taxonID	type	description	language	source
533087D1CB5AFFA6E47BFDB7E8DBFA0B.taxon	type_taxon	Type species Inodesmus jamaicensis Cook, 1896, by monotypy.	en	Romero-Rincon, Juan, Alvear, Santiago, Martínez-Torres, Daniela, Robles-Piñeros, Jairo (2025): First record of the millipede family Haplodesmidae (Diplopoda: Polydesmida) from Colombia: two new species of Inodesmus Cook, 1896 from the Northern Andes. European Journal of Taxonomy 976 (1): 33-46, DOI: 10.5852/ejt.2025.976.2773, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2773/12679
533087D1CB5AFFA3E473F9A2EE24FCA9.taxon	description	urn: lsid: zoobank. org: act: 945 A 115 E- 0 BE 9 - 484 A- 9581 - FF 7 E 09 C 59803 Figs 1 – 4, 8	en	Romero-Rincon, Juan, Alvear, Santiago, Martínez-Torres, Daniela, Robles-Piñeros, Jairo (2025): First record of the millipede family Haplodesmidae (Diplopoda: Polydesmida) from Colombia: two new species of Inodesmus Cook, 1896 from the Northern Andes. European Journal of Taxonomy 976 (1): 33-46, DOI: 10.5852/ejt.2025.976.2773, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2773/12679
533087D1CB5AFFA3E473F9A2EE24FCA9.taxon	diagnosis	Diagnosis Males with head + 19 rings, females with head + 20 rings. This species differs from its congeners in having a coxal projection on male leg 6. Telopodite (Fig. 2) with pp straight; at in transverse plane, large, somewhat fusiform, and slightly bent posteriorly; dp directed laterobasally, basal portion of the dp linguiform; meb curving strongly behind mab, bifurcated basally, with a long, acuminate axial process, and a hook-shaped process, with another, small process at midlength; mab concave medially, broad distally, curved outward, and with a small subtriangular process distally.	en	Romero-Rincon, Juan, Alvear, Santiago, Martínez-Torres, Daniela, Robles-Piñeros, Jairo (2025): First record of the millipede family Haplodesmidae (Diplopoda: Polydesmida) from Colombia: two new species of Inodesmus Cook, 1896 from the Northern Andes. European Journal of Taxonomy 976 (1): 33-46, DOI: 10.5852/ejt.2025.976.2773, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2773/12679
533087D1CB5AFFA3E473F9A2EE24FCA9.taxon	etymology	Etymology The specific epithet, mesobovi (noun, genetive case), is a patronym, honoring Dr. Robert Mesibov for his valuable contributions to the knowledge of the Australian millipede fauna.	en	Romero-Rincon, Juan, Alvear, Santiago, Martínez-Torres, Daniela, Robles-Piñeros, Jairo (2025): First record of the millipede family Haplodesmidae (Diplopoda: Polydesmida) from Colombia: two new species of Inodesmus Cook, 1896 from the Northern Andes. European Journal of Taxonomy 976 (1): 33-46, DOI: 10.5852/ejt.2025.976.2773, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2773/12679
533087D1CB5AFFA3E473F9A2EE24FCA9.taxon	materials_examined	Material examined Holotype COLOMBIA • ♂; Cundinamarca, San Antonio del Tequendama, Parque Natural Chicaque, oak forest; [04 º 37.028 ′ N, 74 ° 18.830 ′ W]; 2232 – 2275 m a. s. l.; 12 Jan. 2023; J. Romero-Rincon and J. Veloza leg.; under a decaying fallen log; daytime hand collection; ICN-MD- 2913. Paratypes (11 ♂♂, 15 ♀♀, 17 immatures) COLOMBIA • 1 ♂ (fragmented), 1 ♀; same data as for holotype; MHN-UPN-MD- 168 • 1 ♂ (fragmented); same data as for holotype; MHN-UPN-MD- 176 • 1 ♂, 1 ♀; same data as for holotype; MHNUC-MD- 276 • 1 ♀; same data as for holotype; with teratologies; MHN-UPN-MD- 177 • 1 ♂, 3 ♀♀, 7 immatures; same data as for holotype; MHN-UPN-MD- 178 • 2 ♂♂, 2 ♀♀; same data as for holotype; ICN-MD- 3130 • 1 ♂; same locality as for holotype; 26 – 29 Sep. 2022; J. Romero-Rincon and L. Poveda leg.; under decaying log; daytime hand collection; MHN-UPN-MD- 92 • 1 ♀, 1 immature (fragmented); same data as for preceding; MHN-UPN- 94 • 1 ♀; same locality as for preceding; 29 May 2022; J. Romero-Rincon and R. De La Cruz leg.; inside a decaying fallen log; daytime hand collection; MHN-UPN-MD- 96 • 1 ♀; same locality as for preceding; 29 Apr. 2022; J. Romero-Rincon leg.; under a decaying fallen log; night time hand collection; MHN-UPN-MD- 99 • 1 ♂, 1 ♀, 4 immatures; same locality as for preceding; 25 – 26 Jun. 2022; J. Romero-Rincon and R. De La Cruz leg.; inside a decaying fallen log; night time hand collection; MHN-UPN-MD- 98 • 1 ♂, 1 immature; same locality as for preceding; 25 – 26 Jun. 2022; J. Romero-Rincon and R. De La Cruz leg.; under a decaying fallen log; daytime hand collection; MHNUC-MD- 412 • 2 ♀♀, 1 immature; same locality as for preceding; 19 Jun. 2023; J. Romero-Rincon leg.; under a decaying fallen log; night time hand collection; MHN-UPN-MD- 163 • 1 ♂, 3 immatures; same locality as for preceding; 16 Aug. 2023; J. Romero-Rincon and H. Reip leg.; under a decaying fallen log; daytime hand collection; MHN-UPNMD- 169 • 1 ♀; Cundinamarca, San Antonio del Tequendama, Parque Natural Chicaque, secondary forest; [04 º 36.854 ′ N, 74 ° 18.859 ′ W]; 2155 m a. s. l.; 28 – 29 Apr. 2022; J. Romero-Rincon leg.; under bark of fallen log; daytime hand collection; MHN-UPN-MD- 97 • 1 ♂; same locality as for preceding; 30 – 31 Jul. 2022; J. Romero-Rincon leg.; under a decaying fallen log; night time hand collection; MHN-UPNMD- 95.	en	Romero-Rincon, Juan, Alvear, Santiago, Martínez-Torres, Daniela, Robles-Piñeros, Jairo (2025): First record of the millipede family Haplodesmidae (Diplopoda: Polydesmida) from Colombia: two new species of Inodesmus Cook, 1896 from the Northern Andes. European Journal of Taxonomy 976 (1): 33-46, DOI: 10.5852/ejt.2025.976.2773, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2773/12679
533087D1CB5AFFA3E473F9A2EE24FCA9.taxon	description	Description MEASUREMENTS. Male with head + 19 rings, female with head + 20 rings (Fig. 1). Average measurement of adult type specimens: male / female ca 10 / 11.5 mm long; maximum width ca 0.9 / 1.2 mm. COLORATION. Metazonites, prozonites, and collum deep reddish brown 41 to deep brown 56; head, legs (prefemur and coxa pale greenish yellow 104), and antennae light reddish brown 42. HEAD. Wider than high; narrower than collum; facing downwards (Fig. 3 A – C); setose clypeus (Fig. 3 A). Vertex microvillose and microgranulate. Antennae (Fig. 3 A – C) short, stout, clavate, densely setose, and held close to head. Antennomere relative lengths as follows: 6> 1> 5> 4 = 2> 3> 7. Antennae with 4 sensory apical cones. TRUNK. Anterior and posterior margins of collum broadly convex; corners rounded and hidden under paranota of second ring, collum with an irregular pattern of two groups of rounded tubercles. At first glance, pattern continues on remaining body rings, with a supposed transverse medial separation on each tergite (Fig. 3 B – C). Ring 2 tergite largest, extending basally, laterally, and posteriorly; extension of paranota of ring 2 incrassate without well-defined tubercles (Fig. 3 B – C). Posterior rings (Fig. 3 G), up to ring 18, with a lateral row of three small tubercles, neither enlarged nor forming pseudo-paranota. Ozopores (Fig. 3 F) very small, internal closing not evident, located just above or in middle of last lateral row of tubercles (Fig. 3 G). Ozopores internally and externally bordered by microtubercules, with neither evident elevation nor porostele. Pore formula normal (5, 7, 9, 10, 12, 13, 15 – 19). Diplosternites (Fig. 3 D) with transverse impression deeper than longitudinal impression, anterior sternites with eight acicular setae, posterior sternites on ring 5 with bumps bearing six acicular setae each. Legs (Fig. 3 B – C, G – H) short and stout; relative lengths of podomeres: tarsus> (prefemur ≥ femur)> tibia> postfemur; claw about as long as postfemur. Spiracles not evident. Telson facing downwards. Paraprocts parallel to substrate and almost flat (Fig. 3 G – H). Epiproct (Fig. 3 H – I) distally projected, but short, flattened dorsoventrally, with four inconspicuous setae (spinnerets), each spinneret with a single low sheath, each seta inside a circular, deep, walled depression (Fig. 3 I). Hypoproct (Fig. 3 H) subtrapeziform with a slightly convex anterior margin. Tegument with microsculpture, especially along posterior edge of metazonite, anterior edge of prozonite, head, collum, lateral part of metazonite of second ring, edges of ozopores, and posterior area of telson (Fig. 3). Microsculpture mostly as a cellular mesh with narrow irregular folds. Integument further elevated into tubercles of different sizes and shapes on head, collum, tergites, metatergites, and telson (Fig. 3 A – C, G – H); some tubercles with a single small and acicular seta. Cell boundaries in posterior part of metazonite not extending basally past limbus (Fig. 3 E). Primary limbus element with a regular set of rounded lobes and tooth-like lobes on the secondary element (Fig. 3 E). Prozonites sharply demarcated from metazonites (Fig. 3 B – C, G). Anterior part of prozonite (a) with small irregular subcardiform units arranged in transverse “ rows ” placed on a smooth background. A transverse ridge (r) is marked by a row of cell-like units. Posterior part of the prozonite (b) with a smooth surface and lacking spherical knobs. GONOPOD. Oval aperture, rim widely raised laterally. Telopodite (Fig. 2) long, reaching base of leg pair 5 (Fig. 3 C). Basal portion of telopodite with a blunt, basally directed projection arising posteromedially to junction with cx, several short setae in basal portion of telopodite, and three large setae in a row on lateral edge of apical tab (Fig. 2).	en	Romero-Rincon, Juan, Alvear, Santiago, Martínez-Torres, Daniela, Robles-Piñeros, Jairo (2025): First record of the millipede family Haplodesmidae (Diplopoda: Polydesmida) from Colombia: two new species of Inodesmus Cook, 1896 from the Northern Andes. European Journal of Taxonomy 976 (1): 33-46, DOI: 10.5852/ejt.2025.976.2773, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2773/12679
533087D1CB5AFFA3E473F9A2EE24FCA9.taxon	discussion	Remarks A notable case of “ helicomerism ” was observed in a female of I. mesibovi sp. nov. in which the segmental anomaly breaks the serial arrangement of the body rings (Demange & Pereira 1980; Minelli & Pasqual 1986; Leśniewska et al. 2009). In this specimen (MHN-UPN-MD- 177), the tergite of ring 8 is entirely divided into two sections, a smaller isolated portion on the left and one on the right side of the metazonite that is completely fused with ring 7, giving a diagonal shape to the posterior part of the segment (Fig. 4 B). Additionally, another teratological feature is observed in the collum, in which an abnormal longitudinal transverse partition is evident (Fig. 4 A). However, it cannot be dismissed that the damage may have been influenced by external factors beyond the scope of development.	en	Romero-Rincon, Juan, Alvear, Santiago, Martínez-Torres, Daniela, Robles-Piñeros, Jairo (2025): First record of the millipede family Haplodesmidae (Diplopoda: Polydesmida) from Colombia: two new species of Inodesmus Cook, 1896 from the Northern Andes. European Journal of Taxonomy 976 (1): 33-46, DOI: 10.5852/ejt.2025.976.2773, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2773/12679
533087D1CB5AFFA3E473F9A2EE24FCA9.taxon	distribution	Distribution Only known from Parque Natural Chicaque, Cundinamarca, Colombia (Fig. 8).	en	Romero-Rincon, Juan, Alvear, Santiago, Martínez-Torres, Daniela, Robles-Piñeros, Jairo (2025): First record of the millipede family Haplodesmidae (Diplopoda: Polydesmida) from Colombia: two new species of Inodesmus Cook, 1896 from the Northern Andes. European Journal of Taxonomy 976 (1): 33-46, DOI: 10.5852/ejt.2025.976.2773, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2773/12679
533087D1CB5FFFAEE471FC05EECDFB07.taxon	description	Figs 5 – 8	en	Romero-Rincon, Juan, Alvear, Santiago, Martínez-Torres, Daniela, Robles-Piñeros, Jairo (2025): First record of the millipede family Haplodesmidae (Diplopoda: Polydesmida) from Colombia: two new species of Inodesmus Cook, 1896 from the Northern Andes. European Journal of Taxonomy 976 (1): 33-46, DOI: 10.5852/ejt.2025.976.2773, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2773/12679
533087D1CB5FFFAEE471FC05EECDFB07.taxon	diagnosis	Diagnosis Males with head + 19 rings, females with head + 20 rings. Differs from other species of Inodesmus in the following telopodite features: pp straight; at large, pointed, and bent posteriorly; dp triangular and concave, directed laterobasally; mab partially curved behind meb (lateral process only), divided into two large processes; the medial process is directed laterally and shows strong torsion, bifurcated distally with two subtriangular processes; lateral process flattened and divided into two processes; the lateral process is formed by three small subtriangular processes, one trifurcated sub-process, and a simple medial process directed mediobasally; meb wide and flattened at its base, directed laterobasally with acuminate apex (Fig. 6).	en	Romero-Rincon, Juan, Alvear, Santiago, Martínez-Torres, Daniela, Robles-Piñeros, Jairo (2025): First record of the millipede family Haplodesmidae (Diplopoda: Polydesmida) from Colombia: two new species of Inodesmus Cook, 1896 from the Northern Andes. European Journal of Taxonomy 976 (1): 33-46, DOI: 10.5852/ejt.2025.976.2773, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2773/12679
533087D1CB5FFFAEE471FC05EECDFB07.taxon	etymology	Etymology The specific epithet, miconiae (noun, genetive case), is derived from the abundance of flowering plants of the genus Miconia in the only known habitat of the species.	en	Romero-Rincon, Juan, Alvear, Santiago, Martínez-Torres, Daniela, Robles-Piñeros, Jairo (2025): First record of the millipede family Haplodesmidae (Diplopoda: Polydesmida) from Colombia: two new species of Inodesmus Cook, 1896 from the Northern Andes. European Journal of Taxonomy 976 (1): 33-46, DOI: 10.5852/ejt.2025.976.2773, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2773/12679
533087D1CB5FFFAEE471FC05EECDFB07.taxon	materials_examined	Material examined Holotype COLOMBIA • ♂; Cauca, Popayán, Hacienda Río Blanco, oak forest; [02 ° 29.235 ′ N, 76 ° 32.101 ′ W]; 1993 m a. s. l.; 9 Sep. 2023; A. S. Alvear leg.; under bark of decaying branch; daytime hand collection; MHNUC-MD- 170. Paratypes (total: 3 ♂♂, 4 ♀♀, 1 immature) COLOMBIA • 1 ♂; same data as for holotype; MHNUC-MD- 176 • 1 ♂; same data as for holotype; MHN-UPN-MD- 198 • 1 ♀, 1 immature; same data as for holotype; MHNUC-MD- 177 • 1 ♂; same locality as for holotype; 10 Sep. 2023; A. S. Alvear and D. Dueñas leg.; ICN-MD- 2914 • 3 ♀♀; same locality as for holotype; 2 Sep. 2023; A. S. Alvear and D. Pancho leg.; MHNUC-MD- 175.	en	Romero-Rincon, Juan, Alvear, Santiago, Martínez-Torres, Daniela, Robles-Piñeros, Jairo (2025): First record of the millipede family Haplodesmidae (Diplopoda: Polydesmida) from Colombia: two new species of Inodesmus Cook, 1896 from the Northern Andes. European Journal of Taxonomy 976 (1): 33-46, DOI: 10.5852/ejt.2025.976.2773, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2773/12679
533087D1CB5FFFAEE471FC05EECDFB07.taxon	description	Description MEASUREMENTS. Male with head + 19 rings, female with head + 20 rings (Fig. 5). Average measurement of adult type specimens: male / female ca 7 / 9 mm long; maximum width ca 0.5 / 0.8 mm. COLORATION. Collum and pleurotergites deep reddish brown 41 to dark reddish brown 44; prefemora moderate yellowish brown 77; remaining podomeres, head, sternites, and antennae yellowish white 92 to moderate yellowish pink 29. HEAD. Head wider than high; narrower than collum; facing downwards (Fig. 7 A – C); clypeus microvillous, with needle-like setae; frons, epicranium, and genae microgranulate (Fig. 7 A). Antennae (Fig. 7 A – C) short, stout, clavate, densely setose, and held close to head. Antennomere relative lengths as follows: 6> 5> 1> 4> 2> 3> 7. Antennae with 4 sensory apical cones. TRUNK. Anterior and posterior margins of collum broadly convex; corners rounded and hidden by paranota of second segment (Fig. 7 B – C). Ring 2 tergite largest, extending laterally and anteriorly; paranota of ring 2 with three faintly defined tubercles (Fig. 7 B – C). Posterior rings (Fig. 7 C, G), up to ring 18, with a lateral row of 3 small tubercles, neither enlarged nor forming pseudo-paranota. Ozopore (Fig. 7 F) very small, of irregular shape, and with double aperture. Internal closing of ozopore traceable, with neither evident elevation nor porostele. Pores internally and externally bordered by microtubercules, located just above or in middle of last lateral row of tubercles (Fig. 7 G). Pore formula normal (5, 7, 9 – 10, 12 – 13, 15 – 19). Diplosternites (Fig. 7 D) with a longitudinal impression slightly deeper than transverse impression. Ring 5 with eight acicular setae on anterior sternites and six on posterior sternites. Legs (Fig. 7 A, C, G) short and stout; relative lengths of podomeres: tarsus> (prefemur ≥ femur)> (tibia ≥ postfemur); claw about half length of postfemur. Spiracles not evident. Telson facing downwards. Paraprocts parallel to substrate and almost flat (Fig. 7 G – H). Epiproct (Fig. 7 H – I) distally projected, but short, flattened dorsoventrally, with four inconspicuous setae (spinnerets), each spinneret with a single low sheath, each seta inside a circular, deep, walled depression (Fig. 7 I). Hypoproct (Fig. 7 H) subtrapeziform with a slightly convex anterior margin. Tegument with microsculpture, especially along posterior edge of metazonites, anterior edge of head, collum, prozonites, lateral part of metazonite of second ring, and telson (Fig. 7), mostly as a cellular mesh with narrow irregular folds. Integument further raised into subcircular tubercles of different sizes arranged in three transverse rows on collum and metatergites in rings 3 – 18 (Fig. 7 B – C, G); all tubercles with a single short, acicular seta. Cell boundaries in posterior part of metazonite not extending basally past limbus (Fig. 7 E). Primary limbus element with a regular set of rounded lobes and on secondary limbus element lobes more tooth-like (Fig. 7 E). Prozonites sharply demarcated from metazonites (Fig. 7 B – C, G). Anterior part of prozonite (a), transverse ridge (r) and posterior part of prozonite (b) not traceable (Fig. 7 E). GONOPOD. Oval aperture, rim slightly raised laterally and posteriorly. Cx (Fig. 6) with microgranular integumental sculpture and two large setae. Telopodite (Fig. 6) short, not reaching metazonite of segment 6 (Fig. 5 A). Basal portion of telopodite with blunt, posteriorly directed projection arising posteromedially to junction with cx; with three large setae basally, increasing in size, positioned in a row on lateral edge of apical tab (Fig. 6).	en	Romero-Rincon, Juan, Alvear, Santiago, Martínez-Torres, Daniela, Robles-Piñeros, Jairo (2025): First record of the millipede family Haplodesmidae (Diplopoda: Polydesmida) from Colombia: two new species of Inodesmus Cook, 1896 from the Northern Andes. European Journal of Taxonomy 976 (1): 33-46, DOI: 10.5852/ejt.2025.976.2773, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2773/12679
533087D1CB5FFFAEE471FC05EECDFB07.taxon	discussion	Remarks Although the type locality of the species has been widely sampled, all specimens originate from a single location within a depression in the oak forest. The specimens were collected in an area measuring about 1.5 × 3.0 m that featured a notable abundance of plants of the genus Miconia (Melastomataceae). All specimens were discovered beneath the bark of small decomposing branches.	en	Romero-Rincon, Juan, Alvear, Santiago, Martínez-Torres, Daniela, Robles-Piñeros, Jairo (2025): First record of the millipede family Haplodesmidae (Diplopoda: Polydesmida) from Colombia: two new species of Inodesmus Cook, 1896 from the Northern Andes. European Journal of Taxonomy 976 (1): 33-46, DOI: 10.5852/ejt.2025.976.2773, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2773/12679
533087D1CB5FFFAEE471FC05EECDFB07.taxon	distribution	Distribution Only known from the Hacienda Río Blanco in Popayán, Cauca, Colombia (Fig. 8).	en	Romero-Rincon, Juan, Alvear, Santiago, Martínez-Torres, Daniela, Robles-Piñeros, Jairo (2025): First record of the millipede family Haplodesmidae (Diplopoda: Polydesmida) from Colombia: two new species of Inodesmus Cook, 1896 from the Northern Andes. European Journal of Taxonomy 976 (1): 33-46, DOI: 10.5852/ejt.2025.976.2773, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2773/12679
