identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
4E4F69489A5CFFA85DD4F974A0A2FB88.text	4E4F69489A5CFFA85DD4F974A0A2FB88.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macutula	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Macutula gen. nov.</p>
            <p> Type species:  Macutula aracoiaba sp. nov.</p>
            <p>Etymology. The name is an arbitrary combination of letters to be treated as feminine.</p>
            <p> Phylogenetic relationship. Although DNA data consistently support the monophyly of amycoids (e.g., Maddison &amp; Hedin 2003; Maddison et al. 2008), morphological synapomorphies remain unknown. The new genus is supposed to belong to Amycoida by similarities with several amycoid lineages. In general appearance, spiders of this group resemble those of  Breda Peckham and Peckham , which could be an amycoid according to unpublished DNA data, by the dark, glabrous carapace. The third leg longest is also common in amycines, such as  Mago O.P.- Cambridge and  Noegus Simon. Cheliceral teeth are similar to the ones present in  Breda and in the  Hurius group, with several teeth on promargin and one on retromargin (see Chickering 1946, fig. 16; Galiano 1985, figs 4, 18). The reduced sperm loop immediately before the embolus is similar to that present in species of  Sitticus Simon. The epigynal structure is also similar to that of many amycoids: the anterior atrium is found in  Breda and many amycines, such as  Mago and  Noegus . Moreover, copulation ducts coiled posteriorly and anterior spermathecae are also found in almost all groups of amycoids. </p>
            <p> Diagnosis.  Macutula gen. nov. differs from the remaining genera of Amycoida by having a very deep prolateral excavation on the dorsal surface of the cymbium of the male palp, forming a dorsal keel (Fig. 7). </p>
            <p>Description. Median sized jumping spiders (4–7mm) with slightly flattened bodies and dark brown, glabrous carapaces (Figs 1–2); chelicera similar in males and females, with an anterior color pattern (Fig. 11), one big tooth on retromargin and five on promargin, the proximal two being the largest (Fig. 12); male endite similar to that of the female (Fig. 13); labium rounded with posterior ventral excavations (Fig. 13) and sternum subpentagonal (Fig. 14); male palp with a straight femur (Fig. 6), short tibia with a curved RTA and a poorly developed RvTA (Figs 3– 5, 24, 26), a triangular cymbium seen in anterior view, with a deep prolateral-dorsal excavation (Fig. 7), small subtegulum and tegulum with a small loop of the sperm duct just before it enters the embolus (Figs 3, 8) and a long curling embolus, whose tip lies in a ventral excavation on the distal cymbium (Figs 3, 23, 25). Leg formula 1342 in males (Figs 15–18) and 3412 in females (Figs 19–22). Spination: male: palpal femur 0; femur I d1-1-1 p1di, II d1- 1-1, p1di, III d1-1-1 p2di, IV d1-1-1; patella I=II 0, III=IV r1; tibia I v2-2 -2, II v1 r-2-2, p1-0-1, III v1 p-0-1r, p1-1, r1-1-1, IV v1 p-0-1r, p1-0, r1-1-1; metatarsus I=II v2-2, III v2-2, p1-2, r1-2, IV v1 p-1r, p0-2, r1-1-2; female: femur I=II d1-1-1, p1di, III d1-1-1 p2di, IV d1-1-1; patella I-IV 0; tibia I v2-2 -2, II v1 r-2-2, III v1 p-0-1r, p0-1, r1-1-1, IV v1 p-0-0, p0, r1-1; metatarsus I=II v2-2, III v2-2, p1-2, r1-2, IV v1 p-1r, p0-2, r1-2. Abdomen about the same length as the carapace with longitudinal dorsal color patterns (Figs 1–2). Epigynum with an anterior atrium (Figs 9, 27), a membranous, spiral initial portion of the copulation duct coiling backwards around the second, sclerotized portion, which coils forward and becomes loose, reaching the anterior spermatheca (Figs 10, 28).</p>
            <p> Composition. Three species:  M. aracoiaba sp. nov. ,  M. santana sp. nov. ,  M. caruaru sp. nov.</p>
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	https://treatment.plazi.org/id/4E4F69489A5CFFA85DD4F974A0A2FB88	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ruiz, Gustavo Rodrigo Sanches	Ruiz, Gustavo Rodrigo Sanches (2011): Description of Macutula, a new genus of jumping spiders from Northeastern Brazil (Araneae: Salticidae: Amycoida). Zootaxa 2785: 53-60, DOI: 10.5281/zenodo.203341
4E4F69489A5FFFAC5DD4FA93A0EEFB66.text	4E4F69489A5FFFAC5DD4FA93A0EEFB66.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macutula aracoiaba	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Macutula aracoiaba sp. nov.</p>
            <p>Figs 1 –22</p>
            <p> Type material: Holotype: male from Campo de Instrução Marechal Newton Cavalcante,  Araçoiaba , Pernambuco, Brazil, coordinates -7.781944, -35.150556, 2009–2010, A.A. Costa (IBSP 160499). Paratypes: 1 female, same data as holotype (IBSP 160500), and 1 male and 1 female, same data as holotype (UFPE). </p>
            <p> Additional material examined. BRAZIL: Pernambuco: 5 males and 11 females,  Araçoiaba , Campo de Instrução Marechal Newton Cavalcante, 2009–2010, A.A. Costa (IBSP 160498, 160501 – 160506, 160740 – 160745, UFPE). </p>
            <p>Etymology. The specific epithet is a toponymic name in apposition.</p>
            <p> Diagnosis. Males of this species differ from those of the other two by having a curling embolus slightly longer than 2.25 turns around the bulb in the male palp, and having the sperm duct loop on the proximal, prolateral region of the tegulum (Fig. 3). Females can be distinguished from those of  M. caruaru sp. nov. by the longer copulation duct, best noticed at the posteriormost region, where the membranous portion becomes the sclerotized portion (Fig. 10). </p>
            <p>Description. Male holotype: Total length: 4.40. Carapace dark brown, glabrous, 2.20 long, 1.60 wide, 1.10 high (Fig. 1). Ocular quadrangle 1.12 long. Anterior eye row 1.50 wide and posterior 1.40 wide. Chelicera, endite, labium and sternum dark brown. Palp dark brown. Legs I and II dark brown, with proximal half of metatarsi and entire tarsi yellow (Figs 15–16), III with coxa and trochanter light brown, femur dark brown with a median dorsal light area, patella and tibia dark brown, metatarsus yellow with proximal and distal portions dark brown and tarsus yellow (Fig. 17), IV with coxa and trochanter light brown, femur and patella dark brown, tibia yellow with proximal and distal portions dark brown, metatarsus yellow with distal dark brown portion and yellow tarsus (Fig. 18). Length of femur: I 1.40, II 1.10, III 1.45, IV 1.25; patella + tibia: I 1.90, II 1.30, III 1.40, IV 1.32; metatarsus + tarsus: I 1.30, II 0.95, III 1.15, IV 1.25. Abdomen dorsally dark brown, variegated, with anterior white border and a median, longitudinal light brown stripe between a pair of longitudinal narrow stripes of white scales (Fig. 1); ventrally dark brown, variegated. Spinnerets dark brown.</p>
            <p> FIGURES 15–18.  Macutula aracoiaba sp. nov. 15– 18 male legs, prolateral (15 I, 16 II, 17 III, 18 IV). </p>
            <p>Female (paratype IBSP 160500): Total length: 5.55. Carapace dark brown, glabrous, 2.35 long, 1.77 wide, 1.10 high (Fig. 2). Ocular quadrangle 1.30 long. Anterior eye row 1.62 wide and posterior 1.57 wide. Chelicera, endite, labium and sternum dark brown. Palp light brown. Leg coxae and trochanters light brown; Legs I and II yellow, with femora, distal portions of patellae, tibiae and metatarsi and proximal portion of tibiae dark (Figs 19–20); III and IV yellow, with femora and distal portion of patellae dark brown (Figs 21–22). Length of femur: I 1.25, II 1.15, III 1.55, IV 1.35; patella + tibia: I 1.65, II 1.35, III 1.50, IV 1.45; metatarsus + tarsus: I 1.15, II 1.00, III 1.22, IV 1.40. Abdomen dorsally dark brown, variegated, with white anterior border and two pairs of lateral light spots, light longitudinal dorsal stripe interrupted at the middle and a pair of narrow, paramedian stripes (Fig. 2); ventrally dark brown. Spinnerets dark brown.</p>
            <p> Variation. Some females examined have an elliptical atrium on the epigyne, similar to that of the paratype of  M. caruaru sp. nov. (Fig. 27), with no posterior prolongation of the atrium as that of the paratype of  M. aracoiaba sp. nov. described (Fig. 9). </p>
            <p>Distribution. Known only from the State of Pernambuco, Brazil.</p>
            <p>Biological note. The specimens mentioned above were collected from the ground using a pitfall trap.</p>
            <p> FIGURES 19–22.  Macutula aracoiaba sp. nov. 19– 22 female legs, prolateral (19 I, 20 II, 21 III, 22 IV). </p>
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	https://treatment.plazi.org/id/4E4F69489A5FFFAC5DD4FA93A0EEFB66	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ruiz, Gustavo Rodrigo Sanches	Ruiz, Gustavo Rodrigo Sanches (2011): Description of Macutula, a new genus of jumping spiders from Northeastern Brazil (Araneae: Salticidae: Amycoida). Zootaxa 2785: 53-60, DOI: 10.5281/zenodo.203341
4E4F69489A59FFAC5DD4FAB0A177F875.text	4E4F69489A59FFAC5DD4FAB0A177F875.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macutula santana	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Macutula santana sp. nov.</p>
            <p>Figs 23–24</p>
            <p> Type material. Holotype: male from Serra de São José, Feira de  Santana , Bahia, Brazil, coordinates -12.133333, - 39.016667, 2009–2010, G. da S.C. Ferreira (IBSP 160507). Paratype: 1 male, same data as holotype (IBSP 160514). </p>
            <p>Etymology. The specific epithet is a toponymic name in apposition.</p>
            <p>Diagnosis. Males of this species differ from those of the other two by having a curling embolus slightly longer than 1.5 turns around the bulb, and having the sperm duct loop on the distal, retrolateral region of the tegulum (Fig. 23).</p>
            <p> Description. Male holotype: Total length: 5.10. Coloration as in  M. aracoiaba sp. nov. , except for legs III and IV. Carapace 2.30 long, 1.70 wide, 1.15 high. Ocular quadrangle 1.20 long. Anterior eye row 1.65 wide and posterior 1.60 wide. Leg III with coxa and trochanter light brown, femur, patella, tibia and distal half of metatarsus dark brown, proximal half of metatarsus and tarsus yellow; IV as in III, except for the light brown median third of the tibia. Length of femur: I 1.40, II 1.15, III 1.40, IV 1.30; patella + tibia: I 1.85, II 1.35, III 1.50, IV 1.45; metatarsus + tarsus: I 1.20, II 1.00, III 1.25, IV 1.40. </p>
            <p>Female: Unknown.</p>
            <p>Distribution. Known only from the State of Bahia, Brazil.</p>
            <p>Biological note. The specimens mentioned above were collected from the ground using a pitfall trap.</p>
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	https://treatment.plazi.org/id/4E4F69489A59FFAC5DD4FAB0A177F875	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ruiz, Gustavo Rodrigo Sanches	Ruiz, Gustavo Rodrigo Sanches (2011): Description of Macutula, a new genus of jumping spiders from Northeastern Brazil (Araneae: Salticidae: Amycoida). Zootaxa 2785: 53-60, DOI: 10.5281/zenodo.203341
4E4F69489A5AFFAE5DD4F9F2A177FC3C.text	4E4F69489A5AFFAE5DD4F9F2A177FC3C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macutula caruaru	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Macutula caruaru sp. nov.</p>
            <p>Figs 25–28</p>
            <p> Type material. Holotype: male from  Caruaru , Pernambuco, Brazil, coordinates -8.266667, -35.966667, 2009– 2010, H.P. Amorim (IBSP 160508). Paratypes: 1 female, same data as holotype (IBSP 160509), and 1 male and 1 female, same data as holotype (UFPE). </p>
            <p> Additional material examined. BRAZIL: Pernambuco: 5 males,  Caruaru, 2009 –2010, H.P. Amorim (IBSP 160510 – 160513, UFPE). </p>
            <p>Etymology. The specific epithet is a toponymic name in apposition.</p>
            <p> Diagnosis. Males of this species differ from those of the other two by having the embolus curling almost two turns around the bulb, and having the sperm duct loop at the center of the distal region of the tegulum (Fig. 25). Females can be distinguished from those of  M. aracoiaba sp. nov. by the shorter copulation duct, best noticed at the posteriormost region, where the membranous portion becomes the sclerotized portion (Fig. 28). </p>
            <p>Description. Male holotype: Total length: 6.30. Carapace dark brown, glabrous, 3.05 long, 2.35 wide, 1.60 high. Ocular quadrangle 1.65 long. Anterior eye row 2.00 wide and posterior 1.90 wide. Chelicera, endite, labium and sternum dark brown. Palp dark brown. Legs I and II dark brown with yellow tarsi, III and IV with yellow coxae and trochanters, dark brown femora, light brown patellae and tibiae and yellow metatarsi and tarsi. Length of femur: I 2.30, II 1.85, III 2.20, IV 1.90; patella + tibia: I 3.40, II 2.30, III 2.35, IV 2.00; metatarsus + tarsus: I 2.20, II 1.55, III 1.85, IV 1.90. Abdomen light brown with white anterior border and a dorsal longitudinal lighter stripe; ventrally light brown. Spinnerets dark brown.</p>
            <p>Female (paratype IBSP 160509): Total length: 6.40. Carapace dark brown, glabrous, 2.90 long, 2.17 wide, 1.40 high, with a light brown area around the ocular quadrangle. Ocular quadrangle 1.55 long. Anterior eye row 1.90 wide and posterior 1.85 wide. Chelicera, endite and labium dark brown, sternum light brown. Palp light brown. Legs I and II yellow, with distal portions of femora, patellae, tibiae and metatarsi dark brown, as well as the proximal portion of tibiae, III and IV yellow with distal portions of femora, patellae, tibiae and metatarsi dark brown, as well as the proximal portion of femora and tibiae. Length of femur: I 1.60, II 1.50, III 1.90, IV 1.70; patella + tibia: I 2.15, II 1.80, III 1.95, IV 1.85; metatarsus + tarsus: I 1.40, II 1.25, III 1.55, IV 1.75. Abdomen dorsally variegated with dark brown and pale yellow, anterior border white and a pair of longitudinal sinuous light lines that fuse at the middle and again twice in the posterior half of the abdomen, being completely fused posteriorly; ventrally light brown. Spinnerets dark brown.</p>
            <p> Variation. Although the female paratype described has an elliptical atrium on the epigyne (Fig. 27), the other female paratype has a posterior prolongation of the atrium similar to that of the paratype of  M. aracoiaba sp. nov. (Fig. 9). </p>
            <p>Distribution. Known only from the State of Pernambuco, Brazil.</p>
            <p>Biological note. The specimens mentioned above were collected from the ground using a pitfall trap.</p>
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	https://treatment.plazi.org/id/4E4F69489A5AFFAE5DD4F9F2A177FC3C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ruiz, Gustavo Rodrigo Sanches	Ruiz, Gustavo Rodrigo Sanches (2011): Description of Macutula, a new genus of jumping spiders from Northeastern Brazil (Araneae: Salticidae: Amycoida). Zootaxa 2785: 53-60, DOI: 10.5281/zenodo.203341
