taxonID	type	description	language	source
4C578782E155FF98FF77FA2A22BAFC58.taxon	description	Pulaeus — Den Heyer, 1980 a: 19; El-Bishlawy & Rakha, 1983: 373 – 375; Bu & Li, 1991: 70 – 72; Barilo, 1991: 132 – 134; Corpus-Raros, 1996: 119 – 138; Lin et al, 2001: 145 – 153 Neocunaxoides — Chaudhri, 1977: 50; Chaudhri et al, 1979: 176 – 177; Tseng, 1980: 267 – 272 Type species: Eupalus pectinatus Ewing, 1909. Historical-taxonomic review. Species belonging to this genus are (re-) described and / or figured by the following: Ewing (1909) describes a new species from Illinois, USA, viz. Eupalus pectinatus. Berlese (1916) describes E. sternalis as a new species from Palermo (Panormitano), Italy but also states that this form differs from E. pectinatus, which he received from Ewing, only in palp length. The co-author in 1975 studied specimens of these species in the Berlese Collection in Florence and concluded that they are conspecific. Berlese describes in the same paper a new species, viz. E. subterraneus, from a nest of field mice at Terrara, Italy. Thor & Willmann (1941) provide redescriptions of all above species. In their work on the Cunaxidae Baker & Hoffmann (1948) provide redescriptions of all above-mentioned species. Muma (1960) gives a short description and drawings of a new species from Florida, USA, viz. C. pectinellus. We regard this species to be conspecific with P. pectinatus mentioned in this paper. Chaudhri et al (1979) redescribed Neocunaxoides krama Chaudhri, 1977; this species very strongly resembles P. longignathus Bu & Li, 1987 and is regarded by us as a new combination. We are also of opinion that the following species remain in this genus: Cunaxoides americanus Baker & Hoffmann, 1948; P. ardeola, Barilo, 1991; P. cebuensis Corpuz-Raros, 2007; P. chongqingensis Bu & Li, 1987; P. longignathus Bu & Li, 1987; P. m u s c i Liang, 1985; P. rimandoi Corpuz- Raros, 1996; P. s a m a re n s i s Corpuz-Raros, 2007 and P. w h a r t o n i (Baker & Hoffmann, 1948). Redescription. Mites of this genus are rather well sclerotized and characterised by the presence of fairly well-demarcated idiosomal shields, of a depression for peg organ on tarsus I proximally positioned; genital setae (g 1 — g 4) in approximately straigth row; palp tibiotarsus with a single pointed process. In his redescription of one of the species, viz. Pulaeus glebulentus Den Heyer, Smiley (1992) mentioned the following numbers of setae on basifemora I — IV: 4 sts — 5 sts — 4 sts — 2 sts; he informed that the redescription presented by him corresponds to the original description of the species. Although, the numbers of the setae he mentioned for the basifemora I — IV do not correspond to those presented in the original description of that species (4 sts — 6 sts — 3 sts — 2 sts) being characteristic of the genus, the taxonomical position of P. glebulentus remains unchanged.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E155FF98FF77FA2A22BAFC58.taxon	etymology	Etymology: The co-author, when he erected the genus Pulaeus Den Heyer, 1980 a, did not explain the name of this genus. The name is an anagram of the name Eupalus, as given by Koch (1838), a name restricted to the subfamily Cunaxoidinae, as is the genus name Lupaeus, as applied in the paper.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E156FF99FF77FA0824EEFF10.taxon	description	Female (n = 1). Idiosoma: length 445 and width 305. Dimensions of sensillae and setae: vi 92, ve 60, sci 40, sce 120, c 1 33, c 2 27, d 1 31, e 1 46, setae f 1 38, f 2 15, h 1 25 and h 2 15. Hypostome 130 long and hg - setae: hg 1 15, hg 2 20, hg 3 30 and hg 4 9. Chelicera 125 long and cheliceral seta 7. Length of legs: I 260, II, 225, III 225, IV 260; length of tarsi: I 102, II 77, III 70, IV 72.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E156FF99FF77FA0824EEFF10.taxon	discussion	Remarks. The Brazilian specimens differ from those redescribed by Den Heyer (1980 a) of P. pectinatus by the presence of 3 solenidia on genu I instead of 2 solenidia and by the reduction number of simple tactile setae on tarsi I and II that is 19 — 19 instead of 21 — 22. As there is just one specimen this will considered as belonging to P. pectiantus. Specimens examined. One female, from leaf litter of Astrocaryum aculeatissimum (Schott) Burret (Arecaceae), 11. X. 2000, Cananéia, A. C. Pereira.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E157FF9EFF77F8A42345FB9A.taxon	diagnosis	Diagnosis. Although this species resembles P. franciscae, it differs by having a dorsal shield strongly, instead of, slightly punctated.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E157FF9EFF77F8A42345FB9A.taxon	description	Female (Figs. 1 – 2) (n = 3). Idiosoma: length 328 (310 – 350) and width 207 (183 – 225). Dorsum (Fig. 1 A). Dorsal shield strongly punctated, sensillae (vi and sce) setose; dimensions: vi 106 (103 – 110), ve 25 (24 – 26), sci 19 (16 – 21), sce 96 (90 – 100), c 1 14 (14 – 15), c 2 15 (15 – 16), d 1 14 (14 – 15) and e 1 14 (13 – 15). Posteriad to the dorsal idiosomal shield two pairs of tiny platelets occur. Each of those immediate posteriad to the idiosomal shield carry setae f 1 18 (17 – 19) and f 2 12 (11 – 12) each whereas each of the most posterior platelets carry setae h 1 21 (18 – 23) and h 2 17 (17 – 18). Ve n t e r (Fig. 1 B). Propodosomal coxal plates not fused. One pair of paracoxal setae occurs on the coxal region IV. The integument bears 1 pair of propodogastral setae and 4 pairs of hysterogastral setae. One pair of paragenital setae occur laterad to the genital valves; 4 pairs of genital setae in approximately almost straight longitudinal row. Gnathosoma (Fig. 1 C – D). Hypostome 151 (145 – 157) long with 4 pairs of hg - setae [hg 1 21 (20 – 22), hg 2 20, hg 3 41 and hg 4 9] and and 2 pairs of adoral setae. Three- jointed palp, 92 (87 – 95) long. Trochanter without setae; femorogenu, 6 sts; tibiotarsus, 4 sts, 1 asl terminal, 1 bladder like apophysis and 1 ventral pointed process. Chelicera 150 long with cheliceral seta 7. Legs (Fig. 2). Length of legs: I 197, II 167, III 185, IV 200; length of tarsi: I 70, II 52, III 45, IV 47. The leg chaetotaxy is as follows: coxae I — IV, 1 peg, 3 sts — 3 sts — 3 sts — 2 sts; trochanters I — IV, 1 sts — 1 sts — 2 sts — 1 sts; basifemora I — IV, 4 sts — 6 sts — 3 sts — 2 sts; telofemora I — IV, 5 sts — 5 sts — 4 sts — 3 sts; genua I — IV, 3 asl, 5 sts — 2 asl, 5 sts — 1 asl, 5 sts — 1 asl, 5 sts; tibiae I — IV, 2 asl, 5 sts — 1 bsl, 5 sts — 1 bsl, 5 sts — 1 T, 4 sts; tarsi I — IV, 1 asl, 3 bsl, 1 dep, 1 dtsl, 2 tsl, 21 sts — 1 bsl, 1 dtsl, 2 tsl, 18 sts — 1 tsl, 15 sts — 15 sts. Male. Unknown.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E157FF9EFF77F8A42345FB9A.taxon	materials_examined	Type material. Holotype female, from soil under Myrcia sp. (Myrtaceae), 27. VII. 2000, Pirassununga, A. R. Oliveira. Paratypes. Pirassununga: 4 females from soil under Myrcia sp., 03. V. 2000, A. R. Oliveira; 1 female from soil under Myrcia sp., 27. VII. 2000, A. R. Oliveira.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E157FF9EFF77F8A42345FB9A.taxon	etymology	Etymology. The species name refers to the plant family Myrtaceae, in which this species is associated.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E151FF9FFF77FF082284F826.taxon	diagnosis	Diagnosis. This species resembles P. franciscae, but differs from the latter by the presence of 4 solenidia on genua I instead of 3.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E151FF9FFF77FF082284F826.taxon	description	Female (Figs. 3 – 4) (n = 13). Idiosoma: length 252 (237 – 287) and width 176 (150 – 212). Dorsum (Fig. 3 A). Dorsal shield slightly punctated with sensillae (vi and sce) setose; dimensions: vi 90 (80 – 107), ve 23 (20 – 27), sci 16 (12 – 22), sce 78 (67 – 87), c 1 12 (10 – 15), c 2 12 (10 – 12), d 1 13 (10 – 17) and e 1 15 (12 – 17). Posteriad to the dorsal idiosomal shield one pair of tiny platelets occur that carring setae f 1 19 (15 – 22) and f 2 11 (10 – 12) each. Setae h 1 22 (17 – 25) and h 2 13 occur on the integument. Ven t er (Fig. 3 B). The venter resembles that of P. myrtaceus sp. nov., but with 7 or 8 hysterogastral setae. Gnathosoma (Fig. 3 C – D). Hypostome 117 (100 – 127) long, bearing 4 pairs of hg setae [hg 1 11 (10 – 15), hg 2 13 (10 – 17), hg 3 35 (30 – 37) and hg 4 10 (7 – 12)] and 2 pairs of adoral setae. Three- jointed palp, 74 (62 – 87) long. Trochanter without setae; femorogenu, 6 sts; tibiotarsus, 4 sts, 1 asl terminal, 1 bladder-like apophysis and 1 ventral pointed process. Chelicera 116 (105 – 120) long with cheliceral seta 7. Legs (Fig. 4 A – D). Length of legs: I 165 (145 – 197), II 140 (120 – 170), III 158 (142 – 195), IV 171 (155 – 202); length of tarsi: I 62 (55 – 72), II 47 (40 – 60), III 44 (37 – 50), IV 45 (37 – 52). The leg chaetotaxy is as follows: coxae I — IV, 1 peg, 3 sts — 3 sts — 3 sts — 2 sts; trochanters I — IV, 1 sts — 1 sts — 2 sts — 1 sts; basifemora I — IV, 4 sts — 6 sts — 3 sts — 2 sts; telofemora I — IV, 5 sts — 5 sts — 1 ms, 3 sts — 1 ms, 2 sts; genua I — IV, 3 asl, {1 asl, 1 sts}, 4 sts — 2 asl, 5 sts — 1 asl, 5 sts — 1 asl, 5 sts; tibiae I — IV, 1 asl, 1 bsl, 5 sts — 1 bsl, 5 sts — 1 bsl, 5 sts — 1 T, 4 sts; tarsi I — IV, 1 asl, 3 bsl, 1 dep + peo, 1 dtsl, 2 tsl, 22 sts — 1 bsl, 1 dtsl, 1 tsl, 19 sts — 1 tsl, 16 sts — 17 sts. Male (Fig. 4). (n = 3). Dimensions. Idiosoma: length 191 (175 – 212) and width 143 (135 – 165). Dorsum (Fig. 4 E). The dorsum is similar to that of the female; dimensions: vi 85 (77 – 90), ve 25 (22 – 27), sci 16 (15 – 17), sce 75 (72 – 80), c 1 12, c 2 12 (10 – 12), d 1 12 and e 1 12 (12 – 15). Posteriad to the dorsal idiosomal shield one pair of tiny platelets occur carring setae f 1 15 and f 2 8 (7 – 10) each. Setae h 1 17 and h 2 13 occur on the integument. Ven ter (Fig. 4 F). Propodosomal coxal plates fused medially. One pair of paracoxal setae occurs on the coxal region IV. The integument bears 1 pair of propodogastral setae and 3 pairs of hysterogastral setae. Paragenital setae are not present. The genital valves are smaller than those of the females, each bearing 4 gsetae. Gnathosoma. The gnathosoma is similar to that of the female. Hypostome 105 (100 – 112) long with 4 pairs of hg - setae [hg 1 13 (12 – 15), hg 2 15 (15 – 17), hg 3 38 (35 – 42) and hg 4 10] and 2 pairs of adoral setae. Threejointed palp, 74 (70 – 77) long. Chelicera 106 (100 – 110) long with cheliceral setae 10. Legs. Length of legs: I 153 (145 – 160), II 128 (125 – 137), III 142 (135 – 150), IV 152 (145 – 157); length of tarsi: I 57 (55 – 60), II 45 (42 – 47), III 39 (37 – 42), IV 41 (40 – 45). The leg chaetotaxy differs from that of female as follows: genua I — IV, 2 asl, 1 bsl, {1 asl, 1 sts}, 4 sts — 1 asl, 1 bsl, 5 sts — 1 asl, 5 sts — 1 asl, 5 sts.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E151FF9FFF77FF082284F826.taxon	materials_examined	Type material. Holotype female, from leaf litter of Euterpe edulis Mart. (Arecaceae), 11. VII. 2000, Pariquera-Açu, A. R. Oliveira. Paratypes. Cananéia: 1 female, from leaf litter A. aculeatissimum, 18. I. 2000, A. R. Oliveira; 1 male, from leaf litter of E. edulis, 12. VII. 2000, A. R. Oliveira; 1 female, from leaf litter of Syagrus romanzoffiana (Cham.) (Euphorbiaceae), 18. I. 00, A. R. Oliveira. Pariquera-Açu: 1 female, from leaf litter of of S. romanzoffiana, 11. VII. 2000, A. R. Oliveira; 1 male, from roots of E. edulis, 18. IV. 2000, A. R. Oliveira; 1 male, from roots of E. edulis, 11. VII. 2001, A. R. Oliveira.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E151FF9FFF77FF082284F826.taxon	description	Pirassununga: 3 females, from soil, 27. VII. 2000, A. R. Oliveira. São Carlos: 2 females, from soil, 25. VII. 2000, A. R. Oliveira. São Pedro: 4 females, from leaf litter of Syagrus oleraceae (Mart.) Becc. (Arecaceae), 10. VIII. 2000, A. R. Oliveira.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E151FF9FFF77FF082284F826.taxon	etymology	Etymology. The species designation refers to the presence of four solenidia on genu I.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E15CFF92FF77FF0822BFFC77.taxon	etymology	Etymology: The genus name is an anagram of the original name Eupalus, provided by Koch (1838), for a three-jointed cunaxid.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E15CFF92FF77FF0822BFFC77.taxon	diagnosis	Diagnosis of the genus Lupaeus. This genus can be distinguished from other genera of the tribe Pulaeini by the presence of tarsus I with depression distally positioned; basifemur IV with 1 sts; genital seta g 3 near outer margin of the genital valve; palp tibiotarsus with two- or three- pointed process.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E15CFF93FF77FA10225EFD4A.taxon	description	Female (n = 9). Idiosoma: length 254 (200 – 287) and width 178 (175 – 205). Dimensions of sensillae and setae: vi 78 (70 – 87), ve 29 (25 – 32), sci 19 (17 – 22), sce 82 (75 – 85), c 1 16 (15 – 20), c 2 16 (15 – 17), d 1 16 (15 – 17), e 1 19 (17 – 20), f 1 25, f 2 12, h 1 24 (22 – 25) and h 2 15. Hypostome 99 (90 – 105) long and hg- setae: hg 1 12, hg 2 12, hg 3 26 (25 – 30) and hg 4 11 (10 – 12). Three-jointed palp 54 (52 – 57) long. Chelicera 100 (92 – 110) long and cheliceral seta 12. Length of legs: I 152 (150 – 155), II 136 (130 – 150), III 154 (150 – 162), IV 172 (165 – 175); length of tarsi: I 55 (52 – 57), II 45 (42 – 50), III 42 (40 – 45), IV 46 (42 – 50). Male. (n = 3). Idiosoma: length 205 (170 – 225) and width 145 (137 – 157). Dimensions of sensillae and setae: vi 69 (62 – 76), ve 25 (20 – 28), sci 17 (15 – 19), sce 68 (62 – 75), c 1 17 (15 – 19), c 2 17 (15 – 18), d 1 17 (15 – 18), e 1 17 (15 – 19), f 1 20 (17 – 21), f 2 9 (9 – 10), h 1 18 (15 – 20) and h 2 12. Hypostome 90 long and hg - setae: hg 1 11 (1 – 12), hg 2 13 (10 – 15), hg 3 30 (25 – 33) and hg 4 10 (10 – 11). Three-jointed palp 56 (55 – 57) long. Chelicera 84 (77 – 87) long and cheliceral setae 10. Length of legs: I 133 (125 – 140), II 116 (110 – 122), III 131 (122 – 140), IV 140 (127 – 152); length of tarsi: I 50 (47 – 52), II 41 (37 – 45), III 39 (35 – 42), IV 41 (37 – 45).	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E15CFF93FF77FA10225EFD4A.taxon	discussion	Remarks. The specimens examined differ from the original description of the species in relation to the number of tactile setae on tarsi II – IV that are 17 — 16 — 16 sts instead of 18 — 17 — 17 sts. Specimens examined. Pariquera-Açu: 1 female, from fruits of A. aculeatissimum, 11. VII. 2000, G. P. Arruda Filho; 2 females, from leaf litter of E. edulis, 13. VII. 2000, A. R. Oliveira; 3 females, from leaf litter of Syagrus romanzoffiana (Cham.) (Euphorbiaceae), 11. VII. 2000, A. R. Oliveira; 2 females, from leaf litter of S. romanzoffiana, 16. I. 2000, A. R. Oliveira; 1 female, from soil under S. romanzoffiana, 16. IV. 2000, A. R. Oliveira; 4 females, from leaf litter of S. romanzoffiana, 12. X. 2000, A. C. Pereira; 3 females, from soil under S. romanzoffiana, 12. X. 2000, A. C. Oliveira; 2 males, from leaf litter of S. romanzoffiana, 12. X. 2000, A. C. Pereira; 3 males, from leaf litter of S. romanzoffiana, 18. IV. 2000, A. C. Pereira; 1 male, from leaf litter of E. edulis, 13. VII. 2000, A. R. Oliveira.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E15CFF93FF77FA10225EFD4A.taxon	description	Cananéia: 4 females, from leaf litter of Bactris setosa Mart. (Arecaceae), 12. VII. 2000, A. R. Oliveira. Piracicaba: 2 females, from soil under Acrocomia aculeata Lodd. Ex Mart. (Arecaceae), 11. XI. 2000, A. R. Oliveira; 3 females, from soil under A. aculeata, 11. VIII. 2000, A. R. Oliveira.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E15DFF93FF77FD43225AF943.taxon	description	Female (n = 13) Idiosoma: length 238 (212 – 257) and width 166 (150 – 187). Dimensions of sensillae and setae: vi 72 (62 – 77), setae ve 34 (30 – 40), setae sci 21 (17 – 25), sensillae sce 74 (72 – 77), c 1 24 (20 – 27), c 2 16 (12 – 20), d 1 24 (22 – 25), e 1 25 (22 – 27), f 1 26 (25 – 32), f 2 12 (10 – 12), h 1 22 (17 – 25) and h 2 13 (12 – 15). Hypostome 88 (77 – 92) long and hg- setae: hg 1 11 (10 – 12), hg 2 12 (12 – 15), hg 3 21 (17 – 22) and hg 4 8 (7 – 10). Three-jointed palp 49 (45 – 55) long. Chelicera 87 (75 – 92) long and cheliceral seta 10 (7 – 12). Length of legs: I 148 (140 – 157), II 128 (125 – 137), III 145 (140 – 150), IV 162 (160 – 172); length of tarsi: I 52 (50 – 55), II 42 (40 – 45), III 41 (40 – 45), IV 43 (40 – 47).	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E15DFF93FF77FD43225AF943.taxon	discussion	Remarks. The specimens examined differ from the original description of the species in relation to the number of tactile setae on tarsi I and II, in the original description it is given as 21 (22) and 18, respectively, instead of 19 (20) and 17. Specimens examined. Cananéia: 1 female, from leaves of B. setosa, 12. VII. 2000, G. P. Arruda Filho; 2 females, from leaves of an unidentified plant, 17. VII. 2002, N. C. Mesa.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E15DFF93FF77FD43225AF943.taxon	description	Luis Antônio: 1 female, from leaf litter, 26. VII. 2000, A. R. Oliveira. Pariquera-Açu: 1 female, from leaves of B. setosa Mart., 16. I. 2000, L. V. F. Silva; 1 female, from leaves of B. setosa, 11. X. 1998, M. G. C. Gondim Júnior; 1 female, from leaves of Alchornea glandulosa Poepp. & Endl. (Euphorbiaceae), 19. IV. 1998; 1 female, from leaves of Aparisthmium cordatum (A. Juss.) Baill. (Euphorbiaceae), 19. IV. 1998, M. S. Zacarias; 1 female, from leaves of A. cordatum, 17. VII. 2002, N. C. Mesa. Piracicaba: 7 females, from leaf litter of E. edulis Mart., 16. V. 2000, A. R. Oliveira. Pirassununga: 1 female, from leaf litter of Psidium guineense Sw. (Myrtaceae), 27. VII. 2000, A. R. Oliveira. São Pedro: 1 female, from leaf litter of S. oleraceae, 16. II. 2000, A. R. Oliveira.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E15DFF96FF77F97A23A9FB57.taxon	diagnosis	Diagnosis. This species resembles L. martini, but differs from the latter by the presence of a dorsal shield with subcuticular, instead of smooth, punctations, and basifemur II with 5, instead of, 6 setae.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E15DFF96FF77F97A23A9FB57.taxon	description	Female (Fig. 5 – 6). (n = 3). Dimensions. Idiosoma: length 231 (225 – 237) and width 173 (167 – 177). Dorsum (Fig. 5 A). The dorsal shield with subcuticular punctations. Around the sensillae vi is provided by a lobed area. Dorsal shield with one pair each of sensillae vi 72 (67 – 75), setae ve 37 (35 – 40), setae sci 22 (20 – 25), sensillae sce 75, c 1 27 (25 – 30), c 2 16 (15 – 17), d 1 25 (22 – 27) and e 1 25 (20 – 30). Posteriad to the dorsal idiosomal shield one pair of tiny platelets occur carring setae f 1 27 and f 2 11 (10 – 12) each. Setae h 1 21 (20 – 22) and h 2 9 occur on the integument. Ve n t e r (Fig. 5 B). The propodosomal coxal region is completely divided. One pair of paracoxal setae occurs on the coxal region IV. The integument bears 1 pair of propodogastral setae, 6 or 7 hyterogastral setae. Gnathosoma (Fig. 5 C – G). Hypostome 86 (80 – 92) long with 4 pairs of hg - setae [hg 1 10, hg 2 10, hg 3 21 (17 – 25) and hg 4 10] and 2 pairs of adoral setae. Three-jointed palp, 50 long. Trochanter without setae; femorogenu, 6 sts; tibiotarsus, 4 sts, 1 asl terminal, a large bladder-like apophysis and 2 pointed process. Chelicera 89 (87 – 92) long with cheliceral seta 10. Legs (Fig. 6 A – D). Length of legs: I 141 (135 – 145), II 123 (122 – 125), III 138 (137 – 140), IV 157 (155 – 160), length of tarsi: I 51 (50 – 52), II 41 (40 – 42), III 39 (37 – 40), IV 40. The leg chaetotaxy is as follows: coxae I — IV, 1 peg, 3 sts — 3 sts — 3 sts — 2 sts; trochanters I — IV, 1 sts — 1 sts — 2 sts — 1 sts; basifemora I — IV, 4 sts — 5 sts — 3 sts — 1 sts; telofemora I — IV, 5 sts — 5 sts — 1 ms, 3 sts — 1 ms, 2 sts; genua I — IV, 3 asl, {1 asl, 1 sts}, 4 sts — 2 asl, 5 sts — 1 asl, 5 sts — 2 asl, 5 sts; tibiae I — IV, 1 asl, 1 bsl, 5 sts — 1 bsl, 5 sts — 1 bsl, 5 sts — 1 T, 4 sts; tarsi I — IV, 4 bsl, 1 dtsl, 1 dep, 2 tsl, 17 sts — 1 bsl, 1 dtsl, 1 tsl, 17 (18) sts — 1 tsl, 16 sts — 15 sts. Male (Fig. 6). (n = 2). Dimensions. Idiosoma: length 175 and width 131 (130 – 132). Dorsum (Fig. 6 E). The dorsum is similar to that of the female. Dorsal shield with one pair of sensillae vi 67 (65 – 70), setae ve 33 (32 – 35), setae sci 18 (17 – 20), sensillae sce 66 (65 – 67), c 1 22 (20 – 25), c 2 13 (12 – 15), d 1 21 (17 – 25) and e 1 18 (17 – 20). Posteriad to the dorsal idiosomal shield one pair of tiny platelets occur carring setae f 1 21 (20 – 22) and f 2 7 each. Setae h 1 17 and h 2 8 occur on the integument. Ve n t e r (Fig. 6 F). The propodosomal coxal plates fused medially. One pair of paracoxal setae occurs on the coxal region IV. The integument bears 1 pair of propodogastral setae and 1 pair of hysterogastral setae. The genital valves are smaller than those of the females, each bearing 4 g- setae. Gnathosoma. The gnathosoma is almost similar to that of the female. Hypostome 75 long with 4 pairs of hg - setae [hg 1 7, hg 2 10, hg 3 20 and hg 4 8]. Three-jointed palp, 47 long. Chelicera 72 (70 – 75) long with cheliceral setae 7. Legs. Length of legs: I 125 (120 – 130), II 108 (105 – 112), III 115, IV 134 (132 – 137), length of tarsi: I 47, II 36 (35 – 37), III 35, IV 36 (35 – 37). The leg chaetotaxy differs from that of the female as follows: genua I — IV, 2 asl, 1 bsl, {1 asl, 1 sts}, 4 sts — 1 asl, 1 bsl, 5 sts — 1 asl, 1 bsl, 5 sts — 1 asl, 1 bsl, 5 sts; tibiae I, 1 asl, 1 bsl.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E15DFF96FF77F97A23A9FB57.taxon	materials_examined	Type material. Holotype female, from leaf litter of Myrcia sp. (Myrtaceae), 27. VII. 2000, Pirassununga, A. R. Oliveira. Paratypes. Pariquera- Açu: 1 female, from leaf litter of A. aculeatissimum, 11. VII. 2000, A. R. Oliveira; 1 male, from leaf litter of A. aculeatissimum, 11. VII. 2000, A. R. Oliveira; 1 female, from leaf litter of A. aculeatissimum, 18. IV. 2000, A. R. Oliveira; 1 male, from leaf litter of A. aculeatissimum, 18. IV. 2000, A. R. Oliveira.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E15DFF96FF77F97A23A9FB57.taxon	etymology	Etymology. The species designation refers to the substrate litter (- lectus) in which the species was found.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E158FF95FF77FB6626D9FC7F.taxon	diagnosis	Diagnosis. This species resembles L. clarae, but differs from the latter by the presence of basifemur II with 5 setae.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E158FF95FF77FB6626D9FC7F.taxon	description	Female (Fig. 7 – 8). (n = 6). Dimensions. Idiosoma: length 187 (175 – 200) and width 129 (117 – 150). Dorsum (Fig. 7 A). The dorsal shield with transverse lobes, except for the region lateral to the sensillae sce, which is smooth. Dorsal shield with one pair each of sensillae vi 63 (60 – 70); setae ve 20 (17 – 22), setae sci 14 (12 – 17), sensillae sce 72 (67 – 75), c 1 12 (10 – 13), c 2 13 (11 – 15), d 1 12 (11 – 13) and e 1 13 (12 – 15). Posteriad to the dorsal idiosomal shield one pair of tiny platelets occur carring setae f 1 16 (15 – 17) and f 2 11 (10 – 12) each. Setae h 1 17 (15 – 20) and h 2 9 occur on the integument. Ve n te r (Fig. 7 B). The propodosomal coxal region is medially completely divided. One pair of paracoxal setae occurs on the coxal region IV. The integument bears 1 pair of propodogastral setae and 3 pairs of hysterogastral setae. Gnathosoma (Fig. 7 C – E). Hypostome 83 (77 – 87) long with 4 pairs of hg - setae [hg 1 8 (7 – 9), hg 2 9 (8 – 10), hg 3 17 (15 – 20), hg 4 7]. Three-jointed palp, 43 (42 – 45) long. Palp tibiotarsi with 3 pointed process. Chelicera 82 (77 – 85) long with cheliceral setae 10 (9 – 10). Legs (Fig. 8 A – D). Length of legs: I 115 (112 – 125), II 101 (97 – 112), III 112 (105 – 117), IV 130 (120 – 147); length of tarsi: I 42 (40 – 45), II 33 (30 – 35), III 29 (25 – 32), IV 32 (27 – 35). The leg chaetotaxy is as follows: coxae I — IV, 1 peg, 3 sts — 3 sts — 3 sts — 2 sts; trochanters I — IV, 1 sts — 1 sts — 2 sts — 1 sts; basifemora I — IV, 4 sts — 5 sts — 3 sts — 1 sts; telofemora I — IV, 5 sts — 5 sts — 1 ms, 3 sts — 1 ms, 2 sts; genua I — IV, 3 asl, {1 asl, 1 sts}, 4 sts — 2 asl, 5 sts — 1 asl, 5 sts — 2 asl, 5 sts; tibiae I — IV, 1 asl, 1 bsl, 5 sts — 1 bsl, 5 sts — 1 bsl, 5 sts — 1 T, 4 sts; tarsi I — IV, 2 asl, 2 bsl, 1 dep, 1 dtsl, 2 tsl, 19 sts — 1 bsl, 1 tsl, 1 dtsl, 18 sts — 1 tsl, 16 sts — 15 sts. Male (Fig. 8). (n = 2). Dimensions. Idiosoma: length 221 (205 – 233) and width 115 (110 – 120). Dorsum (Fig. 8 E). The dorsum is similar to that of the female. Dorsal shield with one pair each of sensillae vi 57 (54 – 61), setae ve 16 (15 – 17), setae sci 13 (12 – 15), sensillae sce 60 (57 – 64), c 1 9 (9 – 10), c 2 11 (10 – 12), d 1 10 and e 1 10 (10 – 11). Posteriad to the dorsal idiosomal shield one pair of tiny platelets occur carring setae f 1 13 (11 – 15) and f 2 7 (7 – 8) each. Setae h 1 11 (11 – 12) and h 2 6 occur on the integument. Ve n t e r (Fig. 8 F). The propodosomal coxal plates fused medially. The integument bears 1 pair of propodogastral setae and 1 pair of paracoxal setae. The genital valves are smaller than those of the females, each bearing 4 g- setae. Gnathosoma. The gnathosoma is similar to that of the female. Hypostome 70 long with 4 pairs of hg - setae: hg 1 7 (7 – 8), hg 2 8 (8 – 9), hg 3 15 and hg 4 6 (6 – 7). Three-jointed palp, 37 (35 – 40) long. Chelicera 63 (62 – 65) long with cheliceral setae 10 (10 – 11). Legs. Length of legs: I 100, II 87 (84 – 90), III 101 (100 – 103), IV 108 (107 – 110), length of tarsi: I 36 (35 – 37), II 29, III 27, IV 29 (28 – 30). The leg chaetotaxy differs from the female as follows: genua II, 1 asl, 1 bsl, 4 sts.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E158FF95FF77FB6626D9FC7F.taxon	materials_examined	Type material. Holotype female, from soil under Campomanesia sp. (Myrtaceae), 27. VII. 2000, Pirassununga, A. R. Oliveira. Paratypes. Luis Antônio: 2 females, from soil under Psidium sp., 26. VII. 2000, A. R. Oliveira; Pirassununga: 2 females, from soil under Myrcia sp., 03. V. 2000, A. R. Oliveira; 1 female, from soil under Myrcia sp., 27. VII. 2000, A. R. Oliveira; 1 female, from soil, 03. V. 2000, A. R. Oliveira; 1 female from soil under Campomanesia sp., 27. VII. 2000, A. R. Oliveira; 5 females, from soil under a Myrtaceae plant, 27. VII. 2000, A. R. Oliveira; 1 male, from soil under Campomanesia sp., 25. VII. 2000, A. R. Oliveira.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E158FF95FF77FB6626D9FC7F.taxon	description	São Carlos: 1 female, from soil under Myrcia sp., 25. VII. 2000, A. R. Oliveira; 1 male, from soil under Myrcia sp., 25. VII. 2000, A. R. Oliveira; 1 female, from soil under Myrtaceae plant, 25. VII. 2000, A. R. Oliveira.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
4C578782E158FF95FF77FB6626D9FC7F.taxon	etymology	Etymology. The species designation refers to the presence of lobes (lobus-) on the dorsal (- dorsalis) shield.	en	Castro, Tatiane Marie Martins Gomes De, Heyer, Jacob Den (2009): A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae). Zootaxa 2141: 20-36, DOI: 10.5281/zenodo.188565
