identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
D1A0C82594575F54AEDB6ED77B84550D.text	D1A0C82594575F54AEDB6ED77B84550D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Agestrata Eschscholtz 1829	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  Agestrata Eschscholtz, 1829</p>
            <p>Type species.</p>
            <p> Agestrata luconica Eschscholtz, 1829 </p>
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	https://treatment.plazi.org/id/D1A0C82594575F54AEDB6ED77B84550D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
3020A26A575B5B66B61A6E8DCCA01060.text	3020A26A575B5B66B61A6E8DCCA01060.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Agestrata orichalca subsp. orichalca (Linnaeus 1769) orichalca (Linnaeus 1769	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Agestrata orichalca orichalca (Linnaeus, 1769) Fig. 11 </p>
            <p> Scarabaeus orichalcus Linnaeus, 1769: 504. </p>
            <p>Distribution.</p>
            <p> North-eastern India (Sikkim and Arunachal Pradesh), the Chinese provinces of Guangdong, Guangxi and Hainan, the Hong Kong SAR and Taiwan. Also widespread in the Oriental Region (  Bezděk 2016), occurring specifically in Myanmar, Vietnam, Laos, Thailand, Malaysia and the Indonesian islands of Sumatra and Java (Sakai and Nagai 1998; Krajcik 2011). </p>
            <p>Material examined.</p>
            <p>  1♂: Macau,  University of East Asia, 28 May 1989, ER Easton leg (UMEC)  ;   1♂:  Coloane , Cheoc Van, 15 Jun 2019, crushed on road under street light, R Perissinotto &amp; L Clennell (MACT); unknown sex: Alto de Coloane, 18 Oct 2020, elytron found under spot-light, R Perissinotto &amp; L Clennell (MACT)  . </p>
            <p>Other Macau records.</p>
            <p> Taipa, University of East Asia Campus, 3 Sep 1991, near library (in Easton 1991: 111, misspelt as  Agestrata orichalea ); No locality and date, 42 mm (in Pun and Batalha 1997: 65, fig. 107, misspelt as  Agestrata orichalcea ); Alto de Coloane, 18 Aug 2020, on spot-light surface, R Perissinotto &amp; L Clennell; St. Francis  Xavier’s Parish [Coloane], 18 Aug 2020, 14:55, L Clennell (https://www.inaturalist.org/observations/56913518). </p>
            <p>Remarks.</p>
            <p> This is by far the largest cetoniine beetle in Macau, reaching a total length of 40-45 mm and a maximum width of 18-20 mm. Although it is regularly recorded in nearby Hong Kong (see e.g., https://www.inaturalist.org/observations?place_id=7613&amp;subview=grid&amp;taxon_id=127588), it is a rare occurrence in Macau. During this study only two males were recorded, one crushed on a road under a street light and a second, which also died after flying into an incandescent spot-light at the Coloane  A-Mà statue. The remnants of a third specimen were also retrieved in October 2020 under the same spot-light. According to Yiu (2010), this species feeds on a variety of fruits in captivity and is attracted to artificial light at night. The 3rd instar larva of this species was comprehensively described and illustrated by Zhang (1984). The larval stage is most probably strictly saproxylic, and thereby depends on availability of decomposing tree trunks, which are rapidly disappearing in the area as more and more parts of the remaining natural vegetation are converted to city parks and gardens. </p>
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	https://treatment.plazi.org/id/3020A26A575B5B66B61A6E8DCCA01060	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
5C4E5377BD035F4FA7C58E4529521255.text	5C4E5377BD035F4FA7C58E4529521255.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cetonia (Calopotosia) Reitter 1899	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Subgenus  Cetonia Calopotosia Reitter, 1899</p>
            <p>Type species.</p>
            <p> Cetonia submarmorea Burmeister, 1842. </p>
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	https://treatment.plazi.org/id/5C4E5377BD035F4FA7C58E4529521255	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
B08623E36E6656A48744FA3308F7133A.text	B08623E36E6656A48744FA3308F7133A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cetonia (Glycyphaniola) Mikšić 1968	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Subgenus  
Cetonia Glycyphaniola 
Miksic
, 1968
</p>
            <p>Type species.</p>
            <p> Cetonia modesta Fabricius, 1792 </p>
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	https://treatment.plazi.org/id/B08623E36E6656A48744FA3308F7133A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
0E294E324D15509EACFFAA30CB116B14.text	0E294E324D15509EACFFAA30CB116B14.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cetonia (Liocola) CG Thomson 1859	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Subgenus  Cetonia Liocola CG Thomson, 1859</p>
            <p>Type species.</p>
            <p> Cetonia marmorata Fabricius, 1792 </p>
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	https://treatment.plazi.org/id/0E294E324D15509EACFFAA30CB116B14	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
2CC77E4D1ABC592CBFE2AFD71C9B897B.text	2CC77E4D1ABC592CBFE2AFD71C9B897B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cetonia (Potosia) Mulsant & Rey 1871	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Subgenus  Cetonia Potosia Mulsant &amp; Rey, 1871</p>
            <p>Type species.</p>
            <p> Cetonia floricola Herbst, 1790 (=  Cetonia metallica Herbst, 1782) </p>
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	https://treatment.plazi.org/id/2CC77E4D1ABC592CBFE2AFD71C9B897B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
AD64D9ADA7A9544296A909B66124ED56.text	AD64D9ADA7A9544296A909B66124ED56.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cetonia (Protaetia) Burmeister 1842	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Subgenus  Cetonia Protaetia Burmeister, 1842</p>
            <p>Type species.</p>
            <p> Cetonia mandarina Weber, 1801 (=  Cetonia fusca Herbst, 1790) </p>
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	https://treatment.plazi.org/id/AD64D9ADA7A9544296A909B66124ED56	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
855B88D131685C63B3187FFC015D2B81.text	855B88D131685C63B3187FFC015D2B81.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euselates (Euselates) J Thomson 1880	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Subgenus  Euselates Euselates J Thomson, 1880</p>
            <p>Type species.</p>
            <p> Euselates magna J Thomson, 1880 </p>
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	https://treatment.plazi.org/id/855B88D131685C63B3187FFC015D2B81	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
A55004E58C9353CDA26A332D6E806AB7.text	A55004E58C9353CDA26A332D6E806AB7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euselates (Euselates) magna J Thomson 1880	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Euselates (Euselates) magna J Thomson, 1880 Fig. 13 </p>
            <p> Euselates magna J. Thomson, 1880: 277 </p>
            <p>Distribution.</p>
            <p> Known in the Palearctic Region from the Chinese provinces of Hainan, Hubei and the Hong Kong SAR (  Bezděk 2016). It is also widespread in the Oriental Region specifically in Vietnam, Laos and Thailand (Sakai and Nagai 1998; Krajcik 2011). </p>
            <p>Material examined.</p>
            <p>  1♀:  Coloane ,  Hác-Sá , 14 May 2019, on flowers of  Psychotria serpens , R Perissinotto (MACT)  ;   1♂, 1♀:  Coloane Village , 2 Jul 2019, dead on roadside, R Perissinotto &amp; L Clennell (MACT)  . </p>
            <p>Other Macau records.</p>
            <p> Coloane,  Hác-Sá , 28 Apr 2019, on flowers of  Lonicera japonica , R Perissinotto &amp; L Clennell; ibidem 3 May 2019, on flowers of  Psychotria serpens , R Perissinotto &amp; L Clennell; ibidem 15 May 2020, R Perissinotto; St. Francis  Xavier’s Parish [Coloane], 16 May 2020 10:35, L Clennell (https://www.inaturalist.org/observations/56121519); ibidem 11 Jul 2020 9:45, Kisu Wong (https://www.inaturalist.org/observations/57338916); ibidem 11 Jul 2020 10:45, Kit Chang (https://www.inaturalist.org/observations/52662946). </p>
            <p>Remarks.</p>
            <p> This species has been previously reported from nearby Hong Kong with its synonymic name of  E. schoenfeldti Kraatz, 1893 (Yiu 2010; Yiu and Yip 2011). In Macau it is occasionally seen between late April and August, but only in the largest patches of natural vegetation. Adults exhibit an approximate size in the range of 19-22 mm TL and 8-10 mm MW. They are very consistent in their colour pattern and are active during the hottest part of the day, but even under overcast conditions. They are typically flower feeders, having been observed on  Litsea glutinosa ,  Lonicera japonica ,  Psychotria serpens and in Hong Kong also on  Cleistocalyx operculatus (Yiu 2010). On one occasion, a male specimen was found inside a fruit-baited trap containing a mixture of fermenting banana, pineapple, brown sugar, and red wine. The larvae are unknown, but most likely saproxylic, as on two occasions females were observed while entering crevices in decomposing tree trunks. </p>
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	https://treatment.plazi.org/id/A55004E58C9353CDA26A332D6E806AB7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
5AA76D5FEDF1535AB3E0D90DC7973426.text	5AA76D5FEDF1535AB3E0D90DC7973426.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euselates J Thomson 1880	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  Euselates J Thomson, 1880</p>
            <p>Type species.</p>
            <p> Euselates magna J Thomson, 1880 </p>
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	https://treatment.plazi.org/id/5AA76D5FEDF1535AB3E0D90DC7973426	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
96856314531059E5937E885D807988DA.text	96856314531059E5937E885D807988DA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gametis bealiae (Gory & Percheron 1833)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Gametis bealiae (Gory &amp; Percheron, 1833) Fig. 3 </p>
            <p> Cetonia bealiae Gory &amp; Percheron, 1833: 282. </p>
            <p>Distribution.</p>
            <p> Within the Palearctic Region,  G. bealiae is known from the Chinese provinces of Fujian, Guangdong, Hebei, Hubei, Jiangxi, Zhejiang and the Hong Kong SAR (Yiu 2010; Lirong et al. 2013;  Bezděk 2016). It is also widely distributed across the Oriental Region, specifically in Myanmar, Laos, northern Vietnam and north-eastern India (Sakai and Nagai 1998; Krajcik 2011). </p>
            <p>Material examined.</p>
            <p>  1♂:  Coloane Village , 29 Jun 2019, in fruit-baited trap, R Perissinotto &amp; L Clennell (MACT)  . </p>
            <p>Other Macau records.</p>
            <p> Taipa Pequena, 4 Mar 2019, on flowers of  Ligustrum sinense , R Perissinotto; Coloane,  Hác-Sá , 4 Apr 2019, on flowers of  L. sinense , R Perissinotto &amp; L Clennell; ibidem 11 Apr 2020, R Perissinotto &amp; L Clennell. </p>
            <p>Remarks.</p>
            <p> In Macau, this species varies in size between 12.5 and 15 mm in TL and between 7 and 9 mm in MW. Colour forms range from black to olive green background, with testaceous to reddish green pronotum and ochreous to testaceous mid-elytral bands. The white maculation on the general surface appears to be consistent. During 2019, one individual was retrieved inside a fruit-baited trap, while four others were observed feeding on flowers of  Ligustrum sinense . In 2020, only one specimen was observed while feeding on flowers of  L. sinense . In Macau, this species appears to have its peak of adult activity between early spring and early summer, while no specimens have been recorded in late summer, autumn, or winter. In nearby Hong Kong, this species has been recorded feeding on flowers of  Viburnum odoratissimum and  Lonicera sp. between March and May (Yiu 2010) as well as on flowers of  Zanthoxylum avicennae in October (https://www.inaturalist.org/observations/62885551). No information is available on larval or pupal stages in the region. </p>
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	https://treatment.plazi.org/id/96856314531059E5937E885D807988DA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
36ACDBAFF53654C2AD0D08838805A569.text	36ACDBAFF53654C2AD0D08838805A569.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gametis Burmeister 1842	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  Gametis Burmeister, 1842</p>
            <p>Type species.</p>
            <p> Cetonia versicolor Fabricius, 1775. </p>
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	https://treatment.plazi.org/id/36ACDBAFF53654C2AD0D08838805A569	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
1431720A06FD5F17A9D2C29EF3512AC2.text	1431720A06FD5F17A9D2C29EF3512AC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gametis jucunda (Faldermann 1835)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Gametis jucunda (Faldermann, 1835) Fig. 4 </p>
            <p> Cetonia jucunda Faldermann, 1835: 386. </p>
            <p>Distribution.</p>
            <p> Mainly a Palearctic species, recorded from the Chinese provinces and municipalities of Beijing, Chongqing, Fujian, Gansu, Guizhou, Guangxi, Hainan, Hebei, Heilongjiang, Hubei, Jiangsu, Jiangxi, Liaoning, Nei Mongol, Sichuan, Shanghai, Shandong, Shanxi, Yunnan, Zhejiang and the Hong Kong SAR (Lirong et al. 2013;  Bezděk 2016). Also found in Pakistan, Nepal, the Indian Sikkim Province, North and South Korea, Mongolia, Russian Far East and Japan (Sakai and Nagai 1998; Krajcik 2011;  Bezděk 2016). According to  Bezděk (2016), it also occurs in the Oriental Region. </p>
            <p>Material examined.</p>
            <p> 1♂, 1♀: Coloane,  Hác-Sá , 8 Apr 2020, on flowers of  Ligustrum sinense , R Perissinotto (MACT). </p>
            <p>Other Macau records.</p>
            <p> No locality and date, 14 mm (in Pun and Batalha 1997: 65, fig. 108 as  Oxycetonia jucunda Faldermann); Taipa Pequena, 11 Mar 2019, on flowers of  Toddalia asiatica , R Perissinotto &amp; L Clennell. </p>
            <p>Remarks.</p>
            <p> A rare species in Macau, despite its widespread distribution and common occurrence in the surrounding regions, such as the Hong Kong SAR (Yiu 2010; iNaturalist observations). The dorsal background colour is always predominantly green, ranging from bright to olive grade, while the white maculation can vary somehow in extent. In particular, the discal spots on both pronotum and elytra can fade completely in exceptional cases. Specimens range in size within the approximate range of 12-15 mm TL and 6-8 mm MW. The period of adult activity appears to peak in the spring and no specimens have been recorded in Macau during summer or autumn yet, although their occurrence during these seasons is well established in nearby Hong Kong (Yiu 2010). In Macau, adults were observed feeding on flowers of  Ligustrum sinense ,  Clausena lansium and  Toddalia asiatica , while in Hong Kong they were also found on  Guilandina bonduc ,  Rhus chinensis and  Schima superba (Yiu 2010). The 1st-3rd instar larvae of  G. jucunda have been comprehensively described and illustrated by Murayama (1931), Medvedev (1952), Zhang (1984) and Sawada (1991) (  Šípek and  Král 2012). </p>
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	https://treatment.plazi.org/id/1431720A06FD5F17A9D2C29EF3512AC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
BFBCBBF527715F59B005C6CBF791338D.text	BFBCBBF527715F59B005C6CBF791338D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Glycyphana) horsfieldii Hope 1831	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Glycyphana (Glycyphana) horsfieldii Hope, 1831 Fig. 5 </p>
            <p> Cetonia horsfieldii Hope, 1831: 25. </p>
            <p>Distribution.</p>
            <p> Known in the Palearctic Region from the Himalayan countries of Nepal, Bhutan, and the Sikkim Province of India, as well as the Chinese provinces of Guizhou, Jiangxi, Yunnan, and the island of Taiwan (  Bezděk 2016). It has only recently been recorded from Hong Kong for the first time (Aston and Melsom 2019). In the Oriental Region, it has been reported from Myanmar, Vietnam, Thailand, Laos, India and Sri Lanka (Sakai and Nagai 1998; Krajcik 2011). </p>
            <p>Material examined.</p>
            <p>  1♂:  Taipa Monument , 27 Sep 2018, dead on path, R Perissinotto (MACT)  ;   1♀:  Coloane Village , 28 Aug 2019, on flowers of  Zanthoxylum avicennae , R Perissinotto &amp; L Clennell (MACT)  . </p>
            <p>Other Macau records.</p>
            <p> Macau, Guia Hill, 25 Oct 2017, R Perissinotto &amp; L Clennell; Taipa Pequena, 4 May 2018, R Perissinotto &amp; L Clennell; ibidem 11 Oct 2018, R Perissinotto &amp; L Clennell; Macau Peninsula, 22 Dec 2019, Angus Chan (pers. comm.); Coloane,  Hác-Sá , 7 Apr 2020, on flowers of  Ligustrum sinense , R Perissinotto &amp; L Clennell. </p>
            <p>Remarks.</p>
            <p> This species appears to be a new record for the broader region, having also been observed for the first time in Hong Kong only in April 2018 (Aston and Melsom 2019). The subspecies  G. h. chinensis , originally described by  Mikšić (1970) based on its wider extent of the red pronotal margin over the nominal subspecies (e.g., Ma 1995), is no longer recognised and has been synonymised with  G. horsfieldii (  Jákl in  Löbl and  Löbl 2016: 18). Specimens in Macau range in size from approximately 13 to 16 mm TL and from 6 to 8 mm MW. The period of adult activity seems to extend throughout the year, but most observations are from the spring (Apr-May) and autumn (Sep-Oct) months. While the larval stages are not known, adults have been recorded on flowers of  Ligustrum sinense ,  Zanthoxylum avicennae ,  Shefflera heptaphylla and  Homalium cochinchinense . </p>
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	https://treatment.plazi.org/id/BFBCBBF527715F59B005C6CBF791338D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
1B90E994423E5E258F46F4B780F76D20.text	1B90E994423E5E258F46F4B780F76D20.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Glycyphaniola) laotica Miksic 1968	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Glycyphana (Glycyphaniola) laotica 
Miksic
, 1968
 Fig. 6 </p>
            <p> Glycyphaniola nicobarica laotica Mikšić , 1968: 48. </p>
            <p>Distribution.</p>
            <p> According to  Jákl (in  Löbl and  Löbl 2016: 18), it is currently known from the Hong Kong SAR and parts of the Oriental Region. It is likely that it occurs through much of southern China. </p>
            <p>Material examined.</p>
            <p>  1♀:  Taipa Central , October 2017, Jeff Lei (MACT)  ;   1♂:  Barra Hill , 5 May 2018, dead on roadside, L Clennell (MACT)  ;   1♂:  Taipa Pequena , 26 Sep 2018, on flowers of  Rhus chinensis by roadside, R Perissinotto &amp; L Clennell (MACT)  ;   1♀:  Coloane Village , 13 Mar 2019, on flowers of  Bidens sp., R Perissinotto (SJCP)  ; 1♂: ibidem 13 Apr 2019, R Perissinotto (SJCP). </p>
            <p>Other Macau records.</p>
            <p> Taipa Pequena, 26 Oct 2017, on flowers of  Rhus chinensis , R Perissinotto &amp; L Clennell; ibidem 2 Mar 2020; Coloane Village, 2 Jul 2019, on flowers of  Bidens sp., R Perissinotto &amp; L Clennell; ibidem, 29 Sep 2018, R Perissinotto &amp; L Clennell; Coloane,  Ká-Hó , 7 Oct 2018, R Perissinotto &amp; L Clennell; Macau, Guia Hill, 14 Mar 2020, on flowers of  Ligustrum sinense , R Perissinotto &amp; L Clennell; St. Francis  Xavier’s Parish, Macau [Coloane], 7 Jul 2019 13:04, Kit Chang (https://www.inaturalist.org/observations/28360614); ibidem 12 Apr 2020 13:29, L Clennell (https://www.inaturalist.org/observations/55132359); ibidem 27 Sep 2020 14:59, L Clennell (https://www.inaturalist.org/observations/60940151); ibidem 30 Sep 2020, L Clennell (https://www.inaturalist.org/observations/61293565); Our Lady of  Carmel’s Parish [Taipa], 17 Aug 2020 14:14, L Clennell (https://www.inaturalist.org/observations/56723739); ibidem 14 Sep 2020 16:07, Kit Chang (https://www.inaturalist.org/observations/59561942); Circuito da Barragem de  Hac-Sá , Coloane, 4 Jul 2020 15:26, Annie Lao (https://www.inaturalist.org/observations/51892005). </p>
            <p>Remarks.</p>
            <p> This taxon has recently been elevated to species rank by  Jákl (in  Löbl and  Löbl 2016: 18), on the basis of comparative studies of numerous specimens from continental Asia and the type material of  Glycyphana nicobarica Janson, 1877. Thus, all previous identifications of the local  G. (Glycyphaniola) species occurring in Macau and nearby Hong Kong almost certainly refer to this species rather than to  G. nicobarica , which is probably an endemic to the Nicobar Islands (S  Jákl , pers. comm.). Specimens range in size between approximately 9-12 mm in TL and 5-6 mm in MW. The background colour of their body surface can vary from bright light green, to dark olive-green (Fig. 6) and in extreme cases even brown to brick-red. The extent of white maculation also varies substantially across both elytral and pronotal surfaces. </p>
            <p> Larvae, cocoons, and freshly emerged adults have been observed inside decomposing tree trunks and branches (pers. obs.), thereby confirming the saproxylic habit of its immature stages. In Hong Kong (then referred to as  Glycyphana nicobarica ), larvae were successfully reared to adulthood on fermented sawdust, and several adult specimens were found inside a compost heap composed of pig waste and sawdust (Yiu 2010). Adults have been recorded feeding on a variety of flowers, including  Rhus chinensis ,  Ligustrum sinense ,  Mallotus paniculatus ,  Murraya paniculata ,  Schefflera heptaphylla ,  Viburnum odoratissimum ,  Zanthoxylum avicennae ,  Bauhinia championii , and even on the alien invasive herbs  Bidens alba and  B. pilosa . They do not seem to be attracted to fruit-baited traps and have not been observed on sap flows either. This is one of the most often encountered species in the Macau SAR, but never in abundance. It occurs all year round, with distinct peaks of adult activity in the spring (Mar-Apr) and autumn (Sep-Oct). </p>
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	https://treatment.plazi.org/id/1B90E994423E5E258F46F4B780F76D20	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
45A7C5B2D95F56888687E5637CB83D3E.text	45A7C5B2D95F56888687E5637CB83D3E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana Burmeister 1842	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Subgenus:  Glycyphana Burmeister, 1842</p>
            <p>Type species.</p>
            <p> Cetonia horsfieldii Hope, 1831. </p>
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	https://treatment.plazi.org/id/45A7C5B2D95F56888687E5637CB83D3E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
C4D5F2D935F55383A13A51D117ADDF8D.text	C4D5F2D935F55383A13A51D117ADDF8D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Calopotosia) orientalis subsp. orientalis (Gory & Percheron 1833) orientalis (Gory & Percheron 1833	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Protaetia (Calopotosia) orientalis orientalis (Gory &amp; Percheron, 1833) Fig. 7 </p>
            <p> Cetonia orientalis Gory &amp; Percheron, 1833: 193 </p>
            <p>Distribution.</p>
            <p> Within the Palearctic Region, this species is known from the Chinese provinces of Chongqing, Fujian, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Sichuan, Shandong, Zhenjiang, the Hong Kong SAR, North and South Korea as well as the Russian Far East (Lirong et al. 2013;  Bezděk 2016). It is also widespread in the Oriental Region, specifically in northern India, the Kashmir region, northern Vietnam, and Laos (Sakai and Nagai 1998; Krajcik 2011). </p>
            <p>Material examined.</p>
            <p>  1♂, 1♀:  Coloane ,  Ká-Hó , 16 Jun 2018, dead on roadside, L Clennell (MACT)  ;   1♂:  Coloane ,  A-Mà Cultural Village, 13-15 Jun 2019, aggregation on sap of  Albizia lebbeck , R Perissinotto &amp; L Clennell (MACT)  ;   1♂: Macau,  University of East Asia Library, 4 May 1990, ER Easton leg (UMEC)  ; 1♀: ibidem, on building, 1 Aug 1989, ER Easton leg (UMEC); 1♂: ibidem 30 Jul 1989, ER Easton leg (UMEC); 1♂: ibidem, on building, 12 Jul 1989, ER Easton leg (UMEC); 1♀: ibidem no data, ER Easton leg (UMEC). </p>
            <p>Other Macau records.</p>
            <p> Taipa, University of East Asia Campus, near library, 28 May 1991 (Easton 1991: 111; 1993: 55); No locality and date, 18 mm (Pun and Batalha 1997: 66, fig. 109); Taipa Grande, 22 May 2018, L Clennell; Coloane, 10 May 2019, R Perissinotto &amp; L Clennell; ibidem 18 May 2019, R Perissinotto &amp; L Clennell; Macau, 15 Jun 2019, Peggi Chao (pers. comm.); ibidem 8 Aug 2019, Ben Wong (pers. comm.); Alto de Coloane, 23 May 2020, R Perissinotto &amp; L Clennell; ibidem 5 Jul 2020, feeding on sap of  Sapium discolor R Perissinotto &amp; L Clennell; Barra Hill, Macau, 9 Jul 2018 12:58, Kisu Wong (https://www.inaturalist.org/observations/23843295); Coloane Alto, Macao, 15 Jun 2019 16:11, L Clennell (https://www.inaturalist.org/observations/56299608); St. Francis  Xavier’s Parish [Coloane], 12 May 2019 10:27, Kit Chang (https://www.inaturalist.org/observations/24989100); ibidem 5 Jul 2020 13:56, L Clennell (https://www.inaturalist.org/observations/56110922); Avenida Doutor Stanley Ho [Macau], 24 May 2020 11:44, Benny Kuok (https://www.inaturalist.org/observations/47261733). </p>
            <p>Remarks.</p>
            <p> This is the only species currently seen in reasonable numbers in the Macau SAR, but only from late spring and throughout the summer. Specimens are generally of a bright green colour with golden sheen and white markings, but the background colour can turn olive-green or even darker in some specimens. Adult size varies within the approximate range of 20-25 mm in TL and 10-13 mm in MW. Its diet appears to be the most variable exhibited by any of the cetoniines encountered in Macau, with adults recorded in aggregations on sap flows of  Albizia lebbeck (Fig. 7) and  Sapium discolor . It also enters regularly into fruit-baited traps and has been observed feeding on wild figs and lychee fruits. Among the flowering trees that attract this species are  Acronychia pedunculata ,  Litsea glutinosa ,  Paliurus spina-christi and  Syzigium buxifolium . </p>
            <p>Easton (1993) reported that this species was regarded a minor pest, as the adults fed on lychees and other soft fruits such as peaches, while their grubs were believed to feed on the roots of grasses. Although adults are typically diurnal, with activity peaking in the hottest part of the day, they have also been found on the walls of buildings illuminated at night (Easton 1993). The 3rd instar larva of this species was comprehensively described and illustrated by Sawada (1991). In Hong Kong, larvae have been successfully reared in captivity using fermented sawdust as food (Yiu 2010).</p>
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	https://treatment.plazi.org/id/C4D5F2D935F55383A13A51D117ADDF8D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
6376EE7A9E1F5B80AB6B78FC9863A039.text	6376EE7A9E1F5B80AB6B78FC9863A039.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Liocola) speculifera (Swartz 1817)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Protaetia (Liocola) speculifera (Swartz, 1817) Fig. 8 </p>
            <p> Cetonia speculifera Swartz, 1817: 53. </p>
            <p>Distribution.</p>
            <p> In the Palearctic Region this species is only known from the Chinese provinces of Hainan, Henan, Hunan and Jiangxi (Sakai and Nagai 1998;  Bezděk 2016). It has also been recorded in the Oriental Region from northern Vietnam and Laos (Krajcik 2011). </p>
            <p>Material examined.</p>
            <p>  1♂:  Coloane ,  A-Mà Cultural Village, 19 Jun 2019, dead under tree, R Perissinotto &amp; L Clennell (MACT)  . </p>
            <p>Other Macau records.</p>
            <p> Alto de Coloane, 14 Jun 2019, on sap flow of  Albizia lebbeck , R Perissinotto &amp; L Clennell; ibidem 5 Jul 2020, on sap flow of  Sapium discolor , R Perissinotto &amp; L Clennell; Coloane Village, 8 Jul 2020, landing on tree trunk, R Perissinotto &amp; L Clennell. </p>
            <p>Remarks.</p>
            <p> This species has a rather sporadic occurrence in Macau, having been observed only twice in 2019 and always on sap flows of  Albizia lebbeck , and then again on another two occasions in 2020, on sap flows of  Sapium discolor and in hovering flight respectively (pers. obs.). Adult size varies in the approximate range of 20-23 mm TL and 11-13 mm MW. Although it has not been recorded formally from nearby Hong Kong, a few observations reported on the citizen science platform iNaturalist from that area (e.g., https://www.inaturalist.org/observations/25994523; https://www.inaturalist.org/observations/24358351; https://www.inaturalist.org/observations/1126433) indicate that this species [or the closely related  P. (L.) brevitarsis (Lewis, 1879)] may occur there too. It has probably been overlooked in the past, as superficially it resembles quite well  P. (C.) orientalis orientalis both in size and general appearance. Even an alerted observer needs to be within close distance in order to be able to appreciate the stockier body shape, the reduction of white maculation on the dorsal surface and the gold-red sheen that allow the diagnosis of this species against  P. (C.) orientalis orientalis . Adult activity of  P. (L.) speculifera in Macau seems to be restricted to the summer months. </p>
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	https://treatment.plazi.org/id/6376EE7A9E1F5B80AB6B78FC9863A039	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
278F35D2947B5C4F89C39516A146AB04.text	278F35D2947B5C4F89C39516A146AB04.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Potosia) intricata WW Saunders 1852	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Protaetia (Potosia) intricata WW Saunders, 1852 Fig. 9 </p>
            <p> Protaetia intricata WW Saunders, 1852: 31 </p>
            <p>Distribution.</p>
            <p> This species appears to be rather restricted geographically, having so far been recorded only in the Chinese provinces of Fujian and Zhejiang (  Bezděk 2016; Krajcik 2011). </p>
            <p>Material examined.</p>
            <p> 1♂: Coloane,  Ká-Hó , 2 Jun 2019, on flowers of  Syzigium buxifolium , R Perissinotto &amp; L Clennell (SJCP). </p>
            <p>Other Macau records.</p>
            <p> Coloane,  Ká-Hó , 29 May 2020, on flowers of  Syzigium buxifolium , R Perissinotto [identification uncertain]. </p>
            <p>Remarks.</p>
            <p> This is certainly the rarest cetoniine recorded so far in Macau, having been observed with certainty only once in June 2019, feeding on flowers of  Syzigium buxifolium . A second potential specimen was observed in a nearby locality on the same flowers in May 2020, but its identification could not be conclusively verified as it was too far above the ground. According to S.  Jákl (pers. comm.), this species is extremely rare throughout its limited distribution range and to our knowledge has not been reported from Hong Kong yet. The approximate size of the 2019 male specimen was 15 mm TL and 9 mm MW. </p>
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	https://treatment.plazi.org/id/278F35D2947B5C4F89C39516A146AB04	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
F5D2CBC4184A585AB5ECE328992F984E.text	F5D2CBC4184A585AB5ECE328992F984E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) fusca (Herbst 1790)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Protaetia (Protaetia) fusca (Herbst, 1790) Fig. 10 </p>
            <p> Cetonia fusca Herbst, 1790: 257 </p>
            <p>Distribution.</p>
            <p> Occurring widely in the Palearctic Region, especially the Chinese provinces and municipalities of Fujian, Guangdong, Guangxi, Hainan, Hubei, Jiangxi, Zhenjang, Shanghai, the Hong Kong SAR and the island of Taiwan as well as Japan and  India’s Sikkim Province (  Bezděk 2016). Also found in the Oriental, Australian, Afrotropical, and Pacific regions (  Bezděk 2016), particularly across SE Asia, Australia, New Guinea, Madagascar, Mauritius, Hawaii, Polynesia, Micronesia, and Melanesia (Sakai and Nagai 1998; Krajcik 2011). It has recently been intercepted in Florida and in the Caribbean countries of Bahamas and Barbados (Woodruff 2006), thus becoming a near-cosmopolitan species. </p>
            <p>Material examined.</p>
            <p>  1♂: Macau,  University of East Asia, Jun 1990, ER Easton leg (UMEC)  ;   1♂:  Coloane , Cheoc Van, 29 Jun 2019, crushed on sidewalk, R Perissinotto &amp; L Clennell (MACT)  . </p>
            <p>Other Macau records.</p>
            <p> Coloane,  Hác-Sá , 4 Apr 2019, on flowers of  Ligustrum sinense , R Perissinotto; Coloane Village, 17 Jul 2020, on building wall, L Clennell; Coloane, Oscar Farm, on rice stem, 24 Oct 2020, Kit Chang (pers. comm.). </p>
            <p>Remarks.</p>
            <p> Despite being one of the most worldwide spread cetoniine, this species is extremely scarce in Macau. Adults are active mainly in spring and summer and range in size from approximately 13 to 15 mm TL and from 8 to 9 mm MW. In Macau, they have been observed feeding only on flowers of  Ligustrum sinense , but the widely used common name of Asian mango flower beetle for the species indicates a diet with this staple component in its natural habitat. Globally, it has actually shown a very variable diet, including a multitude of flowers, fruits and even bee honey. In Hawaii, where it was first recorded in 1949 (Maehler 1950), it is regarded as a pest, causing damage to commercially cultivated roses, maize and a wide variety of flowers and fruits. The damage caused has been regarded significant enough to justify the introduction of parasitic wasps from other regions, in an effort to exert biological control over its rapidly expanding population (Woodruff 2006). The entire life cycle of this species, including egg, 1st-3rd instar larvae and pupa, was comprehensively described and illustrated by Simpson (1990). Larvae are compost feeders and have not been observed to attack roots of healthy plants and thus, unlike their adults, are not regarded as pests (Simpson 1990). </p>
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	https://treatment.plazi.org/id/F5D2CBC4184A585AB5ECE328992F984E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
BEABEF80690251F8BFE46AF6ADDA282E.text	BEABEF80690251F8BFE46AF6ADDA282E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia Burmeister 1842	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  Protaetia Burmeister, 1842</p>
            <p>Type species.</p>
            <p> Cetonia mandarina Weber, 1801 (=  Cetonia fusca Herbst, 1790). </p>
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	https://treatment.plazi.org/id/BEABEF80690251F8BFE46AF6ADDA282E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
B8D93851344556FBB732C0BB898130E2.text	B8D93851344556FBB732C0BB898130E2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thaumastopeus Kraatz 1885	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  Thaumastopeus Kraatz, 1885</p>
            <p>Type species.</p>
            <p> Lomaptera mohnikii J. Thomson, 1877 </p>
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	https://treatment.plazi.org/id/B8D93851344556FBB732C0BB898130E2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
D6D827F23D9C5CA2B1905FA902544A61.text	D6D827F23D9C5CA2B1905FA902544A61.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thaumastopeus shangaicus Neervoort van de Poll 1886	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Thaumastopeus shangaicus Neervoort van de Poll, 1886 Fig. 12 </p>
            <p> Thaumastopeus shangaicus Neervoort van de Poll, 1886: 181. </p>
            <p>Distribution.</p>
            <p> Known in the Palearctic Region from the Chinese provinces of Hainan and Yunnan, the Shanghai Municipality and the Hong Kong SAR (Yiu 2010;  Bezděk 2016). Also widespread in the Oriental Region, specifically in Vietnam, Thailand, Laos, peninsular Malaysia as well as Sumatra and the Nias Islands in Indonesia (Sakai and Nagai 1998; Krajcik 2011). </p>
            <p>Material examined.</p>
            <p>  1♀:  Coloane Village , 31 Mar 2019, extracted prematurely from broken cocoon, R Perissinotto &amp; L Clennell (MACT)  ;   1♀:  Coloane Village , 22 May 2020, on flowers of  Psychotria serpens , R Perissinotto (MACT)  ; 1♂: ibidem 28 May 2020, dead on roadside, R Perissinotto (MACT). </p>
            <p>Other Macau records.</p>
            <p> No locality and date, 23 mm [in Pun and Batalha 1997: 66, fig. 110, reported as  Thaumastopeus nigritus (Frohlic)]; Coloane Village, 14 Jun 2019, on flowers of  Paliurus spina-christi , R Perissinotto &amp; L Clennell; ibidem 30 Apr 2020, on flowers of  Psychotria serpens R Perissinotto &amp; L Clennell; Coloane,  Hác-Sá , 29 Apr 2019, emerged from cocoon found in decomposing wood, R Perissinotto &amp; L Clennell; Coloane, Cheoc Van, 4 May 2019, on flowers of  Litsea glutinosa , R Perissinotto &amp; L Clennell; Alto de Coloane, 23 May 2020, R Perissinotto &amp; L Clennell; ibidem 11 Jul 2020, on sap flow of  Sapium discolor , R Perissinotto &amp; L Clennell; Macau, Barra Hill, 16 Apr 2019, on flowers of  Ligustrum sinense , R Perissinotto &amp; L Clennell; Coloane Village, 1 Jun 2019 13:51, Hannah Leung (https://www.inaturalist.org/observations/27733112); Taipa Pequena, 23 May 2020 9:56, Eric Kwan (https://www.inaturalist.org/observations/47007347). </p>
            <p>Remarks.</p>
            <p> This is the second largest cetoniine species found in Macau, attaining a size in the range of 22-30 mm TL and 9-13 mm MW. Specimens are very stable in their colour, which is generally shiny and black with a dark blue sheen. It has been confused in the past with the closely related  T. nigritus (  Frölich , 1792) (e.g., Pun and Batalha 1997; Yiu 2010; Yiu and Yip 2011), which actually occurs mainly in the Indian subcontinent including the Himalayan region (Krajcik 2011;  Bezděk 2016). The correct identification of  T. shangaicus in the Macau and Hong Kong area has now been conclusively established through analysis of aedeagal material (S.  Jákl , pers. comm.). The species is saproxylic, with larvae, cocoons and even adults having been found regularly inside decomposing tree trunks and branches (pers. obs.). Adults appear to have a very broad diet, feeding on fruits, flowers, and sap flows, but do not seem to be attracted into fruit-baited traps. In Macau, several specimens were observed on sap flows of  Sapium discolor in July 2020 and in Hong Kong this species has often been found feeding on ripe fruits of longan and figs (Yiu 2010). Among the plants where  T. shangaicus has been recorded feeding on flowers are  Litsea glutinosa ,  Psychotria serpens ,  Paliurus spina-christi ,  Ligustrum sinense , and  Acronychia pedunculata . </p>
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	https://treatment.plazi.org/id/D6D827F23D9C5CA2B1905FA902544A61	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Perissinotto, Renzo;Clennell, Lynette	Perissinotto, Renzo, Clennell, Lynette (2021): Census of the fruit and flower chafers (Coleoptera, Scarabaeidae, Cetoniinae) of the Macau SAR, China. ZooKeys 1026: 17-43, DOI: http://dx.doi.org/10.3897/zookeys.1026.60036, URL: http://dx.doi.org/10.3897/zookeys.1026.60036
