taxonID	type	description	language	source
4E6387C65F779D47BE61FB761CCBF93C.taxon	description	urn: lsid: zoobank. org: act: 3 BED 6271 - ED 7 E- 4 CAA- 8 F 69 - 1 DC 48 A 68 D 7 E 1 Figs 3 – 8	en	Jonishi, Taro, Nakano, Takafumi (2023): Two new species of Scolopocryptops centipedes from southern Japan (Chilopoda: Scolopendromorpha: Scolopocryptopidae). European Journal of Taxonomy 908: 155-182, DOI: 10.5852/ejt.2023.908.2345, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2345/10223
4E6387C65F779D47BE61FB761CCBF93C.taxon	diagnosis	Diagnosis Antenna with sparse minute hairs and short setae on dorsal surface of basal two or three articles, subsequent articles densely setose. Cephalic plate with short lateral marginal sulci in posterior half. Forcipular coxosternite with anterior margin weakly bilobed, bearing darkly sclerotized bands almost reaching outer part, with a pair of small teeth. Coxopleuron approx. 1.5 – 1.7 × as long as sternite 23; pleural dorsal margin slightly protruding from lateral margin of tergite 23, posterior and ventral margins forming approx. 70 – 80 ° angle; coxopleural process short.	en	Jonishi, Taro, Nakano, Takafumi (2023): Two new species of Scolopocryptops centipedes from southern Japan (Chilopoda: Scolopendromorpha: Scolopocryptopidae). European Journal of Taxonomy 908: 155-182, DOI: 10.5852/ejt.2023.908.2345, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2345/10223
4E6387C65F779D47BE61FB761CCBF93C.taxon	etymology	Etymology The specific name is dedicated to the late Dr Yasunori Miyosi, who first provided detailed taxonomic accounts for this new species.	en	Jonishi, Taro, Nakano, Takafumi (2023): Two new species of Scolopocryptops centipedes from southern Japan (Chilopoda: Scolopendromorpha: Scolopocryptopidae). European Journal of Taxonomy 908: 155-182, DOI: 10.5852/ejt.2023.908.2345, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2345/10223
4E6387C65F779D47BE61FB761CCBF93C.taxon	vernacular_names	We herein suggest a Japanese name for this species as ‘ Miyosi-akamukade’.	en	Jonishi, Taro, Nakano, Takafumi (2023): Two new species of Scolopocryptops centipedes from southern Japan (Chilopoda: Scolopendromorpha: Scolopocryptopidae). European Journal of Taxonomy 908: 155-182, DOI: 10.5852/ejt.2023.908.2345, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2345/10223
4E6387C65F779D47BE61FB761CCBF93C.taxon	materials_examined	Material examined Holotype JAPAN – Kyushu • ♂; Kagoshima, near Kirishima Mountains (Figs 1, 3 A); 31.88765 ° N, 130.83962 ° E; 616 m alt.; 2 Aug. 2022; T. Jonishi leg.; KUZ Z 4375. Paratypes JAPAN – Kyushu • 1 ♂; same locality as for holotype; 31.88796 ° N, 130.84037 ° E; 626 m alt.; 2 Aug. 2022; T. Jonishi leg.; KUZ Z 4376 • 1 ♀; Kagoshima, Aira; 31.86924 ° N, 130.59942 ° E; 364 m alt.; 2 Aug. 2022; T. Jonishi leg.; KUZ Z 4377. Additional material JAPAN – Kyushu • 1 subadult; Oita, Saeki; 32.93942 ° N, 131.72764 ° E; 71 m alt.; 20 Jun. 2022; Naoto Sawada leg.; KUZ Z 4374. – Amami Island • 1 subadult; Uken; 28.28792 ° N, 129.31997 ° E; 435 m alt.; 13 Mar. 2021; Futaro Okuyama leg.; KUZ Z 4378 • 1 subadult; Uken, Mt. Yuwan-dake; 28.28962 ° N, 129.31441 ° E; 506 m alt.; 20 Jun. 2022; T. Jonishi leg.; KUZ Z 4379 • 2 subadults; Yamato; 28.33112 ° N, 129.36187 ° E; 315 m alt.; 20 Jun. 2022; T. Jonishi leg.; KUZ Z 4380, Z 4381 • 1 ♀; same locality as for preceding; 28.33104 ° N, 129.36158 ° E; 314 m alt.; 21 Jun. 2022; T. Jonishi leg.; KUZ Z 4382 • 1 subadult; same locality as for preceding; 28.33090 ° N, 129.36163 ° E; 317 m alt.; 22 Jun. 2022; T. Jonishi leg.; KUZ Z 4383 • 1 ♀; same locality as for preceding; 28.33040 ° N, 129.36076 ° E; 301 m alt.; 22 Jun. 2022; T. Jonishi leg.; KUZ Z 4384.	en	Jonishi, Taro, Nakano, Takafumi (2023): Two new species of Scolopocryptops centipedes from southern Japan (Chilopoda: Scolopendromorpha: Scolopocryptopidae). European Journal of Taxonomy 908: 155-182, DOI: 10.5852/ejt.2023.908.2345, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2345/10223
4E6387C65F779D47BE61FB761CCBF93C.taxon	description	Description of holotype [variation in other specimens given in square brackets] Body length 54.4 mm [48.0 – 69.1 mm in adults] in 80 % ethanol, 60.3 mm [54.6 – 73.4 mm] before fixation. Body color brownish orange, antennae, legs, and ultimate legs yellowish (Fig. 3 B). Antennae 10.0 mm in length, approx. 0.18 × [0.16 – 0.23 ×] as long as body, composed of 17 articles; basal 3 articles with sparse minute hairs (see Bonato et al. 2010) and short setae dorsally, 4 th and subsequent articles densely setose [3 rd article moderately setose, articles 4 – 17 more densely setose in KUZ Z 4377 and Z 4382] (Fig. 4 A – B). Cephalic plate as long as wide, with sides converging anteriorly (Figs 4 C – D, 5 A); its surface finely punctate [minute setae on each punctum present in KUZ Z 4382 and subadults]; short lateral marginal sulci present in posterior half [marginal sulci significantly longer and more apparent in KUZ Z 4377] (Figs 4 C – D, 5 A). Second maxillae article 2 with elongated and semi-transparent dorsal spur distally; dorsal brush without transparent margin (Fig. 5 B – C); pretarsus consisting of dark brown basal and semi-transparent short apical parts (Fig. 5 B – C). Forcipular coxosternite and trochanteroprefemora sparsely punctate, coxosternite without sutures (Fig. 5 D); forcipular trochanteroprefemur with small and blunt black process, with apparent basal suture (Figs 5 D, 6); anterior margin of coxosternite weakly convex and divided by median diastema; darkly sclerotized bands almost reaching outer part of anterior margin of coxosternite, with a pair of small but prominent teeth at middle of each side [right tooth absent in KUZ Z 4377] (Figs 5 D, 6). Tergites finely punctate; tergite 1 with anterior transverse suture, anterior margin covered by cephalic plate (Figs 5 A, 7 A). Paramedian sutures present on tergites 2 – 22 (Fig. 7 A); lateral marginations complete on tergites 6 – 21 (Fig. 7 A). Sternites lacking paramedian sutures, finely punctate (Fig. 7 B); sternites 3 – 20 with shallow longitudinal sulcus and slight median depression (Fig. 7 B), both shallower and unapparent on anterior sternites. Sides of sternite 23 converging posteriorly, its posterior margin slightly concave (Fig. 7 C). Spiracles ovoid, present on leg-bearing segments 3, 5, 8, 10, 12, 14, 16, 18, 20, and 22. Legs lacking setae; tarsi of legs 1 – 21 undivided; legs 1 – 19 [1 – 20 in KUZ Z 4376] with lateral and ventral tibial spurs and tarsal spur, legs 20 and 21, respectively, with tibial spur and tarsal spur [leg 21 without tibial spur in KUZ Z 4382]; leg 22 without spurs. All legs with two accessory spines. Coxopleuron approx. 1.7 × [1.5 – 1.7 ×] as long as sternite 23 (Fig. 7 D). Dorsal margin of ultimate pleuron slightly protruding from lateral side of tergite 23 (Fig. 7 D – E), posterior margin with minute dark spine (Fig. 7 D). Posterior and ventral margins of coxopleuron converging posteriorly, forming approx. 80 ° [70 – 80] angle (Fig. 7 D); coxopleural process short, tip of process pointed, slightly directed dorsally (Fig. 7 D). Surface of coxopleuron without setae, covered with various sized coxal pores; coxopleural process and dorso-posterior area of coxopleuron poreless (Fig. 7 D). Ultimate leg 14.1 mm [13.9 – 17.3 mm] in length, 0.26 × [0.24 – 0.3 ×] as long as body; all articles lacking setae (Fig. 8 A); prefemur with two conical and pointed spinose processeses, ventral process large, dorso-medial one minute; pretarsus with two short accessory spines. Genital segments occupying approx. ½ length of sternite 23. Sternite of genital segment 1 with sparse minute setae; posterior margin weakly convex (Fig. 8 B). Sternite of genital segment 2 well developed, covered with sparse minute setae; posterior part of genital segment 2 overlapped by lamina subanalis, penis not visible in ventral view (Fig. 8 B); lamina subanalis situated between genital segment 2 and anal valves (Fig. 8 B) [in female paratype KUZ Z 4377, genital segment 1 as described for holotype, genital segment 2 absent; lamina subanalis situated between genital segment 1 and anal valves] (Fig. 8 C).	en	Jonishi, Taro, Nakano, Takafumi (2023): Two new species of Scolopocryptops centipedes from southern Japan (Chilopoda: Scolopendromorpha: Scolopocryptopidae). European Journal of Taxonomy 908: 155-182, DOI: 10.5852/ejt.2023.908.2345, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2345/10223
4E6387C65F779D47BE61FB761CCBF93C.taxon	discussion	Remarks This species has previously been identified as S. curtus (Miyosi 1961, 1971; Ogawa 1961), but S. miyosii sp. nov. can be distinguished by the following characters: the presence of short sulci along the lateral margin of the cephalic plate (Fig. 4 C – D) (vs sulci absent in S. curtus), the convex anterior margin of the forcipular coxosternite with teeth (Figs 5 D, 6) (vs anterior margin virtually straight with teeth almost reduced, forming antero-lateral corner of sclerotized bands; also see Remarks for S. brevisulcatus sp. nov. below), and the dorsal margin of the ultimate pleuron protruding from the lateral side of tergite 23 (Fig. 7 E) (vs dorsal margin flattened) (see Table 3).	en	Jonishi, Taro, Nakano, Takafumi (2023): Two new species of Scolopocryptops centipedes from southern Japan (Chilopoda: Scolopendromorpha: Scolopocryptopidae). European Journal of Taxonomy 908: 155-182, DOI: 10.5852/ejt.2023.908.2345, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2345/10223
4E6387C65F779D47BE61FB761CCBF93C.taxon	distribution	Distribution Known from Kyushu, and Amami Island in the Ryukyu Islands.	en	Jonishi, Taro, Nakano, Takafumi (2023): Two new species of Scolopocryptops centipedes from southern Japan (Chilopoda: Scolopendromorpha: Scolopocryptopidae). European Journal of Taxonomy 908: 155-182, DOI: 10.5852/ejt.2023.908.2345, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2345/10223
4E6387C65F729D59BE0AF91F1BE0F93C.taxon	description	urn: lsid: zoobank. org: act: 3 C 9 DB 52 B- 2691 - 4980 - B 339 - 1 B 31 E 177 F 10 C Figs 9 – 14	en	Jonishi, Taro, Nakano, Takafumi (2023): Two new species of Scolopocryptops centipedes from southern Japan (Chilopoda: Scolopendromorpha: Scolopocryptopidae). European Journal of Taxonomy 908: 155-182, DOI: 10.5852/ejt.2023.908.2345, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2345/10223
4E6387C65F729D59BE0AF91F1BE0F93C.taxon	diagnosis	Diagnosis Antenna with sparse minute hairs and short setae on dorsal surface of basal four articles, subsequent articles densely setose. Cephalic plate with short and shallow lateral marginal sulci in posterior half. Forcipular coxosternite with anterior margin slightly bilobed but median diastema almost lacking, darkly sclerotized bands not reaching outer part of anterior margin, with a pair of almost-reduced teeth. Coxopleuron approx. 1.6 – 1.7 × as long as sternite 23; pleural dorsal margin slightly protruding from lateral margin of tergite 23, posterior and ventral margins forming approx. 70 – 80 ° angle; coxopleural process short.	en	Jonishi, Taro, Nakano, Takafumi (2023): Two new species of Scolopocryptops centipedes from southern Japan (Chilopoda: Scolopendromorpha: Scolopocryptopidae). European Journal of Taxonomy 908: 155-182, DOI: 10.5852/ejt.2023.908.2345, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2345/10223
4E6387C65F729D59BE0AF91F1BE0F93C.taxon	etymology	Etymology The specific name is derived from the Latin words ‘ brevis ’ (‘ short, shallow’) and ‘ sulcus ’ (‘ groove’), referring to the unapparent cephalic marginal sulcus of this new species.	en	Jonishi, Taro, Nakano, Takafumi (2023): Two new species of Scolopocryptops centipedes from southern Japan (Chilopoda: Scolopendromorpha: Scolopocryptopidae). European Journal of Taxonomy 908: 155-182, DOI: 10.5852/ejt.2023.908.2345, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2345/10223
4E6387C65F729D59BE0AF91F1BE0F93C.taxon	vernacular_names	We herein suggest a Japanese name for this species as ‘ Kirekomi-akamukade’.	en	Jonishi, Taro, Nakano, Takafumi (2023): Two new species of Scolopocryptops centipedes from southern Japan (Chilopoda: Scolopendromorpha: Scolopocryptopidae). European Journal of Taxonomy 908: 155-182, DOI: 10.5852/ejt.2023.908.2345, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2345/10223
4E6387C65F729D59BE0AF91F1BE0F93C.taxon	materials_examined	Material examined Holotype JAPAN – Okinawa Island • ♂; Motobu, near Mt. Katsuu-dake (Figs 1, 9 A); 26.63157 ° N, 127.93771 ° E; 311 m alt.; 14 Jun. 2022; T. Jonishi leg.; KUZ Z 4390. Paratypes JAPAN – Okinawa Island • 1 subadult; Kunigami, near Mt. Nishime-dake; 26.80745 ° N, 128.26618 ° E; 298 m alt.; 4 May 2021; F. Okuyama leg.; KUZ Z 4386 • 1 adult, sex undetermined; same locality as for preceding; 26.81195 ° N, 128.28415 ° E; 107 m alt.; 8 Apr. 2022; F. Okuyama leg; KUZ Z 4387 • 1 ♀; same collection data as for holotype; KUZ Z 4389 • 1 ♂; Kunigami, near Mt. Nishime-dake; 26.80080 ° N, 128.27728 ° E; 355 m alt.; 15 Jun. 2022; T. Jonishi leg.; KUZ Z 4391 • 1 subadult; Kunigami, Mt. Fuenchiji; 26.75251 ° N, 128.24303 ° E, 384 m alt.; 16 Jun. 2022; T. Jonishi leg.; KUZ Z 4392. Additional material JAPAN – Okinawa Island • 1 adult, sex undetermined; same locality as for holotype; 26.63145 ° N, 127.93812 ° E; 291 m alt.; 20 Sep. 2019; T. Jonishi leg.; KUZ Z 4385 • 1 subadult; same locality as for holotype; 26.63139 ° N, 127.93792 ° E; 307 m alt.; 14 Jun. 2022; T. Jonishi leg.; KUZ Z 4388 • 1 juv.; Ogimi, near Mt. Nekumachiji; 26.68491 ° N, 128.13430 ° E; 244 m alt.; 16 Jun. 2022; T. Jonishi leg.; KUZ Z 4393. – Izena Island • 1 ♀; Mt. Ufu-yama; 26.94288 ° N, 127.92713 ° E; 48 m alt.; 8 Nov. 2022; Ryobu Fukuyama leg.; KUZ Z 4394.	en	Jonishi, Taro, Nakano, Takafumi (2023): Two new species of Scolopocryptops centipedes from southern Japan (Chilopoda: Scolopendromorpha: Scolopocryptopidae). European Journal of Taxonomy 908: 155-182, DOI: 10.5852/ejt.2023.908.2345, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2345/10223
4E6387C65F729D59BE0AF91F1BE0F93C.taxon	description	Description of holotype [variation in other specimens given in square brackets] Body length 58.0 mm [43.6 – 59.1 mm in adults] in 80 % ethanol, 64.0 mm before fixation. Body color brownish red [brownish orange]; antennae, legs, and ultimate legs yellowish (Fig. 9 B). Antennae damaged, 13 th and subsequent articles lost [9.9 mm in length, approx. 0.2 × as long as body, composed of 17 articles in paratype KUZ Z 4391]; density of setae increasing toward distal articles; basal 4 articles with sparse minute hairs (see Bonato et al. 2010) and short setae dorsally, 5 th and subsequent articles setose, 6 th – 17 th articles more densely covered with setae [basal 2 articles sparsely setose, 3 rd and subsequent articles densely covered with setae in juvenile KUZ Z 4393; 4 th article moderately setose, 5 th – 17 th articles more densely covered with setae in subadults] (Fig. 10 A). Cephalic plate as long as wide, sides converging anteriorly (Figs 10 B, 11 A); its surface finely punctate [minute setae on each punctum present in juvenile and subadults]; short and shallow lateral marginal sulci present in posterior half [marginal sulci absent in juvenile and subadults] (Figs 10 B, 11 A). Second maxillae article 2 with elongated and semi-transparent dorsal spur distally; dorsal brush without transparent margin (Fig. 11 B – C); pretarsus consisting of dark brown basal and semi-transparent short apical parts (Fig. 11 B – C). Forcipular coxosternite and trochanteroprefemora sparsely punctate, coxosternite without sutures (Fig. 11 D); forcipular trochanteroprefemur with small and blunt black process, with apparent basal suture (Figs 11 D, 12); anterior margin of coxosternite almost straight and slightly bilobed but median diastema almost lacking; darkly sclerotized bands not reaching outer part of anterior margin of coxosternite; a pair of teeth almost reduced, forming antero-lateral corner of sclerotized bands [teeth on anterior margin prominent in juvenile and subadults] (Figs 11 D – E, 12). Tergites finely punctate; tergite 1 with anterior transverse suture, anterior margin covered by cephalic plate (Figs 11 A, 13 A). Paramedian sutures present on tergites 2 – 22, lateral marginations complete on tergites 6 – 21 (Fig. 13 A). Sternites lacking paramedian sutures, finely punctate (Fig. 13 B); sternites 2 – 20 with shallow longitudinal sulcus and slight median depression (Fig. 13 B), both shallower and unapparent on anterior sternites. Sternite 23 damaged, right posterior margin lost [sides of sternite 23 converging posteriorly, posterior margin slightly concave in paratype KUZ Z 4387 and other specimens] (Fig. 13 C). Spiracles ovoid, present on leg-bearing segments 3, 5, 8, 10, 12, 14, 16, 18, 20, and 22. Legs lacking setae; tarsi of legs 1 – 21 undivided; legs 1 – 19 with lateral and ventral tibial spurs and tarsal spur, legs 20 and 21, respectively, with tibial spur and tarsal spur; leg 22 without spurs. All legs with two accessory spines. Coxopleuron approx. 1.7 × [1.6 – 1.7 ×] as long as sternite 23 (Fig. 13 D). Dorsal margin of ultimate pleuron slightly protruding from lateral side of tergite 23 (Fig. 13 D – E), posterior margin with minute dark spine (Fig. 13 D). Posterior and ventral margins of coxopleuron converging posteriorly, forming approx. 70 ° [70 – 80 °] angle (Fig. 13 D); coxopleural process short, tip of process broken [tip of process pointed, slightly directed dorsally in KUZ Z 4387] (Fig. 13 D). Surface of coxopleuron without setae, covered with coxal pores of various size; coxopleural process and dorso-posterior area of coxopleuron poreless (Fig. 13 D). Ultimate leg 15.3 mm in length, 0.26 × as long as body; all articles lacking setae (Fig. 14 A); prefemur with two conical and pointed spinose processes, ventral process large, dorso-medial one minute; pretarsus with two accessory spines. Genital segments occupying approx. ½ length of sternite 23. Sternite of genital segment 1 with sparse minute setae; posterior margin weakly convex (Fig. 14 B). Sternite of genital segment 2 well developed, covered with sparse minute setae; posterior part of genital segment 2 overlapped by anal valves, penis not visible in ventral view (Fig. 14 B); lamina subanalis situated between genital segment 2 and anal valves (Fig. 14 B) [in female paratype KUZ Z 4389, genital segment 1 as described for holotype, genital segment 2 absent; lamina subanalis situated between genital segment 1 and anal valves] (Fig. 14 C).	en	Jonishi, Taro, Nakano, Takafumi (2023): Two new species of Scolopocryptops centipedes from southern Japan (Chilopoda: Scolopendromorpha: Scolopocryptopidae). European Journal of Taxonomy 908: 155-182, DOI: 10.5852/ejt.2023.908.2345, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2345/10223
4E6387C65F729D59BE0AF91F1BE0F93C.taxon	discussion	Remarks This species has previously been identified as S. elegans (e. g., Ômine 1987, 2002) and S. curtus (Omine & Ito 1998) and is quite similar to S. miyosii sp. nov. (see above). However, S. brevisulcatus sp. nov. can be distinguished by the combination of the following features: basal 4 antennal articles of adults with sparse setae (Fig. 10 A) (vs only basalmost article sparsely setose or all articles densely setose in S. elegans, and basal 2 or 3 articles with sparse setae in S. miyosii), the presence of short sulci along the lateral margin of the cephalic plate in adults (Fig. 10 B) (vs sulci absent in S. curtus), the anterior margin of coxosternite almost lacking median diastema (Figs 11 D, 12 A) (vs median diastema apparent in S. elegans, S. miyosii, and S. curtus), the sclerotized bands of coxosternite not reaching outer part of anterior margin (Figs 11 D, 12 A) (vs sclerotized bands almost reaching outer part in S. elegans and S. miyosii), the almost-reduced teeth on coxosternite (Figs 11 D, 12 A) (vs teeth prominent in S. miyosii), the dorsal margin of the ultimate pleuron protruding from the lateral margin of tergite 23 (Fig. 13 E) (vs dorsal margin flattened in S. curtus) and the short coxopleural process (Fig. 13 D) (vs the process moderately long in S. elegans) (see Table 3).	en	Jonishi, Taro, Nakano, Takafumi (2023): Two new species of Scolopocryptops centipedes from southern Japan (Chilopoda: Scolopendromorpha: Scolopocryptopidae). European Journal of Taxonomy 908: 155-182, DOI: 10.5852/ejt.2023.908.2345, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2345/10223
4E6387C65F729D59BE0AF91F1BE0F93C.taxon	distribution	Distribution Known from Okinawa Island and its adjacent islet (Izena Island), Ryukyu Islands.	en	Jonishi, Taro, Nakano, Takafumi (2023): Two new species of Scolopocryptops centipedes from southern Japan (Chilopoda: Scolopendromorpha: Scolopocryptopidae). European Journal of Taxonomy 908: 155-182, DOI: 10.5852/ejt.2023.908.2345, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2345/10223
