identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
504F2D6CFFD8FF8D97BEF774FD1C1CBC.text	504F2D6CFFD8FF8D97BEF774FD1C1CBC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myopia alaskana	<div><p>Myopia alaskana (Christiansen and Bellinger, 1980.)</p><p>(Figs. 1–12, 112, 115A)</p><p>Type locality. Lemon Creek Glacier, Juneau, Alaska (Christiansen &amp; Bellinger 1980: 768).</p><p>Material examined. Alaska: Juneau, mountains between Mendenhall Glacier and Montana Creek trail, 58o29'N, 134o40'W, 14.VII.1980, on snow 1,300 m., A. Fjellberg leg.; British Columbia: Vancouver Island, Comox Glacier, 49o33'N, 125o21'W, 26.VIII.1983, on snow 1,950 m., A. Fjellberg leg.; Garibaldi Park, Helm Glacier, 2.IX.1983, on ice 1,930 m., A. Fjellberg leg.; Washington: Mt. Rainier at McClure, 26.VI.1976, on snow, J. Edwards leg.; Mt. Rainier at Anvil and Sugarloaf Mountains, 21.VIII.1980, on snow, A. Fjellberg leg.; Snohomish Co., Columbia Glacier, 30.VII.2002, on ice, P. Hartzell leg.; Snohomish Co., Honeycomb Suiattle Glacier, 48o49'25''N, 121o37'12''W, 12.VIII.2002, on ice, P. Hartzell leg.; Whatcom Co., Lower Curtis Glacier, 48o49'25''N, 121o37'12''W, 6.VIII.2002, on ice, P. Hartzell leg.; Alberta: Bow Glacier (51.64o N, 116.51o W) and Yoho Glacier (51.62o N, 116.55o W), summer 2003, P. Hartzell leg.</p><p>Redescription. Body size up to 1.2 mm. Body shape cylindrical. Appendages relatively short, in particular the tibiotarsi of the two foremost pairs of legs (Fig. 115A). Viewed from above the sides of the head with a small angular projection outside the bases of antennae caused by the forwardly directed two anterior ocelli (Fig. 3). Abd. 5–6 almost fused, with a weak break in setal cover. Colour shiny black or violet black apart from the two distal pure white antennal segments. Second antennal segment often paler in distal part. Occipital area of head with two small white spots. Distal part of tibiotarsi pale. Ocelli 6+6, the two anterior (A and B) enlarged and placed close to antennal bases, directed forward (Figs. 2–3); ocelli G &amp; H invisible. PAO narrow, almost hidden between antennal base and the two anterior ocelli; only one seta below PAO. Ant. 1 on ventral side with two small apical sensilla and up to 13 elongate sensilla (Fig. 4). Ant. 2 with 2–5 enlarged sensilla in lateral position (Fig. 5). Ant. 3 with up to 7 additional long sensilla in addition to the standard set of 4 dorsal and 3 ventrolateral (Fig. 6). Ant. 4 with simple or bifurcate subapical pin-seta. Erect male sensorial setae not present on antennae. Subapical organite small, peg-shaped. Labrum with 4/554 setae, apical edge with 4 small sharp ridges and a composite ventroapical ciliation. Central field of frontoclypeal area with 4 setae. One seta above posterior corner of labrum (Fig. 2). Maxillary palp normally trifurcate with 4 sublobal setae; palps rarely bifurcate, or bifurcate on one side and trifurcate on the other. Labial palps with all papillae and guards present, including guard e7; terminal setae on papillae short. Hypostomal papilla with H much shorter than h1/h2. Basal fields of labium with 4 median and 5 lateral setae. Head with 4+4 (3) postlabial setae. Mandibles normal, with strong teeth. Maxillae unmodified, lamellae covered with fine denticles, Lam.1 not reaching beyond the 3-toothed capitulum. Integument smooth, without visible granulation. Body with a uniform ground cover of short smooth setae, becoming slightly longer towards end of abdomen. Macrochaetae weak, not differentiated from ordinary setae apart from their more erected posture; M1 on abd. 5 is 1.2-1.7 as long as inner edge of last claw. Tergal sensilla short and delicate, distributed as 22/11125 (s) and 10/000 (ms) (Fig. 1). On abd. 1–3 only upper sensillum retained, abd. 5 without anterior sensilla. Abdominal sensilla within the p-row. Ventral tube with 1+1 frontal, 3+3 lateral and 4–5 caudal setae. Retinaculum with 4+4 teeth and 2–4 setae. Manubrium with blunt ventroapical teeth, 1+1(2) short ventroapical setae and fewer than 10 setae in the central ventral field (Fig. 12). Dens crenulated, with 7(6) dorsal setae in proximal 1/3 and a subapical microseta (Fig. 7); ventroapical seta not prolonged. Mucro short and curved, with 5 sharp teeth, no lateral seta. Inner lateral tooth derived from the inner lateral lamella (Fig. 8–9). Tibiotarsi with 11 apical setae (A1–7, T1–4). Legs short, in particular tib.1–2 which have only 3+3 setae along the inner median line (as Fig. 72). Claws small, inner and lateral teeth absent; shape narrow, sides constricted towards base (Fig. 10–11). Unguis only slightly longer than unguiculus. Reproductive males with unmodified setal cover (no epitoky).</p><p>Discussion. The single species in this genus, originally described as a subgenus of Isotoma, was discovered in Alaska and described by Christiansen &amp; Bellinger (1980: 768). These authors suggested that PAO was absent. However, in well cleared specimens the small PAO is visible and it may be that M. alaskana is a specialised Desoria . Molecular genetics may clear up the relationships. The unusual arrangement of the ocelli, however, is unique in the family although there is a gradual transformation from the normal condition to the arrangement shown in Myopia. Some of the new Desoria species described below show an elongation of the eye-field and "wandering" of anterior ocelli (compare Figs. 18 and 41). The low number of tergal sensilla and the few ventral setae on manubrium in Myopia is unlike most Desoria and may be significant.</p><p>The Alberta specimens from Haigh Glacier, Bow Glacier and Yoho Glacier all differ from Alaskan specimens in having three terminal white antennal segments rather than two (Fig. 112). No other consistent differences could be found.</p><p>Distribution and ecology. The species appears to be common on glacial ice and snow fields in the Rocky Mountains from southern Alaska to Washington, extending east to Alberta. For details, see above. On Haig Glacier specimens were observed on snow and ice at a density of 5.000–10.000 inds./m2, covering an area of several hectares. This species could be a North American ecological equivalent to the European "Gletscherflohs" ( Desoria saltans Nicolet, 1841).</p></div>	https://treatment.plazi.org/id/504F2D6CFFD8FF8D97BEF774FD1C1CBC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fjellberg, Arne	Fjellberg, Arne (2010): Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A. Zootaxa 2513: 27-49, DOI: 10.5281/zenodo.196078
504F2D6CFFDDFF8D97BEF214FDAA1F69.text	504F2D6CFFDDFF8D97BEF214FDAA1F69.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Desoria Nicolet 1841	<div><p>Desoria Nicolet, 1841</p><p>Type species: Desoria saltans Nicolet, 1841</p></div>	https://treatment.plazi.org/id/504F2D6CFFDDFF8D97BEF214FDAA1F69	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fjellberg, Arne	Fjellberg, Arne (2010): Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A. Zootaxa 2513: 27-49, DOI: 10.5281/zenodo.196078
504F2D6CFFDDFF8C97BEF2A4FF3F1EC4.text	504F2D6CFFDDFF8C97BEF2A4FF3F1EC4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Desoria uniens Christiansen and Bellinger 1980	<div><p>Desoria uniens Christiansen and Bellinger, 1980</p><p>(Figs. 13–18, 115B)</p><p>Type locality. Allegheny Mountains, Terra Alta, West Virginia (Christiansen &amp; Bellinger 1980: 761)</p><p>Material examined. Alaska: Clearwater Mountains, 63.20o N, 147.04o W, 30.VII.1980, alpine meadow 1,670 m., A. Fjellberg leg.; British Columbia: Vancouver Island, Comox Glacier, 26.VIII.1983, moss at melt water 1,850 m.; Sparwood, Bald Mountain, 7.VI.1983, moss at melt water 2,700m., A. Fjellberg leg.; Garibaldi Park, Panorama Ridge, 1.IXC.1983, wet moss at snow 1,920 m., A. Fjellberg leg.; Washington: Mount Rainier, Sugarloaf Mountain, 21.VIII.1980, moss at melt water, 2,400 m., A. Fjellberg leg.; Olympic National Park, Hurricane Ridge, 10.IX.1983, wet moss at snow 1,940 m., A. Fjellberg leg.; Nisqually Glacier, summer 2003, P. Hartzell leg.; Oregon: Zigzag South, summer 2003, P. Hartzell leg.; Colorado: Boulder County, Upper Diamond Lake, 26.VIII.1980, wet moss at snow 3,550 m., A. Fjellberg leg.</p><p>Redescription. Body size up to 1.6 mm. Body shape slender, normal for the genus (Fig. 115B). Head not prognathous. Abd. 5–6 clearly demarcated. Colour of large individuals uniformly blackish green. Smaller specimens violet green with paler extremities, last abdominal segments (abd. 4–6) and lateral and posterior part of head often paler than rest of body. Ocelli 8+8, G and H very small (Fig. 18). PAO oval, about the size of nearest ocellus, with three associated setae. Ant. 1 on ventral side with up to 6 short blunt apical sensilla and several (up to 10) thin long setaceous sensilla, near base with two setaceous microsensilla. In addition males with 3–4 short erect sensilla on the inner ventrolateral side (Fig. 15). Ant. 2–3 also with groups of short blunt ventroapical sensilla in addition to short setaceous sensilla which are dispersed over the surface (Fig. 16). Ant. 4 with numerous setaceous sensilla, simple pin-seta and a small rod-shaped subapical organite. Labrum with 4/554 normal setae, four sharp apical folds and composite ventroapical ciliation. Frontoclypeal area with 12–15 setae. Maxillary palps bifurcate with 4 sublobal setae. Labial palps complete, guard e7 present. Proximal setae 5. Hypostomal group normal, H stiff and straight. Basal fields of labium with 5 lateral and 5 median setae. Head with 3–4 postlabial setae on each side of the ventral line. Mandibles normal, strong. Maxillae with short densely denticulate lamellae, none of them extending beyond tip of capitulum. Integument smooth, cover of body setae uniform and moderately long, macrochaetae in anterior part of abdomen not longer than ordinary setae, only recognised by their erect posture. Median macrochaeta on abd. 5 about 1.5 as long as inner length of last claw. Macrochaetae on abd. 5–6 smooth or indistinctly serrated. Setaceous sensilla on the tergites distributed as 88/58785, with some variation (Fig. 13). Spine-like microsensilla as 11/111, those on thorax are set outside the setaceous sensilla. Sensilla along the posterior edge of tergites set in front of the p-row. Abd. 4–5 with some anterior sensilla. Ventral tube with 1–5 frontal setae on each side (very variable), lateral setae always 3+3, posterior setae up to 12. Retinaculum with 4+4 teeth and up to 12 setae in large specimens. Manubrium with 3–4 short ventroapical setae on each side, midventral field rather narrow, with 20–25 setae, manubrial teeth blunt. Dens with more than 30 dorsal setae in large specimens, only apical 1/3 free of dorsal setae. Mucro short and compact, apparently with 3 teeth; a fourth apical tooth reduced to a small hook on the ventral edge; subapical tooth with inner lamella ending before tip; inner basal tooth in line with subapical; lateral seta absent (Fig. 17). Tibiotarsi with 9 apical setae (A1–7, T1, T4, Fig. 14), seta A1 slightly prolonged, pointed; in small juveniles T 4 may be absent. Inner side of tib.1–2 with more than 3+3 setae along median line. Upper subcoxa on first leg with two setae. Claws normal, unguis with small lateral teeth, no inner tooth; unguiculus sometimes with a small corner tooth. Males present, reproductive specimens not observed.</p><p>Discussion. The tridentate mucro and the reduced number of apical setae on tibiotarsi (9, not 11) links this species with the pjasini -group (Potapov 2001), unlike all other Desoria of the Rocky Mountains. The mucronal shape and the presence of only 3+3 lateral setae on the ventral tube is similar to species of Isotomurus of the palustris group, but trichobothria are not present.</p><p>The recent samples were compared with a slide with 3 paratypes and the holotype (MCZ) from Terra Alta, West Virginia, and judged to belong to the same species. The type specimens have the same shape of mucro, 8–9 tibiotarsal apical setae, bifurcate maxillary palp and 3+3 laterodistal setae on the ventral tube. Type specimens have slightly longer body setae (M1 on Abd. 5 is 2.0 times as long as inner length of last claw) and their colour may differ as dark pigment is mainly present in dorsomedian part of the anterior body segments. However, at present there seem to be no sharp characters distinguishing the Rocky Mountain samples from the type specimens. Fjellberg (1984) reported D. uniens from the Colorado Front Range Mountains. These specimens were re-examined and found to be conspecific with the more recent samples.</p><p>Distribution and ecology. Common and often abundant on and near snow and ice fields in the Rocky Mountains of British Columbia, Washington, Oregon and Colorado (details above). In the original description of the species Christiansen &amp; Bellinger (1980) also included records from California, Montana and New Mexico.</p></div>	https://treatment.plazi.org/id/504F2D6CFFDDFF8C97BEF2A4FF3F1EC4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fjellberg, Arne	Fjellberg, Arne (2010): Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A. Zootaxa 2513: 27-49, DOI: 10.5281/zenodo.196078
504F2D6CFFDCFF8F97BEF443FAB51E29.text	504F2D6CFFDCFF8F97BEF443FAB51E29.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Desoria albicornis	<div><p>Desoria albicornis sp. n.</p><p>(Figs. 19–29, 111, 115C)</p><p>Type material (all in alcohol, sex not checked). Holotype and 55 paratypes from: "Alaska, mountains between Mendenhall Glacier and Montana Creek Trail, 58o29'N, 134o40'W, on melt-water ponds, 975 m. alt., 13.VII.1980, A. Fjellberg leg." (INHS).</p><p>Description. Body size up to 1.3 mm. Body shape cylindrical, slender, with relatively short legs (Fig. 115 C). Head shape normal. Abd. 5–6 weakly separated, with a clear break in setal cover. Colour uniformly violet black, antennae pure white apart from dark proximal half on ant. 1. Head with a transverse row of four small white spots (Fig. 111). Tibiotarsi paler than rest of the legs. Each side of head with 6 large ocelli and two very small ones. PAO oval, as long as diameter of nearest ocellus, with three associated setae (Fig. 24). Ant. 1 on ventral side with up to 10 long slender sensilla and 2–3 short ones in apical position. Ant. 2 with an enlarged sensillum in lateral position (Fig. 22). Ant. 3 organ normal, with 4 dorsoapical sensilla and 3 ventrolateral (Fig. 23). Ant. 4 with bifurcate subapical pin seta and a short rod-shaped subapical organite in a pit. Labrum with 4/554 slender setae, apical edge with 4 low ridges and a composite ventroapical ciliation (Fig. 29). Central part of frontoclypeal field with 4 (rarely 5) setae (Fig. 28). Maxillary outer lobe with bifurcate palp and 4 sublobal setae. Labial palp with apical papillae A–E present, 4 proximal setae; guard e7 present at base of papilla E; terminal seta of the papillae shorter than the guards. Lateral process (l.p.) on papilla E strong, finger-like. Hypostomal papilla with H shorter than h1/h2. Basal fields of labium with 4 median and 5 lateral setae. Head with 4+4 postlabial setae along ventral line. Mandibles normal, with moderately strong teeth. Maxillae with 3-toothed capitulum and 6 lamellae covered with fine denticles only. Lamellae short, not projecting beyond tip of capitulum. Integument smooth, without visible granulation. The cover of body setae short, dense and uniform. Macrochaetae smooth, acuminate, weakly differentiated from ordinary setae. Median macrochaeta on abd. 5 about half as long as tergite in midline. Tergites with 44/3345 ordinary sensilla n the p-row on abdomen (one anterior s on abd. 5) and slightly in front of the p-row on th. 2–3. Spine-like microsensilla as 11/001 (Fig. 19). Ventral tube with 1+1 frontal setae, usually 5+5 (4–6) lateral and 3–5 caudal setae. Retinaculum with 4+4 teeth and 2–4 setae. No ventral setae on thorax. Manubrium with a variable number of ventral setae, fewer than 15 (Fig. 27); ventroapical thickening with blunt teeth; 1+1 short ventroapical setae. Dens with 7 dorsal setae in the proximal 1/3 and a subapical microseta; ventroapical seta not prolonged. Mucro short and curved, with 5 sharp teeth; a tooth on the inner side derived from the inner lateral lamella (Figs. 25–26); no lateral seta. Tibiotarsi with 11 acuminate apical setae (A1–7, T1–4, Fig. 20). Legs shortened, in particular tib.1–2 with only 3+3 setae along median line (as Fig. 72). Claws narrow, slightly constricted at base, without visible inner and lateral teeth; unguiculus with wide lamellae (Figs. 21– 22). Reproductive males without differentiated setal cover.</p><p>Etymology. The name reflects the strikingly white antennae of the species.</p><p>Discussion. The only described North American Desoria having a 5-toothed mucro is cancellarei Christiansen et Bellinger, 1980, described from California and Iowa (on snow). This species is said to have 6+6 ocelli, otherwise it comes very close to albicornis, having few setae on ventral tube and retinaculum as well as untoothed claws of similar shape. However, cancellarei has much longer abdominal macrochaetae, 2– 3 times as long as inner edge of the claw (1.5 as long in albicornis). D. albicornis resembles Myopia alaskana in colour, except only basal part of ant. 1 is dark, the rest is white.</p><p>Distribution and ecology. So far only recorded from snow and melt water ponds in Alaska (types) and British Columbia (Vancouver Island: Comox Glacier, 49o33'N, 125o21'W, 26.VIII.1983, A. Fjellberg leg.).</p></div>	https://treatment.plazi.org/id/504F2D6CFFDCFF8F97BEF443FAB51E29	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fjellberg, Arne	Fjellberg, Arne (2010): Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A. Zootaxa 2513: 27-49, DOI: 10.5281/zenodo.196078
504F2D6CFFDFFF8E97BEF39CFEBB1F4A.text	504F2D6CFFDFFF8E97BEF39CFEBB1F4A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Desoria triangularis	<div><p>Desoria triangularis sp. n.</p><p>(Figs. 30–34, 110, 115E)</p><p>Type material. Holotype (alc.) from "Alaska. Richardson Hwy., N of Paxson. Triangle Peak, 63o24'N, 145o31'W, 09.VIII.1980. Wet moss near snow, 1,600 m alt. A. Fjellberg 144/80." Paratypes, two specimens (slide), male and female, from the same sample (INHS).</p><p>Description. Body size up to 1.2 mm. Body shape cylindrical, rather stout (Fig. 115E). Head shape normal. Abd. 5–6 fused without clear break in setal cover. Colour bluish or greenish black, legs and furca bluish grey. Antennae white, except the first segment which is bluish grey. Occipital area of head with two large triangular white spots (Fig. 110); in addition head and body with diffuse blue-grey marks in the spots that usually mark muscular incisions. Ocelli 6+6 (A–F), a small G-ocellus may also be present. PAO elongated, as long as diameter of nearest ocellus. Two setae associated with PAO (Fig. 31). Ant. 1 with two short ventroapical sensilla and about 8 longer setaceous sensilla. Ant. 2 with one curved enlarged sensillum in ventroapical position. Ant. 3 organ with 4 dorsal sensilla and 3 (one small) lateral ones. No additional sensilla (Fig. 34). Ant. 4 with simple subapical pin seta, subapical organite as an apically expanded rod set in a pit. Labral chaetotaxy 4/554, setae normal, not spine-like. Apical edge of labrum with 4 sharp folds and a composite ventroapical ciliation. Central frontoclypeal field with 6–8 setae, two setae above posterior corner of labrum. Maxillary palp bifurcate, 4 sublobal setae. Labial palps unmodified, with terminal setae of the papillae as long as the inner guard setae; guard e7 present. Hypostomal setae normal. H long and straight, sub equal to h1 and h2. Basal fields of labrum with 4 median and 5 lateral setae. Head with 4+4 postlabial setae along ventral line. Mandibles normal, strong. Maxillae unmodified, with strong 3-toothed capitulum and 6 finely denticulate lamellae of which Lam.1 reaches tip of capitulum. Body integument smooth, without visible granulation. Setal cover of body very dense, "double", with short and long setae (Fig. 30). Macrochaetae clearly differentiated on abdomen, but not long. Median macrochaetae on abd. 5 about 1.7 as long as inner length of last claw. Sensilla on the tergites hard to distinguish from ordinary short setae and formula unclear, but numbers appear to be low, probably 4–5 on anterior abdominal segments. Formula of spine-like microsensilla appears like 10/001 but must be verified. Ventral tube with 1+1 frontal, 4+4 lateral and 4 caudal setae (only two specimens examined). Retinaculum with 4+4 teeth and 4–5 setae. Manubrium with blunt apical teeth, 1+1(2) short ventroapical setae. Central field of ventral side densely piles (more than 50 setae). Dens with 8–9 dorsal setae in proximal half; ventroapical seta not prolonged. Mucro short and compact, with 5 sharp teeth; inner lateral lamella transformed to a secondary tooth; lateral seta absent (Fig. 32). Tibiotarsi with 11 apical setae (A1–7, T1–4), all acuminate. Legs shortened, inner side of tib.1–2 with 3+3 setae along median line (as Fig. 72). Claws small, inner and lateral teeth absent on unguis (Fig. 33). Males present.</p><p>Etymology. The name refers to both the type locality (Triangle Peak) and the triangular white spots on the head.</p><p>Discussion. The outstanding character of this species is the strikingly "double" cover of seta on the body which separates it from all other cryophilic Desoria / Gnathisotoma which are dark with white antennae. From albicornis it differs by having two triangular pale spots on head, rather than four small spots. Also abd. 5-6 are completely fused while they are weakly separated in albicornis . The absence of the spine-like microsensillum on th. 3 in triangularis may also be a distinctive character, but a verification is necessary.</p><p>Distribution and ecology. So far only known from the type locality in the alpine region of the mountains of southern Alaska.</p></div>	https://treatment.plazi.org/id/504F2D6CFFDFFF8E97BEF39CFEBB1F4A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fjellberg, Arne	Fjellberg, Arne (2010): Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A. Zootaxa 2513: 27-49, DOI: 10.5281/zenodo.196078
504F2D6CFFDEFF8097BEF2BBFADA1DD1.text	504F2D6CFFDEFF8097BEF2BBFADA1DD1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Desoria ater	<div><p>Desoria ater sp. n.</p><p>(Figs. 35–43, 113, 115D)</p><p>Type material (in alcohol, sex not checked). Holotype and 11 paratypes from "Alaska Mountains,. between Mendenhall Glacier and Montana Creek Trail, 58o29'N, 134o40'W, on goat dung in snowfields, 1,000 m alt., 14.VII.1980, A. Fjellberg leg." (INHS).</p><p>Description. Body size up to 1.7 mm. Body shape slender, typical Desoria with abdomen slightly pearshaped (Fig. 115D). Head shape normal, not prognathous/hypognathous. Abd. 5–6 completely fused, without a gap in setal cover. Colour uniformly dark greyish black. Antennae, legs (beyond coxae) and furca paler, greyish green; antennal segments all of same colour. Head with two small white spots between the eye-fields (Fig. 113). Ocelli 8+8, G &amp; H slightly smaller, arrangement characteristic, with ocelli ABCDE forming a triangle. PAO oval, as long as diameter of nearest ocellus. Two setae near ventral edge of PAO (Fig. 41). Ant. 1 with a group of 5–6 short ventroapical sensilla and more than 10 setaceous sensilla (Fig. 36). Ant. 2 and 3 with several lateral and ventrolateral setaceous sensilla (Figs. 37–38). Ant. 4 with bifurcate subapical pin seta, subapical organite small, globular. Labrum with 4/554 slender normal setae, apical edge with 4 sharp folds and a composite ventral ciliation. Frontoclypeal field with more than 10 setae (Fig. 40), two setae present between antennal base and upper corner of labrum. Maxillary palp bifurcate, four sublobal setae. Labial palps complete, guard e7 present, four proximal setae; terminal setae of the papillae as long as guards. Hypostomal papilla with subequal H and h1/h2. Basal fields of labium with 5 medial and 5 lateral setae. Ventral side of head with 4–8 postlabial setae on each side. Mandibles normal, strong. Maxillae unmodified, with 3-toothed capitulum and short lamellae covered with fine denticles. Body integument smooth, macrochaetae very long with fine ciliation, mostly on anterior side (Figs. 35, 39). Median macrochaetae on abd. 5 are 3.5–4.0 as long as inner length of last claw. Ground cover of setae smooth and dense, on head and anterior part of the body notably double with short setae and longer dagger-like setae (as Fig. 30). Sensilla on the tergites hard to separate from ordinary setae, but numbers appear to be low, around 5 on each tergite from th. 2–abd. 5, positioned slightly anterior to the p-row of setae. Anterior sensilla appear to be absent on the tergites. Spinelike microsensilla as 11/100. Ventral tube on each side with 2–4 frontal and 10–12 lateral setae. Posterior side with 6–8 setae (upper four in a transverse row). No ventral setae on thorax. Retinaculum with 4+4 teeth and 15–20 setae. Manubrium with blunt ventroapical teeth and usually 2+2 short ventroapical setae. Ventromedian field with more than 40 setae. Dens crenulated, with 10–12 dorsal setae in proximal half; subapical microseta present; ventroapical seta not prolonged. Mucro short and compact, with 4 sharp teeth (Fig. 42); inner basal tooth small; inner lateral lamella not transformed to a tooth, ending at tip of the subapical tooth; lateral seta absent. Tibiotarsi with 11 apical setae, all acuminate. Inner side of tib.1–2 with more than 3+3 setae along median line. Claws normal, with distinct lateral teeth, no inner tooth; unguiculus with low dorsal lamella, sometimes toothed (Fig. 43). Nine examined specimens examined are all females.</p><p>Etymology. The name ater (Latin: black) reflects the dark colour of the animal.</p><p>Discussion. The dark colour which includes the antennae, short 4-toothed mucro, increased number of antennal sensilla, eye configuration and particularly the very long and densely ciliated macrochaetae serve to distinguish this species.</p><p>Distribution and ecology. Only known from the type locality where large number of specimens occurred on droppings from mountain goat deposited on glacier ice in the mountains east of Juneau, Alaska.</p></div>	https://treatment.plazi.org/id/504F2D6CFFDEFF8097BEF2BBFADA1DD1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fjellberg, Arne	Fjellberg, Arne (2010): Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A. Zootaxa 2513: 27-49, DOI: 10.5281/zenodo.196078
504F2D6CFFD0FF8097BEF134FBA21AE6.text	504F2D6CFFD0FF8097BEF134FBA21AE6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Desoria garibaldii	<div><p>Desoria garibaldii sp. n.</p><p>(Figs. 44–49, 115F)</p><p>Type material (all in alcohol, sex not checked). Holotype and 60 paratypes from " Canada, B.C., Garibaldi Park, Helm Glacier, 49o57'N, 123o0'W, 02.IX.1983. Muddy blue ice at terminal lobe, 1,770 m alt. A. Fjellberg 398/83" (INHS). Other material: USA, Alaska, Ptarmigan Glacier, summer 2003, P. Hartzell leg; Washington, Snohomish Co., Honeycomb Suiattle Glacier, 12.VIII.2002, snow pack on glacier ice, P. Hartzell leg.; same county, Columbia Glacier, 04.VIII.2002, snow pack on glacier ice, P. Hartzell leg; same state, Whatcom Co., Easton Glacier, 48o44''N, 121o50'W, 04.VIII.2002, on snow, P. Hartzell leg.</p><p>Description. Body size up to 1.6 mm. Body shape slender, typical Desoria (Fig. 115F). Head shape normal. Abd. 5–6 almost completely fused, with a weak incision between the tergites and a distinct break in setal cover. Colour completely black, including extremities. Head without white spots between eye-fields. Freshly collected specimens with a greenish lustre. Unlike most species even small juveniles as dark as the adults. Ocelli 8+8, rather small. PAO oval, about 1.5 as long as diameter of nearest ocellus. Three setae associated with PAO (Fig. 48). Ant. 1 with 5–8 short curved ventroapical sensilla and a few curved setaceous sensilla (Fig. 45). Ant. 2 with a single ventrolateral curved sensillum in apical position. Ant. 3 organ, with a few additional sensilla (Fig. 46). Ant. 4 with bifurcate pin seta and a small apically expanded peg in the subapical organ. Labrum with 4/554 normal setae, apical edge with 4 sharp ridges and a composite ventral ciliation. Frontoclypeal field with 10–15 setae, two setae between antennal base and upper corner of labrum. Maxillary palp bifurcate, 4 sublobal setae. Labial palp complete, guard e 7 present; terminal seta on the papillae as long as guards. Hypostomal papilla with subequal H and h1/h2. Labial base with 5(4) median and 5 lateral setae. Head with 4+4(3) postlabial setae. Mandibles normal, strong. Maxillae unmodified, with 3-toothed capitulum and short lamellae densely covered with fine denticles. Integument smooth. Ground cover of setae uniform and dense, not markedly "double" (cf. Fig. 30) except on dorsal side of head and thorax. Macrochaetae smooth, short, poorly differentiated from ordinary setae. Median macrochaeta on abd. 5 only 1.0–1.2 as long as inner length of last claw. Sensillary formula in a juvenile specimen as 55/44456, spine-like microsensilla 10/001 (Fig. 44). On abd. 1–4 sensilla set slightly anterior to p-row; anterior sensilla present on abd. 4–5. Ventral tube on each side with 1+1 frontal and 5–6 lateral setae, posterior side with 4–6 setae. Retinaculum with 4+4 teeth and 9–15 setae. No ventral setae on thorax. Manubrium with blunt apical teeth and 2–3 short ventroapical setae on each side, ventromedial field with many (&gt;40) setae. Dens dorsally crenulated, with 20–30 dorsal setae extending beyond mid point; ventroapical seta not prolonged. Mucro short and compact, with 4 sharp primary teeth. Inner lateral lamella not reaching tip of subapical tooth and so forming a small secondary tooth; lateral seta absent (Fig. 49). Tibiotarsi with 11 apical setae, all acuminate. Inner side of tib.1–2 with 3–4 setae along each side of median line. Claws with small lateral teeth, a weak inner tooth may be present on anterior two pairs of claws; unguiculus with a high dorsal lamella (Fig. 47). Males present, reproductive individuals not seen.</p><p>Etymology. The new species is named after the type locality, Garibaldi Provincial Park in British Columbia. The park itself was named in honour of Guiseppe Garibaldi (1807–1882), the Italian revolutionary who paved the way for a unified Italy.</p><p>Discussion. The species is recognised by the dark colour, short body setae, compact mucro and many dorsal setae on dens. It is close to D. cancellarei Christiansen et Bellinger, 1980, described from snow in Iowa, but this latter species is pale blue and has only 6+6 ocelli and longer abdominal setae (longest abdominal macrochaetae at least twice as long as inner edge of last claw). Also garibaldii has a 4-toothed mucro, while the mucro has 5 teeth in cancellarei .</p><p>Distribution and ecology. Common and abundant on and near snow and glacier ice in the northwestern Rocky Mountains in Alaska, British Columbia and Washington (details see above).</p></div>	https://treatment.plazi.org/id/504F2D6CFFD0FF8097BEF134FBA21AE6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fjellberg, Arne	Fjellberg, Arne (2010): Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A. Zootaxa 2513: 27-49, DOI: 10.5281/zenodo.196078
504F2D6CFFD3FF8397BEF681FE8F1A6B.text	504F2D6CFFD3FF8397BEF681FE8F1A6B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Desoria cryophila	<div><p>The cryophila group</p><p>The following four new species and two probably undescribed species (sp. A and sp. B) form a group characterized by an almost white body with dark eye-spots and a 5-toothed mucro. This grouping may not necessarily have a phylogenetic significance, but the diversity of such similar forms in a limited mountainous area is notable.</p></div>	https://treatment.plazi.org/id/504F2D6CFFD3FF8397BEF681FE8F1A6B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fjellberg, Arne	Fjellberg, Arne (2010): Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A. Zootaxa 2513: 27-49, DOI: 10.5281/zenodo.196078
504F2D6CFFD3FF8397BEF08EFCCA1B45.text	504F2D6CFFD3FF8397BEF08EFCCA1B45.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Desoria pilifrons	<div><p>Desoria pilifrons sp. n.</p><p>(Figs. 50–56)</p><p>Type material. Holotype: One specimen (slide, not sexed) labelled " Canada, B.C. Vancouver Island, Comox Glacier, 49o33'N, 125o21'W, 26.VIII.1983. Wet moss in melt water, 1.850 m. A. Fjellberg 375/83". Paratypes: Four specimens in two slides, same data as holotype (INHS).</p><p>Description. Body size up to 1.6 mm. Body shape slender, with pear-shaped abdomen. Abd. 5–6 fused, but with a gap in setal cover. Head distinctly prognathous (Fig. 50). Colour pale bluish grey, darkest in anterior part of the body. Ocelli 8+8, G &amp; H smaller. PAO oval, about 1.3 times as long as diameter of nearest ocellus. Three setae associated with PAO (Fig. 51). Ant. 1 with 4–5 short ventroapical sensilla and a few curved setaceous sensilla. Ant. 2 with two small setaceous sensilla in lateral position near apex. Ant. 3 organ normal, with several additional small setaceous sensilla (Fig. 54). One or two erect setaceous sensilla ventrally on ant. 2–3 in some individuals. Ant. 4 apically with two blunt lobes and a bifurcate pine seta; subapical organ with a globular organite (Fig. 52). Labrum with 4/554 strong curved setae. Apical edge with 4 sharp lobes and a composite ventral ciliation. Frontoclypeal area with more than 30 setae in adults (Fig. 53). Maxillary palp bifurcate, 4 sublobal setae. Labial palps complete, guard e7 present, 4 proximal setae. Terminal sensilla on labial papillae longer than guards. Basal fields of labium with 5 median and 5 lateral setae. Hypostomal papilla with H longer than h1/h2. Head with 5–7 postlabial setae on each side of ventral line. Mandibles normal, strong. Maxillae unmodified, with 3-toothed capitulum and 6 short lamellae covered with fine denticles. Body integument smooth. Ground cover of setae dense, markedly "double" all over the body with short microchaetae and dagger-like mesochaetae (as Fig. 30). Macrochaetae strong, those on abd. 5–6 finely serrated in apical half. Median macrochaeta on abd. 5 2.5–3.0 times as long as inner edge of last claw. Sensillary equipment on the tergites unclear, but numbers apparently low with five setaceous sensilla on each side of th. 2–abd. 2; spine-like microsensilla as 11/111. On abd. 5 one sensillum in anterior position. Ventral tube on each side with 8–10 frontal and 8–9 lateral setae; posterior side with more than 10 setae in large individuals. No ventral setae on thorax. Retinaculum with 4+4 teeth and 7–9 setae. Manubrium with blunt apical teeth and 3+3(2) short ventroapical setae, ventral field densely pilose, with more than 50 setae. Dens dorsally crenulated, with about 20 dorsal setae in the basal half; ventral subapical seta not prolonged. Mucro 4-toothed, rather slender with relatively large apical tooth (Fig. 55), inner lateral lamella ending at the subapical tooth without forming a secondary tooth; lateral seta absent. Tibiotarsi with 11 apical setae, all acuminate. Inner side of tib.1–2 with more than 3+3 setae along median line. Claws normal, with distinct lateral teeth; inner tooth present on anterior pairs of claws; unguiculus on two last pairs of claws with a high dorsal lamella bearing a small corner tooth (Fig. 56). Males present, reproductive individuals not seen.</p><p>Etymology. The name reflects the unusually dense setal vestiture in the frontoclypeal field.</p><p>Discussion. Differs from the other species with a 4-tothed mucro ( ater and garibaldii) by prognathous head with increased number of setae in frontoclypeal area (&gt;30), the marked "double" setal cover, long macrochaetae and increased number of setae on ventral tube (Table 1).</p><p>Distribution and ecology. Only known from the type locality at Comox Glacier, Vancouver Island where it occurred along a melt water stream near the glacier.</p></div>	https://treatment.plazi.org/id/504F2D6CFFD3FF8397BEF08EFCCA1B45	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fjellberg, Arne	Fjellberg, Arne (2010): Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A. Zootaxa 2513: 27-49, DOI: 10.5281/zenodo.196078
504F2D6CFFD2FF8297BEF08EFDC91BDE.text	504F2D6CFFD2FF8297BEF08EFDC91BDE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Desoria cryophila	<div><p>Desoria cryophila sp. n.</p><p>(Figs. 57–63, 115H)</p><p>Type material. Holotype (alc.) and 33 paratypes (26 in alcohol, 7 in two slides) from " Canada, Alberta, Kananaskis, French creek, 50o55'N, 134o40'W, 16.VI.1984. Wet moss at waterfall, 1,930 m. alt., A. Fjellberg leg." (INHS).</p><p>Description. Size up to 1.2 mm. Body shape normal for the genus, with a slightly pear-shaped abdomen. Head normal, not prognathous/hypognathous. Abd. 5–6 fused without distinct break in setal cover. Colour white with black eye-spots (Fig. 115H). Ocelli 6+6 (G &amp; H absent), PAO oval, slightly shorter than diameter of nearest ocellus. Three setae associated with PAO (Fig. 58). Ant. 1 ventrally with 4–5 short thick apical sensilla and several slender setaceous sensilla; base with a group of ventral microsensilla (Fig. 60). Ant. 2 with 3–4 small and thin sensilla in lateroapical position (Fig. 62). Ant. 3 organ with some additional setaceous sensilla (Fig. 61). No ventral erect sensilla present on ant. 2–3. Ant. 4 with bifurcate pin seta and a small globular organite in the subapical organ. Labrum with 4/554 normal setae, apical edge with 4 sharp folds and a composite ventral ciliation. Frontoclypeal area with 5–9 setae, two setae between antennal base and labral corner. Maxillary palp bifurcate, four sublobal setae. Labial palps unmodified, complete, guard e7 present; terminal sensilla on the papillae as long as guards; four proximal setae present. Hypostomal group with H as long as h1/h2. Basal fields of labium with 4(5) median and 5 lateral setae. Head with 5–6 postlabial setae. Mandibles normal, strong. Maxillae unmodified, with short denticulate lamellae and 3-toothed capitulum. Body integument smooth. Body with a dense markedly "double" setal cover with short ordinary setae and dagger-like mesochaetae some as long as the erect macrochaetae on thorax and abd. 1 (as Fig. 30). Macrochaetae moderately strong on abdomen, distinctly serrated on the last abdominal segments. Median macrochaetae on abd. 5 about 2.5 as long as inner edge of last claw. Sensilla on the tergites hard to see in adults because of dense setal cover, but several half-grown juveniles show distribution as 55/44456 (Fig. 57) with some variation in number on the three last segments; anterior sensilla present on abd. 4–5; spine-like microsensilla distributed as 11/001. Ventral tube on each side with (1)3–5 frontal and up to 15 lateral setae, posterior side with up to 8 setae. Retinaculum with 4+4 teeth and up to 8 setae. Manubrium with blunt apical teeth and 3+3(4) short ventroapical setae; ventromedial field densely pilose, with more than 50 setae. Dens dorsally crenulated, with 12–14 dorsal setae in proximal half and a tiny subapical setula; subapical ventral seta not prolonged. Mucro with 4 sharp primary teeth and a small secondary tooth on the inner side formed by the inner lateral lamella ending before tip of the subapical tooth (Fig. 59); apical and subapical teeth subequal; no lateral seta. Tibiotarsi with 11 apical setae, all acuminate. Legs long and slender, inner side of Tib. 1–2 with 4– 5 setae along each side of median line (Fig. 63). Claws of normal shape, with small laterobasal teeth and a weak tooth in the middle of the inner edge; unguiculus with high basal lamella, usually with a small corner tooth. Males present, reproductive individuals not seen.</p><p>Etymology. The name reflects the cold environment in which the species lives.</p><p>Discussion. Recognised by the white colour, bifurcate maxillary palp, more than 5 lateral setae on the ventral tube, more than 10 dorsal setae on dens, rarely more than 4 basomedian setae on labium, and subcoxa 1 having only one seta. Otherwise it is similar to rosea which has a slightly larger PAO, more compact mucro and a more open cover of setae on Abd. 5–6 which is less rounded and more clearly separated. The erect sensilla which are observed ventrally on ant. 2–3 in rosea, are not seen in cryophila .</p><p>Distribution and ecology. Only known from the type locality which is a cold water stream in the Kananaskis mountains, Alberta.</p></div>	https://treatment.plazi.org/id/504F2D6CFFD2FF8297BEF08EFDC91BDE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fjellberg, Arne	Fjellberg, Arne (2010): Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A. Zootaxa 2513: 27-49, DOI: 10.5281/zenodo.196078
504F2D6CFFD2FF8597BEF734FAB01884.text	504F2D6CFFD2FF8597BEF734FAB01884.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Desoria rosea	<div><p>Desoria rosea sp. n.</p><p>(Figs. 64–70)</p><p>Type material. Holotype (alc.) and 9 paratypes (3 in alc., 6 in two slides) from " Canada, B.C., Sparwood, 49o43'N, 114o57'W, 09.VI.1983. Plant debris in moist avalanche track, 1,900 m., A. Fjellberg 119/83" (INHS).</p><p>Other material: Canada, B.C., Garibaldi Park, Panorama Ridge, 1.IX.1983. Wet moss at snow edge, 1,900 m. A. Fjellberg 387/83; Canada, Alberta, Kananaskis, Marmot Creek at Twin Fork, 29.VI.1983. Litter in spruce Krummholtz at tree line, 2,300 m. A. Fjellberg 193/83; Canada, Alberta, Kananaskis, French Creek, 16.V 1.1984, Moss &amp; litter at stream bank, A. Fjellberg leg.; USA. Alaska, Juneau, mountains. between Mendenhall Glacier and Montana Creek Trail, 13.VII.1980. On ponds at snow bed, 950 m. alt., A. Fjellberg leg; USA, Washington, Wenatchee Mountains, Ingalls Lake, 23.VIII.1980. Hemlock litter, A. Fjellberg 181/ 80.</p><p>Description. Size up to 1.2 mm. Colour white or faintly pinkish grey, eye-spots black. Ocelli 6+6, G and H absent. PAO elongated, slightly longer than diameter of nearest ocellus, with three associated setae (Fig. 64). Body shape normal, head not prognathous or hypognathous. Last two abdominal segments weakly separated. Ant. 2–3 with an increased number of additional sensilla, erect ventral sensilla present in full grown specimens (Figs. 65–66). Ant. 4 with bifurcated pin seta and a small globular organite. Basomedian field of labium usually with five setae (Fig. 67). Details of labrum, frontoclypeal field, maxillary palps, labial palps, mandibles and maxillae as in previous species. Body integument smooth; setal cover open and clearly "double" in anterior part of the body (short ordinary setae, longer dagger-like mesochaetae). Longest macrochaetae on tip of abdomen clearly serrated; median macrochaetae on abd. 5 more than three times as long as inner edge of last claw. Spine-like microsensilla as 11/001, rarely 11/101. Number of ordinary sensilla as 55/44456. Ventral tube on each side with 3 frontal and 5 lateral setae. Posterior side with 6–8 setae. Retinaculum with 3–4 setae. Manubrium with blunt apical teeth and 3+3 short ventroapical setae; ventromedian field densely pilose. Mucro 5-toothed, apical tooth smaller than subapical (Fig. 68). Legs long and slender, inner side of tib. 1–2 with 3–4 pairs of setae along median line. Subcoxa 1 with 1–2 setae. Claw structures as in previous species. Males present, reproductive individuals with unmodified setal cover.</p><p>Etymology. The pinkish colour of the animal is reflected in the name.</p><p>Discussion. Similar to cryophila, but differs most sharply in number of lateral setae on the ventral tube which is 5+ 5 in rosea and up to 15 on each side in cryophila . Also number of dorsal setae on dens is never more than 8 in rosea, while 12–14 in adult cryophila . In general the setal cover of the body is more open and abd. 5-6 less fused than in cryophila . Although variable, rosea usually has five basomedian setae on labium (Fig. 67) while cryophila has four. Erect ventral sensilla on ant. 2–3 are present in full grown rosea (Figs. 65- 66), not in cryophila . PAO is more elongate in rosea, slightly longer than diameter of nearest ocellus (Fig. 64). The serrated macrochaetae on tip of abdomen are slightly longer in rosea, median macrochaetae on abd.5 often more than 3 times as long as inner edge of last claw (less than 3.0 in cryophila). Apical tooth of mucro is relatively smaller in rosea than in cryophila (compare Figs. 59 and 68).</p><p>Distribution and ecology. Judging from present records the species is widely distributed in the Rockies from Alaska to Washington and Alberta, appearing in damp plant litter close to cold water and snowfields.</p></div>	https://treatment.plazi.org/id/504F2D6CFFD2FF8597BEF734FAB01884	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fjellberg, Arne	Fjellberg, Arne (2010): Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A. Zootaxa 2513: 27-49, DOI: 10.5281/zenodo.196078
504F2D6CFFD5FF8497BEF5CEFB431F21.text	504F2D6CFFD5FF8497BEF5CEFB431F21.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Desoria olympica	<div><p>Desoria olympica sp. n.</p><p>(Figs. 71–76)</p><p>Type material (all in slides). Holotype (male) and 10 paratypes (in 4 slides) from " USA, Washington, Olympic National Park, Hurricane Ridge 47o59'N, 123o31'W, 10.IX.1983. Wet moss at snow edge, 1,800 m., A. Fjellberg 422/83" (INHS).</p><p>Description. Size up to 1.0 mm. Colour white, eye-spots black. Body shape as in two previous species, abd. 5–6 weakly separated. Characters of ocelli, labrum, labial and maxillary palps, maxillae and mandibles as in previous two species. PAO roundish, the size of a ocellus (Fig. 75). Frontoclypeal field with three setae (Fig. 76). Ant. 1 with only two ventroapical short sensilla in addition to a few setaceous ones, ant. 2–3 with only the standard set of sensilla. Ventromedian field of labium with 4 setae. Head with 3+3(4) postlabial setae. Body integument smooth; setal cover sparse and uniform. Macrochaetae distinct, smooth. Median macrochaetae on abd. 5 about 2.7 times as long as inner edge of last claw. Number of ordinary sensilla on tergites as 22/11135 (Fig. 71). Microsensilla as 11/001. Posterior sensilla on th. 2–abd. 4 set in the p-row; no ordinary sensillum laterally to the spine-like ms on abd. 3. Ventral tube with only 1+1 frontal setae; retinaculum with only two setae (10 specimens examined); ventral central field of manubrium with fewer than 20 setae (Fig. 74); dens with 7 dorsal setae; mucro 5-toothed (Fig. 73). Inner side of tib. 1-2 with three pairs of setae along median line (Fig. 72). Males present, reproductive specimens not seen.</p><p>Etymology. Named after the type locality, the Olympic National Park.</p><p>Discussion. The character which immediately identifies olympica is the reduced number of tergal sensilla (22/11135), which is probably the lowest number observed in any Desoria . Unlike the two previous species the posterior sensilla on th. 2–abd. 4 are set in the p-row. It also differs by the reduced number of antennal sensilla and the low number of setae on ventral tube and retinaculum (Table 1).</p><p>Distribution and ecology. This distinct species is only known from the type series which appeared in a wet snow edge community in the coastal mountains above Port Angeles, Washington.</p></div>	https://treatment.plazi.org/id/504F2D6CFFD5FF8497BEF5CEFB431F21	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fjellberg, Arne	Fjellberg, Arne (2010): Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A. Zootaxa 2513: 27-49, DOI: 10.5281/zenodo.196078
504F2D6CFFD4FF8497BEF5E4FA8E1A6E.text	504F2D6CFFD4FF8497BEF5E4FA8E1A6E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Desoria	<div><p>Desoria sp. A</p><p>Two specimens from grass and moss cushions in a melt-water stream close to snowfields at Comox Glacier (49o33'N, 125o21'W), Vancouver Island (A. Fjellberg, 26.VIII.1983) are almost identical to olympica with 6+6 ocelli, 4–5 frontoclypeal setae, bifurcate maxillary palp, open uniform setal cover with abd. 1–4 sensilla in prow, smooth long macrochaetae, ventral tube with 1–2 frontal setae and 5–6 lateral setae, retinaculum with 2 setae, manubrium with 10–12 ventral setae in median field and dens with 7 dorsal setae in proximal half; same shape of mucro and sensillary equipment of antennae as olympica . The differences are presence of 5+5 setae in the basomedian field of labium and increased number of macrosensilla on th. 2–abd. 5: 44/ 323454 in sp. A. These specimens may represent an undescribed new species but more specimens need to be examined.</p></div>	https://treatment.plazi.org/id/504F2D6CFFD4FF8497BEF5E4FA8E1A6E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fjellberg, Arne	Fjellberg, Arne (2010): Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A. Zootaxa 2513: 27-49, DOI: 10.5281/zenodo.196078
504F2D6CFFD4FF8497BEF264FD3418A1.text	504F2D6CFFD4FF8497BEF264FD3418A1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Desoria capra	<div><p>Desoria capra sp. n.</p><p>(Fig. 77)</p><p>Type material. Holotype (female, in slide) and paratype (male, slide) from "Alaska: Juneau. Mts. between Mendenhall Glacier and Montana Creek Trail, 58o29'N, 134o40'W, 13.VII.1980. On melt-water ponds, 975 m. alt., A. Fjellberg leg." (INHS).</p><p>Description. Size 1.2 mm (subadult female in slide). Eye-spots black, rest of body white. Ocelli 6+6. Labrum modified, with stiff setae set close together (somewhat like Gnathisotoma spinolabris, described below). Apical edge of labrum membranous, without visible ventral ciliation (more like rough sandpaper). Maxillary palp trifurcate. Maxillae modified, with prolonged lamellae; Lam.1 passes well beyond tip of capitulum and appears almost smooth, without visible ciliation. Third tooth of capitulum thin and membranous (Fig. 77). Labium with 4+4 basomedian setae. Ventral tube with 1+1 frontal and 3+3 lateral setae. Dens with 7 dorsal setae in proximal half. Tib. 1–2 shortened, inner side with 3+3 setae along median line (as Fig. 72). Abd. 5–6 fused.</p><p>Etymology. The name refers to the mountain goat ( Oreamnos americanus) which is abundant in the area where this new species was collected.</p><p>Discussion. The species shares essential characters with cryophila and rosea in the 6+6 ocelli, small roundish PAO with three associated setae, "double" setal cover with distinct macrochaetae (as Fig. 30), abdominal sensilla inserted anterior to the p-row, manubrium with abundant (&gt;50) setae in the ventral median field, mucro compact with 5 teeth. It differs sharply from other species of this group by having a trifurcate maxillary palp and modified labrum and maxillae. Despite that only two specimens are known, the species is so distinct that a formal description seems justified.</p></div>	https://treatment.plazi.org/id/504F2D6CFFD4FF8497BEF264FD3418A1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fjellberg, Arne	Fjellberg, Arne (2010): Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A. Zootaxa 2513: 27-49, DOI: 10.5281/zenodo.196078
504F2D6CFFD6FF8697BEF08EFA871C89.text	504F2D6CFFD6FF8697BEF08EFA871C89.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Desoria	<div><p>Desoria sp. B</p><p>One specimen collected together with olympica at the type locality differs from that species by its larger size (1.2 mm) and a denser uniform cover of setae with macrochaetae hardly differentiated from ordinary setae. Median abd. 5 macrochaeta is only 1.8 times as long as inner edge of last claw, sensilla of the abdominal tergites are inserted in the p-row, but number is unclear (not as reduced as in olympica). None of the other species of the cryophila group have so short uniform setal cover as this probably undescribed species.</p></div>	https://treatment.plazi.org/id/504F2D6CFFD6FF8697BEF08EFA871C89	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fjellberg, Arne	Fjellberg, Arne (2010): Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A. Zootaxa 2513: 27-49, DOI: 10.5281/zenodo.196078
504F2D6CFFD6FF8697BEF1FCFA4B19E1.text	504F2D6CFFD6FF8697BEF1FCFA4B19E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gnathisotoma Cassagnau 1957	<div><p>Gnathisotoma Cassagnau, 1957</p><p>Cassagnau (1957) established the genus Gnathisotoma for the new species bicolor which was found in the central Pyrenees. Later Najt (1981) redescribed bicolor and described a new species G. deharvengi from snowfields in the Pyrenees and reported another possibly new species (known only from ecomorphic specimens) from the Cantabrian mountains in Spain. Yoshii (1990) described two more species from glaciers in Chile, G. patagonica and G. pallens, and also included the Antarctic species Isotoma klovstadi Carpenter, 1902 in Gnathisotoma . Stevens et al. (2006) transferred klovstadi to the genus Desoria . Further species of Gnathisotoma are not known.</p><p>In the original diagnosis of the genus Cassagnau (1957) stressed the modified maxillae of bicolor. Najt (1981) defined Gnathisotoma on the reduced number of sensilla on the tergites, the modified maxilla and the absence of frontal setae on the ventral tube. Yoshii (1990) used the subsegmentation in proximal part of dens as a single diagnostic character for Gnathisotoma . Potapov (2001) used the presence of four long maxillary lamellae and the absence of anterior setae on ventral tube as diagnostic characters separating Gnathisotoma from Desoria . However, none of the above characters are clearly inclusive of all species and unique to the genus. There is only a gradual difference between the maxilla of the Pyreneean Gnathisotoma and some Desoria species (tigrina -group). Also some Desoria have the sensillary equipment of tergites as much reduced as in Gnathisotoma . The significance of the ventral incision (subsegmentation) at base of dens is unclear on a generic level. The presence of the Gnathisotoma described below from Alaska with maxilla almost identical to bicolor but with 1–2 frontal setae on ventral tube and an increased number of tergal sensilla make the situation more confused and questions the status of Gnathisotoma as separate genus from Desoria .</p></div>	https://treatment.plazi.org/id/504F2D6CFFD6FF8697BEF1FCFA4B19E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fjellberg, Arne	Fjellberg, Arne (2010): Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A. Zootaxa 2513: 27-49, DOI: 10.5281/zenodo.196078
504F2D6CFFD6FF9997BEF523FA9C1864.text	504F2D6CFFD6FF9997BEF523FA9C1864.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gnathisotoma spinolabris	<div><p>Gnathisotoma spinolabris sp. n.</p><p>(Figs. 78–92, 114, 115G)</p><p>Type material (all in alcohol, sex not checked). Holotype and 50 paratypes from "Alaska: Juneau. Mts. near Mendenhall Glacier, 58o29'N, 134o40'W, 13.VII.1980. On melt–water ponds, 975 m. alt., A. Fjellberg leg." (INHS).</p><p>Description. Body size up to 1.2 mm. Body shape slender, abdomen slightly pear-shaped, widening towards tip of abdomen, furca long (Fig. 115G). Head distinctly prognathous, with protruding mouth (Fig. 81). Abd. 5–6 fused without incision between the segments but with a clear gap in setal cover. Colour uniformly bluish black with white antennae, including the first segment. Tip of ant. 4 with a diffuse dark spot. Occipital area of head with a large crown-shaped white spot (Figs. 114, 115G). Legs pale beyond coxae, dens pale. Head with 6+6 large ocelli A–F, a small H ocellus also present. PAO elongated, slightly longer than diameter of nearest ocellus. Three setae associated with PAO (Fig. 83). Ant. 1 on ventral side with one short apical and three longer sensilla as well as 2–3 microsetae (Fig. 79). Ant. 2 with two lateral poorly differentiated setaceous sensilla near apex. Ant. 3 organ with two large pointed sensilla and two slender guards (Fig. 84). In lateral position on ant. 3 one small and one large sensillum near apex. Ant. 4 with bifurcate subapical pin seta, the subapical organite as a small peg in a pit. Labrum with 4 prelabral setae and 5,5,4 stiff spine-like setae inserted in distinct papillae (Fig. 80); apical edge with 4 low ridges; ventroapical ciliation very weak, in the shape of a simple irregular row. Central field of frontoclypeal area with 4–5 setae, two setae above posterior corner of labrum. Maxillary palp simple, with 4 stiff sublobal setae set close together. Labial palps modified, each split in two parts consisting of papillae A–B and C–E respectively (Fig. 90). Terminal sensilla of the papillae short; guards b3–b4 enlarged, strongly curved; guard e7 absent; lateral process (l.p.) on papilla E small, finger-like; four proximal setae present. Hypostomal papillae with a tiny seta-like H while h1–h2 are strong, curved (Fig. 91). Basal fields of labium with 4 median and 5 lateral setae. Head with 5–6 postlabial setae on each side of ventral line. Mandibles normal, with strong teeth. Maxillae strongly modified with a narrow 3-toothed capitulum and elongated lamellae passing tip of capitulum (Figs. 88–89); Lam.1 and 4 appear thin and transparent, without visible denticles or ciliations; Lam.2 and 5 have fine denticles like the short Lam. 3 and 6. Body integument smooth without visible granulation. Ground setal cover of body consists of rather long, thick, smooth setae. Macrochaeta strong, smooth, stiff (Fig. 82). Macrosensilla on tergites present as 55/55696, with some variation on abd. 3–5. All abdominal tergites with one or more (abd. 4) sensilla set in anterior position (Fig. 78). Posterior sensilla on abd. 1–4 inserted in the prow. Spine-like microsensilla as 10/000. Ventral tube 1+1(2) frontal, 3–4 lateral and 1–3 caudal setae. No ventral setae on thorax. Retinaculum with 4+4 teeth and 4–5 setae. Manubrium with blunt apical teeth and 0– 3 short ventroapical setae on each side; ventromedian part without or at most with 1–2 setae near base (Fig. 85). Dens dorsally crenulated, usually with 8 dorsal setae in basal 2/3 and one minute subapical setula (Fig. 87); ventroapical seta not differentiated. Mucro short and compact, with 4 sharp teeth and a large outer basal seta; inner lamella extends to tip of subapical tooth, not transformed to a secondary tooth (Fig. 92). Tibiotarsi with 11 apical setae (A1–7, T1–4), all acuminate. Inner side of tib.1–2 with 3+3 setae along the median line (as Fig. 72). Claws small, straight, inner tooth absent, lateral teeth very small or absent; unguiculus with a high dorsal lamella on last claw, less distinct on other claws (Fig. 86). Reproductive males without differentiated setal cover (nonepitokous).</p><p>Etymology. The name reflects the unusually "spiny" appearance of the labrum.</p><p>Discussion. Until the generic relationship of Gnathisotoma and Desoria is resolved, the present species is tentatively placed in the former genus due to similarity in maxilla and the small number of anterior setae on manubrium shared with the type species of the genus. It differs from the two Pyrenean species by presence of a mucronal seta and an increased number of tergal sensilla. The two South American species are not precisely described, but their colour is uniformly black including the antennae in patagonica Yoshii, and both are said to have a 3–toothed mucro apparently without lateral seta. Also one maxilla is figured with a very prominent 3-toothed capitulum, unlike other Gnathisotoma . Ocelli are said to be 5+5, although figured with 6 by Yoshii (1990, Fig. 4 I).</p><p>Distribution and ecology. Only known from the type locality in the coastal mountains above Juneau, Alaska, where a sample was "scooped" from the surface of water draining from the melting snow fields.</p></div>	https://treatment.plazi.org/id/504F2D6CFFD6FF9997BEF523FA9C1864	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fjellberg, Arne	Fjellberg, Arne (2010): Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A. Zootaxa 2513: 27-49, DOI: 10.5281/zenodo.196078
504F2D6CFFC9FF9997BEF5AAFC5C1BF6.text	504F2D6CFFC9FF9997BEF5AAFC5C1BF6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mucronia	<div><p>Mucronia gen. n.</p><p>Type species: Mucronia enigmatica sp. n.</p><p>Diagnosis of the genus. The genus belongs to the family Isotomidae as defined by the following characters: Body elongated with normal setal cover, scales absent. Prothorax without dorsal setae. Mid-dorsal length of abdominal segment 3 slightly longer (1.2) than segment 4. Trochanteral organ absent. The two single characters which separate the new genus from most other genera of the family are presence of a clearly lobed PAO (Fig. 94) and a mucro bearing more than one seta (Fig. 102).</p></div>	https://treatment.plazi.org/id/504F2D6CFFC9FF9997BEF5AAFC5C1BF6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fjellberg, Arne	Fjellberg, Arne (2010): Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A. Zootaxa 2513: 27-49, DOI: 10.5281/zenodo.196078
504F2D6CFFC9FF9A97BEF759FB271BDF.text	504F2D6CFFC9FF9A97BEF759FB271BDF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mucronia enigmatica	<div><p>Mucronia enigmatica sp. n.</p><p>(Figs. 93–109)</p><p>Type material. Holotype: Reproductive male (slide) from "Alaska: Juneau. Mts. between Mendenhall Glacier and Montana Creek Trail, 58o29'N, 134o40'W, 13.VII.1980. On melt-water ponds, 975 m. alt., A. Fjellberg leg." Paratype: Reproductive male (slide, without head) from the same sample. Both specimens are transparent due to long storage in diluted ethanol and details of chaetotaxy are hard to see (INHS).</p><p>Description. Body size 0.95 mm (in slide). Body slender, with long furca (Fig. 93). Eye-spots dark, otherwise almost white. Frons slightly swollen. Abd. 5–6 not fused. Eye-fields with 6+6 large ocelli. PAO about the size of an ocellus, roundish, with four blunt lobes and swollen mid section (Fig. 94). Details of antennal chaetotaxy not seen, apart from ant. 3 organ which has two large sensilla and a ventrolateral spinelike sensillum (Fig. 99). Labrum with 5,5,4 papillate stiff setae, 4 prelabrals, anterior edge with four low ridges followed by a median tubercle (Fig. 97). Ventroapical ciliation not seen (if present presumably very fine). Frontoclypeal field with 10–12 setae. Maxillary outer lobe with trifurcate palp; sublobal setae unclear, but at least two present (Fig. 98). Labial palps modified, with papillae A–B, C–D and E in three separate groups (Figs. 95, 103). Terminal setae of the papillae short, not longer than surrounding guards. All guards present, including e7. Guards b4 and d4 enlarged, recurved at apex. Four proximal setae present. Hypostomal papilla reduced, H, h1 and h2 subequal. Basal fields of labium with 4 median and 5 lateral setae (Fig. 95). Postlabial setae 3–4 on each side. Mandibles normal, with strong molar plate and 4–5 apical teeth. Maxilla highly modified with 6 elongated lamellae and a narrow long stylet representing the capitulum, stylet with a small hook in apical 1/3 (Fig. 96); Lam.1 with weak denticles in apical 1/3; Lam.2 with rows of small sharp teeth along the edges only, extending from base to tip; Lam.3 present as a small flap at base of Lam.2, with delicate serration; Lam.4–5 with fine denticles from base to tip, not clearly defined in rows; Lam.6 short, with few denticles. Quality of body cuticle and setal cover difficult to evaluate because of transparency of the specimens, but setal cover apparently short and uniform. Macrochaetae only differentiated at abd. 5–6. Longest macrochaetae on abd. 5–6 about twice as long as inner length of claw 3. Macrochaetae apparently pointed and smooth, without serrations/ciliations. Because of transparency body sensilla not seen. Ventral tube apparently with 2+2 lateral (distal) setae, no frontals and two caudals (posterior). Retinaculum with 4+4 teeth and a single seta on corpus. Manubrial chaetotaxy not clearly seen, but with many setae both dorsal and ventral (Fig. 104); apical thickening with blunt teeth, short ventroapical setae 1+1. Dens with many (at least 20) ventral setae, dorsal side with fewer than 10 setae (Fig. 105). Mucro elongated, with three visible teeth, one of the basal teeth absent or reduced; a long and distinct inner lateral lamella ending at subapical tooth; two lateral setae present (Fig. 102). Tibiotarsi with 11 apical setae (A1–7, T1–4). Tibiotarsi on first two pairs of legs shortened, each with three regular whorls of setae (Fig. 100). Last pair of tibiotarsi longer, with increased number of setae (more than 30, Fig. 101). Claws small, without visible teeth; unguiculus pointed, with high basal lamella (Fig. 109). Reproductive males with distinct genital papillae, central part cone-shaped and separated from rest of the papilla, surrounded by 6 setae (Figs. 106–107); four associated setae possibly situated outside the papilla. Genital tract with two bulbous swellings (Fig. 108).</p><p>Etymology. Enigmatic: Not clearly understood.</p><p>Discussion. The remarkably transformed maxillae and the modified labial palps may actually not be significant on generic level. Similarly transformed palps and maxillae are commonly seen in several genera of Isotomidae ( Archisotoma, Parisotoma, Desoria) and probably reflects a specialisation to particular food or feeding habits. The simply lobed PAO and the bisetose mucro however, appears unique in the family. In the genera Micrisotoma Bellinger and Heteroisotoma Stach the PAO has a more irregular or complex fashion, different from the simple lobation observed in Mucronia . The complete ring of apical setae (11) on tibiotarsus probably represents a plesiomorphic condition in the family. The male genital papilla on the other hand, is very unlike the multisetosus type commonly found in Isotomidae and more resembles the types seen in the family Entomobryidae .</p><p>From the other side of the Bering Strait, in the alpine regions of the Aborigen mountains in Chukotka (Russian Far East), I have a sample (wet moss among rocks, 1,400 m alt., 23.VII.1979, V. Behan leg.) of what may be another species of the same genus. It has a similarly lobed PAO and bisetose elongated mucro, differing in its simple unmodified mouth parts, very long multiciliate macrochaetae and presence of dark body pigment. Also the mucro has two basal teeth and 1–2 secondary minute teeth between the subapical tooth and the basal teeth.</p><p>I have three specimens from Australia collected by P. Greenslade (NSW, 1.2 km NNE of S. Ramshead, 31o S, 148o E, subalpine pitfall trap, 1,800 m, July 1982, K. Green leg.) of another Isotomidae with elongate mucro bearing three outer setae and a complex set of teeth and spines, dens with many short spines on inner side in basal half, PAO roundish with a tendency to lobation.</p><p>These different species, all living in cold mountainous environments, may represent an old pre-Isotomidae line showing characters resembling both Isotomidae and Tomoceridae (mucro, dental spines).</p></div>	https://treatment.plazi.org/id/504F2D6CFFC9FF9A97BEF759FB271BDF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fjellberg, Arne	Fjellberg, Arne (2010): Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A. Zootaxa 2513: 27-49, DOI: 10.5281/zenodo.196078
504F2D6CFFCAFF9D97BEF736FA5C19D4.text	504F2D6CFFCAFF9D97BEF736FA5C19D4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Collembola	<div><p>Key to cryophilic Collembola of the genera Myopia, Gnathisotoma and Desoria of the North West Rocky Mountains</p><p>1 Body white or with weak pigment only; mucro with 5 teeth ....................................................................................... 2</p><p>- Body distinctly pigmented, in several species bluish black with contrasting white antennae; mucro with 3–5 teeth 7</p><p>2 Maxillary palp bifurcate; labrum and maxilla unmodified .......................................................................................... 3</p><p>- Maxillary palp trifurcate; labrum with stiff spine-like setae set close together; maxilla with elongated lamellae (Fig. 77) ................................................................................................................................................ Desoria capra sp. n.</p><p>3 Ventral tube with 6 or fewer setae on each side of the lateral flaps; dens with 7–8 dorsal setae ................................ 4</p><p>- Ventral tube with more than 6 (up to 15) lateral setae on each side; dens with more than 10 dorsal setae.................... ................................................................................................................................................. Desoria cryophila sp. n.</p><p>4 Ground cover of setae on head and body distinctly "double" (as Fig. 30) with short and long setae mixed .............. 5</p><p>- Ground cover of setae uniform, not clearly "double" .................................................................................................. 6</p><p>5 Abdominal macrochaetae long; M1 on abd. 5 at least 2.5 as long as inner edge of last claw........ Desoria rosea sp. n.</p><p>- Abdominal macrochaeta hardly developed; M1 on abd. 5 less than 2.0 as long as inner edge of last claw .................. .................................................................................................................................................................. Desoria sp. B</p><p>6 Sensilla on th. 2–abd. 5 distributed as 22/11135; basomedian field of labium with 4 setae ... Desoria olympica sp. n.</p><p>- Sensilla on th. 2–abd. 5 distributed as 44/32345; basomedian field of labium with 5 setae ................... Desoria sp. A</p><p>7 Tibiotarsi with 9 apical setae (Fig. 14); mucro apparently 3-toothed (Fig. 17) .................................... Desoria uniens</p><p>- Tibiotarsi with 11 apical setae; mucro with at least 4 distinct teeth ............................................................................ 8</p><p>8. Anterior two ocelli displaced anteriorly; PAO small and narrow, almost hidden, with one associated seta (Figs. 2–3) ............................................................................................................................................................ Myopia alaskana</p><p>- Anterior two ocelli not so strongly displaced; PAO roundish, distinct, with more than one associated seta.............. 9</p><p>9. Manubrium with at most 1–2 midventral setae (Fig. 85); mucro with lateral seta present (Fig. 92); maxillae with 4 lamellae extending beyond tip of capitulum (Figs. 88–89) ....................................... Gnathisotoma spinolabris sp. n.</p><p>- Manubrium with more than two midventral setae; lateral seta absent on mucro; maxillae with at most one lamella extending beyond tip of capitulum ............................................................................................................................ 10</p><p>10. Abdominal macrochaetae very long and densely ciliated, those on abd. 5 more than 3 times as long as inner edge of last claw ........................................................................................................................................... Desoria ater sp. n.</p><p>- Abdominal macrochaetae shorter, not densely ciliated, those on abd. 5 less than 3 times as long as inner edge of last claw ............................................................................................................................................................................. 11</p><p>11. Frontoclypeal field with more than 30 setae (Fig. 53). Mucro with 4 teeth ............................ Desoria pilifrons sp. n.</p><p>- Frontoclypeal field with fewer than 30 setae. Mucro with 4 or 5 teeth...................................................................... 12</p><p>12. Whole body black, including antennae; no white spots on head between antennae. Mucro with 4 teeth...................... ................................................................................................................................................ Desoria garibaldii sp. n.</p><p>- Antennae with at least some segments contrastingly white. Mucro with 5 teeth ....................................................... 13</p><p>13. Ground cover of body setae "double" (as Fig. 30), with short and long setae; head with two large triangular pale spots between eyes (Figs. 110, 115 E) ................................................................................ Desoria triangularis sp. n.</p><p>- Ground cover of body setae simple, of uniform length; head with 4 small pale spots between eyes (Fig. 111)............ ................................................................................................................................................ Desoria albicornis sp. n.</p></div>	https://treatment.plazi.org/id/504F2D6CFFCAFF9D97BEF736FA5C19D4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fjellberg, Arne	Fjellberg, Arne (2010): Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A. Zootaxa 2513: 27-49, DOI: 10.5281/zenodo.196078
