etymology. As for the genus.</p><p>Composition. The subgenus includes three extinct Eocene species: Amelyris (Amelyris) sveneldi sp. nov. (Rovno amber), Amelyris (Amelyris) elongatus sp. nov. (Baltic amber), and Amelyris (Amelyris) tricarinatus sp. nov. (Baltic amber).</p></div>	https://treatment.plazi.org/id/596FA515FFA6FFBC6D96C2D4FE94FBB3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tshernyshev, Sergei E.;Alekseev, Vitalii;Bukejs, Andris;Vasilenko, Dmitry V.;Legalov, Andrej A.;Perkovsky, Evgeny E.	Tshernyshev, Sergei E., Alekseev, Vitalii, Bukejs, Andris, Vasilenko, Dmitry V., Legalov, Andrej A., Perkovsky, Evgeny E. (2025): First record of the family Melyridae (Coleoptera: Cleroidea) in late Eocene European ambers with description of a new tribe, a new genus and seven new species. Zootaxa 5696 (4): 542-566, DOI: 10.11646/zootaxa.5696.4.5, URL: https://doi.org/10.11646/zootaxa.5696.4.5
596FA515FFA6FFBC6D96C4A8FBFCF92F.text	596FA515FFA6FFBC6D96C4A8FBFCF92F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	(Asymelyris) Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov	<div><p>Subgenus Asymelyris Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov &amp; Perkovsky, subgen. nov.</p><p>urn:lsid:zoobank.org:act: 68AA255C-EBB5-4F37-84E8-708CB3E9F622</p><p>Type species: Amelyris (Asymelyris) paucidentatus Tshernyshev,Alekseev, Bukejs, Vasilenko, Legalov &amp; Perkovsky, sp. nov., by present designation</p><p>Diagnosis. Antennal club asymmetrical; sutural margin widened apically on elytral slope; third elytral carina tending to bend and merging with second carina at elytral apex; pygidium and last visible abdominal ventrite simple, lacking tufts of black bristles or hairs.</p><p>Etymology. The name of the new subgenus is a combination of the old Greek prefix ἀ- / ἀν- (meaning “not”), the initial syllable sy- derived from συμμετρικός (meaning “symmetrical”) and Melyris, the type-genus in the family Melyridae . The compound name refers to the asymmetrical antennal club of the subgenus representatives. Gender masculine.</p><p>Composition. The subgenus includes four extinct Eocene species: Amelyris (Asymelyris) paucidentatus sp. nov. (Baltic amber), Amelyris (Asymelyris) lupatus sp. nov. (Baltic amber), Amelyris (Asymelyris) groehni sp. nov. (Baltic amber), and Amelyris (Asymelyris) jutlandicus sp. nov. (Danish amber).</p></div>	https://treatment.plazi.org/id/596FA515FFA6FFBC6D96C4A8FBFCF92F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tshernyshev, Sergei E.;Alekseev, Vitalii;Bukejs, Andris;Vasilenko, Dmitry V.;Legalov, Andrej A.;Perkovsky, Evgeny E.	Tshernyshev, Sergei E., Alekseev, Vitalii, Bukejs, Andris, Vasilenko, Dmitry V., Legalov, Andrej A., Perkovsky, Evgeny E. (2025): First record of the family Melyridae (Coleoptera: Cleroidea) in late Eocene European ambers with description of a new tribe, a new genus and seven new species. Zootaxa 5696 (4): 542-566, DOI: 10.11646/zootaxa.5696.4.5, URL: https://doi.org/10.11646/zootaxa.5696.4.5
596FA515FFA6FFBD6D96C755FA71FC44.text	596FA515FFA6FFBD6D96C755FA71FC44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amelyris Tshernyshev & Alekseev & Bukejs & Vasilenko & Legalov & Perkovsky 2025	<div><p>Key to the species of Amelyris gen. nov. from Baltic, Rovno and Danish amber</p><p>1. Dilated antenomeres forming a loose symmetrical club; sutural margin of equal width along entire elytron ( Amelyris subgen. nov.)............................................................................................... 2</p><p>- Apical antennomeres widened outwards, forming a loose asymmetrical club; sutural margin widened apically on elytral slope ( Asymelyris subgen. nov.).............................................................................. 4</p><p>2. Lateral sides of pronotum bicarinate (with two longitudinal carinae)............................................ 3</p><p>- Lateral sides of pronotum tricarinate (with additional thin third carina between two main carinae).............................................................................................. A. (Amelyris) tricarinatus sp. nov.</p><p>3. Body parallel, narrow, flattened; pronotum appears rounded-square, not narrowed anteriorly; humeral carina thin, curved towards third carina, poorly visible at humeral base; elytral apex flattened............... A. (Amelyris) elongatus sp. nov.</p><p>- Body distinctly widened posteriorly, more or less oval, not flattened ventrally; pronotal disc appears trapezoidal, narrowed anteriorly; humeral carina short, curved, distinct along entire humerus; elytral apex not flattened.................................................................................................. A. (Amelyris) sveneldi sp. nov.</p><p>4. At least one lateral carina of pronotum dentate, lower carina arcuate, not parallel to upper carina; humeral carina straight, distinct along the entire humerus......................................................................... 5</p><p>- Lateral carinae of pronotum not dentate, straight, parallel, lower one not rounded, slightly protruding; humeral carina either curved or visible only at humeral base.................................................................... 6</p><p>5. All carinae of pronotum and elytral margin with small denticles; body narrow, almost parallel-sided; pronotum weakly transverse; sutural interval in elytra as wide as adjacent ones; elytral apex narrowly flattened and not elevated; first tarsomere 2.5× shorter than second tarsomere.............................................. A. (Asymelyris) lupatus sp. nov.</p><p>- Small denticles visible only on lower carina of pronotum, upper carina and elytral margin without denticles; sutural interval of elytra 1.5× wider than neighbouring ones; body looks wide; pronotum distinctly transverse; elytral apex distinctly flattened and elevated upwards; first tarsomere 1.5× shorter than second tarsomere.............. A. (Asymelyris) paucidentatus sp. nov.</p><p>6. Pronotum impressed only laterally at posterior angles, not impressed distally before anterior angles; middle part of disc convex; longitudinal carinae of elytra very thin, only third carina more pronounced; lateral margins widened at middle, not flattened distally. Elytral apices rounded separately. Body appears almost parallel-sided and narrow. Humeral carina very thin, poorly visible. First tarsomere twice shorter than tarsomere 2............................ A. (Asymelyris) jutlandicus sp. nov.</p><p>- Pronotum widely impressed at sides and distally; middle part of disc flattened; longitudinal carinae of elytra strongly expressed, elevated; lateral margins widened from base to apex of elytra. Elytral apices rounded together and straightly cut at apex. Body oval, widened behind. Humeral carina thin, prominent at base of humeral tubercle. First tarsomere 2.5× shorter than tarsomere 2......................................................................... A. (Asymelyris) groehni sp. nov.</p></div>	https://treatment.plazi.org/id/596FA515FFA6FFBD6D96C755FA71FC44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tshernyshev, Sergei E.;Alekseev, Vitalii;Bukejs, Andris;Vasilenko, Dmitry V.;Legalov, Andrej A.;Perkovsky, Evgeny E.	Tshernyshev, Sergei E., Alekseev, Vitalii, Bukejs, Andris, Vasilenko, Dmitry V., Legalov, Andrej A., Perkovsky, Evgeny E. (2025): First record of the family Melyridae (Coleoptera: Cleroidea) in late Eocene European ambers with description of a new tribe, a new genus and seven new species. Zootaxa 5696 (4): 542-566, DOI: 10.11646/zootaxa.5696.4.5, URL: https://doi.org/10.11646/zootaxa.5696.4.5
596FA515FFA7FFBF6D96C3BBFC4FFB0B.text	596FA515FFA7FFBF6D96C3BBFC4FFB0B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amelyris (Amelyris) sveneldi Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov	<div><p>Amelyris (Amelyris) sveneldi Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov &amp; Perkovsky, sp. nov.</p><p>urn:lsid:zoobank.org:act: 77C8564B-E8F3-4670-B6D1-91E6586300CF</p><p>(Figs 1–7)</p><p>Type material. Holotype No SIZK Be-26, adult, male; Rovno amber.</p><p>Locus typicus. Belokorovychi vicinity, Korosten District, Zhitomir Oblast, Ukraine.</p><p>Stratum typicum. Rovno amber, late Eocene.</p><p>Description. External colouration monochrome black-brown, without metallic lustre; dorsal setae dark brown to black. Body elongate, subparallel, flattened dorsally and moderately convex ventrally, slightly expanded and evenly rounded posteriorly. Measurements: body length 3.83 mm; body maximum width (in area of elytral base) 1.1 mm.</p><p>Head strongly prognathous, not sculptured, slightly convex, suboval, very weakly protruding distally and not wider than pronotum at eyes level; compound eyes oval, bean-shaped, large, weakly convex, bare; frons and interocular area flat, without impressions. Surface of head evenly and finely punctured, microsculpture distinct, surface sparsely covered with adpressed short, weakly visible hairs.Antennae 11-segmented, short, reaching to about the middle of the pronotum, attached to the head near the lower edge of the eyes near the clypeus, sparsely covered with short, semi-erect fine hairs. Antennomeres evenly dilated towards the apex forming lose 5-segmented club. Antennomere 1 swollen and enlarged, round-oval, 2.5× as long as antennomere 2; antennomere 2 oval-triangular; antennomere 3 elongate, subcylindrical, as long as antennomere 2, slightly narrower than antennomere 2 and 5 times as long as antennomeres 4, 5 or; antennomeres 7–10 broader, subtriangular, equilateral, forming loose club, approximately equal in lengths; antennomere 11 slightly longer than antennomere 10, flattened and evenly rounded, slightly narrowed apically. Clypeus slightly elongated and narrowed; labrum small, transverse and narrow, slightly emarginate in middle of distal side. Mandibles not covered by labrum, short, strong, rounded, bidentate. Maxillary palpi elongate, narrow, 4-segmented; palpomere 1 enlarged, round-oval, located on lateral side of clypeus below genae and clearly visible from dorsal side of head; three remaining palpomeres distinctly narrower, palpomeres 2 and 3 cylindrical; palpomere 4 elongate, narrowed and flattened towards apex, appearing subcylindrical. Labial palpi 4-segmented, with palpomeres of equal lengths, short; apical palpomere narrow and cylindrical.</p><p>Pronotum convex, slightly longer than wide, slightly narrowed anteriorly, with acute angles and typical bicarinate lateral side: upper carina straight and lower carina roundly prominent. The surface evenly and largely punctured; each puncture surrounded by raised limbus giving to surface cellular appearance; base of each cell distinctly microsculptured; margins of distal and basal sides and upper lateral carinae straight; lower carina provided with small protuberances or denticles. Pubescence small, curved, sparse. Disc distinctly impressed at posterior and anterior angles, narrowly impressed in centre of upper lateral carina.</p><p>Scutellum clearly visible, quadrate, with acute angles and straight sides, with small punctures and fine adpressed pubescence; disc mat, not marginate on sides.</p><p>Elytra carinate, elongate, subparallel, slightly convex just below middle, evenly narrowed posteriorly, with separately rounded apices, completely cover abdomen. Bases of elytra slightly broader than pronotum. Humeri distinct, small, not prominent, slightly impressed, with small short curved carina. Elytral disc probably mat. Each elytron with four full carinae; carina 1 (near-sutural) looks like elevated suture margin; carina 4 situated above flattened lateral side, distinct from basal quarter to apex of elytron. All carinae thin, distinct, not very prominent. Each interspace between carinae microsculptured, with three longitudinal regular rows of sparse, distinct punctures. Dorsal pubescence sparse, short, semi-recumbent, situated mainly on carinae. Elytral epipleura broad, extended to apex, with row of sparse punctures in middle, distinctly microsculptured. Metathoracic wings well developed.</p><p>Legs short, simple, not thickened, evenly flattened, densely and coarsely punctured to appear granular. Femora of all legs flattened and widened, not curved. Tibiae straight, evenly dilated to apex, spurs short and thin, same length as setae surrounding peripheral margin of tibia; surface of tibiae covered with short, black, strong, erect setae. All tarsi 5-segmented; tarsomeres strongly compressed, with short, black, erect hairs on periphery of pedal part. Basal tarsomeres small, triangular, 0.5× as long as corresponding tarsomeres 2 in anterior and posterior legs and 1.5× shorter than tarsomere 2 in intermediate legs; tarsomeres 2–4 in all legs equal in size, triangular, all combined 0.5× as long as tarsomere 5; tarsomere 5 longest. Tarsal claws distinct, narrow, sharp, symmetrical, with elongated basal tooth, giving claws bifid appearance.</p><p>Ventral side moderately convex, weakly shiny, finely and densely punctured, bare or with fine dense adpressed hairs not clearly visible under turbid surface in amber mass. Pygidium and last visible ventrite simple, slender, not depressed, with rectangular central emargination and tufts of several long, strong black bristles on outer margins of lateral sides.</p><p>Etymology. The specific epithet is a patronym; the new species is named after Sveneld, the semi-legendary 10th-century Varangian warlord that got tribute on the Drevlyan land.</p></div>	https://treatment.plazi.org/id/596FA515FFA7FFBF6D96C3BBFC4FFB0B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tshernyshev, Sergei E.;Alekseev, Vitalii;Bukejs, Andris;Vasilenko, Dmitry V.;Legalov, Andrej A.;Perkovsky, Evgeny E.	Tshernyshev, Sergei E., Alekseev, Vitalii, Bukejs, Andris, Vasilenko, Dmitry V., Legalov, Andrej A., Perkovsky, Evgeny E. (2025): First record of the family Melyridae (Coleoptera: Cleroidea) in late Eocene European ambers with description of a new tribe, a new genus and seven new species. Zootaxa 5696 (4): 542-566, DOI: 10.11646/zootaxa.5696.4.5, URL: https://doi.org/10.11646/zootaxa.5696.4.5
596FA515FFA5FFB16D96C570FD36FA9B.text	596FA515FFA5FFB16D96C570FD36FA9B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amelyris (Amelyris) elongatus Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov	<div><p>Amelyris (Amelyris) elongatus Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov &amp; Perkovsky, sp. nov.</p><p>urn:lsid:zoobank.org:act: 4DF7429B-78FE-4E74-AA62-B0892E9B12D8</p><p>(Figs 8–10)</p><p>Type material. Holotype No ISEZh, sch_012, adult, male; Baltic amber. Complete beetle is inside flat, trapezoid semi-transparent piece of amber with approximate dimensions 25 × 15 × 3 mm, with the beetle situated in a middle of the piece. Underside and legs are not clearly visible due to rows of air bubbles in amber matrix beneath the beetle. Syninclusions are represented by a number of stellate Fagaceae trichomes and many detrital particles.</p><p>Locus typicus. Southeastern Baltic Sea coast, the Sambian peninsula, Kaliningrad Oblast .</p><p>Stratum typicum. Baltic amber, late Eocene.</p><p>Description. Integument unicolorous black (as preserved), at certain lighting showing greenish metallic sheen; dorsal setation white and brown; legs dark brown, antennae black. Body elongate, subparallel, flattened dorsally and not convex ventrally, gradually widened from humeri posteriad, with maximum width in about the apical third. Measurements: body length 2.82 mm; body width (in the area of the base of the elytra) 0.95 mm.</p><p>Head prognathous and elongate distally; compound eyes large, oval, convex, bare; frons and interocular area flat, lacking impressions. Antennae 11-segmented, short, reaching base of pronotum; antennomeres evenly dilated apically and forming indistinct loose club. Antennomere 1 swollen, oval-triangular, 1.5× as long as antennomere 2; antennomere 2 oval-triangular, elongate, 1.2× as long as antennomere 3; antennomeres 3–10 transverse, oval-triangular, moniliform; antennomere 3 slightly wider than antennomere 2; each subsequent segment very slightly longer, but distinctly wider than the previous one; apical antennomere 2.0× as long as antennomere 10, flattened, uniformly rounded, slightly narrowed towards apex. Surface of antennae sparsely covered with short, erect fine hairs located on periphery of the antenomeres. Clypeus slightly elongate; labrum distinctly elongate, longitudinal. Maxillary palpi elongate, narrow, palpomere 1 elongate, subcylindrical, palpomere 2 longitudinal, 1.5× as long as palpomere 1, last palpomere strongly narrowed at apex and elongate, narrow, 1.5× as long as palpomere 1.</p><p>Pronotum slightly convex, appears oval-quadrate due to arcuate protruding distal side and rounded low lateral carinae, with slightly rounded distal angles and acute posterior ones; lateral side of pronotum bicarinate: upper carina straight, lower carina slightly roundly prominent. Surface evenly and coarsely punctured; covered with small curved adpressed white hairs, which caused shaping of the cellular appearance of the surface; microsculpture distinct between punctures; margins of basal side and upper lateral carinae straight; lower lateral carina slightly prominent; all margins smooth and without denticles. Pubescence thin, white and curved, sparsely distributed around punctures, poorly visible. Disc clearly impressed at posterior angles, along lateral and distal sides.</p><p>Scutellar shield clearly visible. rectangular, with acute angles and straight lateral sides, distal margin with weak triangular medial emargination; surface appears cellular as in pronotum due to curved fine pubescence associated with air in amber mass; punctures sparse, distinct; sides not marginate; surface matt.</p><p>Elytra carinate, elongate, subparallel, slightly widened just below humeri, with maximum width near apical slope, evenly narrowed towards apex, with apices separately rounded. Base of elytra slightly wider than pronotum; abdomen completely covered by elytra. Elytra disc mat. Sides of elytra flattened from humeri to elytral tips. Humeri small and weakly prominent, distinct, slightly impressed at base, with narrow curved carina bent over humerus to third carina.Elytral epipleura very narrow, distinct along elytra from base to metathorax, with distinct microsculpture; edges of epipleura smooth, without denticles. Each elytron with five thin and not strongly elevated carinae, carina 3 more distinctly protruding. All carinae extending from base to apical part; first carina situated close to suture and complete from base to apex, carinae 2 and 3 reaching apical slope of elytra and not fused; carina 4 situated close to carina 5 and ends at slope of elytron, curving towards the carina 3. Carina 5 coincides with the epipleura and is distinct from base to elytral apex. Surface between carinae with three regular rows of distinct round punctures and distinct microsculpture; pubescence short, sparse and small, situated mainly on carinae and occasionally near punctures. Metathoracic wings well developed.</p><p>Legs elongate, thin and simple, femora slightly thickened and evenly flattened with thin long bases. Tarsomeres strongly compressed and elongate, thin, first tarsomere 2.0× shorter than second tarsomere on all legs, tarsomeres 2 and 3 equal in size; last tarsomere 1.5× as long as tarsomere 4. Tarsal claws thin and long, slightly curved, lacking appendages at base.</p><p>Underside convex, mat, finely and densely punctured, glabrous. Last abdominal ventrite slightly elongate and narrow.</p><p>Etymology. The specific epithet of the new species elongatus (Latin adjective meaning “elongate, oblong”) refers to the general elongate habitus of the species.</p></div>	https://treatment.plazi.org/id/596FA515FFA5FFB16D96C570FD36FA9B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tshernyshev, Sergei E.;Alekseev, Vitalii;Bukejs, Andris;Vasilenko, Dmitry V.;Legalov, Andrej A.;Perkovsky, Evgeny E.	Tshernyshev, Sergei E., Alekseev, Vitalii, Bukejs, Andris, Vasilenko, Dmitry V., Legalov, Andrej A., Perkovsky, Evgeny E. (2025): First record of the family Melyridae (Coleoptera: Cleroidea) in late Eocene European ambers with description of a new tribe, a new genus and seven new species. Zootaxa 5696 (4): 542-566, DOI: 10.11646/zootaxa.5696.4.5, URL: https://doi.org/10.11646/zootaxa.5696.4.5
596FA515FFABFFB36D96C5C0FB82FA53.text	596FA515FFABFFB36D96C5C0FB82FA53.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amelyris (Amelyris) tricarinatus Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov	<div><p>Amelyris (Amelyris) tricarinatus Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov &amp; Perkovsky, sp. nov.</p><p>urn:lsid:zoobank.org:act: ACACD6E7-3857-4AF9-AC7F-AB5810B105C2</p><p>(Figs 11–13)</p><p>Type material. Holotype No ISEZh, sch_013, adult, male; Baltic amber. Complete beetle included in right triangular semi-transparent piece of amber with approximate dimensions 25 × 11 × 4 mm, with the beetle located near right angle of the piece. Underside and legs are not well visible due to milky-white coating and opacities in amber matrix. Syninclusions are represented by a few stellate Fagaceae trichomes and many detrital particles.</p><p>Locus typicus. Southeastern Baltic Sea coast, the Sambian peninsula, Kaliningrad Oblast .</p><p>Stratum typicum. Baltic amber, late Eocene.</p><p>Description. Integument unicolorous black (as preserved); dorsal setae brown to black, legs light brown to dark yellow. Body elongate, parallel, flattened dorsally and convex ventrally, very slightly expanded below the middle, with maximum width at about the middle; elytra slightly narrowed towards the apex and rounded, elytral apices closed. Measurements: body length 2.58 mm; body width (in the area of the base of the elytra) 0.8 mm.</p><p>Head prognathous, not sculptured, convex, subrounded, not protruding distally and not wider than pronotum at eye level. Compound eyes round-oval, protruding, bare; frons and interocular area evenly convex, without impressions. Surface of head densely punctate, with distinct microsculpture and covered with short, adpressed hairs which, in combination with punctures, give the surface a “cellular” structure as on pronotum. Antennae 11-segmented, short, attached to the head near lower edge of eyes near clypeus, extending over middle but not reaching base of pronotum. Last five antennomeres dilated, forming 5-segmented loose club. Antennomeres 1 and 2 slightly swollen and enlarged, round-oval; 1.5× as long as antennomere 2; antennomere 3 slightly elongated, subcylindrical, slightly narrower and shorter than antennomere 2 and 1.5× as long as antennomere 4; antennomeres 4–10 wide, subtriangular, moniliform, symmetrically dilated; antennomeres 7–11 forming loose club; antennomere 10 subquadrate; antennomere 11 slightly longer and wider than antennomere 10, evenly rounded and flattened at apex.</p><p>Pronotum convex, slightly longer than wide, quadrate-rectangular due to straight distal and basal margins and straight upper lateral carina, with slightly rounded angles, laterally with typical double carinate margins: upper carina straight, lower carina rounded, small longitudinal rib present in middle of lateral side between upper and lower carina. Pronotal surface evenly and coarsely punctured and covered with small adpressed hairs; microsculpture distinct; margins of distal and basal sides and upper lateral carinae straight, lower lateral carina widely prominent; all margins smooth without denticles. Pubescence short, curved, sparse, inconspicuous. Disc clearly impressed: at basal angles, near anterior angles, and narrowly along upper carina.</p><p>Scutellar shield clearly visible, rectangular, with acute angles and straight lateral sides; distal margin with triangular emargination in middle; surface with small punctures and fine adpressed pubescence; disc not marginate, mat.</p><p>Elytra carinate, elongate and parallel, slightly widened near middle, evenly weakly narrowed and rounded towards apex, completely cover abdomen, with closed apices. Bases of elytra slightly wider than pronotum. Humeri small and weakly prominent, distinct, slightly impressed at base, with small, narrow, weakly curved carina. Elytral disc probably dull. Each elytron with five thin and not strongly elevated carinae extending from base to apical part; first carina situated near suture, complete from base to apex; second third carinae not merged at elytral apex; fourth carina begins at base of elytra and continues on outside of humerus and along lateral side to elytral slope; fifth carina strongly attached to epipleura and distinct to end of elytral slope. Surface between carinae with three rows of distinct punctures and developed microsculpture, pubescence short, sparse and small, situated mainly on carinae. Epipleura narrow, distinct along elytra from base to apex, with row of sparse punctures in middle and distinct microsculpture; edges of epipleura smooth, lacking denticles. Metathoracic wings well developed.</p><p>Legs short, simple; femora slightly thickened and evenly flattened; tarsomeres 1–4 strongly compressed, first metatarsomere 2.0× shorter than metatarsomere 2.</p><p>Ventral side convex, mat, finely and densely punctate, bare.</p><p>Etymology. The specific epithet of the new species tricarinatus (Latin adjective meaning “having three ridges or keels”) refers to the three lateral carinae of pronotum, characteristical for the species.</p></div>	https://treatment.plazi.org/id/596FA515FFABFFB36D96C5C0FB82FA53	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tshernyshev, Sergei E.;Alekseev, Vitalii;Bukejs, Andris;Vasilenko, Dmitry V.;Legalov, Andrej A.;Perkovsky, Evgeny E.	Tshernyshev, Sergei E., Alekseev, Vitalii, Bukejs, Andris, Vasilenko, Dmitry V., Legalov, Andrej A., Perkovsky, Evgeny E. (2025): First record of the family Melyridae (Coleoptera: Cleroidea) in late Eocene European ambers with description of a new tribe, a new genus and seven new species. Zootaxa 5696 (4): 542-566, DOI: 10.11646/zootaxa.5696.4.5, URL: https://doi.org/10.11646/zootaxa.5696.4.5
596FA515FFA9FFB56D96C588FD6FF809.text	596FA515FFA9FFB56D96C588FD6FF809.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amelyris (Asymelyris) paucidentatus Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov	<div><p>Amelyris (Asymelyris) paucidentatus Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov &amp; Perkovsky, sp. nov.</p><p>urn:lsid:zoobank.org:act: B777E7D2-EEAC-44E8-9CB8-6B5D00D965FB</p><p>(Figs 14–16)</p><p>Type material. Holotype No KAM 8636 [KRAM], adult, male; Baltic amber. Complete beetle included in flat, irregular, transparent amber piece with approximate dimensions 43 × 26 × 7 mm, with the beetle situated in lateral part of the piece. Prosternum, mesosternum and head are not clearly visible due to milky-white coating and opacities in amber matrix. Syninclusions are represented by few stellate Fagaceae trichomes, many detrital particles and small plant fragments, numerous gas vesicles, one specimen of Collembola (Symphypleona), one specimen of Chalcidoidea ( Hymenoptera), one insect larva (?Lepidoptera), and one imago of a beetle ( Coleoptera), densely covered by milky coating.</p><p>Locus typicus. Southeastern Baltic Sea coast, the Sambian peninsula, Kaliningrad Oblast .</p><p>Stratum typicum. Baltic amber, late Eocene.</p><p>Description. Integument unicolorous black, without metallic lustre (as preserved), legs and antennae brown to light brown; dorsal setae brown. Body dilated, moderately elongate, depressed and moderately convex, evenly expanded just behind middle and narrowing at apex, with closed elytral apices. Measurements: body length (combined length of elytra, pronotum, and head along midline, measured dorsally) 3.25 mm; body maximum width across elytra 1.38 mm; pronotum length 0.7 mm; pronotum maximum width (at base) 0.95 mm; elytra width at base (combined) 1.0 mm; elytra length 2.25 mm.</p><p>Head prognathous, glabrous, convex, subrounded, not protruding distally and narrower than pronotum at eye level, transversely rugose in frons (forming shallow cell-like sculpture), finely punctate, covered with adpressed short hairs in vertex; compound eyes round-oval, vertical, bare, finely facetted; frons and interocular area evenly convex, lacking impressions. Antennae 11-segmented, attached to head near lower edge of eyes near clypeus, short, extending over middle but not reaching base of pronotum; ultimate antennomeres asymmetrically dilated, forming 5-segmented loose serrate antennal club). Antennomeres 1 and 2 swollen and enlarged, round-oval; antennomere 2 slightly shorter than antennomere 1; antennomeres 3–10 transversely triangular, with extremely extended outer edges; antennomere 3 small, narrowest; antennomeres 4–7 cylindrical, approximately equal in lengths, gradually widened, 1.5× as long as wide; antennomeres 8–10 triangular, dilated apically, about 1.5× as wide as antennomere 6; antennomere 11 flame-shaped, elongated, longitudinal, narrowed, truncate at apex. Terminal maxillary palpomere large, 3× as long as previous palpomere, narrowly triangular and narrowly rounded apically.</p><p>Pronotum convex, transverse, about 1.4× wider than long, trapezoidal in dorsal view, appears rectangular due to straight distal and basal margins and straight upper lateral carina, with rounded apical angles and acute basal angles, laterally with typical dicarinate margin: upper carina straight, lower carina rounded and distinctly prominent. Surface evenly densely punctured with strong punctures, distinct microsculpture, covered with fine, curved, slightly compressed hairs, giving the surface cellular appearance due to air retention by hairs in amber mass; margins of distal and basal sides and upper lateral carina smooth, lower lateral carina with denticles. Pronotal disc distinctly impressed: at anterior and posterior angles, along lateral upper carina.</p><p>Scutellar shield small, very short, transverse, about 0.5× as long as wide, rectangular, with acute angles and straight sides, surface sparsely punctured with small punctures and covered with fine adpressed pubescence; disc not marginate on sides, mat.</p><p>Elytra carinate, strongly dilated, suboval, slightly expanded just behind middle, about 1.6× as long as wide, evenly rounded and narrowed posteriorly with weakly stretched apex, completely cover abdomen; elytral apices separately rounded. Bases of elytra distinctly wider than pronotum. Humeri small, not prominent, with thin small rib located on each humerus and curving to third elytral carina. Elytral disc shiny. Each elytron with five thin and not strongly elevated carinae reaching from base to apical part; first carina located near suture, complete from base to apex, widened at elytral slope; carinae 2 and 3 reaching elytral slope, approximated but not merged; carina 4 shortened, begins at about humerus and ends at beginning of elytral slope; carina 5 strongly adjacent to epipleura and continues to elytral apex. Intervals between carinae wide; distance between rows of punctures equal to 0.5× diameter of one puncture; first interval 1.5× as wide as second one. Lateral sides of elytra flattened in apical half; lateral margin elevated and curved dorsally at elytral apex. Surface between punctures with distinct microsculpture; pubescence light, short and small, evenly distributed on carinae. Epipleura wide from base to middle of abdominal ventrite 1, then twice narrowed and distinct to apex; row of sparse punctures with microsculpture, distinct in middle of the widened part of epipleura; edges of epipleura smooth lacking denticles. Metathoracic wings well developed.</p><p>Legs short, simple, not flattened; impunctate, sparsely covered with fine, short pubescence. Femora thickened; tibiae evenly widened posteriorly, rounded. Tibial spurs spicular, short and sharp. Tarsomeres compressed; first tarsomere 1.5× as short as tarsomere 2 in anterior and intermediate legs and about 1.2× as short as tarsomere 2 in posterior legs; tarsomeres 2 and 3 equal in size; tarsomere 4 weakly shorter than tarsomere 3 in all legs; tarsomere 5 subcylindrical, as long as tarsomeres 2–4 combined. Tarsal claws broad, short; with not sharp, short and curved basal denticle.</p><p>Ventral side convex, shining, finely and sparsely punctate; pubescence fine, short and adpressed. Abdomen with five visible ventrites, finely and sparsely pubescent; intercoxal process narrowly triangular; ventrite 1 longest; ventrite 5 with wide and rounded emargination apically.</p><p>Etymology. The epithet of the new species paucidentatus is derived from the combined Latin adjective paucus (meaning ‘few, little’) and the Latin adjective dentatus (meaning ‘toothed’). The name refers to the comparatively few number of denticulate pronotal carinae.</p></div>	https://treatment.plazi.org/id/596FA515FFA9FFB56D96C588FD6FF809	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tshernyshev, Sergei E.;Alekseev, Vitalii;Bukejs, Andris;Vasilenko, Dmitry V.;Legalov, Andrej A.;Perkovsky, Evgeny E.	Tshernyshev, Sergei E., Alekseev, Vitalii, Bukejs, Andris, Vasilenko, Dmitry V., Legalov, Andrej A., Perkovsky, Evgeny E. (2025): First record of the family Melyridae (Coleoptera: Cleroidea) in late Eocene European ambers with description of a new tribe, a new genus and seven new species. Zootaxa 5696 (4): 542-566, DOI: 10.11646/zootaxa.5696.4.5, URL: https://doi.org/10.11646/zootaxa.5696.4.5
596FA515FFACFFA86D96C0F1FCC1FDDF.text	596FA515FFACFFA86D96C0F1FCC1FDDF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amelyris (Asymelyris) lupatus Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov	<div><p>Amelyris (Asymelyris) lupatus Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov &amp; Perkovsky, sp. nov.</p><p>urn:lsid:zoobank.org:act: 34AB5AF4-46A7-4C13-95FA-1FD5DB0A5761</p><p>(Figs 17–19)</p><p>Type material: Holotype No KAM 4915 [KRAM], adult, male; Baltic amber. A small elongate piece of transparent amber 21 × 7 × 3 mm in size and preserved without supplementary fixation, with the beetle situated in central part of the piece and diagonally located to the longitudinal axis of the piece. Pronotum, lateral portion of mesothorax and anterior third of elytra are not clearly visible due to milky-white coating and opacities of amber matrix. A thermal processing of the amber piece in an autoclave is assumed based on the dark, semitransparent halo-like coloration in cracks around of the inclusions. Syninclusions consist of few stellate Fagaceae trichomes.</p><p>Locus typicus. Southeastern Baltic Sea coast, the Sambian peninsula, Kaliningrad Oblast .</p><p>Stratum typicum. Baltic amber, late Eocene.</p><p>Description. Body integument unicolorous dark (as preserved), at certain lighting showing greenish metallic sheen; legs brown to light brown; dorsal setae brown. Body elongate and subparallel, depressed, not convex, not expanded and evenly rounded posteriorly with jointly rounded elytral apices. Measurements: body length (combined length of elytra, pronotum, and visible part of declined head along midline, measured dorsally) 2.8 mm; body maximum width across elytra 1.1 mm; pronotum length 0.65 mm; pronotum maximum width 0.8 mm; elytra width at base (combined) 0.9 mm; elytra length 2.15 mm.</p><p>Head prognathous, not sculptured, subrounded, slightly protruding distally and narrower than pronotum at eye level; compound eyes small, round-oval, bare, not protruding, frons and interocular area flat, lacking impressions. The surface of head densely punctured, with distinct micro-sculpture, covered with short, adpressed hairs giving surface cellular appearance in amber mass. Antennae 11-segmented, attached to head near inferior edge of eyes, short, reaching base of pronotum; antennomeres 4–10 asymmetrically dilated; with five apical antennomeres forming indistinctly loose club. Antennomeres 1 and 2 swollen, round-oval; antennomere 2 about 0.5× as long as antennomere 1, moderately narrow; antennomere 3 elongate, subconic, slightly longer as antennomere 2; antennomeres 4–10 cup-shaped and transverse; antennomere 4 about 2.0× as wide as antennomere 3 and 1.5× as narrow as antennomere 5; antennomeres 5 and 6 equal size and 1.2× as narrow as antennomere 7; antennomeres 7–10 equal in size; antennomere 11 elongate-oval, moderately narrow, longer than antennomere 10. Antennae with erect, fine, light pubescence visible on outer margins of antennomeres. Terminal maxillary palpomere large, 3.0× as long as previous palpomere, spindle-form, narrowly rounded apically.</p><p>Pronotum convex, weakly transverse, about 1.2× wider than long, appears rectangular due to straight distal and basal margins and straight upper lateral carina, all angles acute and almost completely right-angled; laterally with typical dicarinate margins: upper carina straight, lower carina roundly prominent. Surface evenly and densely punctate, but poorly visible due to blanking in the amber mass. Disc distinctly impressed at anterior and posterior angles. Margin of upper carina with small denticles, margin of lower carina with strong scalloped denticles. Distal margin of pronotum also rough, with denticles.</p><p>Scutellar shield short, broad, transverse, small, rectangular, with acute angles and straight sides.</p><p>Elytra carinate, parallel, suboval, very slightly expanded posteriorly, evenly rounded and slightly narrowed at apex, completely covering abdomen. Elytral apices jointly rounded. Bases of elytra distinctly wider than pronotum. Humeri small, not prominent, with distinct carina over humerus near base of carina 4. Elytral disc shiny. Each elytron with five thin and not strongly elevated carinae spanning from base to the apical part; first carina close to suture and entire from base to apex of elytra, not coincides with sutural margin; carinae 2 and 3 continued to end of elytral slope and terminate just at flattened apices, parallel and not merged; carina 4 started below humerus and reaching end of elytral slope; carina 5 strongly coincides with upper margin of epipleuron and continues to elytral apex. Intervals between carinae even and broad, with three rows of punctures separated by distance of one-half diameter of each puncture. Sides of elytra narrowly flattened and curved dorsally from level of humeri to elytral apex. Surface between punctures with distinct microsculpture. Epipleura gradually narrowed apical, with row of sparse punctures and distinct microsculpture; margins with small, sharp, distinct denticles. Metathoracic wings well developed.</p><p>Legs short, simple. Femora flattened; tibiae thin, slightly widened anteriorly.Tibial spurs short, sharp.Tarsomeres compressed; tarsomere 1 about 2.5× shorter than tarsomere 2 in anterior and intermediate legs, metatarsomere 1 about 2.0× shorter than metatarsomere 2; tarsomeres 2–4 in all legs equal in size; ultimate tarsomere largest, subcylindrical and equal in length to tarsomeres 3–4 combined. Tarsal claws wide, short, not sharp, denticulate. Surfaces of legs coarsely punctured; covered with fine, short, adpressed pubescence. Pubescence of tarsomeres arranged on apical edge of ventral side.</p><p>Ventral side slightly convex, shiny, densely pubescent, with short adpressed dark hairs. Ventrites 1 and 2 most likely connate and have not changed their configuration after ventrally curved abdomen. Pygidium and apical ventrite simple, transverse, depressed or emarginated.</p><p>Etymology. The epithet of the new species lupatus (Latin adjective meaning ‘jagged’) refers to the denticulated both carinae of pronotum and elytral margin in the species.</p></div>	https://treatment.plazi.org/id/596FA515FFACFFA86D96C0F1FCC1FDDF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tshernyshev, Sergei E.;Alekseev, Vitalii;Bukejs, Andris;Vasilenko, Dmitry V.;Legalov, Andrej A.;Perkovsky, Evgeny E.	Tshernyshev, Sergei E., Alekseev, Vitalii, Bukejs, Andris, Vasilenko, Dmitry V., Legalov, Andrej A., Perkovsky, Evgeny E. (2025): First record of the family Melyridae (Coleoptera: Cleroidea) in late Eocene European ambers with description of a new tribe, a new genus and seven new species. Zootaxa 5696 (4): 542-566, DOI: 10.11646/zootaxa.5696.4.5, URL: https://doi.org/10.11646/zootaxa.5696.4.5
596FA515FFB2FFAA6D96C230FC89F8B8.text	596FA515FFB2FFAA6D96C230FC89F8B8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amelyris (Asymelyris) groehni Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov	<div><p>Amelyris (Asymelyris) groehni Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov &amp; Perkovsky, sp. nov.</p><p>urn:lsid:zoobank.org:act: 7B84671C-B1A5-4DD0-9EDF-70932880A476</p><p>(Figs 20–25)</p><p>Type material. Holotype No. GPIH 5223, CCGG 8709 (ex coll. Jonas Damzen JDC-6437), male; Baltic amber. A flat piece of transparent amber 30×18× 8 mm in size and preserved without supplementary fixation. Ventral side of the specimen partially covered by milky-white coating. Syninclusions consist of few stellate Fagaceae trichomes.</p><p>Paratype No. ABAC 066 [ABAC] (ex coll. Jonas Damzen JDC-8916), female; Baltic amber. A flat piece of transparent amber 48×30× 8 mm in size and preserved without supplementary fixation, with the beetle situated in lateral part of the piece. Syninclusions consist of one small spider specimen ( Araneae), few stellate Fagaceae trichomes and detrital particles.</p><p>Paratype No. GPIH 5225, CCGG C-4746, male; Baltic amber. A flat piece of transparent amber 25×17× 6 mm in size and preserved without supplementary fixation, with the beetle situated in lateral part of the piece. Ventral side of the specimen covered by milky-white coating and opacities of amber matrix. Syninclusions consist of one Brachycera (Diptera) specimen and few stellate Fagaceae trichomes .</p><p>Paratype No. GPIH 5224, CCGG 8710 (ex coll. Jonas Damzen JDC-12535), sex unknown; Baltic amber. A flat piece of transparent amber 25×11× 4 mm in size and preserved without supplementary fixation. Syninclusions consist of few stellate Fagaceae trichomes .</p><p>Locus typicus. Southeastern Baltic Sea coast, the Sambian peninsula, Kaliningrad Oblast .</p><p>Stratum typicum. Baltic amber, late Eocene.</p><p>Description of holotype. Body elongate, suboval, weakly depressed and moderately convex, evenly dilated and rounded posteriorly with straightly cut apex); unicolorous dark brown, without metallic luster; pubescence: head, pronotum and elytra with short, fine, recumbent to semi-erect setae, ventral side of specimen with inconspicuous, recumbent setation. Measurements: body length (pronotum + elytra) 2.54 mm, maximum body width 1.15 mm; pronotal length 0.54 mm, pronotal maximum width 0.84 mm; elytral length 2.0 mm, elytra width at base 0.92, elytral maximum width (in posterior one-third) 1.15 mm.</p><p>Head prognathous, slightly convex, round, not stretched distally and not wider than pronotum at level of eyes; evenly and finely punctate. Compound eyes oval, rather small, slightly prominent, bare. Frons and interocular area flat. Antennae 11-segmented, with lose 5-segmented club, inserted near lower inner edge of eye near clypeus, narrow, slightly elongate, reaching base of elytra, sparsely covered with fine, erect setae; antennomeres 5–10 evenly widened towards apex, subtriangular to triangular, asymmetric, with stretched outer edges; antennomeres 1 and 2 subcylindrical, swollen and slightly enlarged, elongate, 1.7× as long as wide, subeqal in size; antennomere 3 conical, slightly dilated apically, 1.7× shorter and narrower than antennomere 2; antennomere 4 subconical, about as long as antennomere 3; antennomeres 5 and 6 subtriangular, elongate, slightly longer and wider than antennomere 4; antennomeres 7–8 triangular, about as long as wide; antennomeres 9–10 widely triangular, transverse, 1.3× as wide as long; antennomere 11 elongate, 1.5× as long as wide, subconical, wide basally and narrowed apically. Clypeus slightly narrowed anteriad, slightly convex medially. Labrum distinct, slightly transverse, distinctly narrowed anteriad, subtrapezoidal with rounded, not emarginate distal margin, not covers mandibles. Mandibles short, apparently bidentate. Maxillary palpi narrow; ultimate palpomere elongate, 2.3× as long as wide, conical, evenly narrowed to apex, about 2.5× as long as penultimate antennomere.</p><p>Pronotum convex, transverse, 1.6× as wide as long, evenly rounded and narrowed anteriorly, with acute and slightly protruding posterior angles; laterally with double carinate side: upper carina widely rounded, not protruding, distinctly serrate, lower carina almost straight, curved inward medially along disc, apparently smooth, lacking small protuberances or denticles. Surface with large, irregular cells and dense micropunctation. Pronotal disc with weak, narrow, longitudinal impression medially; distinctly impressed posterorolaterally and along lateral sides, weakly impressed along anterior side.</p><p>Scutellar shield rather large, subtrapezoidal, weakly transverse, 1.3× as wide as long, densely covered with fine punctation.</p><p>Elytra carinate, subparallel-sided, elongate and suboval, 1.7× as long as wide, slightly widened at apical one-third, evenly rounded posteriorly with apices closed and straightly cut. Lateral sides strongly flattened from base to apices. Base of elytra slightly wider than pronotum; elytra cover abdomen completely. Humeri small, slightly protruding, not impressed, narrow carina short and distinct at base of humerus only. Each elytron with five narrow, distinctly convex carinae; carina 1 located near elytral suture and almost completely coincides with margin of suture, but distinctly separated at elytral slope forming ‘widened’ appearance to suture; carinae 2 and 3 entire from base to slope of elytra and not merged on slope; carina 4 started under humerus and continued to elytral slope; carina 5 distinct from base to elytral apices. Interspaces between carinae with three rows of rather large punctures and dense micropunctation. Epipleura not vertical, wide, evenly narrowing posteriad, reaching elytral apices, with row of sparse, small punctures and microsculture, lateral margins of epipleura with small inconspicuous denticles. Metaventrite with disc convex, densely covered with fine punctation. Metanepisternum wide, about 4.4× as long as wide; with dense micropunctation. Metathoracic wings well developed and partially exposed from under right elytral apex.</p><p>Legs short, simple, covered with fine recumbent setae. Femora evenly narrowly suboval, flattened, with dense and fine punctation. Tibiae evenly slightly dilated to apex; with fringe of stout, dark setae apically and apparently with two fine apical spurs. Tarsi compact, all tarsi 5-segmented, with setation ventrally, tarsomeres 1–4 subconical, slightly dilated apically, tarsomere 5 subcylindrical, slightly curved, elongate; tarsomere 1 shortest, about 0.8× as long as tarsomere 2; tarsomeres 2–4 nearly equal size; tarsomere 5 longest in all legs, as about as long as tarsomeres 2–4 combined. Claws small, sharp, symmetrical, with elongated basal tooth.</p><p>Abdomen with six visible ventrites, densely covered with fine punctation, ventrites 1 and 2 apparently connate. Pygidium and last ventrite simple, not emarginate or depressed, and lacking tufts of hairs; pygidium slightly elongate and domelike; last ventrite with apical margin widely rounded and covered with fine, erect setae.</p><p>Paratype GPIH 5225, CCGG C-4746, male. Measurements: total body length (pronotum + elytra) 2.92 mm (3.0 mm including visible part of head), body maximum width 1.15 mm. Pronotal length 0.69 mm, pronotal maximum width 0.92 mm. Elytral length 2.23 mm, elytral width at base 1.0 mm, elytral maximum width (in posterior one-third) 1.15 mm. Otherwise, this specimen is similar in all visible morphological characters to holotype No. 5223 [GPIH] .</p><p>Paratype No. ABAC 066, female. Measurements: total body length (pronotum + elytra) 2.46 mm (2.61 mm including visible part of head), maximum body width 1.15 mm; pronotal length 0.54 mm, pronotal maximum width 0.84 mm; elytral length 1.92 mm, elytral width at base 0.92 mm; elytral maximum width (in posterior one-third) 1.15 mm .</p><p>It differs from male by more slender antennae with elongate and more narrower antennomeres forming indistinct club; inner margin of epipleurae with distinct small denticles more distinctly prominent posteriorly (probably due to clear visibility of beetle in amber piece, while epipleura poorly visible in holotype); legs more narrower, especially tibiae, tarsi thinner and shorter, but claws longer. Otherwise, this specimen is similar in all visible morphological characters to holotype No. 5223 [GPIH] .</p><p>Paratype GPIH 5224, CCGG 8710, sex unknown. Measurements: total body length (pronotum + elytra) 3.0 mm (3.1 mm including visible part of head), body maximum width 1.2 mm; pronotal length 0.7 mm, pronotal maximum width 0.8 mm; elytral length 2.3 mm, elytral width at base 0.9 mm, elytral maximum width (in posterior one-third) 1.2 mm. Otherwise, this specimen is similar in all visible morphological characters to holotype No. 5223 [GPIH] .</p><p>Etymology. The specific epithet is a patronym; the new species is named in honor of Mr. Carsten Gröhn (Glinde, Germany), an enthusiast and specialist in Baltic amber.</p></div>	https://treatment.plazi.org/id/596FA515FFB2FFAA6D96C230FC89F8B8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tshernyshev, Sergei E.;Alekseev, Vitalii;Bukejs, Andris;Vasilenko, Dmitry V.;Legalov, Andrej A.;Perkovsky, Evgeny E.	Tshernyshev, Sergei E., Alekseev, Vitalii, Bukejs, Andris, Vasilenko, Dmitry V., Legalov, Andrej A., Perkovsky, Evgeny E. (2025): First record of the family Melyridae (Coleoptera: Cleroidea) in late Eocene European ambers with description of a new tribe, a new genus and seven new species. Zootaxa 5696 (4): 542-566, DOI: 10.11646/zootaxa.5696.4.5, URL: https://doi.org/10.11646/zootaxa.5696.4.5
596FA515FFB1FFAD6D96C0F1FE49FD4B.text	596FA515FFB1FFAD6D96C0F1FE49FD4B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amelyris (Asymelyris) jutlandicus Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov	<div><p>Amelyris (Asymelyris) jutlandicus Tshernyshev, Alekseev, Bukejs, Vasilenko, Legalov &amp; Perkovsky, sp. nov.</p><p>urn:lsid:zoobank.org:act: 6E9CED7A-9E17-4C8E-AF59-FF4E738025F9</p><p>(Figs 26–28)</p><p>Type material. Holotype No NHMD 153481 with label “ Cucujidae B. Mortensen / 11-7-1960 ”; adult, male. Danish amber.</p><p>Locus typicus. Denmark, precise locality unknown.</p><p>Stratum typicum. Danish amber, late Eocene.</p><p>Description. External colouration monochromous black-brown, probably with a faint green metallic lustre; dorsal setae black. Body elongate, subparallel, slightly depressed and moderately convex, slightly expanded around the middle and evenly rounded posteriorly with separately rounded elytral apices. Measurements: body length 3.0 mm; body width (in the area of the base of the elytra) 0.95 mm.</p><p>Head prognathous, not sculptured, evenly convex, slightly impressed above clypeus, suboval, weakly protruding distally and not wider than pronotum at eye level. Compound eyes round, large, convex, bare. Frons and interocular area flat, without impressions. The surface of head evenly and finely punctate, microsculpture distinct, surface sparsely covered with adpressed short, weakly visible hairs, which, in association with air, assembled cellular structure remained in amber mass. Antennae 11-segmented, attached to head near lower inner edge of eyes near clypeus, short, reaching base of pronotum; antennomeres evenly widened towards apex, forming indistinct loose club. First antennomere swollen and enlarged, round-oval, 2.0× as long as antennomere 2; antennomere 2 oval-triangular, asymmetrical with prominent outer margin; antennomere 3 elongate, triangular, as long as antennomere 2, 2.0× as narrow as antennomere 2, 1.2× as long as antennomere 4; antennomere 4 wide, triangular, equilateral; antennomeres 2–10 triangular, asymmetrical with prominent outer margin; apical segments not strongly widened, triangular, transverse; antennomere 11 oval-rectangular with rounded apex. Antennal club indistinct, loose; surface of antennae shiny, sparsely covered with fine, short, light-coloured hairs. Clypeus very short, narrow, transverse and slightly elevated distally; labrum distinct, slightly transverse, distinctly elongate, narrowed anteriorly, not covering mandibles, distal margin straight. Mandibles not protruding, short and bidentate. Maxillary palpi, elongate, narrow, palpomere 1 subtriangular, palpomeres 2 and 3 elongate-triangular, equal in length to palpomere 1; apical palpomere 2.0× as long as palpomere 1, widened and flattened in apical half, narrowing further to sharp tip. Labial palpi of similar shape, cylindrical, narrow and elongate, with apical palpomere moderately widened anteriorly.</p><p>Pronotum convex, distinctly longitudinal, weakly narrowed anteriorly, with acute and slightly stretched posterior angles; laterally with dicarinate side: upper and lower carinae straight, slightly curved inwards distally along disc. Surface evenly punctured with moderate-sized punctures provided with short brown adpressed hairs, surface of pronotum appears cellular, microsculpture distinct in bottom of each cell. Margins of distal and basal sides of pronotum slightly prominent; lateral carinae straight, not prominent but slightly curved distally; all margins smooth, lacking small protuberances or denticles. Pronotal pubescence short, curved, sparse. Disc clearly impressed at posterior angles and near anterior angles, with flattened lateral side along upper carina.</p><p>Scutellar shield distinct, subquadrate, with acute angles and slightly widened distally; all margins straight; punctured as in pronotum; with fine adpressed pubescence.</p><p>Elytra carinate, elongate, subparallel, slightly widened just below middle, evenly rounded posteriorly, completely covering abdomen, with apices separately rounded. Lateral sides flattened from level of humeri towards elytral slope, apices not flattened. Bases of elytra slightly wider than pronotum. Humeri small, slightly protruding and indistinct, very slightly impressed inwards, provided with narrow short curved carina on upper side. Each elytron with five full carinae reaching from base to apex; all carinae thin and not strongly elevated, only carina 3 distinctly protruding; carina 1 close to suture and looks like elevated sutural margin, but distinctly separated from suture margin at elytral slope and forming wide ‘suture’ appearance; carinae 2 and 3 entire from base to slope of elytra, merged on slope; carina 4 begins below humerus and continues along lateral side towards slope of elytron; carina 5 distinct from base to apex of elytron. Carinae thin, not strongly prominent, only carina 3 more distinctly protruding. Surface between carinae with three longitudinal rows of distinct punctures and developed microsculpture; pubescence sparse, short, located mainly on carinae and near punctures. Elytral epipleura wide, distinct along elytra from base to apex, but wider from base to end of first abdominal ventrite, with a row of sparse punctures in the middle from base to ventrite 4, and further without punctures but with transverse wrinkles; microsculpture distinct. Metathoracic wings well developed.</p><p>Legs short, simple, not thickened, densely punctured. Femora of all legs flattened and widened, not curved, reaching epipleura. Tibiae straight, evenly dilated to apex, with two spurs strong slightly curved and sharp, with surface covered by short black hairs along ventral surface and by light erect setae on outer edge near spurs. Tarsi 5-segmented, tarsomeres strongly compressed, not compact or elongate, with short light erect hairs on periphery of pedal part; tarsomeres 1–4 triangular, tarsomere 5 elongate and slightly conical; tarsomere 1 2.0× as short as tarsomere 2 in anterior and intermediate legs and 1.5× as short as tarsomere 2 in posterior legs; tarsomeres 2–4 in all legs equal in size; tarsomere 5 longest in all legs, as long as tarsomeres 2–4 combined. Tarsal claws distinct, narrow, curved and sharp, with strong, sharp tooth at base.</p><p>Ventral side moderately convex, densely punctured, fine dense adpressed hairs. Six abdominal segments visible; ventrites 1 and 2 probably connate. Pygidium (apical tergite) and last abdominal ventrite simple, moderately elongate, narrowed, not depressed; last ventrite with small round emargination in middle and archly impressed distal side.</p><p>Etymology. The toponymical specific epithet refers to the Jutland peninsula where the amber with the studied inclusion was collected.</p></div>	https://treatment.plazi.org/id/596FA515FFB1FFAD6D96C0F1FE49FD4B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tshernyshev, Sergei E.;Alekseev, Vitalii;Bukejs, Andris;Vasilenko, Dmitry V.;Legalov, Andrej A.;Perkovsky, Evgeny E.	Tshernyshev, Sergei E., Alekseev, Vitalii, Bukejs, Andris, Vasilenko, Dmitry V., Legalov, Andrej A., Perkovsky, Evgeny E. (2025): First record of the family Melyridae (Coleoptera: Cleroidea) in late Eocene European ambers with description of a new tribe, a new genus and seven new species. Zootaxa 5696 (4): 542-566, DOI: 10.11646/zootaxa.5696.4.5, URL: https://doi.org/10.11646/zootaxa.5696.4.5
