taxonID	type	description	language	source
5A19A270FF8F7439FF52FF49FE86D903.taxon	etymology	Etymology. The Latin word of “ fortis- ”, which means strong, refers to its strong mandibles and legs.	en	Jia, Ting, Liang, Hongbing, Chang, Huali, Ren, Dong (2011): A new genus and species of fossil Eodromeinae from the Yixian Formation of Western Liaoning, China (Coleoptera: Adephaga: Trachypachidae). Zootaxa 2736: 63-68, DOI: 10.5281/zenodo.203143
5A19A270FF8F7439FF52FF49FE86D903.taxon	materials_examined	Type species. Fortiseode pervalimand Jia & Ren sp. nov.	en	Jia, Ting, Liang, Hongbing, Chang, Huali, Ren, Dong (2011): A new genus and species of fossil Eodromeinae from the Yixian Formation of Western Liaoning, China (Coleoptera: Adephaga: Trachypachidae). Zootaxa 2736: 63-68, DOI: 10.5281/zenodo.203143
5A19A270FF8F7439FF52FF49FE86D903.taxon	diagnosis	Diagnosis. Head transverse, length of head capsule more than half its width; Mandibles with the same length as the head; Pronotum cordiform, widest at about anterior one-fifth of the pronotum, anterior edge of pronotum strongly concave; base of pronotum with similar width as base of elytra; length of head including mandibles nearly equal to width of head capsule; hind coxae strong, transverse; legs with strong femora and tibiae, metatarsus shorter than 1 / 2 of elytra, metatarsus short, with length nearly 1 / 2 metatarsus length; trochanters large, longer than 1 / 3 metafemur length.	en	Jia, Ting, Liang, Hongbing, Chang, Huali, Ren, Dong (2011): A new genus and species of fossil Eodromeinae from the Yixian Formation of Western Liaoning, China (Coleoptera: Adephaga: Trachypachidae). Zootaxa 2736: 63-68, DOI: 10.5281/zenodo.203143
5A19A270FF8F7439FF52FF49FE86D903.taxon	discussion	Remarks. The new genus is assigned to the family Trachypachidae based on the following characters: (1) Antennae smooth and non-pubescent; (2) Hind coxae large, and extend to the lateral margins of body. The attribution of the new genus to the subfamily Eodromeinae Ponomarenko, 1977 is based on the following characters: (1) Clypeus not extending to point of antennal attachment; (2) Forecoxal cavities open; (3) Lateral wall of midcoxal cavities formed by mesosternum, mesepimeron and metasternum; (4) Mesepisternum not extending to midcoxal cavities, minimum separation equal to mesepisternal mesal margin; (5) Hind coxae separating metasternum and abdomen, with large femoral plates. Comparisons. According to the fossil record in the subfamily Eodromeinae, there are seven genera described from Mesozoic Era: Sogdodromeus Ponomarenko, 1977; Platycoxa Ponomarenko, 1977; Unda Ponomarenko, 1977; Psacodromeus Ponomarenko, 1977; Karatoma Ponomarenko, 1977; Karadromeus Ponomarenko, 1977; Eodromeus Ponomarenko, 1977. The new genus, as compared with the genus Sogdodromeus Ponomarenko, 1977 from Triassic of Soviet Central Asia, has the following unique characters: (1) Head transverse, length of head capsule more than half its width; (2) Width of the last sternite nearly equal to that of the base of abdomen; (3) Midcoxal cavities large, oval, and separated from one another. The new genus differs from the genus Platycoxa Ponomarenko, 1977 from Late-Jurassic of Soviet Central Asia by the following features: (1) Head transverse, length of head capsule more than half its width; (2) Pronotum cordiform, widest at about anterior one-fifth of the pronotum, anterior edge of pronotum strongly concave; (3) Width of the last sternite nearly equal to that of the base of abdomen. The new genus differs from the genus Unda Ponomarenko, 1977 from Early-Cretaceous of Transbaikal by the following features: (1) Antennae shorter and thicker; (2) Pronotum cordiform, widest at about anterior one-fifth of the pronotum, anterior edge of pronotum strongly concave; (3) Last sternite anterior margin nearly equal to base of abdomen. Compared with the genus Psacodromeus Ponomarenko, 1977 from Late-Jurassic of South Kazakhstan, the new genus has a distinct teardrop-shaped body and much shorter antennae. Compared with the genus Karatoma Ponomarenko, 1977 from Late-Jurassic of South Kazakhstan, the new species has the following unique characters: (1) Elytra smooth, without large irregularly distributed punctures; (2) Antennae shorter and thicker; (3) Metasternum, distance between middle and hind coxae longer than middle coxae; (4) Length of head including mandibles nearly equal to width of head capsule. The new genus differs from the genus Karadromeus Ponomarenko, 1977 from Late-Jurassic of South Kazakhstan by the following features: (1) Pronotum cordiform, widest at about anterior one-fifth of the pronotum, anterior edge of pronotum strongly concave; (2) Hind coxae strong, transverse. Compared with the genus Eodromeus Ponomarenko, 1977 from Early Cretaceous of Trans-Baikal, the new species has these unique characters: Pronotum cordiform, widest at about anterior one-fifth of the pronotum, anterior edge of pronotum strongly concave; base of pronotum with similar width as base of elytra. According to the fossil record in the family Trachypachidae, there are two genera: Beipiaocarabus Hong, 1983 and Xinbinia Hong, 1983, described from Liaoning Province. The new genus differs from the genus Beipiaocarabus Hong, 1983 from Middle-Jurassic of Haifanggou Formation by the following features: (1) Body larger, length 10 mm, width 3.5 mm; (2) Pronotum cordiform, widest at about anterior one-fifth of the pronotum, anterior edge of pronotum strongly concave, with width 3.1 mm, posterior edge straight, width 2.1 mm, base of pronotum with similar width as base of elytra. Compared with the genus Xinbinia Hong, 1983 from Middle-Jurassic of Houjiatun Formation has these unique characters: Pronotum cordiform, widest at about anterior one-fifth of the pronotum, anterior edge of pronotum strongly concave, with width 3.1 mm, posterior edge straight, width 2.1 mm, base of pronotum with similar width as base of elytra.	en	Jia, Ting, Liang, Hongbing, Chang, Huali, Ren, Dong (2011): A new genus and species of fossil Eodromeinae from the Yixian Formation of Western Liaoning, China (Coleoptera: Adephaga: Trachypachidae). Zootaxa 2736: 63-68, DOI: 10.5281/zenodo.203143
5A19A270FF8D743EFF52FF06FC56D998.taxon	etymology	Etymology. The specific name of pervalida is derived from the Greek word of “ pervalid - ” (meaning strong), referring to the robust mandibles.	en	Jia, Ting, Liang, Hongbing, Chang, Huali, Ren, Dong (2011): A new genus and species of fossil Eodromeinae from the Yixian Formation of Western Liaoning, China (Coleoptera: Adephaga: Trachypachidae). Zootaxa 2736: 63-68, DOI: 10.5281/zenodo.203143
5A19A270FF8D743EFF52FF06FC56D998.taxon	materials_examined	Material. CNU-COL-LB- 2010495, deposited in the Key Lab of Insect Evolution & Environmental Changes, Capital Normal University, Beijing, China. Locality and horizon. Collected from 2 nd bed of Yixian Formation in Huangbanjigou, near Chaomidian Village, Shangyuan County, Beipiao City, Liaoning Province, China; Late Jurassic-Early Cretaceous.	en	Jia, Ting, Liang, Hongbing, Chang, Huali, Ren, Dong (2011): A new genus and species of fossil Eodromeinae from the Yixian Formation of Western Liaoning, China (Coleoptera: Adephaga: Trachypachidae). Zootaxa 2736: 63-68, DOI: 10.5281/zenodo.203143
5A19A270FF8D743EFF52FF06FC56D998.taxon	diagnosis	Diagnosis. Same as the genus.	en	Jia, Ting, Liang, Hongbing, Chang, Huali, Ren, Dong (2011): A new genus and species of fossil Eodromeinae from the Yixian Formation of Western Liaoning, China (Coleoptera: Adephaga: Trachypachidae). Zootaxa 2736: 63-68, DOI: 10.5281/zenodo.203143
5A19A270FF8D743EFF52FF06FC56D998.taxon	description	Description. Body small sized, length 10 mm, width 3.5 mm; elytron 5 mm long, with 8 longitudinal striae. The holotype is a dorso-ventral compression of a carabid. Antennae unisetose, scape long and robust, not completely visible from dorsal side; elytra without large pores. Head. Head sub-triangular, convex; narrower than prothorax; eyes moderately prominent; antennae inserted between eyes and base of mandibles under a frontal ridge, filiform, with eleven antennomere; antennomere 1 (scape) long and robust, antennomere 2 shorter than scape, antennomeres 6 to 10 subequal in length, apical antennomere narrower and longer; mandible falciform, robust, prominent, apices rather blunt, laterally with setae, occlusal margins toothed, length equal to that of head, molar plate visble, outer sides of mandibles not expanded; base of head retracted into pronotum. Thorax. Prothorax width nearly equal to that of paired elytra together, with lateral margins inflexed and a distinct submarginal suture between proepipleuron and propleuron; pronotum cordiform, length 2 mm, width 3.4 mm, widest at about anterior one-fifth of the pronotum, anterior edge of pronotum strongly concave, with width 3.1 mm, posterior edge straight, width 2.1 mm. base of pronotum with similar width as base of elytra; pronotum with lateral margins narrowly reflexed; forecoxal cavities open, confluent, bridged; midcoxal cavities disjunct, confluent; hind coxal cavities incomplete, confluent. Pterothorax. Scutellum triangular, visible; elytra margined laterally and basally, humeri broadly rectangular; posteriorly not sinuate, apical margin narrowly rounded, elytra broadest in the middle, with basal border, width 2.5 mm, wider than that of posterior edge of pronotum, length 6 mm, twice as long as wide, dorsal surface striate, with 8 longitudinal striae, each stria linear, striae absent laterally, surface smooth, without large pores; second elytral interval with similar width as other intervals, elytra margin with internal plica toward apex. Abdomen. Elytra completely cover the last ventrite of abdomen, abdomen with six sterna (2 – 7), sternum 2 (first visible sternum) interrupted by hind coxae, remnants visible only at sides; the last four ventrites of abdomen are much shorter compared with the first two ventrites; the first abdominal segment with length 0.7 mm, the second segment longest, with length 0.9 mm, segments 3 to 5 equal in length 0.3 mm, and the last segment length 0.6 mm. Note “ - ” indicate that it is not sure about the age. Legs. Gressorial, fairly short and thick; legs with strong femora and tibiae, metatarsus shoter than 1 / 2 of elytra, metatarsus short, with length nearly 1 / 2 of metatarsus; trochanters large, longer than 1 / 3 metafemur length, tarsi with 5 segments; tarsomere 5 terminated by pair of claws; claws with inner margins smooth; metatarsi length 2.3 mm, shorter than half of elytra, length 2.3 mm, shorter than half of elytra, nearly equal to half of metatibia; front and middle coxae globular, hind coxae dilated internally, and extended each side to lateral margins of body; metacoxae larger and transverse; metafemur much wider than metatibia; front tibia with large apical spurs, inner margin simple; hind trochanters large, longer than one-third of metafemur.	en	Jia, Ting, Liang, Hongbing, Chang, Huali, Ren, Dong (2011): A new genus and species of fossil Eodromeinae from the Yixian Formation of Western Liaoning, China (Coleoptera: Adephaga: Trachypachidae). Zootaxa 2736: 63-68, DOI: 10.5281/zenodo.203143
