taxonID	type	description	language	source
4F645F78244BFFCFFF4021F572C5D600.taxon	materials_examined	Type: — PANAMÁ. Provincia de Veraguas: Parque Nac. Coiba, Las Salinas, Sur del sendero de los Pozos, 5 m, 7 ˚ 26 ’ 07.6 ” N, 81 ˚ 43 ’ 42.9 ” W, 20 April, 2008, fl., C. Galdames 6098, C. Vasquez, A. Ibáñez & F. Hernández (holotype, PMA!). Shrubs ca. 1.5 – 5 m or small trees to 10 m; bark reddish or brown-reddish; branching dichotomous, with 4 to 6 leaves per branchlet; branches glabrous or with a few hairs; branchlets glabrous to pubescent, compressed at nodes. Leaf blades ovate, narrowly ovate, elliptic, broadly elliptic, or rarely rhombic, 3 – 6 cm × 1.5 – 3.6 cm, 1.5 – 2.5 times as long as wide; base rounded, acute to broadly acute, margins revolute, pubescent or sometimes glabrous; apex acute to broadly acuminate, sometimes rounded; chartaceous or subcoriaceous, when fresh dark green above and light green below, new leaves reddish; drying slightly discolorous, dull greyish to brown-greyish above, brown-greenish, brown to brown-greyish below; the midvein brown with a reddish tint, pubescent or sometimes glabrous, flat or depressed adaxially, convex abaxially, distally attenuating; lateral veins 5 to 8 per side, glabrous or sometimes with a few scattered hairs, straight and ascending, when dry conspicuous on both surfaces (more numerous in young leaves), sometimes inconspicuous above; marginal veins 1 – 2 mm (3 mm in new leaves) separated from the margin, glands numerous on both surfaces, sometimes inconspicuous above; petioles 1 – 3 mm, terete, pubescent or sometimes glabrous, with striate to lightly striate epidermis. Inflorescences axillary, shortly racemose to racemose, solitary; axis 0.5 – 5 mm; flowers 2 to 8; bracts 0.5 – 1.3 × 0.5 – 1 mm, ovate, persistent, margin usually ciliate and scarious, sometimes glabrous and scarious or glabrous; buds 1 – 1.5 mm, globose or ovoid. Flowers pedicellate, pedicels 6 – 16 × 0.5 mm, markedly glandular, glabrous or sometimes pubescent; bracteoles 0.5 – 1 × 0.5 – 0.7 mm, persistent, pubescent like the bracts, ovate, glandular, free in the base; hypanthium ca. 1 mm long, campanulate, prolongated 0.1 – 0.2 mm long above summit of ovary, like a neck, this separates the developing fruit from the calyx; calyx lobes four, membranaceous, with scarious margin, concave, in unequal pairs, the smaller 1.1 – 1.2 × 0.8 – 1 mm, the larger 1.6 – 1.8 × 1.1 – 1.2 mm, broadly elliptic, glandular, the apex cuspidate; petals four, 4 – 5 × 2 – 2.8 mm, ovate or ovoid with rounded apex, disk ca. 1.2 mm in diameter; staminal ring glabrous; stamens 20 to 30, 3 – 4 mm long; style ca. 3.5 mm long, glabrous. Ovary bilocular, with 4 to 8 ovules per locule. Fruits 8 – 10 × 8 – 10 mm, globose, black when mature; pericarp thin-walled (0.5 – 0.8 mm when dry) with glands, glabrous; calyx persistent. Seed one, smooth. Eponymy: — The epithet of this species refers to the Panamanian Province of Veraguas.	en	Flores, Rodolfo, Ibáñez, Alicia, Correa, Mireya D. (2016): Eugenia veraguensis (Myrtaceae), a new species from Golfo de Chiriquí in Veraguas Province, Panama, with notes on Eugenia rhombea. Phytotaxa 270 (3): 217-222, DOI: 10.11646/phytotaxa.270.3.6, URL: http://dx.doi.org/10.11646/phytotaxa.270.3.6
4F645F78244BFFCFFF4021F572C5D600.taxon	distribution	Distribution: — Islands of Western Pacific Panama. Coiba National Park, Bahía Honda region, Cébaco Island. Habitat and Ecology: — E. veraguensis is a shrub or small tree that grows in the understory of coastal scrub and forests on drier areas of Western Pacific Panama (Golfo de Chiriquí). Phenology: — Collected in flower in April; collected in fruit March-June. Conservation status: — Eugenia veraguensis is reported in eight locations, five of which are in protected areas, and therefore we do not consider there to be any threat to the largest populations of the species. For this reason, we believe the category Least Concern (LC) fits the IUCN Red List categories and criteria (IUCN 2012).	en	Flores, Rodolfo, Ibáñez, Alicia, Correa, Mireya D. (2016): Eugenia veraguensis (Myrtaceae), a new species from Golfo de Chiriquí in Veraguas Province, Panama, with notes on Eugenia rhombea. Phytotaxa 270 (3): 217-222, DOI: 10.11646/phytotaxa.270.3.6, URL: http://dx.doi.org/10.11646/phytotaxa.270.3.6
4F645F78244BFFCFFF4021F572C5D600.taxon	discussion	Discussion: — Eugenia veraguensis resembles Eugenia rhombea morphologically as well as in its mainly coastal habitat, but is distinguished from it by the characters listed in the diagnosis. The specimens of E. veraguensis that we studied comprised two groups, one with pubescence on branchlets, petioles, veins, margins, bracts and pedicels, while the other group is nearly glabrous. Since the specimen Castroviejo 14702 SC previously supported the occurrence of E. rhombea in Panama (Landrum et al. 2009) and this specimen is here considered as a paratype of the new E. veraguensis, we conclude that E. rhombea does not occur in Panama. The number of Eugenia species currently known for Panama thus remains as 38. Paratypes: — PANAMÁ. Provincia de Veraguas. P. N. Coiba. Brincanco. Norte de la Isla, 1 m, 7 ˚ 51 ’ 59.05 ” N, 81 ˚ 47 ’ 45.03 ” W, 29 May 2004, fr., A. Ibáñez & al. 3133 AI (PMA!, MO, F, SCZ, UCH); P. N. Coiba. Las Salinas, 1 m, 7 ˚ 26 ’ 6.94 ” N, 81 ˚ 43 ’ 22.61 ” W, 20 April 2008, fr., A. Ibáñez & F. Hernández 5649 AI (PMA!, MO, F, SCZ, UCH); P. N. Coiba. Manila, 5 m, 7 ˚ 22 ’ 34.45 ” N, 81 ˚ 47 ’ 6.51 ” W, 24 March 1998, fr., S. Castroviejo & al. 14702 SC (PMA!, MO, F, MA, SEL); P. N. Coiba. Río Barco Quebrado, 20 m, 7 ˚ 22 ’ 57.87 ” N, 81 ˚ 42 ’ 6.82 ” W, 21 October 2005, sterile, A. Ibáñez & C. Núñez 4453 AI (PMA!); P. N. Coiba, W de Playa Blanca, 20 m, 7 ˚ 22 ’ 59.09 ” N, 81 ˚ 40 ’ 36.08 ” W, 7 December 2005, sterile, A. Ibáñez & C. Núñez 4484 AI (PMA!); P. N. Coiba. Jicarita. Norte de la Isla, 5 m, 7 ˚ 13 ’ 19.27 ” N, 81 ˚ 48 ’ 18.74 ” W, 29 June 2004, sterile, A. Ibáñez & al. 3509 AI (PMA!, SCZ); P. N. Coiba. Río Colorado, 40 m, 7 ˚ 19 ’ 58.80 ” N, 81 ˚ 39 ’ 43.42 ” W, 5 December 2005, sterile, A. Ibáñez & C. Núñez 4528 AI (PMA!); P. N. Coiba. Contreras, Isla Escorpión, 7 ˚ 50 ’ 27.90 ” N, 81 ˚ 45 ’ 14.78 ” W, 11 January 2012, sterile, J. Carrión & al. 526 JFC (PMA!); P. N. Coiba. Islote al SW de Brincanco, 7 ˚ 51 ’ 13.45 ” N, 81 ˚ 47 ’ 51.53 ” W, 13 March 2011, sterile, N. León & al. 379 NL (PMA!); P. N. Coiba. Uva, 5 m, 7 ˚ 48 ’ 52.38 ” N, 81 ˚ 45 ’ 24.11 ” W, 2 june 2004, sterile, A. Ibáñez & al. 3268 AI (PMA!); P. N. Coiba. Río Negro, 20 m, 7 ˚ 23 ’ 10.24 ” N, 81 ˚ 44 ’ 48.21 ” W, 21 March 2006, sterile, A. Ibáñez & al. 4846 AI (PMA!); P. N. Coiba. Brincanco. Costa N de la isla, 1 m, 7 ˚ 51 ’ 51.99 ” N, 81 ˚ 47 ’ 26.14 ” W, 31 May 2004, fr., O. Rodríguez & al. 251 JR (PMA!); P. N. Coiba. Jicarón. Costa SE de la Isla, 1 m, 7 ˚ 14 ’ 44.26 ” N, 81 ˚ 47 ’ 56.69 ” W, 3 July 2004, sterile, O. Rodríguez & al. 398 JR (PMA!); P. N. Coiba. Punta Anegada, 1 m, 7 ˚ 20 ’ 25.76 ” N, 81 ˚ 36 ’ 14.01 ” W, 21 October 2004, sterile, A. Ibáñez & al. 4052 AI (PMA!); P. N. Coiba. Uva. De N al S de la isla, 10 m, 7 ˚ 48 ’ 42.28 ” N, 81 ˚ 45 ’ 27.68 ” W, 3 June 2004, sterile, A. Ibáñez & al. 3325 AI (PMA!); P. N. Coiba. Jicarón. Filo SW de la Isla, 200 m, 7 ˚ 15 ’ 34.39 ” N, 81 ˚ 48 ’ 42.00 ” W, 22 August 2005, sterile, A. Ibáñez & al. 4395 AI (PMA!); P. N. Coiba. Manila. 10 m, 7 ˚ 22 ’ 34.45 ” N, 81 ˚ 47 ’ 6.51 ” W, 23 March 1998, A. Ibáñez & al. 689 AI (F, MA); Golfo de Montijo, Isla Cébaco, 31 m, 7 ˚ 30 ’ 16.02 ” N, 81 ˚ 11 ’ 5.00 ” W, 27 April 2012, fr., M. Ayala & al. 1078 MA (PMA!, MO); Golfo de Montijo. Isla Cébaco, 17 m, 7 ˚ 29 ’ 34.44 ” N, 81 ˚ 12 ’ 16.14 ” W, 6 August 2011, sterile, R. Flores & al. 1070 RF (PMA!, MO); Soná. Bahía Honda. Mitad N de la isla Canales de Tierra, 2 m, 7 ˚ 45 ’ 16.13 ” N, 81 ˚ 34 ’ 42.12 ” W, 3 July 2001, sterile, S. Castroviejo & al. 16214 SC (PMA!, MA); Soná. Bahía Honda. Isla María, 7 ˚ 40 ’ 59.95 ” N, 81 ˚ 28 ’ 41.07 ” W, 13 January 2012, sterile, J. Carrión & al. 589 JC (PMA!).	en	Flores, Rodolfo, Ibáñez, Alicia, Correa, Mireya D. (2016): Eugenia veraguensis (Myrtaceae), a new species from Golfo de Chiriquí in Veraguas Province, Panama, with notes on Eugenia rhombea. Phytotaxa 270 (3): 217-222, DOI: 10.11646/phytotaxa.270.3.6, URL: http://dx.doi.org/10.11646/phytotaxa.270.3.6
