taxonID	type	description	language	source
4E6787A6FFF2FFB0AA9AF928FD6FF891.taxon	materials_examined	Type species: Branchiosyllis oculata Ehlers 1887.	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFF1FFB3AA9AFF57FC74F891.taxon	description	Figs 1 A, C, E	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFF1FFB3AA9AFF57FC74F891.taxon	materials_examined	Material examined. 6 specimens (1 mounted for SEM). Luzón Island, Balayan Bay: 2 specimens (MNCN 16.01 / 14319), “ Koala Point ”, 13 º 41 ' 51 " N – 120 º 49 ' 45 " E, Thalysias sp., Acanthella sp., 5 – 16 m, coral rubble, 1 – 2 m deep, 5 December 2010; 2 specimens (MNCN 16.01 / 14320), Sombrero Island, 13 º 41 ’ 52 ’’ N – 120 º 49 ’ 47 ’’ E, coral rubble, 17 m deep, 6 December 2010; 1 specimen, “ Beatrice Point ”, Sombrero Island, two different unidentified sponges, 5 – 24 m, 9 December 2010. Palawan Island, El Nido: 2 specimens (MNCN 16.01 / 14321), ” Twin Rocks ”, 11 º 17 ' 50 " N – 119 º 19 ' 06 " E, coral rubble, 6 m deep, 17 December, 2010. Comparative material. Branchiosyllis pacifica, Coiba National Park, Chiquirí Gulf, West Panamá: 1 specimen (MNCN 16.01 / 11585), Uvas Island, 7 º 49 ' 00 '' N – 81 º 46 ' 00 '' W, coral rubble, 3 m deep, 6 February 1997; 5 specimens (MNCN 16.01 / 11586), Canal de Afuera Island, 7 º 41 ' 50 '' N – 81 º 38 ' 25 '' W, coral rubble, 6 m deep, 12 February 1997; 1 specimen (MNCN 16.01 / 11588), Uvas Island, 7 º 49 ' 00 '' N – 81 º 46 ' 00 '' W, coral rubble, 3 m deep, 6 February 1997; 1 specimen (MNCN 16.01 / 1706), Granito de Oro Island, 7 º 35 ' 30 '' N – 81 º 42 ' 20 '' W, coral rubble, 4.5 m deep, 19 November 1997; 1 specimen (MNCN 16.01 / 11710), Uvas Island, coral rubble, 16 m deep, 7 February 1997; 1 specimen (MNCN 16.01 / 11725), Jicarita Island, Gulf of Chiriquí, 7 º 37 ' 50 '' N – 81 º 44 ' 30 '' W, 12 m deep, 9 February 1997; 1 specimen (MNCN 16.01 / 11742), Cirilo tip, 7 º 36 ' 10 '' N – 81 º 50 ' 10 '' W, 4 – 9 m deep, 9 February 1997. Branchiosyllis oculata, 1 Syntype (YPMNH 6745), Key West, Florida, USA. Caribbean Sea, Cuba: 12 specimens (1 mounted for SEM) (MNCN 16.01 / 752), Canal de los Vapores, Cayo Bocas de Alonso, Archipiélago de los Canarreos, 21 º 43 ’ 02 ” N – 82 º 28 ’ 50 ’’ W, sponges on Rhizophora mangle roots, 0.5 m deep, April 1984.	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFF1FFB3AA9AFF57FC74F891.taxon	description	Description. Longest complete specimen 3 mm long, 0.5 mm wide, 38 segments. Body strongly dorsoventrally flattened, pigmented on each segment with a dark central wide band. Prostomiun ovate, slightly wider than long, with two pairs of small eyes in trapezoidal arrangement. Antennae inserted on anterior margin of prostomium; median antennae shorter than combined length of prostomium and palps, with nine articles; lateral antennae longer, with 15 articles. Palps triangular, shorter than prostomium, fused at base. Nuchal organs as two ciliated pits on the lateral basis of prostomium. Peristomium dorsally reduced. Dorsal tentacular cirri longer than median antenna with 24 – 30 articles, ventral tentacular cirri shorter than dorsal, with about 10 – 12 articles. Dorsal cirri of anterior four chaetigers longer, with about 20 – 27 articles. Subsequent cirri with 11 – 15 articles. Antennae, tentacular cirri and dorsal cirri spindle-shaped. Parapodia bilobed, prechaetal lobe larger than postchaetal. Small, incipient branchiae, with minutes pores, dorsally on each parapodia, with granular appearance under light microscope (Fig. 1 A). Ventral cirri digitiform, almost as long as parapodial lobe, proximally inserted. Typically three to four compound chaetae per parapodium, smooth, unidentate ungulae, distinctly curved, all of them with ungulae rotated 180 º (Fig. 1 C, E). Ventral chaetae always larger than dorsal ones; all shafts of chaetae thick, ventral ones sligthly thicker than dorsal (Fig. 1 C). Two protruding, slender, straight aciculae, one of them slightly curved on posterior parapodia (Fig. 1 E). Pygidium with two anal cirri, 7 – 9 articles. Pharynx almost as wide as proventricle, through five segments with a small tooth, anteriorly located. Proventricle through eight segments, with 25 – 30 muscle cell rows.	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFF1FFB3AA9AFF57FC74F891.taxon	discussion	Remarks. Branchiosyllis oculata from the Caribbean differs from B. australis n. comb., from Australia and Philippines, in having large and very well developed branchiae (Fig. 1 B), while they are smaller in B. australis; less curved dorsal anterior ungulae (Fig. 1 D) and three parapodial lobes: prechaetal, postchaetal and ventral lobes (Fig. 1 F), instead of only one prechaetal lobe present in B. australis (Fig. 1 E). The Philipinnes specimens slightly differ from the Australian ones in having two aciculae (only one, thick in B. australis), one of the posterior ones slightly curved at the tip. In both populations, however, the aciculae protruded from parapodia (Fig. 1 C; San Martín et al. 2008, Figs 13 E, 15 A). Habitat. Shallow waters, on sponges, algae and coral rubble.	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFF1FFB3AA9AFF57FC74F891.taxon	distribution	Distribution. Philippines Islands and North Western Australia.	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFFFFFBCAA9AFF57FAE6FB12.taxon	materials_examined	Material examined. Holotype (MNHN 143), Djibouti, Gulf of Aden; Mediterranean Sea: 1 specimen (MNCN 16.01 / 6551), Cabo Blanco, west of Mallorca Island, Baleares Islands, 39 º 19 ’ 22 ’’ N – 2 º 59 ’ 01 ’’ E, 10 m deep, coll. Fauna III (229 B 7), 2 July 1994; Philippines Islands: 1 specimen (MNCN 16.01 / 14323), “ Popollcan ”, El Nido, Palawan Island, 11 º 11 ’ 50 ’’ N – 119 º 17 ’ 06 ’’ E, coral rubble, 3 m deep, “ South Intalula ”, El Nido, Palawan Island, 11 º 08 ' 01 " N – 119 º 19 ' 02 " E, sandy bottom, 5 m deep, 15 December 2010; 4 specimens (MNCN 16.01 / 14324), Twin Rocks, El Nido, Palawan Island, 11 º 15 ’ 24 ’’ N – 119 º 41 ’ 41 ’’ E, coral rubble, 6 m deep, 17 December 2010. West Australia: 4 specimens, (AM W 30095), Kimberley region, Bernouli Island, 15 ° S – 124 ° 47 ' E, sandy substrate with coral rubble, intertidal, 12 July 1988; 1 specimen (AM W 30099), SW corner of Lucas Island, 15 ° 13 ' S – 124 ° 31 ' E, dead coral substrate, 2 – 30 m, 24 July 1988; 10 specimens (AM W 30101), inshore reef off Neds Camp, Cape Range National Park, 21 ° 59 ' S – 113 ° 59 ' E, Caulerpa sp., 1 m deep, coll. J. K. Lowry, 2 January 1984; 1 specimen (AM W 33390), Monkey Mia, Shark Bay, 25 º 48 ’ 00 ’’ S – 113 º 45 ’ 41 ’’ E, unidentified sponges, 1 m deep, 17 June 2008; 1 specimen, Ningaloo St., 20, 22 º 50 ’ 40 ’’ S – 113 º 46 ’ 09 ’’ E, coral rubble, 5.2 m deep, 14 June 2008. West Panamá, Coiba National Park: 1 specimen (MNCN 16.01 / 11542), Islote Santa Cruz, 7 º 38 ' 00 '' N – 81 º 47 ' 20 '' W, 16 m deep, 7 February 1997; 1 specimen (MNCN 16.01 / 11583), Playa Rosario, 7 º 38 ' 30 '' N – 81 º 41 ' 40 '' W, coral rubble, 2.5 m deep, 16 November 1997; 1 specimen (MNCN 16.01 / 11584), Isla del Canal de Afuera, 7 º 41 ' 50 '' N – 81 º 38 ' 25 '', coral rubble, 4.5 m deep, 15 November 1997; 7 specimens (MNCN 16.01 / 11587), Isla de Uvas, 7 º 49 ' 00 '' N – 81 º 46 ' 00 ” W, coral rubble, 3 m deep, 6 February, 1997; 1 specimen (MNCN 16.01 / 11726), Playa Rosario, 7 º 38 ' 30 '' N – 81 º 41 ' 40 '' W, coral rubble, 1.2 m deep, 10 September 1998; 2 specimens (MNCN 16.01 / 11727), Isla de Granito de Oro, 7 º 37 ' 50 '' N – 81 º 44 ' 30 '' W, coral rubble, 2 m deep, 6 September 1997; 2 specimens (MNCN 16.01 / 11728), Isla Jicarita, 7 º 37 ' 50 '' N – 81 º 44 ' 30 '' W, 12 m deep, 9 February 1997; 1 specimen (MNCN 16.01 / 11733), Ensenada María, 7 º 24 ' 20 '' N – 81 º 41 ' 00 '' W, algae, 4 m deep, 23 June 1996; 1 specimen (MNCN 16.01 / 11734), Isla de Granito de Oro, 7 º 35 ' 30 '' N – 81 º 42 ' 20 '' W, algae, 3 m deep, 20 June 1996; 4 specimens (MNCN 16.01 / 11735), Isla de Uvas, 7 º 49 ' 00 '' N – 81 º 46 ' 00 '' W, coral rubble, 6 m deep, 6 February 1997; 1 specimen (MNCN 16.01 / 11736), Ensenada María, 7 º 24 ' 20 '' N – 81 º 41 ' 00 '' W, coral rubble, 4 m deep, 23 June 1996; 12 specimens (MNCN 16.01 / 11739), Isla de Uvas, 7 º 49 ' 00 '' N – 81 º 46 ' 00 '' W, coral rubble, 3 m deep, 6 February 1997; 1 specimen (MNCN 16.01 / 11740), Isla del Canal de Afuera, 7 º 41 ' 50 '' N – 81 º 38 ' 25 '' W, coral rubble, 6 m deep, 12 February 1997; 6 specimen (MNCN 16.01 / 11741), Ensenada María, 7 º 24 ' 20 '' N – 81 º 41 ' 00 '' W, coral rubble, 3 m deep, 14 February 1997; 1 specimen (MNCN 16.01 / 11743), Islote Santa Cruz, 7 º 38 ' 00 '' N – 81 º 47 ' 20 '' W, coral rubble, 14 m deep, 6 February, 1997. Caribbean Sea, Cuba: 10 specimens, (16.01 / 751) Cayo Matías, isla de Pinos, archipiélago de los Canarreos, 21 º 34 ’ 28 ’’ N – 82 º 26 ’ 01 ’’ W, Halimeda sp., 3 m deep, 1990. Comparative material. Syllis (Typosyllis) plessisi Rullier, 1972 (Branchiosyllis plessisi sensu Westheide, 1974), Holotype, (MNHN IRFA SYL Y 14) coll. Rullier, New Caledonia, Pins Island, coral, 15 m deep, 21 August 1961.	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFFFFFBCAA9AFF57FAE6FB12.taxon	description	Description. Body slender, cylindrical, variable in pigmentation (i. e. colourless, complety reddish to orange or with black lines in some parapodia). Prostomium rounded with four eyes in open trapezoidal arrangement. Median antenna longer than combined length of prostomium and palps, inserted between anterior eyes; lateral antennae inserted on anterior margin of prostomium, shorter than median antenna. Palps similar in length to prostomium. Dorsal tentacular cirri similar in length to median antenna, ventral ones shorter than dorsal tentacular cirri. Dorsal cirri similar in length or longer than total body width. Parapodial lobes distally bilobed. Compound bidentate chaetae on anterior and midbody parapodia; blades with spines on margin and shafts with distal spines. In posterior chaetigers, falcigers become unidentate with curved tips and distally curved shafts with small spines; most ventral chaetae present the ungulae rotated 180 º, with short, smooth shafts. Four anterior aciculae, three straight, pointed, and one distally curving; only two aciculae on posterior parapodia, one straight and the other one slightly oblique at tip, protruding from parapodia. Pharynx through about six to eight segments; pharyngeal tooth located anteriorly, surrounded by crown of ten soft papillae. Proventricle longer than pharynx, through nine segments. Pygidium small with two anal cirri similar in shape and length to dorsal cirri.	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFFFFFBCAA9AFF57FAE6FB12.taxon	discussion	Remarks. Although there are many small differences among the Branchiosyllis exilis specimens from different localities, all them have anterior compound falcigers and posterior ungulae but no other particular feature, such as branchiae (e. g., B. lamellifera, B. oculata, B. pacifica) or distinct ungulae in all parapodia (e. g., B. baringabooreen, B. carmenroldanae) (Table 1). All these small differences among the studied specimens, though not clear enough to be considered as diagnostic of different taxa may suggest that there are several sibling species traditionally named as Branchiosyllis exilis. Future molecular studies may be able to validate this hypothesis or, by contrast, confirm that it is a unique species with an intraspecific variability associated with different geographical regions. In the meantime, the small morphological differences found during the study of a large amount of material from diverse geographical areas allow us to tentatively join them into two groups. On the one hand, revised specimens from Cuba, Australia and Panamá have slender and large (longer than the total body width, 30 – 40 articles) cirri, and are similar with those reported from the Galapagos Islands by Westheide (1974) and from Indonesia by Aguado et al. (2008). On the other hand, specimens from the Mediterranean Sea and Philippines Islands have cirri with 15 – 25 articles (this paper, San Martín 2003). In addition, the anterior chaetael falcigers (i. e. on segments before proventricle) show a dorso-ventral gradation in falciger length in the specimens from the Mediterranean Sea, Cuba, Australia, and the holotype. These specimens also have bidentate blades, the dorsal ones with distal tooth equal or slightly larger than proximal one, and the ventral ones with a distal tooth much larger than the proximal one (in agreement with Aguado et al. 2008 for Indonesian material). The specimens from Panamá and Philippines Islands have bidentate falcigers similar in length, with distal and proximal teeth equal in size. Other features, such as the first segment with ungulae, the body length, and the colour pattern (often not well preserved) are highly variable and thus not useful to discriminate material (Table 1). The holotype of Syllis (Typosyllis) plessisi Rullier, 1972, from New Caledonia, considered as a synonym of B. exilis (Westheide 1974, Licher 1999; San Martín et al. 2008) has also been examined. Although the slide is in poor condition, the specimen agrees with Australian specimens and belongs to the group with large and slender dorsal cirri and chaetae with dorso-ventral gradation in falciger length. The specimens do not present the bidentate ungulae drawn by Rullier (1972) in anterior segments. Habitat. Shallow waters, on sponges, algae, gorgonians and coral rubble.	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFFFFFBCAA9AFF57FAE6FB12.taxon	distribution	Distribution. Apparently circumtropical, also present in the warmest waters of the Mediterranean.	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFFEFFBFAA9AFAFCFB74FA35.taxon	description	Fig. 2 A	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFFEFFBFAA9AFAFCFB74FA35.taxon	materials_examined	Material examined. Luzón Island, Balayan Bay: 3 specimens (MNCN 16.01 / 14325), " Sepok Wall ", between Balayan Bay and Batangas Bay, 13 º 41 ' 02 " N – 120 º 53 ' 45 " E, unidentified sponge and gogonians, 13 m deep, coral rubble, 6 m deep, 10 December 2010; 3 specimens (MNCN 16.01 / 14326), 13 º 44 ' 26 " N – 120 º 53 ' 34 " E, coral rubble, 2 – 3 m deep, 7 December 2010; 1 specimen (MNCN 16.01 / 14327), " Koala Point ", 13 º 47 ' 56 " N – 120 º 52 ' 10 " E, coral rubble, 1 – 2 m deep, 5 December 2010; 3 specimens (MNCN 16.01 / 14328), 13 º 44 ' 26 " N – 120 º 53 ' 34 " E, hidrozoans, gorgonians (Siphonogorgia sp.), ascidians (Atriolum sp.) and coral rubble, 2 – 4 m, 4 December 2010; 5 specimens (MNCN 16.01 / 14329), “ Sombrero Island ”, 13 º 41 ' 51 " N – 120 º 49 ' 45 " E, unidentified ramified sponges, 2 – 3 m deep, 6 December 2010; Palawan Island, El Nido: 1 specimen (MNCN 16.01 / 14330), “ Popollcan ”, 11 º 11 ' 50 " N – 119 º 17 ' 06 " E, coral rubble, 3 m deep, 15 December 2010; 1 specimen (MNCN 16.01 / 14331), “ South Intalula ”, 11 º 08 ' 01 " N – 119 º 19 ' 02 " E, sandy bottom, 5 m deep, 15 December 2010. Comparative material. Branchiosyllis maculata, 1 specimen (AM W 41617) Cassini, stn 59 / K 10, Q 3, 13 ° 56 ' 01 " S – 125 ° 37 ' 09 " E, mid-littoral reef platform, Flat reef pavement with an extensive cover of Sargassum sp. over a dusting of sediment, 0 m deep, Woodside Kimberley Survey 2010, 25 October 2010; 4 specimens (AM W 30108), inshore reef off Neds Camp, Cape Range National Park, 21 ° 59 ' S – 113 ° 59 ' E, frilly Caulerpa sp., 1 m deep, 2 January 1984; 2 specimens (AM W 30109) Bundegi Reef, Exmouth Gulf, 21 ° 49 ' S – 114 ° 11 ' E, rocky rubble with sediment, brown algae with epiphytic growth, intertidal, 4 January 1984; 3 specimens (AM W 29519), N end of beach, Bundegi Reef, 21 ° 49 ' S – 114 ° 11 ' E, rocky rubble & brown alga with epiphytic growth, sticky sediment, 2 m deep, 4 January 1984. Branchiosyllis cirropunctata, 2 specimens (AM W 30091), Goss Passage, Beacon Island, 28 ° 25 ' 30 " S – 113 ° 47 ' E, dead plates of Acropora sp. covered in coralline algae, 8 m deep, 19 May 1994; 1 specimen (AM W 30092), NE entrance to Goss Passage, Beacon Island, 28 ° 27 ' 54 " S – 113 ° 46 ' 42 " E, dead plate-like Acropora sp., covered in coralline algae, 8 m deep, 25 May 1994; 1 specimen (AM W 21993), Outer Koombana Bay, 33 ° 18 ' 46 " S – 115 ° 38 ' 58 " E, 9.4 m deep, 27 March 1993; 1 specimen (AM W 30094), Pondalowie and Marion Bays, Yorke Peninsula, 35 ° 14 ' S – 136 ° 50 ' E, Caulerpa sp. and green algae 3 m deep, 22 February 1985. Branchiosyllis exilis, 4 specimens (AM W 30095), Kimberley region, Bernouli Island, 15 ° S – 124 ° 47 ' E, sandy substrate with coral rubble, intertidal, 12 July 1988; 1 (AM W 30099), SW corner of Lucas Island, 15 ° 13 ' S – 124 ° 31 ' E, dead coral substrate, 2 – 30 m, 24 July 1988; 10 specimens (AM W 30101), inshore reef off Neds Camp, Cape Range National Park, 21 ° 59 ' S – 113 ° 59 ' E, Caulerpa sp., 1 m deep, coll. J. K. Lowry, 2 January 1984.	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFFEFFBFAA9AFAFCFB74FA35.taxon	description	Description. Body cylindrical, ventrally flattened, pigmentated with one dorsal spot or narrow line of black pigment on middle of each segment from midbody parapodia and black spots on numerous articles of dorsal cirri. Prostomium rounded with four eyes in open trapezoidal arrangement. Median antenna longer than combined length of prostomium and palps, inserted between anterior eyes; lateral antennae inserted on anterior margin of prostomium, shorter than median antenna. Dorsal tentacular cirri similar in length to median antenna, ventral ones shorter than dorsal tentacular cirri. Dorsal cirri of first chaetiger longer than other ones; midbody and posterior dorsal cirri alternating long and short. Parapodial lobes distally bilobed. Anterior and midbody parapodia with dorsal compound bidentate chaetae dorsoventrally decreasing in length, and ventral unidentate chaetae, all of them with marginal spines; midbody parapodia present also two to three unidentate chaetae without spines, short shafts and short blades, some of them rotated 180 º. In posterior parapodia these chaetae are more curved, becoming ungulae. Anterior parapodia each with three slender aciculae, two straight and pointed and one slightly oblique at tip; midbody and posterior segments with two to three aciculae protruding from parapodia, one or two straight and pointed, other slightly oblique at tip. Pharynx through about seven segments; pharyngeal tooth located anteriorly, surrounded by crown of ten soft papillae. Proventricle longer than pharynx, through 12 segments. Pygidium small, with 2 anal cirri similar in length to dorsal cirri. Some specimens with attached pink coloured acephalous stolon, small, short, with 6 – 8 chaetigers (Fig. 2 D).	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFFEFFBFAA9AFAFCFB74FA35.taxon	discussion	Remarks. The studied specimens agree well with the original description of B. maculata, with the Australian material and with those described from Indonesia (Aguado et al. 2008). Branchiosyllis cirropunctata is a similar species, but it possesses ungulae in its far posterior segments with subdistal spurs on the head of the shaft. Branchiosyllis exilis differs in having anterior bidentate falcigers with more spines on edge of blades and larger than those of B. maculata. The colour pattern is also a very distinctive character in well preserved specimens. Habitat. Intertidal to shallow depths, among coral rubble, algae and sponges.	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFFEFFBFAA9AFAFCFB74FA35.taxon	distribution	Distribution. West Pacific Ocean (Southern Japan, Philippines, Australia), Indonesia.	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFFDFFA4AA9AF9D4FDBDFCC2.taxon	description	Figs 2 B, 3 A – G, 4 A – H	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFFDFFA4AA9AF9D4FDBDFCC2.taxon	materials_examined	Material examined. HOLOTYPE (NMA 4434) and 23 paratypes fixed in ethanol (NMA 4435, MNCN 16.01 / 13182) and 708 in formaldehyde (4 mounted for SEM) (NMA 4436, MNCN 16.01 / 13183). Philippines Islands: " Mainif point ", between Balayan Bay and Batangas bay, Luzon Island, 13 º 40 ’ 48 ’’ N – 120 º 51 ’ 20 ’’ E, unidentified orange sponge, 20 m deep, 8 December 2010. Comparative material: Syllis (Typosyllis) plessisi Rullier, 1972 (Branchiosyllis plessisi Westheide, 1974), Holotype (MNHN IRFA SYL Y 14) coll. Rullier, New Caledonia, Pins Island, coral, 15 m deep, 21 August 1961.	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFFDFFA4AA9AF9D4FDBDFCC2.taxon	description	Description. Holotype 1.42 mm long, 0.25 mm wide, 21 segments. Large paratype 1.5 mm long, 0.2 mm wide, 26 segments. Body cylindrical dorsally (Fig. 4 A), orange with black stripes in peristomiun and anterior four to seven segments more pigmented at bases of cirrophores (Figs 2 B, 3 A). Prostomium ovate, with a less pigmented black band, joining the three antennae; four small and red eyes in open trapezoidal arrangement (Figs 2 B, 3 A). Three antennae similar in length slightly longer than combined length of prostomium and palps, with eight articles (Fig. 4 B). Median antenna inserted on middle of prostomium; lateral antennae inserted almost between anterior eyes. Palps triangular, fused at base, slightly shorter than prostomium. Nuchal organs as two ciliated pits on lateral base of prostomium (Fig. 4 B). Peristomiun shorter than subsequent segments. Dorsal tentacular cirri longer than median antenna with 11 – 15 articles, ventral tentacular cirri much shorter than dorsal ones, with 6 articles. Dorsal cirri from one to six chaetigers similar in length with about nine articles; subsequent dorsal cirri shorter, with five or six articles. Dorsal cirri of most posterior segments with three articles. Most distal article of antennae, tentacular cirri and dorsal cirri spindle-shaped. Parapodia blunt, without branchiae. Ventral cirri thick, digitiform, slightly shorter than parapodial lobes, proximally inserted. Typically two compound ungulae per parapodium (occasionally 3) (Fig. 4 E), bidentate (with distal teeth longer than proximal one) (Fig. 4 F, G). Compound chaetae in first four segments occasionally with blades rotated 180 º (Figs 3 D, 4 F); in medium parapodia most dorsal one bidentate, while ventral unidentate, most of them rotated 180 º; unidentate chaetae more curved, shorter and slightly thicker than dorsal one (Fig. 3 E). Posterior segments with two unidentate ungulae with blades always rotated 180 º, ventral one slightly bigger and more curved (Figs 3 F, 4 H). Single slender aciculae per parapodia with slightly oblique tip (Fig. 3 G). Pygidium small, with two anal cirri shorter than dorsal cirri, lost in most specimens (Fig. 4 C). Pharynx almost as wide as proventricle, through five segments; pharyngeal tooth small, anteriorly located, surrounded by crown of 12 soft papillae (Figs 3 B, 4 D). Proventricle through three segments (Fig. 3 A), with 30 – 36 muscle cell rows.	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFFDFFA4AA9AF9D4FDBDFCC2.taxon	discussion	Remarks. Branchiosyllis mayae n. sp., differs from any other species of the genus in the presence of bidentate ungulae, its short dorsal cirri, and the striking colour pattern (Table 1). Branchiosyllis abranchiata Hartmann- Schröder, 1965 (synonymized by Westheide 1974, Licher 1999 and San Martín et al., 2008 with B. exilis) has a similar small size but it has the typical anterior bidentate falcigers of the B. exilis species (see previous discussion) instead of bidentate ungulae. Although bidentate ungulae were illustrated in the original description of B. plessisi, the holotype of this species does not present this kind of chaetae and it is considered a synomy of B. exilis (see previous Discussion).	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFFDFFA4AA9AF9D4FDBDFCC2.taxon	etymology	Etymology. The specific name refers to the cartoons “ Maya the Bee ” due to the colour pattern of the species. Habitat. Inside orange, unidentified sponge, reaching high densities in a single, relatively small host individual.	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFFDFFA4AA9AF9D4FDBDFCC2.taxon	distribution	Distribution. Philippines Islands. Branchiosyllis tagalog n. sp. Figs 2 C – D, 5 A – F, 6 A – G	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFFDFFA4AA9AF9D4FDBDFCC2.taxon	materials_examined	Material examined. Holotype (NMA 4437) and 2 paratypes fixed in ethanol, 1 mounted for SEM (MNCN 16.01 / 11625, 11626). Philippines Islands: Sombrero Island, Balayan Bay, Luzon Island, 13 º 41 ’ 52 ’’ N – 120 º 49 ’ 47 ’’ E, three different unidentified sponges, 2 – 8 m deep, coral rubble, 17 m deep, 6 December 2010. Comparative material. Branchiosyllis baringabooreen, Holotype (AM W 30088) Western Australia: Bernouli Island, 15 ° 00 ' S 124 ° 47 ' E, sandy substrate with coral rubble, intertidal, coll. P. A. Hutchings, 12 July 1988. Paratypes: 1 specimen (AM W 26511) W side of Cassini Island, 13 ° 57 ' S – 125 ° 37 ' E, coralline algae and rubble, low tide, 18 July 1988; 1 specimen (AM W 30089) Bernouli Island, 15 ° 00 ' S – 124 ° 47 ' E, 12 July 1988; 2 specimens (AM W 30090) Reef S of Lucas Island, Brunswick Bay, 15 ° 16 ' S – 124 ° 29 ' E, dead coral with Sargassum with heavy silt loading, 2 m deep, 24 July 1988. One specimen mounted for SEM (AM W 41524), Long Reef, Stn 52 / K 10, Trans T 1, 13 ° 48 ' 36 " S – 125 ° 49 ' 27 " E, mid-littoral reef terrace, honeycombed reef flat with coral rubble and dead coral slabs, 0 m deep, Woodside Kimberley Survey, 23 October 2010; 1 specimen (AM W 41623), Queensland, Watson’s Bay, Lizard Island, 14 º 65 ’ 82 ” S – 145 º 44 ’ 9 ” E, 9 m deep, 30 August 2010.	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFFDFFA4AA9AF9D4FDBDFCC2.taxon	description	Description. Longest complete specimen 8 mm long, 0.72 mm wide, 67 segments (Fig. 2 B). Body strongly dorsoventrally flattened, pigmented on each segment with dark central spot surrounded by two horizontal wide bands and less dense coloured circle on each parapodia. Cirri also with irregular traces of colour (Figs 2 C, 5 A). Prostomiun ovate, slightly wider than long, with two pairs of small red eyes on trapezoidal arragment (Figs 2 C, 5 A). Antennae inserted on anterior margin of prostomium; median antennae shorter than combined length of prostomium and palps, with 10 – 12 articles; lateral antennae longer, with 15 – 17 articles. Palps triangular, shorter than prostomium, fused at base. Peristomiun dorsally reduced (Figs 5 A, 6 A). Nuchal organs not observed. Dorsal tentacular cirri much longer than median antenna with 52 – 54 articles, ventral tentacular cirri shorter than dorsal, with about 18 – 20 articles. Dorsal cirri of all chaetigers similar in length with about 55 – 60 articles, shorter than body width. Tentacular cirri and dorsal cirri spindle-shaped with minute pores irregularly distributed and small papillae organized in lines (Fig. 6 B, C). Parapodia with small post chaetal lobe and without branchiae (Fig. 5 C). Ventral cirri digitiform, as long as parapodial lobe, proximally inserted (Fig. 5 C). Typically two ungulae per parapodium, smooth, unidentate, most with blades rotated 180 º (Figs 5 D, E, 6 D – F). Ventral ungulae always larger than dorsal; shafts of chaetae thick, with small serration on distal part, ventral shafts thicker than dorsal (Fig. 6 E, F). Two slender and straight aciculae per parapodia, different in size (Fig. 5 F). Pygidium only observed in regenerating specimens, without anal cirri. Pharynx almost as wide as proventricle, through 5 segments; pharyngeal tooth small, anteriorly located surrounded by crown of 9 – 10 soft and small papillae. Proventricle through 6 segments, with 36 – 40 muscle cell rows (Fig. 5 B). Some specimens with acephalous stolon, pigmented with dark central spot surrounded by two horizontal lines and one red spot on each parapodia (Figs 2 D, 6 G).	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFFDFFA4AA9AF9D4FDBDFCC2.taxon	discussion	Remarks. The most similar species is B. baringabooreen with similar ungulae and body pigmentation. However, Branchiosyllis tagalog n. sp., differs in having only two curved ungulae per parapodia, ventral one larger than dorsal (Fig. 4 H), longer dorsal cirri with small pores and papillae organized in lines (Figs 6 B, C).	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFFDFFA4AA9AF9D4FDBDFCC2.taxon	etymology	Etymology. The name refers to “ Tagalog ”, the Austronesian language spoken in the Philippines Islands. Habitat. Shallow waters, on sponges and coral rubble.	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
4E6787A6FFFDFFA4AA9AF9D4FDBDFCC2.taxon	distribution	Distribution. Philippines Islands.	en	Álvarez-Campos, Patricia, Martín, Guillermo San, Aguado, M. Teresa (2012): The genus Branchiosyllis Ehlers, 1887 from Philippines Islands, with the description of two new species. Zootaxa 3542: 49-68
