taxonID	type	description	language	source
5667352AE85BFFD4609BFF06FE47FD2F.taxon	diagnosis	Diagnosis. Adults of the genus Liogluta can be distinguished from other athetine genera by combination of the following characters: body length usually more than 3.5 mm; antennomeres not distinctly transverse; α-sensillum of epipharynx long and setaceous; mandible convergent apically; prementum without lateral pseudopores; twin pores of labial palpomere 1 not small; pronotum less than 1.3 times as wide as long, pubescence directed posteriorly in midline; metatarsomeres 1 – 4 subequal in length; abdominal tergite VI not impressed in basal region (Benick & Lohse, 1974; Yosii & Sawada, 1976; Sawada, 1984).	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE85BFFD4609BFF06FE47FD2F.taxon	distribution	Distribution. Worldwide.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE85BFFD0609BFA27FF45FAB7.taxon	description	Description. Length 2.5 – 2.8 mm. Body (Fig. 1 A) parallel-sided; surface fairly glossy and densely pubescent, with microsculpture. Body dark brown to black; head almost black; pronotum and abdomen darker than elytra; legs paler, yellowish brown. Head. Subquadrate, about 1.0 – 1.1 times as wide as long, widest across eyes, slightly narrower than pronotum; eyes large and prominent, about 1.4 – 1.5 times as long as temples; gular sutures moderately separated, more or less diverged basally; infraorbital carina complete; cervical carina complete. Antennae (Fig. 3 A) long and slender; antennomeres 1 – 3 elongate, 1 longest, 2 about as long as 3, 4 – 10 subquadrate to slightly transverse, 11 longer than wide, about as long as preceding two combined. Mouthparts. Labrum (Fig. 2 A) transverse, slightly emarginate in anterior margin, with ε-sensillum and about 9 – 10 macrosetae on each side of midline; epipharynx (Fig. 2 B) with several sensilla, including 2 lateral sensory rows on each side of midline; αsensillum long and setaceous, about 2.0 times as long as ε-sensillum, β-sensillum short, convergent apically, γsensillum reduced. Mandibles (Figs. 2 C – D) asymmetrical, subtriangular, decurved and pointed apically, about 1.5 – 1.6 times as long as basal width; minute denticles present in molar region; right one (Fig. 2 C) with small internal tooth, internal margin slightly serrulate; prostheca developed, composited three portions, second portion slightly longer. Galea and lacinia of maxilla (Fig. 2 E) long and slender; lacinia having seven spines in distal comb region, contiguous with two isolated spines; maxillary palpus elongate, with pubescence and long setae; palpomere 1 smallest, 2 about 2.8 – 3.0 times as long as wide, 3 slightly longer than 2, about 2.7 – 2.9 times as long as wide, 4 digitiform, filamentous sensilla reaching to basal half. Labium (Fig. 2 F) with ligula divided into 2 lobes in basal half; prementum with two medial setae widely separated; two basal pores narrowly separated, about 1.0 – 2.0 times width of basal pore; several medial pseudopores, 1 setal pore and 2 real pores present on each side of midline; labial palpus with many setulae; palpomere 1 largest, about 2.0 times as long as wide, γ-setula contiguous with bseta, 2 shortest, about 1.4 – 1.6 times as long as wide, 3 subparallel-sided, slightly shorter than 1, about 3.0 – 3.5 times as long as wide. Mentum (Fig. 2 F) trapezoidal, anterior margin emarginate; v-seta short, close to u-seta. Thorax. Pronotum slightly transverse, approximately 1.3 times as wide as long, widest in apical third; hypomera fully visible in lateral aspect. Metanotal scutum with 1 long seta and about 4 – 5 relatively short setae on each side of midline. Mesocoxal cavities narrowly separated, mesoventral process pointed at apex, slightly longer than isthmus and metaventral process combined; isthmus slightly longer than metaventral process. Elytra longer and slightly wider than pronotum; elytron approximately 1.5 – 1.6 times as long as wide, pubescence directed posteriorly and postero-laterally; postero-lateral margin almost straight; hind wings fully developed, flabellum composed of about 5 – 6 long setose lobes. Legs. Slender and long, with pubescence and macrosetae; meso- and metatibiae with different length of two spurs at apex; tarsal formula 4 - 5 - 5, length ratio of tarsomeres 25: 27: 30: 67 (protarsus); 31: 36: 38: 39: 68 (mesotarsus); 48: 46: 44: 42: 80 (metatarsus); one empodial seta present, about as long as claw. Abdomen. Parallel-sided; surface distinctly glossy and densely pubescent, with reticulate microsculpture; macrochaetal arrangement of tergites II – VI 01 - 21 - 13 - 13 - 13; male tergite VIII (Fig. 3 B) with 4 macrosetae on each side of midline; broad process present in median region and posterior margin crenate; male sternite VIII (Fig. 3 C) with 9 macrosetae on each side of midline, posterior margin slightly convex, subtriangular, with long marginal setae; posterior margin of female tergite VIII (Fig. 3 D) subtruncate; female sternite VIII (Fig. 3 E) with 7 macrosetae, posterior margin slightly emarginate in median region, with conspicuous marginal setae and minute setae. Aedeagus. Median lobe (Figs. 3 F – G) narrowly ovate and widest in basal fourth, apical process elongate and convergent at apex in ventral aspect; internal sac developed. Apical lobe of paramerites (Fig. 3 H) with four setae; a-seta slightly longer than b- and d-setae subequal in length, c-seta very short and close to d-seta. Spermatheca. Bursa dilated apically and conical shaped umbilicus; duct loosely coiled (Fig. 3 I).	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE85BFFD0609BFA27FF45FAB7.taxon	materials_examined	Type material. Holotype, ♂, labeled as follows: ‘ KOREA: Chungnam Prov., Daejeon-si, Seo-gu, Jangandong, Mt. Jangtaesan, N 36 ° 13 ′ 03.3 ″ E 127 ° 20 ′ 36.2 ″ 258 m, 28 III 2012, DH Lee, TK Kim, SG Lee; HOLOTYPE Liogluta changwhani Lee and Ahn 2016 ’. Desig. S. - G. Lee and K. - J. Ahn 2016. Paratypes, 40 exx. (total): 19 exx. (one on slide), same data as Holotype; 21 exx. (two on slide), Korea. Gyeongbuk prov., Yongjang-ri, Naenammyeon, Gyeongju-si, ex FIT 21. V- 26. VI. 2007, YB Cho coll. E 129 ° 12 ′ 42.9 ″ N 35 ° 46 ′ 19.5 ″. Material examined. SOUTH KOREA: Chungnam prov.: 11 exx. (five in 95 % ETOH), Daejeon-si, Seo-gu, Jangan-dong, Jangtaesan Recreational Forest, N 36 ° 13 ′ 4.32 ″ E 127 ° 20 ′ 34.44 ″ 257 m, 17 III 2011, IS Yoo, YH Kim, SG Lee, leaf litters.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE85BFFD0609BFA27FF45FAB7.taxon	distribution	Distribution. Korea (South).	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE85BFFD0609BFA27FF45FAB7.taxon	discussion	Remarks. Adults are similar to those of L. pyonganica, but can be distinguished by the characters provided in the key and different shape and structure of aedeagus and spermatheca.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE85BFFD0609BFA27FF45FAB7.taxon	etymology	Etymology. Named after the late professor Chang-Whan Kim in honor of his pioneering research on Korean insects.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE85FFFD2609BFA84FAA6F906.taxon	description	Description. Length 2.5 – 3.3 mm. Body (Fig. 1 B) parallel-sided; surface distinctly glossy and densely pubescent, with microsculpture. Body brown to reddish brown; head and abdominal segments V – VIII darker than other parts; legs paler, yellowish brown. Head. Subquadrate, approximately 1.0 – 1.1 times as wide as long, widest across eyes, slightly narrower than pronotum; eyes large and prominent, about 1.5 times as long as temples; gular sutures moderately separated, diverged basally; infraorbital carina complete; cervical carina complete. Antennae (Fig. 4 A) long and slender; antennomeres 1 – 3 elongate, 1 longest, 2 slightly longer than 3, 4 – 10 subquadrate to slightly transverse, 11 about as long as 1, about as long as preceding two combined. Mouthparts. Labrum transverse, slightly emarginate in anterior margin, with ε-sensillum and about 9 macrosetae on each side of midline; epipharynx with several sensilla, including 2 lateral sensory rows on each side of midline; α-sensillum long and setaceous, about 2.0 times as long as ε-sensillum, β- and γ-sensilla very short. Mandibles asymmetrical, subtriangular, decurved and pointed apically, about 1.7 times as long as basal width; minute denticles present in molar region; right one with small internal tooth, internal margin slightly serrulate; prostheca developed, composited three portions. Galea and lacinia of maxilla long and slender; lacinia having seven spines in distal comb region, contiguous with two isolated spines; maxillary palpus elongate, with pubescence and long setae; palpomere 1 smallest, 2 about 2.6 – 2.8 times as long as wide, 3 slightly longer than 2, about 2.7 – 2.8 times as long as wide, 4 digitiform, filamentous sensilla reaching to basal half. Labium with ligula elongate, divided into 2 lobes at base; prementum with two medial setae widely separated; two basal pores moderately separated, about 2.0 times width of basal pore; several medial pseudopores, 1 setal pore and 2 real pores present on each side of midline; labial palpus with many setulae; palpomere 1 largest, about 2.0 times as long as wide, γ-setula contiguous with bseta, 2 shortest, about 1.4 – 1.5 times as long as wide, 3 parallel-sided and about as long as 1, about 3.0 times as long as wide. Mentum trapezoidal, anterior margin slightly emarginate; v-seta very short, more or less equal distant from u- and w-setae. Thorax. Pronotum slightly transverse, approximately 1.2 – 1.3 times as wide as long, widest in apical third; hypomera fully visible in lateral aspect. Metanotal scutum with 2 long setae and about 3 – 6 relatively short setae on each side of midline. Mesocoxal cavities moderately separated, mesoventral process slightly pointed at apex, about as long as or slightly longer than isthmus and metaventral process combined. Elytra longer and slightly wider than pronotum; elytron approximately 1.5 times as long as wide, pubescence directed posteriorly and postero-laterally; postero-lateral margin straight; hind wings fully developed, flabellum composed of about 7 – 9 long setose lobes. Legs. Slender and long, with pubescence and macrosetae; meso- and metatibiae with different length of two spurs at apex; tarsal formula 4 - 5 - 5, length ratio of tarsomeres 24: 27: 30: 74 (protarsus); 34: 40: 42: 40: 74 (mesotarsus); 55: 50: 48: 48: 92 (metatarsus); one empodial seta present, slightly shorter than claw. Abdomen. Parallel-sided; surface distinctly glossy and densely pubescent, with imbricate microsculpture; macrochaetal arrangement of tergites II – VI 01 - 13 - 13 - 13 - 13; male tergite VIII (Fig. 4 B) with 4 macrosetae on each side of midline, broad process present in median region and posterior margin slightly crenate; male sternite VIII (Fig. 4 C) with 9 macrosetae on each side of midline, posterior margin convex, slightly round with long marginal setae; posterior margin of female tergite VIII (Fig. 4 D) subtruncate; female sternite VIII (Fig. 4 E) with 7 macrosetae on each side of midline, posterior margin broadly round, with long and short marginal setae, minute setae in median region. Aedeagus. Median lobe (Figs. 4 F – G) narrowly ovate and widest in basal fourth, apical process convergent at apex in ventral aspect; internal sac developed. Apical lobe of paramerites (Fig. 4 H) with four setae; a-seta longest, c- and d-setae shorter than b-seta, close together and positioned apically. Spermatheca. Reduced.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE85FFFD2609BFA84FAA6F906.taxon	materials_examined	Type material. Lectotype, ♂, labeled as follows: ‘ Type, Japan, Japan G. Lewis, Homalota distans Type D. S., Sharp Coll. 1905 - 313., LECTOTYPE Homalota distans Sharp des. G. Paśnik 2005 ’. Paralectotypes, 10 exx., same data as Lectotypes. Material examined. SOUTH KOREA: Chungnam prov.: 1 ex., Buyeo-gun, Mt. Mansusan, 19 – 20 vii 1999, US Hwang, HJ Kim, FIT; 16 exx., Buyeo-gun, Naesan-myeon, Mt. Wolmyeongsan, Geumgisa, 3 v – 1 vi 2000, US Hwang, HJ Kim, FIT; 2 exx., Daejeon-si, Yuseong-gu, Chungnam nat. univ., 15 v – 4 vi 2003, JH Choi, DH Lee, SM Choi, FIT; 75 exx. (two on slide), Gongju-si, Banpo-myeon, Mt. Gyeryongsan, Woosanbong, 9 v 2000, SJ Park, MS Kim, FIT; 8 exx. (one on slide), same data as former except for ‘ 3 v 2000, US Hwang, SJ Park, HJ Kim, sifting’; 1 ex., Gyeryongsan, Donghaksa, 3 v 1998, KJ Ahn, JH Ahn, on rocks; 2 exx., Nonsan-si, Beolgok-myeon, Mt. Daedunsan, Surak-valley, 2 – 10 V 2000, US Hwang, SJ Park, HJ Kim, FIT. Gangwon Prov.: 1 ex., Pyeongchang-gun, Jinbu-myeon, Mt. Odaesan, Sangwonsa, 22 viii – 20 x 2000, KJ Ahn, FIT; Gyeongbuk Prov.: 1 ex., Ulleung-gun, Taehwa-ri, 17 x 2001, JW Lee; 1 ex., Ulleung-gun, Namyangcheon, 17 x 2001, JW Lee, near stream; 2 exx., Ulleung-gun, Naribunji, 18 x 2001, JW Lee. Jeonbuk Prov.: Jeongeup-si Mt. Naejangsan, Naejangsa, Geumseon valley, 15 vi – 24 vi 2000, US Hwang, HJ Kim, FIT. Jeonnam Prov.: 13 exx. (one on slide), Gurye-gun, Mt. Jirisan, Piagol, 24 – 26 v 2000, HK Choi, JH Song, MS Kim, bait trap.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE85FFFD2609BFA84FAA6F906.taxon	distribution	Distribution. Korea (South) and Japan.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE85FFFD2609BFA84FAA6F906.taxon	discussion	Remarks. Previously known only from Japan, this species is recorded for the first time in Korea.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE85DFFDD609BF8D4FEA0FB2C.taxon	description	Description. Length 3.5 – 4.0 mm. Body (Fig. 1 C) subparallel-sided; surface distinctly glossy and densely pubescent, with fine microsculpture. Body reddish brown; head darker than other parts; elytra and legs paler, yellowish brown to brown. Head. Subquadrate, about as wide as long, widest at middle, slightly narrower than pronotum; eyes moderate in size and slightly prominent, about as long as temples; gular sutures moderately separated, diverged basally; infraorbital carina incomplete. Antennae long and slender, longer than head and pronotum combined; antennomeres 1 – 3 elongate, 1 about as long as 1, 2 slightly shorter than 3, 4 about as long as wide, 5 – 10 subquadrate, 11 longest, about as long as preceding two combined. Thorax. Pronotum slightly transverse, approximately 1.2 times as wide as long, widest in apical third to half, with longitudinally slight impression in median region; pubescence directed posteriorly in midline; hypomera fully visible in lateral aspect. Elytra slightly transverse, slightly wider than pronotum; elytron approximately 1.7 times as long as wide, pubescence directed posteriorly and postero-laterally; postero-lateral margin almost straight; hind wings fully developed. Legs. Slender and long, with pubescence and macrosetae; tibiae with two spurs at apex; tarsal formula 4 - 5 - 5; one empodial seta present, shorter than claw. Abdomen. Subparallel-sided, widest at middle; surface distinctly glossy and densely pubescent; male tergite VIII with 4 macrosetae on each side of midline, posterior margin with broad and pentagon process; posterior margin of male sternite VIII subtruncate; posterior margin of female tergite VIII slightly convex, subtriangular; female sternite VIII with 7 macrosetae on each side of midline, posterior margin distinctly emarginate, with long and short marginal setae. Aedeagus. Median lobe narrowly ovate, apical process subtriangular and pointed apically in ventral aspect. Spermatheca. Bursa subglobular at apex; duct relatively slender and deeply recurved.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE85DFFDD609BF8D4FEA0FB2C.taxon	materials_examined	Material examined. NORTH KOREA: 3 exx., Korea 21 - 25. 5. 74 pr. Ćhongdźin-si Exp. Inst. Zool Cr. [North Korea, Hamgyeongbuk Prov., Cheongjin-si, 21 - 25. v 1974, ISEA]; 4 exx., Korea Rampote-sn, distr. Samdż lg. Pawlowski 9. 71 [North Korea, Yanggang Prov., Mt. Nampodaesan, IX 1971, J. Pawlowski].	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE85DFFDD609BF8D4FEA0FB2C.taxon	distribution	Distribution. Korea (North), Russia (East Siberia, West Siberia) and Europe.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE85DFFDD609BF8D4FEA0FB2C.taxon	discussion	Remarks. This species was recorded by Paśnik (2001) in North Korea and no specimen has been collected from South Korea. We could not describe the mouthparts in detail because no specimens were available for dissection.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE852FFDC609BFB32FEA0FEDF.taxon	description	Description. Length about 3.5 mm. Body (Fig. 1 D) relatively slender and subparallel-sided; surface distinctly glossy and densely pubescent, with fine microsculpture. Body yellowish brown to reddish brown; head and abdomen slightly darker than other parts. Head. Subquadrate, about as wide as long, widest behind eyes, slightly narrower than pronotum; eyes small and slightly prominent, about 0.7 – 0.8 times as long as temples; gular sutures moderately separated, diverged basally; infraorbital carina incomplete. Antennae long and slender, longer than head and pronotum combined; antennomeres 1 – 3 elongate, 1 about as long as 11, 2 slightly shorter than 3, 4 about as long as wide, 5 – 10 subquadrate to slightly transverse, 11 longest, about as long as preceding two combined. Thorax. Pronotum slightly transverse, approximately 1.1 – 1.2 times as wide as long, widest at middle, with longitudinally slight impression in median region; pubescence directed posteriorly in midline; hypomera fully visible in lateral aspect. Elytra slightly transverse, slightly wider than pronotum; elytron approximately 1.8 times as long as wide, pubescence directed posteriorly and postero-laterally; postero-lateral margin almost straight; hind wings fully developed. Legs. Slender and long, with pubescence and macrosetae; tibiae with two spurs at apex; tarsal formula 4 - 5 - 5; one empodial seta present, shorter than claw. Abdomen. Subparallel-sided, widest at middle; surface distinctly glossy and densely pubescent; male tergite VIII with 4 macrosetae on each side of midline, posterior margin with broad and slightly crenate process; posterior margin of male sternite VIII broadly round. Aedeagus. Median lobe narrowly ovate in ventral aspect; apical process long and bent in lateral aspect.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE852FFDC609BFB32FEA0FEDF.taxon	materials_examined	Material examined. NORTH KOREA: 2 ♂♂, Korea 16 - 18.6.74. Kymgang-san Mts. Exp. Inst. Zool. Cr. [North Korea, Gangwon Prov., Mt. Geumgangsan, 16 – 18 VI 1974, ISEA].	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE852FFDC609BFB32FEA0FEDF.taxon	distribution	Distribution. Korea (North) and Russia (East Siberia).	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE852FFDC609BFB32FEA0FEDF.taxon	discussion	Remarks. This species was recorded by Paśnik (2001) in North Korea and no specimen has been collected from South Korea. We could not describe the mouthparts in detail because no specimens were available for dissection.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE853FFDF609BFE6CFA21FE32.taxon	description	Description. Length 2.5 – 2.7 mm. Body (Fig. 1 E) parallel-sided; surface fairly glossy and densely pubescent, with microsculpture. Body reddish black to black; head and abdomen darker than other parts, almost black; antennae and legs paler, reddish brown. Head. Subquadrate, about 1.0 – 1.1 times as wide as long, widest across eyes, slightly narrower than pronotum; eyes moderate in size and slightly prominent, about 1.0 – 1.2 times as long as temples; gular sutures moderately separated, more or less diverged basally; infraorbital carina almost complete; cervical carina complete. Antennae (Fig. 6 A) long and slender; antennomeres 1 – 3 elongate, 1 longest, 2 slightly longer than 3, 4 about as long as wide, 5 – 10 subquadrate to slightly transverse, 11 longer than wide, about as long as preceding two combined. Mouthparts. Labrum (Fig. 5 A) transverse, slightly emarginate in anterior margin, with ε-sensillum and about 8 – 9 macrosetae on each side of midline; epipharynx (Fig. 5 B) with several sensilla, including 2 lateral sensory rows on each side of midline; α-sensillum long and setaceous, about 2.0 times as long as ε-sensillum, β- and c-sensilla very short. Mandibles (Figs. 5 C – D) asymmetrical, subtriangular, decurved and pointed apically, about 1.6 – 1.7 times as long as basal width; minute denticles present in molar region; right one (Fig. 5 C) with small internal tooth, internal margin slightly serrulate; prostheca developed, composited three portions. Galea and lacinia of maxilla (Fig. 5 E) long and slender; lacinia composited seven spines in distal comb region, contiguous with two isolated spines; maxillary palpus elongate, with pubescence and long setae; palpomere 1 smallest, 2 about 2.6 – 2.8 times as long as wide, 3 slightly longer than 2, about 2.6 – 2.8 times as long as wide, 4 digitiform, filamentous sensilla reaching to basal half. Labium (Fig. 5 F) with ligula relatively short, divided into 2 lobes in basal half; prementum with two medial setae widely separated; two basal pores moderately separated, about 2.0 – 3.0 times width of basal pore; several medial pseudopores, 1 setal pore and 2 real pores present on each side of midline; labial palpus with many setulae; palpomere 1 largest, about 1.8 – 2.0 times as long as wide, γ-setula contiguous with b-seta, 2 shortest, about 1.4 – 1.6 times as long as wide, 3 subparallel-sided, about as long as 1, about 3.5 times as long as wide. Mentum (Fig. 5 F) trapezoidal, anterior margin almost straight; v-seta short, close to u-seta. Thorax. Pronotum slightly transverse, approximately 1.2 – 1.3 times as wide as long, widest in apical third; hypomera fully visible in lateral aspect. Metanotal scutum with 2 long setae and about 2 – 4 relatively short setae on each side of midline. Mesocoxal cavities narrowly separated, mesoventral process slightly pointed at apex, length ratio of mesoventral process, isthmus and metaventral process 16: 9: 5. Elytra longer and slightly wider than pronotum; elytron approximately 1.6 times as long as wide, pubescence directed posteriorly and postero-laterally; posterolateral margin slightly sinuate; hind wings fully developed, flabellum composed of about 7 – 9 long setose lobes. Legs. Slender and long, with pubescence and macrosetae; meso- and metatibiae with different length of two spurs at apex; tarsal formula 4 - 5 - 5, length ratio of tarsomeres 27: 30: 33: 68 (protarsus); 35: 39: 39: 36: 70 (mesotarsus); 52: 53: 51: 50: 87 (metatarsus); one empodial seta present, slightly shorter than claw. Abdomen. Parallel-sided; surface distinctly glossy and densely pubescent, with transversely reticulate microsculpture; macrochaetal arrangement of tergites II – VI 01 - 12 - 22 - 23 - 23; male tergite VIII (Fig. 6 B) with 4 macrosetae on each side of midline, posterior margin subtruncate; male sternite VIII (Fig. 6 C) with 9 macrosetae on each side of midline, posterior margin round, with long marginal setae; posterior margin of female tergite (Fig. 6 D) VIII subtruncate; female sternite VIII (Fig. 6 E) with 7 macrosetae on each side of midline, posterior margin more or less triangularly convex, with conspicuous marginal setae, minute setae present in median region. Aedeagus. Median lobe (Figs. 6 F – G) ovate and widest at middle, apical process broad and subtriangular, convergent at apex in ventral aspect; internal sac developed. Apical lobe of paramerites (Fig. 6 H) with four setae; a-seta longest, slightly longer than bseta, c- and d-setae short and subequal in length, contiguous and positioned at apex. Spermatheca. Bursa round at apex, with broad umbilicus; duct slightly sinuate, shortly coiled at apex (Fig. 6 I).	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE853FFDF609BFE6CFA21FE32.taxon	materials_examined	Type material. Holotype, ♂, labeled as follows: ‘ KOREA: Gangwon Prov., Pyeongchang-gun, Jinbu-myeon, Mt. Odaesan, Sangwonsa, N 37 ° 47 ' 10 " E 128 ° 33 ' 57.2 " 10 V 2007, TK Kim, sifting; HOLOTYPE Liogluta jinilli Lee and Ahn 2016 ’. Desig. S. - G. Lee and K. - J. Ahn 2016. Paratypes, 10 exx. (two on slide), same data as Holotype.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE853FFDF609BFE6CFA21FE32.taxon	distribution	Distribution. Korea (South).	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE853FFDF609BFE6CFA21FE32.taxon	discussion	Remarks. This species is similar to L. changwhani sp. nov. and L. pyonganica, but can be distinguished by the characters provided in the key and different shape and structure of aedeagus.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE853FFDF609BFE6CFA21FE32.taxon	etymology	Etymology. Named after the late professor Jin Ill Kim in honor of his pioneering research on Korean beetles.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE856FFDB609BFF4CFE85FCC5.taxon	description	Description. Length 3.0 – 3.8 mm. Body (Fig. 1 F) parallel-sided; surface glossy and densely pubescent, with microsculpture. Body reddish brown to black; head, pronotum, abdomen almost black; elytra and legs paler than other parts. Head. Subquadrate, about as wide as long, widest across eyes, slightly narrower than pronotum; eyes slightly large and prominent, about 1.3 – 1.4 times as long as temples; gular sutures moderately separated; infraorbital carina complete; diverged basally; cervical carina complete. Antennae (Fig. 7 A) long and slender; antennomeres 1 – 3 elongate, 1 longest, 2 about as long as 3, 4 – 10 quadrate to subquadrate, 11 about as long as 1, about as long as preceding two combined. Mouthparts. Labrum transverse, slightly emarginate in anterior margin, with ε-sensillum and about 9 macrosetae on each side of midline; epipharynx with several sensilla, including 2 lateral sensory rows on each side of midline; α-sensillum long and setaceous, about 2.0 times as long as εsensillum, β- and γ-sensilla very short. Mandibles asymmetrical, subtriangular, decurved and pointed apically, about 1.5 – 1.6 times as long as basal width; minute denticles present in molar region; right one with small internal tooth, internal margin slightly serrulate; prostheca developed, composited three portions. Galea and lacinia of maxilla long and slender; lacinia composited seven spines in distal comb region, contiguous with two isolated spines; maxillary palpus elongate, with pubescence and long setae; palpomere 1 smallest, 2 about 2.4 – 2.6 times as long as wide, 3 slightly longer than 2, about 2.8 – 3.0 times as long as wide, 4 digitiform, filamentous sensilla reaching to basal half. Labium with ligula divided into 2 lobes in basal half; prementum with two medial setae widely separated; two basal pores moderately separated, about 1.0 – 2.0 times width of basal pore; several medial pseudopores, 1 setal pore and 2 real pores present on each side of midline; labial palpus with many setulae; palpomere 1 largest, about 1.7 – 1.9 times as long as wide, γ-setula contiguous with b-seta, 2 shortest, about 1.2 – 1.4 times as long as wide, 3 subparallel-sided and about as long as 1, about 3.0 – 3.5 times as long as wide. Mentum trapezoidal, anterior margin slightly emarginate; v-seta very short, close to u-seta. Thorax. Pronotum slightly transverse, approximately 1.1 – 1.2 times as wide as long, widest in apical third to half; hypomera fully visible in lateral aspect. Metanotal scutum with 2 long setae and about 3 – 4 relatively short setae on each side of midline. Mesocoxal cavities narrowly separated, mesoventral process pointed at apex, slightly longer than isthmus and metaventral process combined; isthmus slightly longer than metaventral process. Elytra transverse, slightly wider than pronotum; elytron approximately 1.5 – 1.6 times as long as wide, pubescence directed posteriorly and posterolaterally; postero-lateral margin almost straight; hind wings fully developed, flabellum composed of about 9 – 11 long setose lobes. Legs. Slender and long, with pubescence and macrosetae; meso- and metatibiae with different length of two spurs at apex; tarsal formula 4 - 5 - 5, length ratio of tarsomeres 33: 37: 39: 89 (protarsus); 39: 48: 48: 45: 89 (mesotarsus); 71: 69: 61: 56: 106 (metatarsus); one empodial seta present, slightly shorter than claw. Abdomen. Parallel-sided; surface distinctly glossy and densely pubescent, with transverse and imbricate microsculpture; macrochaetal arrangement of tergites II – VI 01 - 23 - 23 - 23 - 23; male tergite VIII (Fig. 7 B) with 4 – 5 macrosetae on each side of midline, posterior margin slightly crenate in median region; male sternite VIII (Fig. 7 C) with 9 – 10 macrosetae on each side of midline, posterior margin convex, slightly round, with long marginal setae; female tergite VIII (Fig. 7 D) with 4 macrosetae on each side of midline, posterior margin subtruncate; female sternite VIII (Fig. 7 E) with 7 macrosetae on each side of midline, posterior margin broadly round, with conspicuous marginal setae, minute setae present in median region. Aedeagus. Median lobe (Figs. 7 F – G) narrowly ovate and widest in basal fourth, apical process constricted basally and convergent at apex in ventral aspect; internal sac elongate. Apical lobe of paramerites (Fig. 7 H) with four setae; a-seta longest, c- and d-setae shorter than b-seta, close together and positioned apically. Spermatheca. Bursa globular apically, with small umbilicus; duct simply coiled (Fig. 7 I).	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE856FFDB609BFF4CFE85FCC5.taxon	materials_examined	Type material. Holotype, ♂, labeled as follows: ‘ Korea 1981 Vaudo A. Szeptycki; HOLOTYPE Liogluta pyonganica sp. n. Paśnik det. G. Paśnik 2000 ’ [North Korea, Pyeongannam Prov., Nampo-si, Waudo, 1981, A. Szeptycki]. Paratypes, 4 exx., same data as Holotype. Material examined. SOUTH KOREA: Chungbuk Prov.: 13 exx., Danyang-gun, Mt. Sobaeksan, Cheongdong, 7 – 9 v 1999, US Hwang, HJ Kim, FIT. Chungnam Prov.: 1 ex., Buyeo-gun, Naesan-myeon, Mt. Wolmyeongsan, Geumjisa, 3 v – 1 vi 2000, US Hwang, HJ Kim, FIT; 18 exx., Nonsan-si, Beolgok-myeon, Mt. Daedunsan, Surakvalley, 2 – 10 V 2000, US Hwang, SJ Park, HJ Kim, FIT; 1 ex., Gongju-si, Banpo-myeon, Mt. Gyeryongsan, Woosanbong, 10 – 24 vii 2000, SJ Park, MS Kim, FIT; 1 ex., Gongju-si, Mt. Gyeryongsan, Eunseon waterfall, N 36 ° 20 ′ 58.7 ″ E 127 ° 12 ′ 41.3 ″, 1 – 18 vi 2004; SM Choi, JS Park; FIT; 1 ex., same data as the former except for ‘ Nammaetap, N 36 ° 21 ′ 11.8 ″ E 127 ° 13 ′ 20.8 ″ ’; 7 exx., Gongju-si, Sangsin-ri, Mt. Gyeryongsan, N 36 ° 22 ′ 03.2 ″ E 127 ° 12 ′ 50 ″, 31 v – 18 vi 2004, SM Choi, JS Park, FIT. Gangwon Prov.: 2 exx., Hongcheon-gun, Naechon-myeon, Mt. Baekamsan, Garyeong fall, 25 v – 20 vi 2002, KJ Ahn, SJ Park, JS Park, FIT; 1 ex., Hongcheon-gun, Naemyeon, Mt. Gyebangsan, Unduryeong, N 37 ° 42.49.9 ′ E 128 ° 26.40.3 ′, 1100 m, 11 v 2007, TK Kim, YH Kim, fungus on log; 53 exx., Inje-gun, Mt. Jeombongsan, 19 iv – 20 vi 2007, FIT; 7 exx., Pyeongchang-gun, Jinbu-myeon, Dongsan-ri, Mt. Odaesan, Duroryeong, N 37 ° 48.713 ’ E 128 ° 38.422 ’, 16 v 2006 DH Lee, YH Kim, sifting, leaf litter; 105 exx., Pyeongchang-gun, Jinbu-myeon, Dongsan-ri, Mt. Odaesan, Sangwonsa, 30 iv – 4 vi 2001, KJ Ahn, SJ Park, MS Kim, MJ Jeon, FIT; 6 exx., same data as former except for ‘ 4 – 22 vi 2001 ’; 26 exx., same data as former except for ‘ 21 iv – 18 v 2002, SJ Park, CW Shin, FIT’; 3 exx., same data as former except for ‘ 18 vi 2004, SJ Park’; 3 exx., same data as former except for ‘ 8 v 2004, DH Lee, SM Choi, sifting’; 1 ex., same data as former except for ‘ 18 vi – 22 vii 2004, SJ Park, KM Yang, DH Lee, FIT’; 19 exx., same data as former except for ‘ N 37 ° 47.074 ’ E 128 ° 33.735 ’, 15 v 2006, TK Kim, HW Kim, sifting, leaf litter’; 132 exx., same data as former except for, ‘ N 37 ° 47 ' 10 " E 128 ° 33 ' 57.2 ", 19 iv – 11 v 2007, TK Kim, YH Kim, FIT’; 61 exx., Pyeongchang-gun, Jinbu-myeon, Mt. Odaesan, Woljeongsa, 22 viii – 20 x 2000, KJ Ahn, FIT; 2 exx., Pyeongchang-gun, Mt. Odaesan, 25 v 2004, SJ Park, JS Park’; 26 exx., same data as former except for ‘ N 37 ˚ 47 ’ 03 ˝ E 128 ˚ 33 ’ 55 ˝, 9 – 25 v 2004, SJ Park, SM Choi, DH Lee, FIT’; 21 exx., Pyeongchang-gun, Mt. Odaesan, Jeokmyeolbogung, 7 – 9 x 1999, US Hwang, MS Kim, FIT’; 1 ex., same data as former except for ‘ US Hwang, sifting’. Jeonbuk Prov.: 2 exx., Jeongeup-si, Mt. Naejangsan, Naejangsa, Geumseon valley, 15 – 24 vi 2000, US Hwang, HJ Kim, FIT. Jeonnam Prov.: 1 ex., Gurye-gun, Mt. Jirisan, Piagol, 24 v 2000, HJ Kim, MJ Jeon, sifting; 1 ex., Jangseong-gun, Jangseongeup, Yutang-ri, N 35 ° 18 ′ 54.8 ″ E 126 ° 48 ′ 34.0 ″ 90 m, 22 v 2007, TK Kim, YH Kim, dung.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE856FFDB609BFF4CFE85FCC5.taxon	distribution	Distribution. Korea (South, North).	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE856FFDB609BFF4CFE85FCC5.taxon	discussion	Remarks. This species was first recorded by Paśnik (2001) in North Korea and is recorded for the first time in South Korea.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE854FFC5609BFC0FFA82FEA7.taxon	description	Description. Length 2.0 – 2.3 mm. Body (Fig. 1 G) parallel-sided; surface fairly glossy and densely pubescent, with microsculpture. Body reddish brown; head reddish black; elytra and legs paler, yellowish brown; abdominal segments V – VIII darker than other segments. Head. Subquadrate, approximately 1.0 – 1.1 times as wide as long, widest across eyes, slightly narrower than pronotum; eyes moderate in size and slightly prominent, about 1.0 – 1.2 times as long as temples; gular sutures moderately separated, diverged basally; infraorbital carina complete; cervical carina complete. Antennae (Fig. 8 A) long and slender; antennomeres 1 – 3 elongate, 1 longest, 2 slightly longer than 3, 4 – 10 subquadrate to slightly transverse, 11 longer than wide, about as long as preceding two combined. Mouthparts. Labrum transverse, emarginate in anterior margin, with ε-sensillum and about 9 macrosetae on each side of midline; epipharynx with several sensilla, including 2 lateral sensory rows on each side of midline; α-sensillum long and setaceous, about 2.0 times as long as ε-sensillum, β- and γ-sensilla very short. Mandibles asymmetrical, subtriangular, decurved and pointed apically, about 1.5 – 1.6 times as long as basal width; minute denticles present in molar region; right one with small internal tooth, internal margin slightly serrulate; prostheca developed, composited three portions. Galea and lacinia of maxilla long and slender; lacinia composited seven spines in distal comb region, contiguous with two isolated spines; maxillary palpus elongate, with pubescence and long setae; palpomere 1 smallest, 2 about 2.5 – 2.7 times as long as wide, 3 slightly longer than 2, about 2.4 – 2.6 times as long as wide, 4 digitiform, filamentous sensilla not reaching to basal half. Labium with ligula elongate, divided into 2 lobes in basal half; prementum with two medial setae widely separated; two basal pores moderately separated, about 2.0 times width of basal pore; several medial pseudopores, 1 setal pore and 2 real pores present on each side of midline; labial palpus with many setulae; palpomere 1 largest, about 1.5 – 1.7 times as long as wide, γ-setula contiguous with b-seta, 2 shortest, about 1.2 – 1.4 times as long as wide, 3 parallelsided and about as long as 1, about 3.0 times as long as wide. Mentum trapezoidal, anterior margin almost straight; v-seta relatively long, close to u-setae. Thorax. Pronotum slightly transverse, approximately 1.2 – 1.3 times as wide as long, widest in apical third; hypomera fully visible in lateral aspect. Metanotal scutum with 1 long seta and about 2 relatively short setae on each side of midline. Mesocoxal cavities moderately separated, mesoventral process pointed at apex, slightly longer than isthmus and metaventral process combined; isthmus about as long as metaventral process. Elytra longer and slightly wider than pronotum; elytron approximately 1.4 – 1.5 times as long as wide, pubescence directed posteriorly and postero-laterally; postero-lateral margin straight; hind wings fully developed, flabellum composed of about 6 long setose lobes. Legs. Slender and long, with pubescence and macrosetae; tibiae with different length of two spurs at apex; tarsal formula 4 - 5 - 5, meso- and metatarsomere 1 – 4 subequal in length; one empodial seta present, shorter than claw. Abdomen. Parallel-sided; surface fairly glossy and densely pubescent, with imbricate microsculpture; macrochaetal arrangement of tergites II – VI 01 - 13 - 13 - 13 - 13; male tergite VIII (Fig. 8 B) with 4 macrosetae on each side of midline, broad process present in median region and posterior margin denticulate; male sternite VIII (Fig. 8 C) with 9 macrosetae on each side of midline, posterior margin slightly convex, with inconspicuous marginal setae; posterior margin of female tergite VIII subtruncate; posterior margin of female sternite VIII slightly emarginate, with conspicuous and long marginal setae, minute setae in median region. Aedeagus. Median lobe (Figs. 8 D – E) narrowly ovate and widest in basal fourth, apical process elongate and parallel-sided, convergent at apex in ventral aspect; internal sac developed. Apical lobe of paramerites (Fig. 8 F) with four setae; a-seta longest, c- and d-setae shorter than b-seta, close together and positioned apically.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE854FFC5609BFC0FFA82FEA7.taxon	materials_examined	Type material. Holotype, ♂, labeled as follows: ‘ KOREA: Seoul, Dobong-gu, Mt. Bukhansan, 24 III 1988, Y. S. Kim, ex leaf litter; HOLOTYPE Liogluta rufescens Lee and Ahn 2016 ’. Desig. S. - G. Lee and K. - J. Ahn 2016. Paratypes, 3 exx. (one on slide), same data as Holotype.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE854FFC5609BFC0FFA82FEA7.taxon	distribution	Distribution. Korea (South).	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE854FFC5609BFC0FFA82FEA7.taxon	discussion	Remarks. Adults are very similar to those of L. distans, but can be distinguished by the characters provided in the key and different shape and structure of aedeagus.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE854FFC5609BFC0FFA82FEA7.taxon	etymology	Etymology. Named from the Latin rufescens meaning ‘‘ red, reddish ” which refers to the body color.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE84AFFC7609BFEB4FDFCFED9.taxon	description	Description. Length 2.8 – 3.4 mm. Body fusiform; surface fairly glossy and densely pubescent, with fine microsculpture. Body (Fig. 1 H) reddish brown to dark brown; head darker than other parts; elytra and legs paler brown. Head. More or less circular, approximately 1.0 – 1.1 times as wide as long, widest across eyes, distinctly narrower than pronotum; eyes relatively small, about 0.8 – 1.0 times as long as temples; gular sutures moderately separated, diverged basally; infraorbital carina complete; cervical carina absent. Antennae (Fig. 9 A) long and slender; antennomeres 1 – 3 elongate, 2 slightly longer than 3, 4 about as long as wide, 5 – 10 subquadrate to slightly transverse, 11 longest, about as long as preceding two combined. Mouthparts. Labrum transverse, slightly emarginate in anterior margin, with ε-sensillum and about 8 macrosetae on each side of midline; epipharynx with several sensilla, including 2 lateral sensory rows on each side of midline; α-sensillum setaceous, slightly longer than ε-sensillum, β- and γ-sensilla very short, convergent at apex. Mandibles asymmetrical, subtriangular, decurved and pointed apically, about 1.6 times as long as basal width; right one with small internal tooth, internal margin slightly serrulate; prostheca developed, composited three portions, second portion longer. Galea and lacinia of maxilla long and slender; lacinia having seven spines in distal comb region, two isolated spines present; maxillary palpus elongate with pubescence and long setae; palpomere 1 smallest, 2 about 2.8 – 3.0 times as long as wide, 3 slightly longer than 2, about 3.0 times as long as wide, 4 digitiform, filamentous sensilla not reaching to basal half. Labium with ligula long and slender, divided into 2 lobes in basal third; prementum with two medial setae widely separated; two basal pores narrowly separated, about 1.0 times width of basal pore; several medial pseudopores, 1 setal pore and 2 real pores present on each side of midline; labial palpus very elongate, with many setulae; palpomere 1 largest, about 2.0 – 2.4 times as long as wide, γ-setula contiguous with b-seta, 2 shortest, about 1.3 – 1.5 times as long as wide, 3 parallel-sided and shorter than 1, about 3.5 times as long as wide. Mentum trapezoidal, anterior margin truncate; v-seta relatively long, close to u-seta. Thorax. Pronotum approximately 1.4 times as wide as long, widest at middle. Metanotal scutum with 1 long seta and about 5 – 7 relatively short setae on each side of midline. Mesocoxal cavities narrowly separated; mesoventral process pointed at apex; length ratio of mesoventral process, isthmus and metaventral process 6: 4: 1. Elytra slightly longer and slightly wider than pronotum; elytron approximately 1.4 – 1.5 times as long as wide, postero-lateral margin almost straight; pubescence directed posterior and postero-laterally; hind wings fully developed, flabellum composed of about 9 – 10 setose lobes. Legs. Slender and long, with pubescence and macrosetae; tibiae with different length of two spurs at apex; tarsal formula 4 - 5 - 5, length ratio of tarsomeres 32: 36: 40: 90 (protarsus); 42: 51: 52: 48: 90 (mesotarsus); 75: 71: 57: 57: 110 (metatarsus); one empodial seta present, shorter than claw. Abdomen. Narrow apically; surface fairly glossy and densely pubescent, fine and transverse microsculpture; conspicuous macrosetae present in lateral regions; macrochaetal arrangement of tergites II – VI 01 - 12 - 13 - 13 - 13 (or 23); tergite VII subquadrate; male tergite VIII (Fig. 9 B) with 5 or 6 macrosetae on each side of midline, posterior margin truncate; male sternite VIII (Fig. 9 C) with 7 macrosetae on each side of midline, posterior margin subtruncate; posterior margin of female tergite VIII (Fig. 9 D) similar to male; female sternite VIII (Fig. 9 E) with 6 macrosetae on each side of midline, posterior margin subtruncate, with long marginal setae and minute setae in median region. Aedeagus. Median lobe (Figs. 9 F – G) narrowly ovate and widest in basal third, apical process short and convergent at apex in ventral aspect. Paramerite not pointed apically. Apical lobe of paramerites (Fig. 9 H) recurved and pointed apically, with four setae; a-seta longest, other setae subequal in length. Spermatheca. Bursa elongate, with conical shaped umbilicus; duct long and slender, entirely coiled and apex twisted (Fig. 9 I).	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE84AFFC7609BFEB4FDFCFED9.taxon	materials_examined	Material examined. SOUTH KOREA: Gangwon Prov.: 9 exx. (four on slide), Pyeongchang-gun, Jinbumyeon, Mt. Odaesan, N 37 ˚ 47 ' 03 " E 128 ˚ 33 ' 55 ", 18 vi 2004, SJ Park, sifting.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE84AFFC7609BFEB4FDFCFED9.taxon	distribution	Distribution. Korea (South) and Japan.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
5667352AE84AFFC7609BFEB4FDFCFED9.taxon	discussion	Remarks. Liogluta ursi can be distinguished from other Liogluta species by the fusiform body; the macrochaetal arrangement of abdominal tergites; and the different shape and structure of the aedeagus and spermatheca. This species was described by Sawada (1972) and transferred to the genus Liogluta based on their character systems in Yosii & Sawada (1976). The systematic position of L. ursi is doubtful, considering the fusiform body that distinguishes it from other Liogluta species, including the type species. However, the species has most diagnostic characters of Liogluta except for the body form. Therefore, its generic assignment to the Liogluta is maintained here, pending further comprehensive research of this species.	en	Lee, Seung-Gyu, Ahn, Kee-Jeong (2016): A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species. Zootaxa 4193 (2): 285-303, DOI: 10.11646/zootaxa.4193.2.5
