identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
523B87B7FF95270606D6224BFF6AD13E.text	523B87B7FF95270606D6224BFF6AD13E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cebudonus	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Cebudonus n. gen.</p>
            <p> Type species.  Cebudonus poppeorum n. gen. , n. sp., by present designation. </p>
            <p>Diagnosis. Carapace pentagonal; dorsal surface gently convex, covered with numerous small pits (Figs. 1, 2); front with 2 long pseudorostral spines, subparallel, ca. 0.4 times total carapace length, separated by broad U-shaped cleft (Figs. 1, 2, 3 C); anterolateral margin entire, with large, broadly triangular, sharp lateral tooth, projecting laterally, dorsal surface gently convex (Figs. 1, 2, 3 A, 4A); antennae with subrectangular basal article, twice as long as wide (Fig. 3 B); antennules folding at ca. 45° (Fig. 3 B); epistome relatively broad with depressed median part (Fig. 3 A–C); third maxilliped with ischium subrectangular, with deep submedian longitudinal sulcus (Figs. 3 C, 4A), merus subquadrate, anteroexternal angle subauriculiform (Figs. 3 C, 4A); ambulatory legs long, slender (Figs. 1, 2, 4 E); ambulatory merus elongated, slender, surface with margins rounded, not cristate, not distinctly granular, dorsal distal tooth distinct but low (Figs. 1, 2, 4 E); ambulatory propodus, dactylus with ventral margins lined with dense stiff setae, with distinct dactylo-propodal lock, not forming subchelate structure (Figs. 1, 2, 4 D, E); cheliped merus rounded in cross-section; surface covered with numerous small pits (Figs. 1, 2, 3 C); cheliped carpus surface with long, gently curved spine on distal inner angle (Figs. 1, 2, 3 F); chela unarmed, margins not cristate (Fig. 3 F); thoracic sternum relatively narrow transversely, surfaces pitted, sternites 3, 4 completely fused with median depression (Figs. 3 C, 4B); male abdominal locking mechanism present as knob-like tubercle on distal one-third of thoracic sternite 5; male abdomen broadly triangular (Figs. 3 D, 4C); G1 slender, sinuous, distal part elongated (Fig. 4 F, G).</p>
            <p> Etymology. The new genus is named after Cebu City in the Visayas, Philippines; in arbitrary combination with the genus name  Eumedonus . The gender of the genus is masculine. </p>
            <p> Remarks. The unusual combination of carapace, cheliped, ambulatory leg, thoracic sternal and abdominal characters in  Cebudonus poppeorum n. gen. , n. sp. require the establishment of a new genus. While the carapace shape of  Cebudonus n. gen. superficially resembles species associated with echinoids like  Eumedonus intermedius Chia &amp; Ng, 2000 (Madagascar) ,  E. vicinus Rathbun, 1918 (Australia) , and  E. zebra Alcock 1895 (Red Sea to East China Sea), especially with regard to the shape of the laterally directed spines; its front is completely different, with two long pseudorostral spines (Figs. 1, 2) in contrast to  Eumedonus , which has a lobiform pseudorostrum that is bifurcated distally to form two teeth (cf. Chia &amp; Ng 2000: figs. 8, 9A, D, 10, 11, 12A, D, N, 13, 14, 15A, D, Q, R). The lateral carapace tooth and the dorsal surface adjacent to it is gently convex dorsally in  Cebudonus n. gen. (Fig. 3 A) (the tooth is stout with the dorsal surface distinctly convex in  Eumedonus ; cf. Chia et al. 1995: fig. 3A); the cheliped merus is relatively long and unarmed (Figs. 1, 2, 3 C) (relatively short with numerous tubercles and granules in  Eumedonus ; cf. Chia &amp; Ng 2000: figs. 8, 9B, 10, 11, 12B, L, 14, 15B, O); the meri of the ambulatory legs are long, subcylindrical and smooth, without any marginal cristae, and the dorsal distal tooth is relatively low (Figs. 1, 2, 4 E) (short, laterally flattened with distinct marginal cristae and a strong dorsal distal tooth in  Eumedonus ; cf. Chia &amp; Ng 2000: figs. 8, 9E, G, 10, 11, 12E, K, 13, 14, 15E, K); the anterior male thoracic sternum (sternites 1–4) are proportionately narrow transversely (Figs. 3 C, 4B) (proportionately wider transversely in  Eumedonus ; cf. Chia &amp; Ng 2000: figs. 9C, 11B, 12C, 14B, 15N); and the male abdomen is relatively wide (Figs. 3 D, 4C) (proportionately narrower in  Eumedonus ; cf. Chia &amp; Ng 2000: figs. 12J, 14J, 15N). </p>
            <p> The unarmed and non-cristate ambulatory meri of  Cebudonus n. gen. resembles that of the echinoid symbiont  Gonatonotus , but in the new taxon, it is proportionately much longer, more slender and almost smooth (Figs. 1, 2, 4 E) (proportionately shorter and distinctly granular in  Gonatonotus ; cf. Chia &amp; Ng 2000: figs. 17, 18E, G, 20, 21E, K, 22, 23E, K). The differences previously noted in the form of the front, anterolateral region, cheliped, thoracic sternum and abdomen between  Cebudonus n. gen. and  Eumedonus also apply for  Gonatonotus (cf. Chia &amp; Ng 2000: figs. 17, 18A, D, 19A, 20, 21A, D, M, N, 22, 23A, D [front]; figs. 17, 18B, 20, 21B, 22, 23B [chelipeds]; figs. 17B, 19G, 20B, 21C, 22B, 23C [sternum]; figs. 19H, 21J, 22, 23J [abdomen]). </p>
            <p> The form of the front, with the two long pseudorostral spines is similar to that of  Tiaramedon and  Zebrida , but these genera differ markedly from  Cebudonus n. gen. in several other characters.  Cebudonus n. gen. , can be separated from  Tiaramedon by its smooth dorsal surface of the carapace (Figs. 1, 2) (armed with additional gastric and branchial spines in  Tiaramedon ; cf Chia &amp; Ng 1998: figs. 7A, 8A); the two pseudorostral spines joining medially without any trace of a median plate (Figs. 1, 2) (with a small plate present between the pseudorostral spines in  Tiaramedon ; cf. Chia &amp; Ng 1998: fig. 8A, B); the chela is unarmed (Figs. 1, 2, 3 F) (with an additional spine on the dorsal distal margin in  Tiaramedon ; cf. Chia &amp; Ng 1998: figs. 7A, B, 8K); the cheliped carpus has a single long inner spine (Figs. 1, 2, 3 F) (2 large spines in  Tiaramedon ; cf. Chia &amp; Ng 1998: fig. 7A); the ambulatory merus is long and slender with a short dorsal distal tooth (Figs. 1, 2, 4 E) (short and stout with a long distal dorsal tooth in  Tiaramedon ; cf. Chia &amp; Ng 1998: figs. 7, 8C); and the male anterior thoracic sternum is transversely narrow (Figs. 3 C, 4B) (transversely broad in  Tiaramedon ; cf. Chia &amp; Ng 1998: figs. 7B, 8D). The male abdomens of  Cebudonus n. gen. and  Tiaramedon are nevertheless similar in shape (Figs. 3 D, 4C; cf. Chia &amp; Ng 1998: figs. 7B, 8F). The two genera are also different in terms of host specificity, with  Tiaramedon always associated with crinoids (and with young crabs characteristically transversely striped like many other crinoid-associated eumedonines, see Castro et al. 1995; Chia &amp; Ng 1995, 1998) whereas  Cebudonus n. gen. with its longitudinal stripes is probably associated with echinoids. </p>
            <p> Cebudonus n. gen. can be separated from the echinoid symbiont  Zebrida by having two subcylindrical pseudorostral spines (Figs. 1, 2) (distinctly lamelliform in  Zebrida ; cf. Ng &amp; Chia 2000: figs. 2, 3A, B); the lateral carapace tooth and the dorsal surface adjacent to the tooth is dorsally convex (Fig. 3 B) (the area is distinctly dorsally flattened in  Zebrida ; cf. Chia et al. 1995: fig. 3B); the chela and merus are unarmed and the carpus has a long spine at the inner angle (Figs. 1, 2, 3 F) (the chela has a large subdistal dorsal tooth, the carpus and merus each having three large lamelliform teeth in  Zebrida ; cf. Ng &amp; Chia 2000: figs. 2, 3J); the ambulatory merus is long and slender with a short dorsal distal tooth (Figs. 1, 2, 4 E) (short and stout with the margins cristate to dentate and with a large distal dorsal tooth in  Zebrida ; cf. Ng &amp; Chia 2000: figs. 2, 3C, K, M); the ambulatory propodus and dactylus are normal (Figs. 1, 2, 4 D, E) (forming a subchelate clasping structure in  Zebrida ; cf. Ng &amp; Chia 2000: figs. 2, 3C, K, M); and the male anterior thoracic sternum is proportionately narrower transversely (Figs. 3 C, 4B) (proportionately wider in  Zebrida ; cf. Ng &amp; Chia 2000: figs. 2B, 3D). The male abdomens of  Cebudonus n. gen. and  Zebrida are similar, with both relatively wide transversely (Figs. 3 D, 4C; cf. Ng &amp; Chia 2000: figs. 2B, 3F) </p>
            <p> There are also similarities with  Zebridonus but  Cebudonus n. gen. can be separated by its two long pseudorostral spines (Figs. 1, 2) (with a distinctly dorsoventrally flattened lobiform pseudorostrum that is bifurcated distally to form two teeth in  Eumedonus ; cf. Chia et al. 1995: figs. 1, 2A, K); the lateral carapace tooth and the dorsal surface adjacent to it is dorsally convex (Fig. 3 B) (the area is distinctly dorsally flattened in  Zebridonus ; cf. Chia et al. 1995: figs. 1, 2A, 3C); the chela and merus are unarmed, and the carpus has a long spine on the inner angle (Figs. 1, 2, 3 F) (the chela has a distinct distal dorsal tooth, the carpus 2 large lamelliform teeth and the merus several distinct teeth in  Zebridonus ; cf. Chia et al. 1995: figs. 1, 2 I); the ambulatory merus is long, subcylindrical and smooth, without any marginal cristae and the dorsal distal tooth is relatively low (Figs. 1, 2, 4 E) (short, laterally flattened with distinct marginal cristae and a strong dorsal distal tooth in  Zebridonus ; cf. Chia et al. 1995: figs. 1, 2B, C); the anterior male thoracic sternum (sternites 1–4) are proportionately narrower transversely (Figs. 3 C, 4B) (proportionately wider transversely in  Zebridonus ; cf. Chia et al. 1995: fig. 2D); and the male abdomen is proportionately wider (Figs. 3 D, 4C) (proportionately narrower in  Zebridonus ; cf. Chia et al. 1995: fig. 2F). </p>
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	https://treatment.plazi.org/id/523B87B7FF95270606D6224BFF6AD13E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ng, Peter K. L.	Ng, Peter K. L. (2014): Cebudonus poppeorum, a new genus and new species of eumedonine crab (Crustacea: Decapoda: Brachyura: Pilumnidae) from the Philippines. Zootaxa 3815 (1): 94-102, DOI: 10.11646/zootaxa.3815.1.6
523B87B7FF96270306D62558FC1BD05F.text	523B87B7FF96270306D62558FC1BD05F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cebudonus poppeorum	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Cebudonus poppeorum n. gen. , n. sp. </p>
            <p>(Figs. 1–4)</p>
            <p>Material examined. Holotype: male (10.8 × 11.9 mm) (NMCR), Punta Engaño, Mactan Island, Cebu, Visayas, 180–250 m, coll. local trawler, 2013. Paratypes: 1 dried female (10.6 × 12.5 mm) (ZRC 2014.0211), same data as holotype; 1 female (8.1 × 9.2 mm) (ZRC 2014.0212), Punta Engaño, Mactan Island, Cebu, Visayas, 180–210 m, coll. local trawler, 2012; 1 dried male (8.3 × 9.2 mm) (ZRC 2014.0213), Talingting, Siquijor, Visayas, 120 m, coll. local fishermen, tangle nets, 2013; 1 female (10.9 × 12.4 mm) (ZRC 2014.0214), Talingting, Siquijor, Visayas, 120 m, coll. local fishermen, tangle nets, 2013; 1 female (12.5 × 14.5 mm) (GTP), Talingting, Siquijor, Visayas, 150 m, coll. local fishermen, April 2013; 1 male (9.9 × 10.3 mm) (GTP), Basul Island, Surigao, northern Mindanao, 80- 100 m, coll. local trawlers, January 2013. All locations in central Philippines.</p>
            <p> Comparative material. See ZRC material examined in Chia &amp; Ng (1998, 2000), Chia et al. (1995, 1999) and Ng &amp; Chia (1999). In addition:  Zebrida adamsii White, 1847: 1 dried female (9.8 × 9.7 mm) (ZRC), Molocaboc area, north of Negros Island, Philippines 10–25 m, coll. local fisherman, 2014; 1 dried female (12.9 × 12.0 mm) (ZRC), Molocaboc area, north of Negros Island, Philippines, 10–25 m, coll. local fisherman, 2014. </p>
            <p>Diagnosis. As for genus.</p>
            <p>Description. Carapace pentagonal; dorsal surface gently convex, covered with numerous small pits, otherwise smooth; regions poorly demarcated, cardio-gastric groove shallow (Figs. 1, 2). Front with 2 long pseudorostral spines, subparallel, ca. 0.4 times total carapace length; surface finely pitted; tip straight, rounded; spines separated by broad, U-shaped cleft; spines confluent with supraorbital margin (Figs. 1, 2, 3 C). Orbit small, rounded; cornea protrudes beyond edge of carapace when retracted (Figs. 1, 2, 3 A–C). Anterolateral margin concave, entire, demarcated by small, sharp external orbital tooth, large, broadly triangular, sharp lateral tooth, projecting laterally, not upturned, dorsal surfaces adjacent to it gently convex, not flattened (Figs. 1, 2, 3 A, 4A). Posterolateral margin almost straight to gently convex; converging strongly to slightly sinuous posterior carapace margin (Figs. 1, 2, 3 E). Suborbital margin short, concave; inner tooth short, rounded (Fig. 3 B). Pteryogostomial, subhepatic regions pitted, not pubescent (Fig. 3 A–C).</p>
            <p>Antennae short; basal article free, subrectangular, twice as long as wide; flagellum enters orbit but not extending across length (Fig. 3 B). Antennules large, folding at ca. 45°; basal article subtrapezoidal, filling about two-thirds of fossa (Fig. 3 B). Epistome relatively broad longitudinally, covered with small pits except for depressed median part; posterior margin crenulate, with distinct median triangular lobe, separated by distinct fissure (Fig. 3 A–C).</p>
            <p>Outer surfaces of third maxilliped densely covered with small pits (Figs. 3 C, 4A). Ischium subrectangular, with deep submedian longitudinal sulcus; inner margin dentate (Figs. 3 C, 4A). Merus subquadrate; anteroexternal angle subauriculiform (Figs. 3 C, 4A). Carpus short; propodus rectangular; dactylus elongated (Fig. 4 A). Exopod relatively stout, with distinct flagellum (Fig. 4 A).</p>
            <p>Ambulatory legs long, slender; second leg longest, fourth leg shortest (Figs. 1, 2). Merus elongated, slender; surface with scattered pits; margins rounded, not cristate, almost entire, not distinctly granular except for distal margin; dorsal distal tooth distinct but may be low (Figs. 1, 2, 4 E). Carpus smooth, with scattered pits on outer surface (Figs. 1, 2, 4 E). Propodus elongated, dorsoventrally flattened; margins entire, ventral margin with numerous short stiff setae; outer surface with scattered pits (Figs. 1, 2, 4 D, E). Dactylus gently curved; ventral margin with dense mat of short, stiff setae except for corneous tip; forming distinct dactylo-propodal lock (Figs. 1, 2, 4 D, E).</p>
            <p>Chelipeds slender, elongated (Figs. 1, 2). Basis-ischium elongated, covered with small pits (Fig. 3 C). Merus rounded in cross-section; surface covered with numerous small pits, otherswise unarmed (Figs. 1, 2, 3 C). Carpus surface with numerous small pits, subovate; long, gently curved, dorsoventrally flattened spine on distal inner angle (Figs. 1, 2, 3 F). Chela elongated, margins rounded, not cristate; outer surface strongly pitted; fingers stout, ca. half length of palm, not pigmented, cutting edges with low teeth on distal half, proximal half relatively smooth (Fig. 3 F).</p>
            <p>Thoracic sternum relatively narrow transversely; surfaces distinctly pitted; sternites 1, 2 completely fused, forming triangular plate, separated from sternite 3 by distinct sinuous suture; sternites 3, 4 completely fused, demarcated only by lateral cleft, median depression, sutures otherwise not visible (Figs. 3 C, 4B). Male abdomen not spanning width of thoracic sternum; sternite 8 visible between abdominal somites 1, 2, coxae of last pair of ambulatory legs (Fig. 4 E).</p>
            <p>Sternoabdominal cavity deep, reaching to median part of fused sternites 3, 4 (Fig. 3 D). Male abdominal locking mechanism present as knob-like tubercle on distal one-third of sternite 5. Male abdomen broadly triangular, all somites, telson free (Figs. 3 D, 4C). Somite 1 subrectangular, distal margin deeply concave; somite 2 transversely subovate (Figs. 3 E, 4C). Somite 3 broadest, subtrapezoidal; somites 4–6 trapezoidal, lateral margins gently concave to sinuous (Figs. 3 D, 4C). Telson triangular, lateral margins gently convex to sinuous, tip rounded (Figs. 3 C, D, 4B, C). Penis opening on condyle of coxa of fourth ambulatory leg.</p>
            <p>G1 long, slender, sinuous; distal part elongated, straight; distal surfaces with rows of short, to long setae; basal part with plumose setae (Fig. 4 F, G). G2 short, sigmoid; distal segment short (Fig. 4 H).</p>
            <p>Females and variation. The female specimens agree with the male in almost all non-sexual aspects. The pseudorostral spines in some specimens (e.g., a female 10.6 × 12.5 mm, ZRC 2014.0211) converge along the distal half rather than being subparallel. The female abdomen is subovate, broad, and covers the entire surface of the thoracic sternum; with all the somites and telson free (Fig. 3 G). The vulvae are large, ovate, without any opercular cover, with the outer margin forming a low rim; and positioned on the anterior half of thoracic sternite 6 (Fig. 3 H).</p>
            <p>Colour. The striking colour of fresh specimens (Fig. 1) remains even in preserved specimens (Fig. 2). The carapace has two broad longitudinal stripes on a reddish-brown background, with the posterolateral margins also white. The broad reddish-brown median stripe extends to the first two abdominal somites. The frontal parts of the carapace (epistome, suborbital and subhepatic regions) are purplish-pink. The chelipeds are generally white to pinkish-white, with the proximal part of the merus, carpal spine, proximal part of the pollex pink. The ambulatory legs are white to reddish-white.</p>
            <p>Etymology. It is a pleasure to name this new species after the intrepid father and son team of Guido and Philippe Poppe, renowned malacologists and ardent naturalists. Between the two, they have had a significant influence advancing our knowledge of Philippine natural history over the last decade.</p>
            <p> Remarks. The specimens of  Cebudonus poppeorum n. gen. , n. sp. were collected from depths ranging from 80 to 250 m by trawlers and tangle nets. The general features and its close resemblance to  Eumedonus and  Gonatonotus strongly suggest that it is an obligate symbiont of urchins. Like species members of these genera, their dorsal carapace surface is also distinctively longitudinally striped in life (cf. Chia &amp; Ng 2000). It is noteworthy that eumedonines that have their carapaces marked with transverse stripes at some stage of their lives are all associated with crinoids (cf. Chia &amp; Ng 1998). The live colour pattern and host of  Zebridonus (from northern Australia), which is closely allied to  Eumedonus , are not known, but it is also believed to be associated with an echinoid because of its colour pattern. The unusual eumedonine  Tauropus (from the South China Sea and Japan), known only from its type species  T. egeriae (Gordon, 1947; Chia &amp; Ng 1998) is also probably associated with echinoids but known specimens had no distinctive colour pattern (unpublished data). Guido Poppe commented (in litt. 23 April 2014) that large cidaroid sea urchins were collected together with the crab in Mactan Island. Most urchins belonged to  Coelopleurus Agassiz, 1840 (family  Arbaciidae ), with  C. maillardi (Michelin, 1862) the most common one. It is therefore possible that  C. poppeorum n. sp. is associated with this echinoid species. </p>
            <p>Distribution. Known so far from central and southern Philippines.</p>
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	https://treatment.plazi.org/id/523B87B7FF96270306D62558FC1BD05F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ng, Peter K. L.	Ng, Peter K. L. (2014): Cebudonus poppeorum, a new genus and new species of eumedonine crab (Crustacea: Decapoda: Brachyura: Pilumnidae) from the Philippines. Zootaxa 3815 (1): 94-102, DOI: 10.11646/zootaxa.3815.1.6
